Institute for Biodiversity Science and Sustainability
Online Version, Updated 4 November 2019

SEARCH RESULTS FROM

Eschmeyer's Catalog of Fishes
Select the database to search:
GENERA   SPECIES   REFERENCES
Include unavailable names

         Comments: ,

    Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the family or subfamily Zoarcidae: [ 416 ] records


abei, Davidijordania Matsubara [K.] 1936:382, Fig. [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 48 (no. 7); ref. 17610] Off coast of Owase, Kumano-nada Sea, Mie Prefecture, Japan, depth about 200 fathoms. Holotype: FAKU 1805. Paratypes: FAKU 1803-04 (2) [1804 apparently lost]. •Valid as Davidijordania abei Matsubara 1936 -- (Toyoshima in Masuda et al. 1984:305 [ref. 6441], Anderson 1994:116 [ref. 21438], Nakabo 2000:1036 [ref. 25182], Shinohara et al. 2001:328 [ref. 25995], Nakabo 2002:1036 [ref. 26193]). •Valid as Japonolycodes abei (Matsubara 1936) -- (Shinohara et al. 2002:298 [ref. 25992], Anderson & Fedorov 2004:14 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Japonolycodes abei (Matsubara 1936). Zoarcidae: Lycodinae. Distribution: Western North Pacific: Japan. Habitat: marine.

acrogeneius, Opaeophacus Bond [C. E.] & Stein [D. L.] 1984:523, Fig. 1 [Proceedings of the Biological Society of Washington v. 97 (no. 3); ref. 5310] Off Seguam Island, Bering Sea, 52°42'N, 172°15'W, depth 500-800 meters. Holotype: USNM 260320. Paratypes: CAS 52802 (1), OS 10000 (1), USNM 260321 (2, 1 c&s). Type catalog: Springer & Anderson 1997:13 [ref. 22953]. •Valid as Opaeophacus acrogeneius Bond & Stein 1984 -- (Anderson 1994:120 [ref. 21438], Mecklenburg et al. 2002:685 [ref. 25968], Anderson & Fedorov 2004:9 [ref. 27603], Balushkin et al. 2011:1027 [ref. 32449], Parin et al. 2014:427 [ref. 33547]). Current status: Valid as Opaeophacus acrogeneius Bond & Stein 1984. Zoarcidae: Gymnelinae. Distribution: North Pacific: Bering Sea. Habitat: marine.

acropterus, Caneolepis Lahille [F.] 1908:431, Pl. 7 (figs. 1-10) [Anales del Museo Nacional de Historia Natural de Buenos Aires (Serie 3) v. 9; ref. 5869] Cape San Antonio, Argentina, depth 60-80 fathoms. Syntypes: Mus. Nac. Buenos Aires (4, 1 skel.) •Synonym of Iluocoetes fimbriatus Jenyns 1842 -- (Gosztonyi 1977:213 [ref. 6103], Anderson & Fedorov 2004:14 [ref. 27603]). Current status: Synonym of Iluocoetes fimbriatus Jenyns 1842. Zoarcidae: Lycodinae. Habitat: marine.

adocetus, Dieidolycus Anderson [M. E.] 1994:121, Figs. 1-2 [Records of the Australian Museum v. 4 (no. 2); ref. 21460] Bismarck Sea, 3°41.5'S, 151°52.2'E, depth 1957 meters. Holotype: AMS I.32236-001. Paratypes: AMS I.32236-002 (1). •Valid as Dieidolycus adocetus Anderson 1994 -- (Anderson & Pequeño 1998:4 [ref. 23482], Anderson 2001:3494 [ref. 26279], Anderson & Fedorov 2004:12-13 [ref. 27603], Anderson & Møller 2007:183 [ref. 29401], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Dieidolycus adocetus Anderson 1994. Zoarcidae: Lycodinae. Distribution: Western South Pacific: Bismarck Sea. Habitat: marine.

adolfi, Lycodes Nielsen [J. G.] & Fosså [S. A.] 1993:39, Figs. 1-2 [Cybium v. 17 (no. 1); ref. 20655] Greenland, 68°49'N, 19°44'W, depth 1371-1380 meters. Holotype: ZMUC P761186. Paratypes: ZIN 50515 [ex.ZUMC 761188] (1); ZMUC P761187 (1), P751189-90 (3), P761191 (1), P761192 (1). Type catalog: Balushkin et al. 2011:987 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. •Valid as Lycodes adolfi Nielsen & Fosså 1993 -- (Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:23 [ref. 27603], Møller et al. 2010:60 [ref. 31169], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin et al. 2011:987 [ref. 32449], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:140 [ref. 33463], Parin et al. 2014:406 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:63, 195 [ref. 36199], Mecklenburg et al. 2016:160 [ref. 34440], Mecklenburg et al. 2018:321 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes adolfi Nielsen & Fosså 1993. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

agnostus, Lycodes Jensen [A. S.] 1902:209 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Kara-Havet, Kara Sea, depth 46-100 fathoms. Syntypes: ZIN 40714 [ex. ZUMC P22] (1); ZMUC 3, 6, 11, 16, 20, 22, 24-27, 29. Type catalog: Nielsen 1974:76 [ref. 9588], Balushkin et al. 2011:1006 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. Specimens ZMUO J4490 (3) are labeled types but are not types according to Pethon 1969:11 [ref. 19268]. •Synonym of Lycodes polaris (Sabine 1824) -- (Andriashev 1973:545 [ref. 9214], Scott & Scott 1988:410 [ref. 25518], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:710 [ref. 25968], Anderson & Fedorov 2004:28 [ref. 27603], Balushkin et al. 2011:1006 [ref. 32449], Parin et al. 2014:418 [ref. 33547]). Current status: Synonym of Lycodes polaris (Sabine 1824). Zoarcidae: Lycodinae. Habitat: marine.

agulhensis, Lycodes Andriashev [A. P.] 1959:465, Figs. 1-3 [Zoologicheskii Zhurnal v. 38 (no. 3); ref. 11975] Agulhas Bank, 35°44.2-46'S, 22°33.5 -36.2'E, South Africa, southeastern Atlantic, depth 1400 meters. Holotype (unique): ZIN 35812. Type catalog: Balushkin et al. 2011:987 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. •Synonym of Lycodes terraenovae Collett 1896 -- (Møller 1997:48 [ref. 22879], Møller 2000:847 [ref. 24585], Carl 2002:65 [ref. 26925], Anderson & Fedorov 2004:30 [ref. 27603]). •Valid as Lycodes agulhensis Andriachev 1959 -- (Anderson 1986:342 [ref. 5633], Anderson & Almeida 1990:888 [ref. 19328], Anderson 1994:118 [ref. 21438], Balushkin et al. 2011:987 [ref. 32449]). Current status: Valid as Lycodes agulhensis Andriashev 1959. Zoarcidae: Lycodinae. Distribution: Agulhas Bank, South Africa. Habitat: marine.

akuugun, Lycodes Stevenson [D. E.] & Orr [J. W.] 2006:78, Figs. 1-2 [Copeia 2006 (no. 1); ref. 28543] South of Seguam Island, 52.210°N, 172.206°W, central Aleutian Islands, depth 347 meters. Holotype: UW 112519. Paratypes: CAS 222668 (1); LACM 56292-1 (1); UW 112520 (1), 112521 (2), 112522 (1), 112523 (1 c&s), 112524 (1), 112525 (1); USNM 384244 (1). •Valid as Lycodes akuugun Stevenson & Orr 2006 -- (Balushkin et al. 2011:1026 [ref. 32449], Page et al. 2013:162 [ref. 32708]). Current status: Valid as Lycodes akuugun Stevenson & Orr 2006. Zoarcidae: Lycodinae. Distribution: Central Aleutian Islands. Habitat: marine.

alalonga, Bothrocaropsis Garman [S.] 1899:127, Pl. 32 (fig. 2) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Off Gulf of California, Mexico, 27°34'N, 110°53'40"W, Albatross station 3436, depth 905 fathoms. Lectotype: MCZ 28694. Paralectotypes: MCZ 117365 [ex MCZ 28694] (1). Lectotype selected by Anderson et al. 2009:184 [ref. 30175]. •Valid as Bothrocara alalongum (Garman 1899) -- (Anderson 1994:116 [ref. 21438]). •See also Pequeño 1997:83 [ref. 23536]. •Synonym of Bothrocara molle Bean 1890 -- (Anderson & Fedorov 2004:11 [ref. 27603], Anderson 2006:2 [ref. 28576], Anderson et al. 2009:179 [ref. 30175], Parin et al. 2014:385 [ref. 33547]). Current status: Synonym of Bothrocara molle Bean 1890. Zoarcidae: Lycodinae. Habitat: marine.

albeolus, Lycenchelys Andriashev [A. P.] 1958:175, Fig. 1 (no. 1), Fig. 2 [Voprosy Ikhtiologii v. 11; ref. 12042] Northern part of Kuril-Kamchatka Trench, 49°49'N, 157°45'E, depth 3960-4070 meters. Holotype (unique): ZIN 34668. Type catalog: Balushkin et al. 2011:978 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667] •Valid as Lycenchelys albeola Andriashev 1958 -- (Toyoshima 1985:154 [ref. 5722] as albeolus, Anderson 1994:117 [ref. 21438], Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:15 [ref. 27603], Balushkin et al. 2011:978 [ref. 32449], Parin et al. 2014:397 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys albeola Andriashev 1958. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Kuril-Kamchatka Trench. Habitat: marine.

albolineatus, Lycodes Andriashev [A. P.] 1955:397, Figs. 4-6 [Trudy Zoologicheskogo Instituta, Akademii Nauk SSSR v. 21; ref. 9472] Near Cape Lopatka, southeastern coast of Kamchatka, 50°59'N, 157°54"E, depth 241-270 meters. Holotype: ZIN 34582. Paratypes: BMNH [ex ZIN] (1), ZIN 34581 (4, now 3). Type catalog: Balushkin et al. 2011:987 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667].. Holotype established in Fig. 4, 605 mm. •Valid as Lycodes albolineatus Andriashev 1955 -- (Toyoshima 1985:232 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Møller 2001:973 [ref. 25812], Tokranov & Orlov 2002:605 [ref. 26464], Anderson & Fedorov 2004:23 [ref. 27603], Balushkin et al. 2011:987 [ref. 32449], Parin et al. 2014:406 [ref. 33547]). Current status: Valid as Lycodes albolineatus Andriashev 1955. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

albomaculatus, Lycenchelys Toyoshima [M.] in Amaoka et al. 1983:141, Pl. 92; Figs. 16-19 [Fishes from the north-eastern Sea of Japan and the Okhotsk Sea off Hokkaido; ref. 8379] Off Kamaishi, Iwate Prefecture, Japan. Holotype: HUMZ 59531. Paratypes: HUMZ (27 specimens). •Valid as Lycenchelys albomaculata Toyoshima 1983 [albomaculata is correct] -- (Toyoshima in Amaoka et al. 1983:269, 333 [ref. 8379], Toyoshima in Masuda et al. 1984:307 [ref. 6441] as albomaculatus, Toyoshima 1985:159 [ref. 5722] as albomaculatus, Anderson 1994:117 [ref. 21438], Shinohara et al. 1996:180 [ref. 26605], Fedorov 1995:130 [ref. 21958], Nakabo 2000:1035 [ref. 25182], Nakabo 2002:1035 [ref. 26193], Anderson & Fedorov 2004:15 [ref. 27603], Shinohara & Anderson 2007:64 [ref. 29111], Balushkin et al. 2011:978 [ref. 32449], Parin et al. 2014:397 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys albomaculata Toyoshima 1983. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

albonotata, Petroschmidtia Taranetz [A. Ya.] & Andriashev [A. P.] 1934:507, Figs. 1-2 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 2 (no. 8); ref. 4338] Okhotsk Sea, 56°35' to 56°37'N, 142°46' to 142°48'E, depth 202-220 meters. Syntypes: (5) ZIN 24606 (1), 24607 (3). Type catalog: Balushkin et al. 2011:1021 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. Four of the original six types were found, with one paratype as ZIN 24606 and the putative holotype mixed with two paratypes now as ZIN 24607 -- see Anderson & Fedorov 2004:23 [ref. 27603]. •Valid as Lycodes albonotatus (Taranetz & Andriashev 1934) -- (Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:23 [ref. 27603], Tohkairin et al. 2015:12 [ref. 35331]). •Valid as Petroschmidtia albonotata Taranetz & Andriashev 1934 -- (Lindberg & Krasyukova 1975:166 [ref. 7348], Toyoshima in Amaoka et al. 1983:259, 329 [ref. 8379], Toyoshima in Masuda et al. 1984:306 [ref. 6441], Toyoshima 1985:140 [ref. 5722], Nakabo 2000:1030 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1030 [ref. 26193], Fedorov et al. 2003:119 [ref. 27579], Balushkin et al. 2011:1021 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Shinohara et al. 2014:267 [ref. 33330], Parin et al. 2014:427 [ref. 33547]). Current status: Valid as Petroschmidtia albonotatus Taranetz & Andriashev 1934. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

albus, Lycodes Vaillant [L. L.] 1888:309, Pl. 26 (figs. 1-1c) [Expéditions scientifiques du "Travailleur" et du "Talisman"; ref. 4496] Northeastern Atlantic between Azores and France, depth 3975 meters. Lectotype: MNHN 1886-0590 (poor condition). Paralectotypes: MNHN 1886-0591 (1). Type catalog: Bauchot 1967:62 [ref. 20734]. Lectotype selected by Eschmeyer 1998:69 [ref. 23416]. •Valid as Lycenchelys albus (Vaillant 1888) -- (Andriashev 1973:541 [ref. 9214], Andriashev in Whitehead et al. 1986:1133 [ref. 13677]). •Valid as Lycenchelys alba (Vaillant 1888) -- (Anderson 1994:117 [ref. 21438], Møller 1999:327 [ref. 24052], Moore et al. 2003:228 [ref. 27076], Anderson & Fedorov 2004:15 [ref. 27603], Møller et al. 2010:59 [ref. 31169], Balushkin et al. 2011:1026 [ref. 32449], Anderson in Carpenter & De Angelis 2016:2760 [ref. 34618]). Current status: Valid as Lycenchelys alba (Vaillant 1888). Zoarcidae: Lycodinae. Distribution: North Atlantic. Habitat: marine.

alepidotum, Pachycara Anderson [M. E.] & Mincarone [M. M.] 2006:23, Fig. 2 [Zootaxa No. 1177; ref. 28658] Off Rio Grande do Sul state, Brazil, 34°31.4'S, 51°45.8'W, trap in depth of 807 meters. Holotype: MOVI 21033. Paratypes: MOVI 21035 (1); SAIAB 75089 (1). Plus additional material. •Valid as Pachycara alepidotum Anderson & Mincarone 2006 -- (Møller & King 2007:591 [ref. 29259], Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara alepidotum Anderson & Mincarone 2006. Zoarcidae: Lycodinae. Distribution: Southwestern Atlantic: southern Brazil. Habitat: marine.

alepidotus, Derepodichthys Gilbert [C. H.] 1896:456 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Off Queen Charlotte Island, British Columbia, Canada, 52°39'30"N, 132°38'W, Albatross station 3342, depth 1588 fathoms. Holotype (unique): USNM 48615 (poor condition). Type catalog: Springer & Anderson 1997:13 [ref. 22953]. •Valid as Derepodichthys alepidotus Gilbert 1896 -- (Anderson & Hubbs 1981 [ref. 5476], Toyoshima 1985:144 [ref. 5722], McAllister 1990:165 [ref. 14674], Anderson 1994:116 [ref. 21438], Mecklenburg et al. 2002:691 [ref. 25968], Anderson & Fedorov 2004:12 [ref. 27603], Balushkin et al. 2011:975 [ref. 32449]). Current status: Valid as Derepodichthys alepidotus Gilbert 1896. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

altus, Lycenchelys Toyoshima [M.] 1985:158, Figs. 14-15, 31d [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] South of Buldir Island, Aleutian Islands, 52°03.69'N, 175°39.0'E, depth 336 meters. Holotype (unique): HUMZ 88704. •Valid as Lycenchelys alta Toyoshima 1985 [alta is correct] -- (Toyoshima 1985:158 [ref. 5722] as altus, Anderson 1994:117 [ref. 21438], Anderson 1995:59 [ref. 21928], Shinohara & Matsuura 1998:154 [ref. 23382], Mecklenburg et al. 2002:699 [ref. 25968], Anderson & Fedorov 2004:15 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449], Parin et al. 2014:397 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys alta Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: North Pacific. Habitat: marine.

amberensis, Lycenchelys Tomo [A. P.], Marschoff [E.] & Torno [A. E.] 1977:4 [Instituto Antártico Argentino, Direccion Nacional del Antartico, Contribucion No. 221; ref. 8870] Vicinity of Isla Amberes, Antarctic Peninsula, 64°40'S, 63°20'W. Holotype (unique): IAA 2. •Valid as Ophthalmolycus amberensis (Tomo, Marschoff & Torno 1977) -- (Anderson 1988:82 [ref. 7304], Anderson 1990:8 [ref. 16143], Anderson 1990:273 [ref. 21695], Anderson 1992:8 [ref. 18912], Miller 1993:655 [ref. 21297], Anderson 1994:120 [ref. 21438], Pequeño 1997:83 [ref. 23536], Anderson & Fedorov 2004:33 [ref. 27603], Anderson 2006:10 [ref. 28576], Matallanas 2009:55 [ref. 30103], Matallanas 2011:566 [ref. 30839], Balushkin et al. 2011:1019 [ref. 32449]). Current status: Valid as Ophthalmolycus amberensis (Tomo, Marschoff & Torno 1977). Zoarcidae: Lycodinae. Distribution: Southern Ocean: circum-Antarctica. Habitat: marine.

americanus, Blennius Bloch [M. E.] & Schneider [J. G.] 1801:171 [M. E. Blochii, Systema Ichthyologiae; ref. 471] American seas. No types known. •Valid as Macrozoarces americanus (Bloch & Schneider 1801) -- (Robins & Ray 1986:237 [ref. 23100], Scott & Scott 1988:412 [ref. 25518], McAllister 1990:156 [ref. 14674], Balushkin et al. 2011:952 [ref. 32449]). •Valid as Zoarces americanus (Bloch & Schneider 1801) -- (Anderson 1994:120 [ref. 21438], Klein-MacPhee & Collette in Collette & Klein-MacPhee 2002:470 [ref. 26158], Anderson & Fedorov 2004:2 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Chereshev et al. 2007:555 [ref. 29371], Voskoboinikova et al. 2010:704 [ref. 31095], Page et al. 2013:163 [ref. 32708], Afonso et al. 2016:68, 294 [ref. 36199]). Current status: Valid as Zoarces americanus (Bloch & Schneider 1801). Zoarcidae: Zoarcinae. Distribution: Northwestern Atlantic: Labrador (Canada) southward. Habitat: marine.

americanus, Enchelyopus Gronow [L. T.] in Gray 1854:101 [Catalogue of fish collected and described by Laurence Theodore Gronow; ref. 1911] American seas. No types known. If species belongs to Zoarces, the name is preoccupied by Zoarces americanus (Bloch & Schneider 1801) -- see Anderson & Fedorov 2004:48 [ref. 27603]. •Apparently in genus Zoarces -- (Anderson & Fedorov 2004:48 [ref. 27603]). Zoarcidae: Zoarcinae. Habitat: marine.

andersoni, Gymnelus Chernova [N. V.] 1998:738, Figs. 1, 2b, 3a, 4a, 5a [Voprosy Ikhtiologii v. 38 (no. 6); ref. 24169] Barents Sea, Arctic Ocean, 75°00"N, 33°30'E, depth 213 meters. Holotype: ZIN 14140. Paratypes: ZIN 14137 (2), 14156 (5, now 4), 14160 (3), 28252 (1), 29452 (5, now 4), 29455 (1), 32068 (1), 32072 (1), 32770 (1), 51391-92 (2, 1). Plus additional non-type material. Type catalog: Balushkin et al 2011:961 [ref. 32499], Balushkin et al. 2012:156 [ref. 33667]. Anderson in Anderson & Fedorov 2004:5 [ref. 27603] believes this is probably a synonym of G. retrodorsalis. •Synonym of Gymnelus retrodorsalis Le Danois 1913 with question -- (Mecklenburg et al. 2011:124, 128 [ref. 31212]). •Valid as Gymnelus andersoni Chernova 1998 -- (Chernova 1999:306 [ref. 24868], Chernova 2000:7 [ref. 24885], Anderson & Fedorov 2004:5 [ref. 27603], Balushkin et al.2011:961 [ref. 32499], Dolgov 2013:919 [ref. 33112], Parin et al. 2014:391 [ref. 33547]). •Valid as Gymnelus retrodorsalis Le Danois 1913 with question -- (Mecklenburg & Anderson 2015:284 [ref. 33742]), Current status: Valid as Gymnelus andersoni Chernova 1998. Zoarcidae: Gymnelinae. Distribution: Arctic: northern Eurasian seas. Habitat: marine.

andersoni, Ophthalmolycus Matallanas [J.] 2009:56, Figs. 1-2 [Zootaxa No. 2027; ref. 30103] Paradise Bay, Antarctic Peninsula, 64°50'37"S, 62°58'09"W, depth 286 meters. Holotype: UAB B03BSZ25. Paratypes: UAB B03BSZ9 (1), B03BSZ99 (1). •Valid as Ophthalmolycus andersoni Matallanas 2009 -- (Matallanas 2011:566 [ref. 30839]). Current status: Valid as Ophthalmolycus andersoni Matallanas 2009. Zoarcidae: Lycodinae. Distribution: Antarctic Peninsula, Antarctic Ocean. Habitat: marine.

andersoni, Pachycara Møller [P. R.] 2003:357, Fig. 2 [Copeia 2003 (no. 2); ref. 26908] Arabian Sea, 16°13'N, 60°16'E, Indian Ocean, depth 4050 meters. Holotype: ZMUC P764687. Paratypes: SMF 28864 (3); SAIAB [formerly RUSI] 64109 (1); ZMH 9186-9188 (2), 9189 (1), 9190 (1), 9191-9198 (8), 9200-9203 (4); ZMUC P764678-81 (4), 764682 (1), 764684-86 (3), 764688-89 (2). •Valid as Pachycara andersoni Møller 2003 -- (Anderson & Fedorov 2004:34 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Møller & King 2007:588 [ref. 29259], Shinohara 2012:55 [ref. 32102], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Pachycara andersoni Møller 2003. Zoarcidae: Lycodinae. Distribution: Western Indian Ocean. Habitat: marine.

andersoni, Thermarces Rosenblatt [R. H.] & Cohen [D. M.] 1986:78, Fig. 6 [Transactions of the San Diego Society of Natural History v. 21 (no. 4); ref. 5198] Eastern Pacific, 12°48.85'N, 103°56.60'W, depth 2620 meters. Holotype: MNHN 1985-0400. Paratypes: MNHN 1985-0401 (1). •Valid as Thermarces andersoni Rosenblatt & Cohen 1986 -- (Anderson 1994:120 [ref. 21438], Machida & Hashimoto 2002:1 [ref. 25949], Anderson & Fedorov 2004:37 [ref. 27603] with question, Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Thermarces andersoni Rosenblatt & Cohen 1986. Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

andriashevi, Lycodes Fedorov [V. V.] 1966:160, Figs. 1-2 [Voprosy Ikhtiologii v. 6 (no. 1); ref. 20765] Bering Sea, 54°26'N, 116°13'W, depth 550 meters. Holotype (unique): ZIN 37515. Type catalog: Balushkin et al. 2011:992 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. •Valid as Lycodes andriashevi Fedorov 1966 -- (Toyoshima 1985:235 [ref. 5722]). •Synonym of Lycodes concolor Gill & Townsend 1897 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:725 [ref. 25968], Møller 2001:975 [ref. 25812], Anderson & Fedorov 2004:24 [ref. 27603], Balushkin et al. 2011:992 [ref. 32449], Parin et al. 2014:410 [ref. 33547]). Current status: Synonym of Lycodes concolor Gill & Townsend 1897. Zoarcidae: Lycodinae. Habitat: marine.

andriashevi, Zoarces Parin [N. V.], Grigoryev [S. S.] & Karmovskaya [E. S.] 2005:437, Fig. [Voprosy Ikhtiologii v. 45 (no. 4); ref. 28290] Southwestern Kamchatka Island, 52°02'N, 155°01'E, depth 96 meters. Holotype: ZIN 53485. Paratypes: ZIN 53486 (3), IORAN 02600 (3). Type catalog: Balushkin 2011:952 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. •Valid as Zoarces andriashevi Parin, Grigoryev & Karmovskaya 2005 -- (Chereshev et al. 2007:555 [ref. 29371], Voskoboinikova et al. 2010:704 [ref. 31095], Balushkin et al. 2011:952 [ref. 32449], Parin et al. 2014:429 [ref. 33547]). Current status: Valid as Zoarces andriashevi Parin, Grigoryev & Karmovskaya 2005. Zoarcidae: Zoarcinae. Distribution: Western North Pacific: off Kamchatka. Habitat: marine.

anguillaris, Blennius Peck [W. D.] 1804:55, Pl. 2 (fig. 3) [Memoirs of the American Academy of Arts and Sciences v. 2 (pt. 2) (art. 1); ref. 18525] New Hampshire, U.S.A. No types known. •Synonym of Macrozoarces americanus (Bloch & Schneider 1801) -- (Scott & Scott 1988:412 [ref. 25518]). •Synonym of Zoarces americanus (Bloch & Schneider 1801) -- (Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:2 [ref. 27603]). Current status: Synonym of Zoarces americanus (Bloch & Schneider 1801). Zoarcidae: Zoarcinae. Habitat: marine.

anguis, Lycodes Garman [S.] 1899:133, Pl. 30 (fig. 1) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Gulf of California, Mexico, 26°48'N, 110°45'W, Albatross station 3435, depth 1571 meters. Lectotype: MCZ 115068 [ex MCZ 28687]. Paralectotypes: MCZ 28687-88 (2, now 0); ZIN 31124 (1). Type locality: Balushkin et al. 2011:981 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667]. Lectotype selected by Anderson 1995:98 [ref. 21928]. •Synonym of Lycenchelys porifer (Gilbert 1890) -- (Anderson 1994:117 [ref. 21438], Anderson 1995:96 [ref. 21928], Anderson & Fedorov 2004:19 [ref. 27603], Balushkin et al. 2011:981 [ref. 32449]). Current status: Synonym of Lycenchelys porifer (Gilbert 1890). Zoarcidae: Lycodinae. Habitat: marine.

antarctica, Gosztonyia Matallanas [J.] 2009:16, Figs. 1-5 [Polar Biology v. 31 (no. 1); ref. 30174] Bellingshausen Sea, 69°82'S, 77°73'W, Antarctica, depth 615 meters (trap). Holotype: UAB-B03BSZ2. Paratypes: UAB-B03BSZ2B (1), B03BSZ3 (1), B03BSZ4 (1). •Valid as Gosztonyia antarctica Matallanas 2009 -- (Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Gosztonyia antarctica Matallanas 2009. Zoarcidae: Lycodinae. Distribution: Bellingshausen Sea, Antarctica. Habitat: marine.

antarctica, Santelmoa Matallanas [J.], Corbella [C.] & Møller [P. R.] 2012:xx, Figs. 5-6 [Polar Biology v. 35 (no. 9); ref. 31950] Bellingshausen Sea, station MB32A, 69°47'S, 86°27'W, depth 1837 meters. Holotype: UAB B06MB32Z27. Paratypes: UAB B06MB32Z28 (1), B06MB32Z8 (1). •Valid as Santelmoa antarctica Matallanas, Corbella & Møller 2012. Current status: Valid as Santelmoa antarctica Matallanas, Corbella & Møller 2012. Zoarcidae: Lycodinae. Distribution: Bellingshausen Sea, Southern Ocean. Habitat: marine.

antarcticus, Gymnelichthys Fischer [J. G.] 1885:61, Pl. 2 (fig. 9) [Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten v. 2; ref. 1333] Locality unknown, erroneously given as "South Georgia" Island. Syntypes: ZMH 3902 (4). •Synonym of Gymnelus viridis (Fabricius 1780) -- (Andriashev 1973:541 [ref. 9214], Anderson 1982:30 [ref. 5520], Anderson & Fedorov 2004:7 [ref. 27603]). Current status: Synonym of Gymnelus viridis (Fabricius 1780). Zoarcidae: Gymnelinae. Habitat: marine.

antarcticus, Lycenchelys Regan [C. T.] 1913:242, Pl. 9 (fig. 3) [Transactions of the Royal Society of Edinburgh v. 49 (pt 2, no. 2); ref. 3651] Off South Orkney Islands, 62°10'S, 41°21'W, depth 1775 fathoms. Holotype (unique): NMSZ 1921.143.0487. Type catalog: Herman et al. 1990:1 [ref. 20727]. •Valid as Lycenchelys antarctica Regan 1913 [sometimes as antarcticus, should be antarctica] -- (Anderson 1988:88 [ref. 7304], Pequeño 1989:48 [ref. 14125], Anderson 1990:258 [ref. 21695], Miller 1993:660 [ref. 21297], Anderson 1994:117 [ref. 21438], Anderson 1995:60 [ref. 21928], Anderson & Fedorov 2004:15 [ref. 27603], Anderson 2006:4 [ref. 28576], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys antarctica Regan 1913. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific to Weddell Sea, Antarctica. Habitat: marine.

antarcticus, Lycodapus Tomo [A. P.] 1982:84, Figs. 63-64 [Publicacion / Dirección Nacional del Antártico, Instituto Antartico Argentino. No. 14; ref. 20550] Off South Orkney Islands. Holotype (unique): IAA 5. •Valid as Lycodapus antarcticus Tomo 1982 -- (Anderson 1988:104 [ref. 7304], Anderson 1990:266 [ref. 21695], Miller 1993:675 [ref. 21297], Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:21 [ref. 27603], Duhamel et al. 2005:320 [ref. 32059], Balushkin et al. 2011:984 [ref. 32449]). Current status: Valid as Lycodapus antarcticus Tomo 1982. Zoarcidae: Lycodinae. Distribution: Southern Ocean: subantarctic, South Shetland and South Orkney islands to Kerguelen Plateau and southerly seamounts. Habitat: marine.

antarcticus, Lycodichthys Pappenheim [P.] 1911:383 [Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1911 (no. 8) (for 10 Oct. 1911); ref. 6534] Gauss winter station, Wilhelm II Coast, Antarctica, 66°15'S, 88°55'E. Syntypes: BMNH 1913.4.15.60-61 (2); SMNS 4607 (1); ZMB 18941-46 (6, 2, 3, 2, 1, 1); ZMH 8157 (1). Type catalog: Wilkens & Dohse 1993:419 [ref. 21161], Fricke 1995:22 [ref. 28100], Fricke 2005:70 [ref. 29864]. •Valid as Lycodichthys antarcticus Pappenheim 1911 -- (Anderson 1988:78 [ref. 7304], Anderson 1990:267 [ref. 21695], Miller 1993:650 [ref. 21297], Anderson 1994:119 [ref. 21438], Pequeño 1997:83 [ref. 23536], Anderson & Fedorov 2004:31 [ref. 27603], Balushkin et al. 2011:1017 [ref. 32449]). Current status: Valid as Lycodichthys antarcticus Pappenheim 1911. Zoarcidae: Lycodinae. Distribution: Southern Ocean: Antarctica. Habitat: marine.

aptera, Andriashevia Fedorov [V. V.] & Neyelov [A. V.] 1978:953 [847], Figs. 1-2 [Voprosy Ikhtiologii v. 18 (no. 5); ref. 1312] 37°29'N, 142°25'E, Pacific coast of Japan, depth 1025 meters. Holotype (unique): ZIN 43449. Type catalog: Balushkin 2011:958 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667]. On p. 847 of English translation. •Valid as Andriashevia aptera Fedorov & Neyelov 1978 -- (Anderson 1994:116 [ref. 21438], Nakabo 2000:1044 [ref. 25182], Nakabo 2002:1044 [ref. 26193], Anderson & Fedorov 2004:3 [ref. 27603], Balushkin et al. 2011:958 [ref. 32449]). Current status: Valid as Andriashevia aptera Fedorov & Neyelov 1978. Zoarcidae: Gymnelinae. Distribution: Western North Pacific. Habitat: marine.

arabica, Pachycara Møller [P. R.] 2003:362, Fig. 3 [Copeia 2003 (no. 2); ref. 26908] Arabian Sea, 16°13'N, 60°16'E, Indian Ocean, depth 4050 meters. Holotype (unique): ZMH 9199. •Valid as Pachycara arabica Møller 2003 -- (Anderson & Fedorov 2004:34 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Møller & King 2007:588 [ref. 29259], Shinohara 2012:55 [ref. 32102], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Pachycara arabica Møller 2003. Zoarcidae: Lycodinae. Distribution: Western Indian Ocean. Habitat: marine.

aratrirostris, Lycenchelys Andriashev [A. P.] & Permitin [Yu. Ye.] 1968:614, Figs. 2-4 [Voprosy Ikhtiologii v. 8 (no. 4, art. 51); ref. 7858] Scotia Sea, off South Shetland Islands, 60°57'S, 54°49'W, Antarctica, depth 570-583 meters. Holotype (unique): ZIN 38342. Type catalog: Balushkin et al. 2011:978 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667].. •Valid as Lycenchelys aratrirostris Andriashev & Permitin 1968 -- (Anderson 1988:89 [ref. 7304], Anderson 1990:259 [ref. 21695], Miller 1993:662 [ref. 21297], Anderson 1994:117 [ref. 21438], Olaso et al. 1997:83 [ref. 25009], Anderson & Fedorov 2004:15 [ref. 27603], Anderson 2006:5 [ref. 28576], Shinohara & Anderson 2007:61 [ref. 29111], Balushkin et al. 2011:978 [ref. 32449]). Current status: Valid as Lycenchelys aratrirostris Andriashev & Permitin 1968. Zoarcidae: Lycodinae. Distribution: Southern Ocean: western Antarctica. Habitat: marine.

