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Eschmeyer's Catalog of Fishes
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Species in the family or subfamily Psychrolutidae: [ 367 ] records


acuticeps, Oligocottus Gilbert [C. H.] 1896:432 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Unalaska harbor, Unalaska, Aleutian Islands. Syntypes: (4) CAS-SU 3027 (2), USNM 48608 (1). Additional material: (3) Vancouver I. Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Clinocottus acuticeps (Gilbert 1896) -- (Bolin 1944:84 [ref. 6379], Lee et al. 1980:800 [ref. 22416], Eschmeyer & Herald 1983:165 [ref. 9277], Matarese et al. 1989:422 [ref. 26885], McAllister 1990:196 [ref. 14674], Mecklenburg et al. 2002:502 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:85 [ref. 37547], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:39 [ref. 34694], Kells et al. 2016:166 [ref. 35888], Burton & Lea 2019:65 [ref. 37205], Love et al. 2021:119 [ref. 39279]). Current status: Valid as Clinocottus acuticeps (Gilbert 1896). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: Aleutian Islands south to central California (U.S.A.). Habitat: freshwater, brackish, marine.

aenaeus, Cottus Mitchill [S. L.] 1814:8 [Report, in part, of Samuel L. Mitchill; ref. 3030] New York, U.S.A. No types known. Appeared as Cottus aeneus in Mitchill 1815:380 [ref. 13292]. •Possibly valid as Megalocottus aenaeus (Mitchill 1814) -- (Balakirev et al. 2020:11 [ref. 37873]). •Valid as Myoxocephalus aenaeus (Mitchill 1814) -- (Neelov 1979:123 [ref. 3152], Smith 1986:449 [ref. 22186], Robins & Ray 1986:282 [ref. 23100] as aeneus, Scott & Scott 1988:500 [ref. 25518], McAllister 1990:200 [ref. 14674], Klein-MacPhee in Collette & Klein-MacPhee 2002:349 [ref. 26158], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Afonso et al. 2016:46, 225 [ref. 36199], Simian et al. 2022:375 [ref. 40776]). Current status: Valid as Myoxocephalus aenaeus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Atlantic. Habitat: brackish, marine.

affinis, Centridermichthys Steindachner [F.] & Döderlein [L.] 1887:257 [1] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 53 (1. abth.); ref. 4249] Tango, Japan Sea and Kanagawa, Japan. Syntypes: (4) NMW. •Synonym of Pseudoblennius cottoides (Richardson 1848) -- (Watanabe 1960:152 [ref. 11959], Lindberg & Krasyukova 1987:252 [ref. 15964], Kim & Youn 1992:74 [ref. 22287]). Current status: Synonym of Pseudoblennius cottoides (Richardson 1848). Psychrolutidae: Psychrolutinae. Habitat: marine.

albomaculatus, Crossias Schmidt [P. Yu.] 1916:619, Fig. 4 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 3935] Medny Island, Commander Islands, Bering Sea. Lectotype: ZIN 18966. Paralectotypes: ZIN 18964 (lost), 18965 (1), 19193 (2), ?21866 (?). Lectotype established by Neelov 1979:136 [ref. 3152]. •Synonym of Porocottus bradfordi Rutter 1898, but a valid subspecies -- (Neelov 1979:136 [ref. 3152], Muto et al. 2002:231 [ref. 26331]). •Synonym of Porocottus mednius (Bean 1898) -- (Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:486 [ref. 25968], Parin et al. 2002:S84 [ref. 28300], Parin et al. 2014:269 [ref. 33547]). Current status: Synonym of Porocottus mednius (Bean 1898). Psychrolutidae: Myoxocephalinae. Habitat: marine.

alcicornis, Centridermichthys Herzenstein [S. M.] 1890:115 [25] [Mélanges Biologiques, tirés du Bulletin physico-mathématique de l'Académie Impériale des Sciences de St. Pétersbourg v. 13; ref. 2149] Tokyo, Japan. Holotype (unique): ZIN 8718. On p. 25 in Bull. Acad. Imp. Sci. St. Pétersb. (n. s.) v. 2. •Valid as Alcichthys alcicornis (Herzenstein 1890) -- (Watanabe 1960:131 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Kim & Youn 1992:70 [ref. 22287], Shinohara et al. 1996:175 [ref. 26605], Sokolovskaya et al. 1998:9 [ref. 24670], Choi et al. 2003:262, 573 [ref. 26218]). •Synonym of Alcichthys elongatus (Steindachner 1881) -- (Parin et al. 2002:S68 [ref. 28300], Parin et al. 2014:233 [ref. 33547]). Current status: Synonym of Alcichthys elongatus (Steindachner 1881). Psychrolutidae: Psychrolutinae. Habitat: marine.

aleuticus, Thecopterus Smith [H. M.] 1904:163, Fig. 1 [Proceedings of the Biological Society of Washington v. 17; ref. 4040] 150 miles north of Rat Island, Bering Sea, Albatross station 3785, depth 270 fathoms. Holotype (unique): lost. Footnote on p. 164 mentions the mislaying of the specimen. Mecklenburg et al. (2002:514, 519 [ref. 25968]) give nomenclatural history and updated account listing new material. •Valid as Malacocottus aleuticus (Smith 1904) -- (Sheiko & Fedorov 2000:30 [ref. 25650], Mecklenburg et al. 2002:519 [ref. 25968], Parin et al. 2002:S94 [ref. 28300], Fedorov et al. 2003:86 [ref. 27579], Love et al. 2005:95 [ref. 37547], Maslenikov et al. 2013:12 [ref. 37947], Parin et al. 2014:301 [ref. 33547]). •Synonym of Malacocottus zonurus Bean 1890 -- (Stevenson 2015:24 [ref. 33680]). Current status: Synonym of Malacocottus zonurus Bean 1890. Psychrolutidae: Psychrolutinae. Habitat: marine.

allisi, Crossias Jordan [D. S.] & Starks [E. C.] 1904:296, Fig. 28 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate, Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7711. Paratypes: AMS I.6915 (6); BMNH 1905.2.4.385-391 (7); CAS-SU 7579 (34, now 32); USNM 50922 (19), 51240 (2). Type catalog: Böhlke 1953:128 [ref. 12291]. •Valid as Crossias allisi Jordan & Starks 1904 -- (Watanabe 1960:162 [ref. 11959]). •Valid as Porocottus allisi (Jordan & Starks 1904) -- (Neelov 1979:136 [ref. 3152], Sheiko 1983:18 [ref. 37041], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:223 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:646 [ref. 25086], Mecklenburg et al. 2002:404 [ref. 25968], Nakabo 2002:646 [ref. 26001], Choi et al. 2003:263, 577 [ref. 26218], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S83 [ref. 28300], Parin et al. 2014:268 [ref. 33547], Kim et al. 2020:103 [ref. 39118], Dyldin & Orlov 2022:50 [ref. 38972]). Current status: Valid as Porocottus allisi (Jordan & Starks 1904). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific. Habitat: marine.

amissus, Cyclopterichthys Vaillant [L. L.] 1888:33 [Mission scientifique du Cap Horn, 1882-83 v. 6 (Zool.) (pt 1) (art. C); ref. 4495] Tilly Bay, Strait of Magellan. Holotype (unique): whereabouts unknown. Based on a drawing. •Valid, questionably in cyclopterid genus Cyclopterichthys -- (Lindberg & Legeza 1955:420 [ref. 2785]). •Probably a psychrolutid -- (Gill 1891:366 [ref. 26641], Stein et al. 1991:370 [ref. 19979]). Psychrolutidae: Psychrolutinae. Distribution: Chile. Habitat: marine.

anahaze, Pseudoblennius Bleeker [P.] 1879:11 [Verhandelingen der Koninklijke Akademie van Wetenschappen, Afdeeling Natuurkunde (Amsterdam) v. 18; ref. 460] Unexplained new name for Pseudoblennius pseudoclinus Temminck & Schlegel [see under Pseudoblennius] and Pseudoblennius percoides Günther 1861. •Synonym of Pseudoblennius percoides Günther 1861. Replacement Name. Current status: Synonym of Pseudoblennius percoides Günther 1861. Psychrolutidae: Psychrolutinae. Habitat: marine.

analis, Oligocottus Girard [C. F.] 1858:201 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 9 (no. 15) (for 1857); ref. 1811] Monterey, California, U.S.A. Syntypes: FMNH 210 (1); USNM 486 (orig. 7, now 1). •Valid as Oligocottus analis Girard 1858 -- (Herrera 1896:64 [ref. 39336]). •Valid as Clinocottus analis (Girard 1858) -- (Bolin 1944:73 [ref. 6379], Eschmeyer & Herald 1983:166 [ref. 9277], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:85 [ref. 37547], Page et al. 2013:121 [ref. 32708], Kells et al. 2016:166 [ref. 35888], Burton & Lea 2019:66 [ref. 37205], Love et al. 2021:119 [ref. 39279]). Current status: Valid as Clinocottus analis (Girard 1858). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: northern California (U.S.A.) south to Pacific coast of central Baja California (Mexico). Habitat: marine.

anceps, Cottus (Acanthocottus) Sauvage [H.-E.] 1878:145, Pl. 1 (fig. 13) [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris (Série 2) v. 1; ref. 3880] U.S.A. Syntypes: MNHN 0000-7241 (1), 0000-7243 (1). Type catalog: Blanc & Hureau 1968:50 [ref. 20738]. •Synonym of Myoxocephalus aenaeus (Mitchill 1814) -- (Smith 1986:450 [ref. 22186]). Current status: Synonym of Myoxocephalus aenaeus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

angustus, Neophrynichthys Nelson [J. S.] 1977:491, Figs. 3, 5 [Journal of the Royal Society of New Zealand v. 7 (no. 4); ref. 7034] 13 kilometers east of White Island, New Zealand, near 37°31'S, 177°20'E, depth 549-629 meters. Holotype: NMNZ P.3652. Paratypes: AIM uncat. (6, 2, 2); NMNZ P.3301 (1), P.3649 (1), P.5194 (1), P.5234 (2), P.5719 (1), P.5807 (1), P.5871 (1). Type catalog: Hardy 1990:10 [ref. 18942]. •Valid as Neophrynichthys angustus Nelson 1977 -- (Paulin et al. 1989:172 [ref. 24556], Yabe in Amaoka et al. 1990:246 [ref. 16024]). •Valid as Ambophthalmos angustus (Nelson 1977) -- (Jackson & Nelson 1998:1355 [ref. 23806], Jackson & Nelson 1999:428 [ref. 23799], Jackson & Nelson 2000:725 [ref. 25146], Stewart 2015:1123 [ref. 34254]). Current status: Valid as Ambophthalmos angustus (Nelson 1977). Psychrolutidae: Psychrolutinae. Distribution: New Zealand region. Habitat: marine.

anoplus, Thyriscus Gilbert [C. H.] & Burke [C. V.] 1912:43, Fig. 4 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off Attu Island, Bering Sea, Albatross station 4782, depth 57 fathoms. Holotype (unique): USNM 74368. •Valid as Thyriscus anoplus Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:450 [ref. 25968], Parin et al. 2002:S87 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:274 [ref. 33547], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Thyriscus anoplus Gilbert & Burke 1912. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Kuril Islands north to Commander Islands (Russia) and Aleutian Islands (U.S.A.). Habitat: marine.

antilope, Artediellus Schmidt [P. Yu.] 1937:572 [Comptes Rendus (Doklady) de l'Académie des Sciences de l'U.R.S.S. v. 15 (no. 9); ref. 15302] 55°31'N, 149°10'E, Okhotsk Sea, depth 335 meters. Lectotype: ZIN 24463 (female). Paralectotypes: ZIN 24463 (2 males). Lectotype selected by Neelov 1979:160 [ref. 3152]. Appeared first as name only in Schmidt 1935. •Valid as Artediellina antilope (Schmidt 1937) -- (Neelov 1979:160 [ref. 3152], Sheiko & Fedorov 2000:27 [ref. 25650], Parin et al. 2002:S69 [ref. 28300], Fedorov et al. 2003:65 [ref. 27579], Parin et al. 2014:235 [ref. 33547]). Current status: Valid as Artediellina antilope (Schmidt 1937). Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

aporosus, Artediellus Soldatov [V. K.] 1922:321 [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23; ref. 15430] Tatar Strait, northern Sea of Japan; Cape Bellingshausen and Cape Terpeniya, eastern Sakhalin Island, Okhotsk Sea. Lectotype: ZIN 20613 (1 of 6). Paralectotypes: (15) ZIN 12907 (1), 12909 (3), 20613 (5 of 6). Possibly published in 1921. Lectotype selected by Neelov 1979:154 [ref. 3152]. •Valid as Artediellus aporosus Soldatov 1922 -- (Neelov 1979:154 [ref. 3152], Nelson 1986:43 [ref. 5808], Lindberg & Krasyukova 1987:232 [ref. 15964], Sheiko & Fedorov 2000:27 [ref. 25650], Parin et al. 2002:S69 [ref. 28300], Fedorov et al. 2003:65 [ref. 27579], Inagawa & Yabe 2006:192 [ref. 28854], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:235 [ref. 33547], Dyldin & Orlov 2022:39 [ref. 38972]). Current status: Valid as Artediellus aporosus Soldatov 1922. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Northern Sea of Japan, Okhotsk Sea (Russia and Japan). Habitat: marine.

apus, Bunocottus Kner [R.] 1868:28 [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 58 (nos 1-2); ref. 6074] Bordwood Bank at Cape Horn, South America, depth 45 fathoms. Holotype (unique): Whereabouts unknown. Appeared in more detail in Kner 1868:316 [p. 24 of separate], Pl. 3 (fig. 9) [ref. 2646]. •Tentatively the same as Cottunculus granulosus Karrer 1968 -- (Nelson & Nakamura 1980:990 [ref. 26434]). Current status: Synonym of Cottunculus granulosus Karrer 1968. Psychrolutidae: Psychrolutinae. Habitat: marine.

argenteus, Centridermichthys Döderlein [L.] in Steindachner & Döderlein 1887:257 [1] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 53 (1. abth.); ref. 4249] Tokyo Bay, Japan. Holotype (unique): NMW 76418. •Valid as Pseudoblennius argenteus (Döderlein 1887) -- (Nakabo 2000:649 [ref. 25086], Nakabo 2002:649, 1526 [ref. 26001]). Current status: Valid as Pseudoblennius argenteus (Döderlein 1887). Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

armata, Ochotskia Schmidt [P. Yu.] 1916:612, Fig. 1 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 3935] Near St. Jona Island, 56°18'N, 145°04'E, Okhotsk Sea, depth 175 fathoms. Holotype (unique): ZIN 18968. •Valid as Icelus armatus (Schmidt 1916) -- (Nelson 1984:22 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S76 [ref. 28300], Fedorov et al. 2003:71 [ref. 27579], Parin et al. 2014:253 [ref. 33547], Dyldin & Orlov 2022:45 [ref. 38972]). Current status: Valid as Icelus armatus (Schmidt 1916). Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

asperulus, Artedius Starks [E. C.] 1896:553 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 4195] Port Ludlow, Washington, U.S.A. Syntypes: CAS-SU 5046 (3). Type catalog: Böhlke 1953:127 [ref. 12291]. •Synonym of Artedius fenestralis Jordan & Gilbert 1883 -- (Bolin 1944:48 [ref. 6379]). Current status: Synonym of Artedius fenestralis Jordan & Gilbert 1883. Psychrolutidae: Oligocottinae. Habitat: marine.

asprellus, Radulinus Gilbert [C. H.] 1890:88 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off northwestern U.S.A., depth 43-77 fathoms. Lectotype: USNM 43096. Paralectotypes: BMNH 1896.2.10.24 (1), 1900.7.11.11 (1), CAS-SU 25 (4), ZMB 13940 (1), ZMUO J1479 (1). Type catalog: Böhlke 1953:132 [ref. 12291], Paepke & Fricke 1992:286 [ref. 19946]. Lectotype established in Jordan & Evermann 1898:1921 [ref. 2444]. •Valid as Radulinus asprellus Gilbert 1890 -- (Bolin 1944:40 [ref. 6379], Eschmeyer & Herald 1983:181 [ref. 9277], Matarese et al. 1989:406 [ref. 26885], McAllister 1990:201 [ref. 14674], Yabe & Maruyama 2001:59 [ref. 25245], Mecklenburg et al. 2002:446 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:91 [ref. 37547], Maslenikov et al. 2013:11 [ref. 37947], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:44 [ref. 34694], Kells et al. 2016:188 [ref. 35888], Burton & Lea 2019:68 [ref. 37205], Love et al. 2021:125 [ref. 39279]). Current status: Valid as Radulinus asprellus Gilbert 1890. Psychrolutidae: Radulininae. Distribution: Northeastern Pacific: Bering Sea and Aleutian Islands (Alaska, U.S.A.) south to Pacific coast of northern Baja California (Mexico). Habitat: marine.

asundensis, Cottus quadricornis Lönnberg [E.] 1932:17, Pl. (figs. 7 a-b) [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Åsunden, Sweden. Syntypes: NRM 20329-32 (1, 5, 3, 3). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

atlanticus, Artediellus Jordan [D. S.] & Evermann [B. W.] 1898:1906 [Bulletin of the United States National Museum No. 47; ref. 2444] Massachusetts Bay, U.S.A. Syntypes (?): CAS-SU 448 (3), USNM 21069 (?1). Type catalog: Böhlke 1953:126 [ref. 12291]. Based on literature and apparently one specimen; the described specimen would make the best lectotype if identifiable. •Valid as Artediellus atlanticus Jordan & Evermann 1898 -- (Neelov 1973:594 [ref. 7219], Neelov 1979:153 [ref. 3152], Fedorov in Whitehead et al. 1986:1244 [ref. 13677], Nelson 1986:43 [ref. 5808], Scott & Scott 1988:490 [ref. 25518], McAllister 1990:192 [ref. 14674], Yabe in Okamura et al. 1995:161 [ref. 22531], Klein-MacPhee in Collette & Klein-MacPhee 2002:346 [ref. 26158], Parin et al. 2002:S69 [ref. 28300], Moore et al. 2003:222 [ref. 27076], Nelson et al. 2004:118 [ref. 27807], Møller et al. 2010:51 [ref. 31169], Page et al. 2013:121 [ref. 32709], Mecklenburg et al. 2014:135 [ref. 33463], Parin et al. 2014:236 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Afonso et al. 2016:40, 99 [ref. 36199], Mecklenburg et al. 2016:49 [ref. 34440], Mecklenburg et al. 2018:186 [ref. 36370], Love et al. 2021:117 [ref. 39279], Simian et al. 2022:375 [ref. 40776]). Current status: Valid as Artediellus atlanticus Jordan & Evermann 1898. Psychrolutidae: Psychrolutinae. Distribution: North Atlantic and Arctic Ocean, in North Pacific. Habitat: marine.

auriculatus, Artedielloides Soldatov [V. K.] 1922:353, Fig. (p. 353) [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23; ref. 4160] Peter the Great Bay [n. Japanese Sea], Russia, depth 29-30 meters. Holotype (unique): ZIN 20666. See Nakabo 2002:1526 [ref. 26001]. •Valid as Artedielloides auriculatus Soldatov 1922 -- (Neelov 1979:159 [ref. 3152], Lindberg & Krasyukova 1987:236 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Parin et al. 2002:S69 [ref. 28300], Parin et al. 2014:235 [ref. 33547]). Current status: Valid as Artedielloides auriculatus Soldatov 1922. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

australiae, Ebinania Jackson [K. L.] & Nelson [J. S.] 2006:38, Figs. 1-2 [Records of the Australian Museum v. 58; ref. 28707] Off Cape Sorell, Tasmania, 42°16'S, 144°40'E to 42°22'S, 144°42'E, depth 991-548 meters. Holotype: CSIRO T 412. Paratypes: CSIRO T 502 (1), T 504 (1), T 505 (1), T 536 (1); NMV A1977 (1), A2741 (1), A2758 (1). •Valid as Ebinania australiae Jackson & Nelson 2006 -- (Gomon 2008:523 [ref. 30616]). Current status: Valid as Ebinania australiae Jackson & Nelson 2006. Psychrolutidae: Psychrolutinae. Distribution: Southern Australia: Victoria, Tasmania, South Australia to Western Australia. Habitat: marine.

australis, Clinocottus analis Hubbs [C. L.] 1926:11 [Occasional Papers of the Museum of Zoology University of Michigan No. 171; ref. 2235] Point Loma, San Diego, California, U.S.A. Holotype: UMMZ 55002 [?ex FMNH 9256]. Paratypes: AMNH 5600 (1); ANSP; BMNH 1932.2.16.507-516 (40), FMNH 42048-51 [ex UMMZ] (57, 13, 44, 32); MCZ 36085 [ex USNM 66097] (5); CAS-SU (missing); UMMZ 64210 (now 0), 64417-20 (2, 3, 0, 1), 65495-500 (5, 50, 7, 106, 35, 0), 66001-08 (0, 37, 69, 63, 57, 107, 43, 130), 66097 (201); USNM 83501 (13), 117522 (7). Type catalog: Ibarra & Stewart 1987:27 [ref. 12367]. •Synonym of Clinocottus analis (Girard 1858) -- (Bolin 1944:73 [ref. 6379]). Current status: Synonym of Clinocottus analis (Girard 1858). Psychrolutidae: Oligocottinae. Habitat: marine.

axillaris, Boreocottus Gill [T. N.] 1859:166 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 11; ref. 1760] Awatska Bay, Kamchatka, Russia, Bering Strait. Holotype (unique): ?USNM 24027 (poor condition). •Synonym of Myoxocephalus scorpioides (Fabricius 1780) -- (Neelov 1973:599 [ref. 7219], Neelov 1979:122 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:479 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:264 [ref. 33547], Mecklenburg et al. 2016:83 [ref. 34440]). Current status: Synonym of Myoxocephalus scorpioides (Fabricius 1780). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

balushkini, Bathylutichthys Voskoboinikova [O. S.] 2014:607, Figs. 1-2 [Voprosy Ikhtiologii v. 54 (no. 5); ref. 33563] Meteor Shoal, 48°03'S, 8°14'E, southeastern Atlantic, depth 960-970 meters. Holotype: ZIN 50462. Paratypes: ZIN 50461 (1), 50463 (1). •Valid as Bathylutichthys balushkini Voskoboinikova 2014 -- (Voskoboinilova 2015:251 [ref. 33888]). Current status: Valid as Bathylutichthys balushkini Voskoboinikova 2014. Psychrolutidae: Psychrolutinae. Distribution: Meteor Shoal, southeastern Atlantic. Habitat: marine.

bassargini, Eurymen Lindberg [G. U.] in Soldatov & Lindberg 1930:286, Fig. 49 [Izvestiia Tikhookeanskogo Nauchno-Issledovatel'skogo Instituta Rybnogo Khoziaistva i Okeanografii [Proceedings, Pacific Research Institute of Fisheries and Oceanography, TINRO, Moskva] v. 5; ref. 4164] Basargin, Peter the Great Bay, Russia, Japanese Sea. Holotype (unique): not at ZIN. •Synonym of Eurymen gyrinus Gilbert & Burke 1912 -- (Mecklenburg et al. 2002:517 [ref. 25968], Mecklenburg et al. 2016:100 [ref. 34440] needs investigation). •Valid as Eurymen bassargini Lindberg 1930 -- (Watanabe 1960:190 [ref. 11959], Sheiko 1983:18 [ref. 37041], Lindberg & Krasyukova 1987:267 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Parin et al. 2002:S93 [ref. 28300], Parin et al. 2014:300 [ref. 33547]). Current status: Valid as Eurymen bassargini Lindberg 1930. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

bathybius, Cottus Günther [A.] 1878:180 [13] [Annals and Magazine of Natural History (Series 5) v. 2 (nos 7/8/9)(art. 2/22/28); ref. 2010] Off Misaki, Japanese Seas, Challenger station 235, depth 565 fathoms. Holotype (unique): BMNH 1879.5.14.201. On p. 13 of separate. See also Günther 1887:62, Pl. 10 (fig. C) [ref. 2013]. •Valid as Zesticelus bathybius (Günther 1878) -- (Neelov 1979:161 [ref. 3152], Yabe & Okamura 1993:369 [ref. 22281], Yabe 1995:19 [ref. 22666], Nakabo 2000:640 [ref. 25086], Shinohara et al. 2001:319 [ref. 25995], Nakabo 2002:640 [ref. 26001], Parin et al. 2002:S89 [ref. 28300], Parin et al. 2014:279 [ref. 33547]). Current status: Valid as Zesticelus bathybius (Günther 1878). Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific: Bering Sea to Japan. Habitat: marine.

batrachoides, Myoxocephalus Gilbert [C. H.] & Burke [C. V.] 1912:58, Fig. 11 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Codfish banks west of southern part of Kamchatka, Russia, Albatross station 4798, 51°37'N, 156°21'E, depth 25 fathoms. Holotype (unique): USNM 71809. •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Neelov 1979:122 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

beani, Triglops Gilbert [C. H.] 1896:426, Pl. 28 (upper) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Overall localities -- North and south of Aleutian Islands and in Bristol Bay, Alaska, U.S.A., depth 7.25-42 fathoms. Lectotype: ? CAS-SU 2422. Paralectotypes: BMNH 1900.7.11.10 (1); MCZ 28355, 28356 (1); CAS-SU 2418-19 (6, 5), 2421 (5), 2423-24 (2, 3). Type catalog: Böhlke 1953:132 [ref. 12291]. Lectotype established (as figured specimen from Albatross sta. 3220) in caption to Pl. 288, p. 3291 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable; Pietsch 1994:375, caption to fig. 17 regarded a male from SU 2422 as probably the figured specimen. •Valid as Triglops beani Gilbert 1896 -- (Watanabe 1960:16 [ref. 11959]). •Synonym of Triglops pingelii Reinhardt 1837 -- (Neelov 1973:601 [ref. 7219], Lindberg & Krasyukova 1987:174 [ref. 15964], Pietsch 1994:374 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:428 [ref. 25968], Parin et al. 2002:S88 [ref. 28300], Parin et al. 2014:277 [ref. 33547], Mecklenburg et al. 2016:91 [ref. 34440]). Current status: Synonym of Triglops pingelii Reinhardt 1837. Psychrolutidae: Triglopinae. Habitat: marine.

beauchampi, Liparoides Lloyd [R. E.] 1909:163, Fig. 4 [Memoirs of the Indian Museum v. 2 (no. 3); ref. 2814] Bay of Bengal, Investigator station 372, depth 643 fathoms. Holotype: ZSI F2372/1 (destroyed). Type catalog: Talwar 1977:637 [ref. 26599]. Figured in Lloyd 1909: Pl. 45 (fig. 3) [ref. 20539]. •Not a cyclopterid but probably a psychrolutid -- (Stein 1978:377 [ref. 26640], Nelson 1982:1500 [ref. 5470]). Psychrolutidae: Psychrolutinae. Habitat: marine.

bergii, Taurocottus Soldatov [V. K.] & Pavlenko [M. N.] 1915:149-151, Pl. 4 (figs. 1-2) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 4165] 47°55'00"N, 139°39'30"E, Primorye, Russia, Sea of Japan, depth 66 meters. Lectotype: ZIN 18637. Paralectotypes: (7) ZIN 18636-37 (2, 1). Lectotype selected by Neelov 1979:143 [ref. 3152]. •Valid as Taurocottus bergii Soldatov & Pavlenko 1915 -- (Neelov 1979:143 [ref. 3152], Yabe in Masuda et al. 1984:326 [ref. 6441] as bergi, Lindberg & Krasyukova 1987:227 [ref. 15964] as bergi, Kim & Youn 1992:70 [ref. 22287], Sokolovskaya et al. 1998:10 [ref. 24670] as bergi, Nakabo 2000:644 [ref. 25086] as bergi, Mecklenburg et al. 2002:474 [ref. 25968], Nakabo 2002:644 [ref. 26001] as bergi, Choi et al. 2003:263, 578 [ref. 26218], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:273 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Kim et al. 2020:103 [ref. 39118], Dyldin & Orlov 2022:52 [ref. 38972]). Current status: Valid as Taurocottus bergii Soldatov & Pavlenko 1915. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Korea, Russia, Japan. Habitat: marine.

beringi, Crossias Soldatov [V. K.] 1916:218, Pl. 13 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 21; ref. 18735] Abrek Bay and Ayan Bay, Bol. Shantar Island, Okhotsk Sea; also Olga Bay, Sea of Japan. Syntypes: (many) ZIN 19066-67 (3, 4). Two new forms described on p. 221. •Synonym of Porocottus tentaculatus (Kner 1868) -- (Lindberg & Krasyukova 1987:224 [ref. 15964], Parin et al. 2014:270 [ref. 33547] in part). •Synonym of Porocottus minutus (Pallas 1814) or Porocottus tentaculatus (Kner 1868) -- (Neelov 1979:137 [ref. 3152], Parin et al. 2002:S84 [ref. 28300], Parin et al. 2014:269 [ref. 33547]). Current status: Synonym of Porocottus minutus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

beringianum, Stelgistrum Gilbert [C. H.] & Burke [C. V.] 1912:52, Fig. 9 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Petrel Bank, Aleutian Islands, Albatross station 4777, depth 43-52 fathoms. Holotype: USNM 74373. Paratypes: CAS-SU 22245 (1). Type catalog: Böhlke 1953:132 [ref. 12291]. •Valid as Stelgistrum beringianum Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:449 [ref. 25968], Parin et al. 2002:S86 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:272 [ref. 33547], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Stelgistrum beringianum Gilbert & Burke 1912. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: southern Bering Sea, Commander Islands (Russia), and Aleutian Islands (U.S.A.). Habitat: marine.

beringianus, Artediellus scaber Schmidt [P. Yu.] 1927:8 [Proceedings of the United States National Museum v. 71 (no. 2685); ref. 15822] Tkatchen Bay, 64°25'N, 172°48'03"W, northern Bering Sea. Holotype: ZIN 21492 (1 of 2 adults). Paratypes: ZIN 21492 (1 of 2 adults). •Synonym of Artediellus scaber Knipowitsch 1907 -- (Neelov 1979:154 [ref. 3152], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:491 [ref. 25968], Parin et al. 2002:S71 [ref. 28300], Parin et al. 2014:239 [ref. 33547], Mecklenburg et al. 2016:52 [ref. 34440]). Current status: Synonym of Artediellus scaber Knipowitsch 1907. Psychrolutidae: Psychrolutinae. Habitat: brackish, marine.

beringianus, Icelus bicornis Schmidt [P. Yu.] 1927:7 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 19815] Providence Bay, Bering Strait, 63°27' to 64°25'N, 168°48' to 176°27'E, depth 8-40 fathoms. Syntypes: ZIN 21914 (1). •Synonym of Icelus spatula Gilbert & Burke 1912 -- (Neelov 1973:597 [ref. 7219], Nelson 1984:36 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:256 [ref. 33547]). Current status: Synonym of Icelus spatula Gilbert & Burke 1912. Psychrolutidae: Icelinae. Habitat: marine.

bicornis, Cottus Reinhardt [J. C. H.] 1840:9 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) v. 7 (1839); ref. 18617] East Greenland. No types known. Type catalog: Nielsen 1974:81 [ref. 9588]. Published two more times in 1840-41. Cottus bicornis Anonymous (Volta) 1796:clxiv, Pl. 39 (fig. 4) [ref. 30407] from the 'monti del Veronese' (mountains of Province Verona, Italy) is an homonym in fossil fishes; Agassiz 1835:293 [ref. 30408] noted that the original material is lost and the illustration in Volta 1796 is unidentifiable; the name Cottus bicornis Anonymous (Volta) 1796 was not used as valid since 1899, therefore reversal of precedence according to ICZN Art. 23.2 and 23.9 in favor of Cottus bicornis Reinhardt 1840. •Valid as Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:35 [ref. 5391], Fedorov in Whitehead et al. 1986:1250 [ref. 13677], Robins & Ray 1986:282 [ref. 23100], Scott & Scott 1988:498 [ref. 25518], McAllister 1990:198 [ref. 14674], Yabe in Okamura et al. 1995:164 [ref. 22531], Mecklenburg et al. 2002:456 [ref. 25968], Parin et al. 2002:S77 [ref. 28300], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Dolgov 2013:918 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:253 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:43, 173 [ref. 36199], Mecklenburg et al. 2016:67 [ref. 34440], Mecklenburg et al. 2018:205 [ref. 36370], Love et al. 2021:122 [ref. 39279], Zorina & Chernova. 2022:89 [ref. 40474]). 1°Homonym. Current status: Valid as Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Distribution: Circumpolar in Arctic Ocean, North Atlantic and North Pacific. Habitat: marine.

biseriatus, Archaulus Gilbert [C. H.] & Burke [C. V.] 1912:36, Fig. 1 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Petrel Bank, Bering Sea, Albatross station 4778, depth 43 fathoms. Holotype: USNM 74365. Paratypes: CAS-SU 22238 (4); USNM 70879 (1), 74503-04 (3, 3). Type catalog: Böhlke 1953:126 [ref. 12291]. •Valid as Archaulus biseriatus Gilbert & Burke 1912 -- (Orlov et al. 2001:12 [ref. 40755], Parin et al. 2002:S68 [ref. 28300]). •Valid as Archistes biseriatus (Gilbert & Burke 1912) -- (Sheiko & Fedorov 2000:26 [ref. 25650], Mecklenburg et al. 2002:443 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:84 [ref. 37547], Maslenikov et al. 2013:10 [ref. 37947], Page et al. 2013:121 [ref. 32708], Parin et al. 2014:233 [ref. 33547], Kells et al. 2016:162 [ref. 35888], Love et al. 2021:117 [ref. 39279]). Current status: Valid as Archistes biseriatus (Gilbert & Burke 1912). Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Kuril Islands; southern Bering Sea off Aleutian Islands (U.S.A.). Habitat: marine.

bison, Aspicottus Girard [C. F.] 1854:130 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] San Francisco, California, U.S.A. No types known. •Valid as Aspicottus bison Girard 1854 -- (Neelov 1979:148 [ref. 3152]). •Valid as Enophrys bison (Girard 1854) -- (Bolin 1944:91 [ref. 6379], Sandercock & Wilimovsky 1968:844 [ref. 26414], Eschmeyer & Herald 1983:168 [ref. 9277], Matarese et al. 1989:390 [ref. 26885], McAllister 1990:197 [ref. 14674], Mecklenburg et al. 2002:471 [ref. 25968], Nelson et al. 2004:119 [ref. 27807], Love et al. 2005:86 [ref. 37547], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:40 [ref. 34694], Kells et al. 2016:170 [ref. 35888], Burton & Lea 2019:66 [ref. 37205], Love et al. 2021:120 [ref. 39279]). Current status: Valid as Enophrys bison (Girard 1854). Psychrolutidae: Myoxocephalinae. Distribution: Northeastern Pacific: western Gulf of Alaska (U.S.A.) south to central California (U.S.A.). Habitat: marine.

bispinis, Icelus spatula Andriashev [A. P.] 1937:270, Pl. 6 (fig. 5) [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] Penzhina Bay to Chanjanga; northeastern Sachalin Island; and northwestern Kamchatka Peninsula, Okhotsk Sea, depth 55-150 meters and to 240 meters. Syntypes: ZIN 17787 (1), 17790-91 (1, 3), 18060 (1), 21920-23 (5, 6, 6, 2). •Synonym of Icelus bicornis (Reinhardt 1840) -- (Nelson 1984:42 [ref. 5391]). •Synonym of Icelus spatula Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:256 [ref. 33547]). Current status: Synonym of Icelus spatula Gilbert & Burke 1912. Psychrolutidae: Icelinae. Habitat: marine.

boleoides, Radulinus Gilbert [C. H.] in Jordan & Evermann 1898:1919 [Bulletin of the United States National Museum No. 47; ref. 2444] Off Santa Catalina Island, California, U.S.A., Albatross station 3664, depth 59 fathoms. Holotype (unique): USNM 48795 [also = 48879]. Author is C. H. Gilbert. Also appeared as new in Gilbert 1899:26, Pl. 1 [ref. 21322]. •Valid as Radulinus boleoides Gilbert 1898 -- (Bolin 1944:38 [ref. 6379], Eschmeyer & Herald 1983:181 [ref. 9277], Matarese et al. 1989:408 [ref. 26885], McAllister 1990:201 [ref. 14674], Yabe & Maruyama 2001:59 [ref. 25245], Mecklenburg et al. 2002:445 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:91 [ref. 37547], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:44 [ref. 34694], Kells et al. 2016:188 [ref. 35888], Burton & Lea 2019:68 [ref. 37205], Love et al. 2021:125 [ref. 39279]). Current status: Valid as Radulinus boleoides Gilbert 1898. Psychrolutidae: Radulininae. Distribution: Northeastern Pacific: Kodiak Island (Alaska, U.S.A.) south to southern California (U.S.A.). Habitat: marine.

borealis, Crossias beringi forma Soldatov [V. K.] 1916:220, Pl. 13 (upper) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 21; ref. 18735] Okhotsk Sea. Syntypes: ZIN 19066 (1). Treated as infrasubspecific in Eschmeyer et al. 1998; now regarded as subspecific. Muto et al. 2002:231 [ref. 26331] list one syntype. •Synonym of Porocottus minutus (Pallas 1814) -- (Neelov 1979:137 [ref. 3152], Parin et al. 2002:S84 [ref. 28300]). Current status: Synonym of Porocottus minutus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

borealis, Icelinus Gilbert [C. H.] 1896:415, Pl. 25 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] North and south of Aleutians and in Bristol Bay, Alaska, U.S.A. Lectotype: not determined. Paralectotypes (incl. potential lectotype): MCZ 28373 (6); MNHN 1896-0491 [ex USNM] (4); CAS-SU 2291-92 (6, 34); USNM 53037 (6); ZMUO J1454 (3, poor condition) station 3265, ZMUO J6331 (3) station 3213. Type catalog: Böhlke 1953:129 [ref. 12291], Blanc & Hureau 1968:53 [ref. 20738], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen) in caption to Pl. 284, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. •Valid as Icelinus borealis Gilbert 1896 -- (Eschmeyer & Herald 1983:172 [ref. 9277], Peden 1984:79 [ref. 5193], McAllister 1990:198 [ref. 14674], Mecklenburg et al. 2002:441 [ref. 25968], Parin et al. 2002:S76 [ref. 28300], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:252 [ref. 33547], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:174 [ref. 35888], Love et al. 2021:121 [ref. 39279]). Current status: Valid as Icelinus borealis Gilbert 1896. Psychrolutidae: Oligocottinae. Distribution: North Pacific: Bristol Bay and Aleutian Islands (U.S.A.) south to Puget Sound, Washington (U.S.A.). Habitat: marine.

borealis, Oligocottus Jordan [D. S.] & Snyder [J. O.] in Jordan 1896:225 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 2395] Neah Bay, Puget Sound, Washington, U.S.A. Syntypes: ANSP 39141 (1, missing); BMNH 1897.1.12.17-18 (2); CAS-SU 2985 (4), 3396 (27). Type catalog: Böhlke 1953:131 [ref. 12291], Böhlke 1984:65 [ref. 13621]. •Synonym of Oligocottus maculosus Girard 1856 -- (Bolin 1944:65 [ref. 6379], Mecklenburg et al. 2002:500 [ref. 25968]). Current status: Synonym of Oligocottus maculosus Girard 1856. Psychrolutidae: Oligocottinae. Habitat: marine.

borkensis, Cottus quadricornis Lönnberg [E.] 1939:4 [Arkiv för Zoologi v. 31 B (no. 2); ref. 17199] Lake Borken [Bork], parishes of Yxnerum and Björsäter, Osthrogothland Province, Sweden. Syntypes: NRM 20327 (2). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

bradfordi, Porocottus Rutter [C. L.] in Jordan & Evermann 1898:2862 [Bulletin of the United States National Museum No. 47; ref. 2445] Rock pools at Karluk, Kodiak Island, Alaska, U.S.A. Syntypes: BMNH 1900.9.29.209-210 (2), CAS-SU 6063 (57), USNM 60328 (1), USFC. Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Porocottus bradfordi Rutter 1898 -- (Neelov 1979:136 [ref. 3152], Eschmeyer & Herald 1983:179 [ref. 9277], Muto et al. 2002:231 [ref. 26331]). •Synonym of Porocottus mednius (Bean 1898) -- (Mecklenburg et al. 2002:404 [ref. 25968]). Current status: Synonym of Porocottus mednius (Bean 1898). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific: Bering Strait to Commander-Aleutian chain to southeastern Alaska and to Kuril Islands. Habitat: marine.

brandtii, Cottus Steindachner [F.] 1867:119 [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Classe v. 4 (no. 14) (for 16 May 1867); ref. 21724] Mouth of Amur River, Russia. Holotype (unique): NMW 11676. Species also appeared on p. 79 of English version [ref. 18764] as brandtii; later illustrated and described in more detail in Steindachner 1867:706, Pl. 3 (figs. 1-2) [p. 6 of separate] [ref. 6393] as brandti in text and brandtii on plate. •Valid as Myoxocephalus brandtii (Steindachner 1867) -- (Watanabe 1960:73 [ref. 11959], Neelov 1979:121 [ref. 3152] as brandti, Yabe in Masuda et al. 1984:326 [ref. 6441] as brandti, Lindberg & Krasyukova 1987:210 [ref. 15964] as brandti, Sokolovsky & Sokolovskaya 1997:60 [ref. 24691] as brandti, Sokolovskaya et al. 1998:10 [ref. 24670] as brandti, Nakabo 2000:645 [ref. 25086] as brandti, Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:404 [ref. 25968], Nakabo 2002:645 [ref. 26001] as brandti, Parin et al. 2002:S80 [ref. 28300], Fedorov et al. 2003:75 [ref. 27579], Parin et al. 2014:260 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138], Prokofiev 2017:648 [ref. 35721], Dyldin et al. 2021:66 [ref. 38804], Dyldin & Orlov 2022:48 [ref. 38972]). Current status: Valid as Myoxocephalus brandtii (Steindachner 1867). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific. Habitat: brackish, marine.

brashnikovi, Trichocottus Soldatov [V. K.] & Pavlenko [M. N.] 1915:151, Pl. 4 (fig. 3) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 4165] 50°04'50"N, 140°47'E, Tatar Strait, Primorye, Russia, Sea of Japan. Lectotype: ZIN 18640. Paralectotypes: ZIN 18639 (1). Lectotype selected by Neelov 1979:143 [ref. 3152]. •Valid as Trichocottus brashnikovi Soldatov & Pavlenko 1915 -- (Neelov 1979:143 [ref. 3152] as brashnicovi, Lindberg & Krasyukova 1987:228 [ref. 15964] as brashnicovi, Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:474 [ref. 25968], Parin et al. 2002:S87 [ref. 28300], Fedorov et al. 2003:81 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Mecklenburg et al. 2011:123 [ref. 31212], Maslenikov et al. 2013:11 [ref. 37947], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:274 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Mecklenburg et al. 2016:86 [ref. 34440], Mecklenburg et al. 2018:223 [ref. 36370], Love et al. 2021:126 [ref. 39279], Dyldin & Orlov 2022:52 [ref. 38972]). Current status: Valid as Trichocottus brashnikovi Soldatov & Pavlenko 1915. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific and Arctic: Sea of Japan and sea of Okhotsk north to northern Bering Sea and northeastern Chukchi Sea, then east to Alaska (U.S.A.). Habitat: marine.

brephocephalus, Cottunculus Jordan [D. S.] & Starks [E. C.] 1903:689, Fig. [Proceedings of the United States National Museum v. 26 (no. 1326); ref. 15033] Suruga Bay, Japan, Albatross station 3704, depth 94 fathoms. Holotype (unique): USNM 50591. •Valid as Cottunculus brephocephalus Jordan & Starks 1903 -- (Watanabe 1960:195 [ref. 11959], Nelson 1977:508 [ref. 7034]). •Valid as Ebinania brephocephala (Jordan & Starks 1903) -- (Yabe in Amaoka et al. 1983:157, 214 [ref. 8379], Yabe in Masuda et al. 1984:330 [ref. 6441], Nakabo 2000:652 [ref. 25086], Nakabo 2002:652 [ref. 26001], Shinohara et al. 2005:428 [ref. 28370], Jackson & Nelson 2006:42 [ref. 28707], Meguro & Motomura 2011:27 [ref. 32399], Motomura et al. 2015:19 [ref. 34709]). Current status: Valid as Ebinania brephocephala (Jordan & Starks 1903). Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

bryosus, Blennicottus globiceps Jordan [D. S.] & Starks [E. C.] 1895:808 [Proceedings of the California Academy of Sciences (Series 2) v. 5; ref. 2522] Near Neah Bay, Washington, U.S.A. Syntypes: NMW 6549 (1), CAS-SU 3404 (6). Type catalog: Böhlke 1953:127 [ref. 12291]. •Synonym of Clinocottus globiceps (Girard 1858) -- (Bolin 1944:81 [ref. 6379]). Current status: Synonym of Clinocottus globiceps (Girard 1858). Psychrolutidae: Oligocottinae. Habitat: marine.

bubalis, Cottus Euphrasen [B. A.] 1786:65, Pl. 3 (figs. 2-3) [Kongliga Vetenskaps-Academiens Handlingar, Stockholm v. 7 (for 1786); ref. 17465] Bohuslän, Sweden. No types known. Also described by Euphrasen 1787:64, pl. 3, fig. 2-3 [ref. 30157]. •Valid as Enophrys bubalis (Euphrasen 1786) -- (Sandercock & Wilimovsky 1968:841 [ref. 26414]). •Valid as Taurulus bubalis (Euphrasen 1786) -- (Neelov 1973:600 [ref. 7219], Neelov 1979:149 [ref. 3152], Fedorov in Whitehead et al. 1986:1256 [ref. 13677], Parin et al. 2002:S86 [ref. 28300], Fricke 2007:29 [ref. 30577], Parin et al. 2014:273 [ref. 33547], Carneiro et al. 2019:237 [ref. 37250], Bañón & Maño 2021:69 [ref. 38883], Kovačić et al. 2021:68 [ref. 39014]). Current status: Valid as Taurulus bubalis (Euphrasen 1786). Psychrolutidae: Myoxocephalinae. Distribution: Baltic Sea; North Sea; northwestern Mediterranean Sea; northeastern Atlantic: Iceland and Norway south to Gibraltar. Habitat: brackish, marine.

burchami, Icelinus Evermann [B. W.] & Goldsborough [E. L.] 1907:297, Fig. 48 [Bulletin of the Bureau of Fisheries v. 26 (Doc. 624) (for 1906); ref. 6532] Off Loring, Alaska, U.S.A., Albatross station 4228, depth 41-134 fathoms. Holotype: USNM 57822. Paratypes: ANSP 33003 (1, disintegrated); FMNH 6117 (not found); CAS-SU 22614 [not 20010 as published] (1); USNM 57825 (2), 124970 [ex USBF 1637 = 5229]. Type catalog: Böhlke 1953:129 [ref. 12291], Böhlke 1984:115 [ref. 13621]. •Valid as Icelinus burchami Evermann & Goldsborough 1907 -- (Bolin 1944:30 [ref. 6379], Eschmeyer & Herald 1983:172 [ref. 9277], Peden 1984:79 [ref. 5193], McAllister 1990:198 [ref. 14674], Mecklenburg et al. 2002:440 [ref. 25968], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Maslenikov et al. 2013:10 [ref. 37947], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:176 [ref. 35888], Burton & Lea 2019:66 [ref. 37205], Love et al. 2021:121 [ref. 39279]). Current status: Valid as Icelinus burchami Evermann & Goldsborough 1907. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: southeastern Alaska south to southern California (U.S.A.). Habitat: marine.

camchaticus, Artediellus Gilbert [C. H.] & Burke [C. V.] 1912:46, Fig. 6 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off eastern coast of Kamchatka, Russia, 52°46'50"N, Albatross station 4795, depth 48-69 fathoms. Holotype: USNM 74370. Paratypes: CAS-SU 22384 (7); USNM 70927 (10), 74482-83 (27, 23). Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Artediellus camchaticus Gilbert & Burke 1912 -- (Neelov 1979:154 [ref. 3152], Yabe in Masuda et al. 1984:328 [ref. 6441], Nelson 1986:43 [ref. 5808], Nakabo 2000:641 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:493 [ref. 25968], Nakabo 2002:641 [ref. 26001], Fedorov et al. 2003:65 [ref. 27579], Parin et al. 2002:S70 [ref. 28300], Love et al. 2005:84 [ref. 37547], Mecklenburg et al. 2011:122 [ref. 31212], Maslenikov et al. 2013:10 [ref. 37947], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:236 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Love et al. 2021:118 [ref. 39279], Dyldin & Orlov 2022:39 [ref. 38972]). Current status: Valid as Artediellus camchaticus Gilbert & Burke 1912. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Sea of Okhotsk and Kamchatka to Kuril Islands, east to western Aleutian Islands (U.S.A.). Habitat: marine.

camtschaticus, Crossias Schmidt [P. Yu.] 1916:617, Fig. 3 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 3935] Mouth of Osernaya River, western Kamchatka, Russia. Lectotype: ZIN 18963. Paralectotypes: ZIN 18962 (now 7). Also spelled kamtschaticus by authors. Lectotype selected by Neelov 1979:137 [ref. 3152]. •Valid as Porocottus camtschaticus (Schmidt 1916) -- (Neelov 1979:137 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:485 [ref. 25968], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S83 [ref. 28300], Fedorov et al. 2003:79 [ref. 27579], Parin et al. 2014:268 [ref. 33547], Dyldin et al. 2020:114 [ref. 37569]). Current status: Valid as Porocottus camtschaticus (Schmidt 1916). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific: northern Kuril Islands and southern Kamchatka (Russia). Habitat: brackish, marine.

canaliculatus, Icelus Gilbert [C. H.] 1896:412, Pl. 24 (lower) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] North of Unalaska, Albatross station 3329, depth 399 fathoms. Lectotype: USNM or CAS-SU 2182. Paralectotypes (incl. potential lectotype): BMNH 1896.7.23.80 (1); MCZ 28346 (1); CAS-SU 2182 (1, poor condition), 18828 (3); USNM 48590 (1), 126722 (1); ZMUO J1459 (1). Type catalog: Böhlke 1953:130 [ref. 12291], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen from Albatross sta. 3329) in caption to Pl. 286, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable; SU 2182 designated lectotype by Nelson 1984:32 [ref. 5391]. •Valid as Icelus canaliculatus Gilbert 1896 -- (Yabe in Amaoka et al. 1983:281, 337 [ref. 8379], Nelson 1984:32 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:634 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:452 [ref. 25968], Nakabo 2002:634 [ref. 26001], Parin et al. 2002:S77 [ref. 28300], Fedorov et al. 2003:71 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:253 [ref. 33547], Kells et al. 2016:178 [ref. 35888], Dyldin & Orlov 2022:45 [ref. 38972] with question, Love et al. 2021:122 [ref. 39279]). Current status: Valid as Icelus canaliculatus Gilbert 1896. Psychrolutidae: Icelinae. Distribution: North Pacific: northern Hokkaido (Japan) and Sea of Okhotsk north to northern Bering Sea, east to western Aleutian islands (U.S.A.). Habitat: marine.

cataphractus, Agonocottus Pavlenko [M. N.] 1910:23, Figs. 2-3 [Trudy Obshchestva Estestvoispytatelej Imperatorskom' Kazanskom' Universiteti, Kazan v. 42 (no. 2); ref. 3393] Near Cape Povorotnyi, Peter the Great Bay, Russia, Sea of Japan, depth 812-875 feet (= 116-125 Russian fathoms). Holotype (unique): not researched. •Synonym of Icelus spiniger Gilbert 1896, but a valid supspecies -- (Lindberg & Krasyukova 1987:190 [ref. 15964]). •Valid as Icelus cataphractus (Pavlenko 1910) -- (Nelson 1984:28 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Kim & Youn 1992:66 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:633 [ref. 25086], Nakabo 2002:633 [ref. 26001], Choi et al. 2003:262, 575 [ref. 26218], Parin et al. 2002:S77 [ref. 28300], Shinohara et al. 2011:43 [ref. 31715], Song et al. 2012:305 [ref. 36295], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:254 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Kim et al. 2020:103 [ref. 39118], Dyldin et al. 2021:64 [ref. 38804], Dyldin & Orlov 2022:45 [ref. 38972]). Current status: Valid as Icelus cataphractus (Pavlenko 1910). Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

caulias, Sigmistes Rutter [C. L.] in Jordan & Evermann 1898:2863 [Bulletin of the United States National Museum No. 47; ref. 2445] Rock pools at Karluk, Kodiak Island, Alaska, U.S.A. Holotype: CAS-SU 5884 (1 of 4). Paratypes: (5) CAS-SU 5884 (3 of 4), USNM 106612 [ex CAS-SU 5884] (1). Type catalog: Böhlke 1953:132 [ref. 12291]. •Valid as Sigmistes caulias Rutter 1898 -- (Yabe et al. 2001:70 [ref. 25246], Mecklenburg et al. 2002:489 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:272 [ref. 33547], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Sigmistes caulias Rutter 1898. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Pribilof Islands and Aleutian Islands (U.S.A.) to western Gulf of Alaska (U.S.A.). Habitat: marine.

cavifrons, Icelinus Gilbert [C. H.] 1890:83 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., Albatross station 2907 and 2945, depth 30-44 fathoms. Syntypes: CAS-SU 69 (1), 148 (1), 28111 (1); USNM 44405 (2), 48247 (1). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelinus cavifrons Gilbert 1890 -- (Bolin 1944:28 [ref. 6379], Eschmeyer & Herald 1983:172 [ref. 9277], Peden 1984:79 [ref. 5193], Castro-Aguirre et al. 1993:92 [ref. 21807], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:176 [ref. 35888], Burton & Lea 2019:66 [ref. 37205], Love et al. 2021:121 [ref. 39279]). Current status: Valid as Icelinus cavifrons Gilbert 1890. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: central California (U.S.A.) south to Pacific coast of southern Baja California Sur (Mexico). Habitat: marine.

centropomus, Podabrus Richardson [J.] 1848:11, Pl. 1 (figs. 7-11) [The zoology of the voyage of H. M. S. Samarang; ref. 3744] Quelpart, Korea Straits. Holotype (unique): BMNH 1982.6.10.1. •Valid as Vellitor centropomus (Richardson 1848) -- (Watanabe 1960:165 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Kim & Youn 1992:66 [ref. 22287], Nakabo 2000:647 [ref. 25086], Shinohara et al. 2000:177 [ref. 25987], Nakabo 2002:647 [ref. 26001], Choi et al. 2003:263, 579 [ref. 26218], Kwun et al. 2017:141 [ref. 37287], Kim et al. 2020:105 [ref. 39118], Sonoyama et al. 2020:91 [ref. 37637]). Current status: Valid as Vellitor centropomus (Richardson 1848). Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: southern Korea and Japan. Habitat: marine.

cervus, Synanceja Tilesius [W. G. von] 1811:278, Pl. 13 [Mémoires de l'Académie Impériale des Sciences de Sankt Pétersbourg (5e Série) v. 3; ref. 4408] Kamchatka, Russia. No types known. Syntypes: MNHN 0000-7249 (1). Type information: Svetovidov 1978:27 [ref. 6025]. •Synonym of Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414], Neelov 1979:146 [ref. 3152], Parin et al. 2002:S73 [ref. 28300], Shinohara et al. 2014:249 [ref. 33330]). Current status: Synonym of Enophrys diceraus (Pallas 1787). Psychrolutidae: Myoxocephalinae. Habitat: marine.

clarki, Blennicottus Evermann [B. W.] & Goldsborough [E. L.] 1907:323, Fig. 85 [Bulletin of the Bureau of Fisheries v. 26 (Doc. 624) (for 1906); ref. 6532] Bering Sea, Albatross station 3598. Holotype: USNM 57824. Paratypes: USNM 124987 (1). •Synonym of Artediellus pacificus Gilbert 1896 -- (Neelov 1979:154 [ref. 3152], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:492 [ref. 25968], Parin et al. 2002:S71 [ref. 28300], Parin et al. 2014:238 [ref. 33547]). Current status: Synonym of Artediellus pacificus Gilbert 1896. Psychrolutidae: Psychrolutinae. Habitat: marine.

claviger, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:195, Pl. 79 (fig. 2) [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka, Russia. Holotype (unique): BMNH 2011.5.17.1. Type catalog: Blanc & Hureau 1968:60 [ref. 20738]. •Synonym of Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414], Neelov 1979:146 [ref. 3152], Lindberg & Krasyukova 1987:229 [ref. 15964], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:472 [ref. 25968], Parin et al. 2002:S74 [ref. 28300], Parin et al. 2014:245 [ref. 33547]). Current status: Synonym of Enophrys diceraus (Pallas 1787). Psychrolutidae: Myoxocephalinae. Habitat: marine.

concinnum, Stelgistrum Andriashev [A. P.] 1935:292, Fig. 1 [Zoologischer Anzeiger v. 111 (nos 11/12); ref. 13868] Cape Olutorsky, Bering Sea, depth 32 meters. Holotype: ZIN 24934. Paratypes: ZIN 21456 (2). •Valid as Stelgistrum concinnum Andriashev 1935 -- (Mecklenburg et al. 2002:448 [ref. 25968], Parin et al. 2002:S86 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:272 [ref. 33547], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Stelgistrum concinnum Andriashev 1935. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: southern Bering Sea and Aleutian Islands (U.S.A.). Habitat: marine.

corallinus, Allartedius Hubbs [C. L.] 1926:8 [Occasional Papers of the Museum of Zoology University of Michigan No. 170; ref. 6069] California, U.S.A. Holotype: UMMZ 141868. •Valid as Artedius corallinus (Hubbs 1926) -- (Bolin 1944:53 [ref. 6379], Eschmeyer & Herald 1983:161 [ref. 9277], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:84 [ref. 37547], Pietsch & Orr 2015:72 [ref. 34694], Kells et al. 2016:162 [ref. 35888], Burton & Lea 2019:65 [ref. 37205], Love et al. 2021:118 [ref. 39279]). Current status: Valid as Artedius corallinus (Hubbs 1926). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: Washington (U.S.A.) south to Pacific coast of northern Baja California (Mexico). Habitat: marine.

corniger, Artediellus europeus Andriashev [A. P.] 1954:407, Fig. 228 [ ref. 6547] Northern parts of Barents [not Bering], Kara, and Laptev seas. Syntypes: ZIN 32202 (?). •Synonym of Artediellus atlanticus, but a valid subspecies -- (Parin et al. 2002:S70 [ref. 28300]). •Synonym of Artediellus atlanticus Jordan & Evermann 1898 -- (Neelov 1973:594 [ref. 7219], Neelov 1979:153 [ref. 3152], Parin et al. 2014:236 [ref. 33547]). Current status: Synonym of Artediellus atlanticus Jordan & Evermann 1898. Psychrolutidae: Psychrolutinae. Habitat: marine.

coronatus, Porocottus Yabe [M.] 1992:361, Figs. 1-3 [Japanese Journal of Ichthyology v. 38 (no. 4); ref. 19519] Near Konbumori, Pacific coast of Hokkaido, 42°56.5'N, 144°38.5'E, depth 0.5 meters. Holotype: HUMZ 111575. Paratypes: HUMZ 109999-110001 (3), 117471 (1); NSMT-P 35034 (1). •Valid as Porocottus coronatus Yabe 1992 -- (Nakabo 2000:646 [ref. 25086], Nakabo 2002:646 [ref. 26001], Muto et al. 2002:231 [ref. 26331]). Current status: Valid as Porocottus coronatus Yabe 1992. Psychrolutidae: Myoxocephalinae. Distribution: Western North Pacific. Habitat: marine.

costaecanariae, Cottunculus Cervigón [F.] 1961:125, Figs. 5-7 [Investigación Pesquera (Chile) v. 19; ref. 777] Mauritania, western Africa, 20°45'N, depth 450 meters. Holotype (unique): IIPC 33. Originally as costae-canariae, correction mandatory. •Valid as Ebinania costaecanariae (Cervigón 1961) -- (Nelson 1986:491 [ref. 5698] as costaecanarie, Nelson 1990:687 [ref. 17821] as costacanarie, Quéro & Cendrero 2001:177 [ref. 25567], Jackson & Nelson 2006:42 [ref. 28707], Prokofiev & Kukuev 2009:225 [ref. 30172], Nelson 2016:2292 [ref. 34615], Reiner 2019:164 [ref. 38139] as costacanarie, Fermon et al. 2022:296 [ref. 39448]). Current status: Valid as Ebinania costaecanariae (Cervigón 1961). Psychrolutidae: Psychrolutinae. Distribution: Eastern Atlantic: Morocco south to South Africa, including São Tomé and Principe. Habitat: marine.

cottoides, Podabrus Richardson [J.] 1848:13, Pl. 1 (figs. 1-6) [The zoology of the voyage of H. M. S. Samarang; ref. 3744] China Sea. Syntypes: BMNH 1848.3.16.102 (1), 1855.9.19.399 (1). •Valid as Pseudoblennius cottoides (Richardson 1848) -- (Watanabe 1960:150 [ref. 11959], Nakabo in Masuda et al. 1984:330 [ref. 6441], Lindberg & Krasyukova 1987:252 [ref. 15964], Kim & Youn 1992:74 [ref. 22287], Mizuno et al. 1999:7 [ref. 25311], Nakabo 2000:650 [ref. 25086], Shimizu 2001:28 [ref. 25799], Nakabo 2002:650 [ref. 26001], Choi et al. 2003:263, 577 [ref. 26218], Kwun et al. 2017:141 [ref. 37287], Kim et al. 2020:103 [ref. 39118], Sonoyama et al. 2020:90 [ref. 37637]). Current status: Valid as Pseudoblennius cottoides (Richardson 1848). Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: China to Korea and Japan. Habitat: marine.

crassus, Icelus uncinalis Andriashev [A. P.] 1937:265, Pl. 5 (fig. 2) [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] Penzhina Bay, northeastern Sakhalin Island, Okhotsk Sea. Syntypes: ZIN 21900 (1), 21905 (6), 21910 (2), 25281 (1). "Typus" given on caption to Pl. 5 but specimen not distinguished in text. •Synonym in part of Icelus bicornis (Reinhardt 1840) -- (Nelson 1984:42 [ref. 5391]). •Synonym in part of Icelus ochotensis Schmidt 1927 -- (Nelson 1984:50 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300]). •Synonym in part of Icelus spatula Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:28 [ref. 25650]). •Synonym of Icelus ochotensis Schmidt 1927 -- (Parin et al. 2014:255 [ref. 33547]). Current status: Synonym of Icelus ochotensis Schmidt 1927. Psychrolutidae: Icelinae. Habitat: marine.

creaseri, Ruscariops Hubbs [C. L.] 1926:12 [Occasional Papers of the Museum of Zoology University of Michigan No. 170; ref. 6069] Rock pool at Bird Rock, San Diego, California, U.S.A. Holotype: UMMZ 141866. Paratypes: (9) UMMZ 141863-65 (1, 1, 6), ?141865 (6). •Valid as Artedius creaseri (Hubbs 1926) -- (Bolin 1944:43 [ref. 6379], Eschmeyer & Herald 1983:162 [ref. 9277]). •Valid as Ruscarius creaseri (Hubbs 1926) -- (Begle 1989:648 [ref. 12739], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Page et al. 2013:124 [ref. 32708], Kells et al. 2016:190 [ref. 35888], Burton & Lea 2019:68 [ref. 37205], Love et al. 2021:125 [ref. 39279]). Current status: Valid as Ruscarius creaseri (Hubbs 1926). Psychrolutidae: Psychrolutinae. Distribution: Northeastern Pacific: central California (U.S.A.) south to Pacific coast of central Baja California (Mexico). Habitat: marine.

criniger, Cottus Günther [A.] 1860:522 [Catalogue of the fishes in the British Museum v. 2; ref. 1963] Monterey, California, U.S.A. Syntypes: BMNH 1848.7.30.18-20 (3). •Synonym of Clinocottus analis (Girard 1858) -- (Bolin 1944:73 [ref. 6379]). Current status: Synonym of Clinocottus analis (Girard 1858). Psychrolutidae: Oligocottinae. Habitat: marine.

ctenopterygius, Phasmatocottus Bolin [R. L.] 1936:33, Fig. 7 [Proceedings of the United States National Museum v. 84 (no. 3000); ref. 505] Off Sendai Bay, Japan, 38°11'30"N, 142°08'00"E, Albatross station 5050, depth 266 fathoms. Holotype (unique): USNM 102107. •Valid as Phasmatocottus ctenopterygius Bolin 1936 -- (Yabe in Masuda et al. 1984:328 [ref. 6441], Nakabo 2000:639 [ref. 25086], Yabe & Fujii 2001:47 [ref. 25387], Nakabo 2002:639 [ref. 26001], Parin et al. 2014:267 [ref. 33547], Prokofiev 2017:648 [ref. 35721]). Current status: Valid as Phasmatocottus ctenopterygius Bolin 1936. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

decastrensis, Cottus Kner [R.] 1865:2, Pl. 2 (fig. 1) [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 24; ref. 6174] Decastris Bay at Amur River mouth, northeastern Asia. Holotype (unique): NMW 80072. Appeared as Cottus decastrensis n. sp.? •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Berg 1949:1137 [ref. 20662], Neelov 1979:121 [ref. 3152], Lindberg & Krasyukova 1987:207 [ref. 15964], Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:265 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138] with question). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

delacyi, Artedius Hubbs [C. L.] & Schultz [L. P.] 1941:4 [Occasional Papers of the Museum of Zoology University of Michigan No. 431; ref. 10913] Tidepools at Uyak Bay, Kodiak Island, Alaska, U.S.A. Holotype: UMMZ 126490. Paratypes: UMMZ 126491 (1), USNM 117494 (1). •Synonym of Artedius lateralis (Girard 1854) -- (Mecklenburg et al. 2002:459 [ref. 25968]). Current status: Synonym of Artedius lateralis (Girard 1854). Psychrolutidae: Oligocottinae. Habitat: marine.

dentatus, Japonopsychrolutes Nojima [S.] 1936:246 [Transactions of the Sapporo Natural History Society v. 14 (pt 4); ref. 3207] Japan. No types known. •Synonym of Eurymen gyrinus Gilbert & Burke 1912 -- (Watanabe 1960:188 [ref. 11959], Mecklenburg et al. 2002:517 [ref. 25968], Parin et al. 2014:301 [ref. 33547]). Current status: Synonym of Eurymen gyrinus Gilbert & Burke 1912. Psychrolutidae: Psychrolutinae. Habitat: marine.

derjavini, Radulinopsis Soldatov [V. K.] & Lindberg [G. U.] 1930:185, Figs. 30-31 [Izvestiia Tikhookeanskogo Nauchno-Issledovatel'skogo Instituta Rybnogo Khoziaistva i Okeanografii [Proceedings, Pacific Research Institute of Fisheries and Oceanography, TINRO, Moskva] v. 5; ref. 4164] Peter the Great Bay, Russia. Syntypes: (2) not at ZIN. •Valid as Radulinopsis derjavini Soldatov & Lindberg 1930 -- (Lindberg & Krasyukova 1987:187 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670] as derzhavini, Yabe & Maruyama 2001:53 [ref. 25245], Parin et al. 2002:S85 [ref. 28300], Parin et al. 2014:270 [ref. 33547], Dyldin & Orlov 2022:51 [ref. 38972]). Current status: Valid as Radulinopsis derjavini Soldatov & Lindberg 1930. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

derjugini, Icelus bicornis Soldatov [V. K.] in Knipowitsch 1926:124 [Trudy Nauchno-issledovatel'skogo instituta po izucheniya Severea. Moskva [= Transactions of the Institute for Scientific Exploration of the North.] No. 27; ref. 22166] Naiden, White Sea. Holotype: NIISM (whereabouts unknown). •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:36 [ref. 5391], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:253 [ref. 33547]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Habitat: marine.

derjugini, Malacocottus Popov [A. M.] 1931:130, Pl. 1 (fig. 3) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] Okhotsk Sea, 56°08'N, 144°54'E, depth 287 m; 56°09'N, 144°30'E, depth 307 meters. Syntypes: (2+ 1) ZIN 38408 [ex St. Petersburg Univ.] (1). Correct spelling of genus is Malacocottus. •Synonym of Malacocottus zonurus Bean 1890 -- (Sheiko & Fedorov 2000:30 [ref. 25650], Parin et al. 2002:S94 [ref. 28300], Parin et al. 2014:302 [ref. 33547], Stevenson 2015:24 [ref. 33680]). Current status: Synonym of Malacocottus zonurus Bean 1890. Psychrolutidae: Psychrolutinae. Habitat: marine.

derjugini, Radulinopsis Soldatov [V. K.] in Soldatov & Lindberg 1930:189, Figs. 32-33 [Izvestiia Tikhookeanskogo Nauchno-Issledovatel'skogo Instituta Rybnogo Khoziaistva i Okeanografii [Proceedings, Pacific Research Institute of Fisheries and Oceanography, TINRO, Moskva] v. 5; ref. 4164] Peter the Great Bay, Russia. Syntypes: (2) not at ZIN. •Valid as Radulinopsis derjugini Soldatov 1930 -- (Lindberg & Krasyukova 1987:188 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670]). •Synonym of Radulinopsis derjavini Soldatov & Lindberg 1930 -- (Yabe & Maruyama 2001:53 [ref. 25245]). Current status: Synonym of Radulinopsis derjavini Soldatov & Lindberg 1930. Psychrolutidae: Psychrolutinae. Habitat: marine.

detrisus, Gymnocanthus Gilbert [C. H.] & Burke [C. V.] 1912:61, Fig. 13 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off west coast of Kamchatka, Russia, 51°37'N, 156°21'E, Albatross station 4798, depth 25 fathoms. Holotype: USNM 74376. Paratypes: CAS-SU 22361 (2), USNM 74499 (2). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Gymnocanthus detrisus Gilbert & Burke 1912 -- (Watanabe 1960:61 [ref. 11959], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:200 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:629 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Mecklenburg et al. 2002:463 [ref. 25968], Nakabo 2002:629 [ref. 26001], Parin et al. 2002:S74 [ref. 28300] as Gymnacanthus, Fedorov et al. 2003:66 [ref. 27579] in Gymnacanthus, Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:86 [ref. 37547], Page et al. 2013:123 [ref. 32708], Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:246 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Dyldin & Orlov 2022:43 [ref. 38972], Love et al. 2021:120 [ref. 39279]). Current status: Valid as Gymnocanthus detrisus Gilbert & Burke 1912. Psychrolutidae: Icelinae. Distribution: North Pacific: northern Hokkaido (Japan) and Sea of Okhotsk north to Bering Sea, east to Alaska (U.S.A.). Habitat: marine.

diceraus, Cottus Pallas [P. S.] 1787:354, Pl. 10 (fig. 7) [Nova Acta Academiae Scientiarum Imperialis Petropolitanae v. 1 (Physica) (for 1783); ref. 18507] Kamchatka, Russia. Syntypes: ZIN 5380 (1); ZMB 5229 (not found), 6488 (1, dry) Type catalog: Paepke & Fricke 1992:282 [ref. 19946]. Date may be 1788. Type information: Svetovidov 1978:27 [ref. 6025]. •Valid as Ceratocottus diceraus (Pallas 1787) -- (Watanabe 1960:110 [ref. 11959]). •Valid as Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414], Neelov 1979:146 [ref. 3152], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:229 [ref. 15964], Kim & Youn 1992:61 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:630 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:472 [ref. 25968], Nakabo 2002:630 [ref. 26001], Choi et al. 2003:262, 574 [ref. 26218], Parin et al. 2002:S73 [ref. 28300], Fedorov et al. 2003:68 [ref. 27579] dated 1788, Nelson et al. 2004:119 [ref. 27807] dated 1788, Love et al. 2005:86 [ref. 37547], Mecklenburg et al. 2007:174 [ref. 29448], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:245 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Tohkairin et al. 2015:7 [ref. 35331], Mecklenburg et al. 2016:53 [ref. 34440], Kells et al. 2016:170 [ref. 35888], Mecklenburg et al. 2018:192 [ref. 36370], Kim et al. 2020:101 [ref. 39118], Dyldin et al. 2021:63 [ref. 38804], Dyldin & Orlov 2022:43 [ref. 38972], Love et al. 2021:120 [ref. 39279]). Current status: Valid as Enophrys diceraus (Pallas 1787). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific and adjacent Arctic: Sea of Japan and Sea of Okhotsk north to Bering Sea and Chukchi Sea, then east to southeastern Alaska (U.S.A.). Habitat: marine.

distoechus, Stlengis Bolin [R. L.] 1936:328, Pl. 34 (fig. A); Fig. 26 [Proceedings of the United States National Museum v. 83 (no. 2987); ref. 504] Wakayama, Japan, 33°24'50"N, 135°38'40"E, Albatross station 4968, depth 253 fathoms. Holotype: USNM 94728. Paratypes: CAS-SU 28727 (1), USNM 94729 (3). Type catalog: Böhlke 1953:132 [ref. 12291]. •Valid as Stlengis distoechus Bolin 1936 -- (Watanabe 1960:37 [ref. 11959], Fricke & Brunken 1984:46 [ref. 5262], Nakabo 2000:630 [ref. 25086], Shinohara et al. 2001:319 [ref. 25995], Nakabo 2002:630 [ref. 26001]). Current status: Valid as Stlengis distoechus Bolin 1936. Psychrolutidae: Oligocottinae. Distribution: Western North Pacific. Habitat: marine.

dolganovi, Gilbertidia Mandrytsa [S. A.] 1993:288, Fig. 1 [Voprosy Ikhtiologii v. 33 (no. 2); ref. 20793] Off Iturup Island, Kuril Islands, 44°37.8'N, 146°55'E, depth 200 meters. Holotype (unique): ZIN 50136. Generic classification of this species is in need of revision, as Gilbertidia Berg 1898 is now treated as a junior synonym of Psychrolutes Günther 1861, based on the placement of its type species. •Valid as Gilbertidia dolganovi Mandrytsa 1993 -- (Parin et al. 2002:S93 [ref. 28300], Parin et al. 2014:301 [ref. 33547]). •Valid as Psychrolutes dolganovi (Mandrytsa 1993) -- (Nazarkin et al. 2014:177 [ref. 33578]). Current status: Valid as Psychrolutes dolganovi (Mandrytsa 1993). Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

dorothy, Triglops Pietsch [T. W.] & Orr [J. W.] 2006:239, Fig. 1 [Fishery Bulletin v. 104 (no. 2); ref. 28666] Aniva Bay, 46°17'N, 143°09'E, Albatross station 5013, depth 77 meters. Holotype: USNM 74578. Paratypes: CAS-SU 22305 (3); HUMZ 41524 (1), 41525 (1); USNM 74575 (2), 74576 (4), 74577 (1), 74579 (8). •Valid as Triglops dorothy Pietsch & Orr 2006 -- (Parin et al. 2014:275 [ref. 33547], Dyldin & Orlov 2022:52 [ref. 38972]). Current status: Valid as Triglops dorothy Pietsch & Orr 2006. Psychrolutidae: Triglopinae. Distribution: Northwestern Pacific: Sea of Okhotsk, Japan and Kuril Islands north to Sakhalin (Russia). Habitat: marine.

dybowskii, Cottus Hilgendorf [F. M.] 1879:106 [Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1879; ref. 21467] No locality [Japan]. Holotype (unique): ZMB 10628. Type catalog: Paepke & Fricke 1992:282 [ref. 19946]. •Synonym of Myoxocephalus brandtii (Steindachner 1867) -- (Prokofiev 2017:648 [ref. 35721]). Current status: Synonym of Myoxocephalus brandtii (Steindachner 1867). Psychrolutidae: Myoxocephalinae.

dydymovi, Artediellus Soldatov [V. K.] 1915:157, Figs. 1 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 15068] Tatar Strait, 49°28'10"N, 140°39'30"E, Russia, northern Sea of Japan. Holotype: ZIN 18693 (male of 3). Paratypes: ZIN 18653 (6), 18655-63 (6, 6, 4, 3, 2, 1, 1, 1, 1), 18693 (2 females), 19435 (1). •Valid as Artediellus dydymovi Soldatov 1915, subspecies dydymovi Soldatov 1915 -- (Nakabo 2000:642 [ref. 25086], Nakabo 2002:642 [ref. 26001]). •Valid as Artediellus dydymovi Soldatov 1915 -- (Neelov 1979:155 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Nelson 1986:43 [ref. 5808], Lindberg & Krasyukova 1987:235 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Sheiko & Fedorov 2000:27 [ref. 25650], Parin et al. 2002:S70 [ref. 28300], Fedorov et al. 2003:66 [ref. 27579], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:236 [ref. 33547], Dyldin & Orlov 2017:61 [ref. 35138], Sonoyama et al. 2020:89 [ref. 37637], Dyldin et al. 2021:61 [ref. 38804], Dyldin & Orlov 2022:39 [ref. 38972]). Current status: Valid as Artediellus dydymovi Soldatov 1915. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: brackish, marine.

ecornis, Icelus Tsutsui [D.] & Yabe [M.] 1996:302, Fig. 2 [Ichthyological Research v. 43 (no. 3); ref. 22432] Kitami-Yamato Bank, southwestern Okhotsk Sea, off Abashiri, Hokkaido, Japan, 44°30.1'N, 144°14.5'E, depth 208 meters. Holotype: HUMZ 125990. Paratypes: CAS 82510 (2); HUMZ 124100-01 (1, 1), 124326 (1), 124832-34 (1, 1, 1), 124836 (1), 124838 (1), 124845 (1), 124848 (1), 124852 (1), 125966-67 (1, 1), 125975 (1), 125980 (1), 125985 (1), 125991 (1), 125999 (1), 126000 (1), 126002 (1), 126006 (1), 126018 (1), 126020-21 (1, 1); NSMT-P 47286-87 (1, 1); UMMZ 230706 (2). •Valid as Icelus ecornis Tsutsui & Yabe 1996 -- (Nakabo 2000:635 [ref. 25086], Nakabo 2002:635 [ref. 26001], Tsuruoka et al. 2006:51 [ref. 28501], Dyldin & Orlov 2022:46 [ref. 38972]). Current status: Valid as Icelus ecornis Tsutsui & Yabe 1996. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

edomius, Myoxocephalus Jordan [D. S.] & Starks [E. C.] 1904:274, Fig. 18 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Aino village of Edomo, near Mororan, Japan. Holotype: CAS-SU 7707. Paratypes: USNM 50919 (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Myoxocephalus edomius Jordan & Starks 1904 -- (Watanabe 1960:68 [ref. 11959]). •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Neelov 1979:122 [ref. 3152], Lindberg & Krasyukova 1987:212 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:261 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Habitat: marine.

elegans, Antipodocottus Fricke [R.] & Brunken [H.] 1984:41, Figs. 1-2 [Journal of Natural History v. 18 (no. 1); ref. 5262] Off Wollongong, New South Wales, Australia, 34°21'S, 151°24'E, depth 220 fathoms. Holotype: AMS I.18795-007. •Valid as Antipodocottus elegans Fricke & Brunken 1984 -- (Paxton et al. 1989:477 [ref. 12442], Nelson 1990:841 [ref. 15720], Nelson & Carpenter 1999:2426 [ref. 24809], Paxton et al. 2006:952 [ref. 28994], Fricke et al. 2014:56 [ref. 33932], Stewart 2015:1118 [ref. 34254]). Current status: Valid as Antipodocottus elegans Fricke & Brunken 1984. Psychrolutidae: Psychrolutinae. Distribution: Western Pacific: Australia, Papua New Guinea and New Zealand region. Habitat: marine.

elegans, Centridermichthys Steindachner [F.] 1881:185 [7], Pl. 6 (figs. 1, 1a) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 83 (1. Abth.); ref. 4231] Strietok, near Vladivostock, Russia, Sea of Japan. Syntypes: NMW 6374 (3), 6578 (2), 78819 (1). On p. 7 of separate. •Valid as Bero elegans (Steindachner 1881) -- (Watanabe 1960:134 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:250 [ref. 15964], Kim & Youn 1992:72 [ref. 22287], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:648 [ref. 25086], Nakabo 2002:648 [ref. 26001], Choi et al. 2003:262, 573 [ref. 26218], Parin et al. 2002:S72 [ref. 28300], Parin et al. 2014:240 [ref. 33547], Kim et al. 2020:101 [ref. 39118], Dyldin & Orlov 2022:41 [ref. 38972]). Current status: Valid as Bero elegans (Steindachner 1881). Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

elegans, Cottus Gray [J. E.] in Cuvier & Valenciennes 1829:195 [Histoire naturelle des poissons v. 4; ref. 998] Name only mentioned under Cottus claviger. •In the synonymy of Enophrys diceraus (Pallas 1787). Nomen Nudum. Current status: Synonym of Enophrys diceraus (Pallas 1787). Psychrolutidae: Myoxocephalinae.

elongatus, Centridermichthys Steindachner [F.] 1881:186 [8] [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 83 (1. Abth.); ref. 4231] Sea of Japan. On p. 8 of separate. Apparently figured as Centridermichthys glaber n. sp. in Pl. 6 (fig. 2). See account of Centridermichthys glaber Steindachner 1881. •Valid as Alcichthys elongatus (Steindachner 1881) -- (Lindberg & Krasyukova 1987:243 [ref. 15964] as Elophichthys, Nakabo 2000:648 [ref. 25086], Nakabo 2002:648, 1526 [ref. 26001], Parin et al. 2002:S68 [ref. 28300], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:232 [ref. 33547], Kim et al. 2020:101 [ref. 39118], Sonoyama et al. 2020:89 [ref. 37637], Dyldin & Orlov 2022:38 [ref. 38972]). Current status: Valid as Alcichthys elongatus (Steindachner 1881). Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

embryum, Oligocottus Jordan [D. S.] & Starks [E. C.] 1895:808, Pl. 82 [Proceedings of the California Academy of Sciences (Series 2) v. 5; ref. 2522] Several miles east of Neah Bay, Washington, U.S.A. Lectotype: CAS-SU 3128. Paralectotypes: ?BMNH 1897.1.12.16 (1), CAS-SU 3128 (1 of 2). Additional material: (1). Type catalog: Böhlke 1953:131 [ref. 12291]. Lectotype established (as figured specimen) in caption to Pl. 303, p. 3294 in Jordan & Evermann 1900 [ref. 2446]. •Valid as Clinocottus embryum (Jordan & Starks 1895) -- (Bolin 1944:78 [ref. 6379], Eschmeyer & Herald 1983:166 [ref. 9277], Matarese et al. 1989:424 [ref. 26885], McAllister 1990:196 [ref. 14674], Mecklenburg et al. 2002:503 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:85 [ref. 37547], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:39 [ref. 34694], Kells et al. 2016:168 [ref. 35888], Burton & Lea 2019:66 [ref. 37205], Love et al. 2021:119 [ref. 39279]). Current status: Valid as Clinocottus embryum (Jordan & Starks 1895). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: Aleutian Islands (U.S.A.) south to Pacific coast of northern Baja California (Mexico). Habitat: marine.

ensiger, Ainocottus Jordan [D. S.] & Starks [E. C.] 1904:283, Fig. 23 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate,Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7709. Paratypes: USNM 50921 (not found in 1980). Additional material: CAS-SU 7596 (4). Type catalog: Böhlke 1953:126 [ref. 12291]. •Valid as Ainocottus ensiger Jordan & Starks 1904 -- (Watanabe 1960:87 [ref. 11959], Kim & Youn 1992:63 [ref. 22287]). •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Neelov 1979:121 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Choi et al. 2003:576 [ref. 26218], Nakabo 2002:1526 [ref. 26001], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547], Mecklenburg et al. 2016:79 [ref. 34440], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

europaeus, Artediellus Knipowitsch [N. M.] 1907:17 [Zapiski Imperatorskoj Akademii Nauk po Fiziko-Matematiceskomu Otdelniji = Mémoires de l'Académie Impériale des Sciences de St. Petersbourg, Classe Physico-Mathématique (8 Sér.) v. 18 (no. 5); ref. 15152] One of: west coast of Norway, Svalbard, and European Arctic Ocean. Lectotype: ZIN 28393. Paralectotypes: (19) ZIN 28393 (4, with lectotype), 28449 (6+). Lectotype selected by Neelov 1979:154 [ref. 3152]. •Synonym of Artediellus atlanticus Jordan & Evermann 1898, but a valid subspecies -- (Neelov 1973:594 [ref. 7219], Neelov 1979:154 [ref. 3152], Fedorov in Whitehead et al. 1986:1245 [ref. 13677] as europeus, Yabe in Okamura et al. 1995:161 [ref. 22531], Parin et al. 2002:S70 [ref. 28300], Dolgov 2013:918 [ref. 33112] as europeus). •Synonym of Artediellus atlanticus Jordan & Evermann 1898 -- (Parin et al. 2014:236 [ref. 33547]). Current status: Synonym of Artediellus atlanticus Jordan & Evermann 1898. Psychrolutidae: Psychrolutinae. Habitat: marine.

euryops, Icelus Bean [T. H.] 1890:41 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Off Trinity Island, Alaska, U.S.A., Albatross station 2853, 56°N, 154°20'W, depth 159 fathoms. Lectotype: USNM 45367. Paralectotypes: USNM 227363 [ex USNM 45367] (2). Lectotype selected by Nelson 1984:30 [ref. 5391]. •Valid as Icelus euryops Bean 1890 -- (Nelson 1984:30 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:453 [ref. 25968], Parin et al. 2002:S77 [ref. 28300], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Maslenikov et al. 2013:10 [ref. 37947], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:254 [ref. 33547], Love et al. 2021:122 [ref. 39279]). Current status: Valid as Icelus euryops Bean 1890. Psychrolutidae: Icelinae. Distribution: North Pacific: northern Bering Sea to western Gulf of Alaska (U.S.A.). Habitat: marine.

euryptera, Bolinia Yabe [M.] 1991:329, Figs. 1-3 [Copeia 1991 (no. 2); ref. 17338] Near Amukta Pass, Aleutian Islands, 52°51'N, 171°13'W, depth 205-225 meters. Holotype: HUMZ 68566. Paratypes: CAS 68097-98 (2); HUMZ 68293 (1), 68296-98 (3), 68552-53 (2), 68555-60 (6), 68567-68 (2), 88847-48 (2), 88869-72 (4), 89107-111 (5), 89113-123 (11), 89049-53 (5), 110007-10 (4), 110015-17 (3); NSMT-P 45082-83 (2), OSOU 10878-79 (5, 2); plus additional non-type material. •Valid as Bolinia euryptera Yabe 1991 -- (Mecklenburg et al. 2002:488 [ref. 25968], Love et al. 2005:85 [ref. 37547], Love et al. 2021:119 [ref. 39279]). Current status: Valid as Bolinia euryptera Yabe 1991. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Aleutian Islands (U.S.A.) endemic. Habitat: marine.

eurystigmatephoros, Ambophthalmos Jackson [K. L.] & Nelson [J. S.] 1999:428, Figs. 1-4 [Copeia 1999 (no. 2); ref. 23799] Campbell Island Rise, 51°48.5'S, 169°04.9'E, north of Campbell Island, New Zealand, depth 230-282 meters. Holotype: NMNZ P 24882-1. Paratypes: NMNZ P 24882-2 (1), 24882-3 (1). •Valid as Ambophthalmos eurystigmatephoros Jackson & Nelson 1999 -- (Jackson & Nelson 2000:725 [ref. 25146], Stewart 2015:1124 [ref. 34254]). Current status: Valid as Ambophthalmos eurystigmatephoros Jackson & Nelson 1999. Psychrolutidae: Psychrolutinae. Distribution: Southwestern Pacific: New Zealand region (Campbell Plateau). Habitat: marine.

fabricii, Cottus Girard [C. F.] 1850:411 [Proceedings of AAAS (The American Association for the Advancement of Science) 2nd meeting, 1849; ref. 18123] Greenland. Based on Cottus gobio of Fabricius 1780. See also Girard 1850:189 [ref. 1814] and Girard 1851:59 [ref. 18124]. •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Neelov 1973:595 [ref. 7219] as Gymnacanthus). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Psychrolutidae: Icelinae. Habitat: marine.

fasciatus, Ainocottus Pavlenko [M. N.] 1910:30 [Trudy Obshchestva Estestvoispytatelej Imperatorskom' Kazanskom' Universiteti, Kazan v. 42 (no. 2); ref. 3393] Peter the Great Bay, Russia, Sea of Japan. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Neelov 1979:122 [ref. 3152], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

fenestralis, Artedius Jordan [D. S.] & Gilbert [C. H.] 1883:577 [Proceedings of the United States National Museum v. 5 (no. 326); ref. 10614] Commencement Bay near New Tacoma, Washington, U.S.A. Syntypes: USNM 27146 (7), 27206 (2). •Valid as Artedius fenestralis Jordan & Gilbert 1883 -- (Bolin 1944:48 [ref. 6379], Eschmeyer & Herald 1983:162 [ref. 9277], Begle 1989:646 [ref. 12739], Matarese et al. 1989:414 [ref. 26885], McAllister 1990:194 [ref. 14674], Mecklenburg et al. 2002:460 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:85 [ref. 37547], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:164 [ref. 35888], Burton & Lea 2019:65 [ref. 37205], Love et al. 2021:118 [ref. 39279]). Current status: Valid as Artedius fenestralis Jordan & Gilbert 1883. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: eastern Aleutian Islands south to southern California (U.S.A.). Habitat: marine.

filamentosus, Cottus Sauvage [H.-E.] in Vaillant & Sauvage 1875:279 [Revue et Magasin de Zoologie (Sér. 3) v. 3; ref. 17853] "Hawaiian Islands" [error]. Syntypes: MNHN 0000-9090 (2). Type catalog: Blanc & Hureau 1968:51 [ref. 20738]. •Apparently a synonym of Gymnocanthus intermedius (Temminck & Schlegel 1843) -- (see Mundy 2005:336 [ref. 28379]). Current status: Synonym of Gymnocanthus intermedius (Temminck & Schlegel 1843). Psychrolutidae: Icelinae. Habitat: marine.

filamentosus, Icelinus Gilbert [C. H.] 1890:85 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., Albatross station 2893 and 2959, depths 145 and 55 fathoms. Syntypes: CAS-SU 118 (1), 28109 (1); USNM 44407 (1), 48245 (2), 125011 [ex USBF 449] (2). Type catalog: Böhlke 1953:130 [ref. 12291]. Lectotype established (as figured specimen) in Jordan 1896:244 [ref. 2395] if specimen is identifiable. •Valid as Icelinus filamentosus Gilbert 1890 -- (Bolin 1944:24 [ref. 6379], Eschmeyer & Herald 1983:173 [ref. 9277], Peden 1984:79 [ref. 5193], McAllister 1990:198 [ref. 14674], Mecklenburg et al. 2002:439 [ref. 25968], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:176 [ref. 35888], Burton & Lea 2019:67 [ref. 37205], Love et al. 2021:121 [ref. 39279]). Current status: Valid as Icelinus filamentosus Gilbert 1890. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: western Gulf of Alaska south to southern California (U.S.A.). Habitat: marine.

fimbriatus, Icelinus Gilbert [C. H.] 1890:87 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., Albatross station 2893 and 2975, depths 45 and 36 fathoms. Syntypes: USNM 43087 (1), 48246 (1). •Valid as Icelinus fimbriatus Gilbert 1890 -- (Bolin 1944:32 [ref. 6379], Eschmeyer & Herald 1983:173 [ref. 9277], Peden 1984:67 [ref. 5193], McAllister 1990:198 [ref. 14674], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:176 [ref. 35888], Burton & Lea 2019:67 [ref. 37205], Love et al. 2021:121 [ref. 39279]). Current status: Valid as Icelinus fimbriatus Gilbert 1890. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: southern British Columbia (Canada) south to Pacific coast of southern California (U.S.A.). Habitat: marine.

forficata, Elanura Gilbert [C. H.] 1896:430, Pl. 30 (top) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] South of Sannak Island, Aleutian Islands, Albatross station 3214, depth 70 meters. Lectotype: MCZ 28266. Paralectotypes: BMNH 1896.7.23.149 (1); NMW 11468 (2); CAS-SU 2420 (3); USNM 48673 (1), 59404 (1), 119654 (1); ZMUO J1442 (1) station 3214. Type catalog: Böhlke 1953:129 [ref. 12291], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen from Albatross sta. 3214) in caption to Pl. 289, p. 3291 in Jordan & Evermann 1900 [ref. 2446]; this is the specimen figured with the original description and is apparently MCZ 28266. This specimen was also designated lectotype by Pietsch 1994:354 [ref. 21189]. •Valid as Triglops forficatus (Gilbert 1896) -- (Sheiko 1983:18 [ref. 37041], Pietsch 1994:354 [ref. 21189] as forficata, Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:424 [ref. 25968], Parin et al. 2002:S87 [ref. 28300], Fedorov et al. 2003:81 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:93 [ref. 37547], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:275 [ref. 33547], Kells et al. 2016:192 [ref. 35888], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Triglops forficatus (Gilbert 1896). Psychrolutidae: Triglopinae. Distribution: North Pacific: Kuril Islands north to Bering Sea, then east to northwestern Gulf of Alaska; one record from southeastern Alaska (U.S.A.). Habitat: marine.

frykenensis, Cottus quadricornis Lönnberg [E.] 1932:14, Pl. (fig. 5) [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Fryken, Sweden. Syntypes: NRM 30315 (1), Mus. of Karlstad's school (1), Uppsala Univ. Zool. Mus. (1). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

furciger, Icelus Malm [A. W.] 1865:410 [Förhandlingar vid de Skandinaviske Naturforskarnes. v. 9; ref. 17596] Löken, Gullmarfjärden, Sweden. No types known. •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:36 [ref. 5391]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Habitat: marine.

fuscescens, Icelinus Gilbert [C. H.] 1915:340, Pl. 17 (fig. 9) [Proceedings of the United States National Museum v. 48 (no. 2075); ref. 1632] Off Santa Barbara Island, California, U.S.A., Albatross station 4418, depth 260-310 fathoms. Holotype: USNM 75812. Paratypes: CAS-SU 22981 (1), 22985 (1), 22991 (1), 23001 (6). Type catalog: Böhlke 1953:130 [ref. 12291]. •Synonym of Icelinus burchami Evermann & Goldsborough 1907 -- (Bolin 1944:30 [ref. 6379], Mecklenburg et al. 2002:440 [ref. 25968]). •Synonym of Icelinus burchami Evermann & Goldsborough 1907, but a valid subspecies -- (Peden 1984:79 [ref. 5193]). Current status: Synonym of Icelinus burchami Evermann & Goldsborough 1907. Psychrolutidae: Oligocottinae. Habitat: marine.

fuscimentus, Artediellus (Artediellops) Nelson [D. W.] 1986:41, Fig. 10 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 138 (no. 1); ref. 5808] Southern Sea of Japan, 36°34'N, 132°50'20"E, Albatross station 4844, depth 202 meters. Holotype: USNM 74464. Paratypes: CAS-SU 22230 (24, 1 c&s); USNM 74491 (5), 74493-95 (1, 11, 3), 74497-98 (1, 1). •Valid as Artediellus fuscimentus Nelson 1986 -- (Nakabo 2000:642 [ref. 25086], Nakabo 2002:642, 1526 [ref. 26001], Shinohara et al. 2011:43 [ref. 31715], Shinohara et al. 2014:248 [ref. 33330], Shin et al. 2016:209 [ref. 34833], Kim et al. 2020:101 [ref. 39118], Sonoyama et al. 2020:89 [ref. 37637]). Current status: Valid as Artediellus fuscimentus Nelson 1986. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

galatheae, Antipodocottus Bolin [R. L.] 1952:432, Fig. 1 [Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, Kjøbenhavn v. 114; ref. 511] Tasman Sea off New Zealand, 42°10'S, 170°10'E, Galathea station 626, depth 594 [or 610] meters. Holotype: ZMUC P811234. Paratypes: CAS-SU 17360 (1), ZMUC P811235-36 (2). Type catalog: Böhlke 1953:126 [ref. 12291], Nielsen 1974:81 [ref. 9588]. •Valid as Antipodocottus galatheae Bolin 1952 -- (Nelson 1975:80 [ref. 30370], Fricke & Brunken 1984:46 [ref. 5262], Paxton et al. 1989:477 [ref. 12442], Paulin et al. 1989:172 [ref. 24556], Paxton et al. 2006:952 [ref. 28994], Stewart 2015:1119 [ref. 34254], Girard & Smith 2016:556 [ref. 34786]). Current status: Valid as Antipodocottus galatheae Bolin 1952. Psychrolutidae: Psychrolutinae. Distribution: Southwestern Pacific: Australia and New Zealand. Habitat: marine.

galeatus, Gymnacanthus Bean [T. H.] 1881:153 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Iliuliuk, Unalaska Island, Aleutian Islands, Alaska, U.S.A. Lectotype: USNM 28097. Paralectotypes: USNM 28097 (2 of 3, orig. 5). Original genus should have been spelled Gymnocanthus. Lectotype established (as figured specimen) in caption to Pl. 301, p. 3293 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. •Valid as Gymnocanthus galeatus Bean 1881 -- (Watanabe 1960:59 [ref. 11959], Eschmeyer & Herald 1983:169 [ref. 9277], Yabe in Masuda et al. 1984:328 [ref. 6441], McAllister 1990:197 [ref. 14674], Nakabo 2000:629 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Mecklenburg et al. 2002:466 [ref. 25968], Nakabo 2002:629 [ref. 26001], Fedorov et al. 2003:69 [ref. 27579] as Gymnacanthus, Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:86 [ref. 37547], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:247 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Mecklenburg et al. 2016:57 [ref. 34440], Kells et al. 2016:172 [ref. 35888], Balanov & Matveev 2018:367 [431] [ref. 36463], Mecklenburg et al. 2018:196 [ref. 36370], Dyldin & Orlov 2022:43 [ref. 38972], Love et al. 2021:120 [ref. 39279]). Current status: Valid as Gymnocanthus galeatus Bean 1881. Psychrolutidae: Icelinae. Distribution: North Pacific: northern Hokkaido (Japan), northern Sea of Japan and Sea of Okhotsk north to Bering Sea, then south to British Columbia (Canada); also adjacent Arctic Region. Habitat: marine.

gibber, Malacocottus Sakamoto [K.] 1930:16, Fig. 2 [Journal of the Imperial Fisheries Institute v. 26; ref. 16327] Toyama Bay, Japan. Holotype (unique): whereabouts unknown. •Valid as Malacocottus gibber Sakamoto 1930 -- (Watanabe 1960:180 [ref. 11959], Yabe in Amaoka et al. 1983:157, 215 [ref. 8379], Yabe in Masuda et al. 1984:330 [ref. 6441], Lindberg & Krasyukova 1987:271 [ref. 15964], Shinohara et al. 1992 [ref. 19776], Nakabo 2000:651 [ref. 25086], Nakabo 2002:651 [ref. 26001], Parin et al. 2002:S94 [ref. 28300], Shinohara et al. 2011:46 [ref. 31715] dated 1832, Shinohara et al. 2014:254 [ref. 33330], Parin et al. 2014:302 [ref. 33547], Stevenson 2015:31 [ref. 33680], Kim et al. 2020:107 [ref. 39118], Sonoyama et al. 2020:91 [ref. 37637], Dyldin et al. 2021:97 [ref. 38934], Balanov et al. 2022: 119 [109] [ref. 39284], Dyldin & Orlov 2022:55 [ref. 38972]). Current status: Valid as Malacocottus gibber Sakamoto 1930. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia). Habitat: marine.

gilberti, Icelus Taranetz [A. Ya.] 1936:149, Fig. [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (pt 13) (No. 3, art. 107); ref. 6509] Sea of Japan (Vladmir and Olga Bays and Yuzhnyy Cape) and Okhotsk Sea (Aniva Cape), depth 56-180 meters. Syntypes: (many) ZIN 25514 (4), 25517 (1), 25328 (3). •Valid as Icelus gilberti Taranetz 1936 -- (Nelson 1984:21 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:192 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:636 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Nakabo 2002:636 [ref. 26001], Parin et al. 2002:S77 [ref. 28300], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:254 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Dyldin & Orlov 2022:46 [ref. 38972]). Current status: Valid as Icelus gilberti Taranetz 1936. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

gilberti, Oncocottus hexacornis Fowler [H. W.] 1905:366, Figs. 2-5 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 57; ref. 15353] Point Barrow, northern Alaska, U.S.A. Holotype: ANSP not found [ex WIAP 7203]. Paratypes: WIAP (1). Type catalog: Böhlke 1984:65 [ref. 13621] as types not found, specimens perhaps never deposited at ANSP. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies hexacornis (Richardson 1823) -- (Berg 1949:1129 [ref. 20662]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

gilli, Synchirus Bean [T. H.] 1890:642 [Proceedings of the United States National Museum v. 12 (no. 787); ref. 228] Barkley Sound [Barclay Sound], west coast of Vancouver Island, British Columbia, Canada. Syntypes: USNM 41820 (now 4). •Valid as Synchirus gilli Bean 1890 -- (Eschmeyer & Herald 1983:183 [ref. 9277], Matarese et al. 1989:410 [ref. 26885], McAllister 1990:201 [ref. 14674], Mecklenburg et al. 2002:418 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Page et al. 2013:125 [ref. 32708], Pietsch & Orr 2015:45 [ref. 34694], Kells et al. 2016:192 [ref. 35888], Burton & Lea 2019:68 [ref. 37205], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Synchirus gilli Bean 1890. Psychrolutidae: Psychrolutinae. Distribution: Northeastern Pacific: Unalaska Island and Gulf of Alaska (U.S.A.) south to southern California (U.S.A.). Habitat: marine.

glaber, Centridermichthys Steindachner [F.] 1881:219 [41], Pl. 6 (fig. 2) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 83 (1. Abth.); ref. 4231] Appeared as "Fig. 2. Centridermichthys glaber n. sp." in figure captions on p. 41; this corresponds to Centridermichthys elongatus on p. 186 [8]. Regarded as two names for the same taxon; first reviser not researched. •Synonym of Alcichthys elongatus (Steindachner 1881) -- (Parin et al. 2002:S68 [ref. 28300], Parin et al. 2014:233 [ref. 33547]). Current status: Synonym of Alcichthys elongatus (Steindachner 1881). Psychrolutidae: Psychrolutinae. Habitat: marine.

glacialis, Cottus Richardson [J.] 1855:349, Pl. 24 [The last of the Arctic voyages in search of Sir J. Franklin; ref. 18631] Northumberland Sound, 76°53'N, western Greenland, depth 9 fathoms. No types known. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

globiceps, Oligocottus Girard [C. F.] 1858:58 [General report upon zoology of the several Pacific railroad routes, 1857; ref. 4911] Southern Farallones Island, off San Francisco, California, U.S.A. Syntypes: USNM 300 (orig. 4, now 2) Also appeared in Girard 1858:202 [ref. 1811]. •Valid as Clinocottus globiceps (Girard 1858) -- (Bolin 1944:81 [ref. 6379], Eschmeyer & Herald 1983:166 [ref. 9277], Matarese et al. 1989:426 [ref. 26885], McAllister 1990:196 [ref. 14674], Mecklenburg et al. 2002:504 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:86 [ref. 37547], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:39 [ref. 34694], Kells et al. 2016:168 [ref. 35888], Burton & Lea 2019:66 [ref. 37205], Love et al. 2021:119 [ref. 39279]). Current status: Valid as Clinocottus globiceps (Girard 1858). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: western Gulf of Alaska south to southern California (U.S.A.). Habitat: marine.

gomojunovi, Artediellus Taranetz [A. Ya.] 1933:71 [Bulletin of the Far Eastern Branch of the Academy of Sciences U.S.S.R. 1933 (nos 1-2-3); ref. 15743] Gulf of Anadyr and Olytorski, northwestern Bering Sea. Lectotype: ZIN 25337 (male). Paralectotypes: (4+) ZIN 25337 (female). Lectotype selected by Neelov 1979:155 [ref. 3152]. •Valid as Artediellus gomojunovi Taranetz 1933 -- (Neelov 1979:155 [ref. 3152], Nelson 1986:43 [ref. 5808], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:490 [ref. 25968], Parin et al. 2002:S70 [ref. 28300], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:84 [ref. 37547], Mecklenburg et al. 2011:122 [ref. 31212] with comments, Page et al. 2013:121 [ref. 32708], Parin et al. 2014:237 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Love et al. 2021:118 [ref. 39279]). Current status: Valid as Artediellus gomojunovi Taranetz 1933. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Kuril Islands north to Bering Sea off Alaska (U.S.A.) Habitat: marine.

gonez, Cottiusculus Jordan [D. S.] & Starks [E. C.] (ex Schmidt) 1904:298, Fig. 29 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Aniva Bay, Sakhalin Island, Russia. Lectotype: CAS-SU 7713. Paralectotypes: CAS-SU 69887 [ex CAS-SU 7713 [ex ZIN]] (4). Type catalog: Böhlke 1953:127 [ref. 12291]. Appeared without distinguishing features in Schmidt 1903:518 [p. 16 of separate] [ref. 3945]. Apparently appeared first in an available way as above, then in Schmidt 1904:108, Pl. 11 [ref. 3946]. Types are specimens used by Jordan and not by Schmidt; Schmidt's specimens are ZISP 12736-37 (20, 6), 12739-43 (1, 2, 8, 2, 4). Lectotype selected by Kai & Nakabo 2009:223 [ref. 30299]. •Valid as Cottiusculus gonez Jordan & Starks 1904 [sometimes with author as Schmidt] -- (Watanabe 1960:170 [ref. 11959], Neelov 1979:162 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:238 [ref. 15964], Kim & Youn 1992:71 [ref. 22287], Cheng & Zhou 1997:436 [ref. 26385] with author as Schmidt, Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:643 [ref. 25086], Nakabo 2002:643 [ref. 26001], Choi et al. 2003:262, 573 [ref. 26218], Parin et al. 2002:S72 [ref. 28300], Fedorov et al. 2003:67 [ref. 27579] as Schmidt in Jordan & Starks 1904, Kai & Nakabo 2009:222 [ref. 30299], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:240 [ref. 33547], Kim et al. 2020:101 [ref. 39118], Prokofiev 2020:13 [ref. 37765], Dyldin & Orlov 2022:41 [ref. 38972]). Current status: Valid as Cottiusculus gonez Jordan & Starks 1904. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

granulosus, Cottunculus Karrer [C.] 1968:560, Fig. 7 [Zoologische Jahrbücher, Abteilung für Systematik, Ökologie und Geographie der Tiere v. 95 (no. 4); ref. 7898] Off southern coast of Argentina, 48°35'S, 60°34'W, depth 390-400 meters. Holotype (unique): ZMB 21717. Type catalog: Paepke & Fricke 1992:285 [ref. 19946]. •Valid as Cottunculus granulosus Karrer 1968 -- (Nelson 1977:508 [ref. 7034], Nelson & Nakamura 1980:982 [ref. 26434], Nakamura in Nakamura et al. 1986:186 [ref. 14235], Nelson 1989:407 [ref. 12546], Pequeño 1989:57 [ref. 14125], Lima & Mincarone 2004:39 [ref. 27821], Byrkjedal & Orlov 2007:67 [ref. 29312], Costa & Mincarone 2010:348 [ref. 37483], Byrkjedal et al. 2014:1 [ref. 33415], Nión et al. 2016:45 [ref. 35565]). Current status: Valid as Cottunculus granulosus Karrer 1968. Psychrolutidae: Psychrolutinae. Distribution: Southwestern Atlantic; southeastern Pacific. Habitat: marine.

groenlandicus, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:185 [Histoire naturelle des poissons v. 4; ref. 998] Greenland. No types known. Type catalog: Blanc & Hureau 1968:60 [ref. 20738]. Based on Cottus scorpius of Fabricius. •Synonym of Myoxocephalus scorpius (Linnaeus 1758), but a valid subspecies -- (Neelov 1973:599 [ref. 7219], Neelov 1979:122 [ref. 3152], Fedorov in Whitehead et al. 1986:1255 [ref. 13677], Mecklenburg et al. 2007:176 [ref. 29448]). •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Scott & Scott 1988:509 [ref. 25518], Yabe in Okamura et al. 1995:165 [ref. 22531], Mecklenburg et al. 2002:478 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Mecklenburg et al. 2011:123 [ref. 31212], Parin et al. 2014:265 [ref. 33547], Mecklenburg et al. 2016:85 [ref. 34440]). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

groenlandicus, Gymnacanthus tricuspis Schmidt [P. Yu.] 1927:28 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 15821] Arctic. Syntypes: ZMUC (2), ZIN 1930-31 (1, 1). •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Neelov 1973:595 [ref. 7219] as Gymnacanthus, Mecklenburg et al. 2016:62 [ref. 34440]). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Psychrolutidae: Icelinae. Habitat: marine.

gyrinoides, Cottunculus Weber [M.] 1913:505, Fig. 106 [Die Fische der Siboga-Expedition; ref. 4602] Flores Sea, 7°24'S, 118°15.2'E, Siboga station 45, depth 794 meters. Syntypes: ZMA 108188-89 (1, 1). Type catalog: Nijssen et al. 1982:81 [ref. 19236]. •Valid as Cottunculus gyrinoides Weber 1913 -- (Nelson 1977:508 [ref. 7034]). •Valid, possibly in Ebinania -- (Byrkjedal & Orlov 2007:67 [ref. 29312] on Nelson 1989). Current status: Valid as Ebinania gyrinoides (Weber 1913). Psychrolutidae: Psychrolutinae. Distribution: Western Pacific. Habitat: marine.

gyrinops, Besnardia Lahille [F.] 1913:2, Pl. 1 [Anales del Museo Nacional de Historia Natural de Buenos Aires v. 24; ref. 2712] Mar de la Plata, Buenos Aires Province, Argentina, depth 80 meters. Holotype (unique): MACN 3303 [? 3304]. Type catalog: Braga & Piacentino 1994:101 [ref. 23697] as girinops. •Synonym of Neophrynichthys marmoratus Gill 1889 -- (Nelson 1977:501 [ref. 7034]). •Synonym of Psychrolutes marmoratus (Gill 1889). Current status: Synonym of Psychrolutes marmoratus (Gill 1889). Psychrolutidae: Psychrolutinae. Habitat: marine.

gyrinus, Eurymen Gilbert [C. H.] & Burke [C. V.] 1912:64, Fig. 14 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off Avatcha Bay, east coast of Kamchatka, Russia, Albatross station 4795, depth 69 fathoms. Holotype (unique): USNM 74377. •Valid as Eurymen gyrinus Gilbert & Burke 1912 -- (Watanabe 1960:188 [ref. 11959], Eschmeyer & Herald 1983:169 [ref. 9277], Yabe in Masuda et al. 1984:330 [ref. 6441], Lindberg & Krasyukova 1987:265 [ref. 15964], Kim et al. 1993:22 [ref. 22288], Jackson & Nelson 1998:1355 [ref. 23806], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:651 [ref. 25086], Sheiko & Fedorov 2000:30 [ref. 25650], Mecklenburg et al. 2002:517 [ref. 25968], Nakabo 2002:651 [ref. 26001], Choi et al. 2003:276, 583 [ref. 26218], Parin et al. 2002:S93 [ref. 28300], Fedorov et al. 2003:86 [ref. 27579], Nelson et al. 2004:123 [ref. 27807], Love et al. 2005:95 [ref. 37547], Kharin & Cheblukov 2005:567 [ref. 28367], Maslenikov et al. 2013:12 [ref. 37947], Page et al. 2013:126 [ref. 32708], Shinohara et al. 2014:254 [ref. 33330], Parin et al. 2014:300 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Mecklenburg et al. 2016:100 [ref. 34440], Kells et al. 2016:208 [ref. 35888], Mecklenburg et al. 2018:241 [ref. 36370], Kim et al. 2020:107 [ref. 39118], Dyldin & Orlov 2022:55 [ref. 38972], Love et al. 2021:120 [ref. 39279]). Current status: Valid as Eurymen gyrinus Gilbert & Burke 1912. Psychrolutidae: Psychrolutinae. Distribution: North Pacific and adjacent Arctic: Sea of Japan and Sea of Okhotsk north to Bering Sea and northern Chukchi Sea. Habitat: marine.

hamatus, Icelus Krøyer [H. N.] 1845:253 [Naturhistorisk Tidsskrift (Kjøbenhavn) (n. s.) v. 1; ref. 2689] Belsund, Spitsbergen. Lectotype: ZMUC P811869. Paralectotypes: ?ZMUC 10-11 (1 of 2). Type catalog: Nielsen 1974:81 [ref. 9588]. Lectotype selected by Nelson 1984:35 [ref. 5391]. •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:35 [ref. 5391], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:253 [ref. 33547]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Habitat: marine.

hankinsoni, Parartedius Hubbs [C. L.] 1926:4 [Occasional Papers of the Museum of Zoology University of Michigan No. 170; ref. 6069] Tide-pool, near the end of Point Loma, California, U.S.A. Holotype (unique): UMMZ 55001. •Valid as Artedius hankinsoni (Hubbs 1926) -- (Bolin 1944:57 [ref. 6379]). •Synonym of Artedius lateralis (Girard 1854) -- (Begle 1989:648 [ref. 12739]). Current status: Synonym of Artedius lateralis (Girard 1854). Psychrolutidae: Oligocottinae. Habitat: marine.

harringtoni, Axyrias Starks [E. C.] 1896:554, Pl. 74 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 4195] Vicinity of Port Ludlow, Washington, U.S.A. Holotype (unique): CAS-SU 5047. Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Artedius harringtoni (Starks 1896) -- (Bolin 1944:45 [ref. 6379], Eschmeyer & Herald 1983:162 [ref. 9277], Begle 1989:646 [ref. 12739], Matarese et al. 1989:416 [ref. 26885], McAllister 1990:194 [ref. 14674], Mecklenburg et al. 2002:461 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:85 [ref. 37547], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:164 [ref. 35888], Burton & Lea 2019:65 [ref. 37205], Love et al. 2021:118 [ref. 39279]). Current status: Valid as Artedius harringtoni (Starks 1896). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: eastern Aleutian Islands south to southern California (U.S.A.). Habitat: marine.

herzensteini, Gymnocanthus Jordan [D. S.] & Starks [E. C.] 1904:294, Fig. 27 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate,Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7710. Paratypes: CAS-SU 7572 (1). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Gymnocanthus herzensteini Jordan & Starks 1904 -- (Watanabe 1960:51 [ref. 11959], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:197 [ref. 15964], Kim & Youn 1992:62 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:629 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Nakabo 2002:629 [ref. 26001], Choi et al. 2003:262, 574 [ref. 26218], Parin et al. 2002:S74 [ref. 28300] as Gymnacanthus, Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:247 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Kim et al. 2020:101 [ref. 39118], Dyldin et al. 2021:63 [ref. 38804], Dyldin & Orlov 2022:44 [ref. 38972]). Current status: Valid as Gymnocanthus herzensteini Jordan & Starks 1904. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

heterospilos, Neophrynichthys Jackson [K. L.] & Nelson [J. S.] 2000:720, Figs. 1-4 [New Zealand Journal of Marine and Freshwater Research v. 34; ref. 25164] Campbell Rise, New Zealand, 51°57'S, 168°55'E, depth 262-290 meters. Holotype: NMNZ P.12096. Paratypes: NMNZ P.7024 (1), P.7133 (1), P.12015 (1), P.12130 (1), P.22065 (2), P.26798 (1), P.26802 (6), P.34974 (1), P.37144-45 [ex P.12096] (2, 3). •Valid as Neophrynichthys heterospilos Jackson & Nelson 2000 -- (Stewart 2015:1128 [ref. 34254]). Current status: Valid as Neophrynichthys heterospilos Jackson & Nelson 2000. Psychrolutidae: Psychrolutinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

hexacornis, Cottus Richardson [J.] 1823:726 [22] [Narrative of a journey to the shores of the Polar Sea; ref. 18624] Mouth of Tree River near Coppermine River, Arctic Sea about Herschel Island. On p. 22 of separate. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1129 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Mecklenburg et al. 2002:477 [ref. 25968], Parin et al. 2014:262 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

hirundo, Leiocottus Girard [C. F.] 1856:133 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1809] San Miguel Island, near Santa Barbara, California, U.S.A. Syntypes: USNM 313 (1). •Valid as Leiocottus hirundo Girard 1856 -- (Bolin 1944:86 [ref. 6379], Eschmeyer & Herald 1983:175 [ref. 9277], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Page et al. 2013:124 [ref. 32708], Kells et al. 2016:180 [ref. 35888], Love et al. 2021:122 [ref. 39279]). Current status: Valid as Leiocottus hirundo Girard 1856. Psychrolutidae: Psychrolutinae. Distribution: Northeastern Pacific: southern California (U.S.A.) south to Pacific coast of northern Baja California (U.S.A.). Habitat: marine.

hudsonius, Gymnocanthus tricuspis Vladykov [V. D.] 1933:16 [Contributions to Canadian biology and fisheries: being studies from the biological stations of Canada (Ser. A, General) v. 8 (no. 2); ref. 23244] Nottingham Island and Port Burwell, Hudson Bay region, Canada. Syntypes: (4). •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Scott & Scott 1988:496 [ref. 25518], Mecklenburg et al. 2016:62 [ref. 34440]). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Psychrolutidae: Icelinae. Habitat: marine.

humilis, Cottus Bean [T. H.] 1881:149 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Chamisso Island, Spafarief Bay at mouth of Eschscholtz Bay, Kotzebue Sound, Alaska, U.S.A., Chukchi Sea, Arctic Ocean. Holotype (unique): USNM 27972. Non-types: MNHN 1890-0465 and 0466 [ex USNM] (2) station 3242. Type catalog: Blanc & Hureau 1968:51 [ref. 20738]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Mecklenburg et al. 2016:75 [ref. 34440]). •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Neelov 1979:122 [ref. 3152], Mecklenburg et al. 2002:481 [ref. 25968], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Habitat: marine.

hypselopterus, Icelus Fukuzawa [H.], Mori [T.], Matsuzaki [K.] & Kai [Y.] 2022:Figs. [Ichthyological Research v. 70 (no. 1); ref. 38955] Shiretoko Peninsula, Hokkaido Island, Japan. Holotype. Paratypes: (3) •Valid as Icelus hypselopterus Fukuzawa, Mori, Matsuzaki & Kai 2022. Current status: Valid as Icelus hypselopterus Fukuzawa, Mori, Matsuzaki & Kai 2022. Psychrolutidae: Icelinae. Distribution: Northwest Pacific: southern Sea of Okhotsk. Habitat: marine.

ijimai, Porocottus Tanaka [S.] 1908:39 [244], Pl. 9 (figs. 3-4) [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 20; ref. 14360] Mouth of Susuya River, Sakhalin Island, Russia. Holotype: ZUMT 21700 [ex ZUMT 1388]. Paratypes: (3) FMNH 71846 [ex ZUMT 1389] (1), ZUMT (2 or 3, lost). Appeared first as above in Japanese. Then in English version in Tanaka 1908:244, Pl. 9 (figs. 3-4) [see ref. 14360]. •Synonym of Myoxocephalus platycephalus (Pallas 1814), subspecies taeniopterus (Kner 1868) -- (Berg 1949:1128 [ref. 20662]). •Valid as Porocottus ijimai Tanaka 1908 -- (Watanabe 1960:95 [ref. 11959]). •Synonym of Megalocottus platycephalus (Pallas 1814), subspecies taeniopterus (Kner 1868) -- (Neelov 1979:130 [ref. 3152]). •Synonym of Megalocottus platycephalus (Pallas 1814) -- (Parin et al. 2002:S79 [ref. 28300]). •Synonym of Megalocottus taeniopterus (Kner 1868) -- (Parin et al. 2014:258 [ref. 33547], Dyldin & Orlov 2017:64 [ref. 35138] with question). Current status: Synonym of Megalocottus taeniopterus (Kner 1868). Psychrolutidae: Myoxocephalinae. Habitat: marine.

incitus, Myoxocephalus Watanabe [M.] 1958:294, Figs. 121-122, Pl. 21 [Studies of Japanese cottid fishes; ref. 21357] Same, Aomori Prefecture, Japan, depth 10 fathoms. Holotype (unique): not researched. Also described as new in Watanabe 1960:85, Pl. 27 (fig. 1) [ref. 11959]. •Valid as Myoxocephalus incitus Watanabe 1958 -- (Yabe in Masuda et al. 1984:327 [ref. 6441]). •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Distribution: Western North Pacific. Habitat: brackish, marine.

inermis, Cottunculus Vaillant [L. L.] 1888:365, Pl. 28 (fig. 2) [Expéditions scientifiques du "Travailleur" et du "Talisman"; ref. 4496] Coast of Soudan, 22°57'N, 17°31'W, depth 930 meters depth, or "Banc d'Arguin", 20°44'N, 18°07'W, northwestern Africa, depth 1495 meters. Lectotype: MNHN 1887-0207. Paralectotypes: MNHN 1887-0208 (1) Coast of Sudan, nw. Africa; 1 from Banc d'Arguin lost. Type catalog: Blanc & Hureau 1968:50 [ref. 20738]. Lectotype selected by Eschmeyer 1998:773 [ref. 23416]. •Valid as Cottunculoides inermis (Vaillant 1888) -- (Nelson 1977:508 [ref. 7034]). •Valid as Cottunculus inermis (Vaillant 1888) -- (Yamakawa in Okamura et al. 1982:289, 402 [ref. 8256], Yabe in Masuda et al. 1984:331 [ref. 6441], Yatou in Okamura et al. 1985:603, 731 [ref. 8056], Nelson 1986:492 [ref. 5698], Nelson 1990:688 [ref. 17821], Fricke 1990:407 [ref. 15492], Nelson 1995:69 [ref. 22668]). •Valid as Psychrolutes inermis (Vaillant 1888) -- (Nelson 1999:2428 [ref. 24810], Hutchins 2001:29 [ref. 25847] as cf. inermis, Shinohara et al. 2001:320 [ref. 25995], Nelson 2016:2292 [ref. 34615], Fermon et al. 2022:296 [ref. 39448] possible). Current status: Valid as Psychrolutes inermis (Vaillant 1888). Psychrolutidae: Psychrolutinae. Distribution: Eastern Atlantic: Mauritania; southwestern Indian Ocean: South Africa and Mozambique. Habitat: marine.

ingens, Artediellus Nelson [D. W.] 1986:35, Fig. 7 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 138 (no. 1); ref. 5808] Cape Rollin, Simushir Island, Kurils. [Kuril], Albatross station 4804, 46°42'N, 151°47'E, depth 429 meters. Holotype: USNM 74460. Paratypes: CAS-SU 22237 (4); USNM 74484-85 (13, 20). •Valid as Artediellus ingens Nelson 1986 -- (Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:404 [ref. 25968], Parin et al. 2002:S70 [ref. 28300], Parin et al. 2014:237 [ref. 33547]). Current status: Valid as Artediellus ingens Nelson 1986. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

intermedius, Cottus Temminck [C. J.] & Schlegel [H.] 1843:38 [Fauna Japonica Parts 2-4; ref. 4371] Coast of Yezo, Japan. Holotype: RMNH D652 (dry skin, poor condition). Type information: Boeseman 1947:48 [ref. 12876]. •Valid as Gymnocanthus intermedius (Temminck & Schlegel 1843) -- (Watanabe 1960:54 [ref. 11959], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:198 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:629 [ref. 25086], Nakabo 2002:629 [ref. 26001], Choi et al. 2003:262, 575 [ref. 26218], Parin et al. 2002:S74 [ref. 28300] as Gymnacanthus, Mundy 2005:336 [ref. 28379], Shinohara et al. 2014:249 [ref. 33330], Kim et al. 2020:101 [ref. 39118], Dyldin & Orlov 2022:44 [ref. 38972]). Current status: Valid as Gymnocanthus intermedius (Temminck & Schlegel 1843). Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

intermedius, Icelus spiniger Lindberg [G. U.] & Andriashev [A. P.] 1938:517, Figs. 1-2 [Trudy Gidrobiologicheskoi Ekspeditsii Zoologicheskogo instituta AN SSSR No. 1; ref. 19731] Okhotsk Sea, southeastern Kamchatka, Russia, 52°33.3'N, 154°54'E, depth 185-195 meters. Holotype: ZIN 24470 (185 mm). Paratypes: ZIN 24470 (1, with holotype), 27962 (2), 27969 (3). Appeared first as a nomen nudum in Andriashev 1937:301[not researched]. •Synonym of Icelus spiniger Gilbert 1896 -- (Nelson 1984:24 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:257 [ref. 33547]). Current status: Synonym of Icelus spiniger Gilbert 1896. Psychrolutidae: Icelinae. Habitat: marine.

ishikawae, Furcina Jordan [D. S.] & Starks [E. C.] 1904:303, Fig. 32 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Rikuchu, Myiako, Japan. Holotype: CAS-SU 7715. Paratypes: AMS I.6953 (6); BMNH 1905.2.4.365-372 (8?); CAS 31363 [ex IU 10719] (3, c&s), 58402 [ex IU 10719] (6); CAS-SU 7580 (18), 7582 (3, dry), 7587 (4); USNM 50924 (32), 74788 (1). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Furcina ishikawae Jordan & Starks 1904 -- (Watanabe 1960:137 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:244 [ref. 15964], Kim & Youn 1992:72 [ref. 22287], Nakabo 2000:648 [ref. 25086], Shimizu 2001:27 [ref. 25799], Nakabo 2002:648 [ref. 26001], Choi et al. 2003:262, 574 [ref. 26218], Kim et al. 2020:101 [ref. 39118], Sonoyama et al. 2020:89 [ref. 37637]). Current status: Valid as Furcina ishikawae Jordan & Starks 1904. Psychrolutidae: Oligocottinae. Distribution: Northwestern Pacific. Habitat: marine.

islandicus, Triglops pingelii Jensen [A. S.] 1944:16, 29 [Spolia Zoologica Musei Hauniensis v. 4; ref. 13060] Iceland. Syntypes: ZMUC 56 (1), 158 (1), P811667-735 (69). Type catalog: Nielsen 1974:82 [ref. 9588]. •Synonym of Triglops murrayi Günther 1888 -- (Pietsch 1994:369 [ref. 21189], Parin et al. 2014:276 [ref. 33547]). Current status: Synonym of Triglops murrayi Günther 1888. Psychrolutidae: Triglopinae. Habitat: marine.

jaok, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:172 [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka coast, Russia. Holotype (unique): ZMB 31656. Type catalog: Blanc & Hureau 1968:60 [ref. 20738], Paepke & Fricke 1992:284 [ref. 19946]. •Valid as Myoxocephalus jaok (Cuvier 1829) -- (Watanabe 1960:75 [ref. 11959], Neelov 1979:122 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:212 [ref. 15964], Kim & Youn 1992:64 [ref. 22287], Sokolovsky & Sokolovskaya 1997:55 [ref. 24691], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:645 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:481 [ref. 25968], Nakabo 2002:645 [ref. 26001], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:75 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Bogutskaya & Naseka 2004:190 [ref. 28183], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:261 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Tohkairin et al. 2015:7 [ref. 35331], Mecklenburg et al. 2016:76 [ref. 34440], Kells et al. 2016:182 [ref. 35888], Dyldin & Orlov 2017:65 [ref. 35138], Mecklenburg et al. 2018:213 [ref. 36370], Dyldin et al. 2021:66 [ref. 38804], Love et al. 2021:123 [ref. 39279], Dyldin & Orlov 2022:48 [ref. 38972]). Current status: Valid as Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific and adjacent Arctic: northern Sea of Japan and Sea of Okhotsk north to Bering Sea and Chukchi Sea, east to eastern Gulf of Alaska (U.S.A.). Habitat: brackish, marine.

japonicus, Centridermichthys Steindachner [F.] 1870:625 [3], Pl. 1 (fig. 3) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 61 (1. Abth.); ref. 4218] Nagasaki, Japan. Holotype (unique): NMW. On p. 3 of separate. Also described and figured as a new species in Steindachner 1881:187 [9 of separate] [ref. 4231]; 1881 description is based on a different specimen. •Synonym of Pseudoblennius cottoides (Richardson 1848) -- (Watanabe 1960:152 [ref. 11959], Lindberg & Krasyukova 1987:252 [ref. 15964], Kim & Youn 1992:74 [ref. 22287]). Current status: Synonym of Pseudoblennius cottoides (Richardson 1848). Psychrolutidae: Psychrolutinae. Habitat: marine.

japonicus, Dasycottus Tanaka [S.] 1914:308, Pl. 83 (fig. 280) [Figures and descriptions of the fishes of Japan v. 18; ref. 14935] Near Niigata, Echigo Province, Japan. Holotype (unique): ZUMT 3916. Plate is in vol. 17 [ref. 14934]. •Valid as Dasycottus japonicus Tanaka 1914 -- (Watanabe 1960:177 [ref. 11959], Lindberg & Krasyukova 1987:270 [ref. 15964]). •Synonym of Dasycottus setiger Bean 1890 -- (Parin et al. 2002:S92 [ref. 28300], Parin et al. 2014:299 [ref. 33547]). Current status: Synonym of Dasycottus setiger Bean 1890. Psychrolutidae: Psychrolutinae. Habitat: marine.

japonicus, Icelinus Yabe [M.], Tsumura [K.] & Katayama [M.] 1980:106, Figs. 1-2 [Japanese Journal of Ichthyology v. 27 (no. 2); ref. 8640] Mano Bay, off Sado Island, Niigata Prefecture, Japan, 37°57'N, 138°19'E. Holotype: HUMZ 79035. Paratypes: HUMZ 75466 (1), 77562 (1), 77563 (1, dissected); NSMT-P 18575 (1). Other material: HUMZ 77561 (1). •Valid as Icelinus japonicus Yabe, Tsumura & Katayama 1980 -- (Yabe in Masuda et al. 1984:325 [ref. 6441], Peden 1984:79 [ref. 5193], Suzuki & Uno 1992:4 [ref. 24139], Nakabo 2000:636 [ref. 25086], Nakabo 2002:636 [ref. 26001], Rosenblatt & Smith 2004:560 [ref. 27912]). Current status: Valid as Icelinus japonicus Yabe, Tsumura & Katayama 1980. Psychrolutidae: Oligocottinae. Distribution: Western North Pacific. Habitat: marine.

japonicus, Porocottus Schmidt [P. Yu.] in Taranetz 1935:177 [Vestnik Dal'nevostocnogo Filiala Akademii Nauk SSSR = Bulletin of the Far Eastern Branch of the Academy of Sciences of the USSR No. 14; ref. 21496] DeCastri Bay and western coast of Sakhalin. Lectotype: ZIN 26314 or 6314. Paralectotypes: ZIN 18962. Lectotype designation not researched, not designated here. •Valid as Porocottus japonicus Schmidt 1935 -- (Neelov 1979:137 [ref. 3152], Lindberg & Krasyukova 1987:221 [ref. 15964], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S84 [ref. 28300], Parin et al. 2014:268 [ref. 33547], Dyldin & Orlov 2017:67 [ref. 35138], Dyldin et al. 2021:67 [ref. 38804], Dyldin & Orlov 2022:50 [ref. 38972]). Current status: Valid as Porocottus japonicus Schmidt 1935. Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific. Habitat: brackish, marine.

japonicus, Zesticelus Oshima [M.] 1957:2, Fig. 1 [Japanese Journal of Ichthyology v. 6 (nos 1-2); ref. 12161] Offshore of Niigata, Japan. Holotype (unique): Japan Sea Reg. Fish. Lab. Niigata No. 1335 (lost). •Valid as Zesticelus japonicus Oshima 1957 -- (Neelov 1979:161 [ref. 3152], Yabe 1995:19 [ref. 22666]). •Mention as Zeticelus japonicus Oshima 1957 -- (Yabe & Okamura 1993:370 [ref. 22281]). •Status uncertain -- (Nakabo 2002:1526 [ref. 26001]). Current status: Uncertain as Zesticelus japonicus Oshima 1957. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

jordani, Elanura Schmidt [P. Yu.] 1903:518 [16] [Izvestiya Imperatorskago Russkago Geograficheskago Obshchestva v. 38 (pt 5) (for 1902); ref. 3945] On p. 16 of separate. Not available, name and localities but no description. See Prionistius jordani Schmidt in Jordan & Starks 1904. Invalid lectotype designation by Pietsch 1994:351 [ref. 21189] as SU 7704. •Treated as valid as Elanura jordani Schmidt 1903 -- (Kim & Youn 1992:66 [ref. 22287]). •Treated as valid as Triglops jordani (Schmidt 1904) -- (Yabe in Masuda et al. 1984:325 [ref. 6441], Pietsch 1994:351 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Nakabo 2000:628 [ref. 25086], Mecklenburg et al. 2002:422 [ref. 25968], Nakabo 2002:628 [ref. 26001], Choi et al. 2003:263, 578 [ref. 26218], Shinohara et al. 2011:44 [ref. 31715]). Nomen Nudum. Current status: Synonym of Triglops jordani (Schmidt 1904). Psychrolutidae: Triglopinae.

jordani, Prionistius Schmidt [P. Yu.] in Jordan & Starks 1904:252, Fig. 9 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Peter the Great Bay, Vladivostok, Russia. Holotype: CAS-SU 7704 [from Schmidt]. Type catalog: Böhlke 1953:131 [ref. 12291]. Appeared first without distinguishing features in Schmidt 1903:518 (p. 16 of separate) [ref. 3945]. Appeared in 1904 in Jordan & Starks as above and as Prionistius jordani, with the description based solely on one "cotype" received from Schmidt; this cotype is the holotype and authorship is Jordan & Starks as above. Pietsch 1994:351 [ref. 21189] regarded Schmidt 1903 as the original description and named SU 7704 as the lectotype. •Valid as Triglops jordani (Jordan & Starks 1904) -- (Lindberg & Krasyukova 1987:177 [ref. 15964], Pietsch 1994:351 [ref. 21189], Sokolovskaya et al. 1998:10 [ref. 24670], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:422 [ref. 25968], Parin et al. 2002:S87 [ref. 28300], Fedorov et al. 2003:81 [ref. 27579], Love et al. 2005:93 [ref. 37547], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:275 [ref. 33547], Dyldin & Orlov 2017:68 [ref. 35138], Kim et al. 2020:105 [ref. 39118], Dyldin et al. 2021:69 [ref. 38804], Love et al. 2021:126 [ref. 39279], Dyldin & Orlov 2022:52 [ref. 38972]). Current status: Valid as Triglops jordani (Schmidt 1904). Psychrolutidae: Triglopinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia) and Sea of Okhotsk north to southern Kamchatka (Russia). Habitat: brackish, marine.

kallavesensis, Cottus quadricornis race Lönnberg [E.] 1933:7, Pl. 1 (fig. 1) [Arkiv för Zoologi v. 24 A (pt 3) (no. 12) [1932]; ref. 22149] Lake Kallavesi; Lake Niinivesi in Rautalampi; Lake Virmavesi in Karttula, Lake Päijänne basin, Finland. Syntypes: MZH (9). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies onegensis Berg & Popov 1932 -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

karaensis, Icelus Soldatov [V. K.] 1923:31 [English p. 74], Fig. 34 [Russkij gidrobiologiceskij zurnal: organ Obscestva Issledovatelej Vody i ee Zizni = Russische hydrobiologische Zeitschrift - Saratow v. 3; ref. 15569] Kara Sea, 70°08'N, 56°26'E, Russia. Syntypes: ZIN 22027 (3). •Synonym of Icelus spatula Gilbert & Burke 1912 -- (Neelov 1973:597 [ref. 7219], Nelson 1984:42 [ref. 5391], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:256 [ref. 33547]). Current status: Synonym of Icelus spatula Gilbert & Burke 1912. Psychrolutidae: Icelinae. Habitat: marine.

kincaidi, Malacocottus Gilbert [C. H.] & Thompson [J. C.] 1905:979, Fig. 2 [Proceedings of the United States National Museum v. 28 (no. 1414); ref. 1640] Brinnon, Hood's Canal, Washington, U.S.A. Holotype: USNM 53028. Additional material or paratypes: CAS-SU 21406 (10). See Shinohara et al. 1992:422 [ref. 19776]. •Synonym of Malacocottus zonurus Bean 1890 -- (Parin et al. 2002:S94 [ref. 28300]). •Valid as Malacocottus kincaidi Gilbert & Thompson 1905 -- (Watanabe 1960:183 [ref. 11959], Eschmeyer & Herald 1983:175 [ref. 9277], Yabe in Amaoka et al. 1983:157, 215 [ref. 8379], McAllister 1990:202 [ref. 14674], Mecklenburg et al. 2002:521 [ref. 25968], Nelson et al. 2004:123 [ref. 27807], Love et al. 2005:95 [ref. 37547], Page et al. 2013:126 [ref. 32708], Stevenson 2015:26 [ref. 33680], Pietsch & Orr 2015:50 [ref. 34694], Kells et al. 2016:208 [ref. 35888], Love et al. 2021:123 [ref. 39279]). Current status: Valid as Malacocottus kincaidi Gilbert & Thompson 1905. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: southern British Columbia (Canada) south to Oregon (U.S.A.). Habitat: marine.

konstantinovi, Cottunculus Myagkov [N. A.] 1991:37 [Byulleten Moskovskogo Obshchestva Ispytatelei Prirody Otdel Biologicheskii [Bulletin of the Moscow Society of Naturalists Biological Series] v. 96 (no. 3); ref. 20068] Spitsbergen, 80°26'N, 13°03'E, Arctic Ocean, depth 300 meters. Holotype (unique): ZMMU P-16261. Type catalog: Vasil'eva et al. 2001:80 [ref. 25714]. •Questionably a synonym of Cottunculus microps Collett 1875 -- (Parin et al. 2002:S92 [ref. 28300]). •Valid as Cottunculus konstantinovi Myagkov 1991 -- (Byrkjedal & Orlov 2007:67 [ref. 29312], Mecklenburg et al. 2011:124, 125 [ref. 31212] but needing further study). •Synonym of Cottunculus microps Collett 1875 -- (Byrkjedal et al. 2014:14 [ref. 33415], Mecklenburg et al. 2014:137 [ref. 33463], Parin et al. 2014:298 [ref. 33547], Mecklenburg et al. 2016:99 [ref. 34440]). Current status: Synonym of Cottunculus microps Collett 1875. Psychrolutidae: Psychrolutinae. Distribution: Northern Spitsbergen (unique holotype) (if valid). Habitat: marine.

krawchukii, Myoxocephalus quadricornis Mikhalev [Y. V.] 1962:138 [Not found yet; ref. 40286] Reference not traced. •Probably a synonym of Myoxocephalus quadricornis (Linnaeus 1758). Current status: Uncertain as Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish.

labradoricus, Acanthocottus Girard [C. F.] in Storer 1850:247, Pl. 7 (fig. 3) [Boston Journal of Natural History v. 6 (no. 2) (art. 15); ref. 18109] Lost. No types known. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1130 [ref. 20662]). •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Neelov 1973:599 [ref. 7219], Neelov 1979:122 [ref. 3152]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

lateralis, Scorpaenichthys Girard [C. F.] 1854:145 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 5769] Monterey and San Luis Obispo, California, U.S.A. Syntypes: USNM 328 (1). •Valid as Artedius lateralis (Girard 1854) -- (Bolin 1944:55 [ref. 6379], Eschmeyer & Herald 1983:162 [ref. 9277], Begle 1989:648 [ref. 12739], Matarese et al. 1989:418 [ref. 26885], McAllister 1990:194 [ref. 14674], Mecklenburg et al. 2002:459 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:85 [ref. 37547], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:164 [ref. 35888], Burton & Lea 2019:65 [ref. 37205], Love et al. 2021:118 [ref. 39279]). Current status: Valid as Artedius lateralis (Girard 1854). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: western Gulf of Alaska south to northern Baja California (Mexico). Habitat: marine.

laticeps, Acanthocottus Gilbert [C. H.] 1896:422, Pls. 26-27 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Nushagak River [lectotype] and Herendeen Bay, Alaska, U.S.A. Lectotype. Paralectotypes (incl. potential lectotype): BMNH 1896.4.23.48 (1), MCZ 28218 (1), CAS-SU 3016 (4), USNM 48651 (1), ZMUO J1399 (1) station 3232. Type catalog: Böhlke 1953:126 [ref. 12291], Pethon 1969:7 [ref. 19268]. Lectotype established (as figured specimen) in caption to Pl. 299, p. 3293 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable •Synonym of Megalocottus platycephalus (Pallas 1814) -- (Neelov 1979:130 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:475 [ref. 25968], Parin et al. 2002:S79 [ref. 28300], Parin et al. 2014:258 [ref. 33547], Mecklenburg et al. 2016:73 [ref. 34440]). Current status: Synonym of Megalocottus platycephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

latifrons, Cottus Gratzianov [V. I.] 1904:195, Pl. 11 [Zapiski Imperatorskago russkago geograficheskago obshchestva po obshchei geografii v. 41 (no. 1); ref. 21411] Western coast Kanin Peninsula, northern Russia, Barents Sea. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies labradoricus (Girard 1850) -- (Berg 1949:1130 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952], Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Neelov 1973:597 [ref. 7219], Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

latifrons, Stelgidonotus Gilbert [C. H.] & Thompson [J. C.] 1905:977, Fig. 1 [Proceedings of the United States National Museum v. 28 (no. 1414); ref. 1640] Friday Harbor, Washington, U.S.A. Holotype (unique): USNM 53027. •Synonym of Oligocottus rimensis (Greeley 1899) -- (Bolin 1944:63 [ref. 6379], Mecklenburg et al. 2002:499 [ref. 25968]). Current status: Synonym of Oligocottus rimensis (Greeley 1899). Psychrolutidae: Oligocottinae. Habitat: marine.

latus, Psychrolutes Hutton [F. W.] 1875:316 [Annals and Magazine of Natural History (Series 4) v. 16 (no. 95) (art. 41); ref. 18317] Dunedin Harbor, New Zealand. Lectotype: BMNH 1886.11.18.55. Type catalog: Russell 1996:226 [ref. 22562] with a holotype, but second locality (Bluff Harbour) indicates that additional specimens may have contributed to Hutton's understanding of the species. Lectotype designated by Eschmeyer 1998:881 [ref. 23416]. Also in Hutton 1876:214 [ref. 14210]. Lectotype selected by Eschmeyer 1998:881 [ref. 23416]. •Valid as Neophrynichthys latus (Hutton 1875) -- (Nelson 1977:505 [ref. 7034], Paulin et al. 1989:172 [ref. 24556], Yabe in Amaoka et al. 1990:247 [ref. 16024], Jackson & Nelson 2000:719 [ref. 25146], Stewart 2015:1129 [ref. 34254]). Current status: Valid as Neophrynichthys latus (Hutton 1875). Psychrolutidae: Psychrolutinae. Distribution: New Zealand region. Habitat: marine.

leprops, Astrocottus Bolin [R. L.] 1936:331, Pl. 34 (fig. B); Fig. 27 [Proceedings of the United States National Museum v. 83 (no. 2987); ref. 504] Tsugaru Strait, Japan, 41°35'50"N, 140°36'45"E, Albatross station 4808, depth 47 fathoms. Holotype (unique): USNM 94730. •Valid as Astrocottus leprops Bolin 1936 -- (Yabe in Masuda et al. 1984:325 [ref. 6441], Shinohara et al. 1996:176 [ref. 26605], Nakabo 2000:632 [ref. 25086], Yabe & Maruyama 2001:61 [ref. 25245], Nakabo 2002:632 [ref. 26001], Tsuruoka et al. 2008:27 [ref. 29568]). Current status: Valid as Astrocottus leprops Bolin 1936. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

leptosomus, Porocottus Muto [F.], Choi [Y.] & Yabe [M.] 2002:229, Figs. 1-2 [Ichthyological Research v. 49 (no. 3); ref. 26331] Mohang, Taean, Chungnam, western coast of Korea, 36°47'N, 126°10'E, depth 0.2 meters. Holotype: HUMZ 172394. Paratypes: HUMZ 153700-04 (1, 1, 1, 1, 1), 172395-96 (1, 1), 172399 (1); KNUM 1051-52 (1, 1); NSMT-P 61563 (1). •Valid as Porocottus leptosomus Muto, Choi & Yabe 2002 -- (Kim et al. 2020:103 [ref. 39118]). Current status: Valid as Porocottus leptosomus Muto, Choi & Yabe 2002. Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific: Yellow Sea, Sea of Japan (Korea). Habitat: marine.

lilljeborgii, Cottus Collett [R.] 1875:25, Pl. 1 (figs. 4-5) [Norges fiske; ref. 884] Western Norway. Syntypes: ZMUO J4163 (1) Kristiansund, J4160 (1) Hardangerfjord. Type catalog: Pethon 1969:8 [ref. 19268]. •Valid as Enophrys lilljeborgi [sic] (Collett 1875) -- (Sandercock & Wilimovsky 1968:839 [ref. 26414]). •Valid as Tautulus lilljeborgi [sic] (Collett 1875) -- (Neelov 1973:600 [ref. 7219]). •Valid as Micrenophrys lilljeborgii [sometimes as lilljiborgi] (Collett 1875) -- (Neelov 1979:150 [ref. 3152], Fedorov in Whitehead et al. 1986:1252 [ref. 13677], Parin et al. 2002:S80 [ref. 28300], Fricke 2007:29 [ref. 30577], Parin et al. 2014:259 [ref. 33547], Iglésias et al. 2020:294 [ref. 37987]). Current status: Valid as Micrenophrys lilljeborgii (Collett 1875). Psychrolutidae: Psychrolutinae. Distribution: Western Baltic Sea, North Sea, eastern North Atlantic, Arctic Ocean. Habitat: brackish, marine.

limbaughi, Icelinus Rosenblatt [R. H.] & Smith [W. L.] 2004:556, Fig. 1 [Copeia 2004 (no. 3); ref. 27912] La Jolla Submarine Canyon, California, U.S.A., 32°53.0'N, 117°15.0'W, depth 40 meters. Holotype: SIO 62-628. Paratypes: AMNH 233608 [ex SIO 58-510] (2, 1 c&s); CAS 218345 [ex SIO 62-634] (1); SIO 51-247 (1), 51-252 (1), 51-253 (2), 54-112 (18), 58-508 (1), 58-509 (2), 62-632 (1), 62-635 (1), 62-673 (1), 63-239-55A (1), 63-1024-55B (1), 74-164 (1); USNM 374474 [ex SIO 62-636] (1). Plus non-type material. •Valid as Icelinus limbaughi Rosenblatt & Smith 2004 -- (Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Girard & Smith 2016:556 [ref. 34786], Love et al. 2021:121 [ref. 39279]). Current status: Valid as Icelinus limbaughi Rosenblatt & Smith 2004. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: southern California (U.S.A.) endemic. Habitat: marine.

longipinnis, Dasycottus Mori [T.] 1956:16, Fig. 2 [Japanese Journal of Ichthyology v. 5 (nos 1-2); ref. 12213] Off Oki Island, Japan Sea. Holotype (unique): Hyogo Univ. Agric. Psychrolutidae: Psychrolutinae. Habitat: marine.

lonnbergi, Cottus quadricornis Lönnberg [E.] 1919:167, 171 [Fauna och Flora (Uppsala) v. 14 (no. 5); ref. 23061] Lake Mälaren, Sweden. Syntypes: ?NRM. Type catalog: Sideleva & Zhidkov 2021:240 [ref. 38705]. Name should not be changed since it is Swedish (see Koerber 2009:63 [ref. 30671] for spelling of Lönnberg and origin of his name). •Validates the infrasubspecific Myoxocephalus quadricornis morpha lonnbergi Berg 1916. •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Sideleva & Zhidkov 2021:240 [ref. 38705]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

lonnbergi, Myoxocephalus quadricornis morpha Berg [L. S.] 1916:431, Fig. 318 [Les Poissons des eaux douces de la Russie; ref. 277] Lake Ladoga, Russia; lake Mälaren, Sweden. No types known. Original as lönnbergi; should not be changed to loennbergi since the name is not German, but Swedish (see Koerber 2009:63 [ref. 30671] for spelling of Lönnberg and origin of his name.). Not available, as a morpha, infrasubspecific; dates to Lönnberg 1919 or to a later publication by Berg. •In the synonymy of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies as described -- (Berg 1949:1133 [ref. 20662]). •In the synonymy of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Infrasubspecific. Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

lucasi, Ceratocottus Jordan [D. S.] & Gilbert [C. H.] in Jordan & Evermann 1898:1940 [Bulletin of the United States National Museum No. 47; ref. 2444] Near St. Paul Island, Bering Sea (stomach content). Holotype: USNM 48234. Paratypes: CAS-SU 5661 (1), USNM 48235 (1). Type catalog: Böhlke 1953:127 [ref. 12291]. Also appeared in Jordan & Gilbert 1899:459, Pl. 57 [ref. 2478]. •Synonym of Enophrys diceraus (Pallas 1788) -- (Neelov 1979:146 [ref. 3152], Sheiko & Fedorov 2000:27 [ref. 25650], Parin et al. 2002:S74 [ref. 28300]). •Valid as Enophrys lucasi (Jordan & Gilbert 1898) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414], McAllister 1990:197 [ref. 14674], Mecklenburg et al. 2002:473 [ref. 25968], Nelson et al. 2004:119 [ref. 27807], Love et al. 2005:86 [ref. 37547], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:246 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Mecklenburg et al. 2016:55 [ref. 34440], Kells et al. 2016:170 [ref. 35888], Mecklenburg et al. 2018:194 [ref. 36370], Love et al. 2021:120 [ref. 39279]). Current status: Valid as Enophrys lucasi (Jordan & Gilbert 1898). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific: Bering Sea south to northern British Columbia (Canada). Habitat: marine.

macellus, Prionistius Bean [T. H.] 1884:355 [Proceedings of the United States National Museum v. 6 (no. 387); ref. 225] Carter Bay, British Columbia, Canada. Holotype: USNM 31958. Paratypes: USNM 33793 (1). •Valid as Triglops macellus (Bean 1884) -- (Eschmeyer & Herald 1983:184 [ref. 9277], McAllister 1990:202 [ref. 14674], Pietsch 1994:347 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:423 [ref. 25968], Parin et al. 2002:S88 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:93 [ref. 37547], Maslenikov et al. 2013:11 [ref. 37947], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:276 [ref. 33547], Pietsch & Orr 2015:45 [ref. 34694], Kells et al. 2016:192 [ref. 35888], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Triglops macellus (Bean 1884). Psychrolutidae: Triglopinae. Distribution: North Pacific: northeastern Bering Sea, Alaska, to Aleutian Islands, then south to Washington (U.S.A.). Habitat: marine.

macouni, Pterygiocottus Bean [B. A.] & Weed [A. C.] 1920:73, Pl. 3 [Transactions of the Royal Society of Canada (Series 3) v. 13 (sect. 5) (1919); ref. 221] Ucluelet, west coast of Vancouver Island, British Columbia, Canada. Holotype (unique): USNM 82169. •Synonym of Artedius harringtoni (Starks 1896) -- (Bolin 1944:45 [ref. 6379]). Current status: Synonym of Artedius harringtoni (Starks 1896). Psychrolutidae: Oligocottinae. Habitat: marine.

macquariensis, Ebinania Nelson [J. S.] 1982:1485, Figs. 10-11 [Canadian Journal of Zoology v. 60 (no. 6); ref. 5470] Off northeastern Macquarie Island, south of Tasmania, southern Pacific, midpoint 54°31.5'S, 159°01.5'E, depth 494-714 meters. Holotype (unique): LACM 10984-1. •Valid as Ebinania macquariensis Nelson 1982 -- (, Duhamel et al. 2005:298 [ref. 32059], Jackson & Nelson 2006:42 [ref. 28707], Paxton et al. 2006:954 [ref. 28994]). Current status: Valid as Ebinania macquariensis Nelson 1982. Psychrolutidae: Psychrolutinae. Distribution: Southern Ocean: Macquarie Island and Kerguelen Plateau. Habitat: marine.

macrocephalus, Cottunculus Gilchrist [J. D. F.] 1904:7, Pl. 24 [Marine Investigations in South Africa v. 3; ref. 1646] Off Lion's Head, South Africa, depth 230 and 190 fathoms. Syntypes: ?BMNH 1904.3.4.2 (1), SAM 15323 (1) Pieter Faure station 15149, depth 190 fathoms. •Valid as Cottunculoides macrocephalus (Gilchrist 1904) -- (Nelson 1977:508 [ref. 7034]). •Valid as Psychrolutes macrocephalus (Gilchrist 1904) -- (Nelson 1986:492 [ref. 5698], Nelson 1995:69 [ref. 22668], Nelson 1999:2428 [ref. 24810], Nakabo 2000:653 [ref. 25086], Nakabo 2002:653 [ref. 26001], Shinohara et al. 2005:428 [ref. 28370], Nelson 2016:2291 [ref. 34615]). Current status: Valid as Psychrolutes macrocephalus (Gilchrist 1904). Psychrolutidae: Psychrolutinae. Distribution: South Africa and western Pacific. Habitat: marine.

maculatus, Cottus Fischer [J. G.] 1885:78, Pl. 2 (fig. 8) [Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten v. 2; ref. 18084] Reported as Barbados, but that locality in error. Holotype (unique): ZMH H138 [ex ZMH 3523]. Non-types: ZMH H190 [ex 3523] (1). Type catalog: Ladiges et al. 1958:167 [ref. 19181], lkens & Dohse 1993:408 [ref. 21161] with 138 as holotype. Lectotype selected by Ladiges et al. 1958:167 [ref. 19181] but original mentions only one specimen. •Synonym of Enophrys bubalis (Euphrasen 1786) -- (Sandercock & Wilimovsky 1968:842 [ref. 26414]). •Synonym of Taurulus bubalis (Euphrasen 1786) -- (Neelov 1973:600 [ref. 7219], Neelov 1979:149 [ref. 3152]). Current status: Synonym of Taurulus bubalis (Euphrasen 1786). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

maculosus, Oligocottus Girard [C. F.] 1856:133 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1809] Tomales Bay; San Francisco, California, U.S.A. Syntypes: MCZ 13641 (2); USNM 297 (4, not found in 1980), 298 (6). Oligocottus alaskanus Miles 1918:93 was a lapsus calami for Oligocottus maculosus (Bolin 1944:65 [ref. 6379]). •Valid as Oligocottus maculosus Girard 1856 -- (Herrera 1896:64 [ref. 39336], Bolin 1944:65 [ref. 6379], Eschmeyer & Herald 1983:178 [ref. 9277], Yabe in Masuda et al. 1984:328 [ref. 6441], Matarese et al. 1989:430 [ref. 26885], McAllister 1990:201 [ref. 14674], Nakabo 2000:646 [ref. 25086], Mecklenburg et al. 2002:500 [ref. 25968], Nakabo 2002:646, 1526 [ref. 26001], Parin et al. 2002:S83 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:267 [ref. 33547], Pietsch & Orr 2015:43 [ref. 34694], Kells et al. 2016:186 [ref. 35888], Burton & Lea 2019:67 [ref. 37205], Love et al. 2021:124 [ref. 39279]). Current status: Valid as Oligocottus maculosus Girard 1856. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: southeastern Bering Sea south to southern California (U.S.A.). Habitat: marine.

magnicirrus, Neophrynichthys Nelson [J. S.] 1977:497, Fig. 7 [Journal of the Royal Society of New Zealand v. 7 (no. 4); ref. 7034] 2 kilometers east of the northeastern coast of Macquarie Island, 54°30'-34'S, 158°59'E, south of Tasmania. Holotype: LACM 11440-1. Paratypes: LACM 10983-2 (3). •Valid as Ambophthalmos magnicirrus (Nelson 1977) -- (Jackson & Nelson 1998:1355 [ref. 23806], Jackson & Nelson 1999:428 [ref. 23799], Stewart 2015:1125 [ref. 34254]). Current status: Valid as Ambophthalmos magnicirrus (Nelson 1977). Psychrolutidae: Psychrolutinae. Distribution: New Zealand region. Habitat: marine.

malacocephala, Ebinania Nelson [J. S.] 1982:1485, Figs. 12-13 [Canadian Journal of Zoology v. 60 (no. 6); ref. 5470] About halfway between Pitcairn Island and the South Pole, midpoint of haul 54°49.5'S, 129°47'W, depth maximum 603 meters. Holotype (unique): LACM 10936-2. •Valid as Ebinania malacocephala Nelson 1982 -- (Jackson & Nelson 2006:42 [ref. 28707]). Current status: Valid as Ebinania malacocephala Nelson 1982. Psychrolutidae: Psychrolutinae. Distribution: South Pacific. Habitat: marine.

mandibularis, Icelus Yabe [M.] in Amaoka et al. 1983:283, Pl. 66; Figs. 28-30 [Fishes from the north-eastern Sea of Japan and the Okhotsk Sea off Hokkaido; ref. 8379] Okhotsk Sea, 44°47.0'N, 144°26.5'E, depth 905 meters. Holotype: HMMZ 78676. Paratypes: HUMZ 78675 (1), 78677-78 (2). •Valid as Icelus mandibularis Yabe 1983 -- (Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:634 [ref. 25086], Nakabo 2002:634 [ref. 26001], Tsuruoka et al. 2006:51 [ref. 28501]). Current status: Valid as Icelus mandibularis Yabe 1983. Psychrolutidae: Icelinae. Distribution: Western North Pacific. Habitat: marine.

marcidus, Neophrynichthys McCulloch [A. R.] 1926:215, Pl. 55 (fig. 2) [Biological Results Endeavour v. 5 (pt 4); ref. 2947] Deep water off southern Australia or Tasmania, Australia. Holotype (unique): AMS E.6570. See Paulin et al. 1989:259 [ref. 24556]. •Valid as Neophrynichthys marcidus McCulloch 1926 -- (Nelson 1977:504 [ref. 7034]). •Valid as Psychrolutes marcidus (McCulloch 1926) -- (Paxton et al. 1989:478 [ref. 12442], Fricke 1990:407 [ref. 15492], Yabe in Amaoka et al. 1990:248 [ref. 16024], Gomon et al. 1994:527 [ref. 22532], Nelson 1995:69 [ref. 22668], Nelson 1999:2428 [ref. 24810], Hutchins 2001:29 [ref. 25847] as cf. marcidus, Paxton et al. 2006:954 [ref. 28994], Gomon 2008:524 [ref. 30616]). Current status: Valid as Psychrolutes marcidus (McCulloch 1926). Psychrolutidae: Psychrolutinae. Distribution: Australia: southern Queensland to southern Australia and Western Australia. Habitat: marine.

marmoratus, Centridermichthys Döderlein [L.] in Steindachner & Döderlein 1884:210 [42] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 49 (1. Abth.); ref. 4248] Tokyo, Japan. Holotype (unique): NMW 92827. •Valid as Pseudoblennius marmoratus (Döderlein 1884) -- (Watanabe 1960:150 [ref. 11959] as marumoratus, Nakabo in Masuda et al. 1984:330 [ref. 6441], Kimura et al. 1988:19 [ref. 6692], Kim & Youn 1992:73 [ref. 22287], Nakabo 2000:650 [ref. 25086], Nakabo 2002:650 [ref. 26001], Choi et al. 2003:263, 577 [ref. 26218], Yoshigou 2019:200 [ref. 36714], Kwun et al. 2017:141 [ref. 37287], Kim et al. 2020:103 [ref. 39118]). Current status: Valid as Pseudoblennius marmoratus (Döderlein 1884). Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Korea and Japan. Habitat: marine.

marmoratus, Cottus Valenciennes [A.] in Cuvier & Valenciennes 1832:497 [Histoire naturelle des poissons; ref. 1000] Kamchatka, Russia. Type catalog: Blanc & Hureau 1968:61 [ref. 20738]. •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:265 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

marmoratus, Neophrynichthys Gill [T. N.] 1889:327 [Proceedings of the United States National Museum v. 11 (no. 717); ref. 19995] Coast of Patagonia, 53°31'S, 73°44'W. Holotype (unique): BMNH 1880.7.28.5. Based on Neophrynichthys latus (in part) of Günther 1881:20, pl. 1 [ref. 2012]. •Valid as Neophrynichthys marmoratus Gill 1889 -- (Nelson 1977:501 [ref. 7034], Nakamura in Nakamura et al. 1986:188 [ref. 14235], Pequeño 1989:57 [ref. 14125]). •Valid as Psychrolutes marmoratus (Gill 1889) -- (Nelson 1995:69 [ref. 22668], Mabragaña et al. 2011:table S2 [ref. 31800], Nión et al. 2016:45 [ref. 35565] with author as Gill 1889, Kim et al. 2021:122 [ref. 39117]). Current status: Valid as Psychrolutes marmoratus (Gill 1889). Psychrolutidae: Psychrolutinae. Distribution: Southeastern Pacific, southwestern Atlantic: Chile to Argentina. Habitat: marine.

marmoratus, Neophrynichthys Regan [C. T.] 1913:241 [Transactions of the Royal Society of Edinburgh v. 49 (pt 2, no. 2); ref. 3651] Strait of Magellan or Burdwood Bank, southwestern Atlantic. Syntypes: BMNH 1880.7.28.5 (1), 1883.6.28.1 (1); NMSZ 1921.143.454 (1). Based on specimens and description of Neophrynichthys latus of Günther 1881:20, pl. 1 [ref. 2012]. •Permanently invalid, preoccupied by, and also a synonym of Neophrynichthys marmoratus Gill 1889 -- (Nelson 1977:501 [ref. 7034]). •Synonym of Psychrolutes marmoratus (Gill 1889). 1°Homonym. Current status: Synonym of Psychrolutes marmoratus (Gill 1889). Psychrolutidae: Psychrolutinae. Distribution: Southwestern Atlantic. Habitat: marine.

maschalis, Ocynectes Jordan [D. S.] & Starks [E. C.] 1904:307, Fig. 34 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Wakanoura, Wakayama Prefecture, Japan, Inland Sea. Holotype: CAS-SU 7717. Paratypes: USNM 50926 (1). Additional material: CAS-SU 7583 (6), CAS-SU 69836 [ex CAS-SU 7717] (1); USNM 50926. Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Ocynectes maschalis Jordan & Starks 1904 -- (Watanabe 1960:142 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:246 [ref. 15964], Kim & Youn 1992:72 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:647 [ref. 25086], Shinohara et al. 2000:177 [ref. 25987], Nakabo 2002:647 [ref. 26001], Choi et al. 2003:262, 577 [ref. 26218], Parin et al. 2014:266 [ref. 33547], Dyldin & Orlov 2022:50 [ref. 38972]). Current status: Valid as Ocynectes maschalis Jordan & Starks 1904. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

matsubarae, Astrocottus Katayama [M.] 1942:334, Fig. 1 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 54 (no. 8); ref. 22718] Kakogawa, Hyogo Prefecture, Japan, Inland Sea of Japan. Holotype: NSMT-P 18253. Paratypes: (1, missing). The spelling matsubarae is acceptable; early workers almost certainly Latinized the name first, and then added an 'e' (Art. 31.1.1) [WNE, 12 Mar. 2012] -- we suggest using matsubarae when the species is treated as valid. •Valid as Astrocottus matsubarae Katayama 1942 -- (Watanabe 1960:24 [ref. 11959], Yabe in Masuda et al. 1984:325 [ref. 6441], Nakabo 2000:632 [ref. 25086], Yabe & Maruyama 2001:61 [ref. 25245], Nakabo 2002:632 [ref. 26001], Tsuruoka et al. 2008:29 [ref. 29568]). Current status: Valid as Astrocottus matsubarae Katayama 1942. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific: Japan. Habitat: brackish, marine.

matsubarai, Myoxocephalus Watanabe [M.] 1958:289, Figs. 117-118, Pl. 20 [Studies of Japanese cottid fishes; ref. 21357] Funka Bay, Hokkaidô, Japan, depth 35 fathoms. Holotype (unique): not researched. Spelled matsubari on plate caption. Also described as new in Watanabe 1960:80, Pl. 26 (fig. 2) [ref. 11959]. •Valid as Myoxocephalus matsubarai Watanabe 1958 -- (Yabe in Masuda et al. 1984:327 [ref. 6441]). •Synonym of Myxocephalus jaok (Cuvier 1829) -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Distribution: Western North Pacific. Habitat: marine.

matuaensis, Microcottus Yabe [M.] & Pietsch [T. W.] 2003:276, Figs. 1-4 [Ichthyological Research v. 50 (no. 3); ref. 27215] Eastern Matua Island, central Kuril Islands, 48°04.2'N, 153°16.1'E, depth 0.5 meters. Holotype: HUMZ 172380. Paratypes: HUMZ 150599-600 (2), 172381-91 (11); UW 47239 (1), NSMT-P 66123 (1). •Valid as Microcottus matuaensis Yabe & Pietsch 2003 -- (Parin et al. 2014:259 [ref. 33547]). Current status: Valid as Microcottus matuaensis Yabe & Pietsch 2003. Psychrolutidae: Myoxocephalinae. Distribution: Boreal North Pacific, Kuril Archipelago. Habitat: marine.

meanyi, Ruscarius Jordan [D. S.] & Starks [E. C.] 1895:805, Pl. 80 [Proceedings of the California Academy of Sciences (Series 2) v. 5; ref. 2522] Puget Sound, Washington, U.S.A. Lectotype: CAS-SU 3127 (with syntypes). Paralectotypes: CAS-SU 3127 (2). Type catalog: Böhlke 1953:132 [ref. 12291]. Lectotype established (as figured specimen) in caption to Pl. 285, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. •Valid as Artedius meanyi (Jordan & Starks 1895) -- (Eschmeyer & Herald 1983:163 [ref. 9277], Matarese et al. 1989:384 [ref. 26885], McAllister 1990:194 [ref. 14674]). •Valid as Ruscarius meanyi Jordan & Starks 1895 -- (Begle 1989:648 [ref. 12739], Robins et al. 1991:43 [ref. 14237], Mecklenburg et al. 2002:462 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Maslenikov et al. 2013:11 [ref. 37947], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:44 [ref. 34694], Kells et al. 2016:190 [ref. 35888], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Ruscarius meanyi Jordan & Starks 1895. Psychrolutidae: Psychrolutinae. Distribution: Northeastern Pacific: southeastern Alaska (U.S.A.) to northern California (U.S.A.): larvae straying to Aleutian Islands (U.S.A.). Habitat: marine.

mednius, Myoxocephalus Bean [T. H.] in Jordan & Evermann 1898:1983 [Bulletin of the United States National Museum No. 47; ref. 2444] Medny Island, Commander Islands, Bering Sea. Holotype (unique): USNM 33863. Author is T. H. Bean. Also appeared as Bean in Jordan & Gilbert 1899:465, Pl. 64b [ref. 2478]. •Questionably a synonym of Porocottus bradfordi Rutter 1898, subspecies albomaculatus (Schmidt 1916) -- (Neelov 1979:136 [ref. 3152]). •Valid as Porocottus mednius (Bean 1898) -- (Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:486 [ref. 25968], Parin et al. 2002:S84 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:91 [ref. 37547], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:269 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Love et al. 2021:124 [ref. 39279]). Current status: Valid as Porocottus mednius (Bean 1898). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific: Kuril Islands north to Bering ea and Chukchi Sea, then east to southeastern Alaska (U.S.A.) Habitat: marine.

megacephalus, Andriashevicottus Fedorov [V. V.] 1990:746, Fig. [Voprosy Ikhtiologii v. 30 (no. 5); ref. 20136] Simushir Island, Kuril Islands, western Pacific, 46°46'N, 151°51'E, depth 100 meters. Holotype (unique): ZIN 49087. On p. 124 of English translation. •Valid as Andriashevicottus megacephalus Fedorov 1990 -- (Sheiko & Fedorov 2000:26 [ref. 25650], Parin et al. 2002:S68 [ref. 28300], Parin et al. 2014:233 [ref. 33547]). Current status: Valid as Andriashevicottus megacephalus Fedorov 1990. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

megacephalus, Chitonotus Lockington [W. N.] 1879:70 [Mining and Scientific Press. An illustrated journal of mining, popular sciences and general news v. 39 (no. 5) 2 Aug.; ref. 20591] San Francisco Bay, California, U.S.A. Syntypes: CAS (3, lost in 1906). Also appeared in Lockington 1881:142 [ref. 2819]. •Synonym of Chitonotus pugetensis (Steindachner 1876). Current status: Synonym of Chitonotus pugetensis (Steindachner 1876). Psychrolutidae: Icelinae. Habitat: marine.

megalops, Antipodocottus DeWitt [H. H.] 1969:30, Fig. 1 [Copeia 1969 (no. 1); ref. 7811] Near Antipodes Islands, New Zealand Region, 49°40'S, 178°53'E, depth 439-540 meters. Holotype (unique): LACM 10991-1. •Valid as Antipodocottus megalops DeWitt 1969 -- (Fricke & Brunken 1984:46 [ref. 5262], Paulin et al. 1989:172 [ref. 24556], Stewart 2015:1120 [ref. 34254]). Current status: Valid as Antipodocottus megalops DeWitt 1969. Psychrolutidae: Psychrolutinae. Distribution: New Zealand region. Habitat: marine.

meridionalis, Crossias beringi forma Soldatov [V. K.] 1916:220, Pl. 13 (lower) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 21; ref. 18735] Sea of Japan. Lectotype: ZIN 19067. Paralectotypes: ZIN 19067 (3). Treated as infrasubspecific in Eschmeyer et al. 1998; now regarded as subspecific. Muto et al. 2002:231 [ref. 26331] list a lectotype and 3 paralectotypes as ZISP 19067. •Synonym of Porocottus tentaculatus (Kner 1868) -- (Lindberg & Krasyukova 1987:224 [ref. 15964], Parin et al. 2002:S84 [ref. 28300], Parin et al. 2014:270 [ref. 33547], Dyldin & Orlov 2017:67 [ref. 35138]). Current status: Synonym of Porocottus tentaculatus (Kner 1868). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

mertensii, Cottus Valenciennes [A.] in Cuvier & Valenciennes 1832:496 [Histoire naturelle des poissons; ref. 1000] Petropaulski, Kamchatka. Type catalog: Blanc & Hureau 1968:61 [ref. 20738]. •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Neelov 1979:122 [ref. 3152], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:261 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Habitat: marine.

mesembrinus, Stlengis Fricke [R.] & Brunken [H.] 1983:10, Figs. 1-2 [Japanese Journal of Ichthyology v. 30 (no. 1); ref. 8347] Kai Islands, Indonesia, 5°46'00"S, 132°49'35"E, depth 352 meters. Holotype (unique): ZMUC P81868. •Valid as Antipodocottus mesembrinus (Fricke & Brunken 1983) -- (Fricke & Brunken 1984:46 [ref. 5262], Nelson 1990:845 [ref. 15720], Nelson & Carpenter 1999:2426 [ref. 24809]). Current status: Valid as Antipodocottus mesembrinus (Fricke & Brunken 1983). Psychrolutidae: Psychrolutinae. Distribution: Indonesia. Habitat: marine.

metopias, Triglops Gilbert [C. H.] & Burke [C. V.] 1912:50, Fig. 8 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Petrel Bank, Bering Sea, Albatross station 4777, depth 52 fathoms. Holotype: USNM 74372. Paratypes: CAS-SU 22231 (4); USNM 70851 (2), 70870 (1). Type catalog: Böhlke 1953:133 [ref. 12291]. •Valid as Triglops metopias Gilbert & Burke 1912 -- (Pietsch 1994:365 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:427 [ref. 25968], Parin et al. 2002:S88 [ref. 28300], Nelson et al. 2004:122 [ref. 27807], Love et al. 2005:93 [ref. 37547], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:276 [ref. 33547], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Triglops metopias Gilbert & Burke 1912. Psychrolutidae: Triglopinae. Distribution: North Pacific: Bering Sea, Aleutian Islands to southeastern Alaska (U.S.A.). Habitat: marine.

miacanthus, Artediellus Gilbert [C. H.] & Burke [C. V.] 1912:47, Fig. 7 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off east coast of Kamchatka, Russia, 52°46'50"N,148°44'E, Albatross station 4795, depth 48-69 fathoms. Holotype: USNM 74371. Paratypes: CAS-SU 22229 (4), 22243 (4); USNM 74480 (1). Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Artediellus miacanthus Gilbert & Burke 1912 -- (Neelov 1979:154 [ref. 3152], Nelson 1986:43 [ref. 5808], Lindberg & Krasyukova 1987:231 [ref. 15964], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:495 [ref. 25968], Parin et al. 2002:S70 [ref. 28300], Love et al. 2005:84 [ref. 37547], Inagawa & Yabe 2006:196 [ref. 28854], Parin et al. 2014:237 [ref. 33547], Love et al. 2021:118 [ref. 39279] with question). Current status: Valid as Artediellus miacanthus Gilbert & Burke 1912. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: northern Kuril lands north to Bering Sea off Alaska (U.S.A.). Habitat: marine.

microphthalmus, Malacocottus Schmidt [P. Yu.] in Popov 1933:142 [Issledovaniya Morey SSSR, Leningrad No. 19; ref. 6535] Peter the Great Bay, Russia. Holotype (unique). •Synonym of Malacocottus zonurus Bean 1890, but a valid subspecies microphthalmus Schmidt 1933 -- (Sheiko 1983:19 [ref. 37041]). •Synonym of Malacocottus zonurus Bean 1890 -- (Sheiko & Fedorov 2000:30 [ref. 25650]). •Synonym of Malacocottus gibber Sakamoto 1930 -- (Parin et al. 2002:S94 [ref. 28300], Parin et al. 2014:302 [ref. 33547]). Current status: Synonym of Malacocottus gibber Sakamoto 1930. Psychrolutidae: Psychrolutinae. Habitat: marine.

microporos, Psychrolutes Nelson [J. S.] 1995:70, Figs. 1, 2A [Proceedings of the Zoological Society, Calcutta v. 48 (no. 2); ref. 22668] New Zealand. 42°55.1'S, 168°59.0'E, depth 975-986 meters. Holotype: NMNZ 15030. Paratypes: BSKU 23089 (1), 32395 (1); NMNZ 9487 (1), 11348 (1), 11551 (1), 14869 (1), 15029 (1), 15781 (1), 16155 (1), 16157 (1). Plus additional material. •Valid as Psychrolutes microporos Nelson 1995 -- (Nakabo 2000:653 [ref. 25086], Jackson & Nelson 2000:725 [ref. 25146], Nakabo 2002:653 [ref. 26001], Stewart 2015:1130 [ref. 34254]). Current status: Valid as Psychrolutes microporos Nelson 1995. Psychrolutidae: Psychrolutinae. Distribution: New Zealand region. Japan provisional [ref. 34254]. Habitat: marine.

microps, Cottunculus Collett [R.] 1875:20, Pl. 1 (figs. 1-3) [Norges fiske; ref. 884] Hasvig, Sørøy, northern Norway, depth 376 meters. Holotype: ZMUO J4128. Type catalog: Pethon 1969:7 [ref. 19268]. •Valid as Cottunculus microps Collett 1875 -- (Neelov 1973:603 [ref. 7219], Nelson 1977:508 [ref. 7034], Nelson & Nakamura 1980:990 [ref. 26434], Fedorov & Nelson in Whitehead et al. 1986:1262 [ref. 13677], Scott & Scott 1988:492 [ref. 25518], Nelson 1989:407 [ref. 12546], McAllister 1990:202 [ref. 14674], Yabe in Okamura et al. 1995:169 [ref. 22531], Mecklenburg et al. 2002:518 [ref. 25968], Klein-MacPhee in Collette & Klein-MacPhee 2002:357 [ref. 26158], Parin et al. 2002:S92 [ref. 28300], Moore et al. 2003:223 [ref. 27076], Byrkjedal & Orlov 2007:67 [ref. 29312], Møller et al. 2010:54 [ref. 31169], Mecklenburg et al. 2011:125 [ref. 31212], Page et al. 2013:126 [ref. 32708], Dolgov 2013:918 [ref. 33112], Byrkjedal et al. 2014:14 [ref. 33415], Mecklenburg et al. 2014:137 [ref. 33463], Parin et al. 2014:298 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:55, 138 [ref. 36199], Mecklenburg et al. 2016:98 [ref. 34440], Mecklenburg et al. 2018:237 [ref. 36370], Love et al. 2021:120 [ref. 39279], Simian et al. 2022:376 [ref. 40776]). Current status: Valid as Cottunculus microps Collett 1875. Psychrolutidae: Psychrolutinae. Distribution: North Atlantic and adjacent Arctic Ocean. Habitat: marine.

microps, Icelinus Starks [E. C.] 1896:552 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 4195] Name only, not available; mentioned in passing under Icelinus strabo Starks 1896. •In the synonymy of Icelinus borealis Gilbert 1896 -- (Bolin 1936:159 [ref. 506]). Nomen Nudum. Current status: Synonym of Icelinus borealis Gilbert 1896. Psychrolutidae: Oligocottinae.

minor, Cottiusculus Watanabe [M.] 1958:410, Figs. 198-199, Pl. 43 [Studies of Japanese cottid fishes; ref. 21357] Off Funakawa, Akita Prefecture, Japan, depth 70 fathoms. Syntypes: (4). Also described as new in Watanabe 1960:172, Pl. 37 (fig. 2) [ref. 11959]. •Valid as Cottiusculus minor Watanabe 1958 -- (Watanabe 1960:172 [ref. 11959]). •Valid as Artediellus minor (Watanabe 1958) -- (Nakabo 2000:642 [ref. 25086], Nakabo 2002:642, 1526 [ref. 26001], Parin et al. 2014:238 [ref. 33547], Dyldin & Orlov 2022:40 [ref. 38972] with question). Current status: Valid as Artediellus minor (Watanabe 1958). Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

minutus, Cottus Pallas [P. S.] 1814:145 [Zoographia Rosso-Asiatica v. 3; ref. 3351] Talak Island, Okhotsk Sea. No types known. •Valid as Porocottus minutus (Pallas 1814) -- (Neelov 1979:137 [ref. 3152], Sheiko & Fedorov 2000:29 [ref. 25650], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S84 [ref. 28300], Fedorov et al. 2003:79 [ref. 27579], Parin et al. 2014:269 [ref. 33547], Poezzhalova-Chegodaeva 2019:113 [123] [ref. 38518], Dyldin et al. 2021:68 [ref. 38804], Dyldin & Orlov 2022:51 [ref. 38972]). Current status: Valid as Porocottus minutus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific: Sea of Okhotsk. Habitat: marine.

minutus, Vellitor Iwata [A.] 1983:2, Figs. 1, 2 (A-B), 3 (A, C) [Japanese Journal of Ichthyology v. 30 (no. 1); ref. 5794] Kominato, Chiba Prefecture, Japan, 35°07'N, 140°11'E. Holotype: NSMT-P 19799. Paratypes: (39) BMNH 1981.9.5.13-18 (6); HUMZ 92517-21 (5); MTUF 24661-62 (2), 24663 (1, c&s), 24664-66 (3); NSMT-P 19793-98 (6, 1 ea.); USNM 228073-77 (5), YCM P.2372 (1), P.10010-10015 (6); ZUMT 54267-71 (5). Type catalog: Shibata et al. 2011:13 [ref. 31469]. •Valid as Vellitor minutus Iwata 1983 -- (Yabe in Masuda et al. 1984:329 [ref. 6441], Nakabo 2000:647 [ref. 25086], Nakabo 2002:647 [ref. 26001], Shibata et al. 2011:13 [ref. 31469]). Current status: Valid as Vellitor minutus Iwata 1983. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

misakia, Schmidtia Jordan [D. S.] & Starks [E. C.] 1904:237, Fig. 2 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Manazuru Point, Sagami Bay, Japan, Albatross station 3698, depth 153 fathoms. Holotype: USNM 50913. Paratypes: CAS-SU 7506 (1). Type catalog: Böhlke 1953:132 [ref. 12291]. Types: Shinohara & Williams 2006:557 [ref. 31656]. •Valid as Stlengis misakia (Jordan & Starks 1904) -- (Watanabe 1960:35 [ref. 11959], Yamakawa in Okamura et al. 1982:291, 403 [ref. 8256], Yabe in Masuda et al. 1984:325 [ref. 6441], Nakabo 2000:630 [ref. 25086], Shinohara et al. 2001:319 [ref. 25995], Nakabo 2002:630 [ref. 26001], Sonoyama et al. 2020:91 [ref. 37637]). Current status: Valid as Stlengis misakia (Jordan & Starks 1904). Psychrolutidae: Oligocottinae. Distribution: Northwestern Pacific: southern and central Japan. Habitat: marine.

mitchilli, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:188 [Histoire naturelle des poissons v. 4; ref. 998] New York, U.S.A. •Synonym of Myoxocephalus aenaeus (Mitchill 1814) -- (Scott & Scott 1988:502 [ref. 25518]). Current status: Synonym of Myoxocephalus aenaeus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

modestus, Ocynectes Snyder [J. O.] 1911:539 [Proceedings of the United States National Museum v. 40 (no. 1836); ref. 4152] Same, Mutsu, Japan. Holotype: USNM 68240. Paratypes: CAS-SU 21426 (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Ocynectes modestus Snyder 1911 -- (Watanabe 1960:144 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Shiogaki 1987:222 [ref. 6708], Lindberg & Krasyukova 1987:248 [ref. 15964], Nakabo 2000:647 [ref. 25086], Nakabo 2002:647 [ref. 26001]). Current status: Valid as Ocynectes modestus Snyder 1911. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

mororane, Stelgistrum Jordan [D. S.] & Seale [A.] 1906:146, Fig. 4 [Proceedings of the United States National Museum v. 30 (no. 1445); ref. 2496] Mororan harbor, Hokkaido Island, Japan. Holotype (unique): CAS-SU 9260. Type catalog: Böhlke 1953:132 [ref. 12291]. •Synonym of Stelgistrum stejnegeri Jordan & Gilbert 1898 -- (Watanabe 1960:32 [ref. 11959] as morarane, Lindberg & Krasyukova 1987:195 [ref. 15964] as mororanae). •Valid as Icelus mororanis (Jordan & Seale 1906) -- (Tsuruoka et al. 2009:97 [ref. 32107], Parin et al. 2014:255 [ref. 33547], Dyldin & Orlov 2022:46 [ref. 38972]). Current status: Valid as Icelus mororanis (Jordan & Seale 1906). Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Hokkaido (Japan), Kuril Islands and Sakhalin (Russia). Habitat: marine.

mucosus, Acanthocottus Ayres [W. O.] 1854:194 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] New England coast, U.S.A. No types known. Appeared first as above on 13 Oct.; then in Ayres 1854:12 [ref. 18923]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

murrayi, Triglops Günther [A.] 1888:209, Pl. 4 (fig. A) [Proceedings of the Royal Society of Edinburgh v. 15; ref. 14482] Northeastern coast of Scotland, depth 64 fathoms. Lectotype: BMNH 1889.6.28.5. Paralectotypes: BMNH 1889.6.28.2-4 (3). Appeared on Pl. 4 as Inglops murrayi. Lectotype selected by Pietsch 1994:368 [ref. 21189] -- the original figured specimen. •Valid as Triglops murrayi Günther 1888 -- (Neelov 1973:602 [ref. 7219], Fedorov in Whitehead et al. 1986:1257 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:509 [ref. 25518], McAllister 1990:202 [ref. 14674], Pietsch 1994:368 [ref. 21189], Yabe in Okamura et al. 1995:166 [ref. 22531], Klein-MacPhee in Collette & Klein-MacPhee 2002:356 [ref. 26158], Parin et al. 2002:S88 [ref. 28300], Moore et al. 2003:222 [ref. 27076], Nelson et al. 2004:122 [ref. 27807], Fricke 2007:29 [ref. 30577], Møller et al. 2010:53 [ref. 31169], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:276 [ref. 33547], Afonso et al. 2016:51, 289 [ref. 36199], Mecklenburg et al. 2018:225 [ref. 36370], Simian et al. 2022:375 [ref. 40776]). Current status: Valid as Triglops murrayi Günther 1888. Psychrolutidae: Triglopinae. Distribution: North Atlantic and adjacent Arctic Ocean. Habitat: marine.

namiyei, Ceratocottus Jordan [D. S.] & Starks [E. C.] 1904:259, Fig. 13 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Nemuro, Hokkaido, Japan. Holotype: CAS-SU 8108. Paratypes: USNM 50917 (1, dry). Type catalog: Böhlke 1953:127 [ref. 12291]. •Synonym of Ceratocottus diceraus (Pallas 1787) -- (Watanabe 1960:110 [ref. 11959]). •Synonym of Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:836 [ref. 26414], Neelov 1979:146 [ref. 3152], Kim & Youn 1992:62 [ref. 22287], Parin et al. 2002:S74 [ref. 28300], Parin et al. 2014:245 [ref. 33547]). Current status: Synonym of Enophrys diceraus (Pallas 1787). Psychrolutidae: Myoxocephalinae. Habitat: marine.

nebulosus, Hemilepidotus Ayres [W. O.] 1854:166 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] Originally as H. ? nebulosus. Appeared first as above on 25 Aug. and is available from that publication; not published in the Proceedings (Ayres 1854 [ref. 18923]; see Ayres 1855:76 [ref. 159]). •Synonym of Artedius notospilotus Girard 1856. Current status: Synonym of Artedius notospilotus Girard 1856. Psychrolutidae: Oligocottinae. Habitat: marine.

neyelovi, Artediellus Muto [F.], Yabe [M.] & Amaoka [K.] 1994:275, Figs. 1-3 [Japanese Journal of Ichthyology v. 41 (no. 3); ref. 21556] Eastern end of Tsugaru Strait, 41°57'N, 141°08'E, Japan, depth 137 meters. Holotype: HUMZ 103874. Paratypes: HUMZ 93974 (1), 96598-601 (4), 96628 (1), 99540 (1), 100981-88 (8), 100990 (1), 101261-67 (7), 103849 (1), 103876-79 (4), 103881 (1), 103948 (1), 103976 (1), 103990 (1); MNHN 1994-0507 to 0508 (1, 1); NSMT-P 45944-45 (1, 1); USNM 330305-06 (2). •Valid as Artediellus neyelovi Muto, Yabe & Amaoka 1994 -- (Nakabo 2000:641 [ref. 25086], Nakabo 2002:641 [ref. 26001]). Current status: Valid as Artediellus neyelovi Muto, Yabe & Amaoka 1994. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

niger, Cottus Bean [T. H.] 1881:151 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Saint Paul Island, Pribilof Group, Bering Sea, Alaska, U.S.A. Lectotype: USNM 27971. Paralectotypes: MCZ 36021 [ex USNM 27952] (1); USNM 23929 (1), 27952 (orig. 12, now 1). Lectotype established (as figured specimen) in caption to Pl. 298, p. 3293 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. Objectively invalid; preoccupied by Cottus niger Lacepède 1801 but we recommend suppressing that name that has not been used in 100 years and its status is uncertain. •Valid as Myoxocephalus niger (Bean 1881) -- (Watanabe 1960:78 [ref. 11959], Neelov 1979:123 [ref. 3152], Sheiko 1983:18 [ref. 37041], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:213 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:476 [ref. 25968], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:76 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:90 [ref. 37547], Page et al. 2013:124 [ref. 32708], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:262 [ref. 33547], Kells et al. 2016:182 [ref. 35888], Love et al. 2021:123 [ref. 39279], Dyldin & Orlov 2022:49 [ref. 38972] with question). Current status: Valid as Myoxocephalus niger (Bean 1881). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific: northern Sea of Japan and Sea of Okhotsk north to southern Bering Sea, east to western Gulf of Alaska (U.S.A.). Habitat: marine.

nigrescens, Porocottus Tanaka [S.] 1908:37 [241], Pl. 9 (figs. 1-2) [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 20; ref. 14360] Southern part of Sakhalin Island, western Okhotsk Sea. Holotype (unique): ZUMT 1387 [= ZUMT 21699]. Appeared first as above in Japanese. Then in English version in Tanaka 1908:241, Pl. 9 (figs. 1-2) [see ref. 14360]. •Valid as Porocottus nigrescens Tanaka 1908 -- (Watanabe 1960:93 [ref. 11959]). •Synonym of Myoxocephalus platycephalus (Pallas 1814), subspecies taeniopterus (Kner 1868) -- (Berg 1949:1128 [ref. 20662]). •Synonym of Megalocottus platycephalus (Pallas 1814), subspecies taeniopterus (Kner 1868) -- (Neelov 1979:130 [ref. 3152]). •Synonym of Megalocottus platycephalus (Pallas 1814) -- (Parin et al. 2002:S79 [ref. 28300]). •Synonym of Megalocottus taeniopterus (Kner 1868) -- (Parin et al. 2014:258 [ref. 33547], Dyldin & Orlov 2017:64 [ref. 35138] with question). Current status: Synonym of Megalocottus taeniopterus (Kner 1868). Psychrolutidae: Myoxocephalinae. Habitat: marine.

nigricans, Cottus Lesueur [C. A.] in Vaillant 1896:26 [12], Pl. 19 (unedited) [Bulletin de la Société philomathique de Paris (8th Série) v. 8 (no. 1); ref. 13482] No locality. On p. 12 of separate. •Synonym of Myoxocephalus aenaeus (Mitchill 1814). Current status: Synonym of Myoxocephalus aenaeus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

nigripinnis, Artediellus Schmidt [P. Yu.] 1937:571 [Comptes Rendus (Doklady) de l'Académie des Sciences de l'U.R.S.S. v. 15 (no. 9); ref. 15302] Between St. Jona Island (55°40'N, 143°04'E) and western coast of Kamchatka (51°09'N, 154°17'E), Okhotsk Sea, depth 500-592 meters. Lectotype: ZIN 24464 (1 of 3). Paralectotypes: MCZ 34085 [ex ZIN] (1), ZIN (2 of 3). Lectotype selected by Neelov 1979:160 [ref. 3152]. Appeared first as name only in Schmidt 1935. •Valid as Artediellichthys nigripinnis (Schmidt 1937) -- (Sheiko & Fedorov 2000:26 [ref. 25650], Mecklenburg et al. 2002:489 [ref. 25968], Parin et al. 2002:S69 [ref. 28300], Fedorov et al. 2003:64 [ref. 27579], Love et al. 2005:84 [ref. 37547], Parin et al. 2014:234 [ref. 33547], Love et al. 2021:117 [ref. 39279], Dyldin & Orlov 2022:39 [ref. 38972]). Current status: Valid as Artediellichthys nigripinnis (Schmidt 1937). Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Sea of Okhotsk and Kuril Islands north to Bering Sea, Aleutian Islands to eastern Gulf of Alaska. Habitat: marine.

nihonkaiensis, Cottiusculus Kai [Y.] & Nakabo [T.] 2009:215, Figs. 1-3 [Ichthyological Research v. 56 (no. 3); ref. 30299] Off Tango Peninsula, Kyoto Prefecture, Japan. Holotype: FAKU 130664. Paratypes: (31) at FAKU, NSMT. •Valid as Cottiusculus nihonkaiensis Kai & Nakabo 2009 -- (Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:240 [ref. 33547], Kim et al. 2020:101 [ref. 39118], Prokofiev 2020:13 [ref. 37765], Sonoyama et al. 2020:89 [ref. 37637], Dyldin & Orlov 2022:41 [ref. 38972], Jin et al. 2023:143 [ref. 40382]). Current status: Valid as Cottiusculus nihonkaiensis Kai & Nakabo 2009. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia), northern Hokkaido (Japan) and Korean Peninsula. Habitat: marine.

nivosus, Cottus Herzenstein [S. M.] 1890:113 [23] [Mélanges Biologiques, tirés du Bulletin physico-mathématique de l'Académie Impériale des Sciences de St. Pétersbourg v. 13; ref. 2149] St. Olga Bay, Russia, Sea of Japan. Holotype (unique): ZIN 8719. On p. 23 of Bull. Acad. Imp. Sci. St. Petersbourg (n. s.) v. 2. •Synonym of Myoxocephalus brandtii (Steindachner 1867) -- (Neelov 1979:121 [ref. 3152], Lindberg & Krasyukova 1987:210 [ref. 15964] as brandti, Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S80 [ref. 28300], Parin et al. 2014:260 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus brandtii (Steindachner 1867). Psychrolutidae: Myoxocephalinae. Habitat: marine.

notospilotus, Artedius Girard [C. F.] 1856:134 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1809] Tomales Bay, California, U.S.A. Syntypes: USNM 329 (1), 7867 (not found in 1980). •Valid as Artedius notospilotus Girard 1856 -- (Bolin 1944:50 [ref. 6379], Eschmeyer & Herald 1983:163 [ref. 9277], Begle 1989:648 [ref. 12739], Mecklenburg et al. 2002:404 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:85 [ref. 37547], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:164 [ref. 35888], Burton & Lea 2019:65 [ref. 37205], Love et al. 2021:118 [ref. 39279]). Current status: Valid as Artedius notospilotus Girard 1856. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: Puget Sound, Washington (U.S.A.) south to Pacific coast of central Baja California (Mexico). Habitat: marine.

nudithorax, Ricuzenius Bolin [R. L.] 1936:27, Fig. 5 [Proceedings of the United States National Museum v. 84 (no. 3000); ref. 505] Yeso Strait, Japan, 44°04'N, 145°32'E, Albatross station 5031, depth 86 fathoms. Holotype (unique): USNM 102104. •Valid as Ricuzenius nudithorax Bolin 1936 -- (Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:186 [ref. 15964], Nakabo 2000:636 [ref. 25086], Nakabo 2002:636 [ref. 26001], Parin et al. 2002:S85 [ref. 28300], Parin et al. 2014:271 [ref. 33547]). Current status: Valid as Ricuzenius nudithorax Bolin 1936. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

nudus, Centridermichthys Döderlein [L.] in Steindachner & Döderlein 1887:258 [2] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 53 (1. abth.); ref. 4249] Tokyo Bay, Japan. Holotype (unique): NMW. •Synonym of Vellitor centropomus (Richardson 1848) -- (Watanabe 1960:165 [ref. 11959]). Current status: Synonym of Vellitor centropomus (Richardson 1848). Psychrolutidae: Psychrolutinae. Habitat: marine.

nudus, Cottunculus Nelson [J. S.] 1989:402, Figs. 1-4 [Copeia 1989 (no. 2); ref. 12546] South Canterbury Bight, New Zealand, 44°55'S, 174°05'E, depth 1090-1123 meters. Holotype: NMNZ P.15787. Paratypes: NMNZ P.9488 (1), P.11819 (1), P.13137 (2), P.14863 (1), P.15065 (1), P.15071 (1), P.15622 (1), P.15797 (1), P.15799 (1), P.15842 (1), P.16041 (1), P.16046 (1), P.16162 (2), P.17388 (1), P.20881 (2), P.20906 (2). Type catalog: Hardy 1990:10 [ref. 18942]. •Valid as Cottunculus nudus Nelson 1989 -- (Paulin et al. 1989:172 [ref. 24556], Yabe in Amaoka et al. 1990:245 [ref. 16024], Jackson & Nelson 2000:725 [ref. 25146], Byrkjedal & Orlov 2007:67 [ref. 29312], Stewart 2015:1126 [ref. 34254]). Current status: Valid as Cottunculus nudus Nelson 1989. Psychrolutidae: Psychrolutinae. Distribution: New Zealand region. Habitat: marine.

nybelini, Triglops Jensen [A. S.] 1944:24, Pl. 3 (figs. a-b) [Spolia Zoologica Musei Hauniensis v. 4; ref. 13060] Northwestern Greenland, depth 490 meters. Lectotype: ZMUC P81232. Paralectotypes: ZMUC P81182-231 (52), P81233-95 (60). Type catalog: Nielsen 1974:81 [ref. 9588]. Lectotype selected by Pietsch 1994:360 [ref. 21189] -- the figured specimen in Jensen 1944, Pl. 3a. •Valid as Triglops nybelini Jensen 1944 -- (Neelov 1973:602 [ref. 7219], Fedorov in Whitehead et al. 1986:1258 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:510 [ref. 25518], McAllister 1990:202 [ref. 14674], Pietsch 1994:361 [ref. 21189], Yabe in Okamura et al. 1995:167 [ref. 22531], Mecklenburg et al. 2002:426 [ref. 25968], Parin et al. 2002:S88 [ref. 28300], Nelson et al. 2004:122 [ref. 27807], Love et al. 2005:93 [ref. 37547], Møller et al. 2010:53 [ref. 31169], Mecklenburg et al. 2011:124 [ref. 31212], Dolgov 2013:918 [ref. 33112], Maslenikov et al. 2013:12 [ref. 37947], Page et al. 2013:125 [ref. 32708], Mecklenburg et al. 2014:136 [ref. 33463], Parin et al. 2014:277 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:51, 290 [ref. 36199], Mecklenburg et al. 2016:88 [ref. 34440], Mecklenburg et al. 2018:227 [ref. 36370], Love et al. 2021:127 [ref. 39279]). Current status: Valid as Triglops nybelini Jensen 1944. Psychrolutidae: Triglopinae. Distribution: Arctic Ocean and North Atlantic; nearly circumpolar. Habitat: marine.

obtusus, Phallocottus Schultz [L. P.] 1938:188, Fig. 70 [Proceedings of the United States National Museum v. 85 (no. 3038); ref. 3951] Igitkin Island, Aleutian Islands, Alaska, U.S.A. Holotype: USNM 105280. Paratypes: USNM 105281 (5). •Valid as Phallocottus obtusus Schultz 1938 -- (Sheiko & Fedorov 2000:28 [ref. 25650], Yabe et al. 2001:70 [ref. 25246], Mecklenburg et al. 2002:496 [ref. 25968], Parin et al. 2002:S83 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:91 [ref. 37547], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:267 [ref. 33547], Love et al. 2021:124 [ref. 39279]). Current status: Valid as Phallocottus obtusus Schultz 1938. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Kuril Islands; Aleutian Islands and Alaska (U.S.A.). Habitat: marine.

occidentalis, Gymnacanthus tricuspis Schmidt [P. Yu.] 1927:29 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 15821] Barents Sea and White Sea. Syntypes: ZIN 6370 (2), 7083 (1), 10446 (1), 11262 (1), 20380 (1). •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Parin et al. 2002:S75 [ref. 28300] as Gymnacanthus, Parin et al. 2014:249 [ref. 33547], Mecklenburg et al. 2016:62 [ref. 34440]). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Psychrolutidae: Icelinae. Habitat: marine.

occidentalis, Psychrolutes (Cottunculoides) Fricke [R.] 1990:404, Fig. 1 [Japanese Journal of Ichthyology v. 36 (no. 4); ref. 15492] About 35 kilometers east of Rowley Shoals, Western Australia, 17°17'S, 119°57'E, depth 350 meters. Holotype: WAM P.25395-014. Paratypes: AMS I.22809-006 (1), I.22814-003 (3); WAM P.28074-001 (1), P.28133-001 (1). Type catalog: Not treated by Moore et al. 2008 [ref. 30117]. •Valid as Psychrolutes occidentalis Fricke 1990 -- (Nelson 1995:69 [ref. 22668], Nelson 1999:2428 [ref. 24810], Paxton et al. 2006:954 [ref. 28994], Larson et al. 2013:93 [ref. 32988], Fricke et al. 2014:57 [ref. 33932]). Current status: Valid as Psychrolutes occidentalis Fricke 1990. Psychrolutidae: Psychrolutinae. Distribution: Southeastern Indian Ocean, western Pacific: Western Australia and Arafura Sea to Papua New Guinea. Habitat: marine.

ocellatus, Acanthocottus Storer [H. R.] 1850:253 [Boston Journal of Natural History v. 6 (no. 2) (art. 15); ref. 18109] Entrance to the Gut of Canso, Burial Island in Wilmot Harbor, Nova Scotia, Canada. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

ochotensis, Artediellus Gilbert [C. H.] & Burke [C. V.] 1912:44, Fig. 5 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off western coast of Kamchatka, Russia, 51°37'N, Albatross station 4798, depth 25 fathoms. Holotype: USNM 22271 (?74369). Paratypes: CAS-SU 68870 (3), USNM 70910 (3). •Valid as Artediellus ochotensis Gilbert & Burke 1912 -- (Neelov 1979:154 [ref. 3152], Yabe in Masuda et al. 1984:328 [ref. 6441], Nelson 1986:43 [ref. 5808], Lindberg & Krasyukova 1987:233 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:641 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:494 [ref. 25968], Nakabo 2002:641 [ref. 26001], Parin et al. 2002:S71 [ref. 28300], Fedorov et al. 2003:66 [ref. 27579], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:84 [ref. 37547], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:121 [ref. 32709], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:238 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Dyldin & Orlov 2017:62 [ref. 35138], Dyldin et al. 2021:61 [ref. 38804], Love et al. 2021:118 [ref. 39279], Dyldin & Orlov 2022:40 [ref. 38972]). Current status: Valid as Artediellus ochotensis Gilbert & Burke 1912. Psychrolutidae: Psychrolutinae. Distribution: North Pacific and adjacent Arctic: Sea of Japan and Sea of Okhotsk north to Chukchi Sea (Russia). Habitat: brackish, marine.

ochotensis, Gilbertidia Schmidt [P. Yu.] 1916:625, Fig. 6 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 3935] Aniva Bay, southern Sakhalin Island, Russia. Holotype (unique): ZIN 19105. •Valid as Gilbertidia ochotensis Schmidt 1916 -- (Watanabe 1960:193 [ref. 11959]). •Synonym of Eurymen gyrinus Gilbert & Burke 1912 -- (Lindberg & Krasyukova 1987:265 [ref. 15964], Mecklenburg et al. 2002:517 [ref. 25968], Parin et al. 2002:S93 [ref. 28300], Nazarkin et al. 2014:177 [ref. 33578], Parin et al. 2014:301 [ref. 33547]). Current status: Synonym of Eurymen gyrinus Gilbert & Burke 1912. Psychrolutidae: Psychrolutinae. Habitat: marine.

ochotensis, Icelus Schmidt [P. Yu.] 1927:4, Figs. 1-2 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 19815] Northern Okhotsk Sea, between 58°48.5'N to 53°56'N, 151°18.5'-137°32'E, depth 60-105 meters. Syntypes: (45) ZIN 21898-99 (1, 2), 21901-04 (3, 6, 6, 6). •Valid as Icelus ochotensis Schmidt 1927 -- (Nelson 1984:50 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Kim & Youn 1992:67 [ref. 22287], Suzuki & Uno 1992:4 [ref. 24139], Murai et al. 1994:343 [ref. 21908], Nakabo 2000:635 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Nakabo 2002:635 [ref. 26001], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S77 [ref. 28300], Fedorov et al. 2003:71 [ref. 27579], Parin et al. 2014:255 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Kim et al. 2020:103 [ref. 39118], Sonoyama et al. 2020:90 [ref. 37637], Dyldin et al. 2021:65 [ref. 38804], Dyldin & Orlov 2022:46 [ref. 38972]). Current status: Valid as Icelus ochotensis Schmidt 1927. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Sea of Japan to Sea of Okhotsk (Korea, Japan, Russia). Habitat: marine.

ochotensis, Myoxocephalus niger var. Taranetz [A. Ya.] 1937:115 [Izvestiia Tikhookeanskogo Nauchno-Issledovatel'skogo Instituta Rybnogo Khoziaistva i Okeanografii [Proceedings, Pacific Research Institute of Fisheries and Oceanography, TINRO, Moskva] v. 11; ref. 13384] Holotype: ZIN 25468. Objectively invalid; preoccupied by Myoxocephalus verrucosus subspecies ochotensis Schmidt 1929. •Synonym of Myoxocephalus aeneus (Mitchill 1814) -- (Neelov 1979:123 [ref. 3152]). •Synonym of Myoxocephalus niger (Bean 1881) -- (Lindberg & Krasyukova 1987:213 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547]). 1°Homonym. Current status: Synonym of Myoxocephalus niger (Bean 1881). Psychrolutidae: Myoxocephalinae. Habitat: marine.

ochotensis, Myoxocephalus verrucosus Schmidt [P. Yu.] 1929:417, Fig. 3 [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 30 (no. 3); ref. 16059] Penshinskaya Guba, Shantarsky Islands (Russia), Sea of Okhotsk; Karaginsky Island (Russia), western Bering Sea. •Valid as Myoxocephalus ochotensis Schmidt 1929 -- (Neelov 1979:122 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:76 [ref. 27579], Parin et al. 2014:262 [ref. 33547], Mecklenburg et al. 2016:85 [ref. 34440], Dyldin & Orlov 2022:49 [ref. 38972] with question). Current status: Valid as Myoxocephalus ochotensis Schmidt 1929. Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific: Sea of Okhotsk north to western Bering Sea. Habitat: marine.

ochotensis, Porocottus sellaris Schmidt [P. Yu.] 1929:411, Figs. 1-2 [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 30 (no. 3); ref. 16059] Penshinskaya Guba [Penzhinskii Bay], Shantar Islands, northern Okhotsk Sea. Syntypes: ZIN 21934-39 (3, 6, 4, 1, 1, 2). •Synonym of Microcottus sellaris (Gilbert 1896) -- (Neelov 1979:135 [ref. 3152], Lindberg & Krasyukova 1987:219 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S80 [ref. 28300], Parin et al. 2014:260 [ref. 33547]). Current status: Synonym of Microcottus sellaris (Gilbert 1896). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

ochotensis, Zesticelus Yabe [M.] 1995:17, Figs. 1-3 [Japanese Journal of Ichthyology v. 42 (no. 1); ref. 22666] Southwestern Okhotsk Sea off Cape Shiretoko, Japan, 44°27.2'N, 145°06.9'E, depth 1450 meters. Holotype: HUMZ 126133. Paratypes: HUMZ 121626 (1); NSMT-P 46437 (1). •Valid as Zesticelus ochotensis Yabe 1995 -- (Shinohara et al. 1996:176 [ref. 26605], Nakabo 2000:640 [ref. 25086], Sheiko & Fedorov 2000:29 [ref. 25650], Nakabo 2002:640 [ref. 26001], Parin et al. 2002:S90 [ref. 28300], Parin et al. 2014:279 [ref. 33547], Dyldin & Orlov 2022:53 [ref. 38972]). Current status: Valid as Zesticelus ochotensis Yabe 1995. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

octodecemspinosus, Cottus Mitchill [S. L.] 1814:7 [Report, in part, of Samuel L. Mitchill; ref. 3030] New York, U.S.A. No types known. Also in Mitchill 1815:380 [ref. 13292]. Originally as 18-spinosus; correction mandatory; as octodecem-spinosus in Mitchill 1815:380 [ref. 13292]. •Valid as Cottus octodecemspinosus Mitchill 1814 -- (Herrera 1896:63 [ref. 39336] as octodecimspinosus). •Possibly valid as Megalocottus octodecemspinosus (Mitchill 1814) -- (Balakirev et al. 2020:11 [ref. 37873]). •Valid as Myoxocephalus octodecemspinosus (Mitchill 1814) -- (Neelov 1979:123 [ref. 3152], Smith 1986:450 [ref. 22186], Robins & Ray 1986:282 [ref. 23100] as octodecimspinosus, Scott & Scott 1988:502 [ref. 25518], McAllister 1990:200 [ref. 14674], Murdy et al. 1997:147 [ref. 23144], Klein-MacPhee in Collette & Klein-MacPhee 2002:351 [ref. 26158], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Afonso et al. 2016:46, 226 [ref. 36199], Simian et al. 2022:375 [ref. 40776]). Current status: Valid as Myoxocephalus octodecemspinosus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Atlantic. Habitat: brackish, marine.

oculatus, Icelinus Gilbert [C. H.] 1890:87 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., 32°44'30"N, 117°23'W, Albatross station 2935, depth 124 fathoms. Holotype (unique): CAS-SU 80. •Valid as Icelinus oculatus Gilbert 1890 -- (Bolin 1944:19 [ref. 6379], Peden 1984:67 [ref. 5193], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Girard & Smith 2016:556 [ref. 34786], Burton & Lea 2019:67 [ref. 37205], Love et al. 2021:121 [ref. 39279]). Current status: Valid as Icelinus oculatus Gilbert 1890. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: central and southern California (U.S.A.) endemic. Habitat: marine.

oernensis, Cottus quadricornis Lönnberg [E.] 1932:16, Figs. 6 a-b [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Örn, Sweden. Syntypes: (9) ?BMNH 1932.5.17.1 (1), NRM 20323-25 (1, 5, 2). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

okadai, Porocottus Watanabe [M.] 1958:301, Figs. 126, 127, Pl. 22 [Studies of Japanese cottid fishes; ref. 21357] Off Date, Hokkaido, Japan, depth 30 fathoms. Holotype (unique): not researched. Also described as new in Watanabe 1960:90, Pl. 27 (fig. 2) [ref. 11959]. •Valid as Porocottus okadai Watanabe 1958 -- (Watanabe 1960:90 [ref. 11959]). •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Habitat: marine.

okiensis, Alcichthys Mori [T.] 1956:15, Fig. 1 [Japanese Journal of Ichthyology v. 5 (nos 1-2); ref. 12213] Off Oki Island coast, Japan Sea. Holotype (unique): Hyogo Univ. Agric. •Valid as Alcichthys okiensis Mori 1956 -- (Lindberg & Krasyukova 1987:243 [ref. 15964]). •Synonym of Alcichthys elongatus (Steindachner 1881) -- (Nakabo 2002:1526 [ref. 26001], Parin et al. 2002:S68 [ref. 28300], Parin et al. 2014:233 [ref. 33547]). Current status: Synonym of Alcichthys elongatus (Steindachner 1881). Psychrolutidae: Psychrolutinae. Habitat: marine.

ommatistius, Triglops Gilbert [C. H.] 1913:465, Pl. 64 [Proceedings of the United States National Museum v. 44 (no. 1963); ref. 14957] Gloucester, Massachusetts, U.S.A. Holotype: USNM 73921. Paratypes: CAS-SU 23064 (1); USNM 43067-68 (2, 1), 43954 (1), 45547 (6). Type catalog: Böhlke 1953:133 [ref. 12291]. •Synonym of Triglops murrayi Günther 1888, but a valid subspecies -- (Fedorov in Whitehead et al. 1986:1258 [ref. 13677]). •Synonym of Triglops murrayi Günther 1888 -- (Scott & Scott 1988:510 [ref. 25518], Pietsch 1994:368 [ref. 21189], Parin et al. 2014:276 [ref. 33547]). Current status: Synonym of Triglops murrayi Günther 1888. Psychrolutidae: Triglopinae. Habitat: marine.

omodakae, Icelus Tanaka [S.] 1915:616 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 27; ref. 15023] 50 miles off Hinomisaki, Japan, depth 120 fathoms. Syntypes: (3) ZUMT 6420 (1, lost). •Synonym of Icelus cataphractus (Pavlenko 1910) -- (Nelson 1984:28 [ref. 5391]). Current status: Synonym of Icelus cataphractus (Pavlenko 1910). Psychrolutidae: Icelinae. Habitat: marine.

onegensis, Myoxocephalus quadricornis Berg [L. S.] & Popov [A. A.] 1932:158 [Doklady Akademii Nauk SSSR 1932; ref. 16359] Onega Lake and Petrozavodsk, Onega Lake, Russia. Syntypes: ZIN 3842 (1), 9026a (1). Type catalog: Sideleva & Zhidkov 2021:241 [ref. 38705]. Based on Berg 1916:432, fig. 319 and on specimens. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Sideleva & Zhidkov 2021:241 [ref. 38705]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

ontariensis, Triglopsis Jordan [D. S.] & Thompson [W. F.] 1910:75, Fig. 1 [Proceedings of the United States National Museum v. 38 (no. 1728); ref. 10651] Lake Ontario (deep water), off Toronto, Canada. Holotype (unique): USNM 64599. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Scott & Crossman 1998:846 [ref. 24570]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

orientalis, Gymnacanthus tricuspis Schmidt [P. Yu.] 1927:29 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 15821] Kara Sea to Bering Sea. Syntypes: ZIN 14234-35 (1, 2), 20383 (3), 20689-90 (1, 2), 21761-74 (3, 1, 1, 6, 1, 1, 1, 1, 3, 5, 3, 1, 2, 2). •Synonym of Gymnocanthus tricuspis (Reinhardt 1930) -- (Sheiko & Fedorov 2000:27 [ref. 25650] as Gymnacanthus, Parin et al. 2002:S75 [ref. 28300] as Gymnacanthus, Parin et al. 2014:249 [ref. 33547], Mecklenburg et al. 2016:62 [ref. 34440]). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Psychrolutidae: Icelinae. Habitat: marine.

osensis, Stlengis Jordan [D. S.] & Starks [E. C.] 1904:236, Fig. 1 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Ose Point, Suruga Bay, Japan, Albatross station 3788, depth 167 fathoms. Holotype (unique): USNM 50912. •Valid as Stlengis osensis Jordan & Starks 1904 -- (Watanabe 1960:40 [ref. 11959], Fricke & Brunken 1984:46 [ref. 5262], Yabe in Masuda et al. 1984:325 [ref. 6441], Nakabo 2000:630 [ref. 25086], Shinohara et al. 2001:319 [ref. 25995], Nakabo 2002:630 [ref. 26001]). Current status: Valid as Stlengis osensis Jordan & Starks 1904. Psychrolutidae: Oligocottinae. Distribution: Western North Pacific. Habitat: marine.

osimae, Furcina Jordan [D. S.] & Starks [E. C.] 1904:305, Fig. 33 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate, Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7716. Paratypes: USNM 50925 (1). Additional material: BMNH 1905.2.4.364 (1), CAS 58440 [ex IU 10724] (1), CAS-SU 7584 (1). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Furcina osimae Jordan & Starks 1904 -- (Watanabe 1960:139 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:246 [ref. 15964], Kim & Youn 1992:72 [ref. 22287] as osihmae, Nakabo 2000:648 [ref. 25086], Shinohara et al. 2000:177 [ref. 25987], Nakabo 2002:648 [ref. 26001], Choi et al. 2003:262, 574 [ref. 26218], Girard & Smith 2016:556 [ref. 34786], Kwun et al. 2017:141 [ref. 37287], Kim et al. 2020:101 [ref. 39118], Sonoyama et al. 2020:89 [ref. 37637]). Current status: Valid as Furcina osimae Jordan & Starks 1904. Psychrolutidae: Oligocottinae. Distribution: Northwestern Pacific. Habitat: marine.

pachypus, Cottus Günther [A.] 1860:161 [Catalogue of the fishes in the British Museum v. 2; ref. 1963] Port Leopold [Qikiqtaaluk, Somerset Island, Nunavut, Canada]. Syntypes: (2) BMNH 1855.9.19.48 (1), 1855.9.19.761 (1). •Synonym of Myoxocephalus scorpioides (Fabricius 1780) -- (Jordan Evermann 1898:1973 [ref. 2444]). Current status: Synonym of Myoxocephalus scorpioides (Fabricius 1780). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

pacificus, Artediellus Gilbert [C. H.] 1896:416 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bristol Bay; south of Sannak Island; north of Unalaska Island, Alaska. Syntypes: BMNH 1896.7.23.55-58 (4), 1900.7.11.3 (1); MCZ 28328 (12); NMW 6832 (3); CAS-SU 2602 (5), 2604-05 (29, 41); USNM 48605 (1), 48657-58 (6, 3), 48664 (2), 48757 (1), 59410 (5, now 1); ZMUO J1403 (12), J1404 (10). Type catalog: Böhlke 1953:127 [ref. 12291], Pethon 1969:7 [ref. 19268]. •Valid as Artediellus pacificus Gilbert 1896 -- (Neelov 1979:154 [ref. 3152], Eschmeyer & Herald 1983:161 [ref. 9277], Nelson 1986:43 [ref. 5808], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:492 [ref. 25968], Parin et al. 2002:S71 [ref. 28300], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:84 [ref. 37547], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:121 [ref. 32708], Parin et al. 2014:238 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Love et al. 2021:118 [ref. 39279]). Current status: Valid as Artediellus pacificus Gilbert 1896. Psychrolutidae: Psychrolutinae. Distribution: North Pacific and adjacent Arctic: northern Sea of Japan and Sea of Okhotsk north to northern Bering Sea, east to southeastern Alaska (U.S.A.). Habitat: marine.

pacificus, Triglops pingelii Schmidt [P. Yu.] 1930:515, Fig. 1a [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 30 (no. 4) (for 1929); ref. 16144] Okhotsk and Bering seas. Syntypes: (11) ZIN 21462-63 (4, 4), 21468 (1) •Synonym of Triglops pingelii Reinhardt 1837 -- (Pietsch 1994:374 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Parin et al. 2002:S88 [ref. 28300], Parin et al. 2014:277 [ref. 33547]). Current status: Synonym of Triglops pingelii Reinhardt 1837. Psychrolutidae: Triglopinae. Habitat: marine.

paradoxus, Psychrolutes Günther [A.] 1861:516 [Catalogue of the fishes in the British Museum v. 3; ref. 1964] Gulf of Georgia, Vancouver Island, British Columbia, Canada. Holotype (unique): BMNH 1860.8.24.18. •Valid as Psychrolutes paradoxus Günther 1861 -- (Eschmeyer & Herald 1983:180 [ref. 9277], Yabe in Masuda et al. 1984:330 [ref. 6441], Lindberg & Krasyukova 1987:263 [ref. 15964], Matarese et al. 1989:436 [ref. 26885], McAllister 1990:204 [ref. 14674], Fricke 1990:407 [ref. 15492], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:653 [ref. 25086], Sheiko & Fedorov 2000:30 [ref. 25650], Mecklenburg et al. 2002:524 [ref. 25968], Sokolovsky & Sokolovskaya 2001:425 [ref. 25414], Nakabo 2002:653 [ref. 26001], Choi et al. 2003:276, 5846 [ref. 26218], Parin et al. 2002:S94 [ref. 28300], Fedorov et al. 2003:87 [ref. 27579], Nelson et al. 2004:123 [ref. 27807], Love et al. 2005:95 [ref. 37547], Shinohara et al. 2011:46 [ref. 31715], Maslenikov et al. 2013:12 [ref. 37947], Page et al. 2013:126 [ref. 32708], Shinohara et al. 2014:254 [ref. 33330], Mecklenburg & Steinke 2015:164 [ref. 34033], Pietsch & Orr 2015:51 [ref. 34694], Kells et al. 2016:208 [ref. 35888], Kim et al. 2020:107 [ref. 39118], Sonoyama et al. 2020:91 [ref. 37637], Love et al. 2021:125 [ref. 39279], Dyldin & Orlov 2022:55 [ref. 38972]). Current status: Valid as Psychrolutes paradoxus Günther 1861. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Sea of Japan and Sea of Okhotsk north to northern Bering Sea, then south to Puget Sound (Washington, U.S.A.). Habitat: marine.

parvulus, Myoxocephalus Gilbert [C. H.] & Burke [C. V.] 1912:59, Fig. 12 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Tidepool at Preobrazhenskoi Bay, Medny Island, Commander Islands, Bering Sea. Holotype: USNM 74375. Paratypes: CAS-SU 22369 (1), USNM 70829 (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Synonym of Microcottus sellaris (Gilbert 1896) -- (Neelov 1979:135 [ref. 3152], Lindberg & Krasyukova 1987:219 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:483 [ref. 25968], Parin et al. 2002:S80 [ref. 28300], Parin et al. 2014:260 [ref. 33547], Mecklenburg et al. 2016:75 [ref. 34440]). Current status: Synonym of Microcottus sellaris (Gilbert 1896). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

patris, Acanthocottus Storer [H. R.] 1850:250, Pl. 7 (fig. 2) [Boston Journal of Natural History v. 6 (no. 2) (art. 15); ref. 18109] Great Mecatina Island Harbor and Red Bay, Straits of Belle Isle, Labrador, Canada, depth 10-20 fathoms. Syntypes: (2). •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Neelov 1973:595 [ref. 7219] as Gymnacanthus). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Psychrolutidae: Icelinae. Habitat: marine.

percoides, Pseudoblennius Günther [A.] 1861:297 [Catalogue of the fishes in the British Museum v. 3; ref. 1964] Omura, near Nagasaki, Japan. No types known. Based on Temminck & Schlegel 1844: Pl. 79A (figs. 2-3) [ref. 4372]. See Nakabo 2002:1526 [ref. 26001]. •Valid as Pseudoblennius percoides Günther 1861 -- (Watanabe 1960:147 [ref. 11959], Nakabo in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:251 [ref. 15964], Kimura et al. 1988:19 [ref. 6692], Kim & Youn 1992:74 [ref. 22287], Mizuno et al. 1999:8 [ref. 25311], Nakabo 2000:650 [ref. 25086], Shinohara et al. 2000:177 [ref. 25987], Nakabo 2002:650 [ref. 26001], Choi et al. 2003:263, 577 [ref. 26218], Kwun et al. 2017:141 [ref. 37287], Kim et al. 2020:103 [ref. 39118], Sonoyama et al. 2020:90 [ref. 37637]). Current status: Valid as Pseudoblennius percoides Günther 1861. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: southern Korea and Japan. Habitat: marine.

perminovi, Icelus Taranetz [A. Ya.] 1936:151 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (pt 13) (No. 3, art. 107); ref. 6509] Okhotsk Sea, 56°12'N, 143°49'E. Holotype (unique): No types at ZIN. •Valid as Icelus perminovi Taranetz 1936 -- (Nelson 1984:34 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:634 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Nakabo 2002:634 [ref. 26001], Parin et al. 2002:S78 [ref. 28300], Fedorov et al. 2003:72 [ref. 27579], Parin et al. 2014:255 [ref. 33547], Dyldin & Orlov 2022:46 [ref. 38972] with question). Current status: Valid as Icelus perminovi Taranetz 1936. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific. Habitat: marine.

phrictus, Psychrolutes Stein [D. L.] & Bond [C. E.] 1978:3, Figs. 1-2 [Contributions in Science (Los Angeles) No. 296; ref. 8873] North Pacific, 44°44'18"N, 125°41'18"W, depth 2800 meters. Holotype: USNM 216253. Paratypes: CAS 32580 (1); LACM 30807-4 (1), 34185-1 (2), 34338-1 (1), 35230-1 (3), 35232-1 (1), 35234-1 (1), 35561-1 (1); OSUO 1524 (1), 2020 (1), 2021 (1), 2040 (1, c&s), 2029 (1), 2057 (1), 2221 (1). Additional material (3). •Valid as Psychrolutes phrictus Stein & Bond 1978 -- (Eschmeyer & Herald 1983:180 [ref. 9277], Yabe in Amaoka et al. 1983:159, 215, 289, 340 [ref. 8379], Yabe in Masuda et al. 1984:331 [ref. 6441], McAllister 1990:204 [ref. 14674], Fricke 1990:407 [ref. 15492], Nelson 1995:69 [ref. 22668], Shinohara et al. 1996:177 [ref. 26605], Nakabo 2000:653 [ref. 25086], Sheiko & Fedorov 2000:30 [ref. 25650], Mecklenburg et al. 2002:523 [ref. 25968], Nakabo 2002:653 [ref. 26001], Parin et al. 2002:S95 [ref. 28300], Love et al. 2005:95 [ref. 37547], Maslenikov et al. 2013:13 [ref. 37947], Shinohara et al. 2005:428 [ref. 28370], Parin et al. 2014:303 [ref. 33547], Afonso et al. 2016:56, 258 [ref. 36199], Burton & Lea 2019:70 [ref. 37205], Love et al. 2021:125 [ref. 39279], Dyldin & Orlov 2022:55 [ref. 38972]). Current status: Valid as Psychrolutes phrictus Stein & Bond 1978. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Sea of Okhotsk and Pacific coast of Honshu (Japan) north to Bering Sea, then south to southern California (U.S.A.). Habitat: marine.

pietschi, Icelinus Yabe [M.], Soma [A.] & Amaoka [K.] 2001:65, Figs. 1-3 [Ichthyological Research v. 48 (no. 1); ref. 25246] Tide pool on east side of Chirip Peninsula, Okhotsk coast of Iturup Island, 45°19.8'N, 148°00.3'E, depth 0.5-1 meters. Holotype: HUMZ 151942. Paratypes: HUMZ 151943-44 (1, 1), NSMT-P 58459 (1), UW 43755 (1). •Valid as Icelinus pietschi Yabe, Soma & Amaoka 2001 -- (Rosenblatt & Smith 2004:559 [ref. 27912], Balanov & Panchenko 2023:484 [828] [ref. 40446]). Current status: Valid as Icelinus pietschi Yabe, Soma & Amaoka 2001. Psychrolutidae: Oligocottinae. Distribution: Northwestern Pacific: southern Sea of Okhotsk and northern Sea of Japan (Russia), including Kuril Islands. Habitat: marine.

pietschmanni, Triglops pingelii Jensen [A. S.] 1944:15, 29, Pls. 1 (figs. a-b), 2 (fig. a) [Spolia Zoologica Musei Hauniensis v. 4; ref. 13060] Southwestern Greenland. Syntypes: (19) ZMUC P81371 (1), P81374-385 (12), P81482 (1), P81477 (1). Type catalog: Nielsen 1974:82 [ref. 9588]. •Synonym of Triglops murrayi Günther 1888 -- (Neelov 1973:602 [ref. 7219], Pietsch 1994:369 [ref. 21189], Parin et al. 2014:276 [ref. 33547]). Current status: Synonym of Triglops murrayi Günther 1888. Psychrolutidae: Triglopinae. Habitat: marine.

pinetorum, Ricuzenius Jordan [D. S.] & Starks [E. C.] 1904:243, Fig. 5 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Kinkwazan Island, in Matsushima Bay, Japan, Albatross station 3773. Holotype (unique): USNM 50915. •Valid as Ricuzenius pinetorum Jordan & Starks 1904 -- (Watanabe 1960:19 [ref. 11959], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:185 [ref. 15964], Kim & Youn 1992:69 [ref. 22287], Shinohara et al. 1996:176 [ref. 26605], Nakabo 2000:632 [ref. 25086], Nakabo 2002:632 [ref. 26001], Choi et al. 2003:263, 578 [ref. 26218], Shinohara et al. 2014:251 [ref. 33330], Kim et al. 2020:103 [ref. 39118]). Current status: Valid as Ricuzenius pinetorum Jordan & Starks 1904. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Korea, Japan, Russia. Habitat: marine.

pingelii, Triglops Reinhardt [J. C. H.] 1837:114, 118 [32, 36] [Det Kongelige Danske videnskabernes selskabs naturvidenskabelige og mathematiske afhandlinger v. 7; ref. 3691] Quanneoen, south of Frederikshaab, western Greenland. Lectotype: ZMUC 1. Paralectotypes: ZMB 672 [ex ZMUC] (1), ZMUC 2-3 (2). Type catalog: Nielsen 1974:82 [ref. 9588], Paepke & Fricke 1992:284 [ref. 19946]. On pp. 32 & 36 of separate. Described but not named in 1830:17 [ref. 19480]. Lectotype established by Pietsch 1994:374 [ref. 21189]. •Valid as Triglops pingeli Reinhardt 1837, subspecies pingeli Reinhardt 1837 -- (Nakabo 2000:628 [ref. 25086], Nakabo 2002:628 [ref. 26001]). •Valid as Triglops pingelii Reinhardt 1837 -- (Neelov 1973:601 [ref. 7219], Eschmeyer & Herald 1983:184 [ref. 9277], Yabe in Masuda et al. 1984:325 [ref. 6441], Fedorov in Whitehead et al. 1986:1259 [ref. 13677], Robins & Ray 1986:284 [ref. 23100], Lindberg & Krasyukova 1987:174 [ref. 15964], Scott & Scott 1988:511 [ref. 25518], McAllister 1990:202 [ref. 14674], Kim & Youn 1992:65 [ref. 22287], Pietsch 1994:374 [ref. 21189], Yabe in Okamura et al. 1995:168 [ref. 22531], Sokolovskaya et al. 1998:10 [ref. 24670], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:428 [ref. 25968], Choi et al. 2003:263, 578 [ref. 26218] as pingeli, Parin et al. 2002:S88 [ref. 28300], Fedorov et al. 2003:82 [ref. 27579], Nelson et al. 2004:122 [ref. 27807], Love et al. 2005:93 [ref. 37547], Pietsch & Orr 2006:244 [ref. 28666], Mecklenburg et al. 2007:177 [ref. 29448], Møller et al. 2010:53 [ref. 31169], Shinohara et al. 2011:44 [ref. 31715], Page et al. 2013:125 [ref. 32708], Dolgov 2013:918 [ref. 33112], Shinohara et al. 2014:251 [ref. 33330], Mecklenburg et al. 2014:137, 145 [ref. 33463], Parin et al. 2014:277 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Pietsch & Orr 2015:45 [ref. 34694], Tohkairin et al. 2015:7 [ref. 35331], Afonso et al. 2016:52, 291 [ref. 36199], Kells et al. 2016:194 [ref. 35888], Mecklenburg et al. 2016:90 [ref. 34440], Mecklenburg et al. 2018:229 [ref. 36370], Love et al. 2021:127 [ref. 39279], Chernova et al. 2022:6 [ref. 39177], Dyldin & Orlov 2022:53 [ref. 38972], Tokranov et al. 2022:3 [ref. 39739]). Current status: Valid as Triglops pingelii Reinhardt 1837. Psychrolutidae: Triglopinae. Distribution: Circumpolar in Arctic Ocean, adjacent North Pacific and North Atlantic. Habitat: marine.

pistilliger, Cottus Pallas [P. S.] 1814:143, Pl. 20 (figs. 3-4) [Zoographia Rosso-Asiatica v. 3; ref. 3351] Unalaska Island and Port Avatsch. Syntypes: ZMB 8213 (1). Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. Type information: Svetovidov 1978:28 [ref. 6025]. •Valid as Gymnocanthus pistilliger (Pallas 1814) -- (Watanabe 1960:56 [ref. 11959], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:199 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:629 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Mecklenburg et al. 2002:465 [ref. 25968], Nakabo 2002:629 [ref. 26001], Choi et al. 2003:262, 575 [ref. 26218], Parin et al. 2002:S75 [ref. 28300] as Gymnacanthus, Fedorov et al. 2003:69 [ref. 27579] as Gymnacanthus, Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:87 [ref. 37547], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:248 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Mecklenburg et al. 2016:59 [ref. 34440], Kells et al. 2016:172 [ref. 35888], Dyldin & Orlov 2017:63 [ref. 35138], Balanov & Matveev 2018:366 [430] [ref. 36463], Mecklenburg et al. 2018:197 [ref. 36370], Kim et al. 2020:101 [ref. 39118], Dyldin et al. 2021:63 [ref. 38804], Dyldin & Orlov 2022:44 [ref. 38972], Love et al. 2021:121 [ref. 39279]). Current status: Valid as Gymnocanthus pistilliger (Pallas 1814). Psychrolutidae: Icelinae. Distribution: North Pacific and adjacent Arctic: Sea of Japan (Korea, Japan, Russia) and Sea of Okhotsk north to Bering Sea, then south to southeastern Alaska (U.S.A.). Habitat: brackish, marine.

platycephalus, Cottus Pallas [P. S.] 1814:135 [Zoographia Rosso-Asiatica v. 3; ref. 3351] Kamchatka, Russia; America; Okhotsk Sea. Lectotype: ZMB 4756. Syntypes: ZMB 4756 (1, dry). Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. Lectotype designated by Svetovidov 1978:27 [ref. 6025]. •Valid as Myoxocephalus platycephalus (Pallas 1814) -- (Berg 1949:1127 [ref. 20662]). •Valid as Megalocottus platycephalus (Pallas 1814) -- (Neelov 1979:130 [ref. 3152], Pietsch et al. 2001:145 [ref. 25688], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:152 [ref. 23595], Sheiko & Fedorov 2000:28 [ref. 25650], Zemnukhov et al. 2001:420 [ref. 25412], Mecklenburg et al. 2002:475 [ref. 25968], Parin et al. 2002:S79 [ref. 28300], Fedorov et al. 2003:73 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Bogutskaya & Naseka 2004:190 [ref. 28183], Love et al. 2005:89 [ref. 37547], Mecklenburg et al. 2007:183 [ref. 29448], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:257 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Mecklenburg et al. 2016:72 [ref. 34440], Kells et al. 2016:182 [ref. 35888], Mecklenburg et al. 2018:209 [ref. 36370], Love et al. 2021:123 [ref. 39279]). Current status: Valid as Megalocottus platycephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific and Arctic region: northern Sea of Japan and Sea of Okhotsk north to Bering Sea. Habitat: brackish, marine.

pleurostictus, Triglops Cope [E. D.] 1865:81 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 17; ref. 907] Godhaven, western Greenland. Holotype (unique): ANSP (not found). Type catalog: Böhlke 1984:164 [ref. 13621]. •Synonym of Triglops pingelii Reinhardt 1837 -- (Neelov 1973:601 [ref. 7219], Pietsch 1994:374 [ref. 21189]). Current status: Synonym of Triglops pingelii Reinhardt 1837. Psychrolutidae: Triglopinae. Habitat: marine.

plumarius, Archistes Jordan [D. S.] & Gilbert [C. H.] in Jordan & Evermann 1898:1900 [Bulletin of the United States National Museum No. 47; ref. 2444] Ushishir Island, Kurils [Kuril] Islands. Holotype (unique): USNM 48233. Also in Jordan & Gilbert 1899:454, Pl. 53 [ref. 2478]. •Valid as Archistes plumarius Jordan & Gilbert 1898 -- (Sheiko & Fedorov 2000:26 [ref. 25650], Mecklenburg et al. 2002:442 [ref. 25968], Parin et al. 2002:S68 [ref. 28300], Love et al. 2005:84 [ref. 37547], Parin et al. 2014:234 [ref. 33547], Love et al. 2021:117 [ref. 39279]). Current status: Valid as Archistes plumarius Jordan & Gilbert 1898. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Kuril Islands; Commander Islands (Russia). Habitat: marine.

polaris, Cottus Sabine [E.] 1824:213 [ccxiii] [A supplement to the appendix of Captain Parry's voyage; ref. 18649] North Georgia, east side Boothia Peninsula, Northwest Territories, Canada. No types known. •Synonym of Triglopsis quadricornis (Linnaeus 1758), but a valid subspecies -- (Chereshnev 1996:606 [ref. 24659]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Lee et al. 1980:826 [ref. 22416], Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

polyacanthocephalus, Cottus Pallas [P. S.] 1814:133 [Zoographia Rosso-Asiatica v. 3; ref. 3351] No definite locality [= Kodiak, Aleutian Islands]. Holotype (unique): ZMB 4757 (dry). Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. Type information: Svetovidov 1978:27 [ref. 6025]. •Valid as Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Watanabe 1960:66 [ref. 11959], Neelov 1979:121 [ref. 3152], Eschmeyer & Herald 1983:176 [ref. 9277], Yabe in Masuda et al. 1984:326 [ref. 6441], Lindberg & Krasyukova 1987:210 [ref. 15964], McAllister 1990:200 [ref. 14674], Sokolovsky & Sokolovskaya 1997:57 [ref. 24691], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:645 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:482 [ref. 25968], Nakabo 2002:645 [ref. 26001], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:77 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Mecklenburg et al. 2007:176 [ref. 29448], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:261 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Pietsch & Orr 2015:43 [ref. 34694], Tohkairin et al. 2015:7 [ref. 35331], Mecklenburg et al. 2016:78 [ref. 34440], Kells et al. 2016:184 [ref. 35888], Dyldin & Orlov 2017:66 [ref. 35138], Mecklenburg et al. 2018:215 [ref. 36370], Kim et al. 2020:103 [ref. 39118], Dyldin et al. 2021:66 [ref. 38804], Love et al. 2021:123 [ref. 39279], Dyldin & Orlov 2022:49 [ref. 38972]). Current status: Valid as Myoxocephalus polyacanthocephalus (Pallas 1814). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific and adjacent Arctic: eastern Sea of Japan and sea of Okhotsk north to southern Bering Sea, then south to Puget Sound, Washington (U.S.A.). Habitat: marine.

porosus, Cottus Valenciennes [A.] in Cuvier & Valenciennes 1832:498 [Histoire naturelle des poissons; ref. 1000] Baffin Bay, between western Greenland and Baffin Island. Holotype (unique): MNHN 0000-6761. Type catalog: Blanc & Hureau 1968:51 [ref. 20738]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

primoricus, Cottiusculus Prokofiev [A. M.] 2020:11, Figs. 1-5 [Munis Entomology & Zoology v. 15 (no. 1); ref. 37765] Peter the Great Bay, Primorsky Krai, Sea of Japan, Russia. Holotype: IOM "RPR-3098". Paratypes: IOM (1 + 4). •Valid as Cottiusculus primoricus Prokofiev 2020. Current status: Valid as Cottiusculus primoricus Prokofiev 2020. Psychrolutidae: Psychrolutinae. Distribution: Peter the Great Bay, Sea of Japan, Russia. Habitat: marine.

profundorum, Acanthocottus Gilbert [C. H.] 1896:423, Pl. 27 (upper) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bering Sea, north of Unalaska Island, Albatross station 3329, depth 399 fathoms. Syntypes (incl. potential lectotype): USNM 48589 (1), 53030 (1). Lectotype established (as figured specimen) in caption to Pl. 300, p. 3293 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. •Valid as Zesticelus profundorum (Gilbert 1896) -- (Bolin 1944:95 [ref. 6379], Neelov 1979:161 [ref. 3152], Eschmeyer & Herald 1983:184 [ref. 9277], Yabe & Okamura 1993:370 [ref. 22281], Yabe 1995:19 [ref. 22666], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:487 [ref. 25968], Parin et al. 2002:S90 [ref. 28300], Fedorov et al. 2003:83 [ref. 27579], Love et al. 2005:93 [ref. 37547], Parin et al. 2014:279 [ref. 33547], Burton & Lea 2019:68 [ref. 37205], Love et al. 2021:127 [ref. 39279]). Current status: Valid as Zesticelus profundorum (Gilbert 1896). Psychrolutidae: Psychrolutinae. Distribution: North Pacific: northern Kuril islands and Sea of Okhotsk north to southeastern Bering Sea, east to Aleutian Islands (U.S.A.). Habitat: marine.

pugetensis, Artedius Steindachner [F.] 1876:181 [133], Pl. 14 (fig. 2-2a) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 74 (1. Abth.); ref. 4225] Fox Island near near Steilacom, northern Puget Sound; also San Francisco. Syntypes: (several) NMW 6795-96 (2, 3), 11662-66 (4, 2, 2, 2, 1), 11668 (1), 14783-84 (1, 1), 59720 (?), 75544 (1). On p. 133 of separate. •Valid as Chitonotus pugetensis (Steindachner 1876) -- (Bolin 1944:19 [ref. 6379], Eschmeyer & Herald 1983:164 [ref. 9277], Matarese et al. 1989:388 [ref. 26885], McAllister 1990:196 [ref. 14674], Mecklenburg et al. 2002:437 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:85 [ref. 37547], Page et al. 2013:121 [ref. 32709], Pietsch & Orr 2015:39 [ref. 34694], Kells et al. 2016:166 [ref. 35888], Burton & Lea 2019:65 [ref. 37205], Love et al. 2021:119 [ref. 39279]). Current status: Valid as Chitonotus pugetensis (Steindachner 1876). Psychrolutidae: Icelinae. Distribution: Northeastern Pacific: Alaska (U.S.A.) south to Pacific coast of southern Baja California Sur (Mexico). Habitat: marine.

pustulosa, Gilbertidia Schmidt [P. Yu.] 1937:280 [Doklady Akademii Nauk SSSR v. 15 (no. 5); ref. 21497] Okhotsk Sea, 55°31'N, 149°10'N. Syntypes: ZIN 24462 (1), 24830 (1). Could be described earlier as Platycottus pustulosus Schmidt 1935, reference not found. •Valid as Gilbertidia pustulosa Schmidt 1937 -- (Lindberg & Krasyukova 1987:264 [ref. 15964], Nakabo 2000:652 [ref. 25086], Sheiko & Fedorov 2000:30 [ref. 25650], Nakabo 2002:652 [ref. 26001], Parin et al. 2002:S93 [ref. 28300], Parin et al. 2014:301 [ref. 33547], Kim et al. 2020:107 [ref. 39118]). •Valid as Psychrolutes pustulosus (Schmidt 1937) -- (Yabe et al. 1995 [ref. 22110], Nazarkin et al. 2014:178 [ref. 33578], Park et al. 2015:239 [ref. 36311], Dyldin & Orlov 2022:55 [ref. 38972]). Current status: Valid as Psychrolutes pustulosus (Schmidt 1937). Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia) north to Okhotsk Sea. Habitat: marine.

pygmaeus, Cottus quadricornis Lönnberg [E.] 1933:9, Pl. 1 (fig. 2) [Arkiv för Zoologi v. 24 A (pt 3) (no. 12) [1932]; ref. 22149] Lake Puruvesi, Finland. Syntypes: MZH. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies onegensis Berg & Popov 1932 -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

quadratus, Porocottus Bean [T. H.] in Jordan & Evermann 1898:1998 [Bulletin of the United States National Museum No. 47; ref. 2444] Bering Island, Commander Islands. Holotype (unique): USNM 33875. Author is T. H. Bean. Also appeared as Bean in Jordan & Gilbert 1899:466, Pl. 67a [ref. 2478]. •Synonym of Microcottus sellaris (Gilbert 1896) -- (Neelov 1979:134 [ref. 3152], Lindberg & Krasyukova 1987:218 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:483 [ref. 25968], Parin et al. 2002:S80 [ref. 28300], Parin et al. 2014:259 [ref. 33547], Mecklenburg et al. 2016:75 [ref. 34440]). Current status: Synonym of Microcottus sellaris (Gilbert 1896). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

quadricornis, Cottus Linnaeus [C.] 1758:264 [Systema Naturae, Ed. X v. 1; ref. 2787] Baltic Sea. Syntypes: NRM 2491-92 (1, 1). Type catalog: Fernholm & Wheeler 1983:235-236 [ref. 20707] who suggest that the largest syntype would make the best lectotype. •Valid as Triglopsis quadricornis (Linnaeus 1758) -- (Neelov 1979:133 [ref. 3152], Fedorov in Whitehead et al. 1986:1260 [ref. 13677], Kottelat 1997:170 [ref. 22952], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:154 [ref. 23595], Novikov et al. 2000:161 [ref. 40929], Bogutskaya et al. 2001:46 [ref. 26178], Parin et al. 2002:S89 [ref. 28300], Bogutskaya & Naseka 2004:191 [ref. 28183], Fricke 2007:29 [ref. 30577], Kottelat & Freyhof 2007:519 [ref. 29996], Sideleva et al. 2022:3 [ref. 40100]). •Possibly valid as Megalocottus quadricornis (Linnaeus 1758) -- (Balakirev et al. 2020:11 [ref. 37873]). •Valid as Myoxocephalus quadricornis (Linnaeus 1758) -- (Berg 1949:1128 [ref. 20662], Neelov 1973:597 [ref. 7219], Lee et al. 1980:826 [ref. 22416], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:504 [ref. 25518], McAllister 1990:200 [ref. 14674], Page & Burr 1991:245 [ref. 18983], Scott & Crossman 1998:842 [ref. 24570], Mecklenburg et al. 2002:477 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Scharpf 2008:11 [ref. 30399], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212], Page & Burr 2011:468 [ref. 31215], Page et al. 2013:124 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:263 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:46, 227 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:80 [ref. 34440], Mecklenburg et al. 2018:217 [ref. 36370], Love et al. 2021:123 [ref. 39279]). Current status: Valid as Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Distribution: Circumpolar in Arctic Ocean, in Pacific and north Atlantic. Habitat: freshwater, brackish, marine.

quadrifilis, Porocottus Gill [T. N.] 1859:166 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 11; ref. 1760] Bering Strait. Holotype (unique): USNM 6627 (badly damaged). See Mecklenburg et al. 2002:405 and 484 [ref. 25968] •Valid as Porocottus quadrifilis Gill 1859 -- (Neelov 1979:136 [ref. 3152], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:484 [ref. 25968], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S84 [ref. 28300], Love et al. 2005:91 [ref. 37547], Parin et al. 2014:269 [ref. 33547], Love et al. 2021:124 [ref. 39279]). Current status: Valid as Porocottus quadrifilis Gill 1859. Psychrolutidae: Myoxocephalinae. Distribution: North Pacific: probably Bering Strait; known from only one record with uncertain locality. Habitat: marine.

quadriseriatus, Artedius Lockington [W. N.] 1880:330 [Proceedings of the United States National Museum v. 2 (no. 97); ref. 2815] San Francisco, California, U.S.A. Syntypes: USNM 23503 (2). •Valid as Icelinus quadriseriatus (Lockington 1880) -- (Bolin 1944:35 [ref. 6379], Eschmeyer & Herald 1983:173 [ref. 9277], Peden 1984:79 [ref. 5193], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:178 [ref. 35888], Burton & Lea 2019:67 [ref. 37205], Love et al. 2021:121 [ref. 39279]). Current status: Valid as Icelinus quadriseriatus (Lockington 1880). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: northern California (U.S.A.) south to southern Baja California (Mexico). Habitat: marine.

raninus, Myoxocephalus Jordan [D. S.] & Starks [E. C.] 1904:277, Fig. 20 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Aomori, Japan. Holotype: CAS-SU 7708. Paratypes: ?CAS 56330 [ex IU 10595] (1); USNM 50920 (13); BMNH 1905.2.4.373-374 (2). Additional material: CAS-SU 7591 (15, missing), 7592 (5), 7595 (5), 7938 (now 83); USNM 51285-86 (1, 1), 23815 [ex Iowa Mus.] (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Watanabe 1960:82 [ref. 11959], Neelov 1979:121 [ref. 3152], Lindberg & Krasyukova 1987:207 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:480 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:265 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

rastrinoides, Icelus Taranetz [A. Ya.] 1936:150 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (pt 13) (No. 3, art. 107); ref. 6509] Cave Cape; Olympyad Bay; and 45°05'N, 138°06'E, depth 230-411 meters, coast of Primorye, Russia. Syntypes: (8) ZIN 21445 (3). First mentioned by Schmidt 1935:416 [ref. 3939] in the subgenus Ochotskia Schmidt 1915. See also Nakabo 2002:1525 [ref. 26001]. •Valid as Icelus rastrinoides Taranetz 1936 -- (Nelson 1984:29 [ref. 5391], Lindberg & Krasyukova 1987:193 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Parin et al. 2002:S78 [ref. 28300], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:255 [ref. 33547], Dyldin & Orlov 2022:47 [ref. 38972], Kai et al. 2023:16 [ref. 39903]). Current status: Valid as Icelus rastrinoides Taranetz 1936. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Russia, Japan). Habitat: marine.

recalvus, Blennicottus Greeley [A. W.] 1899:9, Fig. 1 [Bulletin of the U. S. Fish Commission v. 19 [1899]; ref. 1889] Pacific Grove, California, U.S.A. Holotype: CAS-SU 6068. Paratypes: (4) CAS-SU 4245 (1, discarded). Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Clinocottus recalvus (Greeley 1899) -- (Bolin 1944:79 [ref. 6379], Eschmeyer & Herald 1983:167 [ref. 9277], Matarese et al. 1989:428 [ref. 26885], Nelson et al. 2004:119 [ref. 27807], Love et al. 2005:86 [ref. 37547], Page et al. 2013:121 [ref. 32708], Kells et al. 2016:168 [ref. 35888], Burton & Lea 2019:66 [ref. 37205], Love et al. 2021:120 [ref. 39279]). Current status: Valid as Clinocottus recalvus (Greeley 1899). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: southern Oregon (U.S.A.) south to Pacific coast of central Baja California (Mexico). Habitat: marine.

regulus, Astrocottus Tsuruoka [O.], Maruyama [S.] & Yabe [M.] 2008:34, Figs. 3C, 6 [Bulletin of the National Museum of Nature and Science (Ser. A) Supplement No. 2; ref. 29568] Off Oumu, Okhotsk coast, Hokkaido, Japan, 44°33.89'N, 143°07.10'E, depth 46 meters. Holotype: HUMZ 190094. Paratypes: CAS 225634 [ex HUMZ 190085, 190087] (2); HUMZ 91981 (1), 91989 (1), 98614-15 (2), 189828-30 (3), 190081-84 (4), 191778 (1), 192515 (1), 195516 (1), 195746 (1), 195747 (1), 195748 (1), 195749-53 (5); NSMT-P 76686-89 [ex HUMZ 195759-62] (4); USNM 384159 [ex HUMZ 190086, 190088] (2). •Valid as Astrocottus regulus Tsuruoka, Maruyama & Yabe 2008 -- (Parin et al. 2014:239 [ref. 33547], Dyldin et al. 2021:61 [ref. 38804], Dyldin & Orlov 2022:40 [ref. 38972]). Current status: Valid as Astrocottus regulus Tsuruoka, Maruyama & Yabe 2008. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: off Usujiri, Okhotsk coast of Hokkaido, and off Miyagi and Fukushima Prefecture, Pacific coast, and off Obira, Hokkaido, Sea of Japan. Habitat: marine.

relicta, Cottus quadricornis var. Lilljeborg [N.] 1891:149 [Sveriges och Norges fauna. Fiskarne. v. 1; ref. 21766] Lake Vättern, Sweden. No types known. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies relictus (Lilljeborg 1891) -- (Berg 1949:1133 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

rhodorus, Ascelichthys Jordan [D. S.] & Gilbert [C. H.] 1880:264 [Proceedings of the United States National Museum v. 3 (no. 144); ref. 2367] Wadda Island, Neah Bay, near Cape Flattery, Washington, U.S.A. Syntypes: (about 200) ANSP 12006-07 (2); MNHN A-3274 (4); MTD F 151 [ex USNM] (3); UMMZ 162182 (3); USNM 27331 (25, 4 in skeletal coll.); ZMB 11607 [ex USNM 27331] (3); ZMUB 1295 [ex USNM 27331] (1). Type catalog: Blanc & Hureau 1968:48 [ref. 20738], Frøiland 1979:148 [ref. 21547], Böhlke 1984:65 [ref. 13621], Paepke & Fricke 1992:282 [ref. 19946], Zarske 2003:31 [ref. 27474]. •Valid as Ascelichthys rhodorus Jordan & Gilbert 1880 -- (Bolin 1944:88 [ref. 6379], Eschmeyer & Herald 1983:163 [ref. 9277], Matarese et al. 1989:386 [ref. 26885], McAllister 1990:194 [ref. 14674], Mecklenburg et al. 2002:419 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:85 [ref. 37547], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:166 [ref. 35888], Burton & Lea 2019:65 [ref. 37205], Love et al. 2021:119 [ref. 39279]). Current status: Valid as Ascelichthys rhodorus Jordan & Gilbert 1880. Psychrolutidae: Psychrolutinae. Distribution: Northeastern Pacific: Prince William Sound, northern Gulf of Alaska (U.S.A.) south to central California (U.S.A.). Habitat: marine.

rimensis, Rusciculus Greeley [A. W.] 1899:13, Fig. 3 [Bulletin of the U. S. Fish Commission v. 19 [1899]; ref. 1889] Point Lobos, near Carmel, California, U.S.A. Holotype: CAS-SU 6067. Paratypes: CAS-SU 6067 (1 of 2). Type catalog: Böhlke 1953:132 [ref. 12291]. Holotype established in table on p. 14 if specimen is identifiable. •Valid as Oligocottus rimensis (Greeley 1899) -- (Bolin 1944:63 [ref. 6379], Eschmeyer & Herald 1983:178 [ref. 9277], McAllister 1990:201 [ref. 14674], Mecklenburg et al. 2002:499 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:43 [ref. 34694], Kells et al. 2016:186 [ref. 35888], Burton & Lea 2019:67 [ref. 37205], Love et al. 2021:124 [ref. 39279]). Current status: Valid as Oligocottus rimensis (Greeley 1899). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: southeastern Alaska (U.S.A.) south to Pacific coast of northern Baja California (Mexico). Habitat: marine.

robustus, Clypeocottus Ayres [W. O.] 1854:194 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] California, U.S.A. No types known. Appeared first as above on 13 Oct.; then in Ayres 1854:11 [ref. 18923] where Ayres mentions that Girard's genus Aspicottus and species A. bison are synonyms and have priority. •Synonym of Enophrys bison (Girard 1854) -- (Bolin 1944:91 [ref. 6379], Sandercock & Wilimovsky 1968:844 [ref. 26414]). Current status: Synonym of Enophrys bison (Girard 1854). Psychrolutidae: Myoxocephalinae. Habitat: marine.

rubellio, Eximia Greeley [A. W.] 1899:18, Fig. 5 [Bulletin of the U. S. Fish Commission v. 19 [1899]; ref. 1889] Monterey Bay, California, U.S.A. Holotype: ? CAS-SU 6066. Paratypes: (5) CAS-SU 3642 (1, discarded), 4048 (2, ? missing). Type catalog: Böhlke 1953:129 [ref. 12291]. Type established in table on p. 19 and in Fig. 5 if specimen is identifiable. •Valid as Oligocottus rubellio (Greeley 1899) -- (Bolin 1944:70 [ref. 6379], Eschmeyer & Herald 1983:178 [ref. 9277], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Page et al. 2013:124 [ref. 32708], Burton & Lea 2019:67 [ref. 37205], Love et al. 2021:124 [ref. 39279]). Current status: Valid as Oligocottus rubellio (Greeley 1899). Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: northern California (U.S.A.) south to Pacific coast of northern Baja California (Mexico). Habitat: marine.

saburrae, Rusulus Starks [E. C.] & Mann [W. M.] 1911:14, Fig. 2 [University of California Publications in Zoology v. 8 (no. 2); ref. 4199] Ballast Point, off mouth of San Diego Bay, California, U.S.A. Holotype (unique): CAS-SU 23615. Type catalog: Böhlke 1953:132 [ref. 12291]. •Synonym of Clinocottus analis (Girard 1858). Current status: Synonym of Clinocottus analis (Girard 1858). Psychrolutidae: Oligocottinae. Habitat: marine.

sadko, Cottunculus Essipov [V. K.] 1937:90, Fig. 2 [Problemy Arktiki: sbornik statej, Arkticeskij i Antarkticeskij Naucno-Issledovatel'skij Institut, Glavnogo Uprevlemija Severnogo Morskogo Puti, Ministerstva Morskogo Flota SSSR. 1937 (no. 4); ref. 20998] Northern Kara Sea, between 78°07.2'-82°09'N, 65°50'-83°08'E, depth 300-690 meters. Syntypes: ZIN 30588-90 (1, 1, 1), 32170 (1, lost). •Valid as Cottunculus sadko Essipov 1937 -- (Neelov 1973:603 [ref. 7219], Nelson 1977:508 [ref. 7034], Nelson & Nakamura 1980:990 [ref. 26434], Fedorov & Nelson in Whitehead et al. 1986:1263 [ref. 13677], Nelson 1989:407 [ref. 12546], Yabe in Okamura et al. 1995:169 [ref. 22531], Mecklenburg et al. 2002:518 [ref. 25968], Parin et al. 2002:S92 [ref. 28300], Love et al. 2005:94 [ref. 37547], Byrkjedal & Orlov 2007:67 [ref. 29312], Mecklenburg et al. 2011:125 [ref. 31212], Dolgov 2013:918 [ref. 33112]). •Synonym of Cottunculus microps Collett 1875 -- (Byrkjedal et al. 2014:14 [ref. 33415], Mecklenburg et al. 2014:137 [ref. 33463], Mecklenburg et al. 2016:99 [ref. 34440]). Current status: Synonym of Cottunculus microps Collett 1875. Psychrolutidae: Psychrolutinae. Habitat: marine.

sagamia, Daruma Jordan [D. S.] & Starks [E. C.] 1904:241, Fig. 4 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Sagami Bay, Japan. Holotype: USNM 50914. Paratypes: CAS-SU 7702 (1). Additional material: CAS-SU 7850 (1), 8557 (1). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Daruma sagamia Jordan & Starks 1904 -- (Watanabe 1960:29 [ref. 11959], Yabe in Masuda et al. 1984:325 [ref. 6441], Nakabo 2000:632 [ref. 25086], Nakabo 2002:632 [ref. 26001]). Current status: Valid as Daruma sagamia Jordan & Starks 1904. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

satoi, Pseudoblennius Watanabe [M.] 1958:391, Figs. 185-186, Pl. 39 [Studies of Japanese cottid fishes; ref. 21357] Toshima-Fukushia, Hokkaido, Japan, depth 20 fathoms. Syntypes: (4). Also described as new in Watanabe 1960:157, Pl. 35 (fig. 2) [ref. 11959]. •Valid as Pseudoblennius satoi Watanabe 1958 -- (Watanabe 1960:157 [ref. 11959]). •Synonym of Pseudoblennius cottoides (Richardson 1848) -- (Nakabo 2002:1526 [ref. 26001]). Current status: Synonym of Pseudoblennius cottoides (Richardson 1848). Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

scaber, Artediellus Knipowitsch [N. M.] 1907:18, Pl. 1 (figs. 7-12) [Zapiski Imperatorskoj Akademii Nauk po Fiziko-Matematiceskomu Otdelniji = Mémoires de l'Académie Impériale des Sciences de St. Petersbourg, Classe Physico-Mathématique (8 Sér.) v. 18 (no. 5); ref. 15152] Northeastern, eastern and southeastern part of European Arctic Ocean, Kara Sea, to Bering Strait. Lectotype: ZIN 14206. Paralectotypes: (79) ZIN 14205-06 (13, 4); ZMUC 22-23 (2). Type catalog: Nielsen 1974:81 [ref. 9588]. Lectotype selected by Neelov 1979:154 [ref. 3152]. •Valid as Artediellus scaber Knipowitsch 1907 -- (Neelov 1973:595 [ref. 7219], Neelov 1979:154 [ref. 3152], Fedorov in Whitehead et al. 1986:1245 [ref. 13677], Nelson 1986:43 [ref. 5808], McAllister 1990:194 [ref. 14674], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:491 [ref. 25968], Parin et al. 2002:S71 [ref. 28300], Nelson et al. 2004:118 [ref. 27807], Love et al. 2005:84 [ref. 37547], Mecklenburg et al. 2007:174 [ref. 29448], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:121 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:239 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Afonso et al. 2016:41, 100 [ref. 36199], Kells et al. 2016:162 [ref. 35888], Mecklenburg et al. 2016:51 [ref. 34440], Mecklenburg et al. 2018:188 [ref. 36370], Love et al. 2021:118 [ref. 39279], Chernova et al. 2022:5 [ref. 39177]). Current status: Valid as Artediellus scaber Knipowitsch 1907. Psychrolutidae: Psychrolutinae. Distribution: Arctic Ocean, North Pacific, North Atlantic: Chukchi and Beaufort seas south to northern Bering Sea, east to Somerset Island, Canada, and west to Barents and Kara seas. Habitat: brackish, marine.

scepticus, Triglops Gilbert [C. H.] 1896:428, Pl. 28 (lower) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Unalaska to Kodiak islands, Alaska, U.S.A., Albatross station 3339, depth 253 meters. Lectotype: USNM 64325. Paralectotypes: BMNH 1900.7.11.6 (1); MCZ 28235 (1), 28353 (1); CAS-SU 2426 (1?), 2427 (1), 2428 (5, 1 + disintegrated specimens), 69686 (5); USNM 125621 (1). Type catalog: Böhlke 1953:133 [ref. 12291]. Lectotype established (as figured specimen from Albatross sta. 3339) in caption to Pl. 288, p. 3291 in Jordan & Evermann 1900 [ref. 2446], which is the same figure as in the original description; Pietsch 1994:358 [ref. 21189] identified that specimen as SU 2426 and designated it as lectotype. •Valid as Triglops scepticus Gilbert 1896 -- (Watanabe 1960:15 [ref. 11959], Yabe in Amaoka et al. 1983:287, 338 [ref. 8379], Yabe in Masuda et al. 1984:325 [ref. 6441], Lindberg & Krasyukova 1987:176 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Pietsch 1994:358 [ref. 21189], Shinohara et al. 1996:176 [ref. 26605], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:628 [ref. 25086], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:425 [ref. 25968], Nakabo 2002:628 [ref. 26001], Choi et al. 2003:263, 579 [ref. 26218], Parin et al. 2002:S89 [ref. 28300], Fedorov et al. 2003:82 [ref. 27579], Nelson et al. 2004:122 [ref. 27807], Love et al. 2005:93 [ref. 37547], Shinohara et al. 2011:44 [ref. 31715], Page et al. 2013:125 [ref. 32708], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:278 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Kells et al. 2016:194 [ref. 35888], Kim et al. 2020:105 [ref. 39118], Love et al. 2021:127 [ref. 39279], Dyldin & Orlov 2022:53 [ref. 38972]). Current status: Valid as Triglops scepticus Gilbert 1896. Psychrolutidae: Triglopinae. Distribution: North Pacific: Bering Sea to southeastern Alaska and to Okhotsk Sea and Japan Sea to Korea. Habitat: marine.

schmidti, Artediellus Soldatov [V. K.] 1915:160, Fig. 2 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 15068] Tatar Strait and Okhotsk Sea, including Aniva Bay. Syntypes: (many) ZIN 12186-87 (6+, 6+), 12190 (2), 12913 (5). •Valid as Artediellus schmidti Soldatov 1915 -- (Lindberg & Krasyukova 1987:235 [ref. 15964], Parin et al. 2002:S71 [ref. 28300], Parin et al. 2014:239 [ref. 33547], Dyldin & Orlov 2022:40 [ref. 38972]). •Synonym of Artediellus dydymovi Soldatov 1915, but a valid subspecies -- (Neelov 1979:155 [ref. 3152], Yabe in Masuda et al. 1984:328 [ref. 6441], Nakabo 2000:642 [ref. 25086], Nakabo 2002:642 [ref. 26001]). •Synonym of Artediellus dydymovi Soldatov 1915 -- (Dyldin & Orlov 2017:61 [ref. 35138] with question). Current status: Synonym of Artediellus dydymovi Soldatov 1915. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific [if valid]. Habitat: marine.

schmidti, Cottiusculus Jordan [D. S.] & Starks [E. C.] 1904:299, Fig. 30 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Kinkazwan [Kinkazan] Island, Matsushima Bay, Japan, depth 79 fathoms. Holotype: USNM 50923. Paratypes: CAS-SU 7714 (4), 7588 (5). Type catalog: Böhlke 1953:128 [ref. 12291]. •Valid as Cottiusculus schmidti Jordan & Starks 1904 -- (Watanabe 1960:168 [ref. 11959], Neelov 1979:162 [ref. 3152], Sheiko 1983:18 [ref. 37041], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:239 [ref. 15964], Kim & Youn 1992:72 [ref. 22287], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:643 [ref. 25086], Choi et al. 2003:262, 574 [ref. 26218], Parin et al. 2002:S72 [ref. 28300], Kai & Nakabo 2009:220 [ref. 30299], Prokofiev 2020:13 [ref. 37765], Dyldin & Orlov 2022:41 [ref. 38972]). Current status: Valid as Cottiusculus schmidti Jordan & Starks 1904. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Pacific coast of Japan to Peter the Great Bay (Russia). Habitat: marine.

schraderi, Sclerocottus Fischer [J. G.] 1885:58, Pl. 1 (figs. 3-4) [Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten v. 2; ref. 1333] Reported as South Georgia Island [locality apparently erroneous, see Berg 1940]. Holotype (unique): ZMH H67 [ex 3888]. Type catalog: Ladiges et al. 1958:165 [ref. 19181], Wilkens & Dohse 1993:407 [ref. 21161]. •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Neelov 1973:595 [ref. 7219] as Gymnacanthus). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Psychrolutidae: Icelinae. Habitat: marine.

scorpioides, Cottus Fabricius [O.] 1780:157 [Fauna groenlandica, systematice sistens animalia Groenlandiae occidentalis, etc.; ref. 17464] Western Greenland. No types known. •Possibly valid as Megalocottus scorpioides (Fabricius 1780) -- (Balakirev et al. 2020:11 [ref. 37873]). •Valid as Myoxocephalus scorpioides (Fabricius 1780) -- (Neelov 1973:599 [ref. 7219], Neelov 1979:122 [ref. 3152], Fedorov in Whitehead et al. 1986:1254 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:505 [ref. 25518], McAllister 1990:200 [ref. 14674], Yabe in Okamura et al. 1995:165 [ref. 22531], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:479 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:264 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:49, 228 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:82 [ref. 34440], Mecklenburg et al. 2018:219 [ref. 36370], Love et al. 2021:123 [ref. 39279]). Current status: Valid as Myoxocephalus scorpioides (Fabricius 1780). Psychrolutidae: Myoxocephalinae. Distribution: Arctic Ocean, northwestern Atlantic and North Pacific: eastern Siberian and Chukchi seas to northern Bering Sea, Beaufort Sea to Greenland and Gulf of St. Lawrence. Habitat: brackish, marine.

scorpius, Cottus Linnaeus [C.] 1758:265 [Systema Naturae, Ed. X v. 1; ref. 2787] European Ocean. Syntypes: BMNH 1853.11.12.17 [Gronovius coll.] (1, skin), LS 17 (left half-skin plus parts), NRM 2357 (2). Type catalog: Goode & Bean 1885:196 [ref. 1841], Wheeler 1958:240 [ref. 13434], Fernholm & Wheeler 1983:234-235 [ref. 20707], Wheeler 1985:46 [ref. 20705]. •Valid as Myoxocephalus scorpius (Linnaeus 1758) -- (Neelov 1973:598 [ref. 7219], Neelov 1979:122 [ref. 3152], Fedorov in Whitehead et al. 1986:1254 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:507 [ref. 25518], Matarese et al. 1989:402 [ref. 26885], McAllister 1990:200 [ref. 14674], Yabe in Okamura et al. 1995:165 [ref. 22531], Klein-MacPhee in Collette & Klein-MacPhee 2002:353 [ref. 26158], Mecklenburg et al. 2002:478 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Fricke 2007:29 [ref. 30577], Mecklenburg et al. 2007:176 [ref. 29448], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212] with comments. Page et al. 2013:124 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:264 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:49, 229 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:84 [ref. 34440], Mecklenburg et al. 2018:221 [ref. 36370], Carneiro et al. 2019:237 [ref. 37250], Love et al. 2021:123 [ref. 39279], Simian et al. 2022:375 [ref. 40776]). Current status: Valid as Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Distribution: Circumpolar, northern Hemisphere. Habitat: brackish, marine.

scutiger, Icelus Bean [T. H.] 1890:41 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Off Trinity Island, Bering Sea, 56°N, 154°20'W, Albatross station 2853, depth 159 fathoms. Holotype: USNM 45368. Paratypes: MCZ 27860 (1), USNM 227362 (2). •Valid as Rastrinus scutiger (Bean 1890) -- (Nelson 1984:17 [ref. 5391], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:457 [ref. 25968], Parin et al. 2002:S85 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Stevenson 2006:792 [ref. 28923], Maslenikov et al. 2013:11 [ref. 37947], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:271 [ref. 33547], Kells et al. 2016:190 [ref. 35888], Love et al. 2021:125 [ref. 39279]). Current status: Valid as Rastrinus scutiger (Bean 1890). Psychrolutidae: Icelinae. Distribution: North Pacific: Commander Islands (Russia) and Aleutian Islands (U.S.A.) to western Gulf of Alaska (U.S.A.). Habitat: marine.

sekii, Icelus Tsuruoka [O.], Munehara [H.] & Yabe [M.] 2006:47, Figs. 1-2 [Ichthyological Research v. 53 (no. 1); ref. 28501] 44°02.1'N, 145°13.2'E, off Rausu, Shiretoko Peninsula, Hokkaido Island, Japan, depth 20 meters. Holotype: HUMZ 187896. Paratypes: HUMZ 179902 (1), 187897 (2), 187898 (1), 188505 (1). •Valid as Icelus sekii Tsuruoka, Munehara & Yabe 2006 -- (Parin et al. 2014:256 [ref. 33547], Dyldin & Orlov 2022:47 [ref. 38972]). Current status: Valid as Icelus sekii Tsuruoka, Munehara & Yabe 2006. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Hokkaido (Japan), Kuril Islands and Sakhalin (Russia). Habitat: marine.

sellaris, Acanthocottus Gilbert [C. H.] 1896:419 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bristol Bay, Alaska, U.S.A., depth 5-17 fathoms. Syntypes: BMNH 1900.7.11.4-5 (2); MCZ 28321 (4), 28351 (2), 28366 (?); MNHN 1896-0461 [ex USNM] (6); CAS-SU 2846-47 (24, 35); USNM 48575 (3, now 4); ZMUO J1401-02 (1, 3). Type catalog: Böhlke 1953:126 [ref. 12291], Blanc & Hureau 1968:48 [ref. 20738], Pethon 1969:7 [ref. 19268]. •Valid as Microcottus sellaris (Gilbert 1896) -- (Neelov 1979:134 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:218 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:644 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:483 [ref. 25968], Nakabo 2002:644 [ref. 26001], Parin et al. 2002:S80 [ref. 28300], Yabe & Pietsch 2003:276 [ref. 27215], Fedorov et al. 2003:74 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:259 [ref. 33547], Mecklenburg et al. 2016:74 [ref. 34440], Kells et al. 2016:182 [ref. 35888], Mecklenburg et al. 2018:211 [ref. 36370], Dyldin et al. 2021:66 [ref. 38804], Love et al. 2021:123 [ref. 39279], Dyldin & Orlov 2022:48 [ref. 38972]). Current status: Valid as Microcottus sellaris (Gilbert 1896). Psychrolutidae: Myoxocephalinae. Distribution: North Pacific and adjacent Arctic: northern Sea of Japan and sea of Okhotsk north to western Bering Sea and Chukchi Sea. Habitat: brackish, marine.

setiger, Dasycottus Bean [T. H.] 1890:42 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Off Sitkalidak Island, Alaska, U.S.A., 57°N, 153°18'W, Albatross station 2855, depth 69 fathoms. Syntypes: (4) MCZ 35983 [ex USNM 45370] (1), USNM 45370 (3). •Valid as Dasycottus setiger Bean 1890 -- (Watanabe 1960:175 [ref. 11959], Eschmeyer & Herald 1983:167 [ref. 9277], Sheiko 1983:18 [ref. 37041], Yabe in Amaoka et al. 1983:155, 214, 287, 339 [ref. 8379], Yabe in Masuda et al. 1984:330 [ref. 6441], Lindberg & Krasyukova 1987:268 [ref. 15964], Matarese et al. 1989:440 [ref. 26885], McAllister 1990:202 [ref. 14674], Shinohara et al. 1996:176 [ref. 26605], Jackson & Nelson 1998:1354 [ref. 23806], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:651 [ref. 25086], Sheiko & Fedorov 2000:30 [ref. 25650], Tokranov & Orlov 2001:482 [516] [ref. 38795], Mecklenburg et al. 2002:516 [ref. 25968], Nakabo 2002:651 [ref. 26001], Choi et al. 2003:276, 583 [ref. 26218], Love et al. 2005:94 [ref. 37547], Parin et al. 2002:S92 [ref. 28300], Fedorov et al. 2003:85 [ref. 27579], Shinohara et al. 2011:46 [ref. 31715], Page et al. 2013:126 [ref. 32708], Shinohara et al. 2014:254 [ref. 33330], Parin et al. 2014:298 [ref. 33547], Pietsch & Orr 2015:50 [ref. 34694], Tohkairin et al. 2015:8 [ref. 35331], Kells et al. 2016:208 [ref. 35888], Kim et al. 2020:107 [ref. 39118], Sonoyama et al. 2020:91 [ref. 37637], Dyldin & Orlov 2022:54 [ref. 38972], Love et al. 2021:120 [ref. 39279]). Current status: Valid as Dasycottus setiger Bean 1890. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Sea of Japan (Korea, Japan, Russia) and Sea of Okhotsk north to Commander Islands (Russia) and Aleutian Islands (U.S.A.), then Alaska (U.S.A.) south to Washington (U.S.A.). Habitat: marine.

shibusawai, Pseudoblennius Watanabe [M.] 1958:395, Figs. 187-188, Pl. 40 [Studies of Japanese cottid fishes; ref. 21357] No locality [Japan]. Syntypes: (4). Also described as new in Watanabe 1960:160, Pl. 36 (fig. 1) [ref. 11959]. •Valid as Pseudoblennius shibusawai Watanabe 1958 -- (Watanabe 1960:152 [ref. 11959]). •Synonym of Pseudoblennius cottoides (Richardson 1848) -- (Nakabo 2002:1526 [ref. 26001]). Current status: Synonym of Pseudoblennius cottoides (Richardson 1848). Psychrolutidae: Psychrolutinae. Habitat: marine.

sigalutes, Gilbertina Jordan [D. S.] & Starks [E. C.] 1895:811, Pl. 86 [Proceedings of the California Academy of Sciences (Series 2) v. 5; ref. 2522] Puget Sound near Port Orchard, Washington, U.S.A. Holotype (unique): CAS-SU 3129. Spelled sigolutes in Jordan & Evermann 1898:2028. •Valid as Gilbertidia sigalutes (Jordan & Starks 1895) -- (Eschmeyer & Herald 1983:169 [ref. 9277], Matarese et al. 1989:434 [ref. 26885], Sheiko & Fedorov 2000:30 [ref. 25650], Parin et al. 2002:S93 [ref. 28300]). •Valid as Psychrolutes sigalutes (Jordan & Starks 1895) -- (McAllister 1990:204 [ref. 14674], Fricke 1990:407 [ref. 15492], Nelson 1995:69 [ref. 22668], Mecklenburg et al. 2002:522 [ref. 25968], Nelson et al. 2004:123 [ref. 27807], Love et al. 2005:95 [ref. 37547], Page et al. 2013:126 [ref. 32708], Nazarkin et al. 2014:177 [ref. 33578]. Parin et al. 2014:301 [ref. 33547], Pietsch & Orr 2015:51 [ref. 34694], Kells et al. 2016:210 [ref. 35888], Love et al. 2021:125 [ref. 39279]). Current status: Valid as Psychrolutes sigalutes (Jordan & Starks 1895). Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Commander Islands (Russia) and Aleutian Islands (U.S.A.) south to Puget Sound (Washington, U.S.A.). Habitat: marine.

sinensis, Cottus (Aspicottus) Sauvage [H.-E.] 1873:55 [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 9 (pt 2, Bulletin); ref. 18661] China. Holotype (unique): MNHN 0000-5265. Type catalog: Blanc & Hureau 1968:52 [ref. 20738]. •Valid as Myoxocephalus sinensis (Sauvage 1873) -- (Zhu 1995:189 [ref. 25213]). Current status: Valid as Myoxocephalus sinensis (Sauvage 1873). Psychrolutidae: Myoxocephalinae. Distribution: China. Habitat: marine.

sinensis, Lepidobero Qin [K.-J.] & Jin [X.-B.] 1992:1 [4] [Journal of Dalian Fisheries College v. 6 (nos 3-4); ref. 22919] Hei-Shi-Jiao, Dalian. Liaoning Province, China. Holotype: Dalian Fish. Coll. 8600051. Paratypes: Dalian Fish Coll. 8603078 (1, allotype); Shanghai Fish. Univ. 81(X)0010-11 (2), 8603976-77 (2). On p. 4 in English. •Valid as Lepidobero sinensis Qin & Jin 1992. Current status: Valid as Lepidobero sinensis Qin & Jin 1992. Psychrolutidae: Psychrolutinae. Distribution: China. Habitat: freshwater.

sio, Psychrolutes Nelson [J. S.] 1980:444, Figs. 1 A-C [Canadian Journal of Zoology v. 58 (no. 3); ref. 8682] Off Arica, Chile, midpoint 18°36.4'S, 70°32.9'W, depth 768-967 meters. Holotype: SIO 72-184. Paratypes: SIO 72-183 (1). •Valid as Psychrolutes sio Nelson 1980 -- (Pequeño 1989:57 [ref. 14125], Fricke 1990:407 [ref. 15492], Nelson 1995:69 [ref. 22668], Robertson et al. 2017:82 [ref. 35676] as cf. sio). Current status: Valid as Psychrolutes sio Nelson 1980. Psychrolutidae: Psychrolutinae. Distribution: Eastern Pacific: El Salvador to Chile. Habitat: marine.

smithi, Sigmistes Schultz [L. P.] 1938:190 [Proceedings of the United States National Museum v. 85 (no. 3038); ref. 3951] Igitkin Island, Aleutian Islands, Alaska, U.S.A. Holotype: USNM 105282. Paratypes: USNM 105283 (2). •Valid as Sigmistes smithi Schultz 1938 -- (Yabe et al. 2001:68 [ref. 25246], Mecklenburg et al. 2002:497 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:92 [ref. 37547], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:272 [ref. 33547], Love et al. 2021:126 [ref. 39279]). Current status: Valid as Sigmistes smithi Schultz 1938. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Kuril Islands; Aleutian Islands (U.S.A.). Habitat: marine.

snyderi, Dialarchus Greeley [A. W.] 1899:14, Fig. 4 [Bulletin of the U. S. Fish Commission v. 19 [1899]; ref. 1889] Pacific Grove, California, U.S.A. Holotype: CAS-SU 5846. Paratypes: CAS-SU 5847 (5). See Oligocottus snyderi Greeley 1898. •Preoccupied by and same as Oligocottus snyderi Greeley 1898. 1°Homonym. Current status: Synonym of Oligocottus snyderi Greeley 1898. Psychrolutidae: Oligocottinae. Habitat: marine.

snyderi, Oligocottus Greeley [A. W.] in Jordan & Evermann 1898:2871 [Bulletin of the United States National Museum No. 47; ref. 2445] Pacific Grove, California, U.S.A. Lectotype: CAS-SU 5846. Paralectotypes: CAS-SU 5847 ( 5). Type catalog: Böhlke 1953:131 [ref. 12291]. Based on Oligogottus maculosus of Jordan & Evermann 1896:2013 [ref. 2444]. Lectotype apparently established by Greeley 1899:16 [ref. 1889]. •Valid as Oligocottus snyderi Greeley 1898 -- (Bolin 1944:67 [ref. 6379], Eschmeyer & Herald 1983:178 [ref. 9277], Matarese et al. 1989:432 [ref. 26885], McAllister 1990:201 [ref. 14674], Mecklenburg et al. 2002:501 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:43 [ref. 34694], Kells et al. 2016:186 [ref. 35888], Burton & Lea 2019:68 [ref. 37205], Love et al. 2021:124 [ref. 39279]). Current status: Valid as Oligocottus snyderi Greeley 1898. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: western Gulf of Alaska (U.S.A.) south to Pacific coast of central Baja California (Mexico). Habitat: marine.

spatula, Icelus Gilbert [C. H.] & Burke [C. V.] 1912:41, Figs. 3, 3a [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off Avatcha Bay, Kamchatka, Russia, 52°47'20"N, 158°44'30"E, Albatross station 4794, depth 58-69 fathoms. Holotype: USNM 74367. Paratypes: CAS-SU 40931 (5), USNM 70904-05 (4, 4). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelus spatula Gilbert & Burke 1912 -- (Neelov 1973:597 [ref. 7219], Nelson 1984:41 [ref. 5391], Fedorov in Whitehead et al. 1986:1251 [ref. 13677], Robins & Ray 1986:282 [ref. 23100], Scott & Scott 1988:499 [ref. 25518], McAllister 1990:199 [ref. 14674], Yabe in Okamura et al. 1995:164 [ref. 22531], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:455 [ref. 25968], Parin et al. 2002:S78 [ref. 28300], Fedorov et al. 2003:72 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Mecklenburg et al. 2007:176 [ref. 29448], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:122 [ref. 31212] with comments, Page et al. 2013:123 [ref. 32708], Dolgov 2013:918 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:256 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:45, 174 [ref. 36199], Kells et al. 2016:178 [ref. 35888], Mecklenburg et al. 2016:69 [ref. 34440], Mecklenburg et al. 2018:207 [ref. 36370], Dyldin & Orlov 2022:47 [ref. 38972], Love et al. 2021:122 [ref. 39279], Zorina & Chernova. 2022:89 [ref. 40474]). Current status: Valid as Icelus spatula Gilbert & Burke 1912. Psychrolutidae: Icelinae. Distribution: Circumpolar, Arctic Ocean, northwestern Atlantic and North Pacific. Habitat: marine.

spiniger, Icelus Gilbert [C. H.] 1896:412, Pl. 24 (top) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Alaska, U.S.A., 54°48'30"N, 165°49'00"W, Albatross station 3225, depth 85 fathoms. Lectotype: ?USNM 164258. Paralectotypes: BMNH 1900.7.11.7 (1?); MCZ 28270 (3); MNHN 1896-0484 to 0486 [ex USNM] (3); CAS-SU 2231 (40), 2266 (2), 2269 (16); ZMUO J1463 (3, poor condition). Type catalog: Böhlke 1953:130 [ref. 12291], Blanc & Hureau 1968:54 [ref. 20738], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen from Albatross sta. 3278) in caption to Pl. 286, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable; a lectotype (USNM 164248, sta. 3225) was also designated by Nelson 1984:24 [ref. 5391]. •Valid as Icelus spiniger Gilbert 1896 -- (Watanabe 1960:42 [ref. 11959], Eschmeyer & Herald 1983:174 [ref. 9277], Nelson 1984:24 [ref. 5391], Peden & Hughes 1986:7 [ref. 9714], McAllister 1990:199 [ref. 14674], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:451 [ref. 25968], Parin et al. 2002:S78 [ref. 28300], Fedorov et al. 2003:73 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Mecklenburg et al. 2011:122 [ref. 31212] as cf. spiniger with comments, Page et al. 2013:123 [ref. 32708], Parin et al. 2014:256 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Mecklenburg et al. 2016:71 [ref. 34440] see remarks, Kells et al. 2016:180 [ref. 35888], Love et al. 2021:122 [ref. 39279]). Current status: Valid as Icelus spiniger Gilbert 1896. Psychrolutidae: Icelinae. Distribution: North Pacific: Sea of Okhotsk north to northern Bering Sea, then south to southern British Columbia (Canada). Habitat: marine.

spinosus, Cottunculus Gilchrist [J. D. F.] 1906:149, Pl. 38 [Marine Investigations in South Africa v. 4; ref. 1647] Cape Point bearing N 70°E distance 40 miles, South Africa, depth about 800 fathoms. Holotype (unique): whereabouts unknown. •Valid as Cottunculoides spinosus (Gilchrist 1906) -- (Nelson 1977:508 [ref. 7034]). •Valid as Cottunculus spinosus Gilchrist 1906 -- (Nelson 1986:491 [ref. 5698], Byrkjedal & Orlov 2007:67 [ref. 29312], Nelson 2016:2291 [ref. 34615]). Current status: Valid as Cottunculus spinosus Gilchrist 1906. Psychrolutidae: Psychrolutinae. Distribution: South Africa. Habitat: marine.

spinosus, Porocottus Watanabe [M.] 1958:311, Figs. 132-133, Pl. 24 [Studies of Japanese cottid fishes; ref. 21357] Kurshiro, Hokaidô, Japan, depth 35 fathoms. Syntypes: (3). Also described as new in Watanabe 1960:98, Pl. 3 (fig. 2) [ref. 11959]. •Valid as Porocottus spinosus Watanabe 1958 -- (Watanabe 1960:98 [ref. 11959]). •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Psychrolutidae: Myoxocephalinae. Habitat: marine.

spitzbergensis, Icelus bicornis Schmidt [P. Yu.] 1927:7 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 19815] Spitsbergen. Syntypes: ZIN 11867 (3), 11900-02 (1, 4, 6+), 12750-52 (3, 2, 5), 12754-55 (13, 3), 12807-08 (1, 1), 13993-97 (6+, 6+, 1, 1, 1). •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neelov 1973:596 [ref. 7219], Nelson 1984:36 [ref. 5391], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:253 [ref. 33547]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Psychrolutidae: Icelinae. Habitat: marine.

stejnegeri, Stelgistrum Jordan [D. S.] & Gilbert [C. H.] in Jordan & Evermann 1898:1921 [Bulletin of the United States National Museum No. 47; ref. 2444] Off Robben Island, Okhotsk Sea [not Bering Sea], Albatross station 3645, depth 10 fathoms. Holotype (unique): USNM 48236. Also appeared in Jordan & Gilbert 1899:456, Pl. 54 [ref. 2478] as steinegeri. •Valid as Stelgistrum stejnegeri Jordan & Gilbert 1898 -- (Watanabe 1960:32 [ref. 11959], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:195 [ref. 15964] as steinegeri, Sokolovskaya et al. 1998:10 [ref. 24670] as steinegeri, Nakabo 2000:636 [ref. 25086], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:447 [ref. 25968], Nakabo 2002:636 [ref. 26001], Parin et al. 2002:S86 [ref. 28300], Fedorov et al. 2003:80 [ref. 27579], Tsuroka et al. 2009:97 [ref. 32107], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:273 [ref. 33547], Dyldin & Orlov 2017:67 [ref. 35138], Dyldin et al. 2021:68 [ref. 38804], Dyldin & Orlov 2022:51 [ref. 38972]). Current status: Valid as Stelgistrum stejnegeri Jordan & Gilbert 1898. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: brackish, marine.

stelleri, Cottus Bloch [M. E.] & Schneider [J. G.] 1801:63 [M. E. Blochii, Systema Ichthyologiae; ref. 471] Mentioned in passing, not available. •In the synonymy of Ceratocottus [now Enophrys] diceraus (Pallas 1787) -- (Watanabe 1960:110 [ref. 11959]). •In the synonymy of Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414]). Nomen Nudum. Current status: Synonym of Enophrys diceraus (Pallas 1787). Psychrolutidae: Myoxocephalinae.

stelleri, Myoxocephalus Tilesius [W. G. von] 1811:273, Pl. 13 (fig. 1) [Mémoires de l'Académie Impériale des Sciences de Sankt Pétersbourg (5e Série) v. 3; ref. 4408] Kamchatka, Russia. No types known. •Valid as Myoxocephalus stelleri Tilesius 1811 -- (Berg 1949:1137 [ref. 20662], Neelov 1979:121 [ref. 3152], Yabe in Masuda et al. 1984:326 [ref. 6441], Lindberg & Krasyukova 1987:207 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Kim 1997:407 [ref. 24032], Sokolovsky & Sokolovskaya 1997:58 [ref. 24691], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:153 [ref. 23595], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:645 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Pietsch et al. 2001:145 [ref. 25688], Zemnukhov et al. 2001:420 [ref. 25412], Kim & Park 2002:327 [ref. 25913], Mecklenburg et al. 2002:480 [ref. 25968], Nakabo 2002:645 [ref. 26001], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S82 [ref. 28300], Hagiwara 2003:37 [ref. 27188], Fedorov et al. 2003:77 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:90 [ref. 37547], Bogutskaya & Naseka 2004:191 [ref. 28183], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:265 [ref. 33547], Kells et al. 2016:186 [ref. 35888], Dyldin & Orlov 2017:66 [ref. 35138], Kim et al. 2020:103 [ref. 39118], Dyldin et al. 2021:67 [ref. 38804], Love et al. 2021:124 [ref. 39279], Dyldin & Orlov 2022:49 [ref. 38972]). Current status: Valid as Myoxocephalus stelleri Tilesius 1811. Psychrolutidae: Myoxocephalinae. Distribution: North Pacific: Sea of Japan (Korea, Japan, Russia) and Sea of Okhotsk north to Commander Islands (Russia) and Aleutian Islands (U.S.A.), then south to southeastern Alaska (U.S.A.). Habitat: brackish, marine.

stenosomus, Icelus uncinalis Andriashev [A. P.] 1937:266, Pl. 5 (fig. 3) [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] 48°08'30"N, 140°08'30'E, Tatar Strait. Holotype: ZIN 17646. Paratypes: ZIN 17641-42 (6, 2), 17645 (4), 17646A (5, with holotype), 17647 (3), 21906-08 (1, 1, 1). "Typus" given on caption to Pl. 5, specimen noted in catalog in 17646 and regarded as holotype. •Synonym of Icelus uncinalis Gilbert & Burke 1912, but a valid subspecies as described -- (Lindberg & Krasyukova 1987:194 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670]). •Valid as Icelus stenosomus Andriashev 1937 -- (Nelson 1984:53 [ref. 5391], Murai et al. 1994 [ref. 21908], Nakabo 2000:635 [ref. 25086], Nakabo 2002:635 [ref. 26001], Parin et al. 2002:S78 [ref. 28300], Ko et al. 2009:61 [ref. 37344], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:257 [ref. 33547], Kim et al. 2020:103 [ref. 39118], Sonoyama et al. 2020:90 [ref. 37637], Dyldin & Orlov 2022:47 [ref. 38972]). Current status: Valid as Icelus stenosomus Andriashev 1937. Psychrolutidae: Icelinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia). Habitat: marine.

stimpsonii, Triglopsis Hoy [P. R.] 1872:99 [Transactions of the Wisconsin Academy of Sciences Arts and Letters 1870-72; ref. 18119] Lake Michigan, off Racine, Milwaukee, U.S.A. (stomach content). Holotype (unique): whereabouts unknown. Apparently not available; appeared above and also as Gill (MS) in Jordan 1878:50 [ref. 10444] as stimpsoni, both without distingusihing features. Subsequent availability not researched. •In the synonymy of Myoxocephalus quadricornis (Linnaeus 1758) -- (Scott & Crossman 1998:846 [ref. 24570]). Nomen Nudum. Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae.

strabo, Icelinus Starks [E. C.] 1896:551 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 4195] Port Ludlow, Washington, U.S.A. Syntypes: BMNH 1897.1.12.19-20 (2), CAS-SU 5045 (about 25, now 19). Type catalog: Böhlke 1953:130 [ref. 12291]. •Synonym of Icelinus borealis Gilbert 1896 -- (Bolin 1936:159 [ref. 506], Rosenblatt & Smith 2004:560 [ref. 27912]). Current status: Synonym of Icelinus borealis Gilbert 1896. Psychrolutidae: Oligocottinae. Habitat: marine.

subborealis, Triglops pingelii var. Le Danois [E.] 1914:11 [Annales de l'Institut Océanographique Monaco v. 7 (no. 2); ref. 21190] Fair Isle, Shetland Islands, North Sea, 59°26'N, 1°30'W, depth 80 meters. Holotype (unique): whereabouts unknown, not at MNHN. •Synonym of Triglops murrayi Günther 1888 -- (Neelov 1973:602 [ref. 7219], Pietsch 1994:369 [ref. 21189]). Current status: Synonym of Triglops murrayi Günther 1888. Psychrolutidae: Triglopinae. Habitat: marine.

subspinosus, Cottunculus Jensen [A. S.] 1902:214 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Off Iceland, depth 750-957 fathoms. Syntypes: ZMUC uncat. Type catalog: Nielsen 1974:81 [ref. 9588]. •Valid as Psychrolutes subspinosus (Jensen 1902) -- (Nelson 1977:508 [ref. 7034], Fedorov & Nelson in Whitehead et al. 1986:1264 [ref. 13677], Nelson 1995:67 [ref. 22668], Møller et al. 2010:54 [ref. 31169], Mecklenburg et al. 2011:125 [ref. 31212]). •Valid as Cottunculus subspinosus Jensen 1902 -- (Neelov 1973:604 [ref. 7219], Parin et al. 2002:S92 [ref. 28300], Parin et al. 2014:299 [ref. 33547], Byrkjedal et al. 2018:14 [ref. 35896], Mecklenburg et al. 2018:239 [ref. 36370]). Current status: Valid as Cottunculus subspinosus Jensen 1902. Psychrolutidae: Psychrolutinae. Distribution: North Atlantic and Arctic Ocean: off eastern Greenland and off northeastern Iceland to off western Norway. Habitat: marine.

taeniopterus, Cottus Kner [R.] 1868:28 [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 58 (nos 1-2); ref. 6074] Decastris Bay near Amur River mouth, northeastern Asia. Holotype (unique): NMW 11612. Appeared in more detail in Kner 1868:310 [p. 18 of separate], Pl. 4 (fig. 10) [ref. 2646]. •Synonym of Myoxocephalus platycephalus (Pallas 1814), but a valid subspecies -- (Berg 1949:1128 [ref. 20662], Zhu 1995:189 [ref. 25213]). •Synonym of Megalocottus platycephalus (Pallas 1814), but a valid subspecies -- (Neelov 1979:130 [ref. 3152], Lindberg & Krasyukova 1987:212 [ref. 15964], Nikiforov et al. 1997:330 [ref. 24694], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:153 [ref. 23595], Sokolovskaya et al. 1998:9 [ref. 24670], Bogutskaya & Naseka 2004:190 [ref. 28183]). •Synonym of Megalocottus platycephalus (Pallas 1814) -- (Mecklenburg et al. 2002:475 [ref. 25968], Parin et al. 2002:S79 [ref. 28300], Mecklenburg et al. 2016:73 [ref. 34440]). •Valid as Megalocottus taeniopterus (Kner 1868) -- (Bogutskaya et al. 2008:352 [ref. 30085], Parin et al. 2014:258 [ref. 33547], Dyldin & Orlov 2017:64 [ref. 35138], Dyldin et al. 2021:65 [ref. 38804], Dyldin & Orlov 2022:47 [ref. 38972]). Current status: Valid as Megalocottus taeniopterus (Kner 1868). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific. Habitat: marine.

taranetzi, Bathylutichthys Balushkin [A. V.] & Voskoboinikova [O. S.] 1990:187, Figs. 1-4 [Voprosy Ikhtiologii v. 30 (no. 2); ref. 20117] South Georgia Island, 52°03'S, 33°09'E, depth 1650 meters. Holotype (unique): ZIN 49301. On p. 72 in English translation. •Valid as Bathylutichthys taranetzi Balushkin & Voskoboinikova 1990 -- (Trunov 1999:465 [ref. 24579], Voskoboinikova 2014:607 [ref. 33563]). Current status: Valid as Bathylutichthys taranetzi Balushkin & Voskoboinikova 1990. Psychrolutidae: Psychrolutinae. Distribution: South Georgia Island, Southern Ocean. Habitat: marine.

taranetzi, Radulinopsis Yabe [M.] & Maruyama [S.] 2001:56, Figs. 3-4 [Ichthyological Research v. 48 (no. 1); ref. 25245] Off Usujiri, 41°54'N, 140°05'E, Pacific coast of southern Hokkaido, Japan, depth 15-20 meters. Holotype: HUMZ 150550. Paratypes: HUMZ 87631 (1), 87770 (1), 92778-79 (2), 92781-83 (3), 92785-88 (4), 113656-61 (6), 121685-86 (2), 121719 (1), 135986 (1), 145741-48 (8), 150549 (1), 150551-52 (2), 150560-61 (2); NSMT-P 57640-41 (1, 1). •Valid as Radulinopsis taranetzi Yabe & Maruyama 2001 -- (Parin et al. 2014:271 [ref. 33547], Dyldin & Orlov 2022:51 [ref. 38972] with question). Current status: Valid as Radulinopsis taranetzi Yabe & Maruyama 2001. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific. Habitat: marine.

taurinus, Enophrys Gilbert [C. H.] 1914:135, Pl. 11 (fig. 1) [Proceedings of the United States National Museum v. 47 (no. 2049); ref. 14153] Monterey Bay, near Pacific Grove, California, U.S.A. Holotype: USNM 75064. Paratypes: CAS-SU 57 (1); plus additional material. Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Aspicottus taurinus (Gilbert 1914) -- (Neelov 1979:148 [ref. 3152]). •Valid as Enophrys taurinus Gilbert 1914 -- (Bolin 1944:93 [ref. 6379]). •Valid as Enophrys taurina Gilbert 1914 -- (Sandercock & Wilimovsky 1968:844 [ref. 26414], Eschmeyer & Herald 1983:169 [ref. 9277], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:86 [ref. 37547], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Kells et al. 2016:170 [ref. 35888], Burton & Lea 2019:66 [ref. 37205], Love et al. 2021:120 [ref. 39279]). Current status: Valid as Enophrys taurina Gilbert 1914. Psychrolutidae: Myoxocephalinae. Distribution: Northeastern Pacific: California (U.S.A.) endemic. Habitat: marine.

taylori, Asemichthys Gilbert [C. H.] 1912:215 [Contributions to Canadian Biology being studies from the Marine Biological Stations of Canada 1906-1910 (art. 12); ref. 9276] Departure Bay, Vancouver Island, British Columbia, Canada. Holotype (unique): CAS-SU 23071. Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Radulinus taylori (Gilbert 1912) -- (Eschmeyer & Herald 1983:181 [ref. 9277], Mecklenburg et al. 2002:444 [ref. 25968], Love et al. 2005:91 [ref. 37547], Page et al. 2013:121 [ref. 32708], Kells et al. 2016:190 [ref. 35888]). •See Yabe & Maruyama 2001:59 [ref. 25245]. •Valid as Asemichthys taylori Gilbert 1912 -- (McAllister 1990:194 [ref. 14674], Coffie 1998:130 [ref. 25001], Nelson et al. 2004:118 [ref. 27807], Pietsch & Orr 2015:39 [ref. 34694], Love et al. 2021:119 [ref. 39279]). Current status: Valid as Asemichthys taylori Gilbert 1912. Psychrolutidae: Radulininae. Distribution: Northeastern Pacific: southeastern Alaska (U.S.A.) to San Juan Islands, northern Washington (U.S.A.). Habitat: marine.

tentaculatus, Cottus Kner [R.] 1868:28 [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 58 (nos 1-2); ref. 6074] Singapore. Holotype (unique): NMW 11621. Appeared in more detail in Kner 1868:314 [p. 22 of separate], Pl. 5 (fig. 12) [ref. 2646]. •Valid as Porocottus tentaculatus (Kner 1868) -- (Neelov 1979:137 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:224 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:646 [ref. 25086], Nakabo 2002:646 [ref. 26001], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S84 [ref. 28300], Fedorov et al. 2003:80 [ref. 27579], Parin et al. 2014:270 [ref. 33547], Dyldin & Orlov 2017:67 [ref. 35138], Dyldin et al. 2021:68 [ref. 38804], Dyldin & Orlov 2022:51 [ref. 38972]). Current status: Valid as Porocottus tentaculatus (Kner 1868). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific. Habitat: brackish, marine.

tenuicaudalis, Malacocottus Watanabe [M.] 1958:427, Figs. 209-210, Pl. 46 [Studies of Japanese cottid fishes; ref. 21357] Abashiri, Hokkaido, Japan, depth 70 fathoms. Syntypes: (5). Also described as new in Watanabe 1960:185, Pl. 39 (fig. 2) [ref. 11959]. Spelled tenuicaudaris in classification (p. 17), on plate caption, and on table 113 caption; tenuicaudalis on p. 427, 432, and twice in list of plates (p. 458, 460). Correct spelling of original genus is Malacocottus. •Valid as Malacottus tenuicaudalis Watanabe 1958 -- (Watanabe 1960:185 [ref. 11959], Lindberg & Krasyukova 1987:273 [ref. 15964]). •Status uncertain -- (Shinohara et al. 1992:422 [ref. 19776]). •Synonym of Malacocottus zonurus Bean 1890 -- (Parin et al. 2002:S94 [ref. 28300], Parin et al. 2014:302 [ref. 33547], Stevenson 2015:24 [ref. 33680]). Current status: Synonym of Malacocottus zonurus Bean 1890. Psychrolutidae: Psychrolutinae. Habitat: marine.

tenuis, Icelinus Gilbert [C. H.] 1890:86 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., 5 Albatross stations, depth 45-150 fathoms. Syntypes: CAS-SU 78 (3), 28110 (1); USNM 43086 (1). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelinus tenuis Gilbert 1890 -- (Bolin 1944:26 [ref. 6379], Eschmeyer & Herald 1983:173 [ref. 9277], Peden 1984:79 [ref. 5193], McAllister 1990:198 [ref. 14674], Mecklenburg et al. 2002:438 [ref. 25968], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:88 [ref. 37547], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:178 [ref. 35888], Burton & Lea 2019:67 [ref. 37205], Love et al. 2021:122 [ref. 39279]). Current status: Valid as Icelinus tenuis Gilbert 1890. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: southeastern Alaska (U.S.A.) south to Pacific coast of central Baja California (Mexico). Habitat: marine.

terraenovae, Triglops ommatistius Gilbert [C. H.] 1913:467 [Proceedings of the United States National Museum v. 44 (no. 1963); ref. 14957] Newfoundland, Canada, 45°09'30"N, 49°48'30"W, Albatross station 2445, depth 39 fathoms. Holotype: USNM 46063. Paratypes: (orig. 57, from 11 stations) USNM 46059 (1), 46061-62 (1, 3), 46064-68 (?36), 46069 (1), 105089 (17). •Synonym of Triglops murrayi Günther 1888 -- (Scott & Scott 1988:510 [ref. 25518], Pietsch 1994:368 [ref. 21189], Parin et al. 2014:276 [ref. 33547]). Current status: Synonym of Triglops murrayi Günther 1888. Psychrolutidae: Triglopinae. Habitat: marine.

thompsonii, Triglopsis Girard [C. F.] 1851:19 [Proceedings of the Boston Society of Natural History v. 4 (1851-1854); ref. 1816] Lake Ontario off Oswego, New York, U.S.A. Syntypes: MCZ 35979 [ex USNM 3175] (1), USNM 3175 (8, poor condition). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Scott & Crossman 1998:843 [ref. 24570], Goto et al. 2014: [6] [ref. 33456]). •Valid as Myoxocephalus thompsonii (Girard 1851) -- (Lee et al. 1980:827 [ref. 22416], Smith 1986:359 [ref. 22186], McAllister 1990:200 [ref. 14674] as thompsoni, Page & Burr 1991:244 [ref. 18983] as thompsoni, Bailey et al. 2004:173 [ref. 27882], Nelson et al. 2004:121 [ref. 27807], Murray et al. 2004:642 [ref. 27932], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:467 [ref. 31215], Page et al. 2013:124 [ref. 32708]). Current status: Valid as Myoxocephalus thompsonii (Girard 1851). Psychrolutidae: Myoxocephalinae. Distribution: Canada and New York (U.S.A.). Habitat: freshwater.

thomsonii, Cottus Günther [A.] 1882:679 [Proceedings of the Royal Society of Edinburgh v. 11; ref. 18204] North Atlantic, 59°33'N, 7°14'W, depth 555 fathoms. Holotype (unique): BMNH 1887.12.9.10. •Valid as Cottunculus thomsonii (Günther 1882) -- (Neelov 1973:603 [ref. 7219] as thomsoni, Nelson 1977:508 [ref. 7034] as thompsoni, Nelson & Nakamura 1980:990 [ref. 26434] as thompsoni, Fedorov & Nelson in Whitehead et al. 1986:1263 [ref. 13677], Nelson 1989:407 [ref. 12546], Scott & Scott 1988:494 [ref. 25518] as thomsoni, McAllister 1990:202 [ref. 14674] as thompsoni, Nelson 1990:687 [ref. 17821], Yabe in Okamura et al. 1995:170 [ref. 22531], Quéro & Cendrero 2001:178 [ref. 25567], Moore et al. 2003:223 [ref. 27076], Byrkjedal & Orlov 2007:67 [ref. 29312], Møller et al. 2010:54 [ref. 31169], Page et al. 2013:126 [ref. 32708], Byrkjedal et al. 2014:1 [ref. 33415], Afonso et al. 2016:55, 139 [ref. 36199], Nelson 2016:2292 [ref. 34615], Carneiro et al. 2019:237 [ref. 37250], Bañón & Maño 2021:69 [ref. 38883], Fermon et al. 2022:296 [ref. 39448] with question). Current status: Valid as Cottunculus thomsonii (Günther 1882). Psychrolutidae: Psychrolutinae. Distribution: North Atlantic. Habitat: marine.

torvus, Cottunculus Goode [G. B.] & Bean [T. H.] 1883:212 [Bulletin of the Museum of Comparative Zoology v. 10 (no. 5); ref. 1838] Atlantic, 33°42'15"-41°32'50"N, 65°55'-76°00'50"W, depth 464-732 fathoms. Syntypes: MCZ 25867 (3), 25868 (1), 25968 (1). Appeared first as name only in Goode 1880:479 [ref. 1833]. •Synonym of Cottunculus thomsonii (Günther 1882) -- (Neelov 1973:603 [ref. 7219] as thomsoni, Fedorov & Nelson in Whitehead et al. 1986:1263 [ref. 13677], Nelson 1990:687 [ref. 17821]). Current status: Synonym of Cottunculus thomsonii (Günther 1882). Psychrolutidae: Psychrolutinae. Habitat: marine.

totomius, Pseudoblennius Jordan [D. S.] & Starks [E. C.] 1904:315, Fig. 36 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Omi Point, Totomi Bay, Japan, Albatross station 3729, depth 34 fathoms. Holotype (unique): USNM 50928. •Valid as Pseudoblennius totomius Jordan & Starks 1904 -- (Nakabo 2000:649 [ref. 25086], Nakabo 2002:649 [ref. 26001]). Current status: Valid as Pseudoblennius totomius Jordan & Starks 1904. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

toyamensis, Ricuzenius (Alloricuzenius) Matsubara [K.] & Iwai [T.] 1951:87, Figs. 1-3 [Miscellaneous Reports of the Research Institute for Natural Resources (Tokyo) Nos 19-21; ref. 2909] Off Uozu, Japan. Holotype: FAKU 14450 (not seen in 1997). Paratypes: FAKU 14451-57 (7, not seen in 1997). •Valid as Ricuzenius toyamensis Matsubara & Iwai 1951 -- (Watanabe 1960:22 [ref. 11959], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:186 [ref. 15964]). •Valid as Icelus toyamensis (Matsubara & Iwai 1951) -- (Nakabo 2000:633 [ref. 25086], Nakabo 2002:633, 1525 [ref. 26001], Tsuruoka et al. 2006:51 [ref. 28501], Shinohara et al. 2011:44 [ref. 31715], Song et al. 2012:304 [ref. 36295], Shinohara et al. 2014:250 [ref. 33330], Kim et al. 2020:103 [ref. 39118]). •Synonym of Icelus rastrinoides Taranetz 1936 -- (Kai et al. 2023:16 [ref. 39903]). Current status: Synonym of Icelus rastrinoides Taranetz 1936. Psychrolutidae: Icelinae. Habitat: marine.

triacis, Orthonopias Starks [E. C.] & Mann [W. M.] 1911:11, Fig. 1 [University of California Publications in Zoology v. 8 (no. 2); ref. 4199] Cortez Bank, off California, U.S.A., depth 11-16 fathoms. Holotype (unique): SIO 60-56. •Valid as Orthonopias triacis Starks & Mann 1911 -- (Bolin 1944:59 [ref. 6379], Eschmeyer & Herald 1983:179 [ref. 9277], Fedorov 1990:745 [ref. 20136], Long 1992:160 [ref. 22238], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:91 [ref. 37547], Page et al. 2013:124 [ref. 32708], Kells et al. 2016:188 [ref. 35888], Burton & Lea 2019:68 [ref. 37205], Love et al. 2021:124 [ref. 39279]). Current status: Valid as Orthonopias triacis Starks & Mann 1911. Psychrolutidae: Oligocottinae. Distribution: Northeastern Pacific: northern California (U.S.A.) south to Pacific coast of central Baja California (Mexico). Habitat: marine.

tribranchius, Atopocottus Bolin [R. L.] 1936:30, Fig. 6 [Proceedings of the United States National Museum v. 84 (no. 3000); ref. 505] Sea of Japan, 38°14'N, 138°54'E, Albatross station 4816, depth 64 fathoms. Holotype: USNM 102105. Paratypes: CAS-SU 30457 (3), USNM 102106 (7). Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Atopocottus tribranchius Bolin 1936 -- (Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:241 [ref. 15964], Nakabo 2000:639 [ref. 25086], Nakabo 2002:639 [ref. 26001], Matsunuma et al. 2017:93 [ref. 35372], Sonoyama et al. 2020:89 [ref. 37637]). Current status: Valid as Atopocottus tribranchius Bolin 1936. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: Japan and East China Sea. Habitat: marine.

tricuspis, Cottus Reinhardt [J. C. H.] 1830:16 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1829-30; ref. 19480] Greenland. Holotype: ZMUC uncat. Type catalog: Nielsen 1974:81 [ref. 9588]. •Valid as Gymnocanthus tricuspis (Reinhardt 1830) -- [sometimes as Gymnacanthus] (Neelov 1973:595 [ref. 7219], Fedorov in Whitehead et al. 1986:1249 [ref. 13677], Robins & Ray 1986:281 [ref. 23100], Scott & Scott 1988:495 [ref. 25518], Matarese et al. 1989:394 [ref. 26885], McAllister 1990:197 [ref. 14674], Yabe in Okamura et al. 1995:163 [ref. 22531], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Klein-MacPhee in Collette & Klein-MacPhee 2002:347 [ref. 26158], Mecklenburg et al. 2002:464 [ref. 25968], Parin et al. 2002:S75 [ref. 28300] as Gymnacanthus Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:87 [ref. 37547], Mecklenburg et al. 2007:174 [ref. 29448], Møller et al. 2010:52 [ref. 31169], Page et al. 2013:123 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:249 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:41, 163 [ref. 36199], Kells et al. 2016:172 [ref. 35888], Mecklenburg et al. 2016:61 [ref. 34440], Mecklenburg et al. 2018:199 [ref. 36370], Love et al. 2021:121 [ref. 39279], Chernova et al. 2022:5 [ref. 39177], Tokranov et al. 2022:888 [ref. 39694]). Current status: Valid as Gymnocanthus tricuspis (Reinhardt 1830). Psychrolutidae: Icelinae. Distribution: North Atlantic and Arctic Ocean; Circumpolar; North Pacific: northern Bering Sea. Habitat: marine.

tuberculatus, Myoxocephalus Soldatov [V. K.] & Pavlenko [M. N.] 1922:342, Fig. 1 [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23; ref. 15431] Many localities, near Shantarskiye [Shantar] Islands, western Okhotsk Sea, depth 10-87 meters. Lectotype: ZIN 20615. Paralectotypes: (26) ZIN 19430 (6), 20614 (2, lost), 20615-24 (2, 4, 1, 2, 4, 2, 2, 2, 1, 1). Lectotype selected by Neelov 1979:122 [ref. 3152]. •Valid as Myoxocephalus tuberculatus Soldatov & Pavlenko 1922 -- (Parin et al. 2014:265 [ref. 33547], Dyldin et al. 2020:114 [ref. 37569], Dyldin et al. 2021:67 [ref. 38804], Dyldin & Orlov 2022:50 [ref. 38972]). Current status: Valid as Myoxocephalus tuberculatus Soldatov & Pavlenko 1922. Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific: northern Okhotsk Sea, including Shantar Islands, Taui Bay and western Kamchatka (Russia) Habitat: brackish, marine.

tubulosus, Cottunculus Byrkjedal [I.] & Orlov [A. M.] 2007:65, Figs. 2-3 [Zootaxa No. 1580; ref. 29312] North Atlantic, 51°45'N, 29°33'E, depth 1950-1872 meters. Holotype (unique): ZMUB 16361. •Valid as Cottunculus tubulosus Byrkjedal & Orlov 2007. Current status: Valid as Cottunculus tubulosus Byrkjedal & Orlov 2007. Psychrolutidae: Psychrolutinae. Distribution: North Atlantic, Mid-Atlantic Ridge. Habitat: marine.

typicus, Triglops pingelii var. Le Danois [E.] 1914:11 [Annales de l'Institut Océanographique Monaco v. 7 (no. 2); ref. 21190] Unneeded name for the typical form pingelii described by Reinhardt. •Synonym of Triglops pingelii Reinhardt 1837 -- (Pietsch 1994:374 [ref. 21189]). Current status: Synonym of Triglops pingelii Reinhardt 1837. Psychrolutidae: Triglopinae. Habitat: marine.

uchidai, Dasycottus Mori [T.] 1955:1, Fig. 1 [Science Reports of the Hyogo University of Agriculture Series Natural Science v. 2 (no. 1); ref. 7766] Coast of Kyogasaki, Kyoto Prefecture, southern Sea of Japan, depth 300-500 meters. Holotype: Hyogo Univ. Agric. Paratypes: (1). Psychrolutidae: Psychrolutinae. Habitat: marine.

uchidai, Triglops Watanabe [M.] 1958:201, Figs. 61-62, Pl. 2 [Studies of Japanese cottid fishes; ref. 21357] Mogami-tai, Yamagata Prefecture, Japan, depth 70 fathoms. Holotype (unique): not researched. Also described as new in Watanabe 1960:11, pl. 16 (fig. 2) [ref. 11959]. •Valid as Triglops uchidai Watanabe 1958 -- (Yabe in Masuda et al. 1984:325 [ref. 6441], Lindberg & Krasyukova 1987:179 [ref. 15964]). •Synonym of Triglops scepticus Gilbert 1896 -- (Pietsch 1994:358 [ref. 21189], Nakabo 2002:1525 [ref. 26001], Parin et al. 2014:278 [ref. 33547]). Current status: Synonym of Triglops scepticus Gilbert 1896. Psychrolutidae: Triglopinae. Habitat: marine.

uncinalis, Icelus Gilbert [C. H.] & Burke [C. V.] 1912:39, Figs. 2, 2a [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Near Attu Island, Aleutian Islands, Bering Sea, 52°55'40"N, 173°26'E, Albatross station 4784, depth 135 fathoms. Holotype: USNM 74366. Paratypes: CAS-SU 25237 (2); USNM 70855 (1), 70885 (2), 70892 (1), 70897 (2), 70912 (3). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelus uncinalis Gilbert & Burke 1912 -- (Nelson 1984:52 [ref. 5391], Kim & Youn 1992:67 [ref. 22287], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:454 [ref. 25968], Choi et al. 2003:262, 576 [ref. 26218], Parin et al. 2002:S79 [ref. 28300], Nelson et al. 2004:120 [ref. 27807], Love et al. 2005:89 [ref. 37547], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:257 [ref. 33547], Love et al. 2021:122 [ref. 39279]). Current status: Valid as Icelus uncinalis Gilbert & Burke 1912. Psychrolutidae: Icelinae. Distribution: North Pacific: Bering Sea north of Commander Islands (Russia) and western Aleutian Islands (U.S.A.). Habitat: marine.

uncinatus, Cottus Reinhardt [J. C. H.] 1834:2-3 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1833-34; ref. 26882] Nanortalik, District of Julianehåb, Greenland. Syntypes: ZMUC (1), plus one syntype lost. Type catalog: Nielsen 1974:81 [ref. 9588]. Also appeared in Kgl. Danske Vidensk. Selsk. Natur. Math. Afhandl., 1837, v. 6:xliv. •Valid as Artediellus uncinatus (Reinhardt 1834) -- (Neelov 1973:593 [ref. 7219], Neelov 1979:153 [ref. 3152], Fedorov in Whitehead et al. 1986:1246 [ref. 13677], Nelson 1986:43 [ref. 5808], Robins & Ray 1986:281 [ref. 23100], Scott & Scott 1988:491 [ref. 25518], McAllister 1990:194 [ref. 14674], Yabe in Okamura et al. 1995:162 [ref. 22531], Mecklenburg et al. 2002:403 [ref. 25968], Nelson et al. 2004:118 [ref. 27807] dating to 1835, Møller et al. 2010:51 [ref. 31169] dated to 1835, Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:121 [ref. 32708], Afonso et al. 2016:41, 101 [ref. 36199], Mecklenburg et al. 2018:190 [ref. 36370], Simian et al. 2022:375 [ref. 40776]). Current status: Valid as Artediellus uncinatus (Reinhardt 1834). Psychrolutidae: Psychrolutinae. Distribution: North Atlantic and Arctic Ocean. Habitat: marine.

vaenernensis, Cottus quadricornis Lönnberg [E.] 1932:8, Fig. 3 [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Vänern, Sweden. Syntypes: NRM 20336-38 (1, 4, 1), 40338 (1). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

vandesandei, Gymnocanthus Poll [M.] 1949:234, Figs. 13-14 [Mémoires, Institut Royal des Sciences Naturelles de Belgique 2.Séries, No. 33; ref. 13028] Pulpito Bay, Rio de Oro, Spanish Sahara, depth 10-15 fathoms. Holotype (unique): IRSNB 91. Type catalog: Walschaerts 1987:25 [ref. 20755], Derijst 1991:43 [ref. 21528]. •Valid as Gymnocanthus vandesandei Poll 1949. Current status: Valid as Gymnocanthus vandesandei Poll 1949. Psychrolutidae: Icelinae. Distribution: Eastern Atlantic. Habitat: marine.

variabilis, Cottus Ayres [W. O.] 1842:68 [Proceedings of the Boston Society of Natural History v. 1 (1841-1844); ref. 15926] Brookhaven, Long Island, New York, U.S.A. Appeared first as name only in Ayres 1842:58 [ref. 15927]. Also in Ayres 1842:259 [ref. 15930]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

ventralis, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:194, Pl. 79 (fig. 1) [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka, Russia. Holotype (unique): BMNH 2011.5.17.3. Type catalog: Blanc & Hureau 1968:60 [ref. 20738]. •Synonym of Gymnocanthus pistilliger (Pallas 1814) -- (Watanabe 1960:56 [ref. 11959], Lindberg & Krasyukova 1987:199 [ref. 15964], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Dyldin & Orlov 2017:63 [ref. 35138]). Current status: Synonym of Gymnocanthus pistilliger (Pallas 1814). Psychrolutidae: Icelinae. Habitat: marine.

vermelnensis, Cottus quadricornis Nybelin [O.] 1940:77, Fig. 1 [Fauna och Flora (Uppsala) v. 35 (no. 2); ref. 23062] Lake Värmeln, Sweden. Syntypes: whereabouts unknown. •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Habitat: freshwater, brackish, marine.

vermiculata, Ebinania Sakamoto [K.] 1932:2, Fig. 1 [Journal of the Imperial Fisheries Institute v. 27 (no. 1); ref. 3860] Iwate Prefecture, Japan. Holotype (unique): whereabouts unknown. •Valid as Ebinania vermiculata Sakamoto 1932 -- (Watanabe 1960:198 [ref. 11959], Yabe in Amaoka et al. 1983:157, 214 [ref. 8379], Yabe in Masuda et al. 1984:330 [ref. 6441], Shinohara et al. 1996:176 [ref. 26605], Nakabo 2000:652 [ref. 25086], Nakabo 2002:652 [ref. 26001], Parin et al. 2002:S92 [ref. 28300], Jackson & Nelson 2006:42 [ref. 28707], Meguro & Motomura 2011:27 [ref. 32399], Parin et al. 2014:300 [ref. 33547]). Current status: Valid as Ebinania vermiculata Sakamoto 1932. Psychrolutidae: Psychrolutinae. Distribution: Western North Pacific. Habitat: marine.

verrucosus, Cottus Bean [T. H.] 1881:152 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Plover Bay, anchorage in Providence Bay, Chukchi Peninsula, northeastern Siberia, Russia, Bering Sea. Holotype (unique): USNM 27547. Non-types: MNHN 1896-0467 (1) station 3232. Type catalog: Blanc & Hureau 1968:52 [ref. 20738]. See nomenclatural comments in Mecklenburg et al. 2007:175-176 [ref. 29448]. •Valid as Myoxocephalus verrucosus (Bean 1881) -- (Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S83 [ref. 28300], Fedorov et al. 2003:78 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Parin et al. 2014:266 [ref. 33547], Chernova et al. 2022:9 [ref. 39177]). •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Mecklenburg et al. 2002:405 [ref. 25968], Mecklenburg et al. 2007:175 [ref. 29448], Mecklenburg et al. 2011:123 [ref. 31212]). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific and adjacent Arctic Ocean (if valid). Habitat: marine.

vicinalis, Icelus Gilbert [C. H.] 1896:413 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bristol Bay, Alaska, U.S.A., 53°33'50"N, 167°46'50"W, Albatross station 3324, depth 200 meters. Lectotype: USNM 48737. Paralectotypes: BMNH 1900.7.11.8 (1); CAS 45714 [ex IU 6887] (2); MCZ 28344 (2); MNHN 1896-0487 and 0488 (2) station 3331; CAS-SU 3018 (12), 3484 (1); USNM 48597 (1), 53038 (2), 103605 (2), 233758-59 (1, 2); ZMUO J1464 (2, dry) station 3331. Type catalog: Böhlke 1953:130 [ref. 12291], Blanc & Hureau 1968:54 [ref. 20738], Pethon 1969:8 [ref. 19268]. Lectotype established by Jordan & Evermann 1898:1917 [ref. 2444], but if USNM 48737 contained more than one specimen at that time then lectotype selected by Nelson 1984:30 [ref. 5391]. •Synonym of Icelus euryops Bean 1890 -- (Nelson 1984:30 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:453 [ref. 25968], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:254 [ref. 33547]). Current status: Synonym of Icelus euryops Bean 1890. Psychrolutidae: Icelinae. Habitat: marine.

vinculus, Radulinus (Radulinellus) Bolin [R. L.] 1950:197, Fig. 1 [Copeia 1950 (no. 3); ref. 510] Bechers Bay, Santa Rosa Island, California, U.S.A., depth 15 fathoms. Holotype (unique): CAS-SU 15491. Type catalog: Böhlke 1953:132 [ref. 12291]. •Valid as Radulinus vinculus Bolin 1950 -- (Eschmeyer & Herald 1983:182 [ref. 9277], Yabe & Maruyama 2001:59 [ref. 25245], Nelson et al. 2004:121 [ref. 27807], Love et al. 2005:91 [ref. 37547], Page et al. 2013:124 [ref. 32708], Love et al. 2021:125 [ref. 39279]). Current status: Valid as Radulinus vinculus Bolin 1950. Psychrolutidae: Radulininae. Distribution: Northeastern Pacific: central to southern California (U.S.A.). Habitat: marine.

virginianus, Cottus Cuvier [G.] (ex Willoughby) 1829:163 [Le Règne Animal (Edition 2) v. 2; ref. 995] Available from footnote as, "(1) C. virginianus, Will. x, 15, ou octodecim spinosus, Mitchill. Trans. New-York, iv, p. 380." •Synonym of Myoxocephalus octodecemspinosus (Mitchill 1814). Current status: Synonym of Myoxocephalus octodecemspinosus (Mitchill 1814). Psychrolutidae: Myoxocephalinae. Habitat: brackish, marine.

wosnessenskii, Oligocottus Schmidt [P. Yu.] 1903:518 [16] [Izvestiya Imperatorskago Russkago Geograficheskago Obshchestva v. 38 (pt 5) (for 1902); ref. 3945] On p. 16 of separate. Not available; appeared without distinguishing features as above and also in synonymy in Schmidt 1904:93 [ref. 3946]. •In the synonymy of Oligocottus maculosus Girard 1856 -- (Bolin 1944:65 [ref. 6379], Parin et al. 2002:S83 [ref. 28300]). Nomen Nudum. Current status: Synonym of Oligocottus maculosus Girard 1856. Psychrolutidae: Oligocottinae.

xenogrammus, Stlegicottus Bolin [R. L.] 1936:36, Fig. 8 [Proceedings of the United States National Museum v. 84 (no. 3000); ref. 505] 150 miles north of Rat Island, Bering Sea, Albatross station 3785, depth 270 fathoms. Holotype (unique): USNM 102108. •Valid as Stlegicottus xenogrammus Bolin 1936 -- (Mecklenburg et al. 2002:458 [ref. 25968], Love et al. 2005:92 [ref. 37547], ). •Synonym of Rastrinus scutiger (Bean 1890) -- (Stevenson 2006:792 [ref. 28923], Parin et al. 2014:271 [ref. 33547]). Current status: Synonym of Rastrinus scutiger (Bean 1890). Psychrolutidae: Icelinae. Habitat: marine.

xenostethus, Triglops Gilbert [C. H.] 1896:429, Pl. 29 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] North of Unalaska Island, 54°15'N, 165°06'W, Alaska, U.S.A., Albatross station 3220, depth 34 fathoms. Holotype (unique): whereabouts unknown. •Valid as Triglops xenostethus Gilbert 1896 -- (Pietsch 1994:344 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:421 [ref. 25968], Parin et al. 2002:S89 [ref. 28300], Nelson et al. 2004:122 [ref. 27807], Love et al. 2005:93 [ref. 37547], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:278 [ref. 33547], Love et al. 2021:127 [ref. 39279]). Current status: Valid as Triglops xenostethus Gilbert 1896. Psychrolutidae: Triglopinae. Distribution: North Pacific: Kuril Islands, Commander Islands (Russia), Aleutian Islands (U.S.A.). Habitat: marine.

yesoensis, Myoxocephalus Snyder [J. O.] 1911:538 [Proceedings of the United States National Museum v. 40 (no. 1836); ref. 4152] Hakodate market, Hokkaido, Japan. Holotype (unique): USNM 68238. •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Berg 1949:1137 [ref. 20662]). •Valid as Myoxocephalus yesoensis Snyder 1911 -- (Watanabe 1960:70 [ref. 11959]). •Synonym of Myxocephalus brandtii (Steindachner 1867) -- (Lindberg & Krasyukova 1987:210 [ref. 15964] as yessoensis and brandti, Parin et al. 2014:261 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus brandtii (Steindachner 1867). Psychrolutidae: Myoxocephalinae. Habitat: marine.

zanclus, Bero Snyder [J. O.] 1911:540 [Proceedings of the United States National Museum v. 40 (no. 1836); ref. 4152] Otaru market, Hokkaido, Japan. Holotype (unique): USNM 68239. •Valid as Bero zanclus Snyder 1911 -- (Yabe in Masuda et al. 1984:329 [ref. 6441]). •Synonym of Alcichthys elongatus (Steindachner 1881) -- (Nakabo 2002:1526 [ref. 26001]). Current status: Synonym of Alcichthys elongatus (Steindachner 1881). Psychrolutidae: Psychrolutinae. Habitat: marine.

zanderi, Argyrocottus Herzenstein [S. M.] 1892:219 [49] [Mélanges Biologiques, tirés du Bulletin physico-mathématique de l'Académie Impériale des Sciences de St. Pétersbourg v. 13 (pt 2); ref. 5037] Korsakov, Sakhalin Island, Sea of Okhotsk, Russia. Holotype (unique): ZIN 9679 (lost). On p. 49 of Bull. Acad. Imp. Sci. St. Petersbourg (n. s.) v. 3. •Valid as Argyrocottus zanderi Herzenstein 1892 -- (Watanabe 1960:128 [ref. 11959], Neelov 1979:141 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:225 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:644 [ref. 25086], Nakabo 2002:644 [ref. 26001], Parin et al. 2002:S68 [ref. 28300], Fedorov et al. 2003:64 [ref. 27579], Parin et al. 2014:234 [ref. 33547], Dyldin & Orlov 2022:39 [ref. 38972]). Current status: Valid as Argyrocottus zanderi Herzenstein 1892. Psychrolutidae: Myoxocephalinae. Distribution: Northwestern Pacific. Habitat: marine.

zebra, Psychrolutes Bean [T. H.] 1890:43 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Between Unga Island and Nagai Island, Alaska, U.S.A., 55°10'N, 160°18'W, Albatross station 2848, depth 110 fathoms. Syntypes: MCZ 27914 (1), USNM 45364 (24). •Synonym of Psychrolutes paradoxus Günther 1861 -- (Mecklenburg et al. 2002:524 [ref. 25968], Parin et al. 2002:S94 [ref. 28300], Parin et al. 2014:303 [ref. 33547]). Current status: Synonym of Psychrolutes paradoxus Günther 1861. Psychrolutidae: Psychrolutinae. Habitat: marine.

zonostigma, Pseudoblennius Jordan [D. S.] & Starks [E. C.] 1904:312, Fig. 35 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Misaki, Japan. Holotype: CAS-SU 7718. Paratypes: USNM 50927 (1). Additional material: CAS-SU 7565 (1). Type catalog: Böhlke 1953:131-132 [ref. 12291]. •Valid as Pseudoblennius zonostigma Jordan & Starks 1904 -- (Watanabe 1960:155 [ref. 11959], Nakabo in Masuda et al. 1984:330 [ref. 6441], Kimura et al. 1988:19 [ref. 6692], Kim & Youn 1992:74 [ref. 22287], Nakabo 2000:649 [ref. 25086], Nakabo 2002:649 [ref. 26001], Choi et al. 2003:263, 577 [ref. 26218], Sonoyama et al. 2020:90 [ref. 37637]). Current status: Valid as Pseudoblennius zonostigma Jordan & Starks 1904. Psychrolutidae: Psychrolutinae. Distribution: Northwestern Pacific: southern Korea and Japan. Habitat: marine.

zonurus, Malacocottus Bean [T. H.] 1890:43 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Off Trinity Island, Alaska, U.S.A., 56°00'N, 154°20'W, Albatross station 2853, depth 159 fathoms. Holotype: USNM 44643. Paratypes: USNM 363488 (6). Author says, "The type of the description and figure is 4 3/4 inches long." That is probably not sufficient to establish a holotype, but the 4 3/4-in. specimen if identifiable and is regarded as the described syntype; it would make the best lectotype [others may regard that specimen as the type]. •Valid as Malacocottus zonurus Bean 1890 -- (Yabe in Amaoka et al. 1983:157, 215, 289, 339 [ref. 8379], Yabe in Masuda et al. 1984:330 [ref. 6441], Lindberg & Krasyukova 1987:271 [ref. 15964], Matarese et al. 1989:442 [ref. 26885], McAllister 1990:204 [ref. 14674] as Malacottus, Shinohara et al. 1992:422 [ref. 19776], Kim et al. 1993:21 [ref. 22288], Shinohara et al. 1996:177 [ref. 26605], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:651 [ref. 25086], Sheiko & Fedorov 2000:30 [ref. 25650], Tokranov & Orlov 2001:482 [516] [ref. 38795], Mecklenburg et al. 2002:520 [ref. 25968], Nakabo 2002:651 [ref. 26001], Choi et al. 2003:276, 583 [ref. 26218], Parin et al. 2002:S94 [ref. 28300], Fedorov et al. 2003:87 [ref. 27579], Nelson et al. 2004:123 [ref. 27807], Love et al. 2005:95 [ref. 37547], Page et al. 2013:126 [ref. 32708], Shinohara et al. 2014:254 [ref. 33330], Parin et al. 2014:302 [ref. 33547], Stevenson 2015:23 [ref. 33680], Tohkairin et al. 2015:8 [ref. 35331], Kells et al. 2016:208 [ref. 35888], Kim et al. 2020:107 [ref. 39118], Sonoyama et al. 2020:91 [ref. 37637], Love et al. 2021:123 [ref. 39279], Dyldin & Orlov 2022:55 [ref. 38972]). Current status: Valid as Malacocottus zonurus Bean 1890. Psychrolutidae: Psychrolutinae. Distribution: North Pacific: Sea of Japan and Sea of Okhotsk north to Commander Islands (Russia) and Aleutian Islands (U.S.A.), then south to Washington (U.S.A.). Habitat: marine.


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