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Eschmeyer's Catalog of Fishes
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Species in the family or subfamily Tripterygiidae: [ 224 ] records


abditus, Gilloblennius Hardy [G. S.] 1986:154, Pl. 1 (bottom 2); figs. 1, 4, 5 [Journal of the Royal Society of New Zealand v. 16 (no. 2); ref. 6139] Snares Island, New Zealand. Holotype: NMNZ P.16367. Paratypes: BMNH 1985.6.19.6 (1); NMNZ P.16368 (2), P.16376 (7), P.16382 (3), P.16385 (3), P.16430 (1), P.16434 (1, c&s), P.17072 (1); USNM 272605 (1). Type catalog: Hardy 1990:13 [ref. 18942], Springer & Orrell 1996:24 [ref. 22274]. •Valid as Gilloblennius abditus Hardy 1986 -- (Paulin et al. 1989:219 [ref. 24556], Paulin & Roberts 1992:80 [ref. 24625], Fricke 1994:353 [ref. 21827], Fricke 1997:575 [ref. 23339], Fricke 2009:33 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Stewart & Clements 2015:1511 [ref. 34299]). Current status: Valid as Gilloblennius abditus Hardy 1986. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand. Habitat: marine.

abeli, Tripterygion Klausewitz [W.] 1960:11, Figs. 1-2 [Senckenbergiana Biologica v. 41 (nos 1/2); ref. 11926] Hurghada, Red Sea Governorate, Egypt, Red Sea. Holotype: SMF 4780. Paratypes: SMF 4781-82 (1, 1). •Valid as Enneapterygius abeli (Klausewitz 1960) -- (Dor 1984:233 [ref. 29757], Holleman 1986:756 [ref. 5671], Fricke 1987:336 [ref. 9108], Winterbottom et al. 1989:62 [ref. 13251], Goren & Dor 1994:61 [ref. 25356], Fricke 1997:565 [ref. 23339], Fricke 1999:466 [ref. 24106], Holleman 2005:4 [ref. 28455], Fricke 2009:38 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Holleman & Bogorodsky 2012:40 [ref. 31735], Fricke et al. 2013:268 [ref. 32706], Fricke et al. 2018:303 [ref. 35805], Golani & Fricke 2018:142 [ref. 36273], Arndt & Fricke 2019:31 [ref. 36883], Zajonz et al. 2019:92 [ref. 36871]). Current status: Valid as Enneapterygius abeli (Klausewitz 1960). Tripterygiidae: Tripterygiinae. Distribution: Red Sea, western Indian Ocean: East Africa, South Africa, Socotra, Seychelles, Comoros, Madagascar and Mauritius (Mascarenes) east to Chagos Archipelago. Habitat: marine.

acanthops, Vauclusella Whitley [G. P.] 1964:15 [Australian Naturalist v. 12 (pt 4); ref. 3721] Heron Island, Capricorn Group, Queensland, Australia. Holotype: AMS IB.4016. Paratypes: AMS IB.4023 (3, now 1), IB.4028 (5, now 7), IB 4076 (1). •Valid as Ceratobregma acanthops (Whitley 1964) -- (Holleman 1987:177 [ref. 6747], Fricke 1991:339 [ref. 19545], Fricke 1994:157 [ref. 21827], Fricke 1997:129 [ref. 23339], Williams & Fricke 2001:3534 [ref. 26288] dated 1965, Hoese 2006:1518 [ref. 28998], Fricke 2009:36 [ref. 30394], Patzner et al. 2009:444 [ref. 30396]). Current status: Valid as Ceratobregma acanthops (Whitley 1964). Tripterygiidae: Tripterygiinae. Distribution: Northeastern Australia. Habitat: marine.

albimacula, Helcogramma Williams [J. T.] & Howe [J. C.] 2003:154, Figs. 1-2 [aqua, Journal of Ichthyology and Aquatic Biology v. 7 (no. 4); ref. 27249] Apo Island, opposite west end of Chanos Pond, 9°04'30"N, 123°16'24"E, Philippines, depth 0-2.5 meters. Holotype: USNM 273934. Paratypes: AMS I.35024-001 (2), BPBM 36434 (2), CAS 81761 (2), USNM 315775 (42). Plus nontype specimens. •Valid as Helcogramma albimacula Williams & Howe 2003 -- (Fricke 2009:47 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Allen & Erdmann 2012:772 [ref. 31980], Fricke et al. 2014:146 [ref. 33932], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma albimacula Williams & Howe 2003. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Eastern Indonesia east to the Philippines, south to Papua New Guinea. Habitat: marine.

alkamr, Helcogramma Holleman [W.] 2007:56, Fig. 2; Pl. 1 [Smithiana, Publications in Aquatic Biodiversity, Bulletin No. 7; ref. 29195] Mayotte, French Territory Community, north coast of Isle Malandzamuzatsinsi, 12°40'19"S, 44°03'26"E, Comoro Islands, southwestern Indian Ocean, depth 1 meter. Holotype: ROM 73734. Paratypes: AMS 42910-001 (5); BMNH 2004.1.6.11-15 (5); ROM 73735 (16), 73736 (23); SAIAB 1915 (5), 70664 (2), 70665 (23); USNM 222359 (6), 222365 (20), 228973 (5), 228984 (7), 261389 (38), 375015 (5), 382859 (3), •Valid as Helcogramma alkamr Holleman 2007 -- (Fricke 2009:47 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Fricke et al. 2018:305 [ref. 35805], Arndt & Fricke 2019:32 [ref. 36883]). Current status: Valid as Helcogramma alkamr Holleman 2007. Tripterygiidae: Tripterygiinae. Distribution: Western Indian Ocean: East Africa, Persian Gulf and Madagascar east to western India. Habitat: marine.

alta, Apopterygion Kuiter [R. H.] 1986:90, Figs. 2-3 [Revue française d'Aquariologie Herpétologie v. 12 (no. 3) (1985); ref. 6026] Rye Pier, Port Phillip Bay, Victoria, Australia, depth 5 meters. Holotype: NMV A3345. Paratypes: AMS I.24817-001 [ex NMV A2215] (2); NMV A3346 (1), A3352-53 (5, 3). •Valid as Apopterygion alta Kuiter 1986 -- (Kuiter 1993:322 [ref. 23929], Fricke 1994:115 [ref. 21827], Gomon et al. 1994:732 [ref. 22532], Fricke 1997:561 [ref. 23339], Kuiter 1997:334 [ref. 25488], Hoese 2006:1518 [ref. 28998], Gomon 2008:685 [ref. 30616], Fricke 2009:32 [ref. 30394], Patzner et al. 2009:443 [ref. 30396]). Current status: Valid as Apopterygion alta Kuiter 1986. Tripterygiidae: Tripterygiinae. Distribution: Australia: Bass Strait, Victoria and Tasmania. Habitat: marine.

altipinnis, Enneapterygius Clark [E.] 1980:99, Figs. 3b, 6c, 11 [Israel Journal of Zoology v. 28 (nos 3-4) [for 1979]; ref. 8837] North of Ras Burka, Egypt, Gulf of Aqaba, Red Sea. Holotype: USNM 205807. Paratypes: HUJ 5481-82 (9, 10), 5501 (1); TAU [= SMNHTAU] 4076b (2), 4398 (5); USNM 205808-09 (33, 2), 212247-48 (2, 1). Type catalog: Springer & Orrell 1996:24 [ref. 22274]. •Synonym of Enneapterygius tutuilae Jordan & Seale 1906 -- (Fricke 1994:285 [ref. 21827], Fricke 1997:337, 572 [ref. 23339], Golani 2006:30 [ref. 28840], Hoese 2006:1523 [ref. 28998], Fricke 2009:42 [ref. 30394]). •Valid as Enneapterygius altipinnis Clark 1980 -- (Dor 1984:233 [ref. 29757], Randall et al. 1990:364 [ref. 15987], Goren & Dor 1994:61 [ref. 25356], Holleman & Bogorodsky 2012:41 [ref. 31735], DiBattista et al. 2015:20 [ref. 35706], Fricke et al. 2018:304 [ref. 35805], Bogorodsky & Randall 2018:[6] 244 [ref. 36492], Golani & Fricke 2018:142 [ref. 36273]). Current status: Valid as Enneapterygius altipinnis Clark 1980. Tripterygiidae: Tripterygiinae. Distribution: Red Sea, western Indian Ocean: East Africa to Seychelles, Comoros and Madagascar. Habitat: marine.

altivelis, Enneanectes Rosenblatt [R. H.] 1960:19, Fig. 4 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 112; ref. 11946] Western tip of Lyford Cay, New Providence Island, Bahamas, 25°02'06"N, 77°34'08"W, depth 25 feet. Holotype: ANSP 74446. Paratypes: ANSP 74440 (3, not 2), 74441-44 (2, 1, 21, 4), 74445 (10, not 9), 85924-26 (7, 1, 23). Type catalog: Böhlke 1984:164 [ref. 13621]. •Valid as Gillias altivelis (Rosenblatt 1960) -- (Fricke 1997:574 [ref. 23339]). •Valid as Enneanectes altivelis Rosenblatt 1960 -- (Greenfield & Johnson 1981:47 [ref. 5580], Robins & Ray 1986:221 [ref. 23100], Schmitter-Soto et al. 2000:165 [ref. 27754], Rocha & Rosa 2001:991 [ref. 25909], Smith & Williams 2002:9 [ref. 26427], Collette et al. 2003:118 [ref. 26784], Williams 2003:1749 [ref. 27109], Moura & Menezes in Menezes et al. 2003:94 [ref. 27192], Smith et al. 2003:47 [ref. 27621], Nelson et al. 2004:161 [ref. 27807], McEachran & Fechhelm 2005:574 [ref. 28815], Fricke 2009:36 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708], Victor 2013:69 [ref. 32783], Smith-Vaniz & Jelks 2014:65 [ref. 33341], Pinheiro et al. 2015:27 [ref. 34324], Victor 2017:68 [ref. 35549], Victor 2019:12 [ref. 36450]). Current status: Valid as Enneanectes altivelis Rosenblatt 1960. Tripterygiidae: Tripterygiinae. Distribution: Western Atlantic: Florida, Bahamas, Caribbean. Habitat: marine.

annulatum, Tripterygium Ramsay [E. P.] & Ogilby [J. D.] 1888:1021 [Proceedings of the Linnean Society of New South Wales (Series 2) v. 2 (pt 4); ref. 18601] Vancluse, Port Jackson, New South Wales, Australia. Syntypes: AMS I.1651 (2). Original genus should have been Tripterygion. •Valid as Enneapterygius annulatus (Ramsay & Ogilby 1888) -- (Randall et al. 1990:364 [ref. 15987], Kuiter 1993:324 [ref. 23929]). •Synonym of Enneapterygius atrogulare (Günther 1873) -- (Fricke 1994:187 [ref. 21827], Fricke 1997:165, 566 [ref. 23339], Johnson 1999:747 [ref. 25471], Randall 2005:471 [ref. 28239], Hoese 2006:1519 [ref. 28998], Fricke 2009:38 [ref. 30394]). Current status: Synonym of Enneapterygius atrogulare (Günther 1873). Tripterygiidae: Tripterygiinae. Habitat: marine.

antarcticum, Tripterygium Tomo [A. P.] 1982:88, Figs. 66-70 [Publicacion / Dirección Nacional del Antártico, Instituto Antartico Argentino. No. 14; ref. 20550] Paradise Bay, 64°S, 62°W, Antarctic Peninsula. Holotype: IAA 4. Paratypes: IAA (4). Original genus should have been Tripterygion. •Valid as Helcogrammoides antarcticus (Tomo 1982) -- (Gon 1990:400 [ref. 21680], Fricke 1997:581 [ref. 23339], Pequeño 1997:84 [ref. 23536], Williams & Springer 2001:118 [ref. 26032], Fricke 2009:47 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Cornejo et al. 2018:17 [ref. 36409]). Current status: Valid as Helcogrammoides antarcticus (Tomo 1982). Tripterygiidae: Tripterygiinae. Distribution: Southern Ocean: western Antarctica. Habitat: marine.

aquila, Helcogramma Williams [J. T.] & McCormick [C. J.] 1990:1021, Fig. 3 [Copeia 1990 (no. 4); ref. 18992] White Beach, past Mahatae, Batan Island, Batanes, Philippines, 20°24'45"N, 121°55'02"E, depth 0-6 meters. Holotype: USNM 298405. Paratypes: AMS I.29427-001 (1); ANSP 164964 (1); BPBM 32855 (1); USNM 293945 (1), 297370 (4, 1 c&s), 298400 (4). Type catalog: Springer & Orrell 1996:24 [ref. 22274]. Species name should not be spelled aquilum -- see account of Helcogramma. •Valid as Helcogramma aquilum [sometimes as aquila] Williams & McCormick 1990 -- (Fricke 1997:409, 577 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Williams & Howe 2003:158 [ref. 27249], Myers & Donaldson 2003:638 [ref. 27495], Holleman 2006:102 [ref. 28864], Fricke 2009:48 [ref. 30394], Patzner et al. 2009:446 [ref. 30396] as aquilum, Allen & Erdmann 2012:772 [ref. 31980], Tashiro & Motomura 2014:103 [ref. 35481], Fricke & Erdmann 2017:41 [ref. 35190]). Current status: Valid as Helcogramma aquila Williams & McCormick 1990. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Ryukyu Islands, Philippines and Guam. Habitat: marine.

ascensionis, Helcogramma Lubbock [R.] 1980:293, Fig. 2 [Journal of Fish Biology v. 17 (no. 3); ref. 8694] Rockpools, Catherine Point, Ascension Island, depth to 2 meters. Holotype: BMNH 1979.1.11.1. Paratypes: BMNH 1979.1.11.2-11 (10), 1979.1.11.12 (1); USNM 219405 (1). Type catalog: Springer & Orrell 1996:24 [ref. 22274]. •Synonym of Helcogramma obtusirostris (Klunzinger 1871) -- (Hansen 1986:341 [ref. 5810]). •Synonym of Helcogramma obtusirostre (Klunzinger 1871) -- (Fricke 1997:579 [ref. 23339]). •Valid as Helcogramma ascensionis Lubbock 1980 -- (Edwards & Glass 1987:657 [ref. 9166], Wirtz 1990:916 [ref. 19374], Fricke 2009:48 [ref. 30394], Wirtz et al. 2014:9 [ref. 33516], Williams in Carpenter & De Angelis 2016:2795 [ref. 34618], Brown et al. 2019:164 [ref. 36444]). Current status: Valid as Helcogramma ascensionis Lubbock 1980. Tripterygiidae: Tripterygiinae. Distribution: Southern central Atlantic: Ascension and Saint Helena islands. Habitat: marine.

atauroensis, Helcogramma Fricke [R.] & Erdmann [M. V.] 2017:36, Figs. 1-3 [Journal of the Ocean Science Foundation v. 26; ref. 35190] Timor-Leste, Ataúro Island, Fatuu, eastern Indian Ocean, 8°08.442'S, 125°36.737'E, depth 3 meters. Holotype: WAM P.34562-003. Paratypes: HUJ, WAM. •Valid as Helcogramma atauroensis Fricke & Erdmann 2017. Current status: Valid as Helcogramma atauroensis Fricke & Erdmann 2017. Tripterygiidae: Tripterygiinae. Distribution: Timor-Leste, eastern Indian Ocean. Habitat: marine.

atlanticus, Tripterygion Wheeler [A. C.] & Dunne [J.] 1975:641, Pl. 1; Fig. 2 [Journal of Fish Biology v. 7 (no. 5); ref. 7482] English Channel off Roschoff, Finisterre Province, France, northeastern Atlantic. Holotype: BMNH 1974.3.19.10. Paratypes: BMNH 1974.3.19.11-14 (1, 1, 1, 1). •Synonym of Tripterygion delaisi Cadenat & Blache 1970 -- (Wirtz 1990:916 [ref. 19374], Fricke 1997:587 [ref. 23339], Fricke 2009:43 [ref. 30394]). Current status: Synonym of Tripterygion delaisi Cadenat & Blache 1970. Tripterygiidae: Tripterygiinae. Habitat: marine.

atriceps, Tripterygium Jenkins [O. P.] 1903:505, Fig. 46 [Bulletin of the U. S. Fish Commission v. 22 (1902); ref. 2341] Honolulu, Oahu Island, Hawaiian Islands. Holotype: USNM 50719. Paratypes: CAS-SU 23305 (9). Non-types: USNM 51165 (3), 126693 (3). Type catalog: Böhlke 1953:99 [ref. 12291], Springer & Orrell 1996:24-25 [ref. 22274]. Original genus should have been Tripterygion. •Valid as Enneapterygius atriceps (Jenkins 1903) -- (Fricke 1997:161, 566 [ref. 23339], Williams & Fricke 2001:3534 [ref. 26288], Mundy 2005:460 [ref. 28379], Fricke 2009:38 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Randall 2007:373 [ref. 30952], Meguro & Motomura 2010:6 [ref. 31252], Motomura et al. 2015:10 [ref. 33825], Fricke & Erdmann 2017:28 [ref. 35129]). Current status: Valid as Enneapterygius atriceps (Jenkins 1903). Tripterygiidae: Tripterygiinae. Distribution: Central Pacific: Hawaiian Ridge and Hawaiian Islands. Habitat: marine.

atrogulare, Tripterygium Günther [A.] 1873:91 [11] [Journal des Museum Godeffroy v. 1 (no. 2); ref. 6424] Bowen, Queensland, Australia. Holotype (unique): BMNH 1873.8.1.47. Original genus should have been Tripterygion. •Valid as Enneapterygius atrogulare (Günther 1873) -- (Fricke 1994:187 [ref. 21827], Fricke 1997:165, 566 [ref. 23339], Randall et al. 1997:364 [ref. 25919], Johnson 1999:747 [ref. 25471], Williams & Fricke 2001:3534 [ref. 26288], Randall et al. 2004:24 [ref. 27624], Randall 2005:471 [ref. 28239], Hoese 2006:1519 [ref. 28998], Fricke 2009:38 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Fricke & Erdmann 2017:29 [ref. 35129]). Current status: Valid as Enneapterygius atrogulare (Günther 1873). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: eastern Australia, Coral Sea and Tonga. Habitat: marine.

atrorus, Enneanectes Rosenblatt [R. H.] 1960:9, Fig. 2 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 112; ref. 11946] Green Cay, Bahamas, 25°07'08"N, 77°11'17"W, depth 48 feet. Holotype (unique): ANSP 85931. Type catalog: Böhlke 1984:164 [ref. 13621]. •Valid as Gillias atrorus (Rosenblatt 1960) -- (Fricke 1997:574 [ref. 23339]). •Valid as Enneanectes atrorus Rosenblatt 1960 -- (Greenfield & Johnson 1981:49 [ref. 5580], Smith & Williams 2002:9 [ref. 26427], Williams 2003:1749 [ref. 27109], Smith et al. 2003:47 [ref. 27621], Nelson et al. 2004:161 [ref. 27807], Fricke 2009:37 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708], Victor 2013:69 [ref. 32783], Victor 2017:68 [ref. 35549], Victor 2019:12 [ref. 36450]). Current status: Valid as Enneanectes atrorus Rosenblatt 1960. Tripterygiidae: Tripterygiinae. Distribution: Western Atlantic: Bahamas, Caribbean. Habitat: marine.

bahasa, Enneapterygius Fricke [R.] 1997:170, Fig. 29 [ ref. 23339] Heron Island, Queensland, Australia, 23°26'S, 151°55'E. Holotype: USNM 259168. Paratypes: SMNS 15934 (3), 15955 (2), 18695 (2); USNM 259129 (31), 209582 (53), 344075 (3), 344087 (1); plus other non-type material. Type catalog: Fricke 2005:64 [ref. 29864]. •Valid as Enneapterygius bahasa Fricke 1997 -- (Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Allen & Adrim 2003:53 [ref. 26830], Mundy 2005:460 [ref. 28379], Hoese 2006:1519 [ref. 28998], Chiang & Chen 2008:190 [ref. 29932], Fricke 2009:38 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Meguro & Motomura 2010:6 [ref. 31252], Motomura et al. 2010:182 [ref. 31256], Allen & Erdmann 2012:765 [ref. 31980], Fricke & Erdmann 2017:23, 25 [ref. 35129]). Current status: Valid as Enneapterygius bahasa Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: southern Japan, Indonesia, Philippines, Papua New Guinea, to Palau and Guam and eastern Australia. Habitat: marine.

bapturum, Tripterygion Jordan [D. S.] & Snyder [J. O.] 1902:447, Fig. 2 [Proceedings of the United States National Museum v. 25 (no. 1293); ref. 2516] Misaki, Japan. Holotype (unique): CAS-SU 7066. Type catalog: Böhlke 1953:99 [ref. 12291]. Genus misspelled Triptergion in original. •Valid as Tripterygion bapturum Jordan & Snyder 1902 -- (Lindberg & Krasyukova 1975:24 [ref. 7348], Yoshino in Masuda et al. 1984:294 [ref. 6441]). •Valid as Enneapterygius bapturum (Jordan & Snyder 1902) -- (Fricke 1987:335 [ref. 9108], Shimizu & Hatooka 1997:5 [ref. 24377]). •Valid as Gracilopterygion bapturum (Jordan & Snyder 1902) -- (Fricke 1994:366 [ref. 21827]). •Valid as Springerichthys bapturus (Jordan & Snyder 1902) -- (Shen 1994:27 [ref. 21605], Fricke 1997:521, 585 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1081 [ref. 25182], Shinohara et al. 2000:181 [ref. 25987], Nakabo 2002:1081 [ref. 26193], Kim et al. 2005:148 [ref. 28305], Fricke 2009:43 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Parin et al. 2014:467 [ref. 33547], Motomura et al. 2015:28 [ref. 34709], Yoshigou 2019:195 [ref. 36714]). Current status: Valid as Springerichthys bapturus (Jordan & Snyder 1902). Tripterygiidae: Tripterygiinae. Distribution: Northwestern Pacific: Sea of Okhotsk, Japan, Taiwan and Korea. Habitat: marine.

bathytaton, Forsterygion Hardy [G. S.] 1989:500, Figs. 1B, 4 [Journal of Natural History v. 23 (no. 3); ref. 12485] Mernoo Bank, 43°06.1'S, 175°20.5'E, depth 153 meters. Holotype: NMNZ P.22058. Paratypes: NMNZ P.5041 (7, 1 c&s), P.7095 (8), P.7096 (3), P.10642 (1), P.10645 (1), P.21407 (2), P.21408 (12), P.21409 (16), P.21574 (5). Type catalog: Hardy 1990:12 [ref. 18942]. •Valid as Forsterygion bathytaton Hardy 1989 -- (Hardy in Amaoka et al. 1990:303 [ref. 16024], Fricke 1994:311 [ref. 21827], Fricke 1997:573 [ref. 23339]). •Valid as Matanui bathytaton (Hardy 1989) -- (Jawad & Clements 2004:97 [ref. 27725] Jawad 2008:2944, 2952 [ref. 29958], Fricke 2009:46 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Stewart & Clements 2015:1514 [ref. 34299]). Current status: Valid as Matanui bathytaton (Hardy 1989). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

bichrous, Enneapterygius Fricke [R.] 1994:195, Fig. 30 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] New Year Island, 10°55'S, 133°01'E, Northern Territory, Australia, depth 0-1 meter. Holotype: NTM S.10600-032. Paratypes: AMS I.21963-001 (1); NTM A.10011-018 (3); ROM 65513 (1), 65515-16 (1, 6), 67974 (2); SMNS 15356 (1); WAM P.29025-039 (13). Plus many more paratype lots at AMS, NTM and WAM; additional non-type material at several museums. Type catalog: Fricke 1995:19 [ref. 28100], Fricke 2005:64 [ref. 29864], Moore et al. 2008:41 [ref. 30117] with remarks. •Synonym of Enneapterygius flavoccipitis Shen 1994 -- (Fricke 1997:199, 567 [ref. 23339], Fricke & Kulbicki 2006:345 [ref. 28867], Hoese 2006:1520 [ref. 28998], Chiang & Chen 2008:195 [ref. 29932], Fricke 2009:39 [ref. 30394], Larson et al. 2013:177 [ref. 32988]). Current status: Synonym of Enneapterygius flavoccipitis Shen 1994. Tripterygiidae: Tripterygiinae. Habitat: marine.

billi, Helcogramma Hansen [P. E. H.] 1986:329, Fig. 10 [Bulletin of Marine Science v. 38 (no. 2); ref. 5810] Rocky area opposite Fort Frederick, Trincomalee (outside harbor), Sri Lanka. Holotype: USNM 222377. Paratypes: USNM 222367 (6, 1 c&s), 222368 (1), 222374 (3), 222380 (17, not 27). Type catalog: Springer & Orrell 1996:25 [ref. 22274]. •Valid as Helcogramma billi Hansen 1986 -- (Fricke 1997:577 [ref. 23339], Manilo & Bogorodsky 2003:S116 [ref. 27377], Holleman 2007:58 [ref. 29195], Fricke 2009:48 [ref. 30394], Patzner et al. 2009:446 [ref. 30396]). Current status: Valid as Helcogramma billi Hansen 1986. Tripterygiidae: Tripterygiinae. Distribution: Northern Indian Ocean: Sri Lanka to southern India. Habitat: marine.

bimaculata, Lepidonectes Allen [G. R.] & Robertson [D. R.] 1992:55, Figs. 4-5 [Revue française d'Aquariologie Herpétologie v. 19 (nos 1-2); ref. 19713] Malpelo Island, eastern Pacific west of Colombia, about 4°00'N, 81°31'W, depth 15 meters. Holotype: ICNMNH 1788. Paratypes: ICNMNH 1789 (1), USNM 321178 (1). Type catalog: Springer & Orrell 1996:25 [ref. 22274], Mojica & Agudelo-Zamora 2018:52 [ref. 36127]. Correct spelling of species is bimaculatus. •Valid as Lepidonectes bimaculata Allen & Robertson 1992 -- (Allen & Robertson 1994:222 [ref. 22193], Fricke 1997:582 [ref. 23339], Patzner et al. 2009:448 [ref. 30396]). •Valid as Lepidonectes bimaculatus Allen & Robertson 1992 -- (Fricke 2009:34 [ref. 30394]). Current status: Valid as Lepidonectes bimaculatus Allen & Robertson 1992. Tripterygiidae: Tripterygiinae. Distribution: Pacific coast of Colombia. Habitat: marine.

boehlkei, Enneanectes Rosenblatt [R. H.] 1960:5, Fig. 1 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 112; ref. 11946] Small lagoon off Salt Cay, Treasure Island, Bahamas, 25°06'16"N, 77°16'22"W. Holotype: ANSP 74447. Paratypes: ANSP 74448 (2), 85927 (4) [not 3], 85928 (5, not 6), 85929-30 (5, 1). Additional original material: ANSP 74449-50 (2, 1), FMNH 61803 (1), USNM 4092 (1), 116806 (3), 117439 (2). Type catalog: Böhlke 1984:164 [ref. 13621]. •Valid as Gillias boehlkei (Rosenblatt 1960) -- (Fricke 1997:575 [ref. 23339]). •Valid as Enneanectes boehlkei Rosenblatt 1960 -- (Greenfield & Johnson 1981:49 [ref. 5580], Robins & Ray 1986:221 [ref. 23100], Cervigón 1994:67 [ref. 24488], Schmitter-Soto et al. 2000:165 [ref. 27754], Smith & Williams 2002:9 [ref. 26427], Collette et al. 2003:118 [ref. 26784], Williams 2003:1749 [ref. 27109], Smith et al. 2003:47 [ref. 27621], Nelson et al. 2004:161 [ref. 27807], McEachran & Fechhelm 2005:575 [ref. 28815], Fricke 2009:37 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708], Victor 2013:63 [ref. 32783], Smith-Vaniz & Jelks 2014:66 [ref. 33341], Robertson et al. 2016:147, 151 [ref. 34961], Victor 2017:69 [ref. 35549], Victor 2019:13 [ref. 36450]). Current status: Valid as Enneanectes boehlkei Rosenblatt 1960. Tripterygiidae: Tripterygiinae. Distribution: Western Atlantic: Florida, Bahamas, North & Western Caribbean, Antilles south only to St. Kitts. Habitat: marine.

brachylepis, Tripterygion Schultz [L. P.] in Schultz et al. 1960:291, Fig. 113 [Bulletin of the United States National Museum No. 202, v. 2; ref. 3972] Coral heads in eastern end of Lagoon, Bikini Atoll, Marshall Islands, western Pacific, depth 20-25 feet. Holotype (unique): USNM 142253. Type catalog: Springer & Orrell 1996:25 [ref. 22274]. •Valid as Norfolkia brachylepis (Schultz 1960) -- (Fricke 1991:340 [ref. 19545], Holleman 1991:233 [ref. 21794], Fricke 1994:465 [ref. 21827], Shen 1994:23 [ref. 21605], Goren & Dor 1994:61 [ref. 25356], Randall 1995:34 [ref. 21853], Randall 1995:310 [ref. 22896], Fricke 1997:504, 583 [ref. 23339], Allen 1997:212 [ref. 23977], Randall et al. 1997:522 [ref. 25919], Myers 1999:217 [ref. 23965], Randall 1999:24 [ref. 25829], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1077 [ref. 25182], Hutchins 2001:41 [ref. 25847], Fricke 2001:3 [ref. 25985], Williams & Fricke 2001:3535 [ref. 26288], Nakabo 2002:1077 [ref. 26193], Fricke 2002:194 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830], Manilo & Bogorodsky 2003:S116 [ref. 27377], Myers & Donaldson 2003:639 [ref. 27495], Randall et al. 2004:24 [ref. 27624], Heemstra & Heemstra 2004:387 [ref. 28072], Randall 2005:478 [ref. 28239], Hoese 2006:1526 [ref. 28998], Gomon 2008:690 [ref. 30616], Fricke 2009:34 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Motomura et al. 2010:189 [ref. 31256], Holleman & Bogorodsky 2012:58 [ref. 31735], Allen & Erdmann 2012:781 [ref. 31980], Fricke et al. 2014:147 [ref. 33932], Motomura et al. 2015:28 [ref. 34709], Fricke et al. 2018:306 [ref. 35805], Golani & Fricke 2018:144 [ref. 36273]). Current status: Valid as Norfolkia brachylepis (Schultz 1960). Tripterygiidae: Tripterygiinae. Distribution: Red Sea, Indo-West Pacific: East and South Africa, Comoros and Madagascar east to Marshall Islands, Samoa and Tonga, north to southern Japan and Ogasawara Islands, south to New Caledonia. Habitat: marine.

bucephalus, Trianectes McCulloch [A. R.] & Waite [E. R.] 1918:53, Pl. 3 (fig. 3) [Records of the South Australian Museum (Adelaide) v. 1 (no. 1); ref. 2950] Spencer Gulf, South Australia. Holotype (unique): SAMA F599. Type catalog: Glover 1976:174 [ref. 19449]. •Valid as Trianectes bucephalus McCulloch & Waite 1918 -- (Kuiter 1993:323 [ref. 23929], Gomon et al. 1994:740 [ref. 22532], Fricke 1994:533 [ref. 21827], Fricke 1997:586 [ref. 23339], Hutchins 2001:42 [ref. 25847], Hoese 2006:1527 [ref. 28998], Gomon 2008:691 [ref. 30616], Fricke 2009:33 [ref. 30394], Patzner et al. 2009:448 [ref. 30396]). Current status: Valid as Trianectes bucephalus McCulloch & Waite 1918. Tripterygiidae: Tripterygiinae. Distribution: Australia: Victoria, Tasmania, South Australia and Western Australia. Habitat: marine.

bucknilli, Tripterygion Griffin [L. T.] 1926:544, Pl. 97 [Transactions of the New Zealand Institute v. 56; ref. 15589] Deep crevices, Mount Maunganui, Bay of Plenty, New Zealand. Holotype: AIM MA 3359. Paratypes: (8) AIM (1), AMS IA.730 (2). Trnski personal communication (12/1/2017) on type status. •Synonym of Notoclinops segmentatus (McCulloch & Phillipps 1923) -- (Hardy 1987:166 [ref. 6027], Fricke 1994:487 [ref. 21827], Fricke 1997:584 [ref. 23339], Fricke 2009:46 [ref. 30394]). Current status: Synonym of Notoclinops segmentatus (McCulloch & Phillipps 1923). Tripterygiidae: Tripterygiinae. Habitat: marine.

caerulepunctus, Notoclinops Hardy [G. S.] 1989:328, Pl. (figs. 1-2) [Journal of the Royal Society of New Zealand v. 19 (no. 3); ref. 17673] Doubtful Sound, Fiordland, 45°18'S, 167°00'E, New Zealand, depth 12 meters. Holotype: NMNZ P.21258. Paratypes: NMNZ (88, 19 lots). Type catalog: Hardy 1990:13 [ref. 18942]. •Valid as Notoclinops caerulepunctus Hardy 1989 -- (Paulin & Roberts 1992:71 [ref. 24625], Fricke 1994:482 [ref. 21827], Fricke 1997:583 [ref. 23339], Fricke 2009:46 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1516 [ref. 34299]). Current status: Valid as Notoclinops caerulepunctus Hardy 1989. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

callionymi, Tripterygium Weber [M.] 1909:147 [Notes from the Leyden Museum v. 31 (note 4); ref. 4600] Sanana, Sulawesi, Indonesia, Siboga station 193. Lectotype: ZMA 112506. Paralectotypes: ZMA 112501-05 (2, 1, 2, 1, 1). Type catalog: Nijssen et al. 1982:95 [ref. 19236]. Lectotype selected by Fricke 1997:173, 280 [ref. 23339]. Original genus should have been Tripterygion. •Synonym of Enneapterygius minutus (Günther 1877) -- (Fricke 1994:243 [ref. 21827], Fricke 1994:4 [ref. 21828]). •Synonym of Enneapterygius philippinus (Peters 1868) -- (Fricke 1997:274, 570 [ref. 23339] after lectotype designation, Fricke 2009:41 [ref. 30394]). Current status: Synonym of Enneapterygius philippinus (Peters 1868). Tripterygiidae: Tripterygiinae. Habitat: marine.

calva, Vauclusella Whitley [G. P.] 1944:272 [Australian Zoologist v. 10 (pt 3); ref. 4705] Cottesloe, 31°59'S, 115°45'E, Western Australia. Holotype: WAM P.1052-001. Paratypes: WAM P.1052-002 (1). Type catalog: Hutchins & Smith 1991:37 [ref. 19335], Moore et al. 2008:42 [ref. 30117]. •Synonym of Helcogramma decurrens McCulloch & Waite 1918 -- (Fricke 1994:418 [ref. 21827], Gomon et al. 1994:734 [ref. 22532], Fricke 1997:578 [ref. 23339], Hoese 2006:1524 [ref. 28998], Fricke 2009:48 [ref. 30394]). Current status: Synonym of Helcogramma decurrens McCulloch & Waite 1918. Tripterygiidae: Tripterygiinae. Habitat: marine.

capense, Tripterygium Gilchrist [J. D. F.] & Thompson [W. W.] 1908:140 [Annals of the South African Museum v. 6 (pt 2, no. 2); ref. 5840] False Bay, about 2.5 miles from Seal Island, depth 14 fathoms; Fish Hoek Bay, depth 5 fathoms, South Africa. Syntypes: (4) SAM 9900-01 (2, 2). Original genus should have been Tripterygion. •Valid as Cremnochorites capensis (Gilchrist & Thompson 1908) -- (Holleman 1986:756 [ref. 5671], Fricke 1997:563 [ref. 23339], Jawad & Clements 2004:84 [ref. 27725], Heemstra & Heemstra 2004:386 [ref. 28072], Fricke 2009:33 [ref. 30394], Patzner et al. 2009:444 [ref. 30396]). Current status: Valid as Cremnochorites capensis (Gilchrist & Thompson 1908). Tripterygiidae: Tripterygiinae. Distribution: Southeastern Atlantic to southwestern Indian Ocean: southern Africa. Habitat: marine.

capidata, Helcogramma Rosenblatt [R. H.] in Schultz et al. 1960:297, Pl. 121 (fig. c) [Bulletin of the United States National Museum No. 202, v. 2; ref. 3972] Presipai region of Saratokmatel reef flat in Teawaitua ship passage, Kapingamarangi Atoll, Caroline Islands, western Pacific. Holotype: CAS-SU 52010. Paratypes: CAS-SU 52011-14 (1, 4, 21, 7); USNM 142249-50 (1, 2), 175255 (2). Additional material: GVF uncat. (3 lots, 8). Type catalog: Springer & Orrell 1996:25 [ref. 22274]. Gender of genus feminine; therefore correct species name capidata. •Valid as Helcogramma capidatum Rosenblatt 1960 -- (Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Myers & Donaldson 2003:638 [ref. 27495], Randall et al. 2004:24 [ref. 27624], Randall 2005:476 [ref. 28239], Allen & Erdmann 2012:772 [ref. 31980]). •Valid as Helcogramma capidata Rosenblatt 1960 -- (Hansen 1986:330 [ref. 5810], Fricke 1994:410 [ref. 21827], Fricke 1997:413, 577 [ref. 23339], Myers 1999:216 [ref. 23965], Fricke 2009:48 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Fricke & Erdmann 2017:43 [ref. 35190]). Current status: Valid as Helcogramma capidata Rosenblatt 1960. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Malaysia to Micronesia and east to Tonga. Habitat: marine.

capito, Tripterygion Jenyns [L.] 1842:94, Pl. 19 (fig. 1) [The zoology of the voyage of H. M. S. Beagle; ref. 2344] Bay of Islands, New Zealand. Syntypes: BMNH 1917.7.14.56-57 (2). See Paulin et al. 1989:264 [ref. 24556]. •Status uncertain -- (Hardy 1989:509 [ref. 12485]). •Valid as Grahamina capito (Jenyns 1842) -- (Fricke 1994:378 [ref. 21827], Fricke 1997:576 [ref. 23339], Clements et al. 2000:373 [ref. 25188], Jawad & Clements 2004:84 [ref. 27725], Fricke 2009:45 [ref. 30394], Patzner et al. 2009:446 [ref. 30396]). •Valid as Forsterygion capito (Jenyns 1842) -- (Jawad 2008:2962 [ref. 29958] dated 1841, Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1503 [ref. 34299]). Current status: Valid as Forsterygion capito (Jenyns 1842). Tripterygiidae: Tripterygiinae. Distribution: New Zealand region. Habitat: marine.

carminale, Tripterygium Jordan [D. S.] & Gilbert [C. H.] 1882:362 [Proceedings of the United States National Museum v. 4 (no. 237); ref. 2470] Mazatlán, Mexico. [Original locality: Deep tide-pool at Mazatlán, Sinaloa, western Mexico.] Syntypes: USNM 28118 (or 120946) (4). Type catalog: Böhlke 1953:99 [ref. 12291], Springer & Orrell 1996:25-26 [ref. 22274]. Original genus should have been Tripterygion. Broch (1940) named a neotype (SU 3854). Syntypes later found (L. P.Schutz) •Valid as Axoclinus carminalis (Jordan & Gilbert 1882) -- (Allen & Robertson 1994:219 [ref. 22193], Thomson et al. 2000:206 [ref. 25640], Nelson et al. 2004:161 [ref. 27807]). •Valid as Enneanectes carminalis (Jordan & Gilbert 1882) -- (De La Cruz Agüero et al. 1997:301 [ref. 24545], Fricke 1997:564 [ref. 23339], Smith & Williams 2002:8 [ref. 26427], Fricke 2009:37 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708], Galván-Villa et al. 2016:148 [ref. 34871], González-Murcia et al. 2019:295 [ref. 36544]). Current status: Valid as Enneanectes carminalis (Jordan & Gilbert 1882). Tripterygiidae: Tripterygiinae. Distribution: Eastern Pacific. Habitat: marine.

cerasina, Helcogramma Williams [J. T.] & Howe [J. C.] 2003:159, Figs. 5-6 [aqua, Journal of Ichthyology and Aquatic Biology v. 7 (no. 4); ref. 27249] Eua Island, fringing reef at base of cliff on northwestern shore, Tonga Islands, 21°18'15"S, 174°26'20"W, depth 0-10.5 meters. Holotype: USNM 329714. Paratypes: USNM 262029 (1), 287093 (1), 329713 (117). •Valid as Helcogramma cerasina Williams & Howe 2003 -- (Fricke 2009:48 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma cerasina Williams & Howe 2003. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: Tonga and Fiji Islands. Habitat: marine.

cerasinus, Enneapterygius Jordan [D. S.] & Seale [A.] 1906:419, Fig. 100 [Bulletin of the Bureau of Fisheries v. 25 (for 1905); ref. 2497] Apia, Upolu Island, Samoa. Holotype (unique): USNM 51802. Type catalog: Springer & Orrell 1996:26 [ref. 22274]. •Questionably a synonym of Enneapterygius minutus (Günther 1877) -- (Fricke 1994:242 [ref. 21827], Fricke 1994:4 [ref. 21828]). •Synonym of Enneapterygius philippinus (Peters 1868) -- (Fricke 1997:274, 570 [ref. 23339], Fricke 2009:41 [ref. 30394]). Current status: Synonym of Enneapterygius philippinus (Peters 1868). Tripterygiidae: Tripterygiinae. Habitat: marine.

cheni, Enneapterygius Wang [S.-C.], Shao [K.-T.] & Shen [S.-C.] 1996:80, Fig. 1 [Acta Zoologica Taiwanica v. 7 (no. 1); ref. 22200] Cheng-Kung, eastern coast of Taiwan, 25°07'N, 121°21'E, depth 8-10 meters. Holotype: ASIZP 057448. Paratypes: ASIZT 057449-50 (1, 2); NTUM 7870 (2). Type catalog: Ho & Shao 2011:52 [ref. 31432]. •Synonym of Enneapterygius niger Fricke 1994 -- (Fricke 1997:248, 569 [ref. 23339]). •Valid as Enneapterygius cheni Wang, Shao & Shen 1996 -- (Shibukawa et al. 2004:2 [ref. 27552], Chiang & Chen 2008:190 [ref. 29932], Fricke 2009:38 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Ho & Shao 2011:52 [ref. 31432], Fricke & Erdmann 2017:20, 26, 29 [ref. 35129]). Current status: Valid as Enneapterygius cheni Wang, Shao & Shen 1996. Tripterygiidae: Tripterygiinae. Distribution: Tropical Western North Pacific: Micronesia to Taiwan. Habitat: marine.

chica, Helcogramma Rosenblatt [R. H.] in Schultz et al. 1960:294, Fig. 144 [Bulletin of the United States National Museum No. 202, v. 2; ref. 3972] Ocean Reef, Hull Island, Phoenix Islands. Holotype: USNM 115516. Paratypes: CAS-SU 52015-16 (8, 10); USNM 115514-15 (19, 5), 142252 (1), 175256 (21). Additional material: USNM 115513 (1), 115517 (1). Type catalog: Springer & Orrell 1996:26 [ref. 22274]. •Valid as Helcogramma chica Rosenblatt 1960 -- (Hansen 1986:331 [ref. 5810], Fricke 1994:414 [ref. 21827], Fricke 1997:422, 577 [ref. 23339], Myers 1999:216 [ref. 23965], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Myers & Donaldson 2003:639 [ref. 27495], Lobel & Lobel 2004:75 [ref. 27576], Randall et al. 2004:24 [ref. 27624], Randall 2005:476 [ref. 28239], Fricke 2009:48 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:773 [ref. 31980], Fricke & Erdmann 2017:43 [ref. 35190]). Current status: Valid as Helcogramma chica Rosenblatt 1960. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: Andaman Sea, Cocos-Keeling and Christmas islands, Papua New Guinea east to Marshall Islands, Wake Island, and north to Guam and Ogasawara Islands. Habitat: marine.

chilensis, Tripterygion Cancino [C.] in de Buen 1960:70 [Revista de Biología Marina, Valparaiso. v. 10 (nos 1-3); ref. 23226] Maitencillo, Chile. Lectotype: MNHNC P-6879. Paralectotypes: MNHNC P-5559 (11), P-6888 (2). Lectotype selected by Williams & Springer 2001:119 [ref. 26032]. •Valid as Tripterygion chilensis Cancino 1960 -- (Pequeño 1989:66 [ref. 14125]). •Valid as Helcogrammoides chilensis (Cancino 1960) -- (Castillo & Pequeño 1998:115 [ref. 23987], Williams & Springer 2001:119 [ref. 26032], Jawad & Clements 2004:85 [ref. 27725], Fricke 2009:47 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Cornejo & Ibáñez 2018:275 [ref. 36410], Cornejo et al. 2018:17 [ref. 36409]). Current status: Valid as Helcogrammoides chilensis (Cancino 1960). Tripterygiidae: Tripterygiinae. Distribution: Southeastern Pacific: Peru south to Chile. Habitat: marine.

clarkae, Enneapterygius Holleman [W.] 1982:121, Fig. 6 [Annals of the Cape Provincial Museums (Natural History) v. 14 (pt 4); ref. 2187] Off Barreira Vermelha, Inhaca Island, Mozambique, southwestern Indian Ocean. Holotype: SAIAB [formerly RUSI] 14175. Paratypes: LACM 31617-27 (1); SAIAB [formerly RUSI] 7938-39 (1, 2), 9842 (2), 14174 (4), 14176 (1), 14177 (2 c&s) not on shelf, 14629 (4); USNM 231378-82 (1, 1, 1, 1, 3). Type catalog: Springer & Orrell 1996:26 [ref. 22274]. •Synonym of Enneapterygius destai Clark 1980 -- (Fricke 1997:566 [ref. 23339]). •Valid as Enneapterygius clarkae Holleman 1982 -- (Holleman 1986:756 [ref. 5671], Goren & Dor 1994:61 [ref. 25356], Holleman 2005:6 [ref. 28455], Fricke 2009:38 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Holleman & Bogorodsky 2012:46 [ref. 31735], Fricke & Erdmann 2017:20, 29 [ref. 35129], Fricke et al. 2018:304 [ref. 35805], Golani & Fricke 2018:142 [ref. 36273]). Current status: Valid as Enneapterygius clarkae Holleman 1982. Tripterygiidae: Tripterygiinae. Distribution: Red Sea, western Indian Ocean: East and South Africa east to Madagascar and Cargados Carajos. Habitat: marine.

clarkei, Triptergium Morton [A.] 1888:78 [Papers and Proceedings of the Royal Society of Tasmania 1887; ref. 3046] Clarke Island, Bass Strait, Australia. Holotype (unique): whereabouts unknown. Genus should have been spelled Tripterygion. •Valid as Norfolkia clarkei (Morton 1888) -- (Fricke 1991:339 [ref. 19545], Kuiter 1993:323 [ref. 23929], Gomon et al. 1994:737 [ref. 22532], Kuiter 1997:334 [ref. 25488]). •Valid as Trinorfolkia clarkei (Morton 1888) -- (Fricke 1994:541 [ref. 21827], Fricke 1997:586 [ref. 23339], Hutchins 2001:42 [ref. 25847], Hoese 2006:1527 [ref. 28998], Gomon 2008:692 [ref. 30616], Fricke 2009:35 [ref. 30394], Patzner et al. 2009:448 [ref. 30396]). Current status: Valid as Trinorfolkia clarkei (Morton 1888). Tripterygiidae: Tripterygiinae. Distribution: Australia: New South Wales, Victoria, Tasmania, South Australia and Western Australia. Habitat: marine.

clarkhubbsi, Lepidonectes Bussing [W. A.] 1991:81, Fig. 2 [Revista de Biología Tropical v. 39 (no. 1); ref. 19493] Off Isla San José, Islas Murciélagos, Costa Rica, depth 10-14 meters. Holotype: LACM 44828-2 [ex UCR 382-5]. Paratypes: LACM (18 lots, 247), UCR (11 lots, 130 or 8 lots, 98). Type catalog: Angulo et al. 2015:377 [ref. 35212]. •Valid as Lepidonectes clarkhubbsi Bussing 1991 -- (Allen & Robertson 1994:222 [ref. 22193], Fricke 1997:582 [ref. 23339], Fricke 2009:34 [ref. 30394], Patzner et al. 2009:448 [ref. 30396]). Current status: Valid as Lepidonectes clarkhubbsi Bussing 1991. Tripterygiidae: Tripterygiinae. Distribution: Pacific coast of Costa Rica to Panama. Habitat: marine.

clea, Enneapterygius Fricke [R.] 1997:176, Fig. 30 [ ref. 23339] Surge Channel, Escape Reef North, Cooktown area, Queensland, Australia, 15°49'S, 145°50'E, depth 5 meters. Holotype: AMS I.22600-056. Paratypes: AMS I.22600-069 (21), SMNS 19120 (2). Type catalog: Fricke 2005:64 [ref. 29864]. •Valid as Enneapterygius clea Fricke 1997 -- (Williams & Fricke 2001:3534 [ref. 26288], Hoese 2006:1520 [ref. 28998], Fricke 2009:38 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Meguro & Motomura 2010:6 [ref. 31252], Fricke & Erdmann 2017:22, 25 [ref. 35129]). Current status: Valid as Enneapterygius clea Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Queensland, Australia. Habitat: marine.

cocoensis, Axoclinus Bussing [W. A.] (ex Rosenblatt) 1991:78, Fig. 1 [Revista de Biología Tropical v. 39 (no. 1); ref. 19493] Eastern side of Isla del Coco, 850 meters north of Cabo Atrevida, 1.2 kilometers south of Punta Ulloa, Costa Rica, depth 7-8 meters. Holotype: LACM 44851-1. Paratypes: LACM (11 lots, 938 spec.); SIO 94-136 (15); UCR (17 lots, 405 spec.). Type catalog: Angulo et al. 2015:369 [ref. 35212]. •Valid as Enneanectes cocoensis (Bussing 1991) -- (Fricke 1997:565 [ref. 23339]). •Valid as Axoclinus cocoensis Bussing 1991 -- (Allen & Robertson 1994:219 [ref. 22193], Smith & Williams 2002:8 [ref. 26427], Fricke 2009:35 [ref. 30394], Patzner et al. 2009:444 [ref. 30396]). Current status: Valid as Axoclinus cocoensis Bussing 1991. Tripterygiidae: Tripterygiinae. Distribution: Costa Rica. Habitat: marine.

compressum, Tripterygium Hutton [F. W.] 1872:32 [Fishes of New Zealand (Hutton); ref. 2287] Stewart Island, New Zealand. Neotype: NMNZ P.27588. Type catalog: Russell 1996:231 [ref. 22562]. Original genus should have been Tripterygion. •Valid as Notoclinus compressus (Hutton 1872) -- (Paulin et al. 1989:219 [ref. 24556], Paulin & Roberts 1992:87 [ref. 24625], Fricke 1994:503 [ref. 21827], Jawad & Clements 2004:86 [ref. 27725], Fricke 2009:32 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Stewart & Clements 2015:1519 [ref. 34299], Cornejo et al. 2018:17 [ref. 36409]). Current status: Valid as Notoclinus compressus (Hutton 1872). Tripterygiidae: Notoclininae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

corallicola, Enneapterygius Kendall [W. C.] & Radcliffe [L.] 1912:153, Pl. 7 (fig. 1) [Memoirs of the Museum of Comparative Zoology v. 35 (no. 3); ref. 14961] San Cristóbal Island [Chatham Island], Galápagos Islands. Holotype: USNM 65484. Paratypes: MCZ 29492 (1) Type catalog: Springer & Orrell 1996:26 [ref. 22274]. •Valid as Lepidonectes corallicola (Kendall & Radcliffe 1912) -- (Allen & Robertson 1994:222 [ref. 22193], Fricke 1997:582 [ref. 23339], Grove & Lavenberg 1997:515 [ref. 24023], Herrera & Lavenberg 2002:4 [ref. 25975], Fricke 2009:34 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], McCosker & Rosenblatt 2010:193 [ref. 30957]). Current status: Valid as Lepidonectes corallicola (Kendall & Radcliffe 1912). Tripterygiidae: Tripterygiinae. Distribution: Eastern Pacific: Galápagos Islands. Habitat: marine.

cristata, Norfolkia Kuiter [R. H.] 1986:92, Figs. 4-5 [Revue française d'Aquariologie Herpétologie v. 12 (no. 3) (1985); ref. 6026] Flinders Island, South Australia, depth 4 meters. Holotype: NMV A3347. Paratypes: AMS I.24788-001 (1), NMV A3348 (3), SAMA F4835 (3), WAM P.27136-016 (3). Type catalog: Hutchins & Smith 1991:37 [ref. 19335], Moore et al. 2008:41 [ref. 30117]. •Valid as Norfolkia cristata Kuiter 1986 -- (Gomon et al. 1994:738 [ref. 22532], Kuiter 1997:334 [ref. 25488]). •Valid as Trinorfolkia cristata (Kuiter 1986) -- (Fricke 1994:547 [ref. 21827], Fricke 1997:586 [ref. 23339], Hoese 2006:1528 [ref. 28998], Gomon 2008:692 [ref. 30616], Fricke 2009:35 [ref. 30394], Patzner et al. 2009:448 [ref. 30396]). Current status: Valid as Trinorfolkia cristata (Kuiter 1986). Tripterygiidae: Tripterygiinae. Distribution: Australia: South Australia. Habitat: marine.

cunninghami, Tripterygium Smitt [F. A.] 1898:26, Pl. 3 (figs. 26-28). [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 24 (afd. 4) (no. 5); ref. 4147] Puerto Madryn, 43°S, Argentina, southwestern Atlantic. Lectotype: NRM 40358. Paralectotypes: NRM 45667 (4). Original genus should have been Tripterygion. •Valid as Tripterygion cunninghami Smitt 1898 -- (Pequeño 1989:66 [ref. 14125], Stepien 1990:600 [ref. 19879]). •Valid as Helcogrammoides cunninghami (Smitt 1898) -- (Gon 1990:400 [ref. 21680], Fricke 1997:581 [ref. 23339], Castillo & Pequeño 1998:113 [ref. 23987], Williams & Springer 2001:121 [ref. 26032], Jawad & Clements 2004:85 [ref. 27725], Fricke 2009:47 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Cornejo & Ibáñez 2018:275 [ref. 36410], Cornejo et al. 2018:17 [ref. 36409]). Current status: Valid as Helcogrammoides cunninghami (Smitt 1898). Tripterygiidae: Tripterygiinae. Distribution: Southeastern Pacific, southwestern Atlantic: southern Chile, southern Argentina. Habitat: marine.

decemdigitatus, Trypterigium Clarke [F. E.] 1879:292, Pl. 15 (lower right) [Transactions and Proceedings of the New Zealand Institute v. 11 (art. 25) (for 1878); ref. 18006] Dusky Sound, New Zealand. Holotype (unique): whereabouts unknown. Correct spelling of the genus is Tripterygion; species name apparently should be spelled decemdigitatum when in Tripterygion. •Valid as Ruanoho decemdigitatus (Clarke 1879) -- (Hardy 1986:158 [ref. 6139], Paulin et al. 1989:219 [ref. 24556], Paulin & Roberts 1992:78 [ref. 24625], Fricke 1994:521 [ref. 21827], Fricke 1997:585 [ref. 23339], Jawad & Clements 2004:87 [ref. 27725], Wellenreuther & Clements 2008:575 [ref. 30937], Fricke 2009:33 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1521 [ref. 34299]). Current status: Valid as Ruanoho decemdigitatus (Clarke 1879). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

decurrens, Helcogramma McCulloch [A. R.] & Waite [E. R.] 1918:52, Pl. 3 (fig. 2) [Records of the South Australian Museum (Adelaide) v. 1 (no. 1); ref. 2950] St. Vincent Gulf, South Australia. Holotype (unique): SAMA F598. Type catalog: Glover 1976:174 [ref. 19449]. •Valid as Helcogramma decurrens McCulloch & Waite 1918 -- (Hansen 1986:333 [ref. 5810], Allen & Swainston 1988:132 [ref. 25477], Fricke 1994:417 [ref. 21827], Gomon et al. 1994:734 [ref. 22532], Fricke 1997:577 [ref. 23339], Allen 1997:212 [ref. 23977], Kuiter 1997:334 [ref. 25488], Hutchins 2001:41 [ref. 25847], Hoese 2006:1524 [ref. 28998], Gomon 2008:687 [ref. 30616], Fricke 2009:48 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Hoschke et al. 2019:159 [ref. 36626]). Current status: Valid as Helcogramma decurrens McCulloch & Waite 1918. Tripterygiidae: Tripterygiinae. Distribution: Southeastern Indian Ocean: Western Australia, South Australia. Habitat: marine.

delaisi, Tripterygion Cadenat [J.] & Blache [J.] 1970:1097, Fig. 1 [Bulletin du Muséum National d'Histoire Naturelle (Série 2) v. 41 (no. 5) (for 1969); ref. 7721] Gorée, Senegal. Holotype: MNHN 1963-210. Paratypes: MNHN 1963-0211 (1), 1963-0970 (1); Marine Station of Gorée 50-866 (1), 50-935-936 (2), 50-966 (1), 50-977 (1), 50-993 to 995 (3). •Valid as Tripterygion delaisi Cadenat & Blache 1970 -- (Zander in Whitehead et al. 1986:1119 [ref. 13677], Wirtz 1990:916 [ref. 19374], Ahnelt 1991:55 [ref. 19120], Fricke 1997:587 [ref. 23339], Santos et al. 1997:98 [ref. 23531], Arruda 1997:108 [ref. 24952], Geertjes et al. 2001:256 [ref. 25579], Bilecenoğlu et al. 2002:119 [ref. 26753], Carreras-Carbonell et al. 2007:76 [ref. 29025] dated 1971, Fricke 2009:43 [ref. 30394], Patzner et al. 2009:449 [ref. 30396], Hastings & Springer 2009:11 [ref. 31156], Lipej & Dulčić 2010:57 [ref. 36649], Williams in Carpenter & De Angelis 2016:2795 [ref. 34618], Cornejo et al. 2018:17 [ref. 36409], Artüz & Fricke 2019:558 [ref. 36612]). Current status: Valid as Tripterygion delaisi Cadenat & Blache 1970. Tripterygiidae: Tripterygiinae. Distribution: Mediterranean Sea, Sea of Marmara, Eastern Atlantic: English Channel south to Senegal, including Azores, Madeira and Canary Islands. Habitat: marine.

deloachorum, Enneanectes Victor [B. C.] 2013:60, Figs. 17-19 [Journal of the Ocean Science Foundation v. 7; ref. 32783] Dominica, Roseau (15.28°, -61.38°). Holotype: UF 158602. Paratypes: UF 185605 (1), 185606 (1), 185612 (2). •Valid as Enneanectes deloachorum Victor 2013 -- (Victor 2017:69 [ref. 35549], Victor 2019:13 [ref. 36450]). Current status: Valid as Enneanectes deloachorum Victor 2013. Tripterygiidae: Tripterygiinae. Distribution: Western Atlantic: Dominica to ABC Netherlands Antilles. Habitat: marine.

desa, Helcogramma Williams [J. T.] & Howe [J. C.] 2003:161, Figs. 8-9 [aqua, Journal of Ichthyology and Aquatic Biology v. 7 (no. 4); ref. 27249] Cocoro Island, 10°52'54"N, 121°12'14"E, Cuyo Islands, Philippines, depth 0-3 meters. Holotype: USNM 222317. Paratypes: AMS I.35025-001 (1); BPBM 36433 (1); USNM 222320 (18), 222898 (5); ROM 73085 (30), 73086 (40). Plus nontypes. •Valid as Helcogramma desa Williams & Howe 2003 -- (Fricke 2009:48 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:773 [ref. 31980], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma desa Williams & Howe 2003. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Philippines and Vietnam. Habitat: marine.

destai, Enneapterygius Clark [E.] 1980:102, Figs. 4b, 13 [Israel Journal of Zoology v. 28 (nos 3-4) [for 1979]; ref. 8837] Northern end of Delemone Island, just east of Ras Coral, Ethiopia, Red Sea. Holotype: USNM 214629. Paratypes: HUJ 5488-90 (1, 1, 3), 5498 (2), 5502 (1), 9210 (20 or 27), 9211 (3); SMF 9494 (1); TAU [= SMNHTAU] N.S. 4397 (1); USNM 205798-99 (14, 2). Type catalog: Springer & Orrell 1996:26 [ref. 22274], Golani 2006:31 [ref. 28840]. Spelled destae in caption to Fig. 13; named for a man so destai is correctly formed. •Valid as Enneapterygius destai Clark 1980 -- (Dor 1984:233 [ref. 29757], Goren & Dor 1994:61 [ref. 25356], Fricke 1997:566 [ref. 23339], Holleman 2005:6 [ref. 28455], Fricke 2009:38 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Holleman & Bogorodsky 2012:46 [ref. 31735], Bogorodsky & Randall 2018:[6] 244 [ref. 36492], Golani & Fricke 2018:143 [ref. 36273]). Current status: Valid as Enneapterygius destai Clark 1980. Tripterygiidae: Tripterygiinae. Distribution: Red Sea endemic. Habitat: marine.

dorsalis, Trypterigium Clarke [F. E.] 1879:291, Pl. 15 (lower left) [Transactions and Proceedings of the New Zealand Institute v. 11 (art. 25) (for 1878); ref. 18006] Mouth of Hokitika, New Zealand. Holotype (unique): whereabouts unknown. Gender of genus Blennodon is masculine; therefore correct species name dorsalis. Original genus should have been Tripterygion. •Valid as Blennodon dorsale (Clarke 1879) -- (Hardy 1987:159 [ref. 5959], Paulin et al. 1989:220 [ref. 24556], Paulin & Roberts 1992:76 [ref. 24625], Fricke 1994:140 [ref. 21827], Fricke 1997:562 [ref. 23339], Patzner et al. 2009:444 [ref. 30396], Feary et al. 2009:1704 [ref. 30938]). •Valid as Blennodon dorsalis (Clarke 1879) -- (Fricke 2009:52 [ref. 30394], Stewart & Clements 2015:1500 [ref. 34299] as dorsale). Current status: Valid as Blennodon dorsalis (Clarke 1879). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

elaine, Enneapterygius Holleman [W.] 2005:7, Fig. 6, Pl. 1 (A,B) [Smithiana, Publications in Aquatic Biodiversity, Bulletin No. 5; ref. 28455] Baladirou, 19°40'S, 63°26'E, Rodrigues Island, Mascarenes, southwestern Indian Ocean. Holotype: SAIAB [SAIAB [formerly RUSI]] 69185. Paratypes: AMS 441476-001 (2); BPBM 39012 (4); BMNH 2002.8.22.1-3 (3); ROM 73204 (4); SAIAB 65577 (3), 65578 (5), 65579 (2), 65580 (3), 65581 (3), 65584 (5), 65586 (5); USNM 370527 (4). •Valid as Enneapterygius elaine Holleman 2005 -- (Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396]). Current status: Valid as Enneapterygius elaine Holleman 2005. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Indian Ocean: Rodrigues (Mascarenes endemic). Habitat: marine.

elegans, Tripterygium Peters [W. (C. H.)] 1876:441 [Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1876; ref. 18495] Mauritius [Île aux Fouquets, 8 kilometers east of Mahébourg, Mascarenes, southwestern Indian Ocean]. Lectotype: ZMB 9465. Paralectotypes: ZMB 32770 [ex ZMB 9465] (2). Original genus should have been Tripterygion. •Valid as Enneapterygius elegans (Peters 1876) -- (Holleman 1986:756 [ref. 5671], Fricke 1987:336 [ref. 9108], Fricke & Randall 1992:3 [ref. 20288], Fricke 1994:203 [ref. 21827], Fricke 1997:179, 566 [ref. 23339], Randall et al. 1997:521 [ref. 25919], Fricke 1999:467 [ref. 24106], Shimojyo & Hayashi 2000:46 [ref. 24903], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1084 [ref. 25182], Fricke 2001:5 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288] dated 1877, Nakabo 2002:1084 [ref. 26193], Fricke 2002:182 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830] dated 1877, Randall et al. 2004:24 [ref. 27624] dated 1877, Heemstra et al. 2004:3327 [ref. 27945], Randall 2005:471 [ref. 28239], Holleman 2005:8 [ref. 28455], Hoese 2006:1520 [ref. 28998], Chiang & Chen 2008:194 [ref. 29932] dated 1877, Fricke 2009:38 [ref. 30394], Fricke et al. 2009:93 [ref. 30213], Patzner et al. 2009:445 [ref. 30396], Allen & Erdmann 2012:765 [ref. 31980], Fricke & Erdmann 2017:28 [ref. 35129], Fricke et al. 2018:304 [ref. 35805], Arndt & Fricke 2019:31 [ref. 36883]). Current status: Valid as Enneapterygius elegans (Peters 1876). Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: East Africa, Madagascar and western Mascarenes east to Samoa and Tonga, south to New Caledonia. Habitat: marine.

ellioti, Tripterygion Herre [A. W. C. T.] 1944:49 [Proceedings of the Biological Society of Washington v. 57; ref. 2137] Maharanepeta Beach, Vizagapatam, India. Lectotype: CAS-SU 38840 (male). Paralectotypes: CAS-SU 38840 (1, female), 38841-45 (1, 1, 8, 28, 114); USNM 123656 [ex SU] (4). Herre considered a male and a female as his types, the rest as paratypes; Böhlke 1953:99 [ref. 12291] treated the male as holotype and the female as allo-paratype; the male was designated as lectotype by Springer & Orrell 1996:26 [ref. 22274]. •Valid as Helcogramma ellioti (Herre 1944) -- (Hansen 1986:331 [ref. 5810], Fricke 1997:578 [ref. 23339], Nakabo 2000:1079 [ref. 25182], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1079 [ref. 26193], Manilo & Bogorodsky 2003:S116 [ref. 27377], Randall 2005:477 [ref. 28239], Holleman 2007:59 [ref. 29195], Fricke 2009:48 [ref. 30394], Patzner et al. 2009:447 [ref. 30396]). Current status: Valid as Helcogramma ellioti (Herre 1944). Tripterygiidae: Tripterygiinae. Distribution: Indian Ocean: southern India. Habitat: marine.

ememes, Helcogramma Holleman [W.] 2007:61, Fig. 5; Pl. 1 [Smithiana, Publications in Aquatic Biodiversity, Bulletin No. 7; ref. 29195] Ibo Island, 12°24'S, 40°34'E, Mozambique. Holotype: SAIAB 70605. Paratypes: BMNH (2004.1.6.7-8 and .9-10), BPBM, ROM, and SAIAB. •Valid as Helcogramma ememes Holleman 2007 -- (Fricke 2009:49 [ref. 30394], Patzner et al. 2009:447 [ref. 30396]). Current status: Valid as Helcogramma ememes Holleman 2007. Tripterygiidae: Tripterygiinae. Distribution: Western Indian Ocean. Habitat: marine.

erythrosoma, Enneapterygius Shen [S.-C.] 1994:7, Fig. 5 [Acta Zoologica Taiwanica v. 5 (no. 2); ref. 21605] Wen-tz-keng, north coast of Taiwan. Holotype: NTUM 07815. Paratypes: NTUM 07816 (1), 07852 (1), 07854 (1); USNM 329658 (1). Non-type material: NTUM. Type catalog: Springer & Orrell 1996:26 [ref. 22274], Ho & Shao 2011:52 [ref. 31432]. Authorship is Shen alone, not Shen & Wu. •Synonym of Enneapterygius rubicauda Shen 1994 -- (Fricke 1997:310, 571 [ref. 23339], Fricke 2009:42 [ref. 30394]). •Valid as Enneapterygius erythrosomus Shen -- (Chiang & Chen 2008:194 [ref. 29932] with authorship as Shen & Wu, Motomura et al. 2010:185 [ref. 31256]). •Valid as Enneapterygius erythrosoma Shen -- (Ho & Shao 2011:52 [ref. 31432], Meguro & Motomura 2010:1 [ref. 31252]). Current status: Synonym of Enneapterygius rubicauda Shen 1994. Tripterygiidae: Tripterygiinae. Distribution: Western North Pacific: Taiwan. Habitat: marine.

etheostoma, Tripterygium Jordan [D. S.] & Snyder [J. O.] 1902:444, Fig. 1 [Proceedings of the United States National Museum v. 25 (no. 1293); ref. 2516] Misaki, Japan. Holotype: CAS-SU 7065. Paratypes: CAS-SU 7100 (27), USNM 50299 (7). Type catalog: Böhlke 1953:99 [ref. 12291], Springer & Orrell 1996:26 [ref. 22274]. Types: Shinohara & Williams 2006:561 [ref. 31656]. Original genus should have been Tripterygion. Treated as a noun in apposition, the spelling etheostoma is correct in Enneapterygius. •Valid as Tripterygion etheostoma (Jordan & Snyder) -- (Lindberg & Krasyukova 1975:24 [ref. 7348]). •Valid as Rosenblatella etheostoma (Jordan & Snyder 1902) -- (Shen 1994:25 [ref. 21605]). •Valid as Enneapterygius etheostomus (Jordan & Snyder 1902) -- (Yoshino in Masuda et al. 1984:294 [ref. 6441] with authors as Jordan & Seale, Fricke 1987:336 [ref. 9108], Fricke 1997:185, 567 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1083 [ref. 25182], Sadovy & Cornish 2000:233 [ref. 25480], Shinohara et al. 2000:181 [ref. 25987], Shimizu 2001:36 [ref. 25799], Nakabo 2002:1083 [ref. 26193], Youn 2002:387, 649 [ref. 26218], Motomura et al. 2005:13 [ref. 28338] as etheostoma, Chiang & Chen 2008:194 [ref. 29932], Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Endo et al. 2010:14 [ref. 30815] as etheostoma, Motomura et al. 2010:183 [ref. 31256], Motomura et al. 2015:27 [ref. 34709] as etheostoma, Fricke & Erdmann 2017:27 [ref. 35129], Cornejo et al. 2018:17 [ref. 36409]). Current status: Valid as Enneapterygius etheostoma (Jordan & Snyder 1902). Tripterygiidae: Tripterygiinae. Distribution: Northwestern Pacific: Vietnam and China east to Taiwan, north to Korea and southern Japan. Habitat: marine.

exsul, Enneanectes Rosenblatt [R. H.], Miller [E. C.] & Hastings [P. A.] 2013:369, Figs. 5-6 [Zootaxa 3636 (no. 2); ref. 32572] North of Caleta Binner, Isla Socorro, Islas Revillagigedo, Mexico, depth 0-3 meters. Holotype: SIO 11-392. Paratypes: CAS 235204 (1); LACM 31783-7 (5), 32096-18 (3), 32097-43 (4); SIO 11-394 (5), 11-395 (1), 11-396 (1), 12-65 (2), SIO 70-392 (9); USNM 410092 (1). •Valid as Enneanectes exsul Rosenblatt, Miller & Hastings 2013 -- (Fourriére et al. 2016:449 [ref. 34668], Del Moral-Flores et al. 2016:610 [ref. 34923]). Current status: Valid as Enneanectes exsul Rosenblatt, Miller & Hastings 2013. Tripterygiidae: Tripterygiinae. Distribution: Revillagigedo Islands (Mexico), eastern Pacific. Habitat: marine.

fasciatum, Tripterygium Weber [M.] 1909:148 [Notes from the Leyden Museum v. 31 (note 4); ref. 4600] Seba, Savu Island; Beo, Karakelang Island, Indonesia. Syntypes: ZMA 112507-08 (1, 2). Type catalog: Nijssen et al. 1982:95 [ref. 19236]. Original genus should have been Tripterygion. •Valid as Enneapterygius fasciatus (Weber 1909) -- (Fricke 1994:4 [ref. 21828], Fricke 1997:193, 567 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Allen & Adrim 2003:53 [ref. 26830], Holleman 2005:9 [ref. 28455], Chiang & Chen 2008:195 [ref. 29932], Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Allen & Erdmann 2012:766 [ref. 31980], Fricke et al. 2014:145 [ref. 33932], Fricke & Erdmann 2017:23, 28, 29 [ref. 35129], Fricke et al. 2018:304 [ref. 35805]). Current status: Valid as Enneapterygius fasciatus (Weber 1909). Tripterygiidae: Tripterygiinae. Distribution: East Africa, Seychelles, Comoros and Madagascar east to Philippines and New Guinea, north to Taiwan, south to Solomon Islands. Habitat: marine.

fasciatus, Brachynectes Scott [T. D.] 1957:181, Fig. 1 [Transactions Royal Society of South Australia v. 80; ref. 3997] Pelican Lagoon, Kangaroo Island, South Australia. Holotype: SAMA F2921. Additional material: WAM P.4846 (2). Type catalog: Glover 1976:174 [ref. 19449], Hutchins & Smith 1991:37 [ref. 19335] with remarks, Moore et al. 2008:41 [ref. 30117] with remarks. •Valid as Brachynectes fasciatus Scott 1957 -- (Kuiter 1993:323 [ref. 23929], Fricke 1994:146 [ref. 21827], Gomon et al. 1994:733 [ref. 22532], Fricke 1997:563 [ref. 23339], Hutchins 2001:41 [ref. 25847], Hoese 2006:1518 [ref. 28998], Gomon 2008:685 [ref. 30616], Fricke 2009:32 [ref. 30394], Patzner et al. 2009:444 [ref. 30396]). Current status: Valid as Brachynectes fasciatus Scott 1957. Tripterygiidae: Notoclininae. Distribution: Australia: Victoria, Tasmania, South Australia and Western Australia. Habitat: marine.

fenestratus, Blennius Forster [J. R.] in Bloch & Schneider 1801:173 [M. E. Blochii, Systema Ichthyologiae; ref. 471] Detention Cove, Dusky Sound, Fiordland, South Island, New Zealand. Holotype (unique): not preserved. Authorship follows Fricke 1994:17 [ref. 21827]. Based on description in manuscript by Forster (Johann Reinhold Forster, 1729-1798, German explorer and naturalist of Cook's second voyage). •Valid as Notoclinus fenestratus (Forster 1801) -- (Paulin et al. 1989:219 [ref. 24556], Paulin & Roberts 1992:86 [ref. 24625], Fricke 1994:507 [ref. 21827], Fricke 1997:584 [ref. 23339], Jawad & Clements 2004:86 [ref. 27725], Fricke 2009:32 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Stewart & Clements 2015:1520 [ref. 34299]). Current status: Valid as Notoclinus fenestratus (Forster 1801). Tripterygiidae: Notoclininae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

flavoccipitis, Enneapterygius Shen [S.-C.] 1994:8, Fig. 6 [Acta Zoologica Taiwanica v. 5 (no. 2); ref. 21605] Ho-bi-hou, southern coast of Taiwan. Holotype: NTUM 07836. Paratypes: NTUM 07837 (1), 07842 (1); USNM 329659 (1). Type catalog: Springer & Orrell 1996:26 [ref. 22274], Ho & Shao 2011:52 [ref. 31432]. Authorship is Shen alone, not Shen & Wu. •Valid as Enneapterygius flavoccipitis Shen 1994 [often with authors as Shen & Wu] -- (Fricke 1997:199, 567 [ref. 23339], Shimojyo & Hayashi 2000:48 [ref. 24903], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1084 [ref. 25182], Hutchins 2001:41 [ref. 25847], Fricke 2001:4, 5 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1084 [ref. 26193], Fricke 2002:184 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830] with authors as Shen & Wu, Randall et al. 2004:24 [ref. 27624] with authors Shen & Wu, Randall 2005:472 [ref. 28239], Hoese 2006:1520 [ref. 28998], Chiang & Chen 2008:195 [ref. 29932] with authorship as Shen & Wu, Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Ho & Shao 2011:52 [ref. 31432], Allen & Erdmann 2012:766 [ref. 31980], Larson et al. 2013:177 [ref. 32988], Fricke et al. 2014:145 [ref. 33932], Fricke & Erdmann 2017:22, 24, 26 [ref. 35129]). Current status: Valid as Enneapterygius flavoccipitis Shen 1994. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Indonesia to Philippines, Papua New Guinea to New Caledonia and Loyalty Islands, Australia, and north to Taiwan and Ryukyu Islands. Habitat: marine.

flavonigrum, Forsterygion Fricke [R.] & Roberts [C. D.] in Fricke 1994:314 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Off Seymore Island, Doubtful Sound, 45°18'S, 167°00'E, Fiordland, South Island, New Zealand, depth 12 meters. Holotype: NMNZ P.16927. Paratypes: AMS uncat. (10); BMNH uncat. (1); NMNZ P.21194 (7), P.15908 (1), P.18255 (1), P.18285 (2), P.18442 (4), P.21653 (1), P.21808 (1), P.21815 (8), P.28284 (8), P.30198 (31); SMNS 24582 (10); USNM uncat. (10). Plus more paratypes at NMNZ. Type catalog: Fricke 1995:20-21 [ref. 28100], Fricke 2005:68 [ref. 29864]. •Valid as Forsterygion flavonigrum Fricke & Roberts 1994 -- (Fricke 1997:573 [ref. 23339], Jawad & Clements 2004:89 [ref. 27725], Jawad 2008:2965 [ref. 29958], Fricke 2009:45 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1504 [ref. 34299]). Current status: Valid as Forsterygion flavonigrum Fricke & Roberts 1994. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

flavus, Enneanectes Victor [B. C.] 2019:3, Figs. 1-6 [Journal of the Ocean Science Foundation v. 32; ref. 36450] North end of Isla Noreste, Los Testigos Archipelago, Venezuela, 11.41255°, -63.04725°. Holotype: AMNH 245238. Paratypes: AMNH, USNM. Plus additional non-type material. •Valid as Enneanectes flavus Victor 2019. Current status: Valid as Enneanectes flavus Victor 2019. Tripterygiidae: Tripterygiinae. Distribution: Western Atlantic: northeastern Venezuela, Tobago and possibly St. Vincent. Habitat: marine.

forsteri, Tripterygion Valenciennes [A.] in Cuvier & Valenciennes 1836:415 [Histoire naturelle des poissons v. 11; ref. 1005] Outer Chetwode Island, Golden Bay and Marlborough sounds, South Island, New Zealand. Neotype: NMNZ P.15419. Type catalog: Bauchot 1967:57 [ref. 20734]. Unneeded new name for Blennius tripennis Forster in Bloch & Schneider 1801; now with same neotype (see tripennis). •Synonym of Gilloblennius tripennis (Forster 1801) -- (Hardy 1986:148 [ref. 6139], Fricke 1997:576 [ref. 23339], Fricke 2009:33 [ref. 30394]). Current status: Synonym of Gilloblennius tripennis (Forster 1801). Tripterygiidae: Tripterygiinae. Habitat: marine.

fuligicauda, Enneapterygius Fowler [H. W.] 1946:188, Fig. 52 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 98; ref. 1456] Aguni Shima, Ryukyu Islands, Japan. Holotype: ANSP 72062. Paratypes: ANSP 72063-64 (2). Type catalog: Böhlke 1984:164 [ref. 13621]. •Synonym of Enneapterygius hemimelas (Kner & Steindachner 1867) -- (Fricke 1994:215 [ref. 21827]). •Synonym of Enneapterygius philippinus (Peters 1868) -- (Fricke 1997:276, 570 [ref. 23339], Fricke 2009:41 [ref. 30394]). Current status: Synonym of Enneapterygius philippinus (Peters 1868). Tripterygiidae: Tripterygiinae. Habitat: marine.

fuscipectoris, Enneapterygius Fowler [H. W.] 1946:186, Figs. 50-51 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 98; ref. 1456] Aguni-jima, Ryukyu Islands, Japan. Holotype: ANSP 72060. Paratypes: ANSP 72061 (1). Type catalog: Böhlke 1984:164 [ref. 13621]. See Fricke 1997:433 [ref. 23339] for selection of fuscipectoris over personatus and quadrimaculatus. •Synonym of Helcogramma obtusirostris (Klunzinger 1871) -- (Hansen 1986:341 [ref. 5810], Shen 1994:21 [ref. 21605]). •Valid as Helcogramma fuscipectoris (Fowler 1946) -- (Fricke 1997:429, 578 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1080 [ref. 25182], Williams & Fricke 2001:3534 [ref. 26288], Allen & Adrim 2003:53 [ref. 26830], Fricke 2009:49 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:774 [ref. 31980], Chiang & Chen 2012:58, 64 [ref. 32986], Tashiro & Motomura 2014:104 [ref. 35481], Koeda et al. 2016:83 [ref. 35333], Fricke & Erdmann 2017:43 [ref. 35190]). Current status: Valid as Helcogramma fuscipectoris (Fowler 1946). Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Malaysia, Indonesia and Philippines to China, Japan and Vanuatu. Habitat: marine.

fuscopinna, Helcogramma Holleman [W.] 1982:115, Fig. 4 [Annals of the Cape Provincial Museums (Natural History) v. 14 (pt 4); ref. 2187] Reef offshore of Sodwana Bay, 27°31'S, 32°41'E, Zululand, KwaZulu-Natal, South Africa, western Indian Ocean. Holotype: SAIAB [formerly RUSI] 954. Paratypes: BMNH 1978.5.30.1-2 (2); BPBM 21164 (5); ROM 38782-38785 (8, 4 lots); SAIAB [formerly RUSI] 490 (28), 489 (2), 955-956 (2, 1); USNM 227738-47 (4, 1, 33, 33, 3, 3, 2, 9, 19, 91); WAM P26507-016 (2). Type catalog: Springer & Orrell 1996:26 [ref. 22274], Ho & Shao 2011:53 [ref. 31432]. •Valid as Helcogramma fuscopinna Holleman 1982 -- (Holleman 1986:757 [ref. 5671], Hansen 1986:337 [ref. 5810], Winterbottom et al. 1989:62 [ref. 13251], Williams & McCormick 1990:1020 [ref. 18992], Fricke & Randall 1992:6 [ref. 20288], Fricke 1997:578 [ref. 23339], Fricke 1999:468 [ref. 24106], Nakabo 2000:1079 [ref. 25182], Allen 2000:116 [ref. 25868], Nakabo 2002:1079 [ref. 26193], Williams & Howe 2003:163 [ref. 27249], Manilo & Bogorodsky 2003:S116 [ref. 27377], Heemstra et al. 2004:3327 [ref. 27945], Heemstra & Heemstra 2004:387 [ref. 28072], Holleman 2006:102 [ref. 28864], Holleman 2007:63 [ref. 29195], Fricke 2009:49 [ref. 30394], Fricke et al. 2009:93 [ref. 30213], Patzner et al. 2009:447 [ref. 30396], Motomura et al. 2010:187 [ref. 31256], Ho & Shao 2011:53 [ref. 31432], Chiang & Chen 2012:58 [ref. 32986], Fricke et al. 2018:305 [ref. 35805]). Current status: Valid as Helcogramma fuscopinna Holleman 1982. Tripterygiidae: Tripterygiinae. Distribution: East and South Africa, Seychelles, Madagascar and Mascarenes east to Maldives and Chagos Archipelago. Habitat: marine.

fuscoventer, Enneapterygius Fricke [R.] 1997:210, Fig. 38 [ ref. 23339] Putic Island, northwestern side (Cuyo Island), Palawan Province, Philippines, 10°55'05"N, 121°02'03"E, depth 0-4.5 meters. Holotype: USNM 259131. Paratypes: SMNS 19172 (2); USNM 259136 (1), 293711 (1), 295566 (9), 345509 (15). Type catalog: Fricke 2005:64 [ref. 29864]. •Valid as Enneapterygius fuscoventer Fricke 1997 -- (Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Chiang & Chen 2008:196 [ref. 29932], Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Allen & Erdmann 2012:766 [ref. 31980], Fricke & Erdmann 2017:24 [ref. 35129], Tashiro & Motomura 2018:[2] 253 [ref. 35918]). Current status: Valid as Enneapterygius fuscoventer Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Philippines and Papua New Guinea east to Society Islands, north to Ryukyu Islands. Habitat: marine.

geminatus, Lepidoblennius Macleay [W.] 1881:13 [Proceedings of the Linnean Society of New South Wales v. 6 (pt 1); ref. 4931] Port Jackson, New South Wales, Australia. Holotype (unique): AMS I.16404-001 [ex MAMU F1026]. Type catalog: Stanbury 1969:209 [ref. 19518]. •Synonym of Lepidoblennius haplodactylus Steindachner 1867 -- (Hardy 1987:254 [ref. 5960], Gomon et al. 1994:736 [ref. 22532], Fricke 1997:581 [ref. 23339], Hoese 2006:1525 [ref. 28998], Fricke 2009:51 [ref. 30394]). Current status: Synonym of Lepidoblennius haplodactylus Steindachner 1867. Tripterygiidae: Tripterygiinae. Habitat: marine.

genamaculatus, Enneapterygius Holleman [W.] 2005:11, Fig. 10 [Smithiana, Publications in Aquatic Biodiversity, Bulletin No. 5; ref. 28455] Southeastern side of Grande Passe, Saint Brandon's Shoals, about 16°28'S, 59°40'E, depth 0-6 m. Holotype: USNM 357600. Paratypes: SAIAB 60503 (6); USNM 357597 (2), 357599 (29), 357602 (12), 357606 (2). •Valid as Enneapterygius genamaculatus Holleman 2005 -- (Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396]). Current status: Valid as Enneapterygius genamaculatus Holleman 2005. Tripterygiidae: Tripterygiinae. Distribution: Western Indian Ocean: Saint Brandon's Shoals. Habitat: marine.

glendae, Enneanectes Rosenblatt [R. H.], Miller [E. C.] & Hastings [P. A.] 2013:362, Figs. 1-2 [Zootaxa 3636 (no. 2); ref. 32572] Ensenada Chacala, off Punta Chacala, Nayarit, Mexico, depth 12 meters. Holotype: SIO 11-394 [ex UCLA 58-10]. Paratypes: CAS 235202 (2); LACM 58001-1 (2); SIO 11-393 (9), 11-394 (6), 70-167 (40); USNM 410090 (2); UW 151893 (1), plus more at SIO. Plus additional non-type material. •Valid as Enneanectes glendae Rosenblatt, Miller & Hastings 2013 -- (Galván-Villa et al. 2016:148 [ref. 34871]). Current status: Valid as Enneanectes glendae Rosenblatt, Miller & Hastings 2013. Tripterygiidae: Tripterygiinae. Distribution: Mexico and Islas Tres Marias, Mexico. Habitat: marine.

gracilis, Crocodilichthys Allen [G. R.] & Robertson [D. R.] 1991:79, Fig. 2 [Revue française d'Aquariologie Herpétologie v. 18 (no. 3); ref. 19723] Los Islotes, Partida Island, Gulf of California, Mexico, about 25°02'N, 110°32'W, depth 10-12 meters. Holotype: USNM 316798. Paratypes: USNM 316799 (1), WAM P.30231-001 (1). Type catalog: Springer & Orrell 1996:26-27 [ref. 22274], Moore et al. 2008:41 [ref. 30117]. •Valid as Crocodilichthys gracilis Allen & Robertson 1991 -- (Allen & Robertson 1994:221 [ref. 22193], De La Cruz Agüero et al. 1997:302 [ref. 24545], Fricke 1997:564 [ref. 23339], Thomson et al. 2000:204 [ref. 25640], Nelson et al. 2004:161 [ref. 27807], Fricke 2009:36 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708]). Current status: Valid as Crocodilichthys gracilis Allen & Robertson 1991. Tripterygiidae: Tripterygiinae. Distribution: Eastern Pacific: Gulf of California. Habitat: marine.

gracilis, Enneapterygius Fricke [R.] 1994:209, Fig. 34 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Cobourg Pennsula, Danger Point, 11°08'S, 132°20'E, Northern Territory, Australia, depth 0-1.5 meters. Holotype: NTM S.10431-027. Paratypes: AMS I.17060-043 (6), IB.7075 (1); NTM S.10005-27 (1); SMNS 14205 (1), 14552 (6), 14612 (8); AMS I.17060-043 (6); USNM 279864 (1). Plus many more paratype lots at NTM and WAM. Type catalog: Fricke 1995:20 [ref. 28100], Springer & Orrell 1996:27 [ref. 22274], Fricke 2005:65 [ref. 29864], Moore et al. 2008:41 [ref. 30117]. •Valid as Enneapterygius gracilis Fricke 1994 -- (Larson & Williams 1997:367 [ref. 23967], Fricke 1997:567 [ref. 23339], Hutchins 2001:41 [ref. 25847], Williams & Fricke 2001:3534 [ref. 26288], Hoese 2006:1520 [ref. 28998], Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Larson et al. 2013:177 [ref. 32988]). Current status: Valid as Enneapterygius gracilis Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Northern Australia. Habitat: marine.

gruschkai, Enneapterygius Holleman [W.] 2005:11, Fig. 11, Pl. 1E [Smithiana, Publications in Aquatic Biodiversity, Bulletin No. 5; ref. 28455] Passe d'Ambulante, Mauritius, Mascarenes, southwestern Indian Ocean, 20°26'10"S, depth 6-8 meters. Holotype: SAIAB 61281. Paratypes: BPBM 35600 (1); CAS 33424 (1); ROM 56795 (4), 56796 (1), 56799 (2), 56800 (1), 56801 (5), 56802 (6); SAIAB 6122 (11), 61283 (11); USNM 357601 (6), 357607 (1), 357611 (2), 357608 (8). Plus non-type material. •Valid as Enneapterygius gruschkai Holleman 2005 -- (Fricke 2009:39 [ref. 30394], Fricke et al. 2009:93 [ref. 30213], Patzner et al. 2009:445 [ref. 30396], Fricke & Erdmann 2017:30 [ref. 35129]). Current status: Valid as Enneapterygius gruschkai Holleman 2005. Tripterygiidae: Tripterygiinae. Distribution: Western Indian Ocean: Comores, Seychelles, Cargados Carajos, Réunion and Mauritius (western Mascarenes), Chagos Archipelago. Habitat: marine.

gymnauchen, Tripterygium Weber [M.] 1909:149 [Notes from the Leyden Museum v. 31 (note 4); ref. 4600] Timor Island, southern Malay Archipelago. Holotype (unique): ZMA 112509. Type catalog: Nijssen et al. 1982:95 [ref. 19236]. Original genus should have been Tripterygion. •Synonym of Helcogramma hudsoni (Jordan & Seale 1906) -- (Hansen 1986:339 [ref. 5810]). •Valid as Helcogramma gymnauchen (Weber 1909) -- (Yoshino in Masuda et al. 1984:293 [ref. 6441], Randall et al. 1990:365 [ref. 15987], Fricke 1994:421 [ref. 21827], Fricke 1997:437, 578 [ref. 23339], Williams & Fricke 2001:3534 [ref. 26288], Allen & Adrim 2003:53 [ref. 26830], Randall 2005:476 [ref. 28239], Hoese 2006:1524 [ref. 28998], Fricke 2009:49 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:774 [ref. 31980], Larson et al. 2013:178 [ref. 32988], Fricke et al. 2014:146 [ref. 33932], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma gymnauchen (Weber 1909). Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Indonesia to Papua New Guinea, northern Australia and New Caledonia. Habitat: marine.

gymnotum, Forsterygion Scott [E. O. G.] 1977:158, Fig. 3 [Papers and Proceedings of the Royal Society of Tasmania v. 111; ref. 7283] Montague Bay, Derwent River, southeastern Tasmania, Australia. Holotype: QVM 1976:5:0204 [ex QVM type 232]. Paratypes: AMS I.20105-001 [ex QVM 1976:5:208] (1); BMNH 1978.3.3.1 (1); QVM 1976:5:205-206 [ex QVM types 233-234] (2); TMH D1310 (1). Type catalog: Andrews 1992:112 [ref. 22689]. See Grahamina signata Fricke & Roberts 1993. •Tripterygiid but not Forsterygion -- (Hardy 1989:509 [ref. 12485]). •Valid as Grahamina gymnota (Scott 1977) -- (Fricke 1994:387 [ref. 21827], Fricke 1997:576 [ref. 23339], Clements et al. 2000:373 [ref. 25188], Jawad & Clements 2004:85 [ref. 27725], Hoese 2006:1524 [ref. 28998], Fricke 2009:45 [ref. 30394], Patzner et al. 2009:446 [ref. 30396]). •Valid as Forsterygion gymnota (Scott 1977) -- (Feary et al. 2009:1704 [ref. 30938]). •Valid as Forsterygion gymnotum Scott 1977 -- (Jawad 2008:2967 [ref. 29958] ? as gymnota, Stewart & Clements 2015:1505 [ref. 34299]). Current status: Valid as Forsterygion gymnotum Scott 1977. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region; introduced in Australia. Habitat: marine.

habena, Helcogramma Williams [J. T.] & McCormick [C. J.] 1990:1026, Fig. 6 [Copeia 1990 (no. 4); ref. 18992] About 1/2 kilometer southeast of Diojo Point, Batan Island, 20°27'55"N, 121°57'12"E, Batanes Province, Philippines, depth 3-6 meters. Holotype: USNM 300194. Paratypes: AMS I.29428-001 (2); ANSP 164965 (2); BPBM 32856 (2); USNM 222333 (3), 222347 (80, not 76), 293946 (10), 300193 (12), 300195-96 (16, 21). Additional material: USNM 222339 (2), 222342 (8), 222920 (2), 297034 (1). Type catalog: Springer & Orrell 1996:27 [ref. 22274], Ho & Shao 2011:53 [ref. 31432]. •Valid as Helcogramma habena Williams & McCormick 1990 -- (Shen 1994:19 [ref. 21605]). •Synonym of Helcogramma inclinatum (Fowler 1946) -- (Fricke 1997:446, 579 [ref. 23339]). •Synonym of Helcogramma inclinata (Fowler 1946) -- (Williams & Howe 2003:151 [ref. 27249], Fricke 2009:49 [ref. 30394], Ho & Shao 2011:53 [ref. 31432], Chiang & Chen 2012:58 [ref. 32986], Tashiro & Motomura 2014:106 [ref. 35481]). Current status: Synonym of Helcogramma inclinata (Fowler 1946). Tripterygiidae: Tripterygiinae. Habitat: marine.

haplodactylus, Lepidoblennius Steindachner [F.] 1867:12 [4], Pl. 1 (figs. 1-1a) [Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften v. 55 (1. Abth.); ref. 4214] Fitzroy River, near Rockhampton, Queensland, Australia. Syntypes: (several) NMW 58145 (3), 78557 (2). On p. 4 of separate. •Valid as Lepidoblennius haplodactylus Steindachner 1867 -- (Hardy 1987:254 [ref. 5960], Kuiter 1993:324 [ref. 23929], Fricke 1994:453 [ref. 21827], Gomon et al. 1994:735 [ref. 22532], Fricke 1997:581 [ref. 23339], Johnson1999:747 [ref. 25471], Williams & Fricke 2001:3534 [ref. 26288], Hoese 2006:1525 [ref. 28998], Gomon 2008:688 [ref. 30616], Fricke 2009:51 [ref. 30394], Patzner et al. 2009:448 [ref. 30396]). Current status: Valid as Lepidoblennius haplodactylus Steindachner 1867. Tripterygiidae: Tripterygiinae. Distribution: Australia: Queensland, New South Wales and Victoria. Habitat: marine.

helenae, Ceratobregma Holleman [W.] 1987:175, Fig. 2 [Cybium v. 11 (no. 2); ref. 6747] Shipwreck on northern coast, Christmas Island, Indian Ocean, 10°29'S, 105°40'E, depth 6-10 meters. Holotype: WAM P.26098-012. Paratypes: AMS I.19456-111 (1); USNM 220065 (6), 224336 (1), 280187-90 (1, 1, 3, 3), 280191 (4, not 2, 1 c&s); WAM P.26080-017 (2), P.26090-011 (1), P.26092-021 (5), P.26098-007 (3), P.26110-015 (2), P.26122-011 (1), P.26113-018 (1). Type catalog: Hutchins & Smith 1991:37 [ref. 19335], Springer & Orrell 1996:27 [ref. 22274], Moore et al. 2008:41 [ref. 30117], Ho & Shao 2011:52 [ref. 31432]. •Valid as Ceratobregma helenae Holleman 1987 -- (Fricke 1991:339 [ref. 19545], Fricke 1994:160 [ref. 21827], Shen 1994:5 [ref. 21605], Fricke 1997:133, 563 [ref. 23339], Randall et al. 1997:521 [ref. 25919], Myers 1999:215 [ref. 23965], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Hutchins 2001:41 [ref. 25847], Fricke 2001:4 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:181 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830], Myers & Donaldson 2003:638 [ref. 27495], Randall et al. 2004:24 [ref. 27624], Randall 2005:471 [ref. 28239], Hoese 2006:1519 [ref. 28998], Fricke 2009:36 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Ho & Shao 2011:52 [ref. 31432], Allen & Erdmann 2012:765 [ref. 31980], Larson et al. 2013:177 [ref. 32988], Fricke et al. 2014:145 [ref. 33932]). Current status: Valid as Ceratobregma helenae Holleman 1987. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: Christmas Island, Malaysia, east to Samoa and Tonga, south to Australia and New Caledonia, north to Ryukyu Islands. Habitat: marine.

hemimelas, Tripterygium Kner [R.] & Steindachner [F.] 1867:371 [16] [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 54 (pt 3); ref. 2640] Fagasa Bay, Tutuila Island, American Samoa, depth 0-3 meters. Neotype: USNM 220068. Type catalog: Springer & Orrell 1996:27 [ref. 22274]. Original genus should have been Tripterygion. Neotype selected by Fricke 1994 [ref. 21827]. •Valid as Enneapterygius hemimelas (Kner & Steindachner 1867) -- (Yoshino in Masuda et al. 1984:294 [ref. 6441], Fricke 1987:335 [ref. 9108], Fricke 1994:215 [ref. 21827], Fricke 1997:214, 567 [ref. 23339], Randall et al. 1997:522 [ref. 25919], Myers 1999:215 [ref. 23965], Johnson 1999:747 [ref. 25471], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1086 [ref. 25182], Allen 2000:116 [ref. 25868], Hutchins 2001:41 [ref. 25847], Fricke 2001:4, 5 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288] dated 1866, Nakabo 2002:1086 [ref. 26193], Fricke 2002:184 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830], Myers & Donaldson 2003:638 [ref. 27495], Randall et al. 2004:24 [ref. 27624], Randall 2005:472 [ref. 28239], Hoese 2006:1520 [ref. 28998], Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Meguro & Motomura 2010:1 [ref. 31252], Motomura et al. 2010:182, 183 [ref. 31256], Allen & Erdmann 2012:767 [ref. 31980], Larson et al. 2013:178 [ref. 32988], Fricke et al. 2014:146 [ref. 33932], Fricke & Erdmann 2017:22, 25, 26 [ref. 35129]). Current status: Valid as Enneapterygius hemimelas (Kner & Steindachner 1867). Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Indonesia, east to Philippines and to American Samoa, north to Taiwan and Japan, south to Australia, Vanuatu, New Caledonia and Tonga. Habitat: marine.

hollemani, Enneapterygius Randall [J. E.] 1995:27, Fig. 1 [Revue française d'Aquariologie Herpétologie v. 22 (no. 1-2); ref. 21853] Northern side of Ra's al Madrakah, central coast of Oman, Arabian Sea, depth 0-0.5 meters. Holotype: BPBM 36078. Paratypes: BPBM 36021 (1), USNM 334523 (1). Type catalog: Springer & Orrell 1996:27 [ref. 22274]. •Valid as Enneapterygius hollemani Randall 1995 -- (Fricke 1997:568 [ref. 23339], Randall 1995:308 [ref. 22896], Manilo & Bogorodsky 2003:S116 [ref. 27377], Holleman 2005:13 [ref. 28455], Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396]). Current status: Valid as Enneapterygius hollemani Randall 1995. Tripterygiidae: Tripterygiinae. Distribution: Northwestern Indian Ocean: Oman. Habitat: marine.

howensis, Enneapterygius Fricke [R.] 1997:224, Fig. 42 [ ref. 23339] South end of middle beach, Lord Howe Island, 31°32'S, 159°04'E, depth 0-3 meters. Holotype: AMS I.17368-036. Paratypes: AMS I.5916 (2), I.17368-053 (10); SMNS 19121 (2). Type catalog: Fricke 2005:65 [ref. 29864]. AMS I.5916 are two syntypes of Tripterygion rufopileum Waite 1904. •Valid as Enneapterygius howensis Fricke 1997 -- (Fricke 1997:568 [ref. 23339], Fricke 2001:4 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:183 [ref. 29865], Hoese 2006:1521 [ref. 28998], Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Fricke & Erdmann 2017:22, 24 [ref. 35129], Tashiro & Motomura 2018:[8] 259 [ref. 35918]). Current status: Valid as Enneapterygius howensis Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: western Coral Sea (Australia), Lord Howe Island and New Caledonia. Habitat: marine.

hsiojenae, Enneapterygius Shen [S.-C.] 1994:11, Fig. 7 [Acta Zoologica Taiwanica v. 5 (no. 2); ref. 21605] Wen-tz-ken [Wen-zi-keng], Gongliao Township, Taipei County, north coast of Taiwan. Holotype: NTUM 07828. Paratypes: NTUM 02829 (1), 07826-27 (1, 1), 07849 (1); USNM 329660 (1). Type catalog: Springer & Orrell 1996:27 [ref. 22274], Ho & Shao 2011:52 [ref. 31432]. Authorship is Shen alone, not Shen & Wu. •Synonym of Enneapterygius vexillarius Fowler 1946 -- (Fricke 1997:355, 572 [ref. 23339]). •Valid as Enneapterygius hsiojenae Shen 1994 -- (Chiang & Chen 2008:196 [ref. 29932] with authorship as Shen & Wu, Endo et al. 2010:14 [ref. 30815], Meguro & Motomura 2010:1 [ref. 31252], Ho & Shao 2011:52 [ref. 31432]). Current status: Valid as Enneapterygius hsiojenae Shen 1994. Tripterygiidae: Tripterygiinae. Distribution: Taiwan. Habitat: marine.

hudsoni, Enneapterygius Jordan [D. S.] & Seale [A.] 1906:419, Fig. 101 [Bulletin of the Bureau of Fisheries v. 25 (for 1905); ref. 2497] Apia, Upolu Island, Samoa; or Pago Pago, Tutuila Island, American Samoa. Holotype: USNM 51798 [not CAS-SU 8691]. Type catalog: Böhlke 1953:95 [ref. 12291], Springer & Orrell 1996:27 [ref. 22274]. Holotype not SU 8691 (that specimen is too small); Fricke 1997:578 [ref. 23339] with incorrect type, see Fricke 1997:444 [ref. 23339]; also see Springer & Eschmeyer 1974 [ref. 21415]. •Valid as Helcogramma hudsoni (Jordan & Seale 1906) -- (Yoshino in Masuda et al. 1984:293 [ref. 6441], Hansen 1986:339 [ref. 5810], Fricke 1994:426 [ref. 21827], Fricke 1997:442, 578 [ref. 23339], Myers 1999:217 [ref. 23965], Nakabo 2000:1080 [ref. 25182], Fricke 2001:4 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1080 [ref. 26193], Fricke 2002:193 [ref. 29865], Randall 2005:476 [ref. 28239], Fricke 2009:49 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Koeda et al. 2016:83 [ref. 35333], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma hudsoni (Jordan & Seale 1906). Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Vanuatu to American Samoa. Habitat: marine.

hystrix, Acanthanectes Holleman [W.] & Buxton [C. D.] 1993:332, Figs. 5, 6B [Cybium v. 17 (no. 4); ref. 21100] Soutrivier, 34°01'42"S, 23°54'19"E, Tsitsikamma National Park, South Africa, depth 24 meters. Holotype: SAIAB [formerly RUSI] 36688. Paratypes: AMS I.33646-001 [ex SAIAB [formerly SAIAB] 36696] (1); NMNZ P.29420 (1); SAIAB [formerly RUSI] 36691-95 (3, 1, 2, 3, 1), 36697 (3), 36699 (1), 37318 (1), 41483 (1); USNM 326000 (2). Type catalog: Springer & Orrell 1996:27 [ref. 22274]. •Valid as Acanthanectes hystrix Holleman & Buxton 1993 -- (Fricke 1997:561 [ref. 23339], Heemstra & Heemstra 2004:387 [ref. 28072], Fricke 2009:32 [ref. 30394], Patzner et al. 2009:443 [ref. 30396]). Current status: Valid as Acanthanectes hystrix Holleman & Buxton 1993. Tripterygiidae: Tripterygiinae. Distribution: South Africa. Habitat: marine.

incisa, Norfolkia Kuiter [R. H.] 1986:93, Figs. 6-7 [Revue française d'Aquariologie Herpétologie v. 12 (no. 3) (1985); ref. 6026] Portsea, Port Phillip Bay, Victoria, Australia, depth 3 meters. Holotype: NMV A3349. Paratypes: AMS I.24789-001 (2); NMV A3350-51 (3, 1); SAMA F4834 (5); WAM P.26005-010 (7), P.27123-012 (4). Type catalog: Hutchins & Smith 1991:37 [ref. 19335], Moore et al. 2008:41 [ref. 30117]. •Valid as Norfolkia incisa Kuiter 1986 -- (Kuiter 1993:322 [ref. 23929], Gomon et al. 1994:739 [ref. 22532]). •Valid as Trinorfolkia incisa (Kuiter 1986) -- (Fricke 1994:549 [ref. 21827], Fricke 1997:586 [ref. 23339], Hutchins 2001:42 [ref. 25847], Hoese 2006:1528 [ref. 28998], Gomon 2008:693 [ref. 30616], Fricke 2009:35 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Hoschke et al. 2019:159 [ref. 36626]). Current status: Valid as Trinorfolkia incisa (Kuiter 1986). Tripterygiidae: Tripterygiinae. Distribution: Southeastern Indian Ocean: Victoria, South Australia and Western Australia. Habitat: marine.

inclinatus, Enneapterygius Fowler [H. W.] 1946:190, Fig. 55 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 98; ref. 1456] Aguni-jima, Ryukyu Islands, Japan. Holotype (unique): ANSP 72067. Type catalog: Böhlke 1984:164 [ref. 13621]. Helcogramma is feminine, based on the lateral line. The spelling inclinata is correct. •Synonym of Helcogramma hudsoni (Jordan & Seale 1906) -- (Hansen 1986:339 [ref. 5810]). •Valid as Helcogramma inclinatum (Fowler 1946) -- (Fricke 1997:446, 579 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288]). •Valid as Helcogramma inclinata (Fowler 1946) -- (Williams & Howe 2003:151 [ref. 27249], Randall 2005:476 [ref. 28239], Motomura et al. 2006:106 [ref. 29490], Fricke 2009:49 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Motomura et al. 2010:187 [ref. 31256], Ho & Shao 2011:53 [ref. 31432], Tashiro & Motomura 2014:106 [ref. 35481], Motomura et al. 2015:28 [ref. 34709], Koeda et al. 2016:84 [ref. 35333], Koeda et al. 2016:14 [ref. 35354], Fricke & Erdmann 2017:41 [ref. 35190]). Current status: Valid as Helcogramma inclinata (Fowler 1946). Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Philippines, Taiwan and southern Japan. Habitat: marine.

indicus, Helcogramma Talwar [P. K.] & Sen [T. K.] 1971:248, Fig. 1 [Records of the Zoological Survey of India v. 65 (1-4) [for 1967]; ref. 21431] Cape Comorin, Tamil Nadu, India. Holotype: ZSI F6074/2. Paratypes: ZSI F6075/2 (3). See Das 2003:371 [ref. 27706]. •Synonym of Helcogramma ellioti (Herre 1944) -- (Hansen 1986:335 [ref. 5810], Fricke 1997:578 [ref. 23339]). Current status: Synonym of Helcogramma ellioti (Herre 1944). Tripterygiidae: Tripterygiinae. Habitat: marine.

ishigakiensis, Lepidoblennius marmoratus Aoyagi [H.] 1954:241 [242], Fig. 3 (not 4) [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 63 (no. 6); ref. 12637] Ishigaki Island, Ishigaki City, Okinawa Prefecture, Yaeyama Islands, southern Ryukyu Islands, Japan. Holotype (unique): YCM-P 31283. Type catalog: Hayashi 1995 [ref. 22717], Shibata et al. 2011:17 [ref. 31469] as isigakiensis. English on p. 242. The genus Helcogramma is feminine, so spelling is inclinata. •Synonym of Helcogramma inclinata (Fowler 1946) -- (Fricke 1997:446, 579 [ref. 23339] as inclinatum, Fricke 2009:49 [ref. 30394]). •Valid as Helcogramma ishigakiensis (Aoyagi 1954) -- (Tashiro & Motomura 2014:98 [ref. 35481]). Current status: Valid as Helcogramma ishigakiensis (Aoyagi 1954). Tripterygiidae: Tripterygiinae. Distribution: Ryukyu Islands, Japan. Habitat: marine.

jenningsi, Tripterygium Hutton [F. W.] 1879:339 [Transactions and Proceedings of the New Zealand Institute v. 11 (art. 38); ref. 18332] Ewing Island, Auckland Island, New Zealand. Lectotype: NMNZ P.20546. Paralectotypes: OM A.81.100 (6). Type catalog: Hardy 1990:14 [ref. 18942], Russell 1996:231 [ref. 22562]. Lectotype selected by Fricke & Roberts 1993:11 [ref. 21298]. Original genus should have been Tripterygion. •See Paulin et al. 1989:264 [ref. 24556]. •Synonym of Grahamina capito (Jenyns 1842) -- (Fricke 1994:378 [ref. 21827], Fricke 1997:576 [ref. 23339], Fricke 2009:45 [ref. 30394]). •Synonym of Forsterygion capito (Jenyns 1842) -- (Jawad 2008:2962 [ref. 29958]). Current status: Synonym of Forsterygion capito (Jenyns 1842). Tripterygiidae: Tripterygiinae. Habitat: marine.

jojettae, Cryptichthys Hardy [G. S.] 1987:266, Fig. 5 [Journal of the Royal Society of New Zealand v. 17 (no. 3); ref. 5960] Mt. Maunganui, New Zealand. Holotype: NMNZ P.18212. Paratypes: NMNZ P.14320 (5), P.14931 (5), P.16377 (6), P.16383-84 (1, 5), P.17071 (6), P.17257 (3), P.18324 (10), P.18367-68 (1, 4), P.18635 (4). Type catalog: Hardy 1990:12 [ref. 18942]. •Valid as Cryptichthys jojettae Hardy 1987 -- (Paulin et al. 1989:220 [ref. 24556], Paulin & Roberts 1992:75 [ref. 24625], Fricke 1994:168 [ref. 21827], Fricke 1997:564 [ref. 23339], Fricke 2009:52 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1501 [ref. 34299]). Current status: Valid as Cryptichthys jojettae Hardy 1987. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

jordani, Gillias Evermann [B. W.] & Marsh [M. C.] 1899:357 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 25 (for 1899) Appendix 7; ref. 1283] Cardona Light-House Reef, Ponce, Puerto Rico. Holotype: USNM 49368. Paratypes: USNM 126096 [ex USFC 840] (1). Type catalog: Springer & Orrell 1996:27 [ref. 22274]. •Valid as Gillias jordani Evermann & Marsh 1899 -- (Fricke 1997:575 [ref. 23339]). •Valid as Enneanectes jordani (Evermann & Marsh 1899) -- (Greenfield & Johnson 1981:50 [ref. 5580], Smith & Williams 2002:9 [ref. 26427], Williams 2003:1749 [ref. 27109], Smith et al. 2003:47 [ref. 27621], Nelson et al. 2004:161 [ref. 27807], McEachran & Fechhelm 2005:576 [ref. 28815], Fricke 2009:37 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708] Victor 2013:69 [ref. 32783], Victor 2017:50 [ref. 35549], Victor 2019:12 [ref. 36450]). Current status: Valid as Enneanectes jordani (Evermann & Marsh 1899). Tripterygiidae: Tripterygiinae. Distribution: Western Atlantic: Florida, Bahamas, Caribbean. Habitat: marine.

kermadecensis, Enneapterygius Fricke [R.] 1994:230, Fig. 42 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Meyer Island, 29°14.9'S, 177°52.2'E, Kermadec Islands, depth 0-2 meters. Holotype: NMNZ P.28611. Paratypes: AMS uncat. (5, not found); BMNH uncat. (5); NMNZ P.13462 (3), P.13498 (1), P.13563 (11), P.17746 (14), P.17749 (8), P.17764 (2), P.28563 (22), P.30185 (13); SMNS 24581 (5); USNM uncat. (5, not received at USNM by 1996). Type catalog: Fricke 1995:20 [ref. 28100], Springer & Orrell 1996:27 [ref. 22274], Fricke 2005:65 [ref. 29864]. •Valid as Enneapterygius kermadecensis Fricke 1994 -- (Fricke 1997:227, 568 [ref. 23339] with note (p. 230) that AMS, BMNH, SMNS and USNM specimens are still with NMNZ P.30185, Fricke 2009:39 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Duffy & Ahyong 2015:117 [ref. 36167], Stewart & Clements 2015:1502 [ref. 34299], Fricke & Erdmann 2017:24 [ref. 35129]). Current status: Valid as Enneapterygius kermadecensis Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: Kermadec Islands endemic. Habitat: marine.

kosiensis, Enneapterygius Holleman [W.] 2005:13, Fig. 12 [Smithiana, Publications in Aquatic Biodiversity, Bulletin No. 5; ref. 28455] Sodwana Bay, KwaZulu-Natal, South Africa, southwestern Indian Ocean. Holotype: SAIAB 59086. Paratypes: SAIAB 9897 (1), 40265 (9). •Valid as Enneapterygius kosiensis Holleman 2005 -- (Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396]). Current status: Valid as Enneapterygius kosiensis Holleman 2005. Tripterygiidae: Tripterygiinae. Distribution: Western Indian Ocean: South Africa. Habitat: marine.

kranos, Helcogramma Fricke [R.] 1997:452, Fig. 92 [ ref. 23339] North of Komodo Island, Tawata Island, northwestern side, Nusa Tenggara Timur Province, Indonesia, depth 1-2 meters. Holotype: BPBM 36503. Paratypes: BPBM uncat. (2), SMNS 18863 (1). Type catalog: Fricke 2005:68 [ref. 29864]. •Valid as Helcogramma kranos Fricke 1997 -- (Williams & Fricke 2001:3534 [ref. 26288], Allen & Adrim 2003:53 [ref. 26830], Fricke 2009:49 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:775 [ref. 31980], Fricke & Erdmann 2017:43 [ref. 35190]). Current status: Valid as Helcogramma kranos Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Indonesia: Lombok and Komodo Islands. Habitat: marine.

kulbickii, Gracilopterygion Fricke [R.] & Randall [J. E.] in Fricke 1994:367, Fig. 77 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Outside barrier reef off Tenia Island, 22°01'06"S, 165°55'30"E, New Caledonia, depth 2-4 meters. Holotype: BPBM 34310. Paratypes: AMS I.18354-051 (1), I.19473-229 (3), I.22616-057 (1), I.30465-106 (2), I.33703-073 (2); BPBM 17536 (1), 36419 (2); ROM 47943 (2), 65520 (1); USNM ?220066 (18), 236396 (1), 283453 (19), 283456 (1), 283476 (1), 289015 (1), 310626 (3), 310883 (1), 323820 (1), 323924 (6); NMNZ P.25295 (1). Plus additional non-type material. Type catalog: Springer & Orrell 1996:27 [ref. 22274]. •Valid as Springerichthys kulbickii (Fricke & Randall 1994) -- (Fricke 1997:525, 585 [ref. 23339], Randall et al. 1997:523 [ref. 25919], Fricke 2001:4 [ref. 25985], Williams & Fricke 2001:3535 [ref. 26288], Fricke 2002:196 [ref. 29865], Randall et al. 2004:24 [ref. 27624], Randall 2005:478 [ref. 28239], Kim et al. 2005:150 [ref. 28305], Hoese 2006:1527 [ref. 28998], Fricke 2009:43 [ref. 30394], Patzner et al. 2009:448 [ref. 30396]). Current status: Valid as Springerichthys kulbickii (Fricke & Randall 1994). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific. Habitat: marine.

lacuna, Helcogramma Williams [J. T.] & Howe [J. C.] 2003:165, Figs. 13-14 [aqua, Journal of Ichthyology and Aquatic Biology v. 7 (no. 4); ref. 27249] Similan Island, Goh Huyong, Thailand, Andaman Sea, 8°29'N, 97°39'E, depth 1-4 meters. Holotype: USNM 222902. Paratypes: BMNH 1983.3.25.1107 (1), USNM 329750 (2), WAM P.26507-016 (2). Type catalog: Moore et al. 2008:41 [ref. 30117]. •Valid as Helcogramma lacuna Williams & Howe 2003 -- (Fricke 2009:49 [ref. 30394], Imamura 2009:244 [ref. 30434], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:775 [ref. 31980], Fricke & Erdmann 2017:41 [ref. 35190]). Current status: Valid as Helcogramma lacuna Williams & Howe 2003. Tripterygiidae: Tripterygiinae. Distribution: Eastern Indian Ocean: East Andaman Sea coast of Thailand. Habitat: marine.

lairdi, Norfolkia Fowler [H. W.] 1953:264, Fig. 12 [Transactions of the Royal Society of New Zealand v. 81 (pt 2); ref. 1463] Kingston, Norfolk Island, southwestern Pacific. Holotype (unique): ANSP 101773. Type catalog: Böhlke 1984:164 [ref. 13621]. •Synonym of Norfolkia squamiceps (McCulloch & Waite 1916) -- (Fricke 1991:339 [ref. 19545], Holleman 1991:236 [ref. 21794], Fricke 1994:473 [ref. 21827], Fricke 1997:512, 583 [ref. 23339], Randall 2005:478 [ref. 28239], Hoese 2006:1526 [ref. 28998], Fricke 2009:34 [ref. 30394]). Current status: Synonym of Norfolkia squamiceps (McCulloch & Waite 1916). Tripterygiidae: Tripterygiinae. Habitat: marine.

lapillum, Forsterygion Hardy [G. S.] 1989:504, Figs. 1 A, 5 [Journal of Natural History v. 23 (no. 3); ref. 12485] Goose Bay, Kaikoura, New Zealand, depth 0-2 meters. Holotype: NMNZ P.21867. Paratypes: BMNH 1988.3.11.1-7 (7); NMNZ P.13283 (8), P.13305 (15), P.13590 (8), P.13593 (1, c&s), P.14113 (29), P.14398 (23, 5 c&s), P.14978 (20, 4 c&s), P.18135 (orig. 28), P.18335 (13), P.19860 (24), P.21041 (6); USNM 295870 [ex NMNZ P.18135] (6). Type catalog: Hardy 1990:12 [ref. 18942], Springer & Orrell 1996:27-28 [ref. 22274]. •Valid as Forsterygion lapillum Hardy 1989 -- (Paulin & Roberts 1992:73 [ref. 24625], Fricke 1994:320 [ref. 21827], Fricke 1997:573 [ref. 23339], Jawad & Clements 2004:89 [ref. 27725], Hoese 2006:1523 [ref. 28998], Jawad 2008:2970 [ref. 29958], Gomon 2008:686 [ref. 30616], Fricke 2009:45 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1506 [ref. 34299]). Current status: Valid as Forsterygion lapillum Hardy 1989. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand. Habitat: marine.

larsonae, Enneapterygius Fricke [R.] 1994:235, Figs. 44-45 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Northwestern tip of Rosemary Island, Dampier Archipelago, Western Australia. Holotype: NTM S.10809-026. Paratypes: NTM S.10809-035 (24), S.10814-036 (7); SMNS 14215 (2), 14575 (3), 14611 (5), 14657 (3); WAM P.25308-016 [not 014] (1), P.26668-011 (5), P.28416-029 (2), P.29883-004 (1), P.29884-015 (1), P.29887-006 (1). Plus more paratypes at AMS, NTM and WAM. Type catalog: Fricke 1995:20 [ref. 28100], Fricke 2005:65 [ref. 29864], Moore et al. 2008:41 [ref. 30117]. •Valid as Enneapterygius larsonae Fricke 1994 -- (Fricke 1997:230, 568 [ref. 23339], Hutchins 2001:41 [ref. 25847], Williams & Fricke 2001:3534 [ref. 26288], Hoese 2006:1521 [ref. 28998], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Allen & Erdmann 2012:767 [ref. 31980], Larson et al. 2013:178 [ref. 32988], Fricke et al. 2014:146 [ref. 33932], Fricke & Erdmann 2017:26, 27, 30 [ref. 35129], Hoschke et al. 2019:159 [ref. 36626]). Current status: Valid as Enneapterygius larsonae Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Eastern Indian Ocean, western Pacific: Papua New Guinea and Australia. Habitat: marine.

larvata, Helcogramma Fricke [R.] & Randall [J. E.] 1992:6, Figs. 5-6 [Stuttgarter Beiträge zur Naturkunde. Serie A (Biologie). No. 484; ref. 20288] Eastern end of Male Island, northern Male Atoll, Maldives, Indian Ocean, depth 1-1.5 meters. Holotype: BPBM 34518. Paratypes: BPBM 34519-20 (1, 1); SMNS 11577 (1). Type catalog: Fricke 1995:21 [ref. 28100], Fricke 2005:69 [ref. 29864]. •Valid as Helcogramma larvata Fricke & Randall 1992 -- (Fricke 1997:579 [ref. 23339], Holleman 2007:65 [ref. 29195], Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396]). Current status: Valid as Helcogramma larvata Fricke & Randall 1992. Tripterygiidae: Tripterygiinae. Distribution: Western Indian Ocean: Maldives. Habitat: marine.

leeuwin, Norfolkia Fricke [R.] 1994:471, Fig. 106 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] 1 kilometer south of Carnac Island, 32°10'S, 115°40'E, Cockburn Sound, Western Australia. Holotype: AMS I.20229-038. Paratypes: AMS I.20222-012 (3), I.20233-023 (3); WAM P.25308-000 (1). Type catalog: Moore et al. 2008:41 [ref. 30117]. •Valid as Norfolkia leeuwin Fricke 1994 -- (Fricke 1997:582 [ref. 23339], Hutchins 2001:41 [ref. 25847], Hoese 2006:1526 [ref. 28998], Gomon 2008:690 [ref. 30616], Fricke 2009:34 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Hoschke et al. 2019:159 [ref. 36626]). Current status: Valid as Norfolkia leeuwin Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Southeastern Indian Ocean: southern Western Australia. Habitat: marine.

lesleyae, Bellapiscis Hardy [G. S.] 1987:263, Fig. 4 [Journal of the Royal Society of New Zealand v. 17 (no. 3); ref. 5960] Mangati Reef, north Taranaki coast, New Zealand. Holotype: NMNZ P.18665. Paratypes: AMS I.26329-001 (9); NMNZ (173, 23 lots); USNM 280259 (16). Type catalog: Hardy 1990:12 [ref. 18942], Springer & Orrell 1996:28 [ref. 22274]. •Valid as Bellapiscis lesleyae Hardy 1987 -- (Paulin et al. 1989:220 [ref. 24556], Paulin & Roberts 1992:90 [ref. 24625], Fricke 1994:126 [ref. 21827], Fricke 1997:562 [ref. 23339], Fricke 2009:52 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1498 [ref. 34299]). Current status: Valid as Bellapiscis lesleyae Hardy 1987. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

leucopunctatus, Enneapterygius Shen [S.-C.] 1994:12, Fig. 8 [Acta Zoologica Taiwanica v. 5 (no. 2); ref. 21605] Wen-tz-keng [Wen-zi-keng], Gongiliao Township, Taipei County, northern coast of Taiwan. Holotype: NTUM 07823. Paratypes: NTUM 7851-52 (1, 1). Non-types: NTUM 07822 (6). Type catalog: Ho & Shao 2011:52 [ref. 31432]. Authorship is Shen alone, not Shen & Wu. •Synonym of Enneapterygius vexillarius Fowler 1946 -- (Fricke 1997:355, 572 [ref. 23339]). •Valid as Enneapterygius leucopunctatus Shen 1994 -- (Chiang & Chen 2008:196 [ref. 29932] with authorship as Shen & Wu, Endo et al. 2010:9 [ref. 30815], Motomura et al. 2010:183 [ref. 31256], Meguro & Motomura 2010:1 [ref. 31252], Ho & Shao 2011:52 [ref. 31432], Koeda et al. 2016:14 [ref. 35354]). Current status: Valid as Enneapterygius leucopunctatus Shen 1994. Tripterygiidae: Tripterygiinae. Distribution: Western North Pacific: Taiwan and Japan. Habitat: marine.

lucillae, Axoclinus Fowler [H. W.] 1944:289, Fig. 242 [Monographs of the Academy of Natural Sciences of Philadelphia No. 6; ref. 1448] Tide pool at Saboga, Perlas Islands, (Pacific) Panama. Holotype (unique): ANSP 70111. Type catalog: Böhlke 1984:61 [ref. 13621]. Genus misspelled Axochnus in figure caption. •Valid as Enneanectes lucillae (Fowler 1944) -- (De La Cruz Agüero et al. 1997:302 [ref. 24545], Fricke 1997:565 [ref. 23339]). •Valid as Axoclinus lucillae Fowler 1944 -- (Allen & Robertson 1994:219 [ref. 22193], Smith & Williams 2002:8 [ref. 26427], Nelson et al. 2004:161 [ref. 27807], Fricke 2009:35 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708]). Current status: Valid as Axoclinus lucillae Fowler 1944. Tripterygiidae: Tripterygiinae. Distribution: Pacific coast of Panama. Habitat: marine.

macleayanum, Tripterygium Lucas [A. H. S.] 1891:12, Pl. 3 (fig. 4) [Proceedings of the Royal Society of Victoria (n. s.) v. 3; ref. 14501] Near Port Phillip Heads, Victoria, Australia. Holotype (unique): whereabouts unknown. Original genus should have been Tripterygion. •Synonym of Norfolkia clarkei (Morton 1888) -- (Gomon et al. 1994:738 [ref. 22532]). •Synonym of Trinorfolkia clarkei (Morton 1888) -- (Fricke 1994:541 [ref. 21827], Fricke 1997:586 [ref. 23339], Hoese 2006:1527 [ref. 28998], Fricke 2009:35 [ref. 30394]). Current status: Synonym of Trinorfolkia clarkei (Morton 1888). Tripterygiidae: Tripterygiinae. Habitat: marine.

macrobrachium, Enneapterygius Fowler [H. W.] 1946:189, Fig. 53 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 98; ref. 1456] Aguni Shima, Ryukyu Islands, Japan. Holotype (unique): ANSP 72065 [not 72055] (missing). Type catalog: Böhlke 1984:164 [ref. 13621]. •Synonym of Enneapterygius hemimelas (Kner & Steindachner 1867) -- (Fricke 1994:216 [ref. 21827]). • Type lost; questionably the same as Enneapterygius ruricauda Shen 1994 -- (Fricke 1997:310, 571 [ref. 23339]). Current status: Uncertain as Enneapterygius rubicauda Shen 1994. Tripterygiidae: Tripterygiinae. Habitat: marine.

macrops, Enneanectes Rosenblatt [R. H.], Miller [E. C.] & Hastings [P. A.] 2013:366, Fig. 3-4 [Zootaxa 3636 (no. 2); ref. 32572] Ensenada Chalcala, off Punta Chacala, Nayarit, Mexico, depth 12 meters. Holotype: SIO 11-394. Paratypes: many at CAS, LACM, SIO, UW, USNM. •Valid as Enneanectes macrops Rosenblatt, Miller & Hastings 2013 -- (Galván-Villa et al. 2016:148 [ref. 34871]). Current status: Valid as Enneanectes macrops Rosenblatt, Miller & Hastings 2013. Tripterygiidae: Tripterygiinae. Distribution: Mexico and Isla Tres Marias, Mexico. Habitat: marine.

macropus, Entomacrodops Fowler [H. W.] 1944:402, Fig. 25 [Monographs of the Academy of Natural Sciences of Philadelphia No. 6; ref. 1448] Maria Madre Island, Tres Marias Islands [Islas Marias], Mexico. Holotype (unique): ANSP 69993. Type catalog: Böhlke 1984:164 [ref. 13621]. In family Tripterygiidae -- (Böhlke 1984:164 [ref. 13621]). Tripterygiidae. Habitat: marine.

malcolmi, Forsterygion Hardy [G. S.] 1987:48, Fig. 1 [National Museum of New Zealand Records v. 3 (no. 5); ref. 9092] Sugar Loaf Islands, New Plymouth, New Zealand, 39°04'S, 174°01'E, depth 14 meters. Holotype: NMNZ P.21064. Paratypes: BMNH 1987.7.14.1-3 (3); NMNZ (153, 43 lots); USNM 289219 [ex NMNZ P.19927] (4). Type catalog: Hardy 1990:12-13 [ref. 18942], Springer & Orrell 1996:28 [ref. 22274]. •Valid as Forsterygion malcolmi Hardy 1987 -- (Hardy 1989:499 [ref. 12485], Paulin et al. 1989:220 [ref. 24556], Paulin & Roberts 1992:79 [ref. 24625], Fricke 1994:331 [ref. 21827], Fricke 1997:573 [ref. 23339], Jawad & Clements 2004:89 [ref. 27725], Jawad 2008:2973 [ref. 29958], Fricke 2009:45 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1507 [ref. 34299]). Current status: Valid as Forsterygion malcolmi Hardy 1987. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

maldivensis, Helcogramma Fricke [R.] & Randall [J. E.] 1992:9, Pl. 1; Figs. 7-8 [Stuttgarter Beiträge zur Naturkunde. Serie A (Biologie). No. 484; ref. 20288] Kandooma Islands, South Male Atoll, Maldives, Indian Ocean, 3°55'38"N, 74°29'33"E, depth 0.5-1.5 meters. Holotype: SMNS 11468. Paratypes: BMNH 1991.4.15.1 (1); BPBM 18889 (2), 32904 (3), 32976 (1), 34521 (1); CAS 75979 (1); MNHN 1991-0700 (1); NSMT-P 34632 (1); SAIAB [formerly RUSI] 36705 (1); SMNS 11469-73 (4, 3, 1, 1, 1); USNM 316488 (1); ZMB 32033 (1). Type catalog: Fricke 1995:21 [ref. 28100], Springer & Orrell 1996:28 [ref. 22274]. Fricke 2005:69 [ref. 29864]. Specific name apparently should be spelled maldivense. •Valid as Helcogramma maldivensis Fricke & Randall 1992 -- (Fricke 1997:579 [ref. 23339], Holleman 2007:65 [ref. 29195], Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396]). Current status: Valid as Helcogramma maldivensis Fricke & Randall 1992. Tripterygiidae: Tripterygiinae. Distribution: Western Indian Ocean: Maldive Islands. Habitat: marine.

marmoratum, Tripterygium Macleay [W.] 1878:34, Pl. 3 (fig. 2) [Proceedings of the Linnean Society of New South Wales v. 3 (pt 1); ref. 2868] King George Sound, Western Australia. Syntypes: AMS I.16403-001 [ex MAMU F1025] (3). Original genus should have been Tripterygion. •Valid as Lepidoblennius marmoratus (Macleay 1878) -- (Hardy 1987:257 [ref. 5960], Fricke 1994:457 [ref. 21827], Kuiter 1993:324 [ref. 23929], Gomon et al. 1994:736 [ref. 22532], Fricke 1997:582 [ref. 23339], Kuiter 1997:332 [ref. 25488], Hutchins 2001:41 [ref. 25847], Hoese 2006:1525 [ref. 28998], Gomon 2008:689 [ref. 30616], Fricke 2009:52 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Hoschke et al. 2019:159 [ref. 36626]). Current status: Valid as Lepidoblennius marmoratus (Macleay 1878). Tripterygiidae: Tripterygiinae. Distribution: Southeastern Indian Ocean: Western Australia, South Australia. Habitat: marine.

maryannae, Obliquichthys Hardy [G. S.] 1987:53, Figs. 2-4 [National Museum of New Zealand Records v. 3 (no. 5); ref. 9092] Seal Rocks, Sugar Loaf Islands, New Plymouth, New Zealand, 39°04'S, 174°01'E, depth 14 meters. Holotype: NMNZ P.20857. Paratypes: BMNH 1987.7.14.4-7 (4); NMNZ (101, 19 lots); USNM 289220 [ex NMNZ P.18119] (3). Type catalog: Hardy 1990:13 [ref. 18942], Springer & Orrell 1996:28 [ref. 22274]. •Valid as Obliquichthys maryannae Hardy 1987 -- (Paulin et al. 1989:220 [ref. 24556], Fricke 1994:514 [ref. 21827], Fricke 1997:584 [ref. 23339], Jawad & Clements 2004:86 [ref. 27725], Fricke 2009:47 [ref. 30394], Patzner et al. 2009:448 [ref. 30396]). •Valid as Forsterygion maryannae (Hardy 1987) -- (Jawad 2008:2974 [ref. 29958], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1508 [ref. 34299]). Current status: Valid as Forsterygion maryannae (Hardy 1987). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand. Habitat: marine.

matador, Enneanectes Victor [B. C.] 2013:49, Figs. 1-9 [Journal of the Ocean Science Foundation v. 7; ref. 32783] Dominica, Roseau (15.28°, -61.38°). Holotype: UF 185600. Paratypes: UF. •Valid as Enneanectes matador Victor 2013 -- (Smith-Vaniz & Jelks 2014:66 [ref. 33341], Victor 2017:68 [ref. 35549], Victor 2019:13 [ref. 36450]). Current status: Valid as Enneanectes matador Victor 2013. Tripterygiidae: Tripterygiinae. Distribution: Western Atlantic: West Caribbean, Greater & Lesser Antilles down to Dominica. Habitat: marine.

medium, Tripterygium Günther [A.] 1861:278 [Catalogue of the fishes in the British Museum v. 3; ref. 1964] New Zealand. Lectotype: BMNH 1855.9.19.854. Paralectotypes: BMNH 1853.2.14.15 (1), 1961.10.20.2-7 [ex 1855.9.19.1487-1489] (6). Lectotype selected by Hardy 1987:263 [ref. 5960]. Original genus should have been Tripterygion. •Valid as Bellapiscis medius (Günther 1861) -- (Hardy 1987:260 [ref. 5960], Paulin et al. 1989:220 [ref. 24556], Paulin & Roberts 1992:88 [ref. 24625], Fricke 1994:132 [ref. 21827], Fricke 1997:562 [ref. 23339], Fricke 2009:52 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1499 [ref. 34299]). Current status: Valid as Bellapiscis medius (Günther 1861). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

melaenocephalus, Tripterygion Cocco [A.] 1829:141 [Giornale di Scienze Lettere e Arti per La Sicilia Anno 7, v. 26 (no. 77); ref. 857] Messina, Sicily, Italy, Mediterranean Sea. No types known. •Synonym of Tripterygion tripteronotus (Risso 1810) -- (Wheeler 1973:531 [ref. 7190], Fricke 1997:587 [ref. 23339] as melanocephalus). •Valid as Tripterygion tripteronotum (Risso 1810) -- (Fricke 2009:44 [ref. 30394]). Current status: Synonym of Tripterygion tripteronotum (Risso 1810). Tripterygiidae: Tripterygiinae. Habitat: marine.

melanolancea, Helcogramma Tashiro [S.] & Motomura [H.] 2018:[2], Figs. [Ichthyological Research v. 66 (no. 1); ref. 36174] Holotype. Paratypes: •Valid as Helcogramma melanolancea Tashiro & Motomura 2018. Current status: Valid as Helcogramma melanolancea Tashiro & Motomura 2018. Tripterygiidae: Tripterygiinae. Distribution: Bali, Indonesia. Habitat: marine.

melanospilus, Enneapterygius Randall [J. E.] 1995:29, Fig. 2 [Revue française d'Aquariologie Herpétologie v. 22 (no. 1-2); ref. 21853] East side of Masirah Island, off central coast of Oman, Arabian Sea, depth 3 meters. Holotype: BPBM 36181. Paratypes: BPBM 35980 (1), USNM 334524 (1). Type catalog: Springer & Orrell 1996:28 [ref. 22274]. •Valid as Enneapterygius melanospilus Randall 1995 -- (Fricke 1997:568 [ref. 23339], Randall 1995:308 [ref. 22896], Manilo & Bogorodsky 2003:S116 [ref. 27377], Holleman 2005:14 [ref. 28455] as melanospilos, Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396]). Current status: Valid as Enneapterygius melanospilus Randall 1995. Tripterygiidae: Tripterygiinae. Distribution: Northwestern Indian Ocean: Oman. Habitat: marine.

melanurus, Tripterygion Guichenot [A.] 1850:75, Pl. 4 (fig. 4) [Histoire naturelle des reptiles et poissons; ref. 1940] Algeria, western Mediterranean Sea. Syntypes: MNHN A-1415 (2). Type catalog: Bauchot 1967:56-57 [ref. 20734]. Gender of genus neuter, therefore correct species name melanurum. •Synonym of Tripterygion tripteronotus Risso 1810 -- (Wheeler 1973:531 [ref. 7190]). •Valid as Tripterygion melanurus Guichenot 1850 -- (Zander in Whitehead et al. 1986:1120 [ref. 13677], Ahnelt 1991:56 [ref. 19120], Fricke 1997:587 [ref. 23339], Geertjes et al. 2001:256 [ref. 25579], Bilecenoğlu et al. 2002:120 [ref. 26753], Carreras-Carbonell et al. 2007:76 [ref. 29025] dated 1845, Fricke et al. 2007:96 [ref. 29533], Patzner et al. 2009:449 [ref. 30396], Hastings & Springer 2009:11 [ref. 31156], Lipej & Dulčić 2010:57 [ref. 36649], Falzon & Falzon 2013:582 [ref. 36181]). •Valid as Tripterygion melanurum Guichenot 1850 -- (Fricke 2009:44 [ref. 30394]). Current status: Valid as Tripterygion melanurum Guichenot 1850. Tripterygiidae: Tripterygiinae. Distribution: Mediterranean Sea endemic. Habitat: marine.

microstigma, Helcogramma Holleman [W.] 2006:92, Figs. 1-2 [aqua, Journal of Ichthyology and Aquatic Biology v. 11 (no. 3); ref. 28864] Off Bazaruto lighthouse, 21°30'S, 35°30'E, Bazaruto Island, Mozambique, depth 9-10 meters. Holotype: SAIAB 73754. Paratypes: ROM 73409 (5), 7310 (1), 76683 (2); SAIAB 73573 (13); USNM 381379 (2). In error USNM 307834 (1). Plus non-type specimens. USNM 307834 was wrongly listed as a paratype of this species; it is a paratype of Helcograma rosea •Valid as Helcogramma microstigma Holleman 2006 -- (Holleman 2007:66 [ref. 29195], Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Fricke & Erdmann 2017:40 [ref. 35190], Fricke et al. 2018:305 [ref. 35805]). Current status: Valid as Helcogramma microstigma Holleman 2006. Tripterygiidae: Tripterygiinae. Distribution: Mozambique to Comoros and Madagascar. Habitat: marine.

minor, Tripterygium Kolombatović [G.] 1892:24 [Godicnje Izviesće C. K. Velike Realke u Splitu za Skolsku Godinu, Split v. 1891-1892; ref. 27542] Dalmatia, Adriatic Sea, Mediterranean Sea. No types known. Original genus should have been Tripterygion. Also appeared in Kolombatović 1904:190[9] [ref. 20233]. Apparently dates to 1892 as above; see footnote on p. 531 in Wheeler 1973 [ref. 7190]. Also in Kolombatović 1894:47 [ref. 18431]. •Synonym of Tripterygion melanurus Guichenot 1850, but a valid subspecies -- (Zander in Whitehead et al. 1986:1120 [ref. 13677], Hastings & Springer 2009:11 [ref. 31156]). •Synonym of Tripterygion melanurus Guichenot 1850 -- (Fricke 1997:587 [ref. 23339]). •Valid as Tripterygion minor Kolombatović 1892 -- (Wheeler 1973:531 [ref. 7190], Fricke et al. 2007:96 [ref. 29533], Fricke 2009:44 [ref. 30394], Patzner et al. 2009:449 [ref. 30396]). Current status: Valid as Tripterygion minor Kolombatović 1892. Tripterygiidae: Tripterygiinae. Distribution: Mediteranian Sea. Habitat: marine.

minutum, Tripterygium Günther [A.] 1877:211, Pl. 118 (fig. D) [Journal des Museum Godeffroy v. 4 (no. 13); ref. 14243] Apia, Upolu Island, Samoa. Lectotype: 1961.10.20.1. Paralectotypes: BMNH 1873.4.3.218 (1). Lectotype selected by Fricke 1994:245 [ref. 21827]. Original genus should have been Tripterygion. •Valid as Enneapterygius minutus (Günther 1877) -- (Yoshino in Masuda et al. 1984:294 [ref. 6441], Fricke 1987:336 [ref. 9108], Fricke 1994:242 [ref. 21827], Shen 1994:14 [ref. 21605], Fricke 1994:4 [ref. 21828], Myers 1999:216 [ref. 23965], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1081 [ref. 25182], Allen 2000:116 [ref. 25868]). •Synonym of Enneapterygius philippinus (Peters 1868) -- (Fricke 1997:274, 570 [ref. 23339], Randall 2005:473 [ref. 28239], Fricke & Kulbicki 2006:345 [ref. 28867], Hoese 2006:1522 [ref. 28998], Fricke 2009:41 [ref. 30394], Motomura et al. 2010:184 [ref. 31256], Larson et al. 2013:178 [ref. 32988]). Current status: Synonym of Enneapterygius philippinus (Peters 1868). Tripterygiidae: Tripterygiinae. Habitat: marine.

mirabilis, Enneapterygius Fricke [R.] 1994:251, Fig. 48 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Escape Reef, 15°49'S, 145°50'E, Cooktown area, Queensland, Australia, depth 10-14 meters. Holotype: AMS I.22581-063. Paratypes: AMS I.20779-169 (2), I.20784-002 (1), I.20956-049 (1), I.20965-022 (1), I.22580-026 (1); USNM 263652 (1); WAM P.28531-012 (1), P.28534-024 (1). Type catalog: Springer & Orrell 1996:28 [ref. 22274], Moore et al. 2008:41 [ref. 30117]. •Valid as Enneapterygius mirabilis Fricke 1994 -- (Fricke 1997:234, 568 [ref. 23339], Shimojyo & Hayashi 2000:41 [ref. 24903], Nakabo 2000:1082 [ref. 25182], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1082 [ref. 26193], Allen & Adrim 2003:53 [ref. 26830], Hoese 2006:1521 [ref. 28998], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Allen & Erdmann 2012:768 [ref. 31980], Fricke & Erdmann 2017:23 [ref. 35129], Tashiro et al. 2018:[6] [ref. 35790]). Current status: Valid as Enneapterygius mirabilis Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: Indonesia (incl. West Papua), Papua New Guinea, Solomon Islands, Vanuatu and eastern Australia. Habitat: marine.

miyakensis, Enneapterygius Fricke [R.] 1987:333, Fig. 1 [Senckenbergiana Maritima v. 19 (nos. 5-6); ref. 9108] Igaya Bay, Miyake-jima, Izu Islands, Japan, 34°05'N, 139°31'E, depth 0-1 meter. Holotype: SMF 21361. Paratypes: SMF 21362 (1), NSMT-P 29837 [ex TMBS 850808-1] (1), ZMB 31655 (1). Type catalog: Moyer & Matsuura 1991:82 [ref. 19927]. •Valid as Enneapterygius miyakensis Fricke 1987 -- (Fricke 1997:237, 568 [ref. 23339], Nakabo 2000:1083 [ref. 25182], Nakabo 2002:1083 [ref. 26193], Motomura et al. 2005:13 [ref. 28338], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Endo et al. 2010:14 [ref. 30815], Fricke & Erdmann 2017:27 [ref. 35129]). Current status: Valid as Enneapterygius miyakensis Fricke 1987. Tripterygiidae: Tripterygiinae. Distribution: Western North Pacific: Izu Islands, Japan. Habitat: marine.

mortenseni, Enneapterygius Rendahl [H.] 1926:11, Figs. 3-4 [Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, Kjøbenhavn. v. 81; ref. 15819] Queen Charlotte Sound, New Zealand. Lectotype: ZMUC P76951. Paralectotypes: ?NRM 10262 (1); ZMUC P76950 (1), P76952-54 (3). Type catalog: Nielsen 1974:74 [ref. 9588]. Lectotype selected by Hardy 1987:170 [ref. 6027]. •Synonym of Notoclinops segmentatus (McCulloch & Phillipps 1923) -- (Hardy 1987:262 [ref. 5960], Hardy 1987:167 [ref. 6027], Fricke 1994:487 [ref. 21827] in part, Fricke 1997:584 [ref. 23339] in part, Fricke 2009:45 [ref. 30394] in part). •Synonym of Forsterygion capito (Jenyns 1842) -- (Jawad 2008:2962 [ref. 29958]). Current status: Synonym of Forsterygion capito (Jenyns 1842). Tripterygiidae: Tripterygiinae. Habitat: marine.

multicinctus, Axoclinus Allen [G. R.] & Robertson [D. R.] 1992:54, Fig. 3 [Revue française d'Aquariologie Herpétologie v. 19 (nos 1-2); ref. 19713] Socorro Island, Revillagigedo Islands, off western Mexico, about 18°48'N, 111°02'W, depth 8 meters. Holotype: USNM 321176. Paratypes: USNM 321177 (2). Type catalog: Springer & Orrell 1996:28 [ref. 22274]. •Valid as Enneanectes multicinctus (Allen & Robertson 1992) -- (Fricke 1997:565 [ref. 23339]). •Valid as Axoclinus multicinctus Allen & Robertson 1992 -- (Allen & Robertson 1994:220 [ref. 22193], Smith & Williams 2002:8 [ref. 26427], Nelson et al. 2004:161 [ref. 27807], Fricke 2009:35 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708], Fourriére et al. 2016:449 [ref. 34668], Del Moral-Flores et al. 2016:610 [ref. 34923]). Current status: Valid as Axoclinus multicinctus Allen & Robertson 1992. Tripterygiidae: Tripterygiinae. Distribution: Eastern Pacific: Revillagigedo Islands. Habitat: marine.

multiradiatum, Forsterygion Scott [E. O. G.] 1977:153, Fig. 2 [Papers and Proceedings of the Royal Society of Tasmania v. 111; ref. 7283] Kangaroo Bay, Bellerive, southeastern Tasmania, Australia, depth 2 fathoms. Holotype: QVM 1976:5:0202 [QVM type no. 230]. Paratypes: QVM 1976/5/203 [ex QVM type 231] (1). Spelled Forsterygion multistriatum in abstract. •Synonym of Forsterygion varium (Forster 1801) -- (Hardy 1989:494 [ref. 12485], Fricke 1994:343 [ref. 21827], Fricke 1997:574 [ref. 23339], Hoese 2006:1523 [ref. 28998], Jawad 2008:2959 [ref. 29958], Fricke 2009:45 [ref. 30394]). Current status: Synonym of Forsterygion varium (Forster 1801). Tripterygiidae: Tripterygiinae. Habitat: marine.

namarrgon, Enneapterygius Fricke [R.] 1997:240, Fig. 48 [ ref. 23339] Gove Peninsula, south of Cape Arnhem, Northern Territory, Australia, depth 5 meters. Holotype: AMS I.21961-014. Paratypes: AMS I.21961-028 (15), SMNS 19119 (2). Type catalog: Fricke 2005:65 [ref. 29864]. •Valid as Enneapterygius namarrgon Fricke 1997 -- (Williams & Fricke 2001:3534 [ref. 26288], Hoese 2006:1521 [ref. 28998], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Meguro & Motomura 2010:6 [ref. 31252], Larson et al. 2013:178 [ref. 32988], Fricke & Erdmann 2017:28 [ref. 35129]). Current status: Valid as Enneapterygius namarrgon Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Northern Territory, Australia. Habitat: marine.

nanus, Tripterygion Schultz [L. P.] in Schultz et al. 1960:288, Fig. 112 [Bulletin of the United States National Museum No. 202, v. 2; ref. 3972] Lagoon reef at Rujoru Island, Eniwetok [Eniwetak] Atoll, western Pacific. Holotype: USNM 142233. Paratypes: USNM 142234-48 (7, 4, 2, 2, 2, 3, 1, 1, 4, 1, 4, 2, 1, 4, 1). Type catalog: Springer & Orrell 1996:28 [ref. 22274]. •Valid as Enneapterygius nanus (Schultz 1960) -- (Fricke 1994:255 [ref. 21827], Shen 1994:16 [ref. 21605], Fricke 1997:242, 569 [ref. 23339], Myers 1999:216 [ref. 23965], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Hutchins 2001:41 [ref. 25847], Fricke 2001:4, 5 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:185 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830], Myers & Donaldson 2003:638 [ref. 27495], Randall et al. 2004:24 [ref. 27624], Randall 2005:472 [ref. 28239], Chiang & Chen 2008:197 [ref. 29932], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Allen & Erdmann 2012:768 [ref. 31980], Fricke & Erdmann 2017:23, 28 [ref. 35129]). Current status: Valid as Enneapterygius nanus (Schultz 1960). Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: East Andaman Sea to Micronesia, New Caledonia, Australia to Taiwan. Habitat: marine.

nasimae, Enneapterygius Hoda [S. M. S.] 1983:116, Figs. 1 (A-D), 2 (A-D), 3 (A-C) [Indian Journal of Fisheries v. 30 (no. 1); ref. 17430] Buleji, Karachi coast, Pakistan, northern Arabian Sea. Holotype: CEMBP 208. Paratypes: (?11) CEMBP 69, 238, 262; ROM 39315. •Valid as Enneapterygius nasimae Hoda 1983 -- (Fricke 1997:569 [ref. 23339], Manilo & Bogorodsky 2003:S116 [ref. 27377]). •Synonym of Enneapterygius ventermaculus Holleman 1982 -- (Holleman 2005:21 [ref. 28455], Holleman & Bogorodsky 2012:53 [ref. 31735]). Current status: Synonym of Enneapterygius ventermaculus Holleman 1982. Tripterygiidae: Tripterygiinae. Habitat: marine.

nasus, Tripterygion Risso [A.] 1827:241 [Histoire naturelle des principales productions de l'Europe méridionale v. 3; ref. 3757] Nice, France, northwestern Mediterranean Sea. No types known. Type catalog: Bauchot 1967:57 [ref. 20734]. •Synonym of Tripterygion tripteronotus (Risso 1810) (sometimes correctly as tripteronotum) -- (Wheeler 1973:531 [ref. 7190], Zander in Whitehead et al. 1986:1120 [ref. 13677], Fricke 1997:587 [ref. 23339], Parin 2003:S23 [ref. 28536], Fricke 2009:44 [ref. 30394], Parin et al. 2014:467 [ref. 33547]). •Synonym of Tripterygion tripteronotum -- (Fricke 2009:44 [ref. 30394]). Current status: Synonym of Tripterygion tripteronotum (Risso 1810). Tripterygiidae: Tripterygiinae. Habitat: marine.

nesion, Helcogramma Williams [J. T.] & Howe [J. C.] 2003:167, Fig. 15 [aqua, Journal of Ichthyology and Aquatic Biology v. 7 (no. 4); ref. 27249] Miyake-jima, Igaya Bay, Izu Islands, Japan, depth 5-10 meters. Holotype: USNM 329645. Paratypes: BPBM 35294 (8); USNM 222324 (6), 222329-32 (2, 2, 1, 2), 222346 (1). Plus nontype material. •Valid as Helcogramma nesion Williams & Howe 2003 -- (Holleman 2006:102 [ref. 28864], Motomura et al. 2006:106 [ref. 29490], Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma nesion Williams & Howe 2003. Tripterygiidae: Tripterygiinae. Distribution: Southern Japan including Ogasawara Islands. Habitat: marine.

niger, Enneapterygius Fricke [R.] 1994:259, Fig. 51 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Grande Terre, Baie de Saint-Vincent, 21°57'24"S, 165°59'54"E, New Caledonia, depth 2-8 meters. Holotype: MNHN 1997-0430. Paratypes: AMS IA.2954 (9), IB.4779 (1); MNHN uncat. (2, 1); NMNZ P.29472 (1), P.29517 (1), P.29558 (1), P.29587 (8), P.29606 (1), P.29608 (4); ROM 65507-08 (10, 8), 65510 (3), 65514 (2), 65523 (1); SMNS 13207-08 (2, 2); USNM 323779 (14). Type catalog: Fricke 1995:20 [ref. 28100], Springer & Orrell 1996:28 [ref. 22274], Fricke 2005:65 [ref. 29864]. •Valid as Enneapterygius niger Fricke 1994 -- (Fricke 1997:248, 569 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Fricke 2001:4, 5, 6 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:186 [ref. 29865], Shibukawa et al. 2004:6 [ref. 27552], Randall 2005:472 [ref. 28239], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Allen & Erdmann 2012:768 [ref. 31980], Fricke & Erdmann 2017:23, 25, 27, 29 [ref. 35129]). Current status: Valid as Enneapterygius niger Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Philippines, Taiwan, Vanuatu, New Caledonia and Pohnpei. Habitat: marine.

nigra, Helcogramma Williams [J. T.] & Howe [J. C.] 2003:169, Fig. 16-18 [aqua, Journal of Ichthyology and Aquatic Biology v. 7 (no. 4); ref. 27249] Rotuma Island, north coast, ocean side of reef crest, about 100 meters from shore, 12°30'N, 177°05'E, Fiji Islands, depth 1-9 meters. Holotype: USNM 329281. Paratypes: AMS I.35026-001 (1); BPBM 36435 (1); CAS 81762 (1); USNM 283420 (19), 283421 (1). Plus nontype material. •Valid as Helcogramma nigra Williams & Howe 2003 -- (Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:775 [ref. 31980], Fricke et al. 2014:146 [ref. 33932], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma nigra Williams & Howe 2003. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Papua New Guinea, Solomon Islands and Fiji Islands, Vanuatu and Rotuma Islands. Habitat: marine.

nigricauda, Enneapterygius Fricke [R.] 1997:255, Fig. 52 [ ref. 23339] E'ua Island, Tongatapu Group, fringing reef at base of cliff on northwestern shore, 21°18'15"S, 174°26'20"W, depth 0-11 meters. Holotype: USNM 331035. Paratypes: BPBM 15516 (7), 29097 (14), 35222 (2); CAS 49701 (10); SMNS 18697 (4); USNM 115526 (30), 142259 (1), 297859 (27), 331031 (51), 343062 (3), 344093 (36), 344097 (55); plus non-type additional material. Type catalog: Fricke 2005:66 [ref. 29864]. •Valid as Enneapterygius nigricauda Fricke 1997 -- (Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Myers & Donaldson 2003:638 [ref. 27495], Lobel & Lobel 2004:75 [ref. 27576], Randall et al. 2004:24 [ref. 27624], Randall 2005:473 [ref. 28239], Chiang & Chen 2008:197 [ref. 29932], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Meguro & Motomura 2010:6 [ref. 31252], Allen & Erdmann 2012:768 [ref. 31980], Fricke & Erdmann 2017:25 [ref. 35129]). Current status: Valid as Enneapterygius nigricauda Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Philippines, north to Taiwan and Ogasawara Islands, east to Vanuatu, islands of Oceania to Line Islands, Samoa, Tonga and Society Islands. Habitat: marine.

nigricaudus, Axoclinus Allen [G. R.] & Robertson [D. R.] 1991:79, Fig. 1 [Revue française d'Aquariologie Herpétologie v. 18 (no. 3); ref. 19723] Near Pichilingue, Baja California, Mexico, about 24°16'N, 110°20'W, depth 0-3 meters. Holotype: USNM 316797. Paratypes: WAM P.30233-001 (1). Type catalog: Springer & Orrell 1996:28 [ref. 22274], Moore et al. 2008:40 [ref. 30117]. •Valid as Enneanectes nigricaudus (Allen & Robertson 1991) -- (De La Cruz Agüero et al. 1997:302 [ref. 24545], Fricke 1997:565 [ref. 23339]). •Valid as Axoclinus nigricaudus Allen & Robertson 1991 -- (Allen & Robertson 1994:220 [ref. 22193], Thomson et al. 2000:206 [ref. 25640], Smith & Williams 2002:8 [ref. 26427], Nelson et al. 2004:161 [ref. 27807], Fricke 2009:35 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708]). Current status: Valid as Axoclinus nigricaudus Allen & Robertson 1991. Tripterygiidae: Tripterygiinae. Distribution: Eastern Pacific: Gulf of California. Habitat: marine.

nigripenne, Tripterygion Valenciennes [A.] in Cuvier & Valenciennes 1836:413, Pl. 339 [Histoire naturelle des poissons v. 11; ref. 1005] New Zealand. Holotype (unique): MNHN A-2153. Type catalog: Bauchot 1967:57 [ref. 20734]. Gender of Forsterygion is neuter, therefore species name correct as nigripenne. •See Paulin et al. 1989:264 [ref. 24556]. •Valid as Forsterygion nigripenne (Valenciennes 1836) -- (Paulin & Roberts 1992:91 [ref. 24625] with question). •Valid as Grahamina nigripenne (Valenciennes 1836) -- (Fricke 1994:391 [ref. 21827], Fricke 1997:576 [ref. 23339], Clements et al. 2000:377 [ref. 25188], Patzner et al. 2009:446 [ref. 30396]). •Valid as Grahamina nigripennis (Valenciennes 1836) -- (Fricke 2009:46 [ref. 30394]). •Valid as Forsterygion nigripenne (Valenciennes 1836) -- (Jawad 2008:2977 [ref. 29958], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1509 [ref. 34299]). Current status: Valid as Forsterygion nigripenne (Valenciennes 1836). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

nikolskii, Tripterygium Maksimov [N. E.] 1909:59-60 [Trudy Obshchestva estestvoispitatelei pri Imperatorskom, Kharkov Universitet v. 42 (for 1907-1908); ref. 14384] Crimea, Ukraine, Black Sea. Syntypes: Mus. Univ. Kharkov 295 (2). Original genus should have been Tripterygion. •Synonym of Tripterygion tripteronotus (Risso 1810) -- (Fricke 1997:587 [ref. 23339], Parin 2003:S24 [ref. 28536]). •Synonym of Tripterygion tripteronotum (Risso 1810) -- (Fricke 2009:44 [ref. 30394]). Current status: Synonym of Tripterygion tripteronotum (Risso 1810). Tripterygiidae: Tripterygiinae. Habitat: brackish, marine.

niue, Enneapterygius Fricke [R.] & Erdmann [M. V.] 2017:16, Figs. 1-4 [Journal of the Ocean Science Foundation v. 25; ref. 35129] Niue, Limu, 18°58'30.12"S, 169°53'58.9"W, intertidal. Holotype: SMF 35952. Paratypes: HUJ, SMF. •Valid as Enneapterygius niue Fricke & Erdmann 2017. Current status: Valid as Enneapterygius niue Fricke & Erdmann 2017. Tripterygiidae: Tripterygiinae. Distribution: Niue and Samoa, southwestern Pacific Ocean. Habitat: marine.

novaecaledoniae, Helcogramma Fricke [R.] 1994:429, Fig. 93 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Grande Terre, Passe Saint Vincent, 22°02'06"S, 165°58'24"E, New Caledonia, depth 3 meters. Holotype: MNHN 1997-0434. Paratypes: MNHN uncat. (2, 1, 1), SMNS 13912-13 (2, 1). Type catalog: Fricke 1995:21-22 [ref. 28100], Fricke 2005:69 [ref. 29864] •Valid as Helcogramma novaecaledoniae Fricke 1994 -- (Fricke 1997:456, 579 [ref. 23339], Fricke 2001:4 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:193 [ref. 29865], Randall 2005:477 [ref. 28239], Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:776 [ref. 31980], Fricke & Erdmann 2017:43 [ref. 35190]). Current status: Valid as Helcogramma novaecaledoniae Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: Solomon Islands and New Caledonia. Habitat: marine.

obscurus, Enneapterygius Clark [E.] 1980:105, Figs. 4d, 15 [Israel Journal of Zoology v. 28 (nos 3-4) [for 1979]; ref. 8837] El Himera, Gulf of Aqaba, Sinai, Egypt, Red Sea. Holotype: USNM 205818. Paratypes: HUJ 5483 (2), 5486 (8); USNM 205819-20 (2, 9). Type catalog: Springer & Orrell 1996:28 [ref. 22274], Golani 2006:31 [ref. 28840]. •Valid as Enneapterygius obscurus Clark 1980 -- (Dor 1984:234 [ref. 29757], Fricke & Randall 1992:5 [ref. 20288], Goren & Dor 1994:61 [ref. 25356], Fricke 1997:569 [ref. 23339], Holleman 2005:15 [ref. 28455], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Holleman & Bogorodsky 2012:48 [ref. 31735], Bogorodsky & Randall 2018:[6] 244 [ref. 36492], Golani & Fricke 2018:143 [ref. 36273]). Current status: Valid as Enneapterygius obscurus Clark 1980. Tripterygiidae: Tripterygiinae. Distribution: Red Sea, western Indian Ocean: East Africa, Maldives. Habitat: marine.

obtusirostre, Tripterygium Klunzinger [C. B.] 1871:498 [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 21; ref. 2622] Al-Qusair, Red Sea Governorate, Egypt, Red Sea. Lectotype: ZMB 31298. Paralectotypes: ZMB 8020 (2). Lectotype selected by Hansen 1986:343 [ref. 5810]. The correct species spelling is obtusirostris as the genus Helcogramma is feminine. Original genus should have been Tripterygion. •Valid as Helcogramma obtusirostris (Klunzinger 1871) [sometimes as obtusirostre] -- (Dor 1984:234 [ref. 29757], Holleman 1986:757 [ref. 5671], Hansen 1986:341 [ref. 5810], Shen 1994:21 [ref. 21605], Goren & Dor 1994:61 [ref. 25356], Randall 1995:32 [ref. 21853], Randall 1995:309 [ref. 22896], Fricke 1997:461, 579 [ref. 23339], Fricke 1999:469 [ref. 24106], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1080 [ref. 26193] as obtusirostris, Manilo & Bogorodsky 2003:S116 [ref. 27377], Heemstra & Heemstra 2004:388 [ref. 28072], Holleman 2007:68 [ref. 29195], Fricke 2009:50 [ref. 30394], Fricke et al. 2009:93 [ref. 30213], Patzner et al. 2009:447 [ref. 30396] as obtusirostre, Motomura et al. 2010:183 [ref. 31256] as obtusirostre, Holleman & Bogorodsky 2012:36 [ref. 31735] as obtusirostris, Allen & Erdmann 2012:776 [ref. 31980], Chiang & Chen 2012:58 [ref. 32986], Fricke & Erdmann 2017:43 [ref. 35190], Golani & Fricke 2018:143 [ref. 36273], Zajonz et al. 2019:92 [ref. 36871] as cf. obtusirostris). Current status: Valid as Helcogramma obtusirostris (Klunzinger 1871). Tripterygiidae: Tripterygiinae. Distribution: Red Sea, northwestern Indian Ocean: Oman. Habitat: marine.

oculus, Apopterygion Fricke [R.] & Roberts [C. D.] in Fricke 1994:118, Fig. 13 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Entrance to Islet Cove, Port Pegasus, Stewart Island, 47°11.3'S, 167°38.1'E, depth 16 meters. Holotype: NMNZ P.23969. Paratypes: AMS uncat. (3, not present in 1997), BMNH uncat. (3), NMNZ P.7309-10 (1, 8), P.18870 (1); SMNS 24580 (3); USNM uncat. (3); plus many more paratype lots at NMNZ. Type catalog: Fricke 1995:19 [ref. 28100], Fricke 2005:63 [ref. 29864]. Second author is C. D. Roberts. •Valid as Apopterygion oculus Fricke & Roberts 1994 -- (Fricke 1997:561 [ref. 23339], Jawad & Clements 2004:83 [ref. 27725], Fricke 2009:33 [ref. 30394], Patzner et al. 2009:443 [ref. 30396], Stewart & Clements 2015:1497 [ref. 34299]). Current status: Valid as Apopterygion oculus Fricke & Roberts 1994. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

ornatus, Enneapterygius Fricke [R.] 1997:262, Fig. 53 [ ref. 23339] Henderson Island, 1/2 mile south of northwestern corner of island, depth 0-1 meter. Holotype: BPBM 17081. Paratypes: BPBM uncat. (16), SMNS 21881 (2). Type catalog: Fricke 2005:66 [ref. 29864]. •Valid as Enneapterygius ornatus Fricke 1997 -- (Randall 1999:24 [ref. 25829], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Fricke & Erdmann 2017:25 [ref. 35129]). Current status: Valid as Enneapterygius ornatus Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Central Pacific: Henderson Island, Pitcairn Group. Habitat: marine.

pallidoserialis, Enneapterygius Fricke [R.] 1997:264, Figs. 54-55 [ ref. 23339] Putic Island, northwestern side (Cuyo Island), Palawan Province, Philippines, 10°55'05"N, 121°02'03"E, depth 0-4.5 meters. Holotype: USNM 279812. Paratypes: SMNS 19171 (2); USNM 227494 (1), 258845 (89), 259354-55 (1, 3), 287086 (7), 345513 (218); plus other non-type material. Type catalog: Fricke 2005:66 [ref. 29864]. •Valid as Enneapterygius pallidoserialis Fricke 1997 -- (Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Chiang & Chen 2008:197 [ref. 29932], Fricke 2009:40 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Allen & Erdmann 2012:769 [ref. 31980], Fricke & Erdmann 2017:24 [ref. 35129]). Current status: Valid as Enneapterygius pallidoserialis Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Malaysia, Philippines, Taiwan, Japan, Vanuatu and Chuuk. Habitat: marine.

pallidus, Enneapterygius Clark [E.] 1980:107, Figs. 4e, 16 [Israel Journal of Zoology v. 28 (nos 3-4) [for 1979]; ref. 8837] El Himeira, Gulf of Aqaba, Sinai, Egypt, Red Sea, depth 21-17 meters. Holotype: USNM 212156. Paratypes: USNM 205829 (1), 212246 (2). Type catalog: Springer & Orrell 1996:29 [ref. 22274]. •Synonym of Enneapterygius pusillus Rüppell 1835 -- (Fricke 1997:570 [ref. 23339], Fricke 2009:41 [ref. 30394]). •Valid as Enneapterygius pallidus Clark 1980 -- (Dor 1984:234 [ref. 29757], Goren & Dor 1994:61 [ref. 25356], Patzner et al. 2009:445 [ref. 30396] with author as Clarke, Holleman & Bogorodsky 2012:48 [ref. 31735], Bogorodsky & Randall 2018:[6] 244 [ref. 36492], Golani & Fricke 2018:143 [ref. 36273], Zajonz et al. 2019:107 [ref. 36871]). Current status: Valid as Enneapterygius pallidus Clark 1980. Tripterygiidae: Tripterygiinae. Distribution: Red Sea, northwestern Indian Ocean: Socotra. Habitat: marine.

pardochir, Enneapterygius Jordan [D. S.] & Seale [A.] 1906:417, Fig. 98 [Bulletin of the Bureau of Fisheries v. 25 (for 1905); ref. 2497] Apia, Upolu Island, Samoa. Holotype: USNM 51799 (poor condition). Paratypes: (14) MCZ 35910 [ex USNM 51799] (2), CAS-SU 8696 (6), USNM 126235 (now 8). Type catalog: Böhlke 1953:95 [ref. 12291], Springer & Orrell 1996:29 [ref. 22274]. •Questionably a synonym of Enneapterygius minutus (Günther 1877) -- (Fricke 1994:242 [ref. 21827], Fricke 1994:4 [ref. 21828]). •Synonym of Enneapterygius philippinus (Peters 1868) -- (Fricke 1997:274, 570 [ref. 23339], Fricke 2009:41 [ref. 30394]). Current status: Synonym of Enneapterygius philippinus (Peters 1868). Tripterygiidae: Tripterygiinae. Habitat: marine.

paucifasciatus, Enneapterygius Fricke [R.] 1994:264, Fig. 52 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Grande Terre, Baie de Saint-Vincent, 100 kilometers northwest of Nouméa, 22°01'06"S, 165°55'30"E, New Caledonia. Holotype: MNHN 1997-0433. Paratypes: SMNS 13868 (1). Other material: BPBM 34312 (1). Type catalog: Fricke 1995:20 [ref. 28100], Fricke 2005:66 [ref. 29864]. Misspelled once on p. 264 as paucifaciatus. •Valid as Enneapterygius paucifasciatus Fricke 1994 -- (Fricke 1997:570 [ref. 23339], Fricke 2001:5, 6 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:186 [ref. 29865], Randall 2005:473 [ref. 28239], Fricke 2009:41 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Fricke & Erdmann 2017:27, 29 [ref. 35129]). Current status: Valid as Enneapterygius paucifasciatus Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Caledonia. Habitat: marine.

pectoralis, Enneapterygius Fowler [H. W.] 1941:96, Figs. 10-12 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 93; ref. 13237] Off Sanibel Island, Florida, U.S.A., depth 10 fathoms. Holotype (unique): ANSP 69728. Type catalog: Böhlke 1984:164 [ref. 13621]. •Valid as Gillias pectoralis (Fowler 1941) -- (Fricke 1997:575 [ref. 23339]). •Valid as Enneanectes pectoralis (Fowler 1941) -- (Greenfield & Johnson 1981:50 [ref. 5580], Robins & Ray 1986:221 [ref. 23100], Cervigón 1994:68 [ref. 24488], Schmitter-Soto et al. 2000:165 [ref. 27754], Smith & Williams 2002:9 [ref. 26427], Williams 2003:1749 [ref. 27109], Smith et al. 2003:47 [ref. 27621], Nelson et al. 2004:161 [ref. 27807], McEachran & Fechhelm 2005:577 [ref. 28815], Fricke 2009:37 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Page et al. 2013:166 [ref. 32708], Victor 2013:69 [ref. 32783], Smith-Vaniz & Jelks 2014:66 [ref. 33341]). •Synonym of Enneanectes jordani (Evermann & Marsh 1899) -- (Victor 2017:50 [ref. 35549]). Current status: Synonym of Enneanectes jordani (Evermann & Marsh 1899). Tripterygiidae: Tripterygiinae. Habitat: marine.

personatus, Enneapterygius Fowler [H. W.] 1946:185, Figs. 48-49 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 98; ref. 1456] Aguni-jima, Ryukyu Islands, Japan. Holotype: ANSP 72057. Paratypes: ANSP 72058-59 (2). Type catalog: Böhlke 1984:164 [ref. 13621]. See Fricke 1997:433 [ref. 23339] for selection of fuscipectoris over personatus and quadrimaculatus. •Synonym of Helcogramma obtusirostris (Klunzinger 1871) -- (Hansen 1986:341 [ref. 5810], Shen 1994:21 [ref. 21605]). •Synonym of Helcogramma fuscipectoris (Fowler 1946) -- (Fricke 1997:429, 578 [ref. 23339], Fricke 2009:49 [ref. 30394], Tashiro & Motomura 2014:104 [ref. 35481]). Current status: Synonym of Helcogramma fuscipectoris (Fowler 1946). Tripterygiidae: Tripterygiinae. Habitat: marine.

philippinum, Tripterygium Peters [W. (C. H.)] 1868:269 [Monatsberichte der Königlichen Preussischen Akademie der Wissenschaften zu Berlin 1868; ref. 3442] Coral reef at Paracale [Camarines Norte Province], Luzon Island, Philippines. Lectotype: ZMB 6708. Paralectotypes: ZMB 32768 [ex ZMB 6708] (1). Lectotype selected by Fricke 1997:274 [ref. 23339]. Original genus should have been Tripterygion. •Valid as Enneapterygius philippinus (Peters 1868) -- (Fricke 1997:279, 570 [ref. 23339], Fricke 1999:467 [ref. 24106], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Hutchins 2001:41 [ref. 25847], Fricke 2001:4 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288] dated 1869, Nakabo 2002:1081 [ref. 26193], Allen & Adrim 2003:53 [ref. 26830] dated 1869, Myers & Donaldson 2003:638 [ref. 27495] dated 1869, Randall et al. 2004:24 [ref. 27624] dated 1869, Heemstra et al. 2004:3327 [ref. 27945], Randall 2005:473 [ref. 28239] dated 1869, Hoese 2006:1522 [ref. 28998], Chiang & Chen 2008:197 [ref. 29932] dated 1869, Fricke 2009:41 [ref. 30394], Fricke et al. 2009:93 [ref. 30213], Imamura 2009:243 [ref. 30434], Patzner et al. 2009:445 [ref. 30396], Motomura et al. 2010:184 [ref. 31256], Allen & Erdmann 2012:769 [ref. 31980], Fricke et al. 2013:268 [ref. 32706], Larson et al. 2013:178 [ref. 32988], Fricke et al. 2014:146 [ref. 33932], Fricke & Erdmann 2017:22, 24 [ref. 35129], Koeda et al. 2016:83 [ref. 35333], Fricke et al. 2018:305 [ref. 35805], Arndt & Fricke 2019:32 [ref. 36883]). Current status: Valid as Enneapterygius philippinus (Peters 1868). Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: Mozambique Channel, Seychelles, Comoros, Europa Island, Madagascar and Mascarenes east to Marshall Islands, Samoa, Tonga and Niue, north to southern Japan, south to northwestern Australia, Queensland (Australia) and New Caledonia. Habitat: marine.

phoenicosoma, Enneapterygius Motomura [H.], Ota [R.] & Meguro [M.] in Motomura, Ota, Meguro & Tashiro 2015:2, Figs. 1-5, 8A-B, 9 [Species Diversity v. 20; ref. 33825] Off Nagasaki-bana, Kushikino-kose, Ichiki-kushikino, Kagoshima, Japan, 31°42'24"N, 130°15'44"E, depth 0.5-1 meters. Holotype: KAUM-I. 21960. Paratypes: BSKU, CMNH-ZF, NSMT-P, KAUM-I, MNHN. Plus non-type specimens. •Valid as Enneapterygius phoenicosoma Motomura, Ota & Meguro 2015 -- (Fricke & Erdmann 2017:28 [ref. 35129]). Current status: Valid as Enneapterygius phoenicosoma Motomura, Ota & Meguro 2015. Tripterygiidae: Tripterygiinae. Distribution: Japan, Caroline Islands and Vanuatu. Habitat: marine.

profundum, Forsterygion Fricke [R.] & Roberts [C. D.] in Fricke 1994:336, Fig. 69 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Off Otago Peninsula, 45°41.85'S, 170°58.72'E, Otago, South Island, New Zealand, depth 97-100 meters. Holotype: NMNZ P.25094. Paratypes: NMNZ P.7088 (2), P.7093-97 (1, 1, 8, 3, 5), P.10642 (1), P.10645-46 (1, 3), P.10705 (5), P.10707-10 (1, 1, 1, 2), plus many more paratype lots at NMNZ. Type catalog: Springer & Orrell 1996:29 [ref. 22274]. •Valid as Forsterygion profundum Fricke & Roberts 1994 -- (Fricke 1997:574 [ref. 23339]). •Valid as Matanui profundum (Fricke & Roberts 1994) -- (Jawad & Clements 2004:100 [ref. 27725], Jawad 2008:2944, 2952 [ref. 29958], Fricke 2009:46 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Stewart & Clements 2015:1515 [ref. 34299]). Current status: Valid as Matanui profundum (Fricke & Roberts 1994). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

punctulatus, Enneapterygius Herre [A. W. C. T.] 1935:432 [Field Museum of Natural History, Publications, Zoölogical Series v. 18 (no. 12); ref. 2109] Wala Island, Vanuatu. Holotype: FMNH 17388. Paratypes: (5) FMNH 17389-91 (1, 0, 1); CAS-SU 24437 (3), USNM 112209 [ex FMNH 17390] (1). Additional material: (4) CAS-SU 25117-18 (1, 2). Type catalog: Böhlke 1953:95-96 [ref. 12291], Ibarra & Stewart 1987:34 [ref. 12367], Springer & Orrell 1996:29-30 [ref. 22274]. •Synonym of Enneapterygius minutus (Günther 1877) -- (Fricke 1994:243 [ref. 21827], Fricke 1994:4 [ref. 21828]). •Synonym of Enneapterygius philippinus (Peters 1868) -- (Fricke 1997:275, 570 [ref. 23339], Fricke 2009:41 [ref. 30394]). Current status: Synonym of Enneapterygius philippinus (Peters 1868). Tripterygiidae: Tripterygiinae. Habitat: marine.

pusillus, Enneapterygius Rüppell [W. P. E. S.] 1835:2, Pl. 1 (fig 2) [Neue Wirbelthiere zu der Fauna von Abyssinien gehörig. Fische des Rothen Meeres; ref. 3844] Massawa, Eritrea, Red Sea. Lectotype: SMF 492. Paralectotypes: SMF 492 (1), 4778-79 (2). Lectotype perhaps first established by Fricke 1997:570 [ref. 23339]; Dor's 1984 use of holotype invalid. •Valid as Enneapterygius pusillus Rüppell 1835 -- (Dor 1984:234 [ref. 29757], Holleman 1986:757 [ref. 5671], Goren & Dor 1994:61 [ref. 25356], Randall 1995:31 [ref. 21853], Randall 1995:308 [ref. 22896], Fricke 1997:570 [ref. 23339], Carpenter et al. 1997:207 [ref. 23922], Manilo & Bogorodsky 2003:S116 [ref. 27377], Fricke 2009:41 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Holleman & Bogorodsky 2012:49 [ref. 31735], Golani & Fricke 2018:143 [ref. 36273], Zajonz et al. 2019:92 [ref. 36871]). Current status: Valid as Enneapterygius pusillus Rüppell 1835. Tripterygiidae: Tripterygiinae. Distribution: Red Sea, western Indian Ocean: East Africa, South Africa, Socotra and Gulf of Oman east to southern India. Habitat: marine.

pyramis, Enneapterygius Fricke [R.] 1994:266, Fig. 53 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Rotuma, Fiji Islands, 12°30'S, 177°05'E, depth 0-12 meters. Holotype: USNM 283069. Paratypes: USNM 283107 (9), 283111 (3), 283442 (12), 339019 (6). Plus non-type material. Type catalog: Springer & Orrell 1996:30 [ref. 22274]. •Valid as Enneapterygius pyramis Fricke 1994 -- (Fricke 1997:289, 570 [ref. 23339], Randall 1999:24 [ref. 25829], Williams & Fricke 2001:3534 [ref. 26288], Myers & Donaldson 2003:638 [ref. 27495], Randall et al. 2004:24 [ref. 27624], Randall 2005:473 [ref. 28239], Hoese 2006:1522 [ref. 28998], Fricke 2009:41 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Fricke & Erdmann 2017:20, 27, 30 [ref. 35129]). Current status: Valid as Enneapterygius pyramis Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Western and South Pacific. Habitat: marine.

qirmiz, Enneapterygius Holleman [W.] & Bogorodsky [S. V.] 2012:509, Figs. 2H, 6-7 [Zootaxa No. 3152; ref. 31735] Uqban, Kamaran Island, Red Sea, Yemen, 15°28.893'N, 42°23.141'E. Holotype: SMF 33093 [ex SMF 29594]. Paratypes: BPBM 35721 (1); NHCY-P1 (1); SAIAB 88720 (2); SMF 29357 (2), 29487 (1), 29492 (2), 29594 (1), 33588 (1), 33589 (1), 33590 (1), 33591 (1), 33592 (1); USNM 205805 (6). Plus non-type material. •Valid as Enneapterygius qirmiz Holleman & Bogorodsky 2012 -- (Bogorodsky & Randall 2018:[6] 244 [ref. 36492], Golani & Fricke 2018:143 [ref. 36273]). Current status: Valid as Enneapterygius qirmiz Holleman & Bogorodsky 2012. Tripterygiidae: Tripterygiinae. Distribution: Red Sea endemic. Habitat: marine.

quadra, Enneanectes Victor [B. C.] 2017:57, Figs. 1-11, 13, 16 [Journal of the Ocean Science Foundation v. 27; ref. 35549] Virgin Islands (USA), St. Croix, off north shore of Buck Island Reef National Monument, 17.78925°N, 64.61789°W, depth 1-3 meters. Holotype: UF 149103. Paratypes: AMNH, ANSP, UF. Plus additional non-type material. •Valid as Enneanectes quadra Victor 2017 -- (Victor 2019:12 [ref. 36450]). Current status: Valid as Enneanectes quadra Victor 2017. Tripterygiidae: Tripterygiinae. Distribution: Western Atlantic: Bahamas, Virgin Islands, Belize, Honduras, Antigua, Yucatán, Providencia. Habitat: marine.

quadrimaculatus, Enneapterygius Fowler [H. W.] 1946:189, Fig. 54 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 98; ref. 1456] Aguni-jima, Ryukyu Islands, Japan. Holotype (unique): ANSP 72066. Type catalog: Böhlke 1984:165 [ref. 13621]. See Fricke 1997:433 [ref. 23339] for selection of fuscipectoris over personatus and quadrimaculatus. •Synonym of Helcogramma obtusirostris (Klunzinger 1871) -- (Hansen 1986:341 [ref. 5810], Shen 1994:21 [ref. 21605]). •Synonym of Helcogramma fuscipectoris (Fowler 1946) -- (Fricke 1997:429, 578 [ref. 23339], Fricke 2009:49 [ref. 30394], Tashiro & Motomura 2014:104 [ref. 35481]). Current status: Synonym of Helcogramma fuscipectoris (Fowler 1946). Tripterygiidae: Tripterygiinae. Habitat: marine.

randalli, Enneapterygius Fricke [R.] 1997:295, Fig. 62 [ ref. 23339] Ilots de Bass (Marotiri), southeastern isle, northwestern side of isle, Austral Islands, depth 6-9 meters. Holotype: BPBM 13410. Paratypes: AMNH 72850-52 (1, 5, 13); BPBM 13001 (2), 13307 (2), 17210 (1), 17230 (5), 17285 (1), uncat. (7); plus other non-type material. •Valid as Enneapterygius randalli Fricke 1997 -- (Williams & Fricke 2001:3534 [ref. 26288], Randall 2005:474 [ref. 28239], Fricke 2009:41 [ref. 30394], Patzner et al. 2009:445 [ref. 30396], Fricke & Erdmann 2017:24 [ref. 35129]). Current status: Valid as Enneapterygius randalli Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Southern central Pacific: Austral Islands. Habitat: marine.

randalli, Helcogramma Williams [J. T.] & Howe [J. C.] 2003:171, Figs. 20-22 [aqua, Journal of Ichthyology and Aquatic Biology v. 7 (no. 4); ref. 27249] Komodo Island, southeastern end at Three Rocks, Indonesia, 8°48'18"S, 119°24'48"E, depth 5-6 meters. Holotype: USNM 366861. Paratypes: BPBM 39284 (2), USNM 363778 (8). •Valid as Helcogramma randalli Williams & Howe 2003 -- (Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:777 [ref. 31980], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma randalli Williams & Howe 2003. Tripterygiidae: Tripterygiinae. Distribution: Eastern Indian Ocean: Lesser Sunda Islands, Indonesia. Habitat: marine.

reticulatus, Enneanectes Allen [G. R.] & Robertson [D. R.] 1991:80, Fig. 3 [Revue française d'Aquariologie Herpétologie v. 18 (no. 3); ref. 19723] Near Pichilingue, Baja California, Mexico, about 24°16'N, 110°20'W, depth 0-3 meters. Holotype: USNM 316796. Paratypes: WAM P.30232-001 (1). Type catalog: Springer & Orrell 1996:30 [ref. 22274], Moore et al. 2008:41 [ref. 30117]. •Valid as Gillias reticulatus (Allen & Robertson 1991) -- (Fricke 1997:575 [ref. 23339]). •Valid as Enneanectes reticulatus Allen & Robertson 1991 -- (Allen & Robertson 1994:221 [ref. 22193], Thomson et al. 2000:205 [ref. 25640], Smith & Williams 2002:9 [ref. 26427], Nelson et al. 2004:161 [ref. 27807], Fricke 2009:37 [ref. 30394], Patzner et al. 2009:444 [ref. 30396], Rosenblatt et al. 2013:372 [ref. 32572], Page et al. 2013:166 [ref. 32708]). Current status: Valid as Enneanectes reticulatus Allen & Robertson 1991. Tripterygiidae: Tripterygiinae. Distribution: Eastern Pacific: Gulf of California to Panama. Habitat: marine.

rhabdotus, Enneapterygius Fricke [R.] 1994:270, Fig. 54 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Anano Bay, Nuku Hiva, 8°56'S, 140°00'W, Marquesas Islands. Holotype: AMS I.21773-009. Paratypes: AMS I.21773-014 (2), I.14317 (1), I.17490-061 (1); BPBM 12109 (2), 12585 (10); plus non-type material. •Valid as Enneapterygius rhabdotus Fricke 1994 -- (Fricke 1997:298, 571 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Randall & Earle 2000:18 [ref. 25806], Fricke 2001:5 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Yoshigou & Yoshino 2002:2 [ref. 26253], Fricke 2002:188 [ref. 29865], Randall et al. 2004:24 [ref. 27624], Randall 2005:474 [ref. 28239], Fricke 2009:41 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Allen & Erdmann 2012:770 [ref. 31980], Delrieu-Trottin et al. 2015:7 [ref. 33988], Koeda et al. 2016:83 [ref. 35333], Fricke & Erdmann 2017:20, 25, 26, 30 [ref. 35129]). Current status: Valid as Enneapterygius rhabdotus Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Gulf of Thailand, Philippines, Taiwan, southern Japan, Papua New Guinea, to Society Islands and Marquesas Islands. Habitat: marine.

rharhabe, Helcogramma Holleman [W.] 2007:69, Fig. 11; Pl. 2 [Smithiana, Publications in Aquatic Biodiversity, Bulletin No. 7; ref. 29195] Sheffield Beach, KwaZulu-Natal, South Africa, depth 0-3 meters. Holotype: SAIAB 70740. Paratypes: many at AMS, BMNH, MNHN, ROM, SAIAB, USNM, and WAM. Type catalog: Not treated by Moore et al. 2008 [ref. 30117]. •Valid as Helcogramma rharhabe Holleman 2007 -- (Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396]). Current status: Valid as Helcogramma rharhabe Holleman 2007. Tripterygiidae: Tripterygiinae. Distribution: Western Indian Ocean: southeastern coast of Africa. Habitat: marine.

rhinoceros, Helcogramma Hansen [P. E. H.] 1986:344, Fig. 7 (left), 15 [Bulletin of Marine Science v. 38 (no. 2); ref. 5810] Cuyo Island, northwestern side of Putuc Island, 10°55'05"N, 121°02'03"E, Palawan Province, Philippines, depth 0-15 feet. Holotype: USNM 222370. Paratypes: USNM 221917 (6, 1 c&s), 221920-21 (2, 6). Type catalog: Springer & Orrell 1996:30 [ref. 22274]. •Valid as Helcogramma rhinoceros Hansen 1986 -- (Fricke 1994:431 [ref. 21827], Fricke 1997:467, 580 [ref. 23339], Suzuki et al. 1998:2 [ref. 24384], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1080 [ref. 25182], Allen 2000:116 [ref. 25868], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1080 [ref. 26193], Allen & Adrim 2003:53 [ref. 26830], Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Motomura et al. 2010:189 [ref. 31256], Allen & Erdmann 2012:777 [ref. 31980], Chiang & Chen 2012:67 [ref. 32986], Tashiro & Motomura 2014:107 [ref. 35481], Fricke & Erdmann 2017:43 [ref. 35190], Tashiro & Motomura 2018:[1] [ref. 36174]). Current status: Valid as Helcogramma rhinoceros Hansen 1986. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: East Andaman Sea, Vietnam, Indonesia, Ryukyu Islands, Philippines, Taiwan, Solomon Islands, Vanuatu and New Caledonia. Habitat: marine.

rhothion, Enneapterygius Fricke [R.] 1997:304, Fig. 64 [ ref. 23339] Touaourou, 6 kilometers southeast of Yaté, Grande Terre, New Caledonia, 22°10'36"S, 166°57'51"E, depth 0-0.6 meters. Holotype: SMNS 18322. Paratypes: SMNS 18698 (1), 18773 (7); USNM 297838 (4), 323778 (5), 323781 (4), 343892 (4); plus non-type material. Type catalog: Fricke 2005:66 [ref. 29864]. •Valid as Enneapterygius rhothion Fricke 1997 -- (Fricke 2001:6 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:188 [ref. 29865], Randall 2005:474 [ref. 28239], Fricke 2009:41 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Fricke & Erdmann 2017:29 [ref. 35129]). Current status: Valid as Enneapterygius rhothion Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: New Caledonia and Vanuatu. Habitat: marine.

robustum, Tripterygium Clarke [F. E.] 1879:292, Pl. 15 (upper left) [Transactions and Proceedings of the New Zealand Institute v. 11 (art. 25) (for 1878); ref. 18006] Jackson's Bay, 500 meters northeast of Jackson's Bay town, 40 kilometers south-southwest of Haast, South Island, New Zealand. Neotype: NMNZ uncat. Original genus should have been Tripterygion., Neotype designated by Fricke 1994:347 [ref. 21827]. See Paulin et al. 1989:264 [ref. 24556]. Clements et al. 2000:379 [ref. 25188] suggest that the neotype designated by Fricke does not represent the species described by Clarke, but they were unable to identify it with certainty with 2 other species and suggest it be regarded as a nomen dubium. •Valid as Forsterygion robustum (Clarke 1879) -- (Paulin & Roberts 1992:93 [ref. 24625]). •Synonym of Forsterygion varium (Forster 1801) -- (Fricke 1994:342 [ref. 21827], Fricke 1997:574 [ref. 23339], Fricke 2009:45 [ref. 30394]). Current status: Synonym of Forsterygion varium (Forster 1801). Tripterygiidae: Tripterygiinae. Habitat: marine.

rosea, Helcogramma Holleman [W.] 2006:95, Figs. 3-4 [aqua, Journal of Ichthyology and Aquatic Biology v. 11 (no. 3); ref. 28864] Ko Hi, Phuket, Thailand, 7°44'31"N, 98°22'32"E, depth 0-5 meters. Holotype: ROM 76679. Paratypes: ROM 76680 (3), 76681 (10), 76682 (11); SAIAB 75326 (4); USNM 222915 (5), 228942 (3), 222919 (3), 222916 (3), 307834 (1). USNM 307834 was wrongly listed as a paratype of H. microstigma Holleman (D. Smith, email 11/16/06). •Valid as Helcogramma rosea Holleman 2006 -- (Holleman 2007:71 [ref. 29195], Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:778 [ref. 31980], Fricke & Erdmann 2017:40 [ref. 35190]). Current status: Valid as Helcogramma rosea Holleman 2006. Tripterygiidae: Tripterygiinae. Distribution: Indian Ocean: Sri Lanka and the Andaman Sea (Thailand). Habitat: marine.

rubicauda, Enneapterygius Shen [S.-C.] 1994:17, Fig. 11 [Acta Zoologica Taiwanica v. 5 (no. 2); ref. 21605] Liu-chiu [Liouciou] Township, Pintung County, off southern coast of Taiwan. Holotype: NTUM 07806. Paratypes: NTUM 07803-04 (1, 1), 07857-59 (1, 1, 1); USNM 329661 (1). Non-types: NTUM 07805 (9). Type catalog: Springer & Orrell 1996:30 [ref. 22274], Ho & Shao 2011:52 [ref. 31432]. Enneapterygius erythrosoma and E. rubricauda were described in the same work, Fricke 1997:313 [ref. 23339] serves as first reviser selecting E rubricauda. •Synonym of Enneapterygius flavoccipitis Shen 1994 -- (Chiang & Chen 2008:195 [ref. 29932] with author as Shen & Wu, Endo et al. 2010:9 [ref. 30815], Meguro & Motomura 2010:1 [ref. 31252], Ho & Shao 2011:52 [ref. 31432]). •Valid as Enneapterygius rubicauda Shen 1994 [often wrongly with Shen & Wu as authors] -- (Fricke 1997:310, 571 [ref. 23339], Shimojyo & Hayashi 2000:49 [ref. 24903], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1085 [ref. 25182], Fricke 2001:5 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1085 [ref. 26193], Fricke 2002:189 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830] as rubricauda, Fricke 2009:42 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Allen & Erdmann 2012:770 [ref. 31980], Fricke & Erdmann 2017:28 [ref. 35129]). Current status: Valid as Enneapterygius rubicauda Shen 1994. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Philippines, Taiwan and Ryukyu Islands, Vanuatu and Loyalty Islands. Habitat: marine.

rubinoffi, Axoclinus Allen [G. R.] & Robertson [D. R.] 1992:53, Fig. 1 [Revue française d'Aquariologie Herpétologie v. 19 (nos 1-2); ref. 19713] Malpelo Island, eastern Pacific, west of Colombia, about 4°00'N, 81°31'W, depth 1-2 meters. Holotype: ICNMNH 1786. Paratypes: ICNMNH 1787 (1), USNM 321175 (1), WAM P.30389-001 (1). Type catalog: Springer & Orrell 1996:30 [ref. 22274], Moore et al. 2008:40 [ref. 30117], Mojica & Agudelo-Zamora 2018:52 [ref. 36127]. •Valid as Enneanectes rubinoffi (Allen & Robertson 1992) -- (Fricke 1997:565 [ref. 23339]). •Valid as Axoclinus rubinoffi Allen & Robertson 1992 -- (Allen & Robertson 1994:220 [ref. 22193], Smith & Williams 2002:8 [ref. 26427], Fricke 2009:36 [ref. 30394], Patzner et al. 2009:444 [ref. 30396]). Current status: Valid as Axoclinus rubinoffi Allen & Robertson 1992. Tripterygiidae: Tripterygiinae. Distribution: Eastern Pacific: Colombia. Habitat: marine.

rufopileum, Tripterygion Waite [E. R.] 1904:182, Pl. 24 (fig. 4) [Records of the Australian Museum v. 5 (no. 3); ref. 4561] Lord Howe Island. Syntypes: (100+) AMS I.5914-15 (6), I.5917-25 (28), I.5926-42 (50), I.5944 (2), I.5945-47 (9). •Valid as Vauclusella rufopilea (Waite 1904) -- (Paulin et al. 1989:219 [ref. 24556]). •Valid as Enneapterygius rufopileus (Waite 1904) -- (Kuiter 1993:325 [ref. 23929], Francis 1993:166 [ref. 25479] as rufopilea, Fricke 1994:273 [ref. 21827], Fricke 1997:314, 571 [ref. 23339], Kuiter 1997:334 [ref. 25488], Fricke 2001:5 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:189 [ref. 29865], Randall et al. 2004:24 [ref. 27624], Randall 2005:474 [ref. 28239], Hoese 2006:1522 [ref. 28998], Fricke 2009:42 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Fricke & Erdmann 2017:28 [ref. 35129]). Current status: Valid as Enneapterygius rufopileus (Waite 1904). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific. Habitat: marine.

rufus, Acanthanectes Holleman [W.] & Buxton [C. D.] 1993:336, Figs. 6A, 8 [Cybium v. 17 (no. 4); ref. 21100] Steilkop Reef, 34°01'40"S, 23°54'30"E, Tsitsikamma National Park, South Africa, depth 20-25 meters. Holotype: SAIAB [formerly RUSI] 36689. Paratypes: AMS I.33647-001 [formerly SAIAB] 36700 (1); NMNZ P.29421 (1); SAIAB [formerly RUSI] 13416 (1, c&s), 36701-03 (2, 1, 2); USNM 326001 (2). Type catalog: Springer & Orrell 1996:30 [ref. 22274]. •Valid as Acanthanectes rufus Holleman & Buxton 1993 -- (Fricke 1997:561 [ref. 23339], Fricke 2009:32 [ref. 30394], Patzner et al. 2009:443 [ref. 30396]). Current status: Valid as Acanthanectes rufus Holleman & Buxton 1993. Tripterygiidae: Tripterygiinae. Distribution: Southeastern Atlantic. Habitat: marine.

segmentatum, Tripterygion McCulloch [A. R.] & Phillipps [W. J.] 1923:20, Pl. 4 (fig. 3) [Records of the Australian Museum v. 14; ref. 15411] Shag Point, Otago, New Zealand, subtidal. Holotype (unique): NMNZ P.225. Type catalog: Hardy 1990:14 [ref. 18942]. •Valid as Notoclinops segmentatus (McCulloch & Phillipps 1923) -- (Hardy 1987:166 [ref. 6027], Paulin et al. 1989:220 [ref. 24556], Paulin & Roberts 1992:72 [ref. 24625], Fricke 1994:487 [ref. 21827], Fricke 1997:584 [ref. 23339], Fricke 2009:46 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1517 [ref. 34299]). Current status: Valid as Notoclinops segmentatus (McCulloch & Phillipps 1923). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand. Habitat: marine.

senoui, Enneapterygius Motomura [H.], Harazaki [S.] & Hardy [G. S.] 2005:6, Figs. 1-3, 5-6 [aqua, Journal of Ichthyology and Aquatic Biology v. 10 (no. 1); ref. 28338] Inside Sokodo fishing port, Hachijo-jima Island, Izu Islands, off Pacific coast of central Honshu Island, Japan, depth 3 meters. Holotype: KPM-NI 9613. Paratypes: AMS I. 43160-001 (1); BPBM 39136 (1); KPM-NI 9606 (1), 9612 (1), 9614 (1). •Valid as Enneapterygius senoui Motomura, Harazaki & Hardy 2005 -- (Fricke 2009:42 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Endo et al. 2010:14 [ref. 30815], Fricke & Erdmann 2017:25, 27 [ref. 35129]). Current status: Valid as Enneapterygius senoui Motomura, Harazaki & Hardy 2005. Tripterygiidae: Tripterygiinae. Distribution: Izu and Ogasawara Islands, Japan. Habitat: marine.

serendip, Helcogramma Holleman [W.] 2007:73, Fig. 13 [Smithiana, Publications in Aquatic Biodiversity, Bulletin No. 7; ref. 29195] Near Fort Frederick, Trincomalee, Sri Lanka. Holotype: SAIAB 73755. Paratypes: SAIAB 73756 (12); USNM 228977 (5), 228981 (3), 385565 (2), 385567 (5). •Valid as Helcogramma serendip Holleman 2007 -- (Fricke 2009:50 [ref. 30394], Patzner et al. 2009:447 [ref. 30396]). Current status: Valid as Helcogramma serendip Holleman 2007. Tripterygiidae: Tripterygiinae. Distribution: Indian Ocean: Sri Lanka. Habitat: marine.

sexmaculatus, Gillias Fowler [H. W.] 1944:286, Fig. 241 [Monographs of the Academy of Natural Sciences of Philadelphia No. 6; ref. 1448] Tide pool at Saboga, Perlas Islands, (Pacific) Panama. Holotype (unique): ANSP 70074. Type catalog: Böhlke 1984:62 [ref. 13621]. •Valid as Enneanectes sexmaculatus (Fowler 1944) -- (Allen & Robertson 1994:221 [ref. 22193], De La Cruz Agüero et al. 1997:302 [ref. 24545], Thomson et al. 2000:205 [ref. 25640], Nelson et al. 2004:161 [ref. 27807]). •Synonym of Enneanectes carminalis (Jordan & Gilbert 1882) -- (Fricke 1997:564 [ref. 23339], Smith & Williams 2002:8 [ref. 26427], Fricke 2009:37 [ref. 30394]). Current status: Synonym of Enneanectes carminalis (Jordan & Gilbert 1882). Tripterygiidae: Tripterygiinae. Habitat: marine.

shaoi, Enneapterygius Chiang [M.-C.] & Chen [I-S.] 2008:189, Figs. 1c, 1d, 3 [Raffles Bulletin of Zoology Suppl. No. 19; ref. 29932] Feng-chui-sha, Hengchun Township, Pingtung County, Taiwan, depth 3-12 meters. Holotype: NTOU-P 2008-06-304. Paratypes: NTOU-P 2008-06-299, -300, -301, -302, -303 (5). Type catalog: Ho & Shao 2011:53 [ref. 31432]. •Valid as Enneapterygius shaoi Chiang & Chen 2008 -- (Ho & Shao 2011:53 [ref. 31432], Meguro & Motomura 2010:1 [ref. 31252], Fricke & Erdmann 2017:20, 26, 31 [ref. 35129]). Current status: Valid as Enneapterygius shaoi Chiang & Chen 2008. Tripterygiidae: Tripterygiinae. Distribution: Western North Pacific: southern Taiwan. Habitat: marine.

sheni, Enneapterygius Chiang [M.-C.] & Chen [I-S.] 2008:185, Figs. 1a, 1b, 2 [Raffles Bulletin of Zoology Suppl. No. 19; ref. 29932] Feng-chui-sha, Hengchun Township, Pingtung County, Taiwan, depth 3-12 meters. Holotype: NTOU-P 208-06-366. Paratypes: NTOU-P 2008-06-359, -363, -367, -368 and -369 (1 ea. = 5). Type catalog: Ho & Shao 2011:53 [ref. 31432]. •Valid as Enneapterygius sheni Chiang & Chen 2008 -- (Ho & Shao 2011:53 [ref. 31432], Meguro & Motomura 2010:1 [ref. 31252], Fricke & Erdmann 2017:26 [ref. 35129]). Current status: Valid as Enneapterygius sheni Chiang & Chen 2008. Tripterygiidae: Tripterygiinae. Distribution: Western North Pacific: Taiwan. Habitat: marine.

shinglensis, Helcogramma Lal Mohan [R. S.] 1971:219, Fig. 1 [Senckenbergiana Biologica v. 52 (no. 3/5); ref. 21432] Shingle Island, Gulf of Mannar, between India and Sri Lanka, 9°12'N, 79°8'E. Holotype: FMRI 165. Paratypes: FMRI F166 (2); SMF 11004 (1). •Synonym of Helcogramma obtusirostre (Klunzinger 1871) -- (Fricke 1997:579 [ref. 23339]). •Valid as Helcogramma shinglensis Lal Mohan 1971 -- (Holleman 2007:74 [ref. 29195], Fricke 2009:51 [ref. 30394], Patzner et al. 2009:447 [ref. 30396]). •Synonym of Helcogramma obtusirostris (Klunzinger 1871) -- (Hansen 1986:341 [ref. 5810], Holleman & Bogorodsky 2012:54 [ref. 31735]). Current status: Synonym of Helcogramma obtusirostris (Klunzinger 1871). Tripterygiidae: Tripterygiinae. Distribution: Indian Ocean: Sri Lanka and Gulf of Mannar [if valid]. Habitat: marine.

signata, Grahamina Fricke [R.] & Roberts [C. D.] 1993:16, Fig. 9 [Stuttgarter Beiträge zur Naturkunde. Serie A (Biologie). No. 504; ref. 21298] Tauranga Bay, on south side, South Island, New Zealand, 41°47'S, 171°27'E. Holotype: NMNZ P.30571 [?P.30569]. Paratypes: AMS I.34234-001 (5); BMNH 1993.9.24:1-5 (5); NMNZ P.14109 (2), P. 17083 (17), P.30570 (5); QM types 233-234 (2); SMNS 13916 (66), 13933 (21), 14047 (1); USNM 326614 (5). Type catalog: Fricke 1995:21 [ref. 28100], Springer & Orrell 1996:30 [ref. 22274], Fricke 2005:68 [ref. 29864]. Clements et al. 2000:381 [ref. 25188] conclude that signata (= gymnota) was introduced to Tasmania by human activity. •Valid as Grahamina signata Fricke & Roberts 1993 -- (Fricke 1994:395 [ref. 21827], Fricke 1997:576 [ref. 23339]). •Synonym of Grahamina gymnota (Scott 1977) -- (Clements et al. 2000:379 [ref. 25188], Fricke 2009:45 [ref. 30394]). •Synonym of Forsterygion gymnota Scott 1977 -- (Jawad 2008:2962 [ref. 29958]). Current status: Synonym of Forsterygion gymnotum Scott 1977. Tripterygiidae: Tripterygiinae. Habitat: marine.

signata, Taboguilla Allen [G. R.] & Robertson [D. R.] 1991:81, Figs. 4-5 [Revue française d'Aquariologie Herpétologie v. 18 (no. 3); ref. 19723] Taboguilla Island, Gulf of Panama, about 8°49'N, 79°31'W, depth 3-10 meters. Holotype: USNM 316800. Paratypes: USNM 316801 (4), WAM P.30229-001 (1). Type catalog: Springer & Orrell 1996:30 [ref. 22274], Moore et al. 2008:41 [ref. 30117]. •Synonym of Lepidonectes clarkhubbsi Bussing 1991 -- (Allen & Robertson 1994:222 [ref. 22193], Fricke 1997:582 [ref. 23339]). Current status: Synonym of Lepidonectes clarkhubbsi Bussing 1991. Tripterygiidae: Tripterygiinae. Habitat: marine.

signicauda, Enneapterygius Fricke [R.] 1997:322, Fig. 67 [ ref. 23339] Small cave about 600 meters north of village, Waisisi, Tanna Island, 19°28'22"S, 169°23'58"E, depth 0-7 meters. Holotype: USNM 344013. Paratypes: SMNS 19173 (1); USNM 331036 (7), 344006 (2), 344008 (1), 344015 (1); plus non-type material. Type catalog: Fricke 2005:67 [ref. 29864]. •Valid as Enneapterygius signicauda Fricke 1997 -- (Shimojyo & Hayashi 2000:51 [ref. 24903], Nakabo 2000:1085 [ref. 25182], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1085 [ref. 26193], Randall et al. 2004:24 [ref. 27624], Fricke 2009:42 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Meguro & Motomura 2010:6 [ref. 31252], Fricke & Erdmann 2017:24 [ref. 35129]). Current status: Valid as Enneapterygius signicauda Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Vanuatu and American Samoa. Habitat: marine.

similis, Enneapterygius Fricke [R.] 1997:326, Fig. 68 [ ref. 23339] Ibahos Island, western side, north end, Batanes Province, Philippines, 20°20'10"N, 121°49'10"E, depth 2-5 meters. Holotype: USNM 293726. Paratypes: AMS (3 lots, 143), BPBM (1), SMNS (4 lots, 24), USNM (22 lots, 347), WAM P.30416-011 (1). Type catalog: Fricke 2005:67 [ref. 29864], Moore et al. 2008:41 [ref. 30117]. •Valid as Enneapterygius similis Fricke 1997 -- (Shimojyo & Hayashi 2000:52 [ref. 24903], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1086 [ref. 25182], Fricke 2001:4 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1086 [ref. 26193], Fricke 2002:190 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830], Randall 2005:475 [ref. 28239], Hoese 2006:1522 [ref. 28998], Fricke 2009:42 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Meguro & Motomura 2010:6 [ref. 31252], Allen & Erdmann 2012:770 [ref. 31980], Fricke & Erdmann 2017:24 [ref. 35129]). Current status: Valid as Enneapterygius similis Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Malaysia, Indonesia, Philippines, Australia, Vanuatu and New Caledonia. Habitat: marine.

smithi, Enneanectes Lubbock [R.] & Edwards [A. J.] 1981:148, Fig. 4 [Journal of Fish Biology v. 18 (no. 2); ref. 8578] North side of Belmonte Islet, St Paul's Rocks, 0°56'N, 29°22'W, mid-Atlantic ridge, rocks and caves, depth 5-10 meters. Holotype: MZUSP 14616. Paratypes: BMNH 1980.1.22.21 (1), SMF 15674 (1). •Valid as Gillias smithi (Lubbock & Edwards 1981) -- (Fricke 1997:575 [ref. 23339]). •Valid as Enneanectes smithi Lubbock & Edwards 1981 -- (Smith & Williams 2002:9 [ref. 26427], Feitoza et al. 2003:69 [ref. 26943], Moura & Menezes in Menezes et al. 2003:94 [ref. 27192], Vaske et al. 2005:78 [ref. 36155], Fricke 2009:37 [ref. 30394], Patzner et al. 2009:444 [ref. 30396]). Current status: Valid as Enneanectes smithi Lubbock & Edwards 1981. Tripterygiidae: Tripterygiinae. Distribution: Central Atlantic: St. Paul's Rocks (Saint Peter and Saint Paul Archipelago). Habitat: marine.

solorensis, Helcogramma Fricke [R.] 1997:472, Fig. 96 [ ref. 23339] Savu Sea, Solor Island, eastern side, off Motton Point, Nusa Tenggara Timur Province, Indonesia, depth 3 meters. Holotype: BPBM 36660. Paratypes: BPBM uncat. (1), SMNS 21880 [ex BPBM uncat.] (1). Type catalog: Fricke 2005:69 [ref. 29864]. •Valid as Helcogramma solorensis Fricke 1997 -- (Williams & Fricke 2001:3534 [ref. 26288], Allen & Adrim 2003:53 [ref. 26830] dated 1994, Fricke 2009:51 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:778 [ref. 31980], Fricke & Erdmann 2017:43 [ref. 35190]). Current status: Valid as Helcogramma solorensis Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Solor Island, Indonesia. Habitat: marine.

springeri, Helcogramma Hansen [P. E. H.] 1986:345, Fig. 16 [Bulletin of Marine Science v. 38 (no. 2); ref. 5810] Off Tandjung Suli, Ambon Island, Molucca Islands, Indonesia, depth less than 2 meters. Holotype: USNM 229368. Paratypes: AMS I.19456-092 (14), 19483-072 (14), 20756-009 (4), 20774-012 (30), 21451-001 (4), 22582-048 (3); USNM 150550 (1), 164600 (1), 210277 (3, not 1), 222349 (5), 222369 (1), 222372 (1), 228903 (22), 228932 (2), 228935 (5), 228940 (1), 263387-89 (1, 1, now 1), 285799 [ex 263389] (1). Type catalog: Springer & Orrell 1996:30 [ref. 22274]. •Valid as Helcogramma springeri Hansen 1986 -- (Fricke 1994:434 [ref. 21827], Fricke 1997:475, 580 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Allen & Adrim 2003:53 [ref. 26830], Hoese 2006:1525 [ref. 28998], Fricke 2009:51 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:779 [ref. 31980], Fricke et al. 2014:147 [ref. 33932], Fricke & Erdmann 2017:43 [ref. 35190]). Current status: Valid as Helcogramma springeri Hansen 1986. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: Andaman Sea, Malaysia, Indonesia, Philippines, Papua New Guinea, Solomon Islands and Australia. Habitat: marine.

springeri, Norfolkia Clark [E.] 1980:95, Figs. 4f, 5d, 9 [Israel Journal of Zoology v. 28 (nos 3-4) [for 1979]; ref. 8837] Northwestern coast just north of Ras Burqa, Sinai, Egypt, Gulf of Aqaba, Red Sea, depth 0-11 m. Holotype: USNM 205793. Paratypes: HUJ 5355 [ex USNM 205796] (2); USNM 205794 (2), 205795 (1, c&s), 205796 (now 0). Type catalog: Springer & Orrell 1996:31 [ref. 22274], Golani 2006:31 [ref. 28840]. •Valid as Norfolkia spinrgeri Clark 1980 -- (Dor 1984:234 [ref. 29757], Holleman 1986:758 [ref. 5671]). •Synonym of Norfolkia brachylepis (Schultz 1960) -- (Fricke 1991:340 [ref. 19545], Holleman 1991:233 [ref. 21794], Fricke 1994:465 [ref. 21827], Fricke 1997:504, 583 [ref. 23339], Fricke 2009:34 [ref. 30394], Golani & Fricke 2018:144 [ref. 36273]). Current status: Synonym of Norfolkia brachylepis (Schultz 1960). Tripterygiidae: Tripterygiinae. Habitat: marine.

squamiceps, Gillias McCulloch [A. R.] & Waite [E. R.] 1916:449, Pl. 41 (fig. 1) [Transactions Royal Society of South Australia v. 40; ref. 15058] Lord Howe Island. Holotype: AMS I.5401 (1 of 2, 55 mm). Paratypes: (2) AMS I.5401 (1 of 2, not 55 mm). •Valid as Norfolkia squamiceps (McCulloch & Waite 1916) -- (Fricke 1991:339 [ref. 19545], Holleman 1991:236 [ref. 21794], Francis 1993:166 [ref. 25479], Fricke 1994:473 [ref. 21827], Fricke 1997:512, 583 [ref. 23339], Randall et al. 1997:522 [ref. 25919], Fricke 2001:3 [ref. 25985], Williams & Fricke 2001:3535 [ref. 26288], Fricke 2002:195 [ref. 29865], Randall 2005:478 [ref. 28239], Hoese 2006:1526 [ref. 28998], Fricke 2009:34 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Holleman & Bogorodsky 2012:58 [ref. 31735]). Current status: Valid as Norfolkia squamiceps (McCulloch & Waite 1916). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific. Habitat: marine.

steinitzi, Helcogramma Clark [E.] 1980:88, Figs. 3d, 5a, 6a, 8; Pls. 2-5 [Israel Journal of Zoology v. 28 (nos 3-4) [for 1979]; ref. 8837] Bay between Marsa Mokrakh and El Himeira, Sinai, Egypt, northwestern coast of Gulf of Aqaba, Red Sea. Holotype: USNM 205787. Paratypes: HUJ 4656 (3), 5351-52 (12, 1), 9209 (2); TAU [= SMNHTAU] N.S.2623a (2), 4076a (5), 4541 (1), 12158b (1), 12161 (1); USNM 205788-90 (2, now 3, 4), 205791 (12, 4 c&s), 205792 (6), 205824 (7), 205826 (1), 205828 (1), 205830 (1), 205832-33 (1, 17). Type catalog: Springer & Orrell 1996:31 [ref. 22274], Golani 2006:31 [ref. 28840]. •Valid as Helcogramma steinitzi Clark 1980 -- (Dor 1984:234 [ref. 29757], Hansen 1986:347 [ref. 5810], Goren & Dor 1994:61 [ref. 25356], Randall 1995:33 [ref. 21853], Randall 1995:310 [ref. 22896], Fricke 1997:580 [ref. 23339], Carpenter et al. 1997:207 [ref. 23922], Manilo & Bogorodsky 2003:S116 [ref. 27377], Holleman 2006:98 [ref. 28864], Holleman 2007:76 [ref. 29195], Fricke 2009:51 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Holleman & Bogorodsky 2012:56 [ref. 31735], Fricke & Erdmann 2017:40 [ref. 35190], Golani & Fricke 2018:143 [ref. 36273], Zajonz et al. 2019:92 [ref. 36871]). Current status: Valid as Helcogramma steinitzi Clark 1980. Tripterygiidae: Tripterygiinae. Distribution: Red Sea, northwestern Indian Ocean: Socotra, Gulf of Oman, Persian Gulf. Habitat: marine.

stewarti, Karalepis Hardy [G. S.] 1984:177, Figs. 1-4 [National Museum of New Zealand Records v. 2 (no. 16); ref. 5340] Princess Bay, Wellington, New Zealand. Holotype: NMNZ P.13996. Paratypes: AIM 5100 (2); BMNH 1984.2.9.1 (1); NMNZ P.13592 (1), P.14340 (4), P.14410-15 (4, 1, 1, 5, 7, 4), P.14988 (13), P.15054 (1), P.15426 (now 1); SAIAB [formerly RUSI] 20010 (2); USNM 265098 (2). Type catalog: Hardy 1990:13 [ref. 18942], Springer & Orrell 1996:31 [ref. 22274]. •Valid as Karalepis stewarti Hardy 1984 -- (Paulin et al. 1989:220 [ref. 24556], Paulin & Roberts 1992:82 [ref. 24625], Fricke 1994:444 [ref. 21827], Fricke 1997:581 [ref. 23339], Fricke 2009:47 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1513 [ref. 34299], Cornejo et al. 2018:17 [ref. 36409]). Current status: Valid as Karalepis stewarti Hardy 1984. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

storeyae, Enneapterygius Brock [V. E.] 1940:33 [Stanford Ichthyological Bulletin v. 2 (no. 1); ref. 13593] Los Frailes, Cape San Lucas, Baja California, depth 15 feet. Holotype (unique): CAS-SU 35045. Type catalog: Böhlke 1953:96 [ref. 12291]. •Synonym of Enneanectes carminalis (Jordan & Gilbert 1882) -- (Fricke 1997:564 [ref. 23339]). •Valid as Axoclinus storeyae (Brock 1940) -- (Smith & Williams 2002:8 [ref. 26427], Fricke 2009:36 [ref. 30394], Patzner et al. 2009:444 [ref. 30396] as storeya, Page et al. 2013:166 [ref. 32708], Galván-Villa et al. 2016:148 [ref. 34871]). Current status: Valid as Axoclinus storeyae (Brock 1940). Tripterygiidae: Tripterygiinae. Distribution: Eastern Pacific: Mexico. Habitat: marine.

striata, Ceratobregma Fricke [R.] 1991:337, Fig. 1 [Japanese Journal of Ichthyology v. 37 (no. 4); ref. 19545] North Oxley Island, Northern Territory, Australia. Holotype (unique): NTM S10605-018. •Valid as Ceratobregma striata Fricke 1991 -- (Fricke 1994:164 [ref. 21827]). •Synonym of Ceratobregma helenae Holleman 1987 -- (Fricke 1997:133, 563 [ref. 23339], Fricke & Kulbicki 2006:345 [ref. 28867], Hoese 2006:1519 [ref. 28988], Fricke 2009:36 [ref. 30394], Larson et al. 2013:177 [ref. 32988]). Current status: Synonym of Ceratobregma helenae Holleman 1987. Tripterygiidae: Tripterygiinae. Habitat: marine.

striata, Helcogramma Hansen [P. E. H.] 1986:349, Fig. 18 [Bulletin of Marine Science v. 38 (no. 2); ref. 5810] Toga Point rocks, Miyakejima, Izu Islands, Japan, 34°07'N, 139°30'E, depth 1-3 meters. Holotype: USNM 221667. Paratypes: AMS I.19456-093 (5), I.22619-024 (1); USNM 221666 (1, lost), 221668-69 (1, 9), 221916 (orig. 13 or 14, now 9) ?USNM ex 222321 (3, c&s), 263386 (8, not 7), 269807 (1, c&s). Type catalog: Springer & Orrell 1996:31-32 [ref. 22274] but types very confusing, Ho & Shao 2011:53 [ref. 31432]. Helcogramma is feminine, so correct spelling is striata. •Valid as Helcogramma striata [sometimes as striatum] Hansen 1986 -- (Randall et al. 1990:365 [ref. 15987], Fricke 1994:437 [ref. 21827], Shen 1994:22 [ref. 21605], Fricke 1997:480, 580 [ref. 23339], Allen 1997:212 [ref. 23977] as striata, Kuiter 1997:334 [ref. 25488] as striata, Randall et al. 1997:365 [ref. 25919], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1078 [ref. 25182], Allen 2000:116 [ref. 25868], Hutchins 2001:41 [ref. 25847], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1078 [ref. 26193], Allen & Adrim 2003:53 [ref. 26830], Shibukawa et al. in Kimura & Matsuura 2003:168 [ref. 27510], Adrim et al. 2004:125 [ref. 27715], Randall 2005:477 [ref. 28239], Hoese 2006:1525 [ref. 28988], Holleman 2007:77 [ref. 29195] as striata, Fricke 2009:51 [ref. 30394] as striata, Imamura 2009:244 [ref. 30434] as striata, Patzner et al. 2009:447 [ref. 30396] as striata, Motomura et al. 2010:189 [ref. 31256] as striata, Ho & Shao 2011:53 [ref. 31432] as striata, Allen & Erdmann 2012:780 [ref. 31980], Chiang & Chen 2012:58, 68 [ref. 32986], Larson et al. 2013:178 [ref. 32988] as striatum, Tashiro & Motomura 2014:108 [ref. 35481], Motomura et al. 2015:28 [ref. 34709], Koeda et al. 2016:84 [ref. 35333], Fricke & Erdmann 2017:41 [ref. 35190]). Current status: Valid as Helcogramma striata Hansen 1986. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: Andaman Sea to Line Islands and Fiji, Australia to Japan and Taiwan; throughout the East Indian region. Habitat: marine.

striaticeps, Tripterygium Ramsay [E. P.] & Ogilby [J. D.] 1888:419 [Proceedings of the Linnean Society of New South Wales (Series 2) v. 3 (pt 2); ref. 18599] Intertidal, Tailor Bay, Port Jackson, New South Wales, Australia. Syntypes: AMS I.1693-94 (1, 1). Original genus should have been Tripterygion. •Valid as Norfolkia striaticeps (Ramasay & Ogilby 1888) -- (Fricke 1991:339 [ref. 19545], Gomon et al. 1994:738 [ref. 22532]). •Synonym of Trinorfolkia clarkei (Morton 1888) -- (Fricke 1994:541 [ref. 21827], Fricke 1997:586 [ref. 23339], Hoese 2006:1527 [ref. 28988], Fricke 2009:35 [ref. 30394]). Current status: Synonym of Trinorfolkia clarkei (Morton 1888). Tripterygiidae: Tripterygiinae. Habitat: marine.

tartessicum, Tripterygion Carreras-Carbonell [J.], Pascual [M.] & Macpherson [E.] 2007:80, Figs. 2, 3a [Scientia Marina v. 71 (no. 1); ref. 29025] Cádiz, Puercas Phare, Spain, 36°18'N, 6°12'W, depth 0-2 meters. Holotype: IIPB 15/2005. Paratypes: Specimens at IIPB, MNCN, and SMNS. •Valid as Tripterygion tartessicum Carreras-Carbonell, Pascual & Macpherson 2007 -- (Fricke 2009:44 [ref. 30394], Patzner et al. 2009:449 [ref. 30396], Cornejo et al. 2018:17 [ref. 36409]). Current status: Valid as Tripterygion tartessicum Carreras-Carbonell, Pascual & Macpherson 2007. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Mediterranean Sea; eastern Atlantic: northern Morocco. Habitat: freshwater.

thomasi, Norfolkia Whitley [G. P.] 1964:193 [Records of the Australian Museum v. 26 (no. 5); ref. 3772] Heron Island, Capricorn Group, 23°26'S, 151°55'E, Queensland, Australia. Holotype: AMS IB.4020. Paratypes: AMS IB.4021 (2), IB.6245 (1). •Valid as Norfolkia thomasi Whitley 1964 -- (Fricke 1991:339 [ref. 19545], Holleman 1991:238 [ref. 21794], Fricke 1994:476 [ref. 21827], Fricke 1997:516, 583 [ref. 23339], Randall et al. 1997:523 [ref. 25919], Johnson 1999:747 [ref. 25471], Nakabo 2000:1077 [ref. 25182], Fricke 2001:3 [ref. 25985], Williams & Fricke 2001:3535 [ref. 26288], Nakabo 2002:1077 [ref. 26193], Fricke 2002:195 [ref. 29865], Randall et al. 2004:24 [ref. 27624], Randall 2005:478 [ref. 28239], Hoese 2006:1526 [ref. 28998], Fricke 2009:34 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Motomura et al. 2010:190 [ref. 31256], Holleman & Bogorodsky 2012:58 [ref. 31735]). Current status: Valid as Norfolkia thomasi Whitley 1964. Tripterygiidae: Tripterygiinae. Distribution: Queensland Australia and southern Pacific to Rapa. Habitat: marine.

trigloides, Tripterygion Bleeker [P.] 1858:234 [Natuurkundig Tijdschrift voor Nederlandsch Indië v. 15 (no. 1); ref. 363] Southeastern edge of Harbor, Port Narevin, Erromango Island, Vanuatu, 18°44'25"S, 169°12'41"E, depth 0-6 meters. Neotype: USNM 343890 [not 343980]. Neotype designated by Fricke 1997:493 [ref. 23339] as 343980; original locality w. Belitung I., Indonesia. Bleeker's unpublished illustration reproduced in Boeseman 1983:v. 11, Pl. 442, fig. 6 [ref. 12564]. •Status uncertain -- (Hansen 1986:352 [ref. 5810]). •Valid as Helcogramma trigloides (Bleeker 1858) -- (Yoshino in Masuda et al. 1984:294 [ref. 6441], Fricke 1997:489, 580 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Fricke 2001:4 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288] with author as Fricke 1997, Fricke 2002:193 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830], Fricke 2009:51 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:780 [ref. 31980], Chaing & Chen 2012:70 [ref. 32986], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma trigloides (Bleeker 1858). Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: East Andaman Sea, Malaysia, Indonesia, Philippines, Taiwan, Papua New Guinea, Solomon Islands and Vanuatu. Habitat: marine.

tripennis, Blennius Forster [J. R.] in Bloch & Schneider 1801:174 [M. E. Blochii, Systema Ichthyologiae; ref. 471] Outer Chetwode Island, Golden Bay and Marlborough sounds, South Island, New Zealand. Neotype: NMNZ P.15419. Authorship follows Fricke 1994:17 [ref. 21827]. Neotype designated by Fricke 1994:362 [ref. 21827]. Based on description in manuscript by Forster (Johann Reinhold Forster, 1729-1798, German explorer and naturalist of Cook's second voyage). •Valid as Gilloblennius tripennis (Forster 1801) -- (Hardy 1986:147 [ref. 6139], Paulin et al. 1989:219 [ref. 24556], Paulin & Roberts 1992:84 [ref. 24625], Fricke 1994:358 [ref. 21827], Fricke 1997:575 [ref. 23339], Fricke 2009:33 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Stewart & Clements 2015:1512 [ref. 34299]). Current status: Valid as Gilloblennius tripennis (Forster 1801). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

tripteronotus, Blennius Risso [A.] 1810:135, Pl. 5 (fig. 14) [Ichthyologie de Nice; ref. 3755] Nice, France, northwestern Mediterranean Sea. No types known. Type catalog: Bauchot 1967:20 [ref. 20734]. Gender of genus neuter, therefore correct species name tripteronotum. •Valid as Tripterygion tripteronotus (Risso 1810) -- (Wheeler 1973:531 [ref. 7190], Zander in Whitehead et al. 1986:1120 [ref. 13677], Fricke 1997:587 [ref. 23339], Al-Hassan 1999:196 [ref. 25945], Geertjes et al. 2001:256 [ref. 25579], Bilecenoğlu et al. 2002:120 [ref. 26753], Parin 2003:S23 [ref. 28536], Carreras-Carbonell et al. 2007:82 [ref. 29024], Fricke et al. 2007:97 [ref. 29533], Vasil'eva 2007:123 [ref. 30517], Kovačić & Golani 2008:423 [ref. 33214], Ninua & Japoshvili 2008:172 [ref. 31251], Patzner et al. 2009:449 [ref. 30396], Lipej & Dulčić 2010:57 [ref. 36649], Artüz & Fricke 2019:558 [ref. 36612]). •Valid as Tripterygion tripteronotum (Risso 1810) -- (Fricke 2009:44 [ref. 30394]). Current status: Valid as Tripterygion tripteronotum (Risso 1810). Tripterygiidae: Tripterygiinae. Distribution: Black Sea and Mediterranean Sea. Habitat: marine.

triserialis, Enneapterygius Fricke [R.] 1994:280, Figs. 56-57 [Tripterygiid fishes of Australia, New Zealand and the southwest Pacific Ocean, with descriptions of 2 new genera and 16 new species (Teleostei) v. 24; ref. 21827] Southwestern side of Ile Longue, Chesterfield Bank, New Caledonia, 19°52'S, 158°19.8'E, eastern Coral Sea, depth 9 meters. Holotype: BPBM 33670. Paratypes: AMS I.30465-108 (14); BPBM 33645 (1), 36434 (1); WAM P.29627-034 (4). Plus many lots of non-type material. Type catalog: Moore et al. 2008:41 [ref. 30117]. •Valid as Enneapterygius triserialis Fricke 1994 -- (Fricke 1997:331, 572 [ref. 23339], Fricke 2001:5 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:190 [ref. 29865], Randall et al. 2004:24 [ref. 27624], Randall 2005:475 [ref. 28239], Hoese 2006:1522 [ref. 28998], Fricke 2009:42 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Fricke & Erdmann 2017:26, 27 [ref. 35129]). Current status: Valid as Enneapterygius triserialis Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Southwestern and South Pacific. Habitat: marine.

trisignatus, Enneapterygius Fricke [R.] 2001:6, Fig. 1 [Stuttgarter Beiträge zur Naturkunde. Serie A (Biologie). No. 630; ref. 25985] Province Nord, northern end of island, 1 kilometer east of Pointe Naharian, 20 kilometers north of Poum, 20°04'18"S, 164°00'15"E, Grande Terre, New Caledonia, depth 1-3 meters. Holotype: SMNS 22074. Paratypes: MNHN 2000-1447 (1), SMNS 22085 (1). Type catalog: Fricke 2005:67 [ref. 29864]. •Valid as Enneapterygius trisignatus Fricke 2001 -- (Fricke 2001:5 [ref. 25985], Fricke 2002:191 [ref. 29865], Fricke 2009:42 [ref. 30394], Patzner et al. 2009:446 [ref. 30396]). Current status: Valid as Enneapterygius trisignatus Fricke 2001. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Caledonia. Habitat: marine.

tusitalae, Enneapterygius Jordan [D. S.] & Seale [A.] 1906:416, Fig. 97 [Bulletin of the Bureau of Fisheries v. 25 (for 1905); ref. 2497] Pago Pago, American Samoa. Holotype: USNM 51800. Paratypes: MCZ 35905 (1), USNM 164583 (4). Type catalog: Springer & Orrell 1996:32 [ref. 22274]. •Synonym of Enneapterygius minutus (Günther 1877) -- (Fricke 1994:242 [ref. 21827], Fricke 1994:4 [ref. 21828]). •Synonym of Enneapterygius philippinus (Peters 1869) -- (Fricke 1997:274, 570 [ref. 23339], Randall 2005:473 [ref. 28239], Fricke 2009:41 [ref. 30394]). Current status: Synonym of Enneapterygius philippinus (Peters 1868). Tripterygiidae: Tripterygiinae. Habitat: marine.

tutuilae, Enneapterygius Jordan [D. S.] & Seale [A.] 1906:418, Fig. 99 [Bulletin of the Bureau of Fisheries v. 25 (for 1905); ref. 2497] Pago Pago, American Samoa. Holotype: USNM 51801. Paratypes: CAS-SU 8695 (2), USNM 164584 (1). Type catalog: Böhlke 1953:96 [ref. 12291], Springer & Orrell 1996:32 [ref. 22274]. •Valid as Enneapterygius tutuilae Jordan & Seale 1906 -- (Fricke 1994:285 [ref. 21827], Fricke 1994:3 [ref. 21828], Fricke 1997:337, 572 [ref. 23339], Randall et al. 1997:364 [ref. 25919], Myers 1999:216 [ref. 23965], Fricke 1999:468 [ref. 24106], Johnson 1999:747 [ref. 25471], Shimojyo & Hayashi 2000:44 [ref. 24903], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1082 [ref. 25182], Hutchins 2001:41 [ref. 25847], Fricke 2001:4 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1082 [ref. 26193], Fricke 2002:191 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830], Manilo & Bogorodsky 2003:S116 [ref. 27377], Myers & Donaldson 2003:638 [ref. 27495], Shibukawa et al. in Kimura & Matsuura 2003:168 [ref. 27510], Randall et al. 2004:24 [ref. 27624], Randall 2005:475 [ref. 28239], Holleman 2005:20 [ref. 28455], Hoese 2006:1523 [ref. 28998], Chiang & Chen 2008:197 [ref. 29932], Fricke 2009:42 [ref. 30394], Fricke et al. 2009:93 [ref. 30213], Imamura 2009:244 [ref. 30434], Patzner et al. 2009:446 [ref. 30396], Motomura et al. 2010:185 [ref. 31256], Allen & Erdmann 2012:771 [ref. 31980], Fricke et al. 2014:146 [ref. 33932], Motomura et al. 2015:28 [ref. 34709], Koeda et al. 2016:83 [ref. 35333], Fricke & Erdmann 2017:23 [ref. 35129], Tashiro et al. 2018:[6] [ref. 35790], Arndt & Fricke 2019:32 [ref. 36883]). Current status: Valid as Enneapterygius tutuilae Jordan & Seale 1906. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: East Africa, Seychelles and western Mascarenes east to Marshall Islands and Society Islands, north to Ryukyu Islands and southern Japan, south to Western Australia and Queensland, New Caledonia and Samoa. Former Red Sea records are now assigned to E. altipinnis Clark 1980. Habitat: marine.

unimaculatus, Enneapterygius Fricke [R.] 1994:5, Figs. 3-4 [Stuttgarter Beiträge zur Naturkunde. Serie A (Biologie). No. 512; ref. 21828] Sanur Beach Reef, Bali, 8°40'S, 115°15'E, Indonesia, depth 0-0.5 meters. Holotype: NTM S.11081-021. Paratypes: NTM S.11081-037 (1). •Valid as Enneapterygius unimaculatus Fricke 1994 -- (Fricke 1997:352, 572 [ref. 23339], Shimojyo & Hayashi 2000:53 [ref. 24903], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1084 [ref. 25182], Williams & Fricke 2001:3534 [ref. 26288], Nakabo 2002:1084 [ref. 26193], Allen & Adrim 2003:53 [ref. 26830], Chiang & Chen 2008:199 [ref. 29932], Fricke 2009:42 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Allen & Erdmann 2012:771 [ref. 31980], Fricke & Erdmann 2017:22, 26 [ref. 35129]). Current status: Valid as Enneapterygius unimaculatus Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: Malaysia, Indonesia, Papua New Guinea, Palau and Taiwan. Habitat: marine.

varius, Blennius Forster [J. R.] in Bloch & Schneider 1801:178 [M. E. Blochii, Systema Ichthyologiae; ref. 471] Northern shore, Ship Cove, outer Queen Charlotte Sound, 41°05'S, 174°14'E, New Zealand, depth 0-2 meters. Neotype: NMNZ P.30201. Authorship follows Fricke 1994:17 [ref. 21827]. Neotype designated by Fricke 1994:347 [ref. 21827]. Based on description in manuscript by Forster (Johann Reinhold Forster, 1729-1798, German explorer and naturalist of Cook's second voyage). •Valid as Forsterygion varium (Forster 1801) -- (Hardy 1989:494 [ref. 12485], Paulin et al. 1989:220 [ref. 24556], Paulin & Roberts 1992:94 [ref. 24625], Fricke 1994:341 [ref. 21827], Fricke 1997:574 [ref. 23339], Jawad & Clements 2004:89 [ref. 27725], Hoese 2006:1523 [ref. 28998], Jawad 2008:2955 [ref. 29958] with Schneider as author, Gomon 2008:686 [ref. 30616], Fricke 2009:45 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1510 [ref. 34299]). Current status: Valid as Forsterygion varium (Forster 1801). Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region; introduced in Tasmania, Australia. Habitat: marine.

velatus, Enneapterygius Tashiro [S.], Senou [H.] & Motomura [H.] 2018:[2], Figs. 1-4 [Ichthyological Research v. 65 (no. 3); ref. 35790] Northeast of Kume-jima Island, Okinawa Islands, Ryukyu Islands, Japan, depth 55 meters. Holotype: KPM-NI 6763. Paratypes: YCM-P 36419 (1). •Valid as Enneapterygius velatus Tashiro, Senou & Motomura 2018. Current status: Valid as Enneapterygius velatus Tashiro, Senou & Motomura 2018. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Ryukyu Islands (Japan); New Britain (Papua New Guinea). Record from new Britain based on G. Allen, pers. comm., Oct. 2018. Habitat: marine.

ventermaculus, Enneapterygius Holleman [W.] 1982:123, Fig. 7 [Annals of the Cape Provincial Museums (Natural History) v. 14 (pt 4); ref. 2187] Rock pool about 10 kilometers south of Sodwana Bay, 27°37'30"S, 33°40'50"E, Zululand, KwaZulu-Natal, South Africa, southwestern Indian Ocean. Holotype: SAIAB [formerly RUSI] 7943. Paratypes: BMNH 1954.4.26.191-196 (6), 1954.4.26.197-208 (12), 1954.4.26.209-214 (6); LACM 38309-5 (6), 38310-10 (43), 38320-9 (2); SAIAB [formerly RUSI] 7944-47 (4, 1, 1, 1). •Valid as Enneapterygius ventermaculus Holleman 1982 -- (Holleman 1986:757 [ref. 5671], Randall 1995:32 [ref. 21853], Randall 1995:309 [ref. 22896], Fricke 1997:572 [ref. 23339], Carpenter et al. 1997:207 [ref. 23922], Manilo & Bogorodsky 2003:S116 [ref. 27377], Holleman 2005:21 [ref. 28455], Fricke 2009:43 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Holleman & Bogorodsky 2012:53 [ref. 31735], Fricke et al. 2013:268 [ref. 32706], Golani & Fricke 2018:143 [ref. 36273]). Current status: Valid as Enneapterygius ventermaculus Holleman 1982. Tripterygiidae: Tripterygiinae. Distribution: Southern Red Sea, western Indian Ocean: East Africa, Transkei (South Africa) to Gulf of Oman east to Pakistan. Habitat: marine.

vexillarius, Enneapterygius Fowler [H. W.] 1946:192, Fig. 56 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 98; ref. 1456] Aguni Shima, Ryukyu Islands, Japan. Holotype (unique): ANSP 72068. Type catalog: Böhlke 1984:165 [ref. 13621]. •Valid as Enneapterygius vexillarius Fowler 1946 -- (Fricke 1997:355, 572 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Motomura et al. 2005:13 [ref. 28338], Fricke 2009:43 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Endo et al. 2010:14 [ref. 30815], Fricke & Erdmann 2017:28 [ref. 35129]). Current status: Valid as Enneapterygius vexillarius Fowler 1946. Tripterygiidae: Tripterygiinae. Distribution: Western North Pacific: Ryukyu Islands, Taiwan and Hong Kong. Habitat: marine.

vulcana, Helcogramma Randall [J. E.] & Clark [E.] 1993:27, Figs. 1-6 [Revue française d'Aquariologie Herpétologie v. 20 (no. 1); ref. 20781] Southern side of Gunung Api, Banda Sea, Indonesia, 6°37'S, 126°38'E, depth 0-1 meter. Holotype: BPBM 35378. Paratypes: AMS I.33528-001 (3); BPBM 32353 (20), 35377 (4); CAS 78521 (3); SMF 26257 (3); USNM 293374 (8). Type catalog: Springer & Orrell 1996:32 [ref. 22274]. See dicussion of gender under Helcogramma. •Valid as Helcogramma vulcanum Randall & Clark 1993 -- (Fricke 1997:496, 580 [ref. 23339], Williams & Fricke 2001:3534 [ref. 26288], Allen & Adrim 2003:53 [ref. 26830]). •Valid as Helcogramma vulcana Randall & Clark 1993 -- (Williams & Howe 2003:174 [ref. 27249], Fricke 2009:51 [ref. 30394], Patzner et al. 2009:447 [ref. 30396], Allen & Erdmann 2012:780 [ref. 31980], Fricke & Erdmann 2017:42 [ref. 35190]). Current status: Valid as Helcogramma vulcana Randall & Clark 1993. Tripterygiidae: Tripterygiinae. Distribution: Banda Sea, Indonesia. Habitat: marine.

waigiensis, Enneapterygius Herre [A. W. C. T.] 1935:434 [Field Museum of Natural History, Publications, Zoölogical Series v. 18 (no. 12); ref. 2109] Near the mouth of Majalibit Inlet, southern coast of Waigeo Island, Indonesia. Holotype: FMNH 17494. Paratypes: CAS-SU 27717 (1). Type catalog: Böhlke 1953:96 [ref. 12291], Ibarra & Stewart 1987:34 [ref. 12367]. •Valid as Enneapterygius waigiensis Herre 1935 -- (Fricke 1994:3 [ref. 21828]). •Synonym of Enneapterygius minutus (Günther 1877) -- (Fricke 1994:243 [ref. 21827]). •Synonym of Enneapterygius philippinus (Peters 1868) -- (Fricke 1997:275, 570 [ref. 23339], Fricke 2009:41 [ref. 30394]). Current status: Synonym of Enneapterygius philippinus (Peters 1868). Tripterygiidae: Tripterygiinae. Habitat: marine.

whero, Ruanoho Hardy [G. S.] 1986:161, Pl. 2 (bottom 2); Fig. 5C [Journal of the Royal Society of New Zealand v. 16 (no. 2); ref. 6139] Te Raekaihau Point, Lyall Bay, Wellington, New Zealand. Holotype: NMNZ P.17240. Paratypes: BMNH 1985.6.19.1-5 (5); SAIAB [formerly RUSI] 23786 (5); NMNZ (113, 15 lots); USNM 272606 (5). Type catalog: Hardy 1990:14 [ref. 18942], Springer & Orrell 1996:32 [ref. 22274]. •Synonym of Ruanoho decemdigitatus (Clarke 1879) as a color morph -- (Fricke 1994:521 [ref. 21827], Fricke 1997:585 [ref. 23339]). •Valid as Ruanoho whero Hardy 1986 -- (Paulin et al. 1989:219 [ref. 24556], Paulin & Roberts 1992:83 [ref. 24625], Jawad & Clements 2004:87 [ref. 27725], Wellenreuther & Clements 2008:575 [ref. 30937], Fricke 2009:33 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1522 [ref. 34299]). Current status: Valid as Ruanoho whero Hardy 1986. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

whitleyi, Tripterygium Scott [E. O. G.] (ex Johnston) 1977:148 [Papers and Proceedings of the Royal Society of Tasmania v. 111; ref. 7283] Leven River, northwestern Tasmania, Australia. No types known. Original genus should have been Tripterygion. •Synonym of Norfolkia clarkei (Morton 1888) -- (Gomon et al. 1994:738 [ref. 22532]). •Synonym of Trinorfolkia clarkei (Morton 1888) -- (Fricke 1994:541 [ref. 21827], Fricke 1997:586 [ref. 23339], Hoese 2006:1527 [ref. 28998], Fricke 2009:35 [ref. 30394]). Current status: Synonym of Trinorfolkia clarkei (Morton 1888). Tripterygiidae: Tripterygiinae. Habitat: marine.

wilki, Enneanectes Victor [B. C.] 2013:55, Figs. 10-16 [Journal of the Ocean Science Foundation v. 7; ref. 32783] Dominica, Roseau (15.28°, -61.38°). Holotype: UF 185610. Paratypes: UF. •Valid as Enneanectes wilki Victor 2013 -- (Victor 2017:69 [ref. 35549], Victor 2019:13 [ref. 36450]). Current status: Valid as Enneanectes wilki Victor 2013. Tripterygiidae: Tripterygiinae. Distribution: Western Atlantic: Dominica to Tobago. Habitat: marine.

williamsi, Enneapterygius Fricke [R.] 1997:361, Fig. 73 [ ref. 23339] Ile Haute, Ouvéa Atoll, western end of island, 20°29'12"S, 166°19'18"E, Loyalty Islands, New Caledonia, depth 3-10 meters. Holotype: USNM 323821. Paratypes: SMNS 18700 (2); USNM 319881 (2), 323786 (2), 343948-52 (27, 5, 21, 1, 21), 344098 (10); plus non-type material. Type catalog: Fricke 2005:68 [ref. 29864]. •Valid as Enneapterygius williamsi Fricke 1997 -- (Fricke 2001:5 [ref. 25985], Williams & Fricke 2001:3534 [ref. 26288], Fricke 2002:192 [ref. 29865], Randall et al. 2004:24 [ref. 27624], Randall 2005:476 [ref. 28239], Fricke 2009:43 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Fricke & Erdmann 2017:25, 27 [ref. 35129]). Current status: Valid as Enneapterygius williamsi Fricke 1997. Tripterygiidae: Tripterygiinae. Distribution: Loyalty Islands, Vanuatu and Tonga. Habitat: marine.

williamsi, Helcogramma Chiang [M.-C.] & Chen [I-S.] 2012:59, Figs. 1-2 [ZooKeys No. 216; ref. 32986] Feng-chui-sha, Hengchun Township, Pingtung County, Taiwan. Holotype: NTOU-P 2012-02-002. Paratypes: NTOU-P 2012002-001 (1), 003 (2), 004 (1), 005 (2). •Valid as Helcogramma williamsi Chiang & Chen 2012 -- (Fricke & Erdmann 2017:43 [ref. 35190]). Current status: Valid as Helcogramma williamsi Chiang & Chen 2012. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific. Habitat: marine.

xanthosoma, Tripterygion Zander [C. D.] & Heymer [A.] 1970:365, Fig. [Vie et Milieu, Série A: Biologie Marine v. 21 (no. 2-A); ref. 394] Banyuls-sur-Mer, France, Mediterranean Sea. Holotype: ZMH 4238. Paratypes: MNHN 1975-0061 (4); ZMH 4239-40 (1, 1). Type catalog: Wilkens 1977:160 [ref. 22013]. May date to 1971. •Valid as Tripterygion xanthosoma Zander & Heymer 1970 -- (Wheeler 1973:531 [ref. 7190], Fricke et al. 2007:97 [ref. 29533], Fricke 2009:44 [ref. 30394], Patzner et al. 2009:449 [ref. 30396]). •Synonym of Tripterygion delaisi Cadenat & Blache 1970, but a valid subspecies -- (Zander in Whitehead et al. 1986:1120 [ref. 13677], Ahnelt 1991:56 [ref. 19120]). •Synonym of Tripterygion delaisi Cadenat & Blache 1970 -- (Wirtz 1990:916 [ref. 19374], Fricke 1997:587 [ref. 23339]). •Status with delaisi is uncertain -- (Hastings & Springer 2009:11 [ref. 31156]). Current status: Synonym of Tripterygion delaisi Cadenat & Blache 1970. Tripterygiidae: Tripterygiinae. Distribution: Mediterranean Sea endemic [if valid]. Habitat: marine.

xenogrammus, Ucla Holleman [W.] 1993:5, Figs. 5-6 [The J.L.B Smith Institute of Ichthyology Special Publication No. 55; ref. 21185] Lizard Island, Great Barrier Reef, Australia, depth 3-15 meters. Holotype: ROM 45272. Paratypes: AMS (13 lots, 64) (1 lot, ?); BPBM (5 lots, 18); CAS (3 lots, 3); ROM (2 lots, 7), (1 lot, ?); USNM (141 spec., 57 lots) [263652 (1) missing]. Type catalog: Springer & Orrell 1996:32-33 [ref. 22274]. Name first used by Rosenblatt 1959 (unpublished dissertation). •Valid as Ucla xenogrammus Holleman 1993 -- (Fricke 1994:554 [ref. 21827], Fricke 1997:530, 587 [ref. 23339], Randall et al. 1997:365 [ref. 25919], Myers 1999:217 [ref. 23965], Johnson 1999:747 [ref. 25471], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Nakabo 2000:1078 [ref. 25182], Allen 2000:116 [ref. 25868], Hutchins 2001:42 [ref. 25847], Fricke 2001:3 [ref. 25985], Williams & Fricke 2001:3535 [ref. 26288], Nakabo 2002:1078 [ref. 26193], Fricke 2002:196 [ref. 29865], Allen & Adrim 2003:53 [ref. 26830], Myers & Donaldson 2003:639 [ref. 27495], Randall et al. 2004:24 [ref. 27624], Randall 2005:479 [ref. 28239], Hoese 2006:1528 [ref. 28998], Fricke 2009:52 [ref. 30394], Imamura 2009:244 [ref. 30434], Patzner et al. 2009:449 [ref. 30396], Motomura et al. 2010:190 [ref. 31256], Allen & Erdmann 2012:781 [ref. 31980], Larson et al. 2013:179 [ref. 32988], Fricke et al. 2014:147 [ref. 33932]). Current status: Valid as Ucla xenogrammus Holleman 1993. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: Cargados Carajos, Christmas Island, east to Andaman Sea to Caroline Islands and Rapa, north to Ryukyu Islands, north to Australia and New Caledonia. Habitat: marine.

yaldwyni, Notoclinops Hardy [G. S.] 1987:170, Figs. 3-4 [Journal of the Royal Society of New Zealand v. 17 (no. 2); ref. 6027] Princess Bay, Wellington, New Zealand, depth 6 meters. Holotype: NMNZ P.16102. Paratypes: BMNH 1986.11.25.1 (1); NMNZ (76, 16 lots); USNM 283758 (1). Type catalog: Hardy 1990:13 [ref. 18942], Springer & Orrell 1996:33 [ref. 22274]. •Valid as Notoclinops yaldwyni Hardy 1987 -- (Paulin et al. 1989:220 [ref. 24556], Paulin & Roberts 1992:95 [ref. 24625], Fricke 1994:494 [ref. 21827], Fricke 1997:584 [ref. 23339], Fricke 2009:46 [ref. 30394], Patzner et al. 2009:448 [ref. 30396], Feary et al. 2009:1704 [ref. 30938], Stewart & Clements 2015:1518 [ref. 34299]). Current status: Valid as Notoclinops yaldwyni Hardy 1987. Tripterygiidae: Tripterygiinae. Distribution: Southwestern Pacific: New Zealand region. Habitat: marine.

ziegleri, Enneapterygius Fricke [R.] 1994:8, Figs. 5-10 [Stuttgarter Beiträge zur Naturkunde. Serie A (Biologie). No. 512; ref. 21828] Sanur Beach Reef, Bali, 8°40'S, 115°15'E, Indonesia. Holotype: NTM S.10689-026. Paratypes: NTM S.10689-041 to 043 (1, 18, 12), S.10695-004 (14), S.11081-032 (85, not 84), S.11127-007 (1); SMNS 15354 (10). Type catalog: Fricke 2005:68 [ref. 29864]. Some type information was corrected from information provided by the NTM. •Valid as Enneapterygius ziegleri Fricke 1994 -- (Fricke 1994:285 [ref. 21827], Fricke 1997:366, 572 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Hutchins 2001:41 [ref. 25847], Williams & Fricke 2001:3534 [ref. 26288], Allen & Adrim 2003:53 [ref. 26830], Hoese 2006:1523 [ref. 28998], Fricke 2009:43 [ref. 30394], Patzner et al. 2009:446 [ref. 30396], Allen & Erdmann 2012:771 [ref. 31980], Fricke & Erdmann 2017:23, 27, 28 [ref. 35129]). Current status: Valid as Enneapterygius ziegleri Fricke 1994. Tripterygiidae: Tripterygiinae. Distribution: Indo-West Pacific: Indonesia, Philippines and Australia. Habitat: marine.