arctica, Lycodes vahlii forma Smitt [F. A.] 1900:57 [Annals and Magazine of Natural History (Series 7) v. 5 (no. 25); ref. 18731] No locality. No types known. Described as "Forma arctica (Vahlii typica)." Also appeared in Smitt 1901 [ref. 18732]. •Incertae sedis in Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Zoarcidae.

arcticus, Lycodes palearis Taranetz [A. Ya.] & Andriashev [A. P.] in Andriashev 1937:333, Pl. 1 (figs. 9-10) [Issled. Morey SSSR, Leningrad v. 25; ref. 20242] Gulf of Anadyr, Bering Sea; Chukchi Sea, depth 84-110 meters. Syntypes: ZIN 26885 (1), 29985 (5). Additional material: ZIN 37927 (9). Type catalog: Balushkin et al. 2011:1002 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. Name mentioned in Taranetz & Andriashev 1935:253 (footnote 4) [ref. 15744]. •Synonym of Lycodes palearis Gilbert 1896 -- (Toyoshima 1985:212 [ref. 5722], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:719 [ref. 25968], Anderson & Fedorov 2004:27 [ref. 27603], Mecklenburg et al. 2007:182 [ref. 29448], Balushkin et al. 2011:1002 [ref. 32449], Parin et al. 2014:417 [ref. 33547]). Current status: Synonym of Lycodes palearis Gilbert 1896. Zoarcidae: Lycodinae. Habitat: marine.

argentinus, Lycenchelys Marschoff [E.], Torno [A. E.] & Tomo [A. P.] 1977:4, Fig. [Instituto Antártico Argentino, Direccion Nacional del Antartico, Contribucion No. 222; ref. 8910] West of South Orkney Islands, 60°39'S, 50°39'W, depth 2120 meters. Holotype: IAA No. 3. •Valid as Lycenchelys argentinus Marschoff, Torno & Tomo 1977 -- (Anderson 1988:92 [ref. 7304]). •Valid as Lycenchelys argentina Marschoff, Torno & Tomo 1977 -- (Anderson 1990:260 [ref. 21695], Miller 1993:663 [ref. 21297], Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:15 [ref. 27603], Anderson 2006:7 [ref. 28576], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys argentina Marschoff, Torno & Tomo 1977. Zoarcidae: Lycodinae. Distribution: Southern Ocean: western Antarctica. Habitat: marine.

atacamensis, Lycenchelys Andriashev [A. P.] 1980:1106, Fig. [Zoologicheskii Zhurnal v. 59 (no. 7) (for July 1980); ref. 8759] Eastern Pacific, Peru-Chile Trench, 6°01°N, 81°48'W, depth 5300-5320 meters. Holotype (unique): ZIN 44940. Type catalog: Balushkin et al. 2011:978 [ref. 32449]. •Synonym of Lycenchelys antarcticus Regan 1913 -- (Anderson 1988:88 [ref. 7304]). •Synonym of Lycenchelys antarctica Regan 1913 -- (Anderson 1990:258 [ref. 21695], Anderson 1994:117 [ref. 21438], Anderson 1995:57 [ref. 21928], Anderson & Fedorov 2004:15 [ref. 27603], Anderson 2006:4 [ref. 28576]). •Valid as Lycenchelys atacamensis Andriashev 1980 -- (Pequeño 1989:48 [ref. 14125], Balushkin et al. 2011:978 [ref. 32449]). Current status: Valid as Lycenchelys atacamensis Andriashev 1980. Zoarcidae: Lycodinae. Distribution: Peru-Chile Trench. Habitat: marine.

atlanticum, Melanostigma Koefoed [E.] 1952:10, Pl. 2 (fig. D) [Scientific Results of the Michael Sars North Atlantic Deep-Sea Expedition 1910 v. 4 (pt 2, no. 2); ref. 2652] North Atlantic, 57°41'N, 11°48'W, station 101, 2500 meters wire out. Holotype: ZMUB 4482. Type catalog: Frøiland 1979:150 [ref. 21547]. •Valid as Melanostigma atlanticum Koefoed 1952 -- (Andriashev 1973:547 [ref. 9214], Andriashev in Whitehead et al. 1986:1148 [ref. 13677], Robins & Ray 1986:236 [ref. 23100], Scott & Scott 1988:414 [ref. 25518], McAllister 1990:165 [ref. 14674], Anderson & Almeida 1990:888 [ref. 19328], Anderson 1994:119 [ref. 21438], Koyanagi in Okamura et al. 1995:206 [ref. 22531], Klein-MacPhee & Collette in Collette & Klein-MacPhee 2002:468 [ref. 26158], Moore et al. 2003:229 [ref. 27076], Anderson & Fedorov 2004:8 [ref. 27603], Lipej & Dulčić 2010:31 [ref. 36649], Møller et al. 2010:64 [ref. 31169], Balushkin et al. 2011:969 [ref. 32449], Page et al. 2013:163 [ref. 32708], Afonso et al. 2016:68, 222 [ref. 36199], Anderson in Carpenter & De Angelis 2016:2760 [ref. 34618], Balushkin & Moganova 2017:131 [ref. 35279], Balushkin & Orlovskaya 2019:135 [ref. 36664], Orlovskaya & Balushkin 2019:505 [ref. 37050]). Current status: Valid as Melanostigma atlanticum Koefoed 1952. Zoarcidae: Gymnelinae. Distribution: Western Mediterranean Sea, eastern North Atlantic: Scotland south to Western Sahara; northwestern Atlantic: Newfoundland (Canada) to southeastern U.S.A. Habitat: marine.

atlanticus, Lycodes Jensen [A. S.] 1902:207 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Coast of North America, off Virginia, U.S.A., 39°34'45"N, 71°21'30"W, depth 516-1423 fathoms. Holotype (unique): ZMUC 536. Type catalog: Nielsen 1974:76 [ref. 9588]. •Valid as Lycodes atlanticus Jensen 1902 -- (Scott & Scott 1988:406 [ref. 25518], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Moore et al. 2003:229 [ref. 27076]). •Synonym of Lycodes terraenovae Collett 1896 -- (Møller 1997:47 [ref. 22879], Møller 2000:847 [ref. 24585], Anderson & Fedorov 2004:30 [ref. 27603], Balushkin et al. 2011:1015 [ref. 32449]). Current status: Synonym of Lycodes terraenovae Collett 1896. Zoarcidae: Lycodinae. Habitat: marine.

atlanticus, Lycodes turneri Vladykov [V. D.] & Tremblay [J.-L.] 1936:38, Pl. 7 (figs. 14-16) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Saint Lawrence River estuary, Quebec, Canada. Syntypes: (18) NMC 60-160 (1). Type catalog: McAllister 1965:9 [ref. 22019] as paratypes. Objectively invalid; preoccupied by Lycodes atlanticus Jensen 1902. •Synonym of Lycodes polaris (Sabine 1824) -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603], Parin et al. 2014:418 [ref. 33547]). Current status: Synonym of Lycodes polaris (Sabine 1824). Zoarcidae: Lycodinae. Habitat: marine.

atratus, Lycodes Vladykov [V. D.] & Tremblay [J.-L.] 1936:26, Pl. 2 (fig. 6) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Saint Lawrence Estuary, Canada. Holotype (unique): whereabouts unknown. •Synonym of Lycodes atlanticus Jensen 1902 -- (Anderson 1994:118 [ref. 21438]). •Synonym of Lycodes terraenovae Collett 1896 -- (Møller 1997:48 [ref. 22879], Møller 2000:847 [ref. 24585], Anderson & Fedorov 2004:30 [ref. 27603]). Current status: Synonym of Lycodes terraenovae Collett 1896. Zoarcidae: Lycodinae. Habitat: marine.

attenuatus, Lycodapus Gilbert [C. H.] 1915:372, Pl. 21 (fig. 22) [Proceedings of the United States National Museum v. 48 (no. 2075); ref. 1632] Monterey Bay, California, U.S.A., Albatross station 4461, depth 285-357 fathoms. Holotype (unique): USNM 75821. Type catalog: Springer & Anderson 1997:14 [ref. 22953]. •Synonym of Lycodapus mandibularis Gilbert 1915 -- (Peden & Anderson 1978:1938 [ref. 8895], Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:22 [ref. 27603]). Current status: Synonym of Lycodapus mandibularis Gilbert 1915. Zoarcidae: Lycodinae. Habitat: marine.

attenuatus, Lycodes Knipowitsch [N. M.] 1906:73, Figs. 13-14 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Laptev Sea, Russia, 75°-77°N, 114°-139°E [3 stations], depth 19-38 meters. Syntypes: ZIN 14249-51 (1, 1, 1). Type catalog: Balushkin et al. 2011:988 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667]. Also described as new in Knipowitsch 1907:36 [ref. 15152]. •Synonym of Lycodes pallidus Collett 1879 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:27 [ref. 27603]). •Valid as Lycodes attenuatus Knipowitsch 1906 -- (Balushkin et al. 2011:988 [ref. 32449], Parin et al. 2014:407 [ref. 33547], but treated as a synonym of Lycodes pallidus (Collett 1879) on p. 417, Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes attenuatus Knipowitsch 1906. Zoarcidae: Lycodinae. Distribution: Arctic and North Pacific, Habitat: marine.

aurantiaca, Lycenchelys Shinohara [G.] & Matsuura [K.] 1998:151, Figs. 1-3 [Ichthyological Research v. 45 (no. 2); ref. 23382] Off Miyagi Prefecture, Honshu, Japan, 37°58.54'N, 142°09.00'E to 37°56.84'N, 142°09.07'E, depth 700 meters. Holotype: NSMT-P 53147. Paratypes: AMS I.38499-001 (1); CAS 98192 (1); NSMT-P 47415-16 (2), 49908 (1), 49911 (1), 53143 (1, c&s), 53144 (1), 53146 (1). •Valid as Lycenchelys aurantiaca Shinohara & Matsuura 1998 -- (Nakabo 2000:1033 [ref. 25182], Nakabo 2002:1033 [ref. 26193], Anderson & Fedorov 2004:15 [ref. 27603], Shinohara & Anderson 2007:64 [ref. 29111], Balushkin et al. 2011:1026 [ref. 32449], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys aurantiaca Shinohara & Matsuura 1998. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Japan. Habitat: marine.

australis, Lycodapus Norman [J. R.] 1937:110, Fig. 59 [Discovery Reports v. 16; ref. 3227] Strait of Magellan, Discovery station WS748, 53°41'30"S, 70°55'00"W, Chile. Holotype: BMNH 1936.8.26.1048. Paratypes: BMNH 1936.8.26.1049-1051 (3). •Valid as Lycodapus australis Norman 1937 -- (Peden & Anderson 1978:1956 [ref. 8895], Pequeño 1989:49 [ref. 14125], Miller 1993:678 [ref. 21297], Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:21 [ref. 27603], McCosker & Rosenblatt 2010:193 [ref. 30957], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycodapus australis Norman 1937. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific. Habitat: marine.

bachmanni, Lycenchelys Gosztonyi [A. E.] 1977:218, Figs. 10-11 [Archiv für Fischereiwissenschaft v. 27 (no. 3); ref. 6103] Continental slope off Río Negro Province, Argentina, 41°05'S, 57°15'W, depth 800 meters. Holotype (unique): ISH 306-1971. Type catalog: Krefft 1978:15 [ref. 9884]. •Valid as Lycenchelys bachmanni Gosztonyi 1977 -- (Nakamura in Nakamura et al. 1986:242 [ref. 14235], Anderson & Gosztonyi 1991:6 [ref. 19348], Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:16 [ref. 27603], Balushkin et al. 2011:979 [ref. 32449], Nión et al. 2016:51 [ref. 35565]). Current status: Valid as Lycenchelys bachmanni Gosztonyi 1977. Zoarcidae: Lycodinae. Distribution: Western South Atlantic: Argentina and Uruguay. Habitat: marine.

barbatum, Lyconema Gilbert [C. H.] 1896:471, Pl. 35 (upper) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Off central California, 36°39'40"N, 122°01'W, Albatross station 3129, depth 204 fathoms. Lectotype: USNM 48582. Paralectotypes: BMNH 1896.7.23.197 (1); MCZ 28317 (1); CAS-SU 3627 (1), 69673 [ex 3627] (4); USNM 53036 (1); ZMB 14336 [or 14386, ex USNM] (1); ZMUO J1684 (1). Type catalog: Böhlke 1953:101 [ref. 12291], Pethon 1969:12 [ref. 19268], Springer & Anderson 1997:14 [ref. 22953]. Lectotype established (as figured specimen) in caption to Pl. 352, p. 3305 in Jordan & Evermann 1900 [ref. 2446] and traced to USNM 45362 by Springer & Anderson 1997:14 [ref. 22953]; lectotype selected by Anderson 1994:76 [ref. 21438] is invalid. •Valid as Lyconema barbatum Gilbert 1896 -- (Eschmeyer & Herald 1983:106 [ref. 9277], Toyoshima 1985:142 [ref. 5722], Anderson 1994:76, 119 [ref. 21438], Anderson & Fedorov 2004:32 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Balushkin et al. 2011:1026 [ref. 32449], Page et al. 2013:163 [ref. 32708], Kells et al. 2016:351 [ref. 35888]). Current status: Valid as Lyconema barbatum Gilbert 1896. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

barsukovi, Gymnelus Chernova [N. V.] 1999:310, Figs. 5-7 [Voprosy Ikhtiologii v. 39 (no. 3); ref. 24868] Bering Sea, Providence Bay, Chukchi Peninsula. Holotype (unique): ZIN 35423. Paratypes: NMC 86-0592B (2); ZIN 25332 (2), 31392 (1), 34984 (1), 43361 (1), 52561-62 (1, 1). Plus non-type material. Type catalog: Balushkin et al 2011:963 [ref. 32499], Balushkin et al. 2012:157 [ref. 33667]. Anderson in Anderson & Fedorov 2004:5 [ref. 27603] believes this is probably a synonym of G. viridis. •Synonym of Gymnelus viridis (Fabricius 1780) -- (Mecklenburg et al. 2011:124, 128 [ref. 31212], Parin et al. 2014:395 [ref. 33547], Mecklenburg & Anderson 2015:278 [ref. 33742]). •Valid as Gymnelus barsukovi Chernova 1999 -- (Chernova 2000:8 [ref. 24885], Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:5 [ref. 27603], Balushkin et al.2011:963 [ref. 32499], Afonso et al. 2016:61, 158 [ref. 36199]). Current status: Valid as Gymnelus barsukovi Chernova 1999. Zoarcidae: Gymnelinae. Distribution: Arctic Ocean, including Bering Sea. Habitat: marine.

bathium, Melanostigma Bussing [W. A.] 1965:219, Fig. 15 [Antarctic Research Series, Washington DC. v. 5; ref. 9201] Off southern Chile, 40°15'S, 78°17'W. Holotype: LACM 10063. Paratypes: LACM 10064 (orig. 2, now 1), 10065-66 (1, 1); ZMUC P761048 [ex LACM 10064] (1). •Valid as Melanostigma bathium Bussing 1965 -- (Anderson 1988:62 [ref. 7304], Pequeño 1989:49 [ref. 14125], Anderson 1990:269 [ref. 21695], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:8 [ref. 27603], McCosker & Rosenblatt 2010:193 [ref. 30957], Balushkin et al. 2011:1026 [ref. 32449], Balushkin & Moganova 2017:131 [ref. 35279], Balushkin & Moganova 2018:136 [ref. 35877], Balushkin 2019:287 [ref. 36807]). Current status: Valid as Melanostigma bathium Bussing 1965. Zoarcidae: Gymnelinae. Distribution: Eastern Pacific to Antarctic Peninsula. Habitat: marine.

bathybius, Lycodes Schmidt [P. Yu.] 1950:105 [91], Pl. 9 (fig. 2) [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] Near southern extremity of Kamchatka, Russia, Okhotsk Sea, 51°09'N, 154°17'E, depth 591 meters. Holotype (unique): ZIN 24829. Type catalog: Balushkin et al. 2011:988 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667]. On p. 91 of English translation. Appeared as name only in Schmidt 1935. •Status uncertain -- (Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:48 [ref. 27603]). •Valid as Lycodes bathybius Schmidt 1950 -- (Toyoshima 1985:203 [ref. 5722], Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:112 [ref. 27579], Balushkin et al. 2011:988 [ref. 32449], Parin et al. 2014:407 [ref. 33547]). Current status: Valid as Lycodes bathybius Schmidt 1950. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

bellingshauseni, Lycenchelys Andriashev [A. P.] & Permitin [Yu. Ye.] 1968:612, Figs. 1-2 [Voprosy Ikhtiologii v. 8 (no. 4, art. 51); ref. 7858] North of South Georgia Island, 53°39'S, 36°20'W, depth 620-640 meters. Holotype (unique): ZIN 37878. Type catalog: Balushkin et al. 2011:979 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667]. •Valid as Lycenchelys bellingshauseni Andriashev & Permitin 1968 -- (Anderson 1988:93 [ref. 7304], Anderson 1990:260 [ref. 21695], Miller 1993:664 [ref. 21297], Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:16 [ref. 27603], Shinohara & Anderson 2007:61 [ref. 29111] once as bellingshausenni, Balushkin et al. 2011:979 [ref. 32449]). Current status: Valid as Lycenchelys bellingshauseni Andriashev & Permitin 1968. Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

beringi, Lycodes (Furcimanus) diapterus Andriashev [A. P.] 1935:115, Fig. 1 [C. R. (Doklady) Acad. Sci. URSS v. 4 [9] (nos 1-2); ref. 13870] Off Commander Island, 6 miles west of Cape Yuchina, depth 200-250 meters. Lectotype: ZIN 29850. Paralectotypes: ZIN 34838 (4), 54299 (now 7). Type catalog: Balushkin et al. 2011:988 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. Subgenus Furcimanus misspelled in original. USNM paralectotypes of Lycodes diapterus are this species. Lectotype selected by Stevenson & Sheiko 2009:132 [ref. 30070]. •Synonym of Lycodes diapterus Gilbert 1892 -- (Toyoshima 1985:230 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:724 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603]). •Valid as Lycodes beringi Andriashev 1935 -- (Stevenson & Sheiko 2009:130 [ref. 30070], Balushkin et al. 2011:988 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:407 [ref. 33547], Pietsch & Orr 2015:58 [ref. 34694]). Current status: Valid as Lycodes beringi Andriashev 1935. Zoarcidae: Lycodinae. Distribution: North Pacific: Kamchatka Peninsula to Puget Sound, Canada. Habitat: marine.

bilabrus, Gymnelis Andriashev [A. P.] 1937:344, Pl. 2 (figs. 20-22) [Issled. Morey SSSR, Leningrad v. 25; ref. 20242] Between Cape Chukotsky [Chukchi] and St. Lawrence Island, Bering Sea, 64°09.5'N, 171°58'W, depth 40 meters. Holotype: ZIN 24943. Paratypes: (2). Type catalog: Balushkin et al 2011:963 [ref. 32499], Balushkin et al. 2012:157 [ref. 33667]. Original genus should have been Gymnelus. •Valid as Gymnelus bilabrus Andriashev 1937 -- (Chernova 1998:182 [ref. 24672], Chernova 1999:306 [ref. 24868], Chernova 2000:8 [ref. 24885], Sheiko & Fedorov 2000:35 [ref. 25650], Balushkin et al.2011:961 [ref. 32499], Afonso et al. 2016:61, 159 [ref. 36199]). •Synonym of Gymnelus viridis (Fabricius 1780) -- (Anderson 1982:30 [ref. 5520], Mecklenburg et al. 2002:690 [ref. 25968], Anderson & Fedorov 2004:7 [ref. 27603], Mecklenburg et al. 2011:124, 128 [ref. 31212], Parin et al. 2014:394 [ref. 33547], Mecklenburg et al. 2016:157 [ref. 34440]). Current status: Synonym of Gymnelus viridis (Fabricius 1780). Zoarcidae: Gymnelinae. Habitat: marine.

birsteini, Lycenchelys Andriashev [A. P.] 1958:178, Figs. 1 (no. 3), 4 [Voprosy Ikhtiologii v. 11; ref. 12042] Northern part of Kuril-Kamchatka Trench, 49°49.5'S, 157°45'E, depth 3960-4070 meters. Holotype: ZIN 34669 (191 mm). Paratypes: ZIN 34669a (1). Type catalog: Balushkin et al. 2011:979 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667]. •Synonym of Lycenchelys plicifera Andriashev 1955 -- (Anderson 1994:117 [ref. 21438], Anderson 1995:94 [ref. 21928], Anderson & Balanov 2000:1056 [ref. 25023], Sheiko & Fedorov 2000:35 [ref. 25650], Mecklenburg et al. 2002:696 [ref. 25968], Anderson & Fedorov 2004:19 [ref. 27603]). •Valid as Lycenchelys birsteini Andriashev 1958 -- (Toyoshima 1985:155 [ref. 5722] as bersteini, Balushkin et al. 2011:979 [ref. 32449], Parin et al. 2014:398 [ref. 33547]). Current status: Valid as Lycenchelys birsteini Andriashev 1958. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Kuril-Kamchatka Trench. Habitat: marine.

blennoides, Zoarcus Griffith [E.] & Smith [C. H.] 1834:opp. 230, Pl. 46 (fig. 3) [The class Pisces, arranged by the Baron Cuvier; ref. 1908] No locality stated. No types known. Genus should have been Zoarces. •Synonym of Zoarces viviparus (Linnaeus 1758). Current status: Synonym of Zoarces viviparus (Linnaeus 1758). Zoarcidae: Zoarcinae. Habitat: marine.

borealis, Lycodes vahlii forma Smitt [F. A.] 1900:57 [Annals and Magazine of Natural History (Series 7) v. 5 (no. 25); ref. 18731] No locality. Not available if regarded as infrasubspecific. Described as "Forma borealis (gracilis, auctt.)." Also appeared in Smitt 1901:6 [ref. 18732]. •Unavailable in family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Zoarcidae.

bothriocephalus, Lycodes Pappenheim [P.] 1912:178, Pl. 10 (fig. 2) [Deutsche Südpolar-Expedition 1901-1903 v. 13 (Zool. 5); ref. 14856] Gauss winter station, Wilhelm II Coast, Antarctica. Holotype: ZMB 18927. Original not seen. •Valid as Ophthalmolycus bothriocephalus (Pappenheim 1912) -- (Anderson 1988:84 [ref. 7304], Anderson 1990:8 [ref. 16143], Anderson 1990:273 [ref. 21695], Anderson 1992:8 [ref. 18912], Miller 1993:657 [ref. 21297], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:33 [ref. 27603], Matallanas 2009:55 [ref. 30103], Struthers & Møller 2009:65 [ref. 30403], Matallanas 2011:566 [ref. 30839], Balushkin et al. 2011:1020 [ref. 32449]). Current status: Valid as Ophthalmolycus bothriocephalus (Pappenheim 1912). Zoarcidae: Lycodinae. Distribution: Southern Ocean: circum-Antarctica. Habitat: marine.

brachycephalus, Lycodes Pappenheim [P.] 1912:179, Pl. 10 (fig. 3) [Deutsche Südpolar-Expedition 1901-1903 v. 13 (Zool. 5); ref. 14856] Gauss winter station, Wilhelm II Coast, Antarctica. Lectotype (?): ZMB 18929. Paralectotypes: ASUM A5.23.2.1a (1); BMNH 1913.4.15 [or 23].58-59 (2); SMNS 4605 (2); ZIN 37929 [ex ZMB 18935] (2); ZMB 24093 [ex ZMB 18929] (4); ZMH 8156 (2). Plus more types at ZMB. Type catalog: Wilkens & Dohse 1993:419 [ref. 21161], Fricke 1995:22 [ref. 28100], Fricke 2005:69-70 [ref. 29864] dated 1913 with syntypes, Balushkin et al. 2011:1020 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667]. Lectotype from the literature, not designated here. •Valid as Pachycara brachycephalum (Pappenheim 1912) -- (Anderson 1988:74 [ref. 7304], Anderson 1989:240 [ref. 13487], Anderson 1990:274 [ref. 21695], Anderson 1991:158 [ref. 20019], Miller 1993:648 [ref. 21297], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:227 [ref. 22911], Pequeño 1997:83 [ref. 23536], Olaso et al. 1997:83 [ref. 25009], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:34 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson 2006:11 [ref. 28576], Møller & King 2007:588 [ref. 29259], Struthers & Møller 2009:65 [ref. 30403], Balushkin et al. 2011:1020 [ref. 32449]). Current status: Valid as Pachycara brachycephalum (Pappenheim 1912). Zoarcidae: Lycodinae. Distribution: Southern Ocean: circum-Antarctic. Habitat: marine.

brachyrhynchus, Lycenchelys Schmidt [P. Yu.] 1904:201, Pl. 6 (fig. 3) [Fishes of the eastern seas of the Russian Empire; ref. 3946] Aniva Bay (sea of Okhotsk) and Terpeniya [Patience] Bay, St. Catherine Cape, Nevelski Straight, Northern Sea of Japan, Russia, depth 91-11 feet (13-17 Russian fathoms). Syntypes: ?ZIN 13089 (1), 13090 (1), ?13091 (3). Type catalog: Balushkin et al. 2011:958 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. Appeared first without distinguishing features in Schmidt 1903:519 (p. 17 of separate) [ref. 3945]. The specific name is a noun. •Valid as Davidojordania [sic] brachyrhyncha (Schmidt 1904) -- (Lindberg & Krasyukova 1975:175 [ref. 7348]). •Valid as Davidijordania brachyrhynchus (Schmidt 1904) (often seen as brachyrhyncha) -- (Toyoshima in Masuda et al. 1984:305 [ref. 6441], Anderson 1994:116 [ref. 21438], Nakabo 2000:1036 [ref. 25182], Sheiko & Fedorov 2000:34 [ref. 25650], Nakabo 2002:1036 [ref. 26193], Fedorov et al. 2003:110 [ref. 27579], Anderson & Fedorov 2004:3 [ref. 27603], Anderson & Imamura 2008:65 [ref. 29573], Shinohara & Kim 2009:232 [ref. 31427], Balushkin et al. 2011:958 [ref. 32449], Parin et al. 2014:387 [ref. 33547], Dyldin & Orlov 2017:73 [ref. 35138] as brachyrhyncha). Current status: Valid as Davidijordania brachyrhynchus (Schmidt 1904). Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk Sea and Sea of Japan. Habitat: brackish, marine.

brashnikovi, Gymnelopsis Soldatov [V. K.] 1922:162 [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23 (no. 2) (for 1917); ref. 4159] Sakalov Island, Cape Eustaphie, Okhotsk Sea, depth 86 meters. Holotype (unique): ZIN 13029. Type catalog: Balushkin et al. 2011:960 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. Published in 1922, not 1917. •Valid as Gymnelopsis brashnikovi Soldatov 1922 -- (Lindberg & Krasyukova 1975:178 [ref. 7348], Toyoshima in Masuda et al. 1984:306 [ref. 6441], Anderson 1994:116 [ref. 21438], Youn 2002:375, 639 [ref. 26218], Nazarkin & Chernova 2003:605 [ref. 27204], Anderson & Fedorov 2004:4 [ref. 27603], Kim et al. 2006:278 [ref. 29191], Balushkin et al. 2011:960 [ref. 32449], Parin et al. 2014:389 [ref. 33547]). Current status: Valid as Gymnelopsis brashnikovi Soldatov 1922. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk Sea and Korea. Habitat: marine.

brashnikovi, Lycodes Soldatov [V. K.] 1918:112, Fig. 1 [An. Mus. Zool. Acad. Sci. Petrograd v. 22; ref. 21830] 53°17'N, 154°47'E and Senyavine Cape, eastern coast of Sakhalin Island, Okhotsk Sea. Syntypes: ZIN 19167 (3), ZIN (1, not found in 1996). Non-types: ZIN 13010 (2) questionably included. Type catalog: Balushkin et al. 2011:989 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. Also in Soldatov 1922:113, Fig. 1 [ref. 15429]. •Synonym of Lycodes palearis Gilbert 1896, subspecies schmidtii Gratzianov 1907 -- (Lindberg & Krasyukova 1975:158 [ref. 7348]). •Synonym of Lycodes fasciatus (Schmidt 1904) -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Synonym of Lycodes schmidti Gratzianov 1907 -- (Toyoshima 1985:214 [ref. 5722], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:29 [ref. 27603]). •Valid as Lycodes brashnikovi Soldatov 1918 -- (Nazarkin 2010:603 [ref. 31019], Balushkin et al. 2011:989 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:408 [ref. 33547]). Current status: Valid as Lycodes brashnikovi Soldatov 1918. Zoarcidae: Lycodinae. Distribution: Okhotsk Sea. Habitat: marine.

brevicauda, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] 1935:248, Figs. 4-5 [Zoologischer Anzeiger v. 112 (nos 9-10); ref. 15744] Okhotsk Sea. Syntypes: (25) ZIN 24825 (1), 29993 (1), ?30015 (2), 30190 (6), 30196 (3), 30986 (3), 31292 (2), 31393 (1). Type catalog: Balushkin et al. 2011:990 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667].. •Synonym of Lycodes tanakai Jordan & Thompson 1914 -- (Anderson 1994:119 [ref. 21438] as tanakae). •Synonym of Lycodes tanakai Jordan & Thompson 1914 -- (Toyoshima 1985:193 [ref. 5722], Anderson & Fedorov 2004:30 [ref. 27603]). •Valid as Lycodes brevicaudus Taranetz & Andriashev 1935 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:113 [ref. 27579], Balushkin et al. 2011:990 [ref. 32449], Parin et al. 2014:408 [ref. 33547], but as a synonym of Lycodes tanakae on p. 423). Current status: Valid as Lycodes bevicaudus Taranetz & Andriashev 1935. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

brevifenestratus, Gymnelopsis Anderson [M. E.] 1982:52, Fig. 33 [National Museum of Canada Publications in Zoology No. 17; ref. 5520] Okhotsk Sea, 57°50'N, 141°47'E, depth 146 meters. Holotype: USNM 92587. Paratypes: ZIN 23944 (1), 23952-53 (1, 3), 33334 (6), 33751 (1), 34842 (1). Type catalog: Springer & Anderson 1997:14 [ref. 22953], Balushkin 2011:960 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. •Valid as Gymnelopsis brevifenestratus Anderson 1982 -- (Anderson 1982:52 [ref. 5520]). •Valid as Gymnelopsis brevifenestrata Anderson 1982 -- (Anderson 1994:116 [ref. 21438], Sheiko & Fedorov 2000:35 [ref. 25650], Nazarkin & Chernova 2003:605 [ref. 27204], Fedorov et al. 2003:110 [ref. 27579], Anderson & Fedorov 2004:4 [ref. 27603], Balushkin et al. 2011:960 [ref. 32449], Parin et al. 2014:389 [ref. 33547]). Current status: Valid as Gymnelopsis brevifenestrata Anderson 1982. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk Sea and Japan. Habitat: marine.

brevimaxillaris, Lycenchelys Toyoshima [M.] 1985:174, Figs. 29-30, 31k [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Pacific coast of Aomori Prefecture, 41°13'N, 141°44'E, northern Japan, depth 690-750 meters. Holotype: HUMZ 189817 [ex ACAP 4681]. Paratypes: NSMT-P 68702 [ex ACAP 4682] (1). See Imamura et al. 2005:1 [ref. 28190] for rediscovery of type specimens. •Valid as Lycenchelys brevimaxillaris Toyoshima 1985 -- (Anderson 1994:117 [ref. 21438], Fedorov 1995:130 [ref. 21958], Nakabo 2000:1033 [ref. 25182], Nakabo 2002:1033 [ref. 26193], Anderson & Fedorov 2004:16 [ref. 27603]). •Synonym of Lycenchelys melanostomias Toyoshima 1983 -- (Imamura et al. 2004:84 [ref. 27600], Imamura et al. 2005:2 [ref. 28190]). Current status: Synonym of Lycenchelys melanostomias Toyoshima 1983. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Japan. Habitat: marine.

brevipes, Lycodes Bean [T. H.] 1890:38 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Between Unga Island and Nagai Island, Aleutian Islands, Alaska, U.S.A., 55°10'N, 160°18'W, Albatross station 2848, depth 110 fathoms [201 meters]. Lectotype: USNM 45362. Paralectotypes: USNM 162712 (16). Type catalog: Springer & Anderson 1997:14-15 [ref. 22953], Balushkin et al. 2011:991 [ref. 32449]. Lectotype selected by Springer & Anderson 1997:14-15 [ref. 22953] (lectotype also figured by them). •Valid as Lycodes brevipes Bean 1890 -- (Eschmeyer & Herald 1983:105 [ref. 9277], Toyoshima 1985:233 [ref. 5722], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:716 [ref. 25968], Anderson & Fedorov 2004:23 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Balushkin et al. 2011:991 [ref. 32449], Page et al. 2013:162 [ref. 32708], Shinohara et al. 2014:265 [ref. 33330], Parin et al. 2014:408 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Pietsch & Orr 2015:58 [ref. 34694], Kells et al. 2016:272 [ref. 35888]). Current status: Valid as Lycodes brevipes Bean 1890. Zoarcidae: Lycodinae. Distribution: North Pacific. Habitat: marine.

brevis, Maynea Norman [J. R.] 1937:108, Fig. 57 [Discovery Reports v. 16; ref. 3227] Discovery station WS825, 50°50'S, 57°15'15"W, Patagonia-Falklands region, depth 135-144 meters. Holotype: BMNH 1936.8.26.1042. Paratypes: BMNH 1936.8.26.1043-1045 (3). •Valid as Oidiphorus brevis (Norman 1937) -- (Gosztonyi 1977:228 [ref. 6103], Anderson & Gosztonyi 1991:9 [ref. 19348], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:33 [ref. 27603], Balushkin et al. 2011:1019 [ref. 32449], Nión et al. 2016:51 [ref. 35565]). Current status: Valid as Oidiphorus brevis (Norman 1937). Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

brunnea, Maynea Bean [T. H.] 1890:39 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Off San Clemente Island, southern California, U.S.A., 33°08'N, 118°40'W, Albatross station 2839, depth 414 fathoms. Holotype (unique): USNM 53029 (nearly disintegrated). Type catalog: Springer & Anderson 1997:15 [ref. 22953]. •Valid as Bothrocara brunneum (Bean 1890) -- (Eschmeyer & Herald 1983:104 [ref. 9277], McAllister 1990:156 [ref. 14674], Anderson 1994:116 [ref. 21438], Sheiko & Fedorov 2000:34 [ref. 25650], Mecklenburg et al. 2002:731 [ref. 25968], Fedorov et al. 2003:109 [ref. 27579], Anderson & Fedorov 2004:10 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Stevenson & Anderson 2005:53 [ref. 28569], Anderson 2006:3 [ref. 28576], Anderson et al. 2009:173 [ref. 30175], Balushkin et al. 2011:973 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:384 [ref. 33547], Kells et al. 2016:270 [ref. 35888]). Current status: Valid as Bothrocara brunneum (Bean 1890). Zoarcidae: Lycodinae. Distribution: Northern Pacific and eastern Pacific: Sea of Okhotsk to Gulf of California (Mexico) and Gulf of Panama. Habitat: marine.

brunneofasciatus, Lycodes Suvorov [E. K.] 1935:437 [English p. 440], Fig. 3 [Bulletin de l'Academie des Sciences de l'U.R.S.S., Classe des Sciences Mathématiques et Naturelles 1935; ref. 4300] Western Kamchatka, Russia, 53°35'N, 154°22'E, depth 250 meters. Holotype (unique): ZIN 25074. Type catalog: Balushkin et al. 2011:991 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. •Valid as Lycodes brunneofasciatus Suvorov 1935 -- (Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:234 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1037 [ref. 25182], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:715 [ref. 25968], Nakabo 2002:1037 [ref. 26193], Tokranov & Orlov 2002:605 [ref. 26464], Fedorov et al. 2003:113 [ref. 27579], Anderson & Fedorov 2004:24 [ref. 27603], Balushkin et al. 2011:991 [ref. 32449], Shinohara et al. 2014:265 [ref. 33330], Parin et al. 2014:409 [ref. 33547]). Current status: Valid as Lycodes brunneofasciatus Suvorov 1935. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

brunneus, Lycodes Fowler [H. W.] 1944:73, Fig. [The Fish Culturist v. 23 (no. 10); ref. 17879] North of Bahamas, east of Florida, Albatross station 2660, 28°40'N, 78°46'W, depth 504 fathoms. Holotype: USNM 119446. Paratypes: USNM 119447 (1). Type catalog: Springer & Anderson 1997:15 [ref. 22953]. •Synonym of Lycodes atlanticus Jensen 1902 -- (Anderson 1994:118 [ref. 21438]). •Synonym of Lycodes terraenovae Collett 1896 -- (Møller 1997:48 [ref. 22879], Møller 2000:847 [ref. 24585], Anderson & Fedorov 2004:30 [ref. 27603]) Current status: Synonym of Lycodes terraenovae Collett 1896. Zoarcidae: Lycodinae. Habitat: marine.

brychios, Leucogrammolycus Mincarone [M. M.] & Anderson [M. E.] 2008:66, Figs. 1-2 [Zootaxa No. 1852; ref. 29768] Off southeastern Brazil, 23°53.137'S, 42°31.271'W, depth 615 meters. Holotype: MOVI 39079. Paratypes: MOVI 38816 (1), 38817 (1), 39079 (1), 39081 (1), 39082 (1), 39083 (1), 39085 (1); RUSI 80187 (1) •Valid as Leucogrammolycus brychios Mincarone & Anderson 2008 -- (Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Leucogrammolycus brychios Mincarone & Anderson 2008. Zoarcidae: Lycodinae. Distribution: Southwestern Atlantic: Brazil. Habitat: marine.

bulbiceps, Maynea Garman [S.] 1899:140, Pl. E (fig. 1) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] East of Cocos Island, off Panama Bay, 6°10'N, 83°06'W, Albatross station 3361, depth 1471 fathoms. Holotype (unique): MCZ 28681 (poor condition). •Valid as Pachycara bulbiceps (Garman 1899) -- (Anderson & Peden 1988:84 [ref. 7215], Anderson 1989:227 [ref. 13487], McAllister 1990:165 [ref. 14674], Anderson 1991:158 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:226 [ref. 22911], Mecklenburg et al. 2002:727 [ref. 25968], Møller 2003:359 [ref. 26908], Moore et al. 2003:229 [ref. 27076], Anderson & Fedorov 2004:34 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Møller & King 2007:588 [ref. 29259], Shinohara 2012:55 [ref. 32102], Anderson 2012:39 [32270], Balushkin et al. 2011:1027 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448], Anderson in Carpenter & De Angelis 2016:2760 [ref. 34618]). Current status: Valid as Pachycara bulbiceps (Garman 1899). Zoarcidae: Lycodinae. Distribution: Eastern Pacific and North Atlantic. Habitat: marine.

bullisi, Lycenchelys Cohen [D. M.] 1964:113, Figs. 1-3 [Proceedings of the Biological Society of Washington v. 77; ref. 1675] Off mouth of Mississippi River, Gulf of Mexico, 28°51'N, 88°41'W, Oregon station 4038, depth 400-410 fathoms. Holotype: USNM 188232. Paratypes: UF 205363 [ex UMML 5363] (1). Type catalog: Springer & Anderson 1997:15 [ref. 22953]. •Valid as Lycenchelys bullisi Cohen 1964 -- (Anderson 1994:117 [ref. 21438], Møller 1999:327 [ref. 24052], Anderson 2003:1741 [ref. 27106], Anderson & Fedorov 2004:16 [ref. 27603], McEachran & Fechhelm 2005:548 [ref. 28815], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys bullisi Cohen 1964. Zoarcidae: Lycodinae. Distribution: Western Atlantic. Habitat: marine.

californica, Maynea Starks [E. C.] & Mann [W. M.] (ex Gilbert) 1911:16 [University of California Publications in Zoology v. 8 (no. 2); ref. 4199] Off San Nicolas Island, southern California, U.S.A., Albatross station 4421, depth 229-298 fathoms. Holotype: USNM 75819. Paratypes: CAS-SU 22399 (1). Type catalog: Springer & Anderson 1997:15-16 [ref. 22953]. Also appeared as new in Gilbert 1915:362 [ref. 1632]. Three specimens from off San Diego used by Starks & Mann are considered paratypes (whereabouts of two are unknown). •Valid as Maynea californica Starks & Mann 1911 -- (Eschmeyer & Herald 1983:106 [ref. 9277], Toyoshima 1985:143 [ref. 5722]). •Valid as Eucryphycus californicus (Starks & Mann 1911) -- (Anderson 1988:94 [ref. 6021], Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:13 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Balushkin et al. 2011:976 [ref. 32449], Page et al. 2013:162 [ref. 32708], Kells et al. 2016:350 [ref. 35888]). Current status: Valid as Eucryphycus californicus (Starks & Mann 1911). Zoarcidae: Lycodinae. Distribution: Eastern North Pacific: California, U.S.A. Habitat: marine.

callista, Lycenchelys Anderson [M. E.] 1995:61, Fig. 1 [Proceedings of the California Academy of Sciences v. 49 (no. 2); ref. 21928] Off Pt. Sur, California, U.S.A., 36°15.1'N, 122°15.5'W, depth 1580-1622 meters. Holotype: CAS 55412. Paratypes: CAS 39849 (5), 55062 (2), 80630 (29); OS 14376 (8); SIO 84-249 (21), 85-51 (22), 71-141 (3). •Valid as Lycenchelys callista Anderson 1995 -- (Anderson & Fedorov 2004:16 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys callista Anderson 1995. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

camchaticus, Lycodes Gilbert [C. H.] & Burke [C. V.] 1912:89, Fig. 34 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off Avatcha Bay, east coast of Kamchatka, Russia, 52°37'30"N, 158°50'E, Albatross station 4797, depth 682 fathoms. Holotype: USNM 74396. Paratypes: CAS-SU 22366 (1), USNM 70928 (2). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:16 [ref. 22953]. •Valid as Lycenchelys camchaticus (Gilbert & Burke 1912) -- (Anderson et al. 1979:262 [ref. 16035], Toyoshima 1985:164 [ref. 5722], Balushkin et al. 2011:979 [ref. 32449], Parin et al. 2014:398 [ref. 33547]). •Valid as Lycenchelys camchatica (Gilbert & Burke 1912) -- (Anderson 1994:117 [ref. 21438], Anderson 1995:63 [ref. 21928], Sheiko & Fedorov 2000:35 [ref. 25650], Mecklenburg et al. 2002:700 [ref. 25968], Anderson & Fedorov 2004:16 [ref. 27603], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys camchatica (Gilbert & Burke 1912). Zoarcidae: Lycodinae. Distribution: North Pacific. Habitat: marine.

campbellensis, Ophthalmolycus (Lacrimolycus) Andriashev [A. P.] & Fedorov [V. V.] 1986:28, Figs. 3-4 [Voprosy Ikhtiologii v. 26 (no. 1); ref. 6133] New Zealand Plateau, 51°44'S, 169°36'E, depth 230-286 meters. Holotype (unique): ZIN 45284. Type catalog: Balushkin et al. 2011:1020 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. •Valid as Ophthalmolycus campbellensis Andriashev & Fedorov 1986 -- (Paulin et al. 1989:211 [ref. 24556], Anderson 1990:5 [ref. 16143], Anderson 1992:8 [ref. 18912], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:33 [ref. 27603], Anderson & Møller 2007:183 [ref. 29401], Matallanas 2009:55 [ref. 30103], Matallanas 2011:566 [ref. 30839], Balushkin et al. 2011:1020 [ref. 32449], Møller & Anderson 2015:1431 [ref. 34289]). Current status: Valid as Ophthalmolycus campbellensis Andriashev & Fedorov 1986. Zoarcidae: Lycodinae. Distribution: New Zealand region. Habitat: marine.

caribbaeum, Pachycara Anderson [M. E.], Somerville [R.] & Copley [J. T.] 2016:72, Figs. 2-4 [Zootaxa 4066 (no. 1); ref. 34148] Von Damm Vent Field, Mid-Cayman Spreading Centre, 18°22.503'N, 81°47.858'W, depth 2313 meters. Holotype: BMNH 2015.9.10.1. Paratypes: MCZ. Plus tentatively referred specimen MCZ. •Valid as Pachycara caribbaeum Anderson, Somerville & Copley 2016. Current status: Valid as Pachycara caribbaeum Anderson, Somerville & Copley 2016. Zoarcidae: Lycodinae. Distribution: Caribbean Sea. Habitat: marine.

carmenae, Santelmoa Matallanas [J.] 2010:[2], Figs. 1-4 [Polar Biology v. 33; ref. 30666] Gerlache Strait, 64°32'58"S, 61°97'58"W, depth 1056 meters. Holotype: UAB BO3GSZ51. Paratypes: UAB B03GSZ33 (1), -Z34 (1), -Z42 (1), -Z43 (1), -Z44 (1), -Z59 (1), -Z60 (1), -Z344 (1). Appeared first electronically, available in 2010. •Valid as Santelmoa carmenae Matallanas 2010 -- (Matallanas et al. 2012:1402 [ref. 31950], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Santelmoa carmenae Matallanas 2010. Zoarcidae: Lycodinae. Distribution: Gerlache Strait, Southern Ocean. Habitat: marine.

caudimaculatus, Lycodes Matsubara [K.] 1936:115, Fig. 1 [Journal of the Imperial Fisheries Institute Tokyo v. 31 (no. 2); ref. 13718] Off Owase, Mie Prefecture, Japan, depth 80-160 fathoms. Holotype: not at FAKU. •Valid as Lycodes caudimaculatus Matsubara 1936 -- (Lindberg & Krasyukova 1975:144 [ref. 7348], Yamakawa in Okamura et al. 1982:243, 385 [ref. 8256], Toyoshima in Amaoka et al. 1983:139, 209 [ref. 8379], Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:220 [ref. 5722], Anderson 1994:118 [ref. 21438], Shinohara et al. 1996:182 [ref. 26605], Nakabo 2000:1031 [ref. 25182], Shinohara et al. 2001:328 [ref. 25995], Nakabo 2002:1031 [ref. 26193], Anderson & Fedorov 2004:24 [ref. 27603], Anderson & Fedorov 2004:24 [ref. 27603], Balushkin et al. 2011:992 [ref. 32449], Shinohara et al. 2014:265 [ref. 33330]). Current status: Valid as Lycodes caudimaculatus Matsubara 1936. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Japan. Habitat: marine.

celatus, Lycodes Jensen [A. S.] 1902:208 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Kara-Havet, depth 59 fathoms. Syntypes: ZMUC. Type catalog: Nielsen 1974:76 [ref. 9588]. •Synonym of Lycodes rossi Malmgren 1865 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603], Balushkin et al. 2011:1010 [ref. 32449], Parin et al. 2014:420 [ref. 33547]). Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae. Habitat: marine.

cerberus, Thermarces Rosenblatt [R. H.] & Cohen [D. M.] 1986:74, Figs. 1-5, 7-8 [Transactions of the San Diego Society of Natural History v. 21 (no. 4); ref. 5198] Hydrothermal vent site (20°51'N, 109°04'W), off Mexico, eastern Pacific, depth 2600 meters. Holotype: SIO 81-155. Paratypes: LACM 43719-1 [ex SIO 82-45] (1); SIO 81-155 (2). Additional material: LACM 43531-1 (1); SIO 82-46 (1, partial, c&s). •Valid as Thermarces cerberus Rosenblatt & Cohen 1986 -- (Anderson 1994:120 [ref. 21438], Geistdoerfer & Seuront 1996:167 [ref. 22308], Geistdoerfer 1999:8 [ref. 23832], Machida & Hashimoto 2002:1 [ref. 25949], Chernova & Geistdorfer 2003:153 [ref. 26887], Anderson & Fedorov 2004:37 [ref. 27603], McCosker & Rosenblatt 2010:193 [ref. 30957], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Thermarces cerberus Rosenblatt & Cohen 1986. Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

chauliodus, Lycenchelys Anderson [M. E.] 1995:66, Fig. 2 [Proceedings of the California Academy of Sciences v. 49 (no. 2); ref. 21928] Off Trujillo, Peru, 7°58'S, 80°37'W, Anton Bruun station 88, depth 1105-1124 meters. Holotype (unique): LACM 45600-1. •Valid as Lycenchelys chauliodus Anderson 1995 -- (Anderson & Fedorov 2004:16 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys chauliodus Anderson 1995. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific. Habitat: marine.

chilensis, Crossolycus Regan [C. T.] 1913:247 [Transactions of the Royal Society of Edinburgh v. 49 (pt 2, no. 2); ref. 3651] Cape Espíritu Santo, Tierra del Fuego, Chile. Holotype (unique): ZMB 16693. •Valid as Crossostomus chilensis (Regan 1913) -- (Gosztonyi 1977:205 [ref. 6103], Pequeño 1989:48 [ref. 14125], Anderson & Gosztonyi 1991:5 [ref. 19348], Anderson 1994:116 [ref. 21438], López et al. 1996:31 [ref. 23681], Pequeño 1999:174 [ref. 24566], Anderson & Fedorov 2004:12 [ref. 27603], Anderson 2006:12 [ref. 28576], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Crossostomus chilensis (Regan 1913). Zoarcidae: Lycodinae. Distribution: Eastern South Pacific and western South Atlantic. Habitat: marine.

chilensis, Ophthalmolycus Anderson [M. E.] 1992:3, Pl. 1 (fig. A) [The J.L.B Smith Institute of Ichthyology Special Publication No. 53; ref. 18912] Off Tocopilla, Chile, 22°42'S, 70°35'W, depth 1000 meters. Holotype (unique): CAS 63058. •Valid as Ophthalmolycus chilensis Anderson 1992 -- (Anderson 1994:120 [ref. 21438], Pequeño 1997:83 [ref. 23536], Anderson & Fedorov 2004:34 [ref. 27603], Matallanas 2009:55 [ref. 30103], Matallanas 2011:566 [ref. 30839], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Ophthalmolycus chilensis Anderson 1992. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific. Habitat: marine.

cicatrifer, Lycodes Garman [S.] 1899:136, Pl. 31 (fig. 1) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Off Mariato Point, Panama Bay, 6°17'N, 82°5'W, Albatross station 3360, depth 1672 fathoms. Holotype (unique): MCZ 28684 (in 2 pieces). •Valid as Lycenchelys cicatrifer (Garman 1899) -- (Anderson 1994:117 [ref. 21438], Anderson 1995:68 [ref. 21928], Anderson & Fedorov 2004:16 [ref. 27603], Balushkin et al. 2011:980 [ref. 32449], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys cicatrifer (Garman 1899). Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

ciliatus, Blennius Mitchill [S. L.] 1814:6 [Report, in part, of Samuel L. Mitchill; ref. 3030] New York, U.S.A. No types known. Also in Mitchill 1815:374 [ref. 13292]. •Synonym of Zoarces americanus (Bloch & Schneider 1801) -- (Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:2 [ref. 27603]). Current status: Synonym of Zoarces americanus (Bloch & Schneider 1801). Zoarcidae: Zoarcinae. Habitat: marine.

cinerea, Bentartia Matallanas [J.] 2010:[7], Figs. 5-8 [Polar Biology v. 33; ref. 30666] Gerlache Strait, Southern Ocean, 64°32'58"S, 61°97'38"W, depth 1056 meters. Holotype: UAB B03GSZ11. Paratypes: UAB B03GSZ30 (1), -Z47 (1). Appeared first electronically, available in 2010. •Valid as Bentartia cinerea Matallanas 2010 -- (Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Bentartia cinerea Matallanas 2010. Zoarcidae: Lycodinae. Distribution: Gerlache Strait, Southern Ocean. Habitat: marine.

coccineus, Lycodes Bean [T. H.] 1881:144 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Big Diomede Island [Chukotsky District, Chukotka Autonomous Okrug, Russia], Bering Strait. Holotype (unique): USNM 27748. Type catalog: Springer & Anderson 1997:16 [ref. 22953]. •Questionably a synonym of Lycodes mucosus Richardson 1855 (Balushkin et al. 2011:1002 [ref. 32449]). •Synonym of Lycodes mucosus Richardson 1855 -- (Toyoshima 1985:200 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:708 [ref. 25968], Anderson & Fedorov 2004:26 [ref. 27603], Parin et al. 2014:417 [ref. 33547]). Current status: Synonym of Lycodes mucosus Richardson 1855. Zoarcidae: Lycodinae. Habitat: marine.

colletti, Lycodes Popov [A. M.] 1931:143, Pl. 2 (fig. 6) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] Okhotsk Sea, 59°11.5'N, 148°19.5'E, depth 141 meters, 57°40'N, 147°50.6'E, depth 144 meters. Syntypes: ZIN 23758 (1), 29996 (1). Type catalog: Balushkin et al. 2011:1015 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667], •Valid as Lycodes colletti Popov 1931 -- (Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:113 [ref. 27579], Anderson & Fedorov 2004:30 [ref. 27603]), •Synonym of Lycodes uschakovi Popov 1931 -- (Lindberg & Krasyukova 1975:147 [ref. 7348], Anderson 1994:119 [ref. 21438], Balushkin et al. 2011:1015 [ref. 32449], Parin et al. 2014:423 [ref. 33547]). Current status: Synonym of Lycodes uschakovi Popov 1931. Zoarcidae: Lycodinae. Habitat: marine.

concolor, Lycodes Roule [L.] & Despax [R.] 1911:280 [Bulletin du Muséum National d'Histoire Naturelle (Série 1) v. 17 (no. 5); ref. 14869] South Shetland Islands, Antarctic. Holotype (unique): MNHN 1911-0099 (poor condidtion). Type catalog: Bauchot 1967:62-63 [ref. 20734]. Objectively invalid; preoccupied by Lycodes concolor Gill & Townsend 1897, replaced by Austrolycichthys dolloi Andriashev & Fedorov 1986. Described as new again in Roule 1913:19, Pl. 2 (fig. 3), Pl. 3 (fig. 6) [ref. 3815]. •Valid as Austrolycichthys concolor (Roule & Despax 1911) -- (Pequeño 1989:48 [ref. 14125]). •Synonym of Ophthalmolycus amberensis (Tomo, Marschoff & Torno 1977) -- (Anderson 1988:82 [ref. 7304], Anderson 1990:273 [ref. 21695], Miller 1993:655 [ref. 21297], Anderson & Fedorov 2004:33 [ref. 27603], Balushkin et al. 2011:1019 [ref. 32449]). Current status: Synonym of Ophthalmolycus amberensis (Tomo, Marschoff & Torno 1977). Zoarcidae: Lycodinae. Habitat: marine.

concolor, Lycodes Gill [T. N.] & Townsend [C. H.] 1897:233 [Proceedings of the Biological Society of Washington v. 11; ref. 1749] Bering Sea, 55°19'N, 168°11'W, Albatross station 3608, depth 276 fathoms. Holotype (unique): USNM 48764. Type catalog: Springer & Anderson 1997:16 [ref. 22953]. •Valid as Lycodes concolor Gill & Townsend 1897 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:725 [ref. 25968], Møller 2001:975 [ref. 25812], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Balushkin et al. 2011:991 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:409 [ref. 33547], Kells et al. 2016:272 [ref. 35888]). Current status: Valid as Lycodes concolor Gill & Townsend 1897. Zoarcidae: Lycodinae. Distribution: North Pacific. Habitat: marine.

conorhynchus, Gymnelis Garman [S.] 1899:131, Pl. 31 (fig. 2) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Off (Pacific) Panama Bay, 6°21'N, 80°41'W, Albatross station 3382, depth 1793 fathoms. Syntypes: MCZ 28690 (3, poor condition). •Valid as Ophthalmolycus conorhynchus (Garman 1899) -- (Anderson 1990:8 [ref. 16143], Anderson 1992:6 [ref. 18912], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:34 [ref. 27603], Matallanas 2009:55 [ref. 30103], Matallanas 2011:566 [ref. 30839], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Ophthalmolycus conorhynchus (Garman 1899). Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

corteziana, Aprodon Gilbert [C. H.] 1890:107 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Cortez Banks off San Diego, California, U.S.A., 33°53'30"N, 119°41'30"W, Albatross station 2948, depth 266 fathoms. Lectotype: USNM 46457. Paralectotypes: (5) CAS-SU 2780 (1); USNM 47056-57 (1, 1), 47180 (1), 47198 (1). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:16 [ref. 22953]. Lectotype established by Jordan & Evermann 1898:2461 [ref. 2445] and as figured specimen in caption to Pl. 349, p. 3304 in Jordan & Evermann 1900 [ref. 2446]. •Valid as Aprodon cortezianus Gilbert 1890 -- (Toyoshima 1985:237 [ref. 5722], Toyoshima 1985:237 [ref. 5722], Balushkin et al. 2011:973 [ref. 32499]). •Valid as Lycodes cortezianus (Gilbert 1890) -- (Eschmeyer & Herald 1983:105 [ref. 9277], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:705 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Page et al. 2013:162 [ref. 32708], Pietsch & Orr 2015:58 [ref. 34694], Kells et al. 2016:272 [ref. 35888]). Current status: Valid as Lycodes cortezianus (Gilbert 1890). Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

cousinsi, Pachycara Møller [P. R.] & King [N.] 2007:590, Fig. 3 [Copeia 2007 (no. 3); ref. 29259] Crozet Plateau, 45°40'S, 56°33'E, southern Indian Ocean, depth 4267-4270 meters. Holotype (unique): ZMUC P766470. •Valid as Pachycara cousinsi Møller & King 2007 -- (Shinohara 2012:55 [ref. 32102]). Current status: Valid as Pachycara cousinsi Møller & King 2007. Zoarcidae: Lycodinae. Distribution: Crozet Plateau, southern Indian Ocean. Habitat: marine.

crassiceps, Lycenchelys Roule [L.] 1916:16 [Bulletin de l'Institut Océanographique (Monaco) No. 320; ref. 3818] Gulf of Gascony, 45°02'N, 3°16'W, station 1453, depth 1455 meters. Syntypes: MOM 0091-0656 (2). Type catalog: Belloc 1949:14 [ref. 21544]. See also Roule 1919:64, Pl. 5 (fig. 1) [ref. 19805]. •Valid as Lycodes crassiceps (Roule 1916) -- (Andriashev in Whitehead et al. 1986:1137 [ref. 13677]). •Valid as Pachycara crassiceps (Roule 1916) -- (Anderson 1989:228 [ref. 13487], Anderson & Almeida 1990:889 [ref. 19328], Anderson 1991:157 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:227 [ref. 22911], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:34 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson & Mincarone 2006:25 [ref. 28658], Balushkin et al. 2011:1020 [ref. 32449], Anderson in Carpenter & De Angelis 2016:2760 [ref. 34618]). Current status: Valid as Pachycara crassiceps (Roule 1916). Zoarcidae: Lycodinae. Distribution: Eastern Atlantic. Habitat: marine.

crassilabris, Lycodopsis Gilbert [C. H.] 1890:106 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., 33°08'N, 118°40'W, Albatross station 2839, depth 414 fathoms. Holotype (unique): USNM 44280 (deteriorated). Type catalog: Springer & Anderson 1997:16 [ref. 22953]. •Synonym of Lycenchelys crotalinus (Gilbert 1890) -- (Anderson 1994:117 [ref. 21438], Anderson 1995:69 [ref. 21928], Anderson & Fedorov 2004:16 [ref. 27603], Parin et al. 2014:398 [ref. 33547]). Current status: Synonym of Lycenchelys crotalinus (Gilbert 1890). Zoarcidae: Lycodinae. Habitat: marine.

crossacanthum, Pachycara Anderson [M. E.] 1989:230, Figs. 7-8 [Proceedings of the California Academy of Sciences v. 46 (no. 10); ref. 13487] Off Senegal, 12°58.0'N, 17°41.8'W, depth 900 meters. Holotype: CAS 55586. Paratypes: CAS 62408 (1); MNHN 1988-1169 (1), 1988-1170 (3). •Valid as Pachycara crossacanthum Anderson 1989 -- (Anderson 1991:157 [ref. 20019], Anderson 1994:120 [ref. 21438], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson 2006:12 [ref. 28576], Anderson & Mincarone 2006:25 [ref. 28658], Tweddle & Anderson 2008:11 [ref. 29604], Balushkin et al. 2011:1027 [ref. 32449], Anderson in Carpenter & De Angelis 2016:2760 [ref. 34618]). Current status: Valid as Pachycara crossacanthum Anderson 1989. Zoarcidae: Lycodinae. Distribution: Eastern Atlantic: Senegal to Angola. Habitat: marine.

crotalinus, Lycodopsis Gilbert [C. H.] 1890:105 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off Santa Barbara Island, 33°49'45"N, 119°24'30"W, Albatross station 2980, depth 603 fathoms. Holotype (unique): USNM 44279. Type catalog: Springer & Anderson 1997:17 [ref. 22953]. •Valid as Embryx crotalinus (Gilbert 1890) -- (Eschmeyer & Herald 1983:104 [ref. 9277], Toyoshima 1985:178 [ref. 5722], Sheiko & Fedorov 2000:34 [ref. 25650], Balushkin et al. 2011:975 [ref. 32449]). •Valid as Lycenchelys crotalinus (Gilbert 1890) -- (McAllister 1990:157 [ref. 14674] as crotalina, Anderson 1994:117 [ref. 21438], Anderson 1995:69 [ref. 21928], Mecklenburg et al. 2002:693 [ref. 25968], Anderson & Fedorov 2004:16 [ref. 27603], Anderson 2006:186 [ref. 29415], Parin et al. 2014:398 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys crotalinus (Gilbert 1890). Zoarcidae: Lycodinae. Distribution: North Pacific. Habitat: marine.

crystallonota, Lycogramma Popov [A. M.] (ex Schmidt) 1933:151 [Issledovaniya Morey SSSR, Leningrad No. 19; ref. 6535] Sea of Japan, east of Petrov's Cape, Russia, depth 1225-1980 meters. Syntypes: (2) ZIN 24499 (now 3). Type catalog: Balushkin et al. 2011:972 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. •Valid as Allopis crystallonota Schmidt 1933 -- (Balushkin et al. 2011:972 [ref. 32449] with author as Schmidt). •Synonym of Bothrocara hollandi (Jordan & Hubbs 1925) -- (Anderson & Fedorov 2004:10 [ref. 27603], Anderson et al. 2009:179 [ref. 30175], Parin et al. 2014:385 [ref. 33547]). Current status: Synonym of Bothrocara hollandi (Jordan & Hubbs 1925). Zoarcidae: Lycodinae. Habitat: marine.

daidaleus, Exechodontes DeWitt [H. H.] 1977:790, Figs. 1-3 [United States National Marine Fisheries Service Fishery Bulletin v. 75 (no. 4); ref. 1124] Gulf of Mexico off Tampa Bay [not "east-southeast of Tampa Bay"], Florida, U.S.A., 27°01'N, 84°55'W, Oregon II station 10632, depth 275 fathoms. Holotype (unique): USNM 211797. Type catalog: Springer & Anderson 1997:17 [ref. 22953]. •Valid as Exechodontes daidaleus DeWitt 1977 -- (Anderson 1994:116 [ref. 21438], Anderson 2003:1741 [ref. 27106], Anderson & Fedorov 2004:13 [ref. 27603], McEachran & Fechhelm 2005:547 [ref. 28815], Balushkin et al. 2011:976 [ref. 32449]). Current status: Valid as Exechodontes daidaleus DeWitt 1977. Zoarcidae: Lycodinae. Distribution: Western Atlantic. Habitat: marine.

dearborni, Rhigophila DeWitt [H. H.] 1962:821, Figs. 1c-d, 2a-c, 3a-b [Copeia 1962 (no. 4); ref. 1122] McMurdo Sound, Antarctica, 77°53'S, 166°44'E, depth 585 meters. Holotype: CAS-SU 54145. Paratypes: ANSP (6, 4 lots); BMNH 1965.1.21.1-5 (5); FMNH 63867 (6); IRSNB 141-142 (1, 1); LACM (6, 6 lots); MCZ 41611 (6); MNHN 1962-0788 to 0789 (1, 1); NMNZ (2); CAS-SU 59564-73 and 59586 (36), 68929 (3); UMMZ 179558 (6); USNM 196978-79 (1, 4); ZIN 37037 (4); ZMB 27530 (2). Type catalog: Bauchot 1967:63 [ref. 20734], Böhlke 1984:165 [ref. 13621], Walschaets 1987:23-24 [ref. 20755], Hardy 1990:11 [ref. 18942], Ibarra & Stewart 1987:75 [ref. 12367], Springer & Anderson 1997:17 [ref. 22953], Balushkin et al. 2011:1017 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. •Valid as Lycodichthys dearborni (DeWitt 1962) -- (Anderson 1988:80 [ref. 7304], Anderson 1990:268 [ref. 21695], Miller 1993:652 [ref. 21297], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:31 [ref. 27603], Struthers & Møller 2009:65 [ref. 30403], Balushkin et al. 2011:1017 [ref. 32449]). Current status: Valid as Lycodichthys dearborni (DeWitt 1962). Zoarcidae: Lycodinae. Distribution: Southern Ocean: Ross Sea, Antarctica. Habitat: marine.

depressiceps, Austrolycus Regan [C. T.] 1913:245, Pl. 5 (fig. 1) [Transactions of the Royal Society of Edinburgh v. 49 (pt 2, no. 2); ref. 3651] Magellan Strait. Lectotype: BMNH 1868.1.15.25. Paralectotypes: (numerous) BMNH 1868.9.16.22-23 (2); NMSZ 1921.143.485 (13 in 4 lots, plus 1 lot disentigrated). Type catalog: Herman et al. 1990:2-3 [ref. 20727]. Lectotype selected by Gosztonyi 1977:200 [ref. 6103]. •Valid as Austrolycus depressiceps Regan 1913 -- (Gosztonyi 1977:200 [ref. 6103], Pequeño 1986:441 [ref. 6718], Pequeño 1989:48 [ref. 14125], Lloris & Rucabado 1991:66 [ref. 22530], Anderson 1994:116 [ref. 21438], López et al. 1996:30 [ref. 23681], Pequeño 1999:174 [ref. 24566] as deprecisseps, López et al. 2002:64 [ref. 26808], Anderson & Fedorov 2004:10 [ref. 27603], Balushkin et al. 2011:973 [ref. 32449]). Current status: Valid as Austrolycus depressiceps Regan 1913. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific and western South Atlantic. Habitat: marine.

derjugini, Lycodapus Andriashev [A. P.] 1935:422, Fig. [C. R. (Doklady) Acad. Sci. URSS v. 3 [8] (no. 9 [69]); ref. 13869] Ozernovski Bay, northeastern Kamchatka, Russia, Bering Sea, 57°31'N, 163°14.5'E. Holotype: ZIN 25107. Paratypes: ZIN 25106 (1). Type catalog: Balushkin et al. 2011:985 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. •Valid as Lycodapus derjugini Andriashev 1935 -- (Peden & Anderson 1978:1947 [ref. 8895], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Anderson & Fedorov 2004:21 [ref. 27603], Balushkin et al. 2011:985 [ref. 32449], Parin et al. 2014:404 [ref. 33547]). Current status: Valid as Lycodapus derjugini Andriashev 1935. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

dermatinus, Lycodapus Gilbert [C. H.] 1896:471, Pl. 35 (middle) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Off California coast, 37°54'10"N, 123°30'00"W, Albatross station 3162, depth 552 fathoms. Holotype (unique): USNM 53035. Type catalog: Springer & Anderson 1997:17 [ref. 22953]. •Valid as Lycodapus dermatinus Gilbert 1896 -- (Peden & Anderson 1978:1948 [ref. 8895], Anderson 1989:152 [ref. 13419], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:740 [ref. 25968], Anderson & Fedorov 2004:21 [ref. 27603], Balushkin et al. 2011:985 [ref. 32449]). Current status: Valid as Lycodapus dermatinus Gilbert 1896. Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

diapteroides, Lycodes brevipes Taranetz [A. Ya.] & Andriashev [A. P.] in Andriashev 1937:253 [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] Bering Sea and Bering Sea at 62°08'N, 178°32'W, depth 98 meters. Syntypes: ZIN 26886 (1), 30011 (3). Type catalog: Balushkin et al. 2011:991 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. •Valid as Lycodes diapteroides Taranetz & Andriashev 1937 -- (Toyoshima 1985:191 [ref. 5722]). •Synonym of Lycodes brevipes Bean 1890 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650] with question, Mecklenburg et al. 2002:716 [ref. 25968], Anderson & Fedorov 2004:23 [ref. 27603], Balushkin et al. 2011:991 [ref. 32449], Parin et al. 2014:409 [ref. 33547]). Current status: Synonym of Lycodes brevipes Bean 1890. Zoarcidae: Lycodinae. Habitat: marine.

diapterus, Lycodes Gilbert [C. H.] 1892:564 [Proceedings of the United States National Museum v. 14 (no. 880); ref. 1626] Off California, 33°55'30"N, 120°28'00"W, Albatross station 2896, depth about 688 meters. Lectotype: USNM 44385. Paralectotypes: BMNH 1900.9.29.245 (1) from Jordan; CAS-SU 176 (2); USNM 46716 (1), 125538 (1). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:17 [ref. 22953]. Lectotype established by Jordan & Evermann 1898:2473 [ref. 2445]. •Valid as Lycodes diapterus Gilbert 1892 -- (Lindberg & Krasyukova 1975:150 [ref. 7348], Eschmeyer & Herald 1983:105 [ref. 9277], Toyoshima 1985:230 [ref. 5722], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:724 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Shinohara & Shirai 2005:172 [ref. 28232], Stevenson & Sheiko 2009:127 [ref. 30070], Balushkin et al. 2011:993 [ref. 32449], Pietsch & Orr 2015:72 [ref. 34694], Kells et al. 2016:272 [ref. 35888]). Current status: Valid as Lycodes diapterus Gilbert 1892. Zoarcidae: Lycodinae. Distribution: North Pacific: western North Pacific to California (U.S.A.). Habitat: marine.

digitatus, Lycodes Gill [T. N.] & Townsend [C. H.] 1897:232 [Proceedings of the Biological Society of Washington v. 11; ref. 1749] Bering Sea, 56°14'N, 164°09'W, Albatross station 3541, depth 49 fathoms. Holotype (unique): USNM 48765. Type catalog: Springer & Anderson 1997:17 [ref. 22953]. •Synonym of Lycodes palearis Gilbert 1896 -- (Toyoshima 1985:212 [ref. 5722], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:719 [ref. 25968], Anderson & Fedorov 2004:27 [ref. 27603], Parin et al. 2014:417 [ref. 33547]). Current status: Synonym of Lycodes palearis Gilbert 1896. Zoarcidae: Lycodinae. Habitat: marine.

diporus, Gymnelus Chernova [N. V.] 2000:11 [7], Figs. 2b, 4 [Voprosy Ikhtiologii v. 40 (no. 1); ref. 24885] Bearing sea, Bering Island, Komandorskii Islands, depth 123 meters. Holotype: ZIN 25243. Paratypes: ZIN 34990 (1). Type catalog: Balushkin et al 2011:963 [ref. 32499], Balushkin et al. 2012:159 [ref. 33667]. Anderson in Anderson & Fedorov 2004:5 [ref. 27603] believes this is probably a synonym of G. hemifascaitus. On p. 7 of English translation. •Valid as Gymnelus diporus Chernova 2000 -- (Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:5 [ref. 27603], Balushkin et al.2011:963 [ref. 32499], Parin et al. 2014:391 [ref. 33547]). •Synonym of Gymnelus hemifasciatus Andriashev 1937 -- (Mecklenburg & Anderson 2015:277 [ref. 33742], Mecklenburg et al. 2016:154 [ref. 34440]). Current status: Synonym of Gymnelus hemifasciatus Andriashev 1937. Zoarcidae: Gymnelinae. Distribution: Western North Pacific [if valid]. Habitat: marine.

dolichaulus, Pachycara Anderson [M. E.] 2006:187, Figs. 3-5 [Species Diversity v. 11 (no. 3); ref. 29415] West of Trujillo, 8°18.8'S, 80°18.2'W, off Peru, depth 666-718 meters. Holotype (unique): HUMZ 167323. •Valid as Pachycara dolichaulus Anderson 2006 -- (Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara dolichaulus Anderson 2006. Zoarcidae: Lycodinae. Distribution: Southeastern Pacific: off Peru. Habitat: marine.

dolloi, Austrolycichthys Andriashev [A. P.] & Fedorov [V. V.] 1986:31 [Voprosy Ikhtiologii v. 26 (no. 1); ref. 6133] South Shetland Islands, Antarctic. Holotype (unique): MNHN 1911-0099. Replacement name for Lycodes concolor Roule & Despax 1911, preoccupied by Lycodes concolor Gill & Townsend 1897; however, an older synonym, Lycenchelys ambererensis Tomo, Marschoff & Torno 1977, was available. •Synonym of Ophthalmolycus amberensis (Tomo, Marschoff & Torno 1977) -- (Anderson 1988:82 [ref. 7304], Anderson 1990:273 [ref. 21695], Anderson & Fedorov 2004:33 [ref. 27603]). Current status: Synonym of Ophthalmolycus amberensis (Tomo, Marschoff & Torno 1977). Zoarcidae: Lycodinae. Habitat: marine.

dorsoscutatus, Lycodonus Oshima [M.] 1957:7, Fig. 5 [Japanese Journal of Ichthyology v. 6 (nos 1-2); ref. 12161] Offshore of Kamo, Yamagata Prefecture, Japan. Holotype (unique): Fish. Exp. Sta. Yamagata Pref. (lost). •Valid as Lycodonus dorsoscutatus Oshima 1957 -- (Koyanagi in Okamura et al. 1995:205 [ref. 22531]). •Status uncertain, not a zoarcid -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:31 [ref. 27603]). Zoarcidae. Habitat: marine.

eastmani, Ophthalmolycus Matallanas [J.] 2011:562, Figs. 1-2 [Journal of the Marine Biological Association of the United Kingdom v. 91 (no. 2); ref. 30839] Baited trap, 68°48'S, 86°38'W, Bellingshausen Sea, Southern Ocean, depth 1837 meters. Holotype: Uncat. Paratypes: Uncataloged (2). •Valid as Ophthalmolycus eastmani Matallanas 2011. Current status: Valid as Ophthalmolycus eastmani Matallanas 2011. Zoarcidae: Lycodinae. Distribution: Bellingshausen Sea, Southern Ocean. Habitat: marine.

effusus, Phucocoetes variegatus forma Smitt [F. A.] 1898:43, Pl. 5 (fig. 32) [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 24 (afd. 4) (no. 5); ref. 4147] Off River Grande, Terra del Fuego, Argentina, southwestern Atlantic. Syntypes: (2) NRM 11156 (2). •Synonym of Iluocoetes elongatus (Smitt 1898) -- (Gosztonyi 1977:215 [ref. 6103], Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:14 [ref. 27603], Matallanas & Corbella 2012:1 [ref. 31967]). Current status: Synonym of Iluocoetes elongatus (Smitt 1898). Zoarcidae: Lycodinae. Habitat: marine.

elegans, Pogonolycus Norman [J. R.] 1937:106, Fig. 55, Pl. 1 (fig. 3) [Discovery Reports v. 16; ref. 3227] Patagonian-Falkland region, 52°25'S, 61°00'W, Discovery station WS 246, depth 267-208 meters. Holotype: BMNH 1936.8.26.1031. Paratypes: (4) BMNH?. •Valid as Pogonolycus elegans Norman 1937 -- (Gosztonyi 1977:238 [ref. 6103], Pequeño 1989:49 [ref. 14125], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:37 [ref. 27603], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Pogonolycus elegans Norman 1937. Zoarcidae: Lycodinae. Distribution: Southern South America. Habitat: marine.

elongata, Bothrocaropsis Garman [S.] 1899:129, Pl. 33 (fig. 2) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Gulf of Panama, eastern Pacific, 7°N, 79-80°W, Albatross station 3353 and 3393, depth 695-1020 fathoms. Lectotype: MCZ 28691. Paralectotypes: MCZ 28692 (1). Lectotype selected by Anderson et al. 2009:179 [ref. 30175]. •Valid as Bothrocara elongatum (Garman 1899) -- (Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:10 [ref. 27603], Anderson et al. 2009:177 [ref. 30175]). •Valid as Bentartia elongata (Garman 1899) -- (Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Bentartia elongata (Garman 1899). Zoarcidae: Lycodinae. Distribution: Eastern Pacific: Gulf of Panama to off Chile. Habitat: marine.

elongatus, Nalbantichthys Schultz [L. P.] 1967:2, Figs. 1-3 [Proceedings of the United States National Museum v. 122 (no. 3598); ref. 3974] South-central Bering Sea, depth 300 meters. Holotype: USNM 200671. Paratypes: Inst. Cercitari Piscicole, Bucharest (1). Type catalog: Springer & Anderson 1997:17 [ref. 22953]. •Valid as Nalbantichthys elongatus Schultz 1967 -- (Anderson 1994:120 [ref. 21438], Mecklenburg et al. 2002:686 [ref. 25968], Anderson & Fedorov 2004:8 [ref. 27603], Balushkin et al. 2011:970 [ref. 32449], Parin et al. 2014:425 [ref. 33547]). Current status: Valid as Nalbantichthys elongatus Schultz 1967. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Bering Sea. Habitat: marine.

elongatus, Phucocoetes variegatus forma Smitt [F. A.] 1898:44, Pl. 5 [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 24 (afd. 4) (no. 5); ref. 4147] Puerto Madryn, 43°S, Argentina, sw Atlantic. Holotype (unique): NRM 11158. •Valid as Iluocoetes elongatus (Smitt 1898) -- (Gosztonyi 1977:215 [ref. 6103], Lloris & Rucabado 1987:56 [ref. 15955], Pequeño 1989:48 [ref. 14125], Anderson 1994:117 [ref. 21438], Pequeño 2000:88 [ref. 25224], Anderson & Fedorov 2004:14 [ref. 27603], Reyes 2007:75 [ref. 31564] as elongatum, Matallanas & Corbella 2012:1 [ref. 31967], Balushkin et al. 2011:976 [ref. 32449]). •Valid as Argentinolycus elongatus (Smitt 1898) -- (Matallanas & Corbella 2012:6 [ref. 31967]). Current status: Valid as Argentinolycus elongatus (Smitt 1898). Zoarcidae: Lycodinae. Distribution: Southwestern Atlantic. Habitat: marine.

elongatus, Zoarces Kner [R.] 1868:30 [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 58 (nos 1-2); ref. 6074] Decastris Bay, Tatar Strait, Sea of Japan. No types known. Appeared in more detail in Kner 1868:344 [52 of separate], Pl. 7 (fig. 22) [ref. 2646]. •Valid as Enchelyopus elongatus (Kner 1868) -- (Cheng & Zhou 1997:335 [ref. 26385]). •Valid as Zoarces elongatus Kner 1868 -- (Lindberg & Krasyukova 1975:132 [ref. 7348], Toyoshima in Masuda et al. 1984:305 [ref. 6441], Anderson 1994:120 [ref. 21438], Sokolovskaya et al. 1998:12 [ref. 24670], Nakabo 2000:1028 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Wang et al. 2001:246 [ref. 26566], Zemnukhov et al. 2001:420 [ref. 25412], Nakabo 2002:1028 [ref. 26193], Fedorov et al. 2003:120 [ref. 27579], Anderson & Fedorov 2004:2 [ref. 27603], Bogutskaya & Naseka 2004:208 [ref. 28183], Parin et al. 2005:439 [ref. 28290], Chereshev et al. 2007:555 [ref. 29371], Voskoboinikova et al. 2010:704 [ref. 31095], Balushkin et al. 2011:953 [ref. 32449], Parin et al. 2014:429 [ref. 33547], Dyldin & Orlov 2017:74 [ref. 35138]). Current status: Valid as Zoarces elongatus Kner 1868. Zoarcidae: Zoarcinae. Distribution: Western North Pacific. Habitat: brackish, marine.

elvirae, Santelmoa Matallanas [J.] 2011:1232, Figs. 1-4 [Journal of the Marine Biological Association of the United Kingdom v. 91 (Sp. issue 6); ref. 31444] Bellingshausen Sea, 68°48'S, 86°38'W, Antarctic, baited trap in 1837 meters. Holotype: UAB:B06MBZ39. Paratypes: UAB:B06MBZ36, -37 and -38 (3). •Valid as Santelmoa elvirae Matallanas 2011 -- (Matallanas et al. 2012:1402 [ref. 31950]) Current status: Valid as Santelmoa elvirae Matallanas 2011. Zoarcidae: Lycodinae. Distribution: Bellingshausen Sea, Antarctica. Habitat: marine.

endemoscotus, Lycodapus Peden [A. E.] & Anderson [M. E.] 1978:1936, Fig. 14 [Canadian Journal of Zoology v. 56 (no. 9) (for Sept. 1978); ref. 8895] North Pacific, 45°44.4'N, 125°26.6'W, depth 2225 meters. Holotype: USNM 216471. Paratypes: AMNH 12838 (1); CAS 36826 (2), 36829-30 (1, 2); LACM 31922-1 (1); NMC (2 lots); OS (2 lots); SIO (8 lots, 17 spec.); UBC (2 lots); USNM 216472-73 (2, 1); UW (3 lots). Type catalog: Springer & Anderson 1997:17 [ref. 22953]. •Valid as Lycodapus endemoscotus Peden & Anderson 1978 -- (Anderson 1989:151 [ref. 13419], McAllister 1990:158 [ref. 14674], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:738 [ref. 25968], Balanov 2003:565 [ref. 27175] as cf endemoscotus, Anderson & Fedorov 2004:22 [ref. 27603], Balushkin et al. 2011:985 [ref. 32449], Parin et al. 2014:404 [ref. 33547]). Current status: Valid as Lycodapus endemoscotus Peden & Anderson 1978. Zoarcidae: Lycodinae. Distribution: North Pacific Ocean: Okhotsk Sea; British Columbia to northern Baja California. Habitat: marine.

esipovi, Gymnelus Chernova [N. V.] 1999:306, Figs. 1-2 [Voprosy Ikhtiologii v. 39 (no. 3); ref. 24868] Kara Sea, 78°53'N, 80°34'E, depth 40 meters. Holotype: ZIN 30558. Paratypes: ZIN 14139 (3), 29453 (2), 32073 (1), 51568 (1), 51569 (1). Plus non-type material. Type datalog: Balushkin et al 2011:963 [ref. 32499], Balushkin et al. 2012:159 [ref. 33667]. Anderson in Anderson & Fedorov 2004:5 [ref. 27603] believes this is probably a synonym of G. retrodorsalis. •Synonym of Gymnelus retrodorsalis Le Danois 1913 with que`stion -- (Mecklenburg et al. 2011:124, 128 [ref. 31212], Mecklenburg & Anderson 2015:284 [ref. 33742]), •Valid as Gymnelus esipovi Chernova 1999 -- (Chernova 2000:8 [ref. 24885], Anderson & Fedorov 2004:5 [ref. 27603], Balushkin et al.2011:963 [ref. 32499], Dolgov 2013:919 [ref. 33112], Parin et al. 2014:391 [ref. 33547]). Current status: Valid as Gymnelus esipovi Chernova 1999. Zoarcidae: Gymnelinae. Distribution: Arctic Ocean: northern Kara Sea, Barents Sea, and off Spitzbergen [if valid]. Habitat: marine.

esmarkii, Lycodes Collett [R.] 1875:95 [Norges fiske; ref. 884] Varanger Fjord, northern Norway. Syntypes: BMNH 1890.11.1.1 (1) from Collett; MNHN 1890-0369 [ex ZMUO] (1); ZMUO J4496 (1), J4498 (1). Type catalog: Pethon 1969:11 [ref. 19268]. •Valid as Lycodes esmarkii Collett 1875 -- (Andriashev 1973:543 [ref. 9214] as esmarki, Andriashev in Whitehead et al. 1986:1138 [ref. 13677] as esmarki, Scott & Scott 1988:407 [ref. 25518] as esmarki, McAllister 1990:159 [ref. 14674] as esmarki, Anderson 1994:118 [ref. 21438], Saito & Okamura in Okamura et al. 1995:194 [ref. 22531], Møller 2000:847 [ref. 24585], Moore et al. 2003:228 [ref. 27076], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Møller et al. 2010:60 [ref. 31169], Balushkin et al. 2011:993 [ref. 32449], Page et al. 2013:162 [ref. 32708], Dolgov 2013:920 [ref. 33112], Parin et al. 2014:410 [ref. 33547], Afonso et al. 2016:63, 196 [ref. 36199], Mecklenburg et al. 2018:322 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes esmarkii Collett 1875. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

eudipleurostictus, Lycodes Jensen [A. S.] 1902:206 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Faroe Islands to Spitzbergen, northern Atlantic, depth 350-471 fathoms. Syntypes: NRM 25527 (1); ZMUC 7, 138; ZMUO (4). Type catalog: Nielsen 1974:77 [ref. 9588]. •Valid as Lycodes eudipleurostictus Jensen 1902 -- (Andriashev 1973:543 [ref. 9214], McAllister et al. 1981:830 [ref. 5431], Andriashev in Whitehead et al. 1986:1138 [ref. 13677], Anderson 1994:118 [ref. 21438], Saito & Okamura in Okamura et al. 1995:195 [ref. 22531], Mecklenburg et al. 2002:723 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603], Shinohara & Shirai 2005:171 [ref. 28232], Møller et al. 2010:60 [ref. 31169], Balushkin et al. 2011:993 [ref. 32449], Page et al. 2013:162 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:410 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:63, 197 [ref. 36199], Mecklenburg et al. 2016:162 [ref. 34440], Mecklenburg et al. 2018:324 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes eudipleurostictus Jensen 1902. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

extensus, Lycodapus Gilbert [C. H.] 1896:455 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] North of Unalaska Island, Alaska, U.S.A., 53°33'50"N, 167°46'50"W, Albatross station 3324, depth 109 fathoms. Holotype (unique): USNM 48638 (poor condition). Type catalog: Springer & Anderson 1997:18 [ref. 22953]. •Status uncertain, nomen dubium -- (Peden & Anderson 1978:1957 [ref. 8895], Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:48 [ref. 27603]). Zoarcidae: Lycodinae. Habitat: marine.

facali, Iluocoetes Lloris [D.] & Rucabado [J. A.] 1987:54, Fig. 1 [Thalassas: An International Journal of Marine Sciences v. 5 (no. 1); ref. 15955] Puerto Harberton, Beagle Canal, 54°52'42"S, 67°20'00"W, Tierra del Fuego, Argentina. Holotype (unique): IIPB 114/1987. Type catalog: Allué et al. 2000:75 [ref. 30499]. •Synonym of Iluocoetes fimbriatus Jenyns 1842 -- (Anderson & Gosztonyi 1991:15 [ref. 19348], Anderson & Fedorov 2004:14 [ref. 27603]). •Valid as Iluocoetes facali Lloris & Rucabado 1987 -- (Lloris & Rucabado 1991:73 [ref. 22530], López et al. 1996:32 [ref. 23681], Pequeño 1997:83 [ref. 23536], Reyes 2007:71 [ref. 31564]). Current status: Valid as Iluocoetes facali Lloris & Rucabado 1987. Zoarcidae: Lycodinae. Distribution: Southern South America: Argentina and Chile. Habitat: marine.

fasciatus, Ilucocoetes fimbriatus Lönnberg [E.] 1905:20 [Wissenschaftliche Ergebnisse der Schwedischen Südpolar-Expedition, 1901-1903 v. 5 (no. 6); ref. 2839] Greenpatch, Port Louis, Falkland Islands, depth 7 meters. Holotype (unique): NRM 11089. Original genus should have been Iluocoetes. •Valid as Crossostomus fasciatus (Lönnberg 1905) -- (Gosztonyi 1977:207 [ref. 6103], Nakamura in Nakamura et al. 1986:240 [ref. 14235], Pequeño 1989:48 [ref. 14125], Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:12 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Crossostomus fasciatus (Lönnberg 1905). Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

fasciatus, Lycenchelys Schmidt [P. Yu.] 1904:203, Pl. 6 (fig. 2) [Fishes of the eastern seas of the Russian Empire; ref. 3946] Aniva Bay, southern Sakhalin Island, Russia, Okhotsk Sea, depth 126-161 feet (17-22 Russian fathoms). Syntypes: ZIN 12446 (1, lost), 13092-93 (1, 1). Type catalog: Balushkin et al. 2011:993 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. Appeared first without distinguishing features in Schmidt 1903:520 (18 of separate) [ref. 3945]. •Synonym of Lycodes palearis Gilbert 1896, but a valid subspecies -- (Lindberg & Krasyukova 1975:157 [ref. 7348], Sokolovskaya et al. 1998:12 [ref. 24670], Shinohara et al. 2011:51 [ref. 31715], Shinohara et al. 2014:266 [ref. 33330]). •Valid as Lycodes fasciatus (Schmidt 1904) -- (Toyoshima 1985:214 [ref. 5722], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:720 [ref. 25968], Sheiko & Fedorov 2000:36 [ref. 25650], Møller 2001:111 [ref. 25378], Fedorov et al. 2003:113 [ref. 27579], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Balushkin et al. 2011:993 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:410 [ref. 33547], Mecklenburg et al. 2016:173 [ref. 34440]). Current status: Valid as Lycodes fasciatus (Schmidt 1904). Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

fedorovi, Lycenchelys Anderson [M. E.] & Balanov [A. A.] 2000:1056, Figs. 1-2 [Copeia 2000 (no. 4); ref. 25023] Bank east of Krusenstern Strait, Kuril Islands, 47°50'N, 154°25'E, depth 550 meters. Holotype: MIMB 2382. Paratypes: HUMZ 161930-36 (7); MIMB 2380-81 (1, 1), 3019-23 (now 8, 3, now 6, 1, 1), 3171-73 (1, 1, 1); NSMT P-57330-32 (6, 1, 1); ZIN 53489 (2), 53490 (1). Type catalog: Balushkin et al. 2011:980 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. •Valid as Lycenchelys fedorovi Anderson & Balanov 2000 -- (Anderson & Imamura 2002:357 [ref. 26499], Anderson & Fedorov 2004:17 [ref. 27603], Balushkin et al. 2011:980 [ref. 32449], Thiel et al. 2018:52 [ref. 35743] as federovi). Current status: Valid as Lycenchelys fedorovi Anderson & Balanov 2000. Zoarcidae: Lycodinae. Distribution: North Pacific: Kuril Islands and Kamchatka. Habitat: marine.

fedorovi, Zoarces Chereshnev [I. A.], Nazarkin [M. V.] & Chegodaeva [E. A.] 2007:556, Figs 1a, 2-3 [Journal of Ichthyology v. 47 (no. 8); ref. 29371] Mouth of Kulkuta River, Odyan Bay, Taui Bay, Sea of Okhotsk, Russia. Holotype: ZIN 54146. Paratypes: ZIN 54147 (8). Plus non-type material. Type catalog: Balushkin 2011:955 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. •Valid as Zoarces fedorovi Chereshnev, Nazarkin & Chegodaeva 2007 -- (Voskoboinikova et al. 2010:704 [ref. 31095], Balushkin et al. 2011:955 [ref. 32449], Parin et al. 2014:430 [ref. 33547]). Current status: Valid as Zoarces fedorovi Chereshnev, Nazarkin & Chegodaeva 2007. Zoarcidae: Zoarcinae. Distribution: Western North Pacific: Sea of Okhotsk. Habitat: marine.

fierasfer, Lycodapus Gilbert [C. H.] 1890:108 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Gulf of California, 27°23'45"N, 111°25'00"W, Albatross station 3010 depth 1105 fathoms. Lectotype: CAS-SU 845. Paralectotypes: USNM 43094 (3) station 3072, 53063 (6?, in pieces). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:18 [ref. 22953]. Lectotype established (as figured specimen) in Jordan 1896:244 [ref. 2395] from station 2980, but that specimen (from station 2980) is apparently now missing. A lectotype was designated by Peden & Anderson 1978:1953 [ref. 8895]. •Valid as Lycodapus fierasfer Gilbert 1890 -- (Peden & Anderson 1978:1953 [ref. 8895], Anderson 1989:152 [ref. 13419], McAllister 1990:158 [ref. 14674], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:739 [ref. 25968], Anderson & Fedorov 2004:22 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Balushkin et al. 2011:986 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:405 [ref. 33547]). Current status: Valid as Lycodapus fierasfer Gilbert 1890. Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

fimbriatus, Iluocoetes Jenyns [L.] 1842:166, Pl. 29 (figs. 2, 2a) [The zoology of the voyage of H. M. S. Beagle; ref. 2344] Chiloé Archipelago, Chile. Holotype (unique): BMNH 1917.7.14.69. •Valid as Iluocoetes fimbriatus Jenyns 1842 -- (Gosztonyi 1977:213 [ref. 6103], Nakamura in Nakamura et al. 1986:232 [ref. 14235], Lloris & Rucabado 1987:56 [ref. 15955], Pequeño 1989:48 [ref. 14125], Anderson 1994:117 [ref. 21438], López et al. 1996:32 [ref. 23681], Figueiredo in Menezes et al. 2003:93 [ref. 27192], Anderson & Fedorov 2004:14 [ref. 27603], Reyes 2007:74 [ref. 31564], Mabragaña et al. 2011:table S2 [ref. 31800], Matallanas & Corbella 2012:2 [ref. 31967], Balushkin et al. 2011:977 [ref. 32449], Nión et al. 2016:51 [ref. 35565] dated 1942). Current status: Valid as Iluocoetes fimbriatus Jenyns 1842. Zoarcidae: Lycodinae. Distribution: Southeastern Pacific to western South Atlantic off South America. Habitat: marine.

flaccidum, Melanostigma Waite [E. R.] 1914:129, Pl. 6 [Transactions of the New Zealand Institute v. 46 (art. 22); ref. 14178] Kaikoura, New Zealand, at surface. Holotype (unique): whereabouts unknown. •Synonym of Melanostigma gelatinosum Günther 1881 -- (Anderson 1988:62 [ref. 7304], Anderson 1990:270 [ref. 21695], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:8 [ref. 27603]). •Valid as Melanostigma flaccidum Waite 1914 -- (Balushkin 2019:287 [ref. 36807]). Current status: Valid as Melanostigma flaccidum Waite 1914. Zoarcidae: Gymnelinae. Distribution: East coast of South Island, New Zealand. Habitat: marine.

flagellicauda, Lycenchelys Jensen [A. S.] 1902:210 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Spitzbergen to Faroe Islands, depth 459-1089 fathoms. Syntypes: ZIN 40722 [ex. ZMUC 28]; ZMUC 21-31; ZMUO J4589 (1), J4590 (2). Type catalog: Pethon 1969:12 [ref. 19268], Nielsen 1974:76 [ref. 9588], Balushkin et al. 2011:1017 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. •Valid as Lycodonus flagellicauda (Jensen 1902) -- (Andriashev 1973:546 [ref. 9214], Gosztonyi 1981:156 [ref. 8596], Toyoshima 1985:143 [ref. 5722], Andriashev in Whitehead et al. 1986:1147 [ref. 13677], Anderson 1994:119 [ref. 21438], Koyanagi in Okamura et al. 1995:204 [ref. 22531], Anderson & Fedorov 2004:31 [ref. 27603], Møller et al. 2010:64 [ref. 31169], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin et al. 2011:1017 [ref. 32449], Parin et al. 2014:424 [ref. 33547], Mecklenburg et al. 2018:364 [ref. 36370] see remarks). Current status: Valid as Lycodonus flagellicauda (Jensen 1902). Zoarcidae: Lycodinae. Distribution: Arctic and North Atlantic. Habitat: marine.

flavus, Lycodes Boulenger [G. A.] 1900:53 [Annals and Magazine of Natural History (Series 7) v. 6 (no. 31); ref. 14720] Falkland Islands. Holotype: BMNH 1900.1.31.9. •Synonym of Phucocoetes latitans Jenyns 1842 -- (Lloris & Rucabado 1991:75 [ref. 22530], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:36 [ref. 27603]). Current status: Synonym of Phucocoetes latitans Jenyns 1842. Zoarcidae: Lycodinae. Habitat: marine.

folletti, Lycenchelys Anderson [M. E.] 1995:72, Fig. 4 [Proceedings of the California Academy of Sciences v. 49 (no. 2); ref. 21928] Guaymas basin, Gulf of California, Mexico, 27°40.0'N, 111°22.6'W, depth 931-953 meters. Holotype: SIO 68-83. Paratypes: SIO 68-83 (7). •Valid as Lycenchelys folletti Anderson 1995 -- (Anderson & Fedorov 2004:17 [ref. 27603], Shinohara & Anderson 2007:62 [ref. 29111], Balushkin et al. 2011:1026 [ref. 32449], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys folletti Anderson 1995. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific: Gulf of California. Habitat: marine.

frigida, Lycodes reticulatus forma Smitt [F. A.] 1901:29 [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 27 (no. 4); ref. 18732] Not intended as new; a subsequent treatment of Lycodes frigidus Collett 1879 as a subspecific name. •In the synonymy of Lycodes pallidus Collett 1879 -- (Møller 2001:983 [ref. 25812]). •Unavailable, in family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Zoarcidae: Lycodinae.

frigidus, Lycodes Collett [R.] 1879:45 [Forhandlinger i Videnskabs-selskabet i Christiania (for 1878) No. 14; ref. 885] Bear Island and Spitzbergen, Arctic Ocean, depth 475-2438 meters. Syntypes: BMNH 1885.12.2.1 (1) from Collett; MNHN 1890-0370 (1); NMW 77810 (1); USNM 22977 (1), ZMUB 1688 (1), 2223 (1); ZMUC 24 (1); ZMUO J4517-22 (6), plus 1 destroyed and some not traced. Type catalog: Pethon 1969:11 [ref. 19268], Nielsen 1974:77 [ref. 9588], Frøiland 1979:150 [ref. 21547], Springer & Anderson 1997:18 [ref. 22953]. •Valid as Lycodes frigidus Collett 1879 -- (Andriashev 1973:543 [ref. 9214], McAllister et al. 1981:829 [ref. 5431], Andriashev in Whitehead et al. 1986:1139 [ref. 13677], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Saito & Okamura in Okamura et al. 1995:196 [ref. 22531], Mecklenburg et al. 2002:726 [ref. 25968], Anderson & Fedorov 2004:25 [ref. 27603], Møller et al. 2010:60 [ref. 31169], Balushkin et al. 2011:995 [ref. 32449], Mecklenburg et al. 2014:144 ,145 [ref. 33463], Parin et al. 2014:411 [ref. 33547], Parin et al. 2014:411 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:63, 198 [ref. 36199], Mecklenburg et al. 2016:164 [ref. 34440], Mecklenburg et al. 2018:326 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes frigidus Collett 1879. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

fulvus, Lycodes Toyoshima [M.] 1985:216, Fig. 52 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Okhotsk Sea, 55°19'N, 142°34'E, depth 178 meters. Holotype: HUMZ 60246. Paratypes: HUMZ 58291-93 (3), 60244 (1), 60248-52 (5), 60254-58 (5), 60278 (1), 61434 (1), 61466 (1). •Valid as Lycodes fulvus Toyoshima 1985 -- (Anderson 1994:118 [ref. 21438], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:114 [ref. 27579], Anderson & Fedorov 2004:25 [ref. 27603], Balushkin et al. 2012:995 [ref. 32449], Parin et al. 2014:411 [ref. 33547]). Current status: Valid as Lycodes fulvus Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

fusca, Santelmoa Matallanas [J.], Corbella [C.] & Møller [P. R.] 2012:xx, Figs. 1-4 [Polar Biology v. 35 (no. 9); ref. 31950] Gerlache Strait, station 24, 64°32'S, 61°97'W, Southern Ocean, depth 1056 meters. Holotype: UAB B03GZS51. Paratypes: UAB B03GSZ10 (1) -31 (1), -49 (1), -50 (1); ZMUC P766589 (1), P766593 (1), P766784 (1), P766788 (1). •Valid as Santelmoa fusca Matallanas, Corbella & Møller 2012. Current status: Valid as Santelmoa fusca Matallanas, Corbella & Møller 2012. Zoarcidae: Lycodinae. Distribution: Southern Ocean. Habitat: marine.

garricki, Pachycara Anderson [M. E.] 1990:9, Fig. 3 [The J.L.B Smith Institute of Ichthyology Special Publication No. 50; ref. 16143] Hikurangi Trough, southwestern Pacific, 42°11'S, 175°11'E to 42°11'S, 175°05'E, depth 2602-2619 meters. Holotype (unique): LACM 43730-1. •Valid as Pachycara garricki Anderson 1990 -- (Anderson 1991:157 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:227 [ref. 22911], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson & Møller 2007:183 [ref. 29401], Balushkin et al. 2011:1027 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448], Møller & Anderson 2015:1432 [ref. 34289]). Current status: Valid as Pachycara garricki Anderson 1990. Zoarcidae: Lycodinae. Distribution: Western South Pacific: New Zealand Plateau. Habitat: marine.

gelatinosum, Melanostigma Günther [A.] 1881:21, Pl. 2 (fig. A) [Proceedings of the Zoological Society of London v. 49, 1881 (pt 1) (art. 1) (for 4 Jan. 1881); ref. 2012] Tilly Bay, Strait of Magellan, Chile, depth 24 fathoms. Holotype (unique): BMNH 1880.7.28.4 [or 28.7]. •Valid as Melanostigma gelatinosum Günther 1881 -- (Anderson 1986:343 [ref. 5633], Nakamura in Nakamura et al. 1986:238 [ref. 14235], Anderson 1988:63 [ref. 7304], Paxton et al. 1989:322 [ref. 12442], Pequeño 1989:49 [ref. 14125], Paulin et al. 1989:211 [ref. 24556], Anderson 1990:4 [ref. 16143], Anderson 1990:270 [ref. 21695], Miller 1993:642 [ref. 21297], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:8 [ref. 27603], Duhamel et al. 2005:322 [ref. 32059], Paxton et al. 2006:964 [ref. 28994], Anderson & Møller 2007:183 [ref. 29401], Balushkin et al. 2011:970 [ref. 32449], Møller & Anderson 2015:1428 [ref. 34289], Balushkin & Moganova 2017:131 [ref. 35279], Balushkin & Moganova 2018:136 [ref. 35877], Balushkin & Orlovskaya 2019:136 [ref. 36664], Balushkin 2019:287 [ref. 36807], Orlovskaya & Balushkin 2019:505 [ref. 37050]). Current status: Valid as Melanostigma gelatinosum Günther 1881. Zoarcidae: Gymnelinae. Distribution: Pacific coast of South America. Habitat: marine.

gigantea, Bilabria Anderson [M. E.] & Imamura [H.] 2008:60, Figs. 1-4 [Bulletin of the National Museum of Nature and Science (Ser. A) Supplement No. 2; ref. 29573] Off Kushiro, Hokkaido, Japan, 42°43.0'N, 144°30.2'E, depth 150-156 meters. Holotype: HUMZ 194804. Paratypes: HUMZ 36833 (1, c&s), 36834 (1), 36843 (1), 45746 (1), 45750 (1), 75984-85 (2), 87596 (1), 88101 (1), 92655 (1), 92658-59 (2), 92661 (1), 92915 (1), 92965 (1), 98034 (1), 98328 (1), 150395 (1); NSMT-P 76021 [ex HUMZ 92654] (1), 76022 [ex HUMZ 92657] (1). •Valid as Bilabria gigantea Anderson & Imamura 2008 -- (Balushkin et al. 2011:958 [ref. 32449], Parin et al. 2014:383 [ref. 33547]). Current status: Valid as Bilabria gigantea Anderson & Imamura 2008. Zoarcidae: Gymnelinae. Distribution: From off Cape Erimo to Sarufutsu, Pacific and Okhotsk coasts of Hokkaido, Japan. Habitat: marine.

gillii, Zoarces Jordan [D. S.] & Starks [E. C.] 1905:212, Fig. 11 [Proceedings of the United States National Museum v. 28 (no. 1391); ref. 2530] Pusan, South Korea. Holotype (unique): USNM 45355. Type catalog: Springer & Anderson 1997:18 [ref. 22953]. As gilli in caption for Fig. 11. •Valid as Zoarces gillii Jordan & Starks 1905 -- (Lindberg & Krasyukova 1975:133 [ref. 7348], Toyoshima in Masuda et al. 1984:305 [ref. 6441] as gilii, Anderson 1994:120 [ref. 21438], Nakabo 2000:1028 [ref. 25182] as gilli, Nakabo 2002:1028 [ref. 26193] as gilli, Youn 2002:375, 640 [ref. 26218], Anderson & Fedorov 2004:2 [ref. 27603], Parin et al. 2005:440 [ref. 28290], Kim et al. 2006:278 [ref. 29191], Chereshev et al. 2007:555 [ref. 29371], Voskoboinikova et al. 2010:704 [ref. 31095], Balushkin et al. 2011:956 [ref. 32449], Parin et al. 2014:430 [ref. 33547]). Current status: Valid as Zoarces gillii Jordan & Starks 1905. Zoarcidae: Zoarcinae. Distribution: Western North Pacific. Habitat: marine.

goni, Pachycara Anderson [M. E.] 1991:155, Figs. 3-5 [Cybium v. 15 (no. 2); ref. 20019] Weddell Sea, off Kapp Norvegia, Queen Maud Land, Antarctica, 71°08.8'S, 13°48.1'W to 71°09.4'S, 13°49.1'W, depth 2025-2037 meters. Holotype (unique): MNHN 1990-0644. •Valid as Pachycara goni Anderson 1991 -- (Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:226 [ref. 22911], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Anderson 2006:12 [ref. 28576], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Pachycara goni Anderson 1991. Zoarcidae: Lycodinae. Distribution: Southern Ocean: Weddell Sea, Antarctica. Habitat: marine.

gosztonyii, Dieidolycus Anderson [M. E.] & Pequeño [G.] 1998:2, Fig. 1 [The J.L.B Smith Institute of Ichthyology Special Publication No. 61; ref. 23482] Off Isla Nueva, Chile, 55°31.6'S, 65°56.8'W to 55°30.7'S, 65°58.8'W, depth 2165-2008 meters. Holotype (unique): MNHNC 7089. •Valid as Dieidolycus gosztonyii Anderson & Pequeño 1998 -- (Anderson & Fedorov 2004:13 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Dieidolycus gosztonyii Anderson & Pequeño 1998. Zoarcidae: Lycodinae. Distribution: Western South Atlantic: Chile. Habitat: marine.

gracilis, Gymnelus Chernova [N. V.] 2000:13 [8], Figs. 2c, 5 [Voprosy Ikhtiologii v. 40 (no. 1); ref. 24885] Pacific coast of Kamchatka near the Cape of Africa, depth 105 meters. Holotype: ZIN 51565. Paratypes: ZIN 51566 (5). Type catalog: Balushkin et al 2011:964 [ref. 32499], Balushkin et al. 2012:160 [ref. 33667]. On p. 8 of English translation. •Valid as Gymnelus gracilis Chernova 2000 -- (Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:5 [ref. 27603], Balushkin et al.2011:964 [ref. 32499], Parin et al. 2014:392 [ref. 33547]). •Synonym of Gymnelus viridis (Fabricius 1780) -- (Mecklenburg & Anderson 2015:278 [ref. 33742]). Current status: Synonym of Gymnelus viridis (Fabricius 1780). Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Kamchatka (if valid). Habitat: marine.

gracilis, Lycodes Sars [M.] 1867:40, 45, Pl. 1 (figs. 1-3) [Forhandlinger i Videnskabs-selskabet i Christiania (for 1866); ref. 18657] Drøbaksundet, Oslo Fjord, Norway, depth 94-113 meters. Holotype: ZMUO J4525. Type catalog: Pethon 1969:11 [ref. 19268]. •Synonym of Lycodes vahlii Reinhardt 1831, but a valid subspecies -- (Andriashev 1973:542 [ref. 9214] as vahli, Andriashev in Whitehead et al. 1986:1146 [ref. 13677] as vahli, Saito & Okamura in Okamura et al. 1995:201 [ref. 22531]). •Synonym of Lycodes vahlii Reinhardt 1831 -- (Anderson 1994:119 [ref. 21438]). •Valid as Lycodes gracilis Sars 1867 -- (Carl 2002:75 [ref. 26925], Anderson & Fedorov 2004:25 [ref. 27603], Møller et al. 2010:61 [ref. 31169], Mecklenburg 2011:124 [ref. 31212] as vahli, Balushkin et al. 2011:995 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:411 [ref. 33547], Mecklenburg et al. 2018:328 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes gracilis Sars 1867. Zoarcidae: Lycodinae. Distribution: Arctic and North Atlantic. Habitat: marine.

grigorjewi, Nemalycodes Herzenstein [S. M.] 1896:14 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 1 (no. 1); ref. 2151] Malye Karmakuly, southwestern Novaya Zemlya. Holotype (unique): ZIN 9688 (in parts, poor condition). Type catalog: Balushkin et al. 2011:966 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. •Synonym of Gymnelus viridis (Fabricius 1780) -- (Andriashev 1973:541 [ref. 9214]). •Doubtful status, genus Gymnelus -- (Anderson 1982:55 [ref. 5520], Balushkin et al. 2011:966 [ref. 32449]). •Possibly the same as Gymnelus knipowitschi Chernova 1999 -- (Chernova 1999:9 [ref. 24863]). •Nomen dubium in Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603], Mecklenburg & Anderson 2015:284 [ref. 33742]). Zoarcidae: Gymnelinae. Habitat: marine.

gronovii, Zoarces Valenciennes [A.] in Cuvier & Valenciennes 1836:469 [Histoire naturelle des poissons v. 11; ref. 1005] America. No types known. Type catalog: Bauchot 1967:63 [ref. 20734]. Based on Gronow 1763, no. 266. •Synonym of Zoarces americanus (Bloch & Schneider 1801) -- (Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:2 [ref. 27603]). Current status: Synonym of Zoarces americanus (Bloch & Schneider 1801). Zoarcidae: Zoarcinae. Habitat: marine.

grossidens, Lycodapus Gilbert [C. H.] 1915:373 [Proceedings of the United States National Museum v. 48 (no. 2075); ref. 1632] Bering Sea, 57°18'N, 171°18'W, Albatross station 3483, depth 56 fathoms. Holotype: USNM 75824. Paratypes: CAS-SU 25634 (2), USNM 135596 (3). Type catalog: Böhlke 1953:102 [ref. 12291], Springer & Anderson 1997:18 [ref. 22953]. •Synonym in part [paratypes] of Lycodapus mandibularis Gilbert 1915 -- (Peden & Anderson 1978:1938 [ref. 8895], Mecklenburg et al. 2002:737 [ref. 25968]). •Synonym of Lycodapus fierasfer Gilbert 1890 -- (Peden & Anderson 1978:1953 [ref. 8895] on holotype, Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:739 [ref. 25968], Anderson & Fedorov 2004:22 [ref. 27603]). Current status: Synonym of Lycodapus fierasfer Gilbert 1890. Zoarcidae: Lycodinae. Habitat: marine.

guentheri, Gymnelopsis ocellatus Popov [A. M.] in Schmidt 1950:125 [112] [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] Near Ayan, Okhotsk Sea, 57°47'N, 141°42'E, depth 150 meters. Holotype (unique): ZIN 23949. Type catalog: Balushkin et al 2011:961 [ref. 32499], Balushkin et al. 2012:162 [ref. 33667]. Originally güntheri in English translation. •Synonym of Gymnelopsis ocellatus Soldatov 1922 -- (Lindberg & Krasyukova 1975:178 [ref. 7348]). •Synonym of Gymnelopsis ocellata Soldatov 1922 -- (Anderson 1982:53 [ref. 5520], Anderson & Fedorov 2004:4 [ref. 27603], Balushkin et al.2011:961 [ref. 32499]). Current status: Synonym of Gymnelopsis ocellata Soldatov 1922. Zoarcidae: Gymnelinae. Habitat: marine.

gymninium, Pachycara Anderson [M. E.] & Peden [A. E.] 1988:88, Fig. 5 [Proceedings of the California Academy of Sciences (Series 4) v. 46 (no. 3); ref. 7215] West of Tasu Sound, Queen Charlotte Islands, British Columbia, 52°38.0'N, 132°05.8'W, depth 2744 meters. Holotype: USNM 280121. Paratypes: BCPM 979-11365 (2), 979-11369 (6), 980-100 (1), 980-121 (1); CAS 60524-25 (2); NMC 86-0444 (1); OSUO 2558 (1); SIO 66-69 (3), 67-121 (36), 68-120 (6), 70-12 (3); USNM 221113 [ex BCPM 979-11369] (1). Type catalog: Springer & Anderson 1997:18 [ref. 22953]. •Valid as Pachycara gymninium Anderson & Peden 1988 -- (Anderson 1989:234 [ref. 13487], Anderson 1991:158 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:227 [ref. 22911], Mecklenburg et al. 2002:728 [ref. 25968], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson 2012:41 [ref. 32270], Balushkin et al. 2011:1027 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448], Anderson et al. 2016:76 [ref. 34148]). Current status: Valid as Pachycara gymninium Anderson & Peden 1988. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

hacheyi, Lycodes reticulatus Vladykov [V. D.] 1933:25 [Contributions to Canadian biology and fisheries: being studies from the biological stations of Canada (Ser. A, General) v. 8 (no. 2); ref. 23244] Hudson Bay, Canada. Syntypes: (2) whereabouts unknown. Also in Vladykov & Tremblay 1936:37, Pl. 6 (fig. 13) [ref. 11811]. •Synonym of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Habitat: marine.

hadrogeneia, Lycenchelys Anderson [M. E.] 1995:74, Fig. 5 [Proceedings of the California Academy of Sciences v. 49 (no. 2); ref. 21928] Off Gulf of Guayaquil, Ecuador, 2°25'S, 81°10'W, depth 700-1000 meters. Holotype: CAS 55589. Paratypes: CAS 81540 (4). •Valid as Lycenchelys hadrogeneia Anderson 1995 -- (Anderson & Fedorov 2004:17 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys hadrogeneia Anderson 1995. Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

heinemanni, Lycodes Soldatov [V. K.] 1916:215, Fig. (p. 215) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 21; ref. 15428] Northern Okhotsk Sea, 58°17'N, 143°15'E, depth 60 meters. Syntypes: ZIN 19169 (3). Type catalog: Balushkin et al. 2011:997 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. •Valid as Lycodes heinemanni Soldatov 1916 -- (Toyoshima 1985:202 [ref. 5722], Anderson 1994:118 [ref. 21438], Fedorov et al. 2003:114 [ref. 27579] dated 1917, Anderson & Fedorov 2004:25 [ref. 27603], Balushkin et al. 2011:997 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:412 [ref. 33547]). Current status: Valid as Lycodes heinemanni Soldatov 1916. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

hemifasciatus, Gymnelis Andriashev [A. P.] 1937:345, Pl. 2 (figs. 23-24) [Issled. Morey SSSR, Leningrad v. 25; ref. 20242] Northern Bering Sea, between Chukchi Peninsula and St. Lawrence Island, 64°15.5'N, 172°27.0'W, depth 55 meters. Holotype: ZIN 24944. Paratypes: (1) not at ZIN. Type catalog: Balushkin et al. 2011:964 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. Original genus should have been Gymnelus. •Valid as Gymnelus hemifasciatus Andriashev 1937 -- (Anderson 1982:38 [ref. 5520], McAllister 1990:165 [ref. 14674], Anderson 1994:117 [ref. 21438], Chernova 1999:11 [ref. 24863], Chernova 2000:8 [ref. 24885], Sheiko & Fedorov 2000:35 [ref. 25650], Mecklenburg et al. 2002:689 [ref. 25968], Anderson & Fedorov 2004:5-6 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Balushkin et al.2011:964 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:392 [ref. 33547], Mecklenburg & Anderson 2015:263 [ref. 33742], Mecklenburg & Steinke 2015:165 [ref. 34033], Mecklenburg et al. 2016:153 [ref. 34440], Kells et al. 2016:270 [ref. 35888], Mecklenburg et al. 2018:307 [ref. 36370] see remarks). Current status: Valid as Gymnelus hemifasciatus Andriashev 1937. Zoarcidae: Gymnelinae. Distribution: Arctic Ocean and North Pacific Ocean. Habitat: marine.

hippopotamus, Lycenchelys Schmidt [P. Yu.] 1950:106 [93], Pl. 9 (fig. 1) [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] East coast of Sakhalin, central Okhotsk Sea, 53°09.5'N, 149°52'E, depth 1000-1150 meters. Lectotype: ZIN 24826. Paralectotypes: ZIN 24826 (3). Type catalog: Balushkin et al. 2011:980 [ref. 32449] dating to 1833 with lectotype and 3 paralectotypes, Balushkin et al. 2012:160 [ref. 33667] dating to 1833 with lectotype and 3 paralectotypes. On p. 93 of English translation. •Valid as Lycenchelys hippopotamus Schmidt 1950 -- (Lindberg & Krasyukova 1975:164 [ref. 7348], Toyoshima in Amaoka et al. 1983:267, 332 [ref. 8379], Toyoshima in Masuda et al. 1984:306 [ref. 6441], Toyoshima 1985:169 [ref. 5722], Anderson 1994:117 [ref. 21438], Anderson 1995:69 [ref. 21928], Nakabo 2000:1032 [ref. 25182], Sheiko & Fedorov 2000:35 [ref. 25650], Mecklenburg et al. 2002:701 [ref. 25968], Nakabo 2002:1032 [ref. 26193], Fedorov et al. 2003:112 [ref. 27579] dated 1835, Anderson & Fedorov 2004:17 [ref. 27603], Shinohara & Anderson 2007:64 [ref. 29111], Balushkin et al. 2011:980 [ref. 32449], Parin et al. 2014:399 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys hippopotamus Schmidt 1950. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

hollandi, Allolepis Jordan [D. S.] & Hubbs [C. L.] 1925:323, Pl. 12 (fig. 2) [Memoirs of the Carnegie Museum v. 10 (no. 2); ref. 2486] Near Fukui, Sea of Japan, Japan. Holotype: FMNH 58842 [ex CM 7952]. Paratypes: CAS-SU 23535 (1). Type catalog: Henn 1928:98 [ref. 12305], Böhlke 1953:101 [ref. 12291], Ibarra & Stewart 1987:7 [ref. 12367]. •Valid as Allolepis hollandi Jordan & Hubbs 1925 -- (Lindberg & Krasyukova 1975:181 [ref. 7348], Toyoshima in Amaoka et al. 1983:334 [ref. 8379], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Sokolovskaya et al. 1998:12 [ref. 24670], Sheiko & Fedorov 2000:34 [ref. 25650], Youn 2002:375, 639 [ref. 26218], Fedorov et al. 2003:109 [ref. 27579], Balushkin et al. 2011:972 [ref. 32449]). •Valid as Bothrocara hollandi (Jordan & Hubbs 1925) -- (Anderson 1994:116 [ref. 21438], Nakabo 2000:1040 [ref. 25182], Nakabo 2002:1040 [ref. 26193], Anderson & Fedorov 2004:10 [ref. 27603], Kim et al. 2006:277 [ref. 29191], Anderson et al. 2009:179 [ref. 30175], Shinohara et al. 2011:50 [ref. 31715], Shinohara et al. 2014:263 [ref. 33330], Parin et al. 2014:384 [ref. 33547], Tohkairin et al. 2015:11 [ref. 35331]). Current status: Valid as Bothrocara hollandi (Jordan & Hubbs 1925). Zoarcidae: Lycodinae. Distribution: North Pacific: Sea of Japan and Okhotsk Sea east to eastern Bering Sea. Habitat: marine.

hubbsi, Lycodes Matsubara [K.] 1955:776 [Fish morphology and hierarchy Part 1; ref. 18463] Off Kinkazan and off Kushiro, Japan. No types known. •Status uncertain -- (Anderson 1994:118 [ref. 21438]). •Valid as Lycodes hubbsi Matsubara 1955 -- (Toyoshima in Amaoka et al. 1983:139, 208, 261, 330 [ref. 8379], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:226 [ref. 5722], Nakabo 2000:1029 [ref. 25182], Sheiko & Fedorov 2000:36 [ref. 25650], Nakabo 2002:1029 [ref. 26193], Anderson & Fedorov 2004:25 [ref. 27603], Shinohara & Shirai 2005:171 [ref. 28232], Balushkin et al. 2011:997 [ref. 32449], Parin et al. 2014:412 [ref. 33547]). Current status: Valid as Lycodes hubbsi Matsubara 1955. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

hubbsi, Lycozoarces Popov [A. M.] 1935:303, Fig. [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (no. 6-7); ref. 3546] Okhotsk Sea, 54°14'N, 143°45'E. Holotype (unique): ZIN 26566. Type catalog: Balushkin et al. 2011:951 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. •Valid as Lycozoarces hubbsi Popov 1935 -- (Lindberg & Krasyukova 1975:125 [ref. 7348]). •Valid, with Lycozoarces regani as unavailable -- (Anderson 1994:24, 119 [ref. 21438]). •Should be treated as a synonym of Lycozoarces regani Popov 1933. •Synonym of Lycozoarces regani Popov 1933 -- (Sheiko & Fedorov 2000:37 [ref. 25650], Anderson & Fedorov 2004:2 [ref. 27603], Balushkin et al. 2011:951 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330]). Current status: Synonym of Lycozoarces regani Popov 1935. Zoarcidae: Lycozoarcinae. Habitat: marine.

humilis, Gymnelopsis Nazarkin [M. V.] & Chernova [N. V.] 2003:603 [578], Figs. 1a-b, 2a-b [Voprosy Ikhtiologii v. 43 (no. 5); ref. 27204] Northern Okhotsk Sea, 57°30.6'N, 146°00.2'E, R/V *Zodiak*, bottom trawler survey-2000, trawl 4, 153 meters. Holotype: ZIN 52831. Paratypes: ZIN 52832 (2). Type catalog: Balushkin 2011:961 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667] as ZIN 53831 and ZIN 52832. On p. 578 of English translation. •Valid as Gymnelopsis humilis Nazarkin & Chernova 2003 -- (Anderson & Fedorov 2004:4 [ref. 27603], Balushkin et al. 2011:961 [ref. 32449], Parin et al. 2014:390 [ref. 33547]). Current status: Valid as Gymnelopsis humilis Nazarkin & Chernova 2003. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

hureaui, Apodolycus Andriashev [A. P.] 1979:30 [482], Figs. 1-2 [Biologiya Morya (Vladivostok) v. 1979 (no. 6); ref. 120] Kerguelen Plateau, 49°36'S, 71°28"E, depth 610 meters. Holotype (unique): ZIN 44333. Paratypes: ZIN 44334 (1). Type catalog: Balushkin et al. 2011:973 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. On p. 482 of English translation. •Valid as Lycenchelys hureaui (Andriashevi 1979) -- (Anderson 1988:94 [ref. 7304], Anderson 1990:261 [ref. 21695], Miller 1993:666 [ref. 21297], Anderson 1994:117 [ref. 21438], Anderson & Balanov 2000:1056 [ref. 25023], Anderson & Fedorov 2004:17 [ref. 27603], Duhamel et al. 2005:318 [ref. 32059]). •Valid as Apodolycus hureaui Andriachev 1979 -- (Balushkin et al. 2011:973 [ref. 32449]). Current status: Valid as Apodolycus hureaui Andriashev 1979. Zoarcidae: Lycodinae. Distribution: Souther Indian Ocean: Kerguelen Plateau. Habitat: marine.

imamurai, Lycenchelys Anderson [M. E.] 2006:184, Figs. 1-2 [Species Diversity v. 11 (no. 3); ref. 29415] Southwest of Punta Negra, 6°39.4'S, 81°04.6'W to 6°41.3'S, 81°04.9'W, off Peru, depth 675-714 meters. Holotype: HUMZ 175168. Paratypes: (many) all at HUMZ. •Valid as Lycenchelys imamurai Anderson 2006. Current status: Valid as Lycenchelys imamurai Anderson 2006. Zoarcidae: Lycodinae. Distribution: Southeastern Pacific: off Peru. Habitat: marine.

incisus, Lycodes Garman [S.] 1899:135, Pl. 30 (fig. 2) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Panama Bay, off Cape Mala, 7°06'15"N, 80°34'W, Albatross station 3353, depth 695 fathoms. Lectotype: MCZ 28685. Paralectotypes: MCZ 122484 [ex MCZ 28685] (1). Lectotype selected by Anderson 1995:80 [ref. 21928]. •Valid as Lycenchelys incisa (Garman 1899) -- (Anderson 1994:117 [ref. 21438], Anderson 1995:77 [ref. 21928], Pequeño 1997:83 [ref. 23536], Anderson & Fedorov 2004:17 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys incisa (Garman 1899). Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

inexpectatum, Melanostigma Parin [N. V.] 1977:63, Fig. 2 [Trudy Instituta Okeanologii Imeni P.P. Shirshova v. 107; ref. 7274] Western equatorial Pacific, 1°15'S, 142°46'E, estimated depth 1000 meters (3300 meters wire out). Holotype (unique): ZIN 42640. Type catalog: Balushkin et al. 2011:970 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. •Valid as Melanostigma inexpectatum Parin 1977 -- (Anderson 1990:2 [ref. 16143], Anderson 1994:120 [ref. 21438], Anderson 2001:3494 [ref. 26279], Anderson & Fedorov 2004:8 [ref. 27603], Anderson & Møller 2007:183 [ref. 29401], Balushkin et al. 2011:970 [ref. 32449], Møller & Anderson 2015:1429 [ref. 34289], Balushkin & Moganova 2017:131 [ref. 35279], Balushkin & Moganova 2018:136 [ref. 35877], Balushkin & Orlovskaya 2019:135 [ref. 36664], Balushkin 2019:287 [ref. 36807]). Current status: Valid as Melanostigma inexpectatum Parin 1977. Zoarcidae: Gymnelinae. Distribution: Southwestern Pacific. Habitat: marine.

ingolfianus, Lycenchelys Jensen [A. S.] 1902:210 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Davis Straits, 64°54'N, 55°10'W, depth 393 fathoms. Holotype (unique): ZMUC 767 [not ZMUC 20]. Type catalog: Nielsen 1974:76 [ref. 9588]. •Status uncertain, possibly a synonym of Lycenchelys paxillus (Goode & Bean 1879) -- (Anderson 1994:117 [ref. 21438]). •Valid as Lycenchelys ingolfianus Jensen 1902 -- (Koyanagi in Okamura et al. 1995:190 [ref. 22531]). •Synonym of Lycenchelys paxillus (Goode & Bean 1879) -- (Møller 1999:323 [ref. 24052], Anderson & Fedorov 2004:19 [ref. 27603], Møller et al. 2010:59 [ref. 31169]). Current status: Synonym of Lycenchelys paxillus (Goode & Bean 1879). Zoarcidae: Lycodinae. Habitat: marine.

insignis, Platea Steindachner [F.] 1898:323, Pl. 20 (figs. 12-12b) [Zoologische Jahrbücher. Jena, Supplementband v. 4; ref. 4238] Cape Espíritu Santo, Tierra del Fuego, Chile. Holotype (unique): ZMB 15694. •Valid as Dadyanos insignis (Steindachner 1898) -- (Gosztonyi 1977:210 [ref. 6103], Pequeño 1989:48 [ref. 14125], Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:12 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Dadyanos insignis (Steindachner 1898). Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

intermedia, Lycodes rossi forma Knipowitsch [N. M.] 1906:28 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Not available, infrasubspecific; publication in an available way as a species or subspecies not investigated [Knipowitsch used v. [variety] and f. [forma] -- in his work, the former is considered subspecific the latter infrasubspecific; see Article 45(g)]. From Barents Sea, Russia. •In the synonymy of Lycodes rossii Malmgren 1865 -- (Anderson 1994:119 [ref. 21438] as rossi, Anderson & Fedorov 2004:48 [ref. 27603], Parin et al. 2014:420 [ref. 33547]). Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae.

japonica, Gengea Katayama [M.] 1943:101, Fig. 1 [Annotationes Zoologicae Japonenses v. 22 (no. 2); ref. 2567] Moroyose, Hyogo-ken, Japan. Holotype (unique): whereabouts unknown. •Valid as Gengea japonica Katayama 1943 -- (Lindberg & Krasyukova 1975:180 [ref. 7348]). •Synonym of Gymnelopsis ochotensis (Popov 1931) -- (Anderson 1982:51 [ref. 5520], Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:4 [ref. 27603]). •Valid as Derjuginia japonica (Katayama 1943) -- (Toyoshima in Masuda et al. 1984:306 [ref. 6441], Nakabo 2000:1042 [ref. 25182], Nakabo 2002:1042 [ref. 26193], Shinohara et al. 2011:51 [ref. 31715], Balushkin 2011:960 [ref. 32449], Shinohara et al. 2014:264 [ref. 33330]). •Valid as Gymnelopsis japonicus (Katayama 1943) -- (Nazarkin & Chernova 2003:605 [ref. 27204], Parin et al. 2014:390 [ref. 33547]). Current status: Valid as Gymnelopsis japonica (Katayama 1943). Zoarcidae: Gymnelinae. Distribution: Western North Pacific. Habitat: marine.

japonica, Lycogramma Oshima [M.] 1957:7 [Japanese Journal of Ichthyology v. 6 (nos 1-2); ref. 12161] Japan. No types known. •Status uncertain, holotype lost, may have been a Bothrocara -- (Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:48 [ref. 27603], Anderson et al. 2009:172 [ref. 30175]). Zoarcidae: Lycodinae. Habitat: marine.

japonicum, Melanostigma Balushkin [A. V.] 2019:284, Figs. 1-2 [Voprosy Ikhtiologii v. 59 (no. 3); ref. 36807] Tosa Bay, southern Japan, 32°59'19"N, 133°35'52"E, depth 700 - 720 meters. Holotype (unique): BSKU 44840. •Valid as Melanostigma japonicum Balushkin 2019. Current status: Valid as Melanostigma japonicum Balushkin 2019. Zoarcidae: Gymnelinae. Distribution: Pacific coast of Japan. Habitat: marine.

japonicus, Lycodes Matsubara [K.] & Iwai [T.] 1951:368, Figs. 1-3 [Japanese Journal of Ichthyology v. 1 (no. 6); ref. 12785] Near the sea-coast of Uozu, southeast of Toyama Bay, Japan, depth 303 meters. Holotype: FAKU 13117 (not seen in 1997). Paratypes: (not seen in 1997) FAKU 12773-87 (15), 13115-16 (2), 13118-38 (21), 13150 (1). •Valid as Lycodes japonicus Matsubara & Iwai 1951 -- (Lindberg & Krasyukova 1975:143 [ref. 7348], Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:191 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1037 [ref. 25182], Mecklenburg et al. 2002:714 [ref. 25968], Nakabo 2002:1037 [ref. 26193], Anderson & Fedorov 2004:25 [ref. 27603], Kim et al. 2006:274 [ref. 29191], Nazakin & Fedorov 2007:705 [ref. 29424], Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:998 [ref. 32449], Shinohara et al. 2014:265 [ref. 33330], Parin et al. 2014:412 [ref. 33547]). Current status: Valid as Lycodes japonicus Matsubara & Iwai 1951. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Sea of Japan, Kuril Islands and Korea. Habitat: marine.

jenseni, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] 1935:243, Figs. 1-2 [Zoologischer Anzeiger v. 112 (nos 9-10); ref. 15744] Okhotsk Sea, east of Sakhalin Island, 50°03'N, 144°08'E. Holotype: ZIN 24836. Paratypes: ZIN 36937 (1). Type catalog: Balushkin et al. 2011:997 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. •Valid as Lycodes jenseni Taranetz & Andriashev 1935 -- (Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:211 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1035 [ref. 25182], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:36 [ref. 25650], Nakabo 2002:1035 [ref. 26193], Anderson & Fedorov 2004:25 [ref. 27603], Nazarkin 2010:9 [ref. 30733], Balushkin et al. 2011:998 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:413 [ref. 33547]). Current status: Valid as Lycodes jenseni Taranetz & Andriashev 1935. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

jordani, Lycodes Evermann [B. W.] & Goldsborough [E. L.] 1907:343, Fig. 120 [Bulletin of the Bureau of Fisheries v. 26 (Doc. 624) (for 1906); ref. 6532] Off Oregon, U.S.A., 43°01'00"N, 125°30'00"W, Albatross station 3788, depth 1946 meters. Holotype: USNM 57828. Paratypes: CAS-SU 20014 (1). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:18-19 [ref. 22953]. •Valid as Lycenchelys jordani (Evermann & Goldsborough 1907) -- (McAllister 1990:157 [ref. 14674], Anderson 1994:117 [ref. 21438], Anderson 1995:80 [ref. 21928], Mecklenburg et al. 2002:694 [ref. 25968], Anderson & Fedorov 2004:17 [ref. 27603], Balushkin et al. 2011:980 [ref. 32449], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys jordani (Evermann & Goldsborough 1907). Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

jordaniana, Davidojordania Schmidt [P. J.] (ex Popov) 1936:99 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad (sér. 2) v.; ref. 3942] Tatar Strait, northern Sea of Japan, 51°26.5'N, 141°29.5'E, depth 37 meters. Syntypes: ZIN 23946 (2), 23497 (4). Type catalog: Balushkin et al. 2011:959 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. Original genus should have been Davidijordania. •Valid as Davidojordania jordaniana Schmidt 1936 -- (Lindberg & Krasyukova 1975:172 [ref. 7348], Sokolovskaya et al. 1998:12 [ref. 24670]). •Valid as Davidijordania jordaniana Schmidt 1936 -- (Toyoshima in Masuda et al. 1984:305 [ref. 6441], Anderson 1994:116 [ref. 21438], Nakabo 2000:1036 [ref. 25182], Nakabo 2002:1036 [ref. 26193], Anderson & Fedorov 2004:3 [ref. 27603], Anderson & Imamura 2008:65 [ref. 29573], Shinohara & Kim 2009:232 [ref. 31427], Balushkin et al. 2011:959 [ref. 32449], Parin et al. 2014:388 [ref. 33547]). Current status: Valid as Davidijordania jordaniana Schmidt 1936. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Sea of Japan and Okhotsk Sea. Habitat: marine.

jugoricus, Lycodes Knipowitsch [N. M.] 1906:18, Figs. 4-5 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Proliv Yugorsky Shar, strait between Kara Sea and Barents Sea, Novaya Zemlya, Russia, depth 9 meters. Holotype (unique): ZIN 13506. Type catalog: Balushkin et al. 2011:997 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. •Valid as Lycodes jugoricus Knipowitsch 1906 -- (Andriashev 1973:543 [ref. 9214], Andriashev in Whitehead et al. 1986:1139 [ref. 13677], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:706 [ref. 25968], Anderson & Fedorov 2004:25 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Balushkin et al. 2011:998 [ref. 32449], Page et al. 2013:162 [ref. 32708], Dolgov 2013:920 [ref. 33112], Parin et al. 2014:413 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:63, 199 [ref. 36199], Mecklenburg et al. 2016:166 [ref. 34440], Mecklenburg et al. 2018:330 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes jugoricus Knipowitsch 1906. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and Canada. Habitat: marine.

karenae, Pachycara Anderson [M. E.] 2012:41, Fig. 1-2 [Zootaxa No. 3559; ref. 32270] Off Monterey Bay, California, USA, 36°33.626'N, 122°2.601'W to 36°35.165'N, 122°44.880'W, depth 3100-2780 meters. Holotype: CAS 233971. Paratypes: CAS 233972 (1), USNM 405384 (1). •Valid as Pachycara karenae Anderson 2012 -- (Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara karenae Anderson 2012. Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

kharini, Melanostigma Balushkin [A. V.] & Moganova [M. V.] 2018:137, Figs. 1, 2c-d, 3-4 [Voprosy Ikhtiologii v. 58 (no. 2); ref. 35877] South Pacific Ocean, Hercules Ridge, north of Antarctic-South Pacific Rise, 53°45'S, 140°12'W, depth 290-600 meters. Holotype: ZIN 56163. Paratypes: ZIN 56164 (2). Plus nontype specimens. •Valid as Melanostigma kharini Balushkin & Moganova 2018 -- (Balushkin & Orlovskaya 2019:136 [ref. 36664], Balushkin 2019:287 [ref. 36807]). Current status: Valid as Melanostigma kharini Balushkin & Moganova 2018. Zoarcidae: Gymnelinae. Distribution: South Pacific Ocean: Hercules Ridge. Habitat: marine.

knipowitschi, Gymnelus Chernova [N. V.] 1999:5 [1], Figs. 1-3 [Voprosy Ikhtiologii v. 39 (no. 1); ref. 24863] Bay of Kazarinov, Novaya Zemlya, Kostin Strait, depth 9-17.5 meters. Holotype: ZIN 14143. Paratypes: ZIN 14153-54 (1, 2), 14157 (3), 32077 (2), 32763 (1), 34106 (1), 35641 (3). Type catalog: Balushkin et al 2011:964 [ref. 32499], Balushkin et al. 2012:160 [ref. 33667]. On p. 1 of English translation. •Valid as Gymnelus knipowitschi Chernova 1999 -- (Chernova 1999:306 [ref. 24868], Chernova 2000:8 [ref. 24885], Sheiko & Fedorov 2000:35 [ref. 25650], Balushkin et al.2011:964 [ref. 32499], Afonso et al. 2016:61, 160 [ref. 36199]). •Synonym of Gymnelus hemifasciatus Andriashev 1937 -- (Anderson & Fedorov 2004:6 [ref. 27603], Mecklenburg et al. 2011:124, 128 [ref. 31212], Parin et al. 2014:392 [ref. 33547], Mecklenburg & Anderson 2015:277 [ref. 33742], Mecklenburg et al. 2016:154 [ref. 34440]). Current status: Synonym of Gymnelus hemifasciatus Andriashev 1937. Zoarcidae: Gymnelinae. Distribution: Arctic Ocean, including Bering Sea. Habitat: marine.

knipowitschi, Lycodes Popov [A. M.] 1931:138, Pl. 2 (fig. 4); Fig. 1 [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] Okhotsk Sea, 59°31.2'N, 150°29.9'E, depth 40 meters. Holotype: ZIN 30009. Paratypes: ? ZIN 30009 (2 juv.). Type catalog: Balushkin et al. 2011:999 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. •Synonym of Lycodes mucosus Richardson 1855 -- (Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:708 [ref. 25968]). •In part synonym of Lycodes raridens Taranetz & Andriashev 1937 -- (Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:711 [ref. 25968]). •Valid as Lycodes knipowitschi Popov 1931 -- (Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:195 [ref. 5722], Nakabo 2000:1038 [ref. 25182], Sheiko & Fedorov 2000:36 [ref. 25650], Nakabo 2002:1038 [ref. 26193], Fedorov et al. 2003:114 [ref. 27579], Anderson & Fedorov 2004:27 [ref. 27603], Balushkin et al. 2011:999 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:413 [ref. 33547]). Current status: Valid as Lycodes knipowitschi Popov 1931. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

kolthoffi, Lycenchelys Jensen [A. S.] 1904:88, Pl. 10 (fig. 2); Figs. 23-25 [Danish Ingolf-Expedition v. 2 (no. 4); ref. 15186] Greenland. No types known. •Valid as Lycenchelys kolthoffi Jensen 1904 -- (Andriashev 1973:542 [ref. 9214], Andriashev in Whitehead et al. 1986:1133 [ref. 13677], McAllister 1990:157 [ref. 14674], Anderson 1994:117 [ref. 21438], Koyanagi in Okamura et al. 1995:191 [ref. 22531], Møller 1999:327 [ref. 24052], Anderson & Balanov 2000:1060 [ref. 25023], Anderson & Fedorov 2004:17 [ref. 27603], Møller et al. 2010:59 [ref. 31169], Balushkin et al. 2011:980 [ref. 32449], Dolgov 2013:919 [ref. 33112], Parin et al. 2014:399 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:62, 191 [ref. 36199], Mecklenburg et al. 2016:158 [ref. 34440], Mecklenburg et al. 2018:313 [ref. 36370]). Current status: Valid as Lycenchelys kolthoffi Jensen 1904. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

kreffti, Aiakas Gosztonyi [A. E.] 1977:199, Fig. 2 [Archiv für Fischereiwissenschaft v. 27 (no. 3); ref. 6103] Continental slope off Golfo San Jorge, Argentina, 46°05'S, 59°54'W, depth 800 meters. Holotype (unique): ISH 385-1971. Type catalog: Krefft 1978:14 [ref. 9884]. •Valid as Aiakas kreffti Gosztonyi 1977 -- (Anderson & Gosztonyi 1991:4 [ref. 19348], Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:9 [ref. 27603], Balushkin et al. 2011:971 [ref. 32449]). Current status: Valid as Aiakas kreffti Gosztonyi 1977. Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

labradorensis, Lycenchelys Geistdoerfer [P.], Hureau [J.-C.] & Rannou [M.] 1970:452, Fig. 1 [Bulletin du Muséum National d'Histoire Naturelle (Série 2) v. 42 (no. 3); ref. 7739] Labrador Sea, 58°51.6'N, 53°04.3'W, depth 3365 meters. Holotype (unique): MNHN 1970-0031. •Valid as Lycenchelys labradorensis Geistdoerfer, Hureau & Rannou 1970 -- (Scott & Scott 1988:403 [ref. 25518], McAllister 1990:157 [ref. 14674], Koyanagi in Okamura et al. 1995:190 [ref. 22531]). •Synonym of Lycenchelys alba (Vaillant 1888) -- (Andriashev in Whitehead et al. 1986:1133 [ref. 13677] as albus, Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:15 [ref. 27603], Møller et al. 2010:59 [ref. 31169]). Current status: Synonym of Lycenchelys alba (Vaillant 1888). Zoarcidae: Lycodinae. Habitat: marine.

labrosus, Blennius Mitchill [S. L.] 1815:375, Pl. 1 (fig. 7) [Transactions of the Literary and Philosophical Society of New-York v. 1 (art. 5) (for 1814); ref. 13292] New York, U.S.A. No types known. •Synonym of Zoarces americanus (Bloch & Schneider 1801) -- (Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:2 [ref. 27603]). Current status: Synonym of Zoarces americanus (Bloch & Schneider 1801). Zoarcidae: Zoarcinae. Habitat: marine.

lacertinus, Lycenchelys Pavlenko [M. N.] 1910:53, Figs. 10-11 [Trudy Obshchestva Estestvoispytatelej Imperatorskom' Kazanskom' Universiteti, Kazan v. 42 (no. 2); ref. 3393] Peter the Great Bay, Russia, Sea of Japan. Syntypes: (3) whereabouts unknown. Type catalog: Balushkin et al. 2011:959 [ref. 32449]. •Valid as Davidojordania [sic] lacertianus (Pavlenko 1910) -- (Lindberg & Krasyukova 1975:174 [ref. 7348], Sokolovskaya et al. 1998:12 [ref. 24670]). •Valid as Davidijordania lacertina (Pavlenko 1910) -- (Toyoshima in Masuda et al. 1984:305 [ref. 6441], Anderson 1994:116 [ref. 21438], Nakabo 2000:1036 [ref. 25182], Nakabo 2002:1036 [ref. 26193], Anderson & Fedorov 2004:4 [ref. 27603], Anderson & Imamura 2008:65 [ref. 29573], Shinohara & Kim 2009:228 [ref. 31427], Balushkin et al. 2011:959 [ref. 32449], Shinohara et al. 2014:264 [ref. 33330], Parin et al. 2014:388 [ref. 33547]). Current status: Valid as Davidijordania lacertina (Pavlenko 1910). Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Sea of Japan. Habitat: marine.

laevifasciatus, Oidiphorus Torno [A. E.], Tomo [A. P.] & Marschoff [E.] 1977:4, Figs. [Instituto Antártico Argentino, Direccion Nacional del Antartico, Contribucion No. 223; ref. 8869] South Sandwich Islands, southeastern Atlantic, 59°25'S, 26°54'W, depth 1080 meters. Holotype: IAA no. 3. •Valid as Seleniolycus laevifasciatus (Torno, Tomo & Marschoff 1977) -- (Anderson 1988:68 [ref. 7304], Anderson 1990:275 [ref. 21695], Miller 1993:644 [ref. 21297], Anderson 1994:120 [ref. 21438], Pequeño 1997:83 [ref. 23536], Anderson & Fedorov 2004:9 [ref. 27603], Anderson 2006:12 [ref. 28576], Møller & Stewart 2006:61 [ref. 28888], Balushkin et al. 2011:971 [ref. 32449]). Current status: Valid as Seleniolycus laevifasciatus (Torno, Tomo & Marschoff 1977). Zoarcidae: Gymnelinae. Distribution: Southern Ocean. Habitat: marine.

laticinctus, Lycodus Berg [C.] 1895:71, Pl. 1 (fig. 2) [Anales del Museo Nacional de Historia Natural de Buenos Aires v. 4; ref. 261] Mouth of Río Santa Cruz, Santa Cruz Province, Argentina. Holotype (unique): MACN 3003. Type catalog: Braga & Piacentino 1994:102 [ref. 23697]. •Valid as Austrolycus laticinctus (Berg 1895) -- (Gosztonyi 1977:202 [ref. 6103], Nakamura in Nakamura et al. 1986:234 [ref. 14235], Pequeño 1989:48 [ref. 14125], Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:10 [ref. 27603], Mabragaña et al. 2011:table S2 [ref. 31800], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Austrolycus laticinctus (Berg 1895). Zoarcidae: Lycodinae. Distribution: Southwestern Atlantic. Habitat: marine.

latitans, Phucocoetes Jenyns [L.] 1842:168, Pl. 29 (figs. 3, 3a) [The zoology of the voyage of H. M. S. Beagle; ref. 2344] Falkland Islands. Lectotype: BMNH 1917.7.14.67 (?=BMNH 1842.2.12.2). Paralectotypes: BMNH 1917.7.14.68 (1) (?=BMNH 1842.2.12.3). Lectotype selected by Gosztonyi 1977:233 [ref. 6103]. •Valid as Phucocoetes latitans Jenyns 1842 -- (Gosztonyi 1977:233 [ref. 6103], Pequeño 1989:49 [ref. 14125], Lloris & Rucabado 1991:75 [ref. 22530], Anderson 1994:120 [ref. 21438], López et al. 1996:33 [ref. 23681], Anderson & Fedorov 2004:36 [ref. 27603], Balushkin et al. 2011:1021 [ref. 32449]). Current status: Valid as Phucocoetes latitans Jenyns 1842. Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

laurentianus, Lycodes reticulatus Vladykov [V. D.] & Tremblay [J.-L.] 1936:34, Pls. 5 (fig. 11), 6 (fig. 12) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Saint Lawrence River estuary, Quebec, Canada. Syntypes: (8) NMC 60-159 (1). Type catalog: McAllister 1965:9 [ref. 22019] as paratypes. •Synonym of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Habitat: marine.

lavalaei, Lycodes Vladykov [V. D.] & Tremblay [J.-L.] 1936:31, Pls. 4 (fig. 9), 5 (figs. 10-11) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Gulf of Saint Lawrence and Newfoundland, Canada. Syntypes: (11) NMC 60-157 (1), 60-158 (1). Type catalog: McAllister 1965:9 [ref. 22019] as paratypes. Misspelled lavalei in earlier editions of the Catalog of Fishes. •Valid as Lycodes lavalaei Vladykov & Tremblay 1936 -- (Andriashev 1973:544 [ref. 9214], Andriashev in Whitehead et al. 1986:1140 [ref. 13677], Robins & Ray 1986:237 [ref. 23100], Scott & Scott 1988:408 [ref. 25518], McAllister 1990:161 [ref. 14674], Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:26 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Mecklenburg et al. 2011:128 [ref. 31212], Balushkin et al. 2011:999 [ref. 32449], Page et al. 2013:162 [ref. 32708], Afonso et al. 2016:64, 200 [ref. 36199], Mecklenburg et al. 2018:331 [ref. 36370]). Current status: Valid as Lycodes lavalaei Vladykov & Tremblay 1936. Zoarcidae: Lycodinae. Distribution: Northwestern Atlantic and adjacent Arctic Region. Habitat: marine.

lenzeni, Lycenchelys Thiel [R.], Knebelsberger [T.] & Eidus [I.] 2018:46, Figs. 2-3 [Zootaxa 4370 (no. 1); ref. 35743] Southwest of Simushir Island, Kuril Archipelago, Russia, 46°36.465'N, 151°34.615'E, depth 2350 meters. Holotype: ZMH 26255. Paratypes: ZMH. •Valid as Lycenchelys lenzeni Thiel, Knebelsberger & Eidus 2018. Current status: Valid as Lycenchelys lenzeni Thiel, Knebelsberger & Eidus 2018. Zoarcidae: Lycodinae. Distribution: North Pacific: Kuril Archipelago. Habitat: marine.

lepinium, Pachycara Anderson [M. E.] & Peden [A. E.] 1988:91, Fig. 6 [Proceedings of the California Academy of Sciences (Series 4) v. 46 (no. 3); ref. 7215] West of Tasu Sound, Queen Charlotte Islands, British Columbia, 52°38.0'N, 132°05.8'W, depth 2744 meters. Holotype: USNM 280120. Paratypes: BCPM 980-98 (1), 980-121 (1); CAS 60526-30 (1 ea.); LACM 32114-2 (1, missing), 44315-1 (1); NMC 86-0445 (1); OSUO 2563 (1); SIO 59-288 (1), 59-289 (4 not 3), 59-365 (5), 60-475 (1), 65-452 (4), 66-50 (1); USNM 221114 (1). Type catalog: Springer & Anderson 1997:19 [ref. 22953]. •Valid as Pachycara lepinium Anderson & Peden 1988 -- (Anderson 1989:227 [ref. 13487], Anderson 1991:158 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:227 [ref. 22911], Mecklenburg et al. 2002:729 [ref. 25968], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson 2006:189 [ref. 29415], Balushkin et al. 2011:1027 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara lepinium Anderson & Peden 1988. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

leptodermatus, Dieidolycus Anderson [M. E.] 1988:72, Fig. 15 [Antarctic Research Series, Washington DC. v. 47; ref. 7304] South of South Georgia Island, Scotia Sea, 55°01'-10'S, 39°55'-46'W, depth 2886-3040 meters. Holotype: LACM 10772-7. Paratypes: LACM 11062-6 (1), 11351-6 (1). •Valid as Dieidolycus leptodermatus Anderson 1988 -- (Anderson 1990:257 [ref. 21695], Miller 1993:646 [ref. 21297], Anderson 1994:116 [ref. 21438], Anderson & Pequeño 1998:4 [ref. 23482], Anderson & Fedorov 2004:13 [ref. 27603], Anderson 2006:3 [ref. 28576], Struthers & Møller 2009:65 [ref. 30403], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Dieidolycus leptodermatus Anderson 1988. Zoarcidae: Lycodinae. Distribution: Southern Ocean: circum-Antarctic. Habitat: marine.

leptopus, Hadropogonichthys Machida [Y.], Shinohara [G.] & Ohta [S.] 2004:194, Figs. 1-4 [Species Diversity v. 9 (no. 3); ref. 27944] Off Ibaraki Prefecture, Honshu Island, Japan, 36°45.41'N, 141°43.16'E to 36°45.06'N, 141°42.78'E, depth 1408 meters. Holotype: NSMT-P 57227. Paratypes: BSKU 19655 (1), 19848 (1), 19989 (1), 20235 (1), 82229 (1); NSMT-P 49050 (11), 55828 (6), 55914 (2). •Valid as Hadropogonichthys leptopus Machida, Shinohara & Ohta 2004. Current status: Valid as Hadropogonichthys leptopus Machida, Shinohara & Ohta 2004. Zoarcidae: Lycodinae. Distribution: Off Japan. Habitat: marine.

leptus, Lycodapus Peden [A. E.] & Anderson [M. E.] 1981:671, Fig. 5 [Canadian Journal of Zoology v. 59 (no. 4); ref. 5531] Bering Sea, 59°00.3'N, 178°21.6'W, depth 568-590 meters. Holotype: USNM 222660 (emaciated). Paratypes: CAS 45730 (2), 45731 (5 not 4); HUMZ 83839 (1, not found in 1997), 86666 (1), 86673 (1), 86681 (1), 86702 (1), 86704 (1); USNM 221054-55 (24, 1), 221066 (3), 222659 (1), 222661-62 (1, 1). Type catalog: Springer & Anderson 1997:19 [ref. 22953]. •Valid as Lycodapus leptus Peden & Anderson 1981 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:736 [ref. 25968], Anderson & Fedorov 2004:22 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449], Parin et al. 2014:405 [ref. 33547], Parin et al. 2014:405 [ref. 33547]). Current status: Valid as Lycodapus leptus Peden & Anderson 1981. Zoarcidae: Lycodinae. Distribution: North Pacific: Bering Sea. Habitat: marine.

lindbergi, Hadropogonichthys Fedorov [V. V.] 1982:724 [19], Figs. 1-2 [Voprosy Ikhtiologii v. 22 (no. 5); ref. 1311] Kuril Islands, Fourth Kuril Strait, 49°35'N, 155°12'E, depth 600-615 meters. Holotype: ZIN 45321. Paratypes: ZIN 45322 (1). Type catalog: Balushkin et al. 2011:976 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. On p. 19 of English translation. •Valid as Hadropogonichthys lindbergi Fedorov 1982 -- (Toyoshima 1985:142 [ref. 5722], Anderson 1994:117 [ref. 21438], Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:13 [ref. 27603], Machida et al. 2004:193 [ref. 27944], Balushkin et al. 2011:976 [ref. 32449], Parin et al. 2014:395 [ref. 33547]). Current status: Valid as Hadropogonichthys lindbergi Fedorov 1982. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea and Kuril Islands. Habitat: marine.

lindbergi, Lycodes Popov [A. M.] 1931:142, Pl. 2 (fig. 5) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] 58°11.5'N, 148°19.5'N, Okhotsk Sea, depth 141 meters. Holotype (unique): whereabouts unknown. •Synonym of Lycodes semenovi Popov 1931 -- (Balushkin et al. 2011:1011 [ref. 32449]). •Synonym of Lycodes uschakovi Popov 1931 -- (Lindberg & Krasyukova 1975:147 [ref. 7348], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:30 [ref. 27603], Parin et al. 2014:423 [ref. 33547]). Current status: Synonym of Lycodes uschakovi Popov 1931. Zoarcidae: Lycodinae. Habitat: marine.

lonchoura, Lycenchelys Anderson [M. E.] 1995:82, Fig. 8 [Proceedings of the California Academy of Sciences v. 49 (no. 2); ref. 21928] Southeastern Pacific, northwest of Pt. Coles, 17°08.5'S, 72°07.4'W, depth 860 meters. Holotype (unique): CAS 58150. •Valid as Lycenchelys lonchoura Anderson 1995 -- (Anderson & Fedorov 2004:17 [ref. 27603], Shinohara & Anderson 2007:62 [ref. 29111], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys lonchoura Anderson 1995. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific. Habitat: marine.

longirostris, Lycenchelys Toyoshima [M.] 1985:166, Figs. 22-23, 31g [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Bering Sea, 58°33.16'N, 175°05.3'W, depth 895-910 meters. Holotype: HUMZ 81914. Paratypes: HUMZ 83948 (1). •Valid as Lycenchelys longirostris Toyoshima 1985 -- (Sheiko & Fedorov 2000:35 [ref. 25650]). •Synonym of Lycenchelys ratmanovi Andriashev 1955 -- (Anderson 1994:118 [ref. 21438], Anderson 1995:100 [ref. 21928], Anderson & Balanov 2000:1056 [ref. 25023], Mecklenburg et al. 2002:702 [ref. 25968], Anderson & Fedorov 2004:20 [ref. 27603], Parin et al. 2014:402 [ref. 33547]). Current status: Synonym of Lycenchelys ratmanovi Andriashev 1955. Zoarcidae: Lycodinae. Habitat: marine.

luetkenii, Lycodes Collett [R.] 1880:103, Pl. 3 (fig. 25) [The Norwegian North-Atlantic Expedition; ref. 887] 115 kilometers west of Norskøerne, Spitsbergen, North Atlantic, depth 459 fathoms. Holotype (unique): ZMUO J4536. Type catalog: Pethon 1969:12 [ref. 19268]. Original as lütkenii. •Valid as Lycodes luetkenii Collett 1880 -- (Andriashev 1973:544 [ref. 9214] as leutkeni, Andriashev in Whitehead et al. 1986:1140 [ref. 13677] as leutkeni, Anderson 1994:118 [ref. 21438], Saito & Okamura in Okamura et al. 1995:197 [ref. 22531] as luetkeni, Møller & Petersen 1997:290 [ref. 23332] as luetkeni, Anderson & Fedorov 2004:26 [ref. 27603], Møller et al. 2010:61 [ref. 31169], Balushkin et al. 2011:999 [ref. 32449], Page et al. 2013:162 [ref. 32708], Dolgov 2013:920 [ref. 33112] as leutkeni, Parin et al. 2014:413 [ref. 33547], Afonso et al. 2016:64, 201 [ref. 36199], Mecklenburg et al. 2018:333 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes luetkenii Collett 1880. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

lugubris, Lycodes Lütken [C. F.] 1880:315 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1879-1880; ref. 19479] Iceland. Syntypes: ZMUC 10 (1), 11 (skeleton), 13 (1), 14 (1). Type catalog: Nielsen 1974:77 [ref. 9588]. •Synonym of Lycodes vahlii Reinhardt 1831 -- (Andriashev 1973:542 [ref. 9214], Anderson 1994:119 [ref. 21438]). •Synonym of Lycodes gracilis Sars 1867 -- (Carl 2002:75 [ref. 26925], Anderson & Fedorov 2004:25 [ref. 27603]). Current status: Synonym of Lycodes gracilis Sars 1867. Zoarcidae: Lycodinae. Habitat: marine.

lycodon, Lycodapus Gilbert [C. H.] 1915:371, Pl. 21 (fig. 21) [Proceedings of the United States National Museum v. 48 (no. 2075); ref. 1632] Monterey Bay, California, U.S.A., Albatross station 4509, depth 152-286 fathoms. Holotype (unique): USNM 75822. Type catalog: Springer & Anderson 1997:19 [ref. 22953]. •Synonym of Lycodapus mandibularis Gilbert 1915 -- (Peden & Anderson 1978:1938 [ref. 8895], Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:22 [ref. 27603]). Current status: Synonym of Lycodapus mandibularis Gilbert 1915. Zoarcidae: Lycodinae. Habitat: marine.

lyoderma, Taranetzella Andriashev [A. P.] 1952:416, Fig. [Trudy Zoologicheskogo Instituta, Akademii Nauk SSSR v. 12; ref. 6593] Olyutorsk [Olutorsky] Bay, western Bering Sea, 59°55'N, 168°14'E, depth 986-1150 meters. Holotype (unique): ZIN 32813. Type catalog: Balushkin et al. 2011:1022 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. •Valid as Taranetzella lyoderma Andriashev 1952 -- (Toyoshima 1985:145 [ref. 5722], Peden & Jamieson 1988:493 [ref. 12864], McAllister 1990:163 [ref. 14674], Anderson 1994:120 [ref. 21438], Anderson 1995:107 [ref. 21928], Machida & Ohta 1996 [ref. 22117], Nakabo 2000:1044 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:692 [ref. 25968], Nakabo 2002:1044 [ref. 26193], Anderson & Fedorov 2004:37 [ref. 27603], Balushkin et al. 2011:1022 [ref. 32449], Parin et al. 2014:428 [ref. 33547]). Current status: Valid as Taranetzella lyoderma Andriashev 1952. Zoarcidae: Lycodinae. Distribution: North Pacific: Bering Sea and Japan to Mexico. Habitat: marine.

macrocephalus, Lycodes reticulatus var. Jensen [A. S.] 1904:258, Pl. 13 (figs. 2 a, b) [Meddelelser om Grønland v. 29; ref. 20203] Eastern Greenland. Syntypes: (11) whereabouts unknown. •Synonym of Lycodes reticulatus Reinhardt 1835, but a valid subspecies as described -- (Andriashev 1973:545 [ref. 9214], Andriashev in Whitehead et al. 1986:1143 [ref. 13677]). •Synonym of Lycodes reticulatus Reinhardt 1835 -- (Anderson & Fedorov 2004:28 [ref. 27603], Balushkin et al. 2011:1009 [ref. 32449]), •In the synonymy of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Parin et al. 2014:419 [ref. 33547]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Habitat: marine.

macrochir, Lycodes Schmidt [P. J.] in Taranetz 1937:162, 164 [Izvestiia Tikhookeanskogo nauchnogo instituta rybnogo khoziaistva [Bulletins of the Pacific Science Institute] v. 11; ref. 13384] Okhotsk Sea. Syntypes: ZIN 25273 (1), 29092 (1), 29142 (1). Type catalog: Balushkin et al. 2011:999 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667].. •Valid as Lycodes macrochir Schmidt 1937 -- (Toyoshima 1985:205 [ref. 5722], Anderson 1994:118 [ref. 21438], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:114 [ref. 27579], Anderson & Fedorov 2004:26 [ref. 27603], Shinohara & Shirai 2005:171 [ref. 28232], Balushkin et al. 2011:999 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:414 [ref. 33547]). Current status: Valid as Lycodes macrochir Schmidt 1937. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

macrolepis, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] 1935:251, Figs. 6-7 [Zoologischer Anzeiger v. 112 (nos 9-10); ref. 15744] Okhotsk Sea, northwest of St. Jonas Island, 57°02'N, 141°40'E, depth 147 meters. Holotype: ZIN 24837. Paratypes: ZIN 24837 (2 of 3), 29998 (1). Type catalog: Balushkin et al. 2011:1000 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. •Valid as Lycodes macrolepis Taranetz & Andriashev 1935 -- (Lindberg & Krasyukova 1975:153 [ref. 7348], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:204 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Nakabo 2002:1039 [ref. 26193], Fedorov et al. 2003:115 [ref. 27579], Anderson & Fedorov 2004:26 [ref. 27603], Nazarkin 2010:596 [ref. 31019], Balushkin et al. 2011:1000 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Shinohara et al. 2014:265 [ref. 33330]). Current status: Valid as Lycodes macrolepis Taranetz & Andriashev 1935. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk and Japan seas. Habitat: marine.

macrops, Lycodes Günther [A.] 1880:21, Pl. 11 (fig. B) [Report on the shore fishes procured during the voyage of H. M. S. Challenger in the years 1873-1876 v. 1 (pt 6); ref. 2011] Challenger station 309, depth 40-140 fathoms [= Magellan Straits, near Puerto Bueno, Chile, 50°56'S, 74°15'-14'W]. Holotype (unique): BMNH 1879.5.14.48. •Valid as Ophthalmolycus macrops (Günther 1880) -- (Gosztonyi 1977:229 [ref. 6103], Pequeño 1989:49 [ref. 14125], Anderson 1990:8 [ref. 16143], Anderson & Gosztonyi 1991:12 [ref. 19348], Anderson 1992:8 [ref. 18912], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:34 [ref. 27603], Pequeño & Sáez 2007:373 [ref. 36439], Matallanas 2009:55 [ref. 30103], Matallanas 2011:566 [ref. 30839], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Ophthalmolycus macrops (Günther 1880). Zoarcidae: Lycodinae. Distribution: Southeastern Pacific: southern Chile. Habitat: marine.

macropus, Phucocoetes variegatus forma Smitt [F. A.] 1898:44, Pl. 5 (figs. 35, 36, 36a-b) [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 24 (afd. 4) (no. 5); ref. 4147] Off River Grande, Tierra del Fuego, southwestern Atlantic. Syntypes: NRM 11159-60 (1, 2). •Synonym of Austrolycus laticinctus (Berg 1895) -- (Gosztonyi 1977:202 [ref. 6103] as Phycocoetes variegatus macropus). •In part synonym of Iluocoetes fimbriatus Jenyns 1842 -- (Gosztonyi 1977:213 [ref. 6103]). •Synonym in part of Austrolycus laticinctus (Berg 1895) -- (Anderson 1994:116 [ref. 21438]). •Synonym in part of Ilucoetes fimbriatus Jenyns 1842 -- (Anderson 1994:117 [ref. 21438]). •Synonym of Austrolycus laticinctus (Berg 1895) -- (Anderson & Fedorov 2004:10 [ref. 27603]). Current status: Synonym of Austrolycus laticinctus (Berg 1895). Zoarcidae: Lycodinae. Habitat: marine.

maculata, Krusensterniella Andriashev [A. P.] 1938:118 [Vestnik Dal'nevostocnogo Filiala Akademii Nauk SSSR = Bulletin of the Far Eastern Branch of the Academy of Sciences of the USSR v. 32 (no. 5); ref. 19938] Tatar Strait to Furugelm Island, Sea of Japan, 51°04'N, 140°49'E, depth 53-150 meters. Syntypes: ZIN 29989 (2). Type catalog: Balushkin et al. 2011:969 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667].. •Valid as Krusensterniella maculata Andriashev 1938 -- (Lindberg & Krasyukova 1975:129 [ref. 7348], Toyoshima in Masuda et al. 1984:305 [ref. 6441], Anderson 1994:117 [ref. 21438], Sokolovskaya et al. 1998:12 [ref. 24670], Nakabo 2000:1044 [ref. 25182], Nakabo 2002:1044 [ref. 26193], Anderson & Fedorov 2004:7 [ref. 27603], Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:969 [ref. 32449], Shinohara et al. 2014:264 [ref. 33330], Parin et al. 2014:396 [ref. 33547]). Current status: Valid as Krusensterniella maculata Andriashev 1938. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Japan Sea. Habitat: marine.

maculatus, Lycenchelys Toyoshima [M.] 1985:149, Figs. 8-9a, 31a [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Off Onahama, Pacific coast of Fukushima Prefecture, Japan, depth 200-300 meters. Holotype: HUMZ 71361. Paratypes: HUMZ 71217 (1), 71362-64 (3), 71391-92 (2), 71537 (1). •Valid as Lycenchelys maculata Toyoshima 1985 -- (Anderson 1994:117 [ref. 21438], Fedorov 1995:130 [ref. 21958], Nakabo 2000:1035 [ref. 25182], Nakabo 2002:1035 [ref. 26193], Anderson & Fedorov 2004:17 [ref. 27603], Shinohara & Anderson 2007:64 [ref. 29111], Balushkin et al. 2011:980 [ref. 32449], Parin et al. 2014:400 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys maculata Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

maculatus, Lycodes vahlii Vladykov [V. D.] & Tremblay [J.-L.] 1936:27, Pl. 3 (figs. 7-8) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Saint Lawrence River estuary, Quebec, Canada. Syntypes: (219) NMC 60-161 (1), 60-162 (orig. 23, now 21); ROM (1); UBC (1). Type catalog: McAllister 1965:10 [ref. 22019] as paratypes. •Synonym of Lycodes vahlii Reinhardt 1831 -- (Anderson 1994:119 [ref. 21438], Carl 2002:69 [ref. 26925] as vahli, Anderson & Fedorov 2004:31 [ref. 27603], Balushkin et al. 2011:1016 [ref. 32449]). Current status: Synonym of Lycodes vahlii Reinhardt 1831. Zoarcidae: Lycodinae. Habitat: marine.

magellanicus, Letholycus Anderson [M. E.] 1988:275, Fig. 1C [Proceedings of the California Academy of Sciences (Series 4) v. 45 (no. 11); ref. 6334] Argentine slope, 45°59.3'S, 59°57.1'W, depth 580 meters. Holotype: ISH 164-1978. Paratypes: ISH 173-1978 (1). •Valid as Letholycus magellanicus Anderson 1988 -- (Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:14 [ref. 27603]). •Valid as Japonolycodes magellanicus (Anderson 1988) -- (Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Japonolycodes magellanicus (Anderson 1988). Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

makushok, Lycenchelys Fedorov [V. V.] & Andriashev [A. P.] 1993:133 [130], Figs. 1-2 [Voprosy Ikhtiologii v. 33 (no. 1); ref. 20790] Of Kuril Islands, off Itrup Island, 44°39.2'N, 149°02.2'E, depth 800 meters. Holotype (unique): ZIN 42290. Type catalog: Balushkin et al. 2011:981 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. On p. 130 of English translation. •Valid as Lycenchelys makushok Fedorov & Andriashev 1993 -- (Anderson 1994:117 [ref. 21438], Shinohara et al. 1996:180 [ref. 26605], Nakabo 2000:1032 [ref. 25182], Sheiko & Fedorov 2000:35 [ref. 25650], Nakabo 2002:1032 [ref. 26193], Anderson & Fedorov 2004:18 [ref. 27603], Shinohara & Anderson 2007:63 [ref. 29111], Balushkin et al. 2011:281 [ref. 32449], Parin et al. 2014:400 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys makushok Fedorov & Andriashev 1993. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

malvinensis, Lycodonus Gosztonyi [A. E.] 1981:153, Figs. 1-2 [Archiv für Fischereiwissenschaft v. 31 (no. 3); ref. 8596] Southwestern Atlantic, 49°58'S, 55°00'W, Walther Herwig station 219/71, depth 1000 meters. Holotype: ISH 200-1971b. Paratypes: ISH 200-1971a (1), 374-1971 (1). Type catalog: Krefft 1987:85 [ref. 20751]. •Valid as Lycodonus malvinensis Gosztonyi 1981 -- (Anderson & Gosztonyi 1991:7 [ref. 19348], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:32 [ref. 27603], Balushkin et al. 2011:1017 [ref. 32449]). Current status: Valid as Lycodonus malvinensis Gosztonyi 1981. Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

mandibularis, Lycodapus Gilbert [C. H.] 1915:369, Pl. 20 (fig. 20) [Proceedings of the United States National Museum v. 48 (no. 2075); ref. 1632] Monterey Bay, California, 36°40.2'N, 122°06.0'W, U.S.A., Albatross station 4533, depth 144-293 fathoms. Holotype: USNM 75823. Paratypes: CAS-SU 22990 (1), 25765 (5, now 4); USNM 149514 (2). Type catalog: Böhlke 1953:102 [ref. 12291], Springer & Anderson 1997:19 [ref. 22953]. •Valid as Lycodapus mandibularis Gilbert 1915 -- (Peden & Anderson 1978:1938 [ref. 8895], Anderson 1981 [ref. 20873], Eschmeyer & Herald 1983:104 [ref. 9277], McAllister 1990:158 [ref. 14674], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:737 [ref. 25968], Anderson & Fedorov 2004:22 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Balushkin et al. 2011:986 [ref. 32449], Page et al. 2013:162 [ref. 32708], Pietsch & Orr 2015:57 [ref. 34694]). Current status: Valid as Lycodapus mandibularis Gilbert 1915. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

manusanus, Pyrolycus Machida [Y.] & Hashimoto [J.] 2002:3, Figs. 1-13 [Ichthyological Research v. 49 (no. 1); ref. 25949] Manus basin, Bismarck Sea, 3°43.66'S, 151°40.39'E, depth 1694 meters. Holotype: BSKU 87741. Paratypes: BSKU 87003 (1), 87004 (1, damaged and skeletonized), 87008 (1, skeletonized), 82228 (1), 87742 (1); NSMT-P 59364-65 (1, 1). •Valid as Pyrolycus manusanus Machida & Hashimoto 2002 -- (Anderson & Fedorov 2004:37 [ref. 27603], Anderson 2006:64 [ref. 28709], Anderson & Møller 2007:183 [ref. 29401], Balushkin et al. 2011:1027 [ref. 32449], Fricke et al. 2019:246 [ref. 36673]). Current status: Valid as Pyrolycus manusanus Machida & Hashimoto 2002. Zoarcidae: Lycodinae. Distribution: Western Pacific: Bismarck Sea off New Ireland (Papua New Guinea). Habitat: marine.

maoriensis, Lycenchelys Andriashev [A. P.] & Fedorov [V. V.] 1986:24, Figs. 1-2 [Voprosy Ikhtiologii v. 26 (no. 1); ref. 6133] New Zealand, Campbell Plateau, 44°30'S, 174°23'E, depth 710-730 meters. Holotype: ZIN 45285. Paratypes: ZIN 45285a (1). Type catalog: Balushkin et al. 2011:981 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. Also see Lycenchelys maoriorum Anderson 1990, which is an unjustified emendation. •Valid as Lycenchelys maoriensis Andriashev & Fedorov 1986 -- (Paulin et al. 1989:211 [ref. 24556], Anderson 1990:12 [ref. 16143] as maoriorum, Anderson & Fedorov 2004:18 [ref. 27603], Shinohara & Anderson 2007:62 [ref. 29111], Anderson & Møller 2007:183 [ref. 29401], Balushkin et al. 2011:981 [ref. 32449], Møller & Anderson 2015:1426 [ref. 34289]). Current status: Valid as Lycenchelys maoriensis Andriashev & Fedorov 1986. Zoarcidae: Lycodinae. Distribution: New Zealand region. Habitat: marine.

maoriorum, Lycenchelys Anderson [M. E.] 1990:12, 15 [The J.L.B Smith Institute of Ichthyology Special Publication No. 50; ref. 16143] New Zealand, 44°30'S, 174°23'E, depth 710-730 meters. Holotype: ZIN 45285. Paratypes: ZIN 45285a (1). An unjustified emendation of and objective synonym of Lycenchelys maoriensis Andriashev & Fedorov 1986. •Objective synonym of Lycenchelys maoriensis Andriashev & Fedorov 1986 -- (Anderson & Fedorov 2004:18 [ref. 27603]). Current status: Synonym of Lycenchelys maoriensis Andriashev & Fedorov 1986. Zoarcidae: Lycodinae. Habitat: marine.

marginalis, Lycodes brevipes Taranetz [A. Ya.] & Andriashev [A. P.] in Taranetz 1937:163 [Izvestiia Tikhookeanskogo nauchnogo instituta rybnogo khoziaistva [Bulletins of the Pacific Science Institute] v. 11; ref. 13384] Not available, no distinguishing features. Apparently misprint for Lycodes breviceps diapteroides. •Nomen nudum, in the synonymy of Lycodes brevipes Bean 1890 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Anderson & Fedorov 2004:48 [ref. 27603], Parin et al. 2014:408 [ref. 33547]). Current status: Synonym of Lycodes brevipes Bean 1890. Zoarcidae: Lycodinae.

marinae, Haushia Lloris [D.] 1988:246, Fig. 1 [Miscellània Zoològica v. 12 (1988); ref. 15965] Staten Island, 54°46.3'S, 63°48.0'W, off Tierra del Fuego, depth 130 meters. Holotype (unique): IIPB 104/1987. Type catalog: Allué et al. 2000:75 [ref. 30499]. •Valid as Pogonolycus marinae (Lloris 1988) -- (Anderson & Gosztonyi 1991:13 [ref. 19348], Lloris & Rucabado 1991:70 [ref. 22530], Anderson 1994:120 [ref. 21438], López et al. 1996:32 [ref. 23681], Pequeño 1997:83 [ref. 23536], Anderson & Fedorov 2004:37 [ref. 27603], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Pogonolycus marinae (Lloris 1988). Zoarcidae: Lycodinae. Distribution: South Atlantic: southern South America. Habitat: marine.

marisalbi, Lycodes Knipowitsch [N. M.] 1906:48, Figs. 9-12 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] White Sea, Russia, 65°13'N, 39°07'E, depth 91 meters. Lectotype: ZIN 40511. Paralectotypes: ZIN 13500-502 (4, 3, 3). Type catalog: Balushkin et al. 2011:1000 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. Originally as maris-albi. Lectotype selected by Andriashev 1973:545 [ref. 7214]. •Synonym of Lycodes pallidus Collett 1879, but a valid subspecies -- (Andriashev 1973:545 [ref. 9214], Andriashev in Whitehead et al. 1986:1142 [ref. 13677]). •Synonym of Lycodes pallidus Collett 1879 -- (Anderson 1994:119 [ref. 21438]). •Valid as Lycodes marisalbi Knipowitsch 1906 -- (Møller 2001:111 [ref. 25378], Anderson & Fedorov 2004:26 [ref. 27603], Balushkin et al. 2011:1000 [ref. 32449], Page et al. 2013:163 [ref. 32708], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:414 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:64, 202 [ref. 36199], Mecklenburg et al. 2016:168 [ref. 34440] see remarks, Mecklenburg et al. 2018:335 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes marisalbi Knipowitsch 1906. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

matallanasi, Pachycara Corbella [C.] & Møller [P. R.] 2014:181, Figs. 1-5 [Marine Biology Research; ref. 33428] Solomon Sea, 7°49'04"S, 156°02'08"E, depth 4350-4450 meters. Holotype: ZMUC P766564. Paratypes: ZMUC. •Valid as Pachycara matallanasi Corbella & Møller 2014. Current status: Valid as Pachycara matallanasi Corbella & Møller 2014. Zoarcidae: Lycodinae. Distribution: Solomon Sea. Habitat: marine.

matsubarai, Lycodes Toyoshima [M.] 1985:218, Fig. 53 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Kitami-yamato Bank, 44°47'N, 144°01.5'E, southern Okhotsk Sea. depth 200 meters. Holotype: HUMZ 33970. Paratypes: HUMZ 33974 (1), 33948 (1), 33971-72 (2), 33974-79 (6), 33981 (1), 33983 (1), 33950 (2), 33960 (1), 33968-69 (2), 40945-46 (2). Lycodes Matsubarai and L. yamatoi were simutaneouly described in ref. 5722; Nazarkin & Shinohara 2012:34 [ref. 32106] act as first revisers selecting matsubarai over yamatoi. •Synonym of Lycodes microlepidotis Schmidt 1950 -- (Balushkin et al. 2011:1000 [ref. 24994], Nazarkin et al. 2014:192 [ref. 33213]). •Valid as Lycodes matsubarai Toyoshima 1985 -- (Anderson 1994:118 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:36 [ref. 25650], Nakabo 2002:1039 [ref. 26193], Anderson & Fedorov 2004:26 [ref. 27603], Nazarkin 2010:9 [ref. 30733], Nazarkin & Shinohara 2012:34 [ref. 32106], Shinohara et al. 2014:265 [ref. 33330], Parin et al. 2014:414 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331]). Current status: Valid as Lycodes matsubarai Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea and Sea of Japan to Peter the Great Bay, Yamato Bank: Japan and Russia. Habitat: marine.

mcallisteri, Lycodes Møller [P. R.] 2001:112, Figs. 2-4 [Ichthyological Research v. 48 (no. 2); ref. 25378] Western Baffin Bay, 70°02'N, 65°32'W, eastern Arctic Canada, depth 579 meters. Holotype: ZMUC P764746. Paratypes: CMNFI 2000-0055(1); HUMZ 172115 (1); NMC 90-0223 (1); USNM 362128 (1); ZIN 52125 (1); ZMUC P765161 (1), P765163 (1), P765168 (1). •Valid as Lycodes mcallisteri Møller 2001 -- (Anderson & Fedorov 2004:26 [ref. 27603], Møller et al. 2010:61 [ref. 31169], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin et al. 2011:1000 [ref. 32449], Afonso et al. 2016:65, 203 [ref. 36199], Mecklenburg et al. 2018:337 [ref. 36370]). Current status: Valid as Lycodes mcallisteri Møller 2001. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and Labrador (Canada). Habitat: marine.

mcallisteri, Oidiphorus Anderson [M. E.] 1988:102, Figs. 8B, 38 [Antarctic Research Series, Washington DC. v. 47; ref. 7304] Off South Georgia Island, Scotia Sea, 53°23'S, 37°11'-21'W, depth 1299-1400 meters. Holotype: LACM 10608-2. Paratypes: LACM 11062-7 (1, c&s). •Valid as Oidiphorus mcallisteri Anderson 1988 -- (Anderson 1990:271 [ref. 21695], Miller 1993:673 [ref. 21297], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:33 [ref. 27603], Anderson 2006:9 [ref. 28576], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Oidiphorus mcallisteri Anderson 1988. Zoarcidae: Lycodinae. Distribution: Southern Ocean: Scotia Sea. Habitat: marine.

megalocephala, Lycodes rossi forma Knipowitsch [N. M.] 1906:27 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Syntypes: ZIM 13406 (1), 13407 (5), 13411 (3), 13412 (2), 13420 (1), 13421 (3), 13422 (3), 13423 (1), 13434 (2). Type cataqlog: Balushkin et al. 2011:1010 [ref. 32449], Balushkin et al. 2012:164 [ref. 33667]. Not available, infrasubspecific; publication in an available way as a species or subspecies not investigated [Knipowitsch used v. [variety] and f. [forma] -- in his work, the former is considered subspecific the latter infrasubspecific; see Article 45(g)]. From Barents Sea, Russia. •Infrasubspecific, in the synonymy of Lycodes rossii Malmgren 1865 -- (Anderson 1994:119 [ref. 21438] as rossi, Anderson & Fedorov 2004:48 [ref. 27603], Balushkin et al. 2011:1010 [ref. 32449], Parin et al. 2014:420 [ref. 33547]). Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae.

melanostomias, Lycenchelys Toyoshima [M.] in Amaoka et al. 1983:271, Pl. 157; Figs. 25-27 [Fishes from the north-eastern Sea of Japan and the Okhotsk Sea off Hokkaido; ref. 8379] Off Shiretoko Peninsula, Hokkaido, Okhotsk Sea, 44°19.5'N, 145°01'E, depth 915-925 meters. Holotype (unique): HUMZ 77572. •Valid as Lycenchelys melanostomias Toyoshima 1983 -- (Toyoshima in Masuda et al. 1984:306 [ref. 6441], Toyoshima 1985:170 [ref. 5722], Anderson 1994:117 [ref. 21438], Fedorov 1995:130 [ref. 21958], Nakabo 2000:1032 [ref. 25182], Nakabo 2002:1032 [ref. 26193], Anderson & Fedorov 2004:18 [ref. 27603], Imamura et al. 2004:84 [ref. 27600], Imamura et al. 2005:3 [ref. 28190], Shinohara & Anderson 2007:64 [ref. 29111], Balushkin et al. 2011:1026 [ref. 32449], Parin et al. 2014:400 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys melanostomias Toyoshima 1983. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

melastomus, Patagolycus Matallanas [J.] & Corbella [C.] 2012:8, Figs. 3-9 [Zootaxa No. 3296; ref. 31967] Southwestern Atlantic, 46°26'50"S, 60°21'37"W, depth 253 meters. Holotype: UAB.ZM2. Paratypes: BMNH 1936.8.26.984-986 (3), 1936.8.26.962 (1); plus several lots at UAB. •Valid as Patagolycus melastomus Matallanas & Corbella 2012. Current status: Valid as Patagolycus melastomus Matallanas & Corbella 2012. Zoarcidae: Lycodinae. Distribution: Southwestern Atlantic: off Argentina and the Falkland Islands. Habitat: marine.

mesoporum, Pachycara Anderson [M. E.] 1989:237, Fig. 13 [Proceedings of the California Academy of Sciences v. 46 (no. 10); ref. 13487] Off Antofagasta, Chile, depth probably about 300 meters. Holotype: CAS 62406. Paratypes: LACM 43717-1 (5), MNHNC P.6653 (1), ZIN 48394 (1). Type catalog: Meléndez C. et al. 1993:173 [ref. 24124], Balushkin et al. 2011:1020 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. •Valid as Pachycara mesoporum Anderson 1989 -- (Anderson 1991:157 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:226 [ref. 22911], Pequeño 1997:83 [ref. 23536], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson 2006:12 [ref. 28576], Anderson & Mincarone 2006:25 [ref. 28658], Møller & King 2007:588 [ref. 29259], Balushkin et al. 2011:1020 [ref. 32449], Shinohara 2012:55 [ref. 32102], Balushkin et al. 2011:1020 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara mesoporum Anderson 1989. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific. Habitat: marine.

meteori, Melanostigma Balushkin [A. V.] & Orlovskaya [M. V.] 2019:136, Figs. 1-2 [Voprosy Ikhtiologii v. 59 (no. 2); ref. 36664] Meteor Bank, southeastern Atlantic Ocean, 48°02'S, 08°24'E, depth 950 - 1000 meters. Holotype: ZIN 56303. Paratypes: ZIN. •Valid as Melanostigma meteori Balushkin & Orlovskaya 2019 -- (Orlovskaya & Balushkin 2019:505 [ref. 37050]). Current status: Valid as Melanostigma meteori Balushkin & Orlovskaya 2019. Zoarcidae: Gymnelinae. Distribution: Meteor Bank, southeastern Atlantic Ocean. Habitat: marine.

microcephalus, Bothrocarichthys Schmidt [P. Yu.] 1938:653 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 19 (no. 8) [1938]; ref. 3943] West coast of Kamchatka, Russia, 58°21'N, 153°30'E, Okhotsk Sea, depth 70-190 meters. Syntypes: (5) ZIN 24847 (2, now 1), 31170 (1). Thpe catalog: Balushkin et al. 2011:974 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. Appear first as name only in Schmidt in Taranetz 1937. •Valid as Bothrocarichthys microcephalus Schmidt 1938 -- (Sheiko & Fedorov 2000:34 [ref. 25650], Fedorov et al. 2003:110 [ref. 27579]). •Valid as Bothrocarina microcephala (Schmidt 1938) -- (Toyoshima in Amaoka et al. 1983:275, 334 [ref. 8379], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Anderson 1994:116 [ref. 21438], Shinohara et al. 1996:180 [ref. 26605], Nakabo 2000:1041 [ref. 25182], Nakabo 2002:1041 [ref. 26193], Anderson & Fedorov 2004:11 [ref. 27603], Balushkin et al. 2011:974 [ref. 32449], Parin et al. 2014:386 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331]). Current status: Valid as Bothrocarina microcephala (Schmidt 1938). Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

microcephalus, Lycodes Jensen [A. S.] 1902:206 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Reykjanes Ridge, south-west of Iceland, 60°37'N, 27°52'W, depth 799 fathoms. Holotype (unique): ZMUC 78. Type catalog: Nielsen 1974:77 [ref. 9588]. •Valid as Lycodes microcephalus Jensen 1902 -- (Andriashev 1973:544 [ref. 9214], Andriashev in Whitehead et al. 1986:1141 [ref. 13677], Anderson 1994:118 [ref. 21438]). •Valid as Pachycara microcephalum (Jensen 1902) -- (Møller & Anderson 2000:62 [ref. 24495], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Pachycara microcephalum (Jensen 1902). Zoarcidae: Lycodinae. Distribution: Eastern North Atlantic. Habitat: marine.

microchir, Lycodapus Schmidt [P. Yu.] 1950:108 [95], Pl. 10 (fig. 1) [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] Off Sakhalin Island, 52°47'30"N, 144°51'E, Okhotsk Sea, depth 1150 meters. Holotype: ZIN 24848. Type catalog: Balushkin et al. 2011:986 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. On p. 95 of English translation. Appeared as name only in Schmidt 1935. •Status uncertain -- (Peden & Anderson 1978:1957 [ref. 8895]). •Valid as Lycodapus microchir Schmidt 1950 -- (Toyoshima in Masuda et al. 1984:309 [ref. 6441], Anderson 1989:150 [ref. 13419], Anderson 1994:118 [ref. 21438], Nakabo 2000:1040 [ref. 25182], Sheiko & Fedorov 2000:36 [ref. 25650], Nakabo 2002:1040 [ref. 26193], Anderson & Fedorov 2004:22 [ref. 27603], Shinohara et al. 2005:436 [ref. 28370], Balushkin et al. 2011:986 [ref. 32449], Parin et al. 2014:405 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331]). Current status: Valid as Lycodapus microchir Schmidt 1950. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

microlepidotus, Lycodes Schmidt [P. Yu.] 1950:91 [78], Pl. 6 (fig. 1) [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] East coast of Sakhalin and west coast of Kamchatka, Russia, Okhotsk Sea. Syntypes: ZIN 24827 (4), 24844 (1), 25268 (1), Type catalog: Balushkin et al. 2011:1001 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. On p. 78 of English translation. •Valid as Lycodes microlepidotus Schmidt 1950 -- (Toyoshima 1985:210 [ref. 5722], Anderson 1994:118 [ref. 21438], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:37 [ref. 25650], Anderson & Fedorov 2004:26 [ref. 27603], Shinohara & Shirai 2005:171 [ref. 28232], Nazarkin 2010:4 [ref. 30733], Balushkin et al. 2011:1001 [ref. 32449], Parin et al. 2014:415 [ref. 33547]). Current status: Valid as Lycodes microlepidotus Schmidt 1950. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

microphthalmus, Melanostigma Norman [J. R.] 1937:110, Fig. 58 [Discovery Reports v. 16; ref. 3227] South of the Falkland Islands, 52°40'S, 58°30'W, Discovery station WS248, depth 210-242 meters. Holotype: BMNH 1936.8.26.1047. Paratype: BMNH 1936.8.26.1046 (1). •Valid as Maynea microphthalmus (Norman 1937) -- (Gosztonyi 1977:221 [ref. 6103] as microphthalma). •Valid as Letholycus microphthalmus (Norman 1937) -- (Anderson 1988:273 [ref. 6334], Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:14 [ref. 27603], Balushkin et al. 2011:977 [ref. 32449]). Current status: Valid as Letholycus microphthalmus (Norman 1937). Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

microporus, Lycenchelys Andriashev [A. P.] 1955:367, Fig. 2 (no. 5), Figs. 11-12 [Trudy Instituta Zoologii / Akademiia Nauk, Azerbaidzhanskoi SSR. v. 18; ref. 12261] Northern part of central depression of the Bering Sea, 59°43'N, 179°39'E, depth 3120 meters. Holotype (unique): ZIN 32963. Type catalog: Balushkin et al. 2011:981 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667] •Valid as Lycenchelys microporus Andriashev 1955 -- (Toyoshima 1985:152 [ref. 5722], Nakabo 2000:1034 [ref. 25182]). •Valid as Lycenchelys micropora Andriashev 1955 -- (Anderson 1994:117 [ref. 21438], Anderson 1995:84 [ref. 21928], Sheiko & Fedorov 2000:35 [ref. 25650], Mecklenburg et al. 2002:695 [ref. 25968], Anderson & Imamura 2002:357 [ref. 26499], Anderson & Fedorov 2004:18 [ref. 27603], Balushkin et al. 2011:981 [ref. 32449], Parin et al. 2014:401 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys micropora Andriashev 1955. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific: Bering Sea. Habitat: marine.

microporus, Lycodes Toyoshima [M.] in Amaoka et al. 1983:263, Pl. 152: Figs. 23-24 [Fishes from the north-eastern Sea of Japan and the Okhotsk Sea off Hokkaido; ref. 8379] Abashiri, northeastern Hokkaidoo, Okhotsk Sea, 44°11'N, 144°47'E, depth 650-700 meters. Holotype: HUMZ 77795. Paratypes: HUMZ 77783-87 (5), 77807 (1), 78782 (1), 79662-63 (2). •Valid as Lycodes microporus Toyoshima 1983 -- (Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:221 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1034 [ref. 26193], Fedorov et al. 2003:115 [ref. 27579], Anderson & Fedorov 2004:26 [ref. 27603], Ikeda et al. 2008:357 [ref. 29896], Balushkin et al. 2011:1001 [ref. 32449], Parin et al. 2014:415 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331]). Current status: Valid as Lycodes microporus Toyoshima 1983. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Hokkaido, southern Okhotsk Sea and northern Okhotsk Sea off Russia. Habitat: marine.

micropus, Phucocoetes variegatus forma Smitt [F. A.] 1898:43, Pl. 5 (fig. 33) [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 24 (afd. 4) (no. 5); ref. 4147] Puerto Gallegos, 51.5°S, Argentina, southwestern Atlantic. Syntypes: (2) [whereabouts unknown]. •Synonym of Iluocoetes elongatus (Smitt 1898) -- (Gosztonyi 1977:215 [ref. 6103], Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:14 [ref. 27603], Matallanas & Corbella 2012:1 [ref. 31967]). Current status: Synonym of Iluocoetes elongatus (Smitt 1898). Zoarcidae: Lycodinae. Habitat: marine.

middendorffii, Hadropareia Schmidt [P. Yu.] 1904:205, Fig. 14 [Fishes of the eastern seas of the Russian Empire; ref. 3946] Big Shantarsky Island, Shantarsky Islands, Russia, Okhotsk Sea. Syntypes: ZIN 12994 (2). Type catalog: Balushkin et al. 2011:967 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. •Valid as Hadropareia middendorffii Schmidt 1904 -- (Lindberg & Krasyukova 1975:169 [ref. 7348], Matyushin 1989:529 [ref. 17696], Anderson 1994:117 [ref. 21438], Sheiko & Fedorov 2000:35 [ref. 25650], Fedorov et al. 2003:111 [ref. 27579], Anderson & Fedorov 2004:7 [ref. 27603], Shinohara et al. 2004:137 [ref. 27581] as middendorfii, Balushkin et al. 2011:967 [ref. 32449], Parin et al. 2014:395 [ref. 33547], Radchenko et al. 2018:314 [ref. 36243]). Current status: Valid as Hadropareia middendorffii Schmidt 1904. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

mirabilis, Lycodonus Goode [G. B.] & Bean [T. H.] 1883:208 [Bulletin of the Museum of Comparative Zoology v. 10 (no. 5); ref. 1838] Atlantic, 38°20'08"N, 73°23'20"W, Blake station 337, depth 740 fathoms. Holotype (unique): MCZ 25859. •Valid as Lycodonus mirabilis Goode & Bean 1883 -- (Gosztonyi 1981:156 [ref. 8596], Toyoshima 1985:143 [ref. 5722], Anderson 1994:119 [ref. 21438], Koyanagi in Okamura et al. 1995:205 [ref. 22531], Moore et al. 2003:229 [ref. 27076], Anderson & Fedorov 2004:32 [ref. 27603], Møller et al. 2010:63 [ref. 31169], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin et al. 2011:1018 [ref. 32449], Afonso et al. 2016:68, 214 [ref. 36199], Mecklenburg et al. 2018:366 [ref. 36370]). Current status: Valid as Lycodonus mirabilis Goode & Bean 1883. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and northwestern Atlantic. Habitat: marine.

moelleri, Pachycara Shinohara [G.] 2012:52, Figs. 1-5 [Bulletin of the National Museum of Nature and Science (Ser. A) Supplement No. 6; ref. 32103] Off Kii Peninsula, 32°00.7'N, 135°57.2'E to 32°00.3'N, 136°02.7'E, Wakayama, Honshu, Japan, depth 4093 meters. Holotype (unique): NSMT-P 105631. •Valid as Pachycara moelleri Shinohara 2012 -- (Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara moelleri Shinohara 2012. Zoarcidae: Lycodinae. Distribution: Western North Pacific: off Kii Peninsula, Japan. Habitat: marine.

moelleri, Pyrolycus Anderson [M. E.] 2006:64, Figs. 1-3 [Journal of the Royal Society of New Zealand v. 36 (no. 2); ref. 28709] Hydrothermal vent field on Brothers Seamount, Kermadec Ridge, 34°52.7'S, 179°04.3'E, depth 1336 meters. Holotype: NMNZ P.42182. Paratypes: NMNZ P.42183 (1). •Valid as Pyrolycus moelleri Anderson 2006 -- (Anderson & Møller 2007:183 [ref. 29401], Møller & Anderson 2015:1433 [ref. 34289]). Current status: Valid as Pyrolycus moelleri Anderson 2006. Zoarcidae: Lycodinae. Distribution: Kermadec Ridge, southwestern Pacific. Habitat: marine.

mollis, Bothrocara Bean [T. H.] 1890:39 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Off Cape St. James, Queen Charlotte Islands, British Columbia, Canada, northeastern Pacific, 51°23'N, 130°34'W, Albatross station 2860, depth 876 fathoms. Holotype (unique): USNM 45359. Type catalog: Springer & Anderson 1997:19 [ref. 22953]. No parentheses are needed around the author and date in the combination Bothrocara molle (Art. 51.3.1 of the Code). •Valid as Bothrocara mollis Bean 1890 -- (Sheiko & Fedorov 2000:34 [ref. 25650]). •Valid as Bothrocara molle Bean 1890 -- (Toyoshima in Amaoka et al. 1983:137, 208, 277, 335 [ref. 8379], Toyoshima in Masuda et al. 1984:309 [ref. 6441], McAllister 1990:156 [ref. 14674], Anderson 1994:116 [ref. 21438], Shinohara et al. 1996:179 [ref. 26605], Nakabo 2000:1041 [ref. 25182], Mecklenburg et al. 2002:732 [ref. 25968], Shinohara et al. 2001:328 [ref. 25995], Nakabo 2002:1041 [ref. 26193], Anderson & Fedorov 2004:11 [ref. 27603], Stevenson & Anderson 2005:53 [ref. 28569], Anderson 2006:2 [ref. 28576], Anderson et al. 2009:182 [ref. 30175], Balushkin et al.2011:973 [ref. 32449], Parin et al. 2014:385 [ref. 33547], Kells et al. 2016:270 [ref. 35888], Robertson et al. 2017:92 [ref. 35676]). Current status: Valid as Bothrocara molle Bean 1890. Zoarcidae: Lycodinae. Distribution: Northern and eastern Pacific: Kamchatka to off Chile; western South Atlantic: South Georgia. Habitat: marine.

monstrosa, Lycenchelys Anderson [M. E.] 1982:208, Figs. 1-2 [Bulletin of Marine Science v. 32 (no. 1); ref. 8517] Gulf of Panama, eastern Pacific, 6°52'N, 79°28'W, depth 3200-3229 meters. Holotype: USNM 224467. Paratypes: CAS 47127 (2), LACM 39977-1 (2), MCZ 56940 (2), UF 233483 [ex UMML 33483] (1), USNM 224466 (2). Type catalog: Springer & Anderson 1997:19 [ref. 22953]. •Valid as Lycenchelys monstrosa Anderson 1982 -- (Anderson 1994:117 [ref. 21438], Anderson 1995:86 [ref. 21928], Anderson & Balanov 2000:1056 [ref. 25023], Anderson & Fedorov 2004:18 [ref. 27603], Anderson 2006:12 [ref. 28576], Balushkin et al. 2011:1026 [ref. 32449], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys monstrosa Anderson 1982. Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

montagui, Cepolophis Kaup [J. J.] (ex Montagu) 1856:97 [Archiv für Naturgeschichte v. 22 (pt. 1); ref. 2574] South coast of Devon. Holotype (unique): whereabouts unknown. Misspelled montgui and as a name in synonymy in Kaup 1856:156 [ref. 2573]. Based on the single specimen described and illustrated by Montagu 1811:95-97, Pl. 4 (fig. 2) [ref. 3038]; description was nearly the same as that provided by Montagu. •Not treated in Zoarcidae by Anderson & Fedorov 2004 [ref. 27603]. Zoarcidae: Gymnelinae. Habitat: marine.

morenoi, Lycodalepis Lahille [F.] 1908:413, Pl. 6 [Anales del Museo Nacional de Historia Natural de Buenos Aires (Serie 3) v. 9; ref. 5869] Cape San Antonio, 36°20'S, Argentina, depth 15-20 meters. Holotype: Mus. Nac. Buenos Aires. •Synonym of Austrolycus laticinctus (Berg 1895) -- (Gosztonyi 1977:202 [ref. 6103], Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:10 [ref. 27603]). Current status: Synonym of Austrolycus laticinctus (Berg 1895). Zoarcidae: Lycodinae. Habitat: marine.

mucosus, Lycodes Richardson [J.] 1855:362, Pl. 26 [The last of the Arctic voyages in search of Sir J. Franklin; ref. 18631] Northumberland Sound, Perry Island, Arctic Canada. Syntypes: (2) BMNH 1855.9.19.760. •Valid as Lycodes mucosus Richardson 1855 -- (Toyoshima 1985:200 [ref. 5722], McAllister 1990:161 [ref. 14674], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:708 [ref. 25968], Anderson & Fedorov 2004:26-27 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Mecklenburg et al. 2007:181 [ref. 29448], Møller et al. 2010:61 [ref. 31169], Balushkin et al. 2011:1002 [ref. 32449], Page et al. 2013:163 [ref. 32708], Parin et al. 2014:416 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:65, 204 [ref. 36199], Kells et al. 2016:272 [ref. 35888], Mecklenburg et al. 2016:170 [ref. 34440] see remarks, Mecklenburg et al. 2018:338 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes mucosus Richardson 1855. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and adjacent North Pacific and North Atlantic. Habitat: marine.

multifasciatus, Lycodes palearis Schmidt [P. Yu.] 1950:95 [82], Pl. 7 (fig. 2) [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] West coast of Kamchatka, Russia, opposite the mouths of the Ozernaya and Bol'shaya rivers, depth 97-250 meters. Syntypes: ZIN 29102 (1), 29102a (3). Type catalog: Balushkin et al. 2011:989 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. On p. 82 of English translation. •Valid as Lycodes multifasciatus Schmidt 1950 -- (Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:214 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Møller 2001:111 [ref. 25378], Nakabo 2002:1039 [ref. 26193]). •Questionably a synonym of Lycodes fasciatus (Schmidt 1904) -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Synonym of Lycodes fasciatus (Schmidt 1904) -- (Mecklenburg et al. 2002:720 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603]). •Synonym of Lycodes brashnikovi Soldatov 1918 -- (Nazarkin 2010:603 [ref. 31019], Balushkin et al. 2011:989 [ref. 32449], Parin et al. 2014:408 [ref. 33547], Mecklenburg et al. 2016:173 [ref. 34440]). Current status: Synonym of Lycodes brashnikovi Soldatov 1918. Zoarcidae: Lycodinae. Habitat: marine.

multispinosa, Krusensterniella Soldatov [V. K.] 1922:158, Fig. [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23 (no. 2) (for 1917); ref. 18736] 55°57'N, 138°13'E, Okhotsk Sea, depth 87 meters. Syntypes: ZIN 19961 (2, now 1). Type locality: Balushkin et al 2011:969 [ref. 32499], Balushkin et al. 2012:162 [ref. 33667] with ZIN 19961 as 'lectotype' (?). •Valid as Krusensterniella multispinosa Soldatov 1922 -- (Lindberg & Krasyukova 1975:129 [ref. 7348], Toyoshima in Masuda et al. 1984:305 [ref. 6441], Anderson 1994:117 [ref. 21438], Nakabo 2000:1044 [ref. 25182], Sheiko & Fedorov 2000:35 [ref. 25650], Nakabo 2002:1044 [ref. 26193], Fedorov et al. 2003:111 [ref. 27579], Anderson & Fedorov 2004:7 [ref. 27603], Balushkin et al. 2011:969 [ref. 32449], Parin et al. 2014:396 [ref. 33547]). Current status: Valid as Krusensterniella multispinosa Soldatov 1922. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

muraena, Lycodes Collett [R.] 1878:15 [Forhandlinger i Videnskabs-selskabet i Christiania (for 1878) No. 4; ref. 21043] 325 kilometers west-southwest of Bodø, Norway, depth 640 meters. Holotype (unique): ZMUO J4538. Type catalog: Pethon 1969:12 [ref. 19268]. Also appeared as new in Collett 1880:116, Pl. 4 (figs. 29-31) [ref. 887]. •Valid as Lycenchelys muraena (Collett 1878) -- (Andriashev 1973:541 [ref. 9214], Andriashev in Whitehead et al. 1986:1134 [ref. 13677], Anderson 1994:117 [ref. 21438], Koyanagi in Okamura et al. 1995:192 [ref. 22531], Møller 1999:327 [ref. 24052], Anderson & Fedorov 2004:18 [ref. 27603], Møller et al. 2010:59 [ref. 31169], Balushkin et al. 2011:981 [ref. 32449], Parin et al. 2014:401 [ref. 33547], Afonso et al. 2016:62, 192 [ref. 36199], Mecklenburg et al. 2018:315 [ref. 36370]). Current status: Valid as Lycenchelys muraena (Collett 1878). Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

nakamurae, Furcimanus Tanaka [S.] 1914:303, Pl. 82 (fig. 276) [Figures and descriptions of the fishes of Japan v. 18; ref. 14935] Off Niigata, Japan, Sea of Japan. Holotype (unique): ZUMT (lost). The spelling nakamurae is acceptable; early workers almost certainly Latinized the name first, and then added an 'e' (Art. 31.1.1) [WNE, 12 Mar. 2012] -- we suggest using nakamurae. Plate is in vol. 17 [ref. 14934]. •Synonym of Lycodes diapterus Gilbert 1892, but a valid subspecies -- (Lindberg & Krasyukova 1975:149 [ref. 7348]). •Valid as Furcimanus nakamurae Tanaka 1914 -- (Anderson 1994:119 [ref. 21438]). •Valid as Lycodes nakamurai (Tanaka 1914) -- (Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:224 [ref. 5722], Sokolovskaya et al. 1998:12 [ref. 24670], Nakabo 2000:1029 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:724 [ref. 25968], Nakabo 2002:1029 [ref. 26193], Youn 2002:375, 640 [ref. 26218], Anderson & Fedorov 2004:27 [ref. 27603], Shinohara & Shirai 2005:172 [ref. 28232], Kim et al. 2006:276 [ref. 29191], Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:1002 [ref. 32449], Nazarkin et al. 2014:177 [ref. 33213], Shinohara et al. 2014:265 [ref. 33330], Saveliev et al. 2014:125 [ref. 33340], Parin et al. 2014:416 [ref. 33547]). Current status: Valid as Lycodes nakamurae (Tanaka 1914). Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

nanospinata, Lycenchelys Anderson [M. E.] 1988:96, Fig. 35 [Antarctic Research Series, Washington DC. v. 47; ref. 7304] Southeastern basin, Scotia Sea, 59°08'-01'S, 36°57'-50'W, depth 2815-2818 meters. Holotype (unique): LACM 10816-2. •Valid as Lycenchelys nanospinata Anderson 1988 -- (Anderson 1990:262 [ref. 21695], Miller 1993:667 [ref. 21297], Anderson 1994:117 [ref. 21438], Anderson & Balanov 2000:1056 [ref. 25023], Anderson & Fedorov 2004:18 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys nanospinata Anderson 1988. Zoarcidae: Lycodinae. Distribution: Southern Ocean: Scotia Sea, Antarctica. Habitat: marine.

nazca, Pachycara Anderson [M. E.] & Bluhm [H.] 1997:220, Figs. 1-3 [Transactions of the Royal Society of South Africa v. 51 (1996); ref. 22911] Peru Basin, 7°03.2'S, 88°24.5'W, depth 4107 meters. Holotype: SMF 27979. Paratypes: SMF 27585 (1), 27587 (1), 27982 (8), 27984 (1); SAIAB [formerly RUSI] 49603 (1). •Valid as Pachycara nazca Anderson & Bluhm 1997 -- (Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Møller & King 2007:588 [ref. 29259], Shinohara 2012:55 [ref. 32102], Anderson 2012:41 [ref. 32270], Balushkin et al. 2011:1027 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara nazca Anderson & Bluhm 1997. Zoarcidae: Lycodinae. Distribution: Eastern Pacific: Peru basin. Habitat: marine.

nazumii, Allolepis Mori [T.] 1956:29 [Science Reports of the Hyogo University of Agriculture Series Natural Science v. 2 (no. 2); ref. 7765] Yamato Bank, central Japan Sea, 38°-39°N, 134°E, depth 300-400 meters. Holotype: not researched. Paratypes: (1). •Valid as Allolepis nazumii Mori 1956 -- (Lindberg & Krasyukova 1975:183 [ref. 7348]). •Synonym of Bothrocara hollandi (Jordan & Hubbs 1925) -- (Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:11 [ref. 27603], Anderson et al. 2009:179 [ref. 30175], Parin et al. 2014:385 [ref. 33547]). Current status: Synonym of Bothrocara hollandi (Jordan & Hubbs 1925). Zoarcidae: Lycodinae. Distribution: Western North Pacific: eastern Bering Sea to northwestern Pacific. Habitat: marine.

nebulosus, Lycodes Krøyer [H. N.] 1845:140 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1844 (no. 8); ref. 19477] Greenland. No types known. •Status uncertain, nomen dubium in Zoarcidae -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:48 [ref. 27603]). Zoarcidae: Lycodinae. Habitat: marine.

nigricans, Lycodes Jensen [A. S.] 1952:21, Pl. 1 (fig. 1) [Meddelelser om Grønland v. 142 (no. 7); ref. 12776] Amerdlok fjord near Holsteinsborg, Greenland, 340-473 meters. Holotype: ZMUC P76803. Paratypes: ZMUC P76799-802 (4), P76804-15 (12). Type catalog: Nielsen 1974:77 [ref. 9588]. •Synonym of Lycodes seminudus Reinhardt 1837 -- (Andriashev 1973:546 [ref. 9214], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:29 [ref. 27603], Balushkin et al. 2011:1011 [ref. 32449], Mecklenburg et al. 2014:142 [ref. 33463], Parin et al. 2014:421 [ref. 33547]). Current status: Synonym of Lycodes seminudus Reinhardt 1837. Zoarcidae: Lycodinae. Habitat: marine.

nigripalatum, Lycenchelys DeWitt [H. H.] & Hureau [J.-C.] 1980:812, Figs. 8-9 [Bulletin du Museum National d'Histoire Naturelle, Série 4: Section A: Zoologie, Biologie et Écologie Animales v. 1 (no. 3) (for 1979); ref. 1125] Bellingshausen Sea, southwest of Adelaide Island, 67°40.2'S, 70°16.2'W, depth 580-650 meters. Holotype (unique): MNHN 1974-0086. Appeared first as name only in Gosztonyi 1977:196 [ref. 6103]. •Valid as Lycenchelys nigripalatum DeWitt & Hureau 1980 -- (Pequeño 1989:48 [ref. 14125], Anderson 1990:263 [ref. 21695], Miller 1993:668 [ref. 21297], Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:18 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys nigripalatum DeWitt & Hureau 1980. Zoarcidae: Lycodinae. Distribution: Southern Ocean: Bellingshausen Sea, Antarctica. Habitat: marine.

nigrocaudata, Bothrocarina Suvorov [E. K.] 1935:435 [English p. 439], Figs. 1-2 [Bulletin de l'Academie des Sciences de l'U.R.S.S., Classe des Sciences Mathématiques et Naturelles 1935; ref. 4300] Okhotsk Sea, region of Ozernaya, west Kamchatka, Russia, 51°24.5'N, 156°17'E, depth 235-134 meters. Holotype (unique): ZIN 25072. Type catalog: Balushkin et al. 2011:975 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. •Valid as Lycogrammoides nigrocaudatus (Suvorov 1935) -- (Sheiko & Fedorov 2000:37 [ref. 25650], Fedorov et al. 2003:118 [ref. 27579]). •Valid as Bothrocarina nigrocaudata Suvorov 1935 -- (Toyoshima in Masuda et al. 1984:308 [ref. 6441], Anderson 1994:116 [ref. 21438], Nakabo 2000:1041 [ref. 25182], Nakabo 2002:1041 [ref. 26193], Anderson & Fedorov 2004:11 [ref. 27603], Balushkin et al. 2011:975 [ref. 32449], Parin et al. 2014:386 [ref. 33547]). Current status: Valid as Bothrocarina nigrocaudata Suvorov 1935. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

nishimurai, Lycodes Shinohara [G.] & Shirai [S. M.] 2005:172, Figs. 1-3 [Ichthyological Research v. 52 (no. 2); ref. 28232] Yamato Bank, 39°28.36'N, 135°0.31'E to 39°27.24'N, 134°59.02'E, Sea of Japan, depth 642-669 meters. Holotype: NSMT-P 63795. Paratypes: NSMT-P 63797-99 (3). •Valid as Lycodes nishimurai Shinohara & Shirai 2005 -- (Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:1002 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Saveliev et al. 2014:152 [ref. 33340], Parin et al. 2014:416 [ref. 33547]). Current status: Valid as Lycodes nishimurai Shinohara & Shirai 2005. Zoarcidae: Lycodinae. Distribution: Sea of Japan. Habitat: marine.

normani, Melanostigma Vladykov [V. D.] in Vladykov & McAllister 1961:71 [Nature Canada v. 88 (no. 3); ref. 21358] Not available, proposed conditionally after 1960; also without distinguishing features. •Nomen nudum, in the family Zoarcidae -- (Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:48 [ref. 27603]). Current status: Unknown. Zoarcidae: Gymnelinae.

notabilis, Krusensterniella Schmidt [P. Yu.] 1904:198, Fig. 12 [Fishes of the eastern seas of the Russian Empire; ref. 3946] Bellingshausen Cape and Popov Cape, eastern Sakhalin Island, Russia, Okhotsk Sea. Syntypes: USNM 92591 [ex ZIN 13011] (1), ZIN 13011 (now 6), 13032 (3). Type catalog: Springer & Anderson 1997:19 [ref. 22953], Balushkin et al. 2011:969 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. •Valid as Krusensterniella notabilis Schmidt 1904 -- (Lindberg & Krasyukova 1975:127 [ref. 7348], Toyoshima in Masuda et al. 1984:305 [ref. 6441], Anderson 1994:117 [ref. 21438], Nakabo 2000:1044 [ref. 25182], Sheiko & Fedorov 2000:35 [ref. 25650], Nakabo 2002:1044 [ref. 26193], Fedorov et al. 2003:112 [ref. 27579], Anderson & Fedorov 2004:7 [ref. 27603], Balushkin et al. 2011:969 [ref. 32449], Shinohara et al. 2014:264 [ref. 33330], Parin et al. 2014:396 [ref. 33547]). Current status: Valid as Krusensterniella notabilis Schmidt 1904. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk and Japan seas. Habitat: marine.

novaezealandiae, Lycenchelys Anderson [M. E.] & Møller [P. R.] 2007:176, Figs. 1-2 [Species Diversity v. 12 (no. 3); ref. 29401] Challenger Plateau, 37°23'S, 168°18'E, southwestern Pacific. Holotype: NMNZ P.37713. Paratypes: NMNZ P.12895 (1), P.14772 (1). •Valid as Lycenchelys novaezealandiae Anderson & Møller 2007 -- (Møller & Anderson 2015:1427 [ref. 34289]). Current status: Valid as Lycenchelys novaezealandiae Anderson & Møller 2007. Zoarcidae: Lycodinae. Distribution: New Zealand region. Habitat: marine.

nyx, Bothrocara Stevenson [D. E.] & Anderson [M. E.] 2005:54, Figs. 1-3 [Zootaxa No. 1094; ref. 28569] Bering Sea [North Pacific], 58.512°N, 175.063°W, depth 1085 meters. Holotype: UW 111798. Paratypes: Many lots at CAS, LACM, SAIAB, USNM, and UW. •Valid as Bothrocara nyx Stevenson & Anderson 2005 -- (Anderson et al. 2009:184 [ref. 30175]). •Valid as Bentartia nyx (Stevenson & Anderson 2001) -- (Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Bentartia nyx (Stevenson & Anderson 2005). Zoarcidae: Lycodinae. Distribution: North Pacific: eastern Bering Sea. Habitat: marine.

obesa, Pachycara Zugmayer [E.] 1911:12 [Bulletin de l'Institut Océanographique (Monaco) No. 193; ref. 6161] Bay of Biscay, eastern Atlantic, 45°27'N, 6°05'W, station 1554, depth 4780 meters. Holotype (unique): MOM 0091-0831. Type catalog: Belloc 1949:13-14 [ref. 21544]. Original should have been Pachycara obesum. Species illustrated and described in more detail in Zugmayer 1911:134, Pl. 6 (fig. 6) [ref. 4846]. •Synonym of Pachycara bulbiceps (Garman 1899) -- (Anderson & Peden 1988:84 [ref. 7215], Anderson 1989:227 [ref. 13487], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:34 [ref. 27603]). •Valid as Pachycara obesum Zugmayer 1911 -- (Andriashev 1973:547 [ref. 9214] as obesa, Toyoshima 1985:144 [ref. 5722] as obesa, Andriashev in Whitehead et al. 1986:1149 [ref. 13677] as obesa, Anderson & Almeida 1990:889 [ref. 19328], Anderson 2012:39 [ref. 32270]). Current status: Synonym of Pachycara bulbiceps (Garman 1899). Zoarcidae: Lycodinae. Habitat: marine.

obscurus, Gymnelus Chernova [N. V.] 2000:14 [9], Figs. 6-7 [Voprosy Ikhtiologii v. 40 (no. 1); ref. 24885] Emma Bay, Providence Bay, northern part of Bering Sea. Holotype: ZIN 34840. Paratypes: ZIN 35479 (1), 51560 (1). Type catalog: Balushkin 2011:965 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. Anderson in Anderson & Fedorov 2004:6 [ref. 27603] believes this is probably a synonym of G. viridis. On p. 9 of English translation. •Valid as Gymnelus obscurus Chernova 2000 -- (Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:6 [ref. 27603], Balushkin 2011:965 [ref. 32449]). •Synonym of Gymnelus viridis (Fabricius 1780) with question -- (Mecklenburg et al. 2011:124, 128 [ref. 31212], Parin et al. 2014:393 [ref. 33547] with question, Mecklenburg & Anderson 2015:278 [ref. 33742]). Current status: Synonym of Gymnelus viridis (Fabricius 1780). Zoarcidae: Gymnelinae. Distribution: North Pacific: Bering Sea (if valid). Habitat: marine.

obscurus, Lycodes Toyoshima [M.] 1985:235, Figs. 65-66 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Southern Okhotsk Sea, 44°59.7'N, 144°21.2'E, depth 340 meters. Holotype: HUMZ 76006. Paratypes: HUMZ 92885-87 (3). •Valid as Lycodes obscurus Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1034 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1034 [ref. 26193], Anderson & Fedorov 2004:27 [ref. 27603], Ikeda et al. 2008:360 [ref. 29896]). •Synonym of Lycodes microporus Toyoshima 1983 -- (Balushkin et al. 2011:1001 [ref. 32449], Parin et al. 2014:416 [ref. 33547]). Current status: Synonym of Lycodes microporus Toyoshima 1983. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea (if valid). Habitat: marine.

ocellatus, Gymnelopsis Soldatov [V. K.] 1922:161, Fig. 1 [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23 (no. 2) (for 1917); ref. 4159] Western Okhotsk Sea, about 55°30'N, 138°30'E, about 70-90 meters. Syntypes: ZIN 20167 (now 2). Type catalog: Balushkin et al. 2011:961 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. Published 1922, not 1917. The ending opsis is feminine. •Valid as Gymnelopsis ocellatus Soldatov 1922 -- (Lindberg & Krasyukova 1975:178 [ref. 7348], Anderson 1982:53 [ref. 5520], Toyoshima in Masuda et al. 1984:306 [ref. 6441], Nazarkin & Chernova 2003:605 [ref. 27204]). •Valid as Gymnelopsis ocellata Soldatov 1922 -- (Anderson 1994:116 [ref. 21438], Nakabo 2000:1042 [ref. 25182], Sheiko & Fedorov 2000:35 [ref. 25650], Nakabo 2002:1042 [ref. 26193], Fedorov et al. 2003:111 [ref. 27579], Anderson & Fedorov 2004:4 [ref. 27603], Balushkin et al. 2011:961 [ref. 32449], Parin et al. 2014:390 [ref. 33547]). Current status: Valid as Gymnelopsis ocellata Soldatov 1922. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

ocellatus, Lycodes Toyoshima [M.] 1985:223, Fig. 57 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Off Kushiro, eastern coast of Hokkaido, Japan, depth 600 meters. Holotype: HUMZ 36817. Paratypes: HUMZ 36811 (1), 36815 (1), 36823 (1) 36824 (1, missing in 1997), 36826 (1), 36829 (1), 36830 (1, missing in 1997), 36844 (1). •Valid as Lycodes ocellatus Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1037 [ref. 25182], Nakabo 2002:1037 [ref. 26193], Anderson & Fedorov 2004:27 [ref. 27603], Ikeda et al. 2008:361 [ref. 29896], Balushkin et al. 2011:1002 [ref. 32449]). Current status: Valid as Lycodes ocellatus Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Hokkaido, Japan. Habitat: marine.

ochotensis, Derjuginia Popov [A. M.] 1931:137, Pl. 1 (fig. 1) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] Northern Okhotsk Sea, 56°08'N, 144°55'E, depth 287 meters. Holotype (unique): ZIN 30359. Type catalog: Balushkin 2011:960 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. •Valid as Derjuginia ochotensis Popov 1931 -- (Toyoshima in Masuda et al. 1984:306 [ref. 6441], Nakabo 2000:1042 [ref. 25182], Sheiko & Fedorov 2000:34 [ref. 25650], Nakabo 2002:1042 [ref. 26193], Fedorov et al. 2003:110 [ref. 27579], Balushkin 2011:960 [ref. 32449], Parin et al. 2014:390 [ref. 33547]). •Valid as Gymnelopsis ochotensis (Popov 1931) -- (Anderson 1982:51 [ref. 5520], Anderson 1994:116 [ref. 21438], Nazarkin & Chernova 2003:605 [ref. 27204], Anderson & Fedorov 2004:4 [ref. 27603], Tohkairin et al. 2015:12 [ref. 35331]). Current status: Valid as Gymnelopsis ochotensis (Popov 1931). Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

ochotensis, Gymnelis viridis morpha Soldatov [V. K.] & Lindberg [G. U.] 1930:506 [Izvestiia Tikhookeanskogo nauchnogo instituta rybnogo khoziaistva [Bulletins of the Pacific Science Institute] v. 5; ref. 4164] Okhotsk Sea, northern Pacific. Syntypes: ZIN 20165-66 (1, 2). Called a "morpha" on p. 506 and 507, as a subspecies [podvid] in text on p. 507. •Synonym of Gymnelus hemifasciatus Andriashev 1937 -- (Anderson & Fedorov 2004:5 [ref. 27603]). Current status: Synonym of Gymnelus hemifasciatus Andriashev 1937. Zoarcidae: Gymnelinae. Habitat: marine.

ochotensis, Lycodes brevipes Schmidt [P. Yu.] 1950:89 [76] [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] East coast of Sakhalin, 53°17'N, 144°10'E, Okhotsk Sea, depth 165 meters. Holotype (unique): ZIN 25271. Type catalog: Balushkin et al. 2011:1001 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. On p. 76 of English translation. Appeared first without description as Schmidt in Taranetz 1937:163 [ref. 13384]. •Synonym of Lycodes brevipes Bean 1890, but a valid subspecies -- (Lindberg & Krasyukova 1975:155 [ref. 7348]). •Valid as Lycodes ochotensis Schmidt 1950 -- (Toyoshima 1985:211 [ref. 5722], Sheiko & Fedorov 2000:37 [ref. 25650], Møller 2001:111 [ref. 25378], Fedorov et al. 2003:115 [ref. 27579]). •Synonym of Lycodes brevipes Bean 1890 -- (Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:23 [ref. 27603], Nazarkin 2010:4 [ref. 30733] as Lycodes breviceps ochotensis on p. 2). •Synonym of Lycodes microlepidotus Schmidt 1950 -- (Balushkin et al. 2011:1001 [ref. 32449], Parin et al. 2014:415 [ref. 33547]). Current status: Synonym of Lycodes microlepidotes Schmidt 1950. Zoarcidae: Lycodinae. Habitat: marine.

olasoi, Bellingshausenia Matallanas [J.] 2009:874, Figs. 1-5 [Polar Biology v. 32; ref. 30073] Bellingshausen Sea, 69°49'34"S, 77°49'92"W, Southern Ocean, depth 602 meters. Holotype: UAB B03BSZ14. Paratypes: UAB B03BSZ22 (1), B03BSZ 14B (1), B30BSZ40 (1), B30BSZ41 (1) •Valid as Bellingshausenia olasoi Matallanas 2009 -- (Balushkin 2011:1026 [ref. 32449] as alasoi). Current status: Valid as Bellingshausenia olasoi Matallanas 2009. Zoarcidae: Lycodinae. Distribution: Bellingshausen Sea. Habitat: marine.

olgae, Melanostigma Balushkin [A. V.] & Moganova [M. V.] 2017:132, Fig. on p. 134 [Voprosy Ikhtiologii v. 57 (no. 2); ref. 35279] Southern Ocean, South Sandwich Islands, 55°49'S, 27°44'W, depth 800-850 meters. Holotype: ZIN 56159. Paratypes: ZIN. •Valid as Melanostigma olgae Balushkin & Moganova 2017 -- (Balushkin & Moganova 2018:136 [ref. 35877], Balushkin & Orlovskaya 2019:135 [ref. 36664]). Current status: Valid as Melanostigma olgae Balushkin & Moganova 2017. Zoarcidae: Gymnelinae. Distribution: Southern Ocean, South Sandwich Islands. Habitat: marine.

ophidium, Lycenchelys Jensen [A. S.] 1902:212 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Iceland, 61°33'N, 19°00'W, depth 1089 fathoms. Holotype: ZMUC 32. Type catalog: Nielsen 1974:76 [ref. 9588]. •Valid as Lycodonus ophidium (Jensen 1902) -- (Andriashev 1973:546 [ref. 9214], Toyoshima 1985:143 [ref. 5722], Andriashev in Whitehead et al. 1986:1147 [ref. 13677]). •Synonym of Lycodonus flagellicauda (Jensen 1902) -- (Andriashev 1973:547 [ref. 9214], Anderson 1994:119 [ref. 21438], Koyanagi in Okamura et al. 1995:205 [ref. 22531], Anderson & Fedorov 2004:31 [ref. 27603]). Current status: Synonym of Lycodonus flagellicauda (Jensen 1902). Zoarcidae: Lycodinae. Distribution: Iceland. Habitat: marine.

orientale, Melanostigma Tominaga [Y.] 1971:151, Figs. 1-3 [Japanese Journal of Ichthyology v. 18 (no. 4); ref. 7709] Sagami Bay, Japan, 35°08'N, 139°16'E, depth about 600-0 meters. Holotype: ZUMT 52454. Paratypes: ZUMT 52455 (1, c&s), 52456 [ex NMC 71-208] (1). •Valid as Melanostigma orientale Tominaga 1971 -- (Toyoshima in Masuda et al. 1984:309 [ref. 6441], Anderson 1994:120 [ref. 21438], Nakabo 2000:1040 [ref. 25182], Shinohara et al. 2001:328 [ref. 25995], Nakabo 2002:1040 [ref. 26193], Anderson & Fedorov 2004:8 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449], Balushkin & Moganova 2017:131 [ref. 35279], Balushkin & Orlovskaya 2019:136 [ref. 36664], Balushkin 2019:287 [ref. 36807]). Current status: Valid as Melanostigma orientale Tominaga 1971. Zoarcidae: Gymnelinae. Distribution: Western North Pacific. Habitat: marine.

ornatus, Lycenchelys Soldatov [V. K.] 1922:162, Fig. 2 [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23 (no. 2) (for 1917); ref. 4159] Tatar Strait near Cape Grasevich, northern Japan Sea, depth 10 fathoms. Holotype (unique): ZIN 20168 (destroyed). Published 1922, not 1917. The holotype was destroyed during World War II, but a radiograph from the specimen is on file at ZIN (Anderson 1994:111 [ref. 21438]). •Valid as Bilabria ornatus (Soldatov 1922) -- (Lindberg & Krasyukova 1975:169 [ref. 7348]). •Valid as Bilabria ornata (Soldatov 1922) -- (Toyoshima in Masuda et al. 1984:306 [ref. 6441], Anderson 1994:116 [ref. 21438], Nakabo 2000:1030 [ref. 25182], Sheiko & Fedorov 2000:34 [ref. 25650], Nakabo 2002:1030 [ref. 26193], Anderson & Fedorov 2004:3 [ref. 27603], Anderson & Imamura 2008:63 [ref. 29573], Balushkin et al. 2011:958 [ref. 32449], Shinohara et al. 2014:263 [ref. 33330], Parin et al. 2014:383 [ref. 33547]). Current status: Valid as Bilabria ornata (Soldatov 1922). Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Okhotsk and Japan seas. Habitat: marine.

ovoviviparus, Blennius Lacepède [B. G. E.] 1800:459, 496 [Histoire naturelle des poissons (Lacepède) v. 2; ref. 2709] European seas. No types known. Possible syntypes: BMNH 1853.11.12.122 [Gronovius coll.] (1, skin), LS 28 (left half-skin). Type catalog: Bauchot 1967:18 [ref. 20734]. Unnecessary replacement name for Blennius viviparus Linnaeus 1758. Also appeared in Sonnini 1803:85, 134 (vol. 7) [ref. 30473]. •Nomen dubium in family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). •Synonym of Zoarces viviparus (Linnaeus 1758). Current status: Synonym of Zoarces viviparus (Linnaeus 1758). Zoarcidae: Zoarcinae. Habitat: marine.

paamiuti, Lycodes Møller [P. R.] 2001:977, Figs. 4-5 [Copeia 2001 (no. 4); ref. 25812] Davis Strait, 66°25'N, 57°04'W, Atlantic, depth 678 meters. Holotype: ZMUC P762724. Paratypes: BMNH 1999.7.9.1-2 (2); BSKU 47297 (1), 47551 (1), 80158 (1), 80527 (1), 80530 (1); CNMFI 1999-0020 (4); ISH 804-1964 (2), 807-1964 (12), 147-1973 (3), 149-1973 (3), 6-1993 (10), 7-1994 (5); ROM 72564 (2); USNM 357279 (2); ZIN 51961 (2); ZMH 8421 (7); ZMUC P761327 (1), P761364 (1), P761646 (1), P761664-68 (5), P762215 (1), P762697-98 (2, the larger c&s), P762699-705 (7), P762707-17 (11), P762718-20 (3), P762723 (1), P762725-28 (4), P762730-32 (3), P763053-56 (4), 763057 (1), P763182 (1), P763184 (1), P763185-86 (2), P764086-91 (6, 1c&s). Additional paratypes at ISH, NHMT, ZMUB, ZMUC and ZMUO. Type catalog: Balushkin et al. 2011:1002 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. •Valid as Lycodes paamiuti Møller 2001 -- (Anderson & Fedorov 2004:27 [ref. 27603], Møller et al. 2010:62 [ref. 31169], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin et al. 2011:1002 [ref. 32449], Afonso et al. 2016:65, 205 [ref. 36199], Mecklenburg et al. 2018:340 [ref. 36370]). Current status: Valid as Lycodes paamiuti Møller 2001. Zoarcidae: Lycodinae. Distribution: North Atlantic and Arctic Ocean: Davis Strait to Greenland and Norwegian seas. Habitat: marine.

pachysoma, Lycodapus Peden [A. E.] & Anderson [M. E.] 1978:1946 [1944], Fig. 16 [Canadian Journal of Zoology v. 56 (no. 9) (for Sept. 1978); ref. 8895] North Pacific, 43°23'00"N, 125°18'30"W, depth 2000 meters. Holotype: USNM 216468. Paratypes: CAS 36827-28 (1, 1); NMC 76-0440 (1); UBC 64-444 (2); USNM 216469-70 (1, 1); UW 19256 (1). Type catalog: Springer & Anderson 1997:19-20 [ref. 22953]. On p. 1944 of corrected pages distributed by the authors. •Valid as Lycodapus pachysoma Peder & Anderson 1978 -- (Anderson 1988:106 [ref. 7304], McAllister 1990:158 [ref. 14674], Anderson 1990:266 [ref. 21695], Miller 1993:676 [ref. 21297], Anderson 1994:118 [ref. 21438], Trunov 1999:463 [ref. 24579], Mecklenburg et al. 2002:735 [ref. 25968], Anderson & Fedorov 2004:22 [ref. 27603], Anderson 2006:8 [ref. 28576], Balushkin et al. 2011:986 [ref. 32449]). Current status: Valid as Lycodapus pachysoma Peden & Anderson 1978. Zoarcidae: Lycodinae. Distribution: Eastern Pacific and Southern Ocean: British Columbia to Oregon; Scotia Sea and subantarctic sector of Indian Ocean (Kerguelen Plateau). Habitat: marine.

pacificus, Lycodes Collett [R.] 1879:381, Fig. [Proceedings of the Zoological Society of London 1879 (pt 2) (art. 4); ref. 886] Said to be from Japan (probably from Alaska). Holotype (unique): ZMB 8280. Described in Lycodes, placed in Lycodopsis on p. 382. •Valid as Lycodopsis pacificus (Collett 1879) -- (Toyoshima 1985:238 [ref. 5722], Balushkin et al. 2011:1018 [ref. 32449] as pacifica, Page et al. 2013:163 [ref. 32708]). •Valid as Lycodes pacificus Collett 1879 -- (Eschmeyer & Herald 1983:105 [ref. 9277], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:704 [ref. 25968], Anderson & Fedorov 2004:27 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Pietsch & Orr 2015:58 [ref. 34694], Kells et al. 2016:274 [ref. 35888]). Current status: Valid as Lycodes pacificus Collett 1879. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

palearis, Lycodes Gilbert [C. H.] 1896:454 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bristol Bay, Alaska, U.S.A., Albatross station 3253, 3254, depth 36-46 fathoms. Syntypes: CAS-SU 3165 (1), 3486 (1); USNM 48592 (1). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:20 [ref. 22953]. •Valid as Lycodes palearis Gilbert 1896 -- (Lindberg & Krasyukova 1975:157 [ref. 7348], Eschmeyer & Herald 1983:106 [ref. 9277], Toyoshima 1985:212 [ref. 5722], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:719 [ref. 25968], Møller 2001:111 [ref. 25378], Fedorov et al. 2003:115 [ref. 27579], Anderson & Fedorov 2004:27 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Mecklenburg et al. 2007:182 [ref. 29448], Nazarkin 2010:603 [ref. 31019], Balushkin et al. 2011:1002 [ref. 32449], Page et al. 2013:163 [ref. 32708], Parin et al. 2014:417 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Pietsch & Orr 2015:58 [ref. 34694], Mecklenburg et al. 2016:172 [ref. 34440], Kells et al. 2016:274 [ref. 35888], Mecklenburg et al. 2018:342 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes palearis Gilbert 1896. Zoarcidae: Lycodinae. Distribution: North Pacific and adjacent Arctic. Habitat: marine.

pallida, Lycodes vahlii forma Smitt [F. A.] 1901:24 [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 27 (no. 4); ref. 18732] Not intended as new; this is a subsequent treatment of Lycodes pallidus Collett 1879 as a subspecific name. •In the synonymy of Lycodes vahlii Reinhardt 1831 -- (Anderson 1994:119 [ref. 21438]). •In the synonymy of Lycodes pallidus Collett 1879 -- (Møller 2001:983 [ref. 25812], Carl 2002:65 [ref. 26925]). •In the synonymy of Lycodes vahlii or Lycodes pallidus Collett 1879 -- (Anderson & Fedorov 2004:48 [ref. 27603]). Current status: Synonym of Lycodes. Zoarcidae: Lycodinae.

pallidus, Lycodes Collett [R.] 1879:70 [Forhandlinger i Videnskabs-selskabet i Christiania (for 1878) No. 14; ref. 885] West of Norskøyane, Spitzbergