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Eschmeyer's Catalog of Fishes
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Species in the family or subfamily Squatinidae: [ 40 ] records


acephala, Squalraia De la Pylaie [A. J. M.] 1835:527 [Congrès Scientifique de France v. 2 (Sept. 1834) (art. 5); ref. 1086] . No distinguishing features, not available. France. •Questionably in the synonymy of Squatina squatina (Linnaeus 1758) -- (Compagno 1984:150 [ref. 6474]). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae.

aculeata, Squatina Cuvier [G.] (ex Duméril) 1829:394 [Le Règne Animal (Edition 2) v. 2; ref. 995] Off Marseille, France, western Mediterranean Sea. Holotype (unique): MNHN 0000-1218. Type catalog: Bertin 1939:77 [ref. 20739] but dated to Duméril 1865. Available from footnote as, "Squat. aculaeta, Dumer., de la Méditerranée, une ranagée de fortes épines le long du dos." •Valid as Squatina aculeata Cuvier 1829 -- (Krefft 1973:49 [ref. 7166], Compagno 1984:140 [ref. 6474], Roux in Whitehead et al. 1984:148 [ref. 13675], Springer 1990:77 [ref. 19315], Bianchi & Carpenter in Bianchi et al. 1993:85 [ref. 25472], Golani 1996:20 [ref. 22271], Compagno 1999:475 [ref. 25589], Bilecenoğlu et al. 2002:18 [ref. 26753], Golani 2005:12 [ref. 37112], Fricke et al. 2007:16 [ref. 29533], Last & White 2008:2 [ref. 29746], Weigmann 2016:69 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Carneiro et al. 2019:39 [ref. 37250], Compagno 2016:1212 [ref. 34544] with author as Duméril in Cuvier, Almeida & Biscoito 2019:116 [ref. 37302], Giovos et al. 2019:4 [ref. 37099], Bariche & Fricke 2020:29 [ref. 37515]). Current status: Valid as Squatina aculeata Cuvier 1829. Squatinidae. Distribution: Mediterranean Sea; eastern Atlantic: southern Portugal south to Namibia, including Selvagens Islands (Portugal). Habitat: marine.

africana, Squatina Regan [C. T.] 1908:248, Pl. 38 [Annals of the Natal Museum v. 1 (pt 3); ref. 14355] Bird Island, Durban Bay, KwaZulu-Natal, South Africa, southwestern Indian Ocean. Holotype (unique): BMNH 1906.11.19.21. •Valid as Squatina africana Regan 1908 -- (Compagno 1984:141 [ref. 6474], Bass 1986:107 [ref. 5635], Gubanov & Timoxin 1989:226 [ref. 36138], Fricke 1999:28 [ref. 24106], Compagno 1999:475 [ref. 25589], Manilo & Bogorodsky 2003:S93 [ref. 27377], Last & White 2008:2 [ref. 29746], Theiss & Ebert 2013:83 [ref. 33034], Weigmann 2016:69 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Fricke et al. 2018:21 [ref. 35805]). Current status: Valid as Squatina africana Regan 1908. Squatinidae. Distribution: Western Indian Ocean: East Africa, South Africa to Madagascar. Habitat: marine.

albipunctata, Squatina Last [P. R.] & White [W. T.] 2008:5, Figs. 2-3, 8a [Zootaxa No. 1734; ref. 29746] South of Lakes Entrance, Victoria, Australia, 38°42'S, 148°18'E, depth 84-92 meters. Holotype: CSIRO H 6561-01. Paratypes: 12 specimens at CSIRO. Plus non-type material. •Valid as Squatina albipunctata Last & White 2008 -- (Theiss & Ebert 2013:83 [ref. 33034], Weigmann 2016:69 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531]). Current status: Valid as Squatina albipunctata Last & White 2008. Squatinidae. Distribution: Australia: Queensland to Victoria. Habitat: marine.

angelina, Squatina Gray [J. E.] 1851:80 [List of the specimens of fish in the collection of the British Museum.; ref. 4939] Not available, no distinguishing features provided; specimen from Suriname. Squatinidae.

angelorum, Cestracion Swainson [W.] 1838:137 [The natural history and classification of fishes, amphibians, & reptiles, or monocardian animals. v. 1; ref. 4302] Europe. No types known. Treated as Squatina angelorum on p. 184. •Synonym of Squatina squatina (Linnaeus 1758) -- (Dyldin 2015:58 [ref. 34524]). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae. Habitat: marine.

angelorum, Squatina Swainson [W.] 1838:184, Figs. 22-23 [The natural history and classification of fishes, amphibians, & reptiles, or monocardian animals. v. 1; ref. 4302] European seas. No types known. Treated as C. angelorum on p. 137. Perhaps best considered an unjustified emendation. •Synonym of Squatina squatina (Linnaeus 1758). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae.

angelus, Squatina Blainville [H. de] 1825:53 [Faune Française; ou histoire naturelle Livr. 13 & 14; ref. 4991] France. Apparently not an original description; if it is then it is a new name for Squalus squatina Linnaeus based on footnote on p. 54. Preoccupied by Squatina angelus Duméril 1808 [not investigated]. •In the synonymy of Squatina squatina (Linnaeus 1758) -- (Bilecenoğlu et al. 2002:19 [ref. 26753], Compagno 2016:1218 [ref. 34544] dated 1816). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae.

angelus, Squatina Gronow [L. T.] in Gray 1854:14 [Catalogue of fish collected and described by Laurence Theodore Gronow; ref. 1911] European seas. Syntypes: BMNH 1853.11.12.207 [Gronovius coll.] (1, skin). Type catalog: Wheeler 1958:204-205 [ref. 13434] but as holotype. Objectively invalid; preoccupied. •Synonym of Squatina squatina (Linnaeus 1758) -- (Wheeler 1958:204 [ref. 13434], Lipej & Dulčić 2010:11 [ref. 36649], Dyldin 2015:58 [ref. 34524]). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae. Habitat: marine.

argentina, Rhina Marini [T. L.] 1930:5, Fig. 1 [Physis (Buenos Aires) v. 10 (no. 35); ref. 16108] Off Province of Buenos Aires, about 39°S, 56°W, Argentina. Holotype (unique): MACN 574. Type catalog: Braga & Piacentino 1994:99 [ref. 23697]. •Valid as Squatina argentina (Marini 1930) -- (Compagno 1984:142 [ref. 6474], Menni 1986:433 [ref. 11487], Vooren & da Silva 1992:597 [ref. 21060], Compagno 1999:475 [ref. 25589], Cousseau & Perrotta 2000:38 [ref. 25942], Soto 2001:66, 96 [ref. 26637], López et al. 2002:62 [ref. 26808], Gadig & Gomes in Menezes et al. 2003:27 [ref. 27192], Castro-Aguirre et al. 2007:1039 [ref. 29408], Last & White 2008:2 [ref. 29746], Vaz & Carvalho 2013:34 [ref. 32825], Weigmann 2016:69 [ref. 34211], Estalles et al. 2016:1 [ref. 34759], Nión et al. 2016:21 [ref. 35565]). Current status: Valid as Squatina argentina (Marini 1930). Squatinidae. Distribution: Southwestern Atlantic: Brazil to Uruguay and Argentina. Habitat: marine.

armata, Rhina Philippi [R. A.] 1887:561 [29], Pl. 7 (fig. 1) [Anales de la Universidad de Chile Sec. 1, v. 71; ref. 3464] Iquique, Chile. Holotype (unique): DBCUCH. On p. 29 of separate. •Valid as Rhina armata Philippi 1887 -- (Pequeño 1989:12 [ref. 14125]). •Synonym of Squatina californica Ayres 1859 -- (Castro 2011:163 [ref. 31457]). •Questionably a synonym of Squatina californica Ayres 1859 -- (Compagno 1984:144 [ref. 6474], Chirichigno F. & Vélez D. 1998:30 [ref. 24555], Last & White 2008:2 [ref. 29746]). •Valid as Squatina armata (Philippi 1887) -- (Weigmann 2016:69 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531]). Current status: Valid as Squatina armata (Philippi 1887). Squatinidae. Distribution: Southeastern Pacific. Habitat: marine.

australis, Squatina Regan [C. T.] 1906:438 [Annals and Magazine of Natural History (Series 7) v. 18 (no. 108) (art. 70); ref. 15711] Port Jackson, New South Wales, Australia. Holotype (unique): BMNH 1890.9.23.181. •Valid as Squatina australis Regan 1906 -- (Compagno 1984:143 [ref. 6474], Paxton et al. 1989:62 [ref. 12442], Kuiter 1993:4 [ref. 23929], Gomon et al. 1994:152 [ref. 22532], Last & Stevens 1994:280 [ref. 23873], Kuiter 1997:12 [ref. 25488], Compagno & Niem 1998:1237 [ref. 23787], Compagno 1999:475 [ref. 25589], Hutchins 2001:15 [ref. 25847], Hoese & Gates 2006:159 [ref. 28997], Last & White 2008:2 [ref. 29746], White 2008:97 [ref. 30617], Theiss & Ebert 2013:83 [ref. 33034], Weigmann 2016:69 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Hoschke et al. 2019:153[ref. 36626]). Current status: Valid as Squatina australis Regan 1906. Squatinidae. Distribution: Southeastern Indian Ocean: Victoria, Tasmania, South Australia and Western Australia. Habitat: marine.

caillieti, Squatina Walsh [J. H.], Ebert [D. A.] & Compagno [L. J. V.] 2011:50, Figs. 1-2, 4A, 5A [Zootaxa No. 2759; ref. 31151] Luzon, 13°08.98-13'09.84'N, 124°04.72-124°00.01'E, Philippines, depth 363-385 meters. Holotype: CAS 226473. •Valid as Squatina caillieti Walsh, Ebert & Compagno 2011 -- (Theiss & Ebert 2013:83 [ref. 33034], Weigmann 2016:69 [ref. 34211]). Current status: Valid as Squatina caillieti Walsh, Ebert & Compagno 2011. Squatinidae. Distribution: Philippines. Habitat: marine.

californica, Squatina Ayres [W. O.] 1859:29, Fig. 7 (p. 56) [Proceedings of the California Academy of Sciences (Series 1) v. 2 (1858-1862); ref. 155] San Francisco Bay, California, U.S.A. Holotype (unique): CAS (lost). As Rhina californica in caption to Fig. 7 (on p. 54). •Valid as Squatina californica Ayres 1859 -- (Eschmeyer & Herald 1983:44 [ref. 9277], Compagno 1984:144 [ref. 6474], Allen & Robertson 1994:18 [ref. 22193], Bussing & López S. 1994:24 [ref. 23101], Compagno et al. in Fischer et al. 1995:729 [ref. 22829], Gaida 1995:532 [ref. 22403], Chirichigno F. & Vélez D. 1998:30 [ref. 24555], Compagno 1999:475 [ref. 25589], Mecklenburg et al. 2002:89 [ref. 25968], Espinosa Pérez et al. 2004:105 [ref. 27705], Nelson et al. 2004:50 [ref. 27807], Love et al. 2005:4 [ref. 37547], Castro 2011:163 [ref. 31457], Page et al. 2013:54 [ref. 32708], Pietsch & Orr 2015:16 [ref. 34694], Del Moral-Flores et al. 2016:66 [ref. 34398], Weigmann 2016:69 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Kells et al. 2016:76 [ref. 35888], Ebert et al. 2017:52 [ref. 35618], Ehemann et al. 2018:24 [ref. 36194], Burton & Lea 2019:30 [ref. 37205]). Current status: Valid as Squatina californica Ayres 1859. Squatinidae. Distribution: Eastern Pacific: Puget Sound (Washington, U.S.A.) south to Pacific coast of Baja California Sur (Mexico); Ecuador south to Chile (needs confirmation [ref. 35618]; questionable from Alaska (U.S.A.)). Habitat: marine.

cervicata, Squalraia De la Pylaie [A. J. M.] 1835:527 [Congrès Scientifique de France v. 2 (Sept. 1834) (art. 5); ref. 1086] No distinguishing features, not available. •Questionably in the synonymy of Squatina squatina (Linnaeus 1758) -- (Compagno 1984:150 [ref. 6474]). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae.

david, Squatina Acero P. [A.], Tavera [J. J.], Anguila [R.] & Hernández [L.] 2016:579, Figs. 1-5 [Copeia v. 104 (no. 2); ref. 34531] Bocas de Ceniza, Colombia, 11°00'33"N, 75°03'19"W, depth 144 meters. Holotype: INVEMAR PEC8250. Paratypes: INVEMAR PEC. •Valid as Squatina david Acero P., Tavera, Anguila & Hernández 2016 -- (Vaz & Carvalho 2018:159 [ref. 35832], Ehemann et al. 2019:[5] [ref. 36861]). Current status: Valid as Squatina david Acero P., Tavera, Anguila & Hernández 2016. Squatinidae. Distribution: Western Atlantic: Panama, Colombia, Venezuela (southern Caribbean); Suriname. Habitat: marine.

dumeril, Squatina Lesueur [C. A.] 1818:225, Pl. 10 [Journal of the Academy of Natural Sciences, Philadelphia v. 1 (pt 2); ref. 2735] East coast of U.S.A., off New York city, 40°N, 67°W. Lectotype: MNHN A-9692 (dry). Paralectotype: (2) missing. Type catalog: Bertin 1939:77 [ref. 20739]. Lectotype selected by Vaz & Carvalho 2013:50 [ref. 32825]. •Valid as Squatina dumerili Lesueur 1818 -- (Robins & Ray 1986:34 [ref. 23100], Castro 2011:167 [ref. 31457]). •Valid as Squatina dumeril Lesueur 1818 -- (Compagno 1984:145 [ref. 6474], Boschung 1992:25 [ref. 23239], Cervigón 1992:190 [ref. 23827], Murdy et al. 1997:35 [ref. 23144], McEachran & Fechhelm 1998:128 [ref. 23897], Cervigón & Alcalá 1999:133 [ref. 24490], Compagno 1999:475 [ref. 25589], Soto 2001:66, 96 [ref. 26637], Compagno 2003:415 [ref. 26984], Moore et al. 2003:170 [ref. 27076], Gadig & Gomes in Menezes et al. 2003:27 [ref. 27192], Moore et al. 2003:170 [ref. 27076], Espinosa Pérez et al. 2004:106 [ref. 27705], Nelson et al. 2004:50 [ref. 27807], Castro-Aguirre et al. 2007:1039 [ref. 29408], Page et al. 2013:54 [ref. 32708], Vaz & Carvalho 2013:50 [ref. 32825], Del Moral-Flores et al. 2016:66 [ref. 34398], Weigmann 2016:69 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Ehemann et al. 2018:24 [ref. 36194], Vaz & Carvalho 2018:159 [ref. 35832], Ehemann et al. 2019:[5] [ref. 36861]). Current status: Valid as Squatina dumeril Lesueur 1818. Squatinidae. Distribution: Western Atlantic (including Caribbean Sea). Habitat: marine.

europaea, Squatina Swainson [W.] 1839:321 [The natural history and classification v. 2; ref. 4303] No types known. Original genus as Squatinae. Based on Bl. 116 -- which is Bloch 1785:Pl. 118 [ref. 4866]. •Synonym of Squatina squatina (Linnaeus 1758) -- (Compagno 1984:150 [ref. 6474], Dyldin 2015:58 [ref. 34524]). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae. Habitat: marine.

fimbriata, Squatina Müller [J.] & Henle [F. G. J.] 1839:101, 192, [Pl. 35 (head)] [Systematische Beschreibung der Plagiostomen; ref. 3069] [Mittelmeer] Mediterranean Sea. No types known. Syntypes: (2) ZMB uncat. (1) plus 1 missing. Type catalog: Paepke & Schmidt 1988:161 [ref. 21041]. •Questionably a synonym of Squatina aculeata Cuvier 1829 -- (Krefft 1973:49 [ref. 7166], Compagno 1984:140 [ref. 6474]). Current status: Uncertain as Squatina aculeata Cuvier 1829. Squatinidae. Habitat: marine.

formosa, Squatina Shen [S.-C.] & Ting [W.-H.] 1972:21, Fig. 4 [Bulletin of the Institute of Zoology Academia Sinica (Taipei) v. 11 (no. 1); ref. 7645] Tung-kong [Tongkang], 20°28'N, 120°26.3'E, Pingtung, southwestern Taiwan, depth 180 meters. Holotype: NTT 7213130 [now NTUM 01329]. Paratypes: NTUM 7041631-32 (1, 1), 7222433 (1). Type catalog: Ho & Shao 2011:19 [ref. 31432]. •Valid as Squatina formosa Shen & Ting 1972 -- (Compagno 1984:146 [ref. 6474], Compagno 1999:475 [ref. 25589], Compagno in Randall & Lim 2000:581 [ref. 25122], Compagno et al. 2005:56 [ref. 29145], Walsh & Ebert 2007:35 [ref. 29245] with paratypes as nebulosa, Kriwet et al. 2010:117 [ref. 30825], Walsh 2011:58 [ref. 31151], Ho & Shao 2011:19 [ref. 31432], Yamashita et al. 2012:127 [ref. 32408], Theiss & Ebert 2013:83 [ref. 33034], Ebert et al. 2013:300 [ref. 33045], Weigmann 2016:70 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531]). Current status: Valid as Squatina formosa Shen & Ting 1972. Squatinidae. Distribution: Western North Pacific. Habitat: marine.

guggenheim, Squatina Marini [T. L.] 1936:23, Fig. 2 [Physis (Buenos Aires) v. 12; ref. 13716] Necochea, Buenos Aires Province, Argentina. Holotype (unique): MACN 938. Type catalog: Braga & Piacentino 1994:101 [ref. 23697], Soto & Mincarone 2004:84 [ref. 37067]. •Synonym of Squatina argentina (Marini 1930) -- (Compagno 1984:142 [ref. 6474]). •Valid as Squatina guggenheim Marini 1936 -- (Vooren & da Silva 1992:595 [ref. 21060], Compagno 1999:475 [ref. 25589], Soto 2001:66, 97 [ref. 26637], Gadig & Gomes in Menezes et al. 2003:27 [ref. 27192], Castro-Aguirre et al. 2007:1039 [ref. 29408], Last & White 2008:2 [ref. 29746], Mabragaña et al. 2011:table S2 [ref. 31800], Vaz & Carvalho 2013:19 [ref. 32825], Weigmann 2016:70 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Estalles et al. 2016:1 [ref. 34759], Petry et al. 2016:299 [ref. 34966], Nión et al. 2016:21 [ref. 35565], Spier et al. 2018:4 [ref. 36103]). Current status: Valid as Squatina guggenheim Marini 1936. Squatinidae. Distribution: Southwestern Atlantic: Brazil south to Argentina. Habitat: marine.

heteroptera, Squatina Castro-Aguirre [J. L.], Espinosa Pérez [H.] & Huidobro Campos [L.] 2007:1036, Figs. 5-8 [Revista de Biología Tropical v. 54 (no. 3) [for Sept. 2006]; ref. 29408] Off Tamaulipas, 22°44'01"N, 97°24'07"W, depth 157-164 meters. Holotype: IBUNAM-P 8307. Paratypes: IBUNAM-P 12722 (1). •Valid as Squatina heteroptera Castro-Aguirre, Espinosa Pérez & Huidobro Campos 2007 -- (Castro 2011:170 [ref. 31457], Page et al. 2013:54 [ref. 32708], Vaz & Carvalho 2013:74 [ref. 32825], Del Moral-Flores et al. 2016:67 [ref. 34398], Ehemann et al. 2018:24 [ref. 36194]). •Synonym of Squatina dumeril Lesueur 1818 -- (Weigmann 2016:69 [ref. 34211]). Current status: Synonym of Squatina dumeril Lesueur 1818. Squatinidae. Distribution: Western Gulf of Mexico (if valid). Habitat: marine.

japonica, Squatina Bleeker [P.] 1858:40 [Acta Societatis Regiae Scientiarum Indo-Neêrlandicae v. 3 (art. 10); ref. 360] Nagasaki, Japan. Holotype (unique): RMNH 7416 (largest of 2 specimens). •Valid as Squatina japonica Bleeker 1858 -- (Compagno 1984:147 [ref. 6474], Nakaya & Shirai in Masuda et al. 1984:11 [ref. 6441], Cheng & Zhou 1997:41 [ref. 26385], Compagno & Niem 1998:1237 [ref. 23787], Senou & Masuda 1998:5 [ref. 24388], Sokolovskaya et al. 1998:7 [ref. 24670], Compagno 1999:475 [ref. 25589], Nakabo 2000:157 [ref. 25086], Compagno in Randall & Lim 2000:581 [ref. 25122], Shinohara et al. 2001:290 [ref. 25995], Nakabo 2002:157 [ref. 26001], Choi et al. 2003:62, 479 [ref. 26218], Compagno et al. 2005:56 [ref. 29145], Walsh & Ebert 2007:40 [ref. 29245], Kriwet et al. 2010:120 [ref. 30825], Walsh 2011:58 [ref. 31151], Yamashita et al. 2012:127 [ref. 32408], Theiss & Ebert 2013:83 [ref. 33034], Ebert et al. 2013:300 [ref. 33045], Shinohara et al. 2014:234 [ref. 33330], Parin et al. 2014:28 [ref. 33547], Dyldin 2015:58 [ref. 34524], Weigmann 2016:70 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Sonoyama et al. 2020:11 [ref. 37637]). Current status: Valid as Squatina japonica Bleeker 1858. Squatinidae. Distribution: Northwestern Pacific. Habitat: marine.

laevis, Squatina Cuvier [G.] 1816:131 [Le Règne Animal v. 2; ref. 993] No locality stated. On Squalus squatina of Bloch, pl. 116. •Synonym of Squatina squatina (Linnaeus 1758) -- (Compagno 1984:150 [ref. 6474], Roux in Whitehead et al. 1984:150 [ref. 13675], Dyldin 2015:58 [ref. 34524], Compagno 2016:1218 [ref. 34544] dated 1817). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae. Habitat: marine.

legnota, Squatina Last [P. R.] & White [W. T.] 2008:12, Figs. 4-5, 8b [Zootaxa No. 1734; ref. 29746] Cilacap landing site, central Java, Indonesia, 7°44'S, 109°00'E. Holotype: MZB 15500. Paratypes: CSIRO H 6565-01 (1). Plus one non-type specimen. •Valid as Squatina legnota Last & White 2008 -- (Kriwet et al. 2010:121 [ref. 30825], Walsh 2011:58 [ref. 31151], Theiss & Ebert 2013:83 [ref. 33034], Weigmann 2016:70 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531]). Current status: Valid as Squatina legnota Last & White 2008. Squatinidae. Distribution: Off southern Indonesia. Habitat: marine.

lewis, Squatina Couch [J.] 1825 Apparently not an original description or perhaps a manuscript name or common name. •In the synonymy of Squatina squatina (Linnaeus 1758) -- (Compagno 1984:150 [ref. 6474]). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae.

machuelo, Raia Osbeck [P.] 1770:99 [Nova Acta Physico-Medica Academiae Caesareae Leopoldina-Carolinae Naturae Curiosorum v. 4; ref. 18497] Hispanicae (Spain). No types known. Original genus should have been spelled Raja. Raja osbeckii Walbaum 1792 [ref. 4572] is based on this species. •Synonym of Squatina aculeata Cuvier 1829. Current status: Synonym of Squatina aculeata Cuvier 1829. Squatinidae. Habitat: marine.

mexicana, Squatina Castro-Aguirre [J. L.], Espinosa Pérez [H.] & Huidobro Campos [L.] 2007:1032, Figs. 1-4 [Revista de Biología Tropical v. 54 (no. 3) [for Sept. 2006]; ref. 29408] Off Tamaulipas, 22°58'05"N, 97°28'07"W, depth 71 meters. Holotype: IBUNAM-P 8306. Paratypes: IBUNAM-P 11136 (2), P-12723 (1), 12724 (1). •Valid as Squatina mexicana Castro-Aguirre, Espinosa Pérez & Campos 2007 -- (Castro 2011:172 [ref. 31457], Page et al. 2013:54 [ref. 32708], Vaz & Carvalho 2013:74 [ref. 32825], Del Moral-Flores et al. 2016:68 [ref. 34398], Ehemann et al. 2018:24 [ref. 36194]). •Synonym of Squatina dumeril Lesueur 1818 -- (Weigmann 2016:69 [ref. 34211]). Current status: Synonym of Squatina dumeril Lesueur 1818. Squatinidae. Distribution: Western Atlantic: Gulf of Mexico (if valid). Habitat: marine.

nebulosa, Squatina Regan [C. T.] 1906:439 [Annals and Magazine of Natural History (Series 7) v. 18 (no. 108) (art. 70); ref. 15711] Japan. Holotype (unique): BMNH 1862.11.1.89. •Valid as Squatina nebulosa Regan 1906 -- (Compagno 1984:148 [ref. 6474], Nakaya in Okamura & Kitajima 1984:61, 305 [ref. 8057], Nakaya & Shirai in Masuda et al. 1984:11 [ref. 6441], Chen et al. 1997:6 [ref. 26476], Senou & Masuda 1998:5 [ref. 24388], Compagno 1999:475 [ref. 25589], Nakabo 2000:157 [ref. 25086], Compagno in Randall & Lim 2000:581 [ref. 25122], Shinohara et al. 2001:290 [ref. 25995], Nakabo 2002:157 [ref. 26001], Choi et al. 2003:62, 479 [ref. 26218], Shinohara et al. 2005:397 [ref. 28370], Walsh & Ebert 2007:42 [ref. 29245], Kriwet et al. 2010:120 [ref. 30825], Walsh 2011:58 [ref. 31151], Theiss & Ebert 2013:83 [ref. 33034], Ebert et al. 2013:301 [ref. 33045], Shinohara et al. 2014:234 [ref. 33330], Weigmann 2016:70 [ref. 34211]). Current status: Valid as Squatina nebulosa Regan 1906. Squatinidae. Distribution: Western North Pacific. Habitat: marine.

occulta, Squatina Vooren [C. M.] & da Silva [K. G.] 1991:592, Fig. 2 [Revista Brasileira de Biologia v. 51 (no. 3) [1991]; ref. 21060] Southern Brazil, 33°06'S, 51°40'W. Holotype: MOVI 10195 [ex MOFURG 06589]. Paratypes: MCP 13999 (1), MZUSP 41518 (1). •Synonym of Squatina guggenheim (Marini 1936) -- (Soto 2001:97 [ref. 26637] with occulta dated 1991 and author as Vooren). •Valid as Squatina occulta Vooren & da Silva 1991 -- (Compagno 1999:475 [ref. 25589], Castro-Aguirre et al. 2007:1039 [ref. 29408], Vaz & Carvalho 2013:5 [ref. 32825], Faria et al. 2014:1 [ref. 33251] with date correction, Weigmann 2016:70 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Estalles et al. 2016:1 [ref. 34759], Nión et al. 2016:21 [ref. 35565], Spier et al. 2018:4 [ref. 36103]). Current status: Valid as Squatina occulta Vooren & da Silva 1991. Squatinidae. Distribution: Southwestern Atlantic: Brazil, Uruguay and Argentina. Habitat: marine.

oculata, Squatina Bonaparte [C. L.] 1840:fasc. 28, punt. 147, Pl. 143 (fig. 2) [Iconografia della fauna italica; ref. 514] Italy, western Mediterranean Sea. No types known. Type catalog: Böhlke 1984:158 [ref. 13621]. •Valid as Squatina oculata Bonaparte 1840 -- (Krefft 1973:50 [ref. 7166], Compagno 1984:149 [ref. 6474], Roux in Whitehead et al. 1984:149 [ref. 13675], Springer 1990:77 [ref. 19315], Bianchi & Carpenter in Bianchi et al. 1993:85 [ref. 25472], Golani 1996:21 [ref. 22271], Compagno 1999:475 [ref. 25589], Bilecenoğlu et al. 2002:19 [ref. 26753], Golani 2005:12 [ref. 37112], Fricke et al. 2007:17 [ref. 29533], Last & White 2008:2 [ref. 29746], Lipej & Dulčić 2010:11 [ref. 36649], Weigmann 2016:71 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Compagno 2016:1217 [ref. 34544], Zava et al. 2017:341 [ref. 35032], Almeida & Biscoito 2019:116 [ref. 37302], Carneiro et al. 2019:39 [ref. 37250], Giovos et al. 2019:4 [ref. 37099], Bariche & Fricke 2020:30 [ref. 37515], Kovačić et al. 2020:7 [ref. 37519]). Current status: Valid as Squatina oculata Bonaparte 1840. Squatinidae. Distribution: Mediterranean Sea; eastern Atlantic: Portugal south to Namibia. Habitat: marine.

osbeckii, Raja Walbaum [J. J.] 1792:532 [Petri Artedi sueci genera piscium Part 3; ref. 4572] Spain, western Mediterranean Sea. No types known. This species is based on Raia machuelo Osbeck 1770. •Synonym of Squatina aculeata Cuvier 1829 -- (P. Parenti, pers. comm.). Current status: Synonym of Squatina aculeata Cuvier 1829. Squatinidae. Habitat: marine.

philippi, Rhina Garman [S.] 1913:254 [Memoirs of the Museum of Comparative Zoology v. 36; ref. 1545] Mejillones, Chile. Syntypes: MCZ 531-S (1). Type catalog: Hartel & Dingerkus 1997:xli [ref. 23119]. •Questionably a synonym of Squatina californica Ayres 1859 -- (Compagno 1984:144 [ref. 6474]). Current status: Uncertain as Squatina californica Ayres 1859. Squatinidae. Distribution: Eastern Pacific. Habitat: marine.

pseudocellata, Squatina Last [P. R.] & White [W. T.] 2008:16, Figs. 6-7, 8c [Zootaxa No. 1734; ref. 29746] West of Dorre Island, Western Australia, 25°09'S, 112°09'E, depth 312 meters. Holotype: CSIRO H 2567-01. Paratypes: CSIRO CA 3291 (1), CA 3663 (1), CA 4474 (1), H 4070-01(1), H 5188-03 (1); WAM P 26195-001 (1). •Valid as Squatina pseudocellata Last & White 2008 -- (Theiss & Ebert 2013:83 [ref. 33034], Weigmann 2016:71 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531]). Current status: Valid as Squatina pseudocellata Last & White 2008. Squatinidae. Distribution: Australia: Western Australia. Habitat: marine.

punctata, Squatina Marini [T. L.] 1936:22, 27, Fig. 4 [Physis (Buenos Aires) v. 12; ref. 13716] Argentina. No types known. Appeared as Squatina puntata twice in text, as punctata in key and caption to Fig. 4. First reviser not researched; punctata was selected by Eschmeyer 1998:1398 [ref. 23416]. Available from key and as name plus figure, with "Su descriptión en el próximo número" [not found]. •Synonym of Squatina argentina Marini 1936 -- (Compagno 1984:142 [ref. 6474]). •Valid as Squatina punctata Marini 1936 -- (Soto 2001:66, 97 [ref. 26637], Gadig & Gomes in Menezes et al. 2003:27 [ref. 27192], Last & White 2008:2 [ref. 29746]). •Synonym of Squatina guggenheim Marini 1936 -- (Vaz & Carvalho 2013:19 [ref. 32825], Weigmann 2016:70 [ref. 34211]). Current status: Synonym of Squatina guggenheim Marini 1936. Squatinidae. Habitat: marine.

squatina, Squalus Linnaeus [C.] 1758:233 [Systema Naturae, Ed. X v. 1; ref. 2787] Mediterranean Sea and northeastern Atlantic [original: "In Oceano Europæo"]; localities include Cornwall, England; Bordeaux, France, Bay of Biscay; Marseille, France; Genoa and Rome, Italy; Lesbos Island, Greece; Syria, Mediterranean Sea. Possible syntypes: ZMUU Linn. coll. 161 (1, dry) and NRM 87 (1). Type catalog: Lönnberg 1896:38 [ref. 23530]. Originally based at least 31 pre-Linnaean sources including Artedi 1738:67 [ref. 30578] and Artedi 1738:95 [ref. 30349] as Squalus pinna ani carens, ore in apice capitis; Rondelet 1554:367 [ref. 30354] as Squatina, Salviani 1558:152, Pl. (p. 151) [ref. 30229] as Squatina, Willughby 1686:79 [ref. 30164] and Ray 1713:26 [ref. 30231] as Squatina. •Valid as Squatina squatina (Linnaeus 1758) -- (Krefft 1973:49 [ref. 7166], Compagno 1984:150 [ref. 6474], Roux in Whitehead et al. 1984:150 [ref. 13675], Dor 1984:12 [ref. 29757], Springer 1990:77 [ref. 19315], Goren & Dor 1994:4 [ref. 25356], Compagno 1999:475 [ref. 25589], Bilecenoğlu et al. 2002:19 [ref. 26753], Golani 2005:12 [ref. 37112], Fricke 2007:25 [ref. 30577], Fricke et al. 2007:17 [ref. 29533], Vasil'eva 2007:19 [ref. 30517], George 2009:56 [ref. 30539], Golani & Bogorodsky 2010:57 [ref. 35264], Lipej & Dulčić 2010:11 [ref. 36649], Parin et al. 2014:28 [ref. 33547], Dyldin 2015:58 [ref. 34524], Weigmann 2016:71 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531], Compagno 2016:1218 [ref. 34544], Almeida & Biscoito 2019:115 [ref. 37302], Carneiro et al. 2019:40 [ref. 37250], Giovos et al. 2019:4 [ref. 37099], Bariche & Fricke 2020:31 [ref. 37515], Kovačić et al. 2020:7 [ref. 37519]). Current status: Valid as Squatina squatina (Linnaeus 1758). Squatinidae. Distribution: Western Baltic Sea, North Sea, Mediterranean Sea, Black Sea, eastern Atlantic: Norway south to Western Sahara, including Canary Islands. Habitat: marine.

tergocellata, Squatina McCulloch [A. R.] 1914:84, Fig. 2; Pl. 15. [Biological Results Endeavour v. 2 (pt 3); ref. 2940] Great Australian Bight, 127°-128°E, South Australia, depth 160-200 fathoms. Holotype (unique): AMS E.4286 (missing). •Valid as Squatina tergocellata McCulloch 1914 -- (Compagno 1984:151 [ref. 6474], Allen & Swainston 1988:26 [ref. 25477], Paxton et al. 1989:62 [ref. 12442], Gomon et al. 1994:153 [ref. 22532], Last & Stevens 1994:281 [ref. 23873], Compagno 1999:475 [ref. 25589], Hutchins 2001:15 [ref. 25847], Hoese & Gates 2006:159 [ref. 28997], Last & White 2008:2 [ref. 29746], White 2008:98 [ref. 30617], Theiss & Ebert 2013:83 [ref. 33034], Weigmann 2016:71 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531]). Current status: Valid as Squatina tergocellata McCulloch 1914. Squatinidae. Distribution: Southern and western Australia. Habitat: marine.

tergocellatoides, Squatina Chen [J. T. F.] 1963:98, Fig. 28 [Biological Bulletin, Tunghai University, Department of Biology, College of Science, Ichthyology Series No. 1; ref. 4449] Taiwan Strait, western Taiwan. Holotype (unique): THUP 00348. Type catalog: Ho & Shao 2011:19 [ref. 31432]. Holotype was thought to be lost by Walsh & Ebert 2007 [ref. 29245], but found by Ho & Shao 2011:19 [ref. 31432]. •Valid as Squatina tergocellatoides Chen 1963 -- (Compagno 1999:475 [ref. 25589], Compagno in Randall & Lim 2000:581 [ref. 25122], Walsh & Ebert 2007:45 [ref. 29245], Kriwet et al. 2010:120 [ref. 30825], Last el al. 2010:42 [ref. 32461], Walsh 2011:58 [ref. 31151], Ho & Shao 2011:19 [ref. 31432], Theiss & Ebert 2013:74 [ref. 33034], Ebert et al. 2013:301 [ref. 33045], Weigmann 2016:71 [ref. 34211], Acero P. et al. 2016:583 [ref. 34531]). Current status: Valid as Squatina tergocellatoides Chen 1963. Squatinidae. Distribution: North Pacific: Taiwan Straits, Vietnam, Hong Kong, Malaysia. Habitat: marine.

varii, Squatina Vaz [D. F. B.] & Carvalho [M. R. de] 2018:145, Figs. 1-8 [Copeia 106 (no. 1); ref. 35832] Continental slope of Espírito Santo state, Brazil, southwestern Atlantic Ocean, 19°42'54"S, 39°25'57"W, depth 195 meters. Holotype: MNRJ 43106. Paratypes: MNRJ. Plus additional non-type material. •Valid as Squatina varii Vaz & Carvalho 2018. Current status: Valid as Squatina varii Vaz & Carvalho 2018. Squatinidae. Distribution: Brazil. Habitat: marine.

vulgaris, Squatina Risso [A.] 1810:45 [Ichthyologie de Nice; ref. 3755] Nice, Fance, northwestern Mediterranean Sea. No types known. •Synonym of Squatina squatina (Linnaeus 1758) -- (Krefft 1973:49 [ref. 7166], Compagno 1984:150 [ref. 6474], Dyldin 2015:58 [ref. 34524], Compagno 2016:1218 [ref. 34544]). Current status: Synonym of Squatina squatina (Linnaeus 1758). Squatinidae. Habitat: marine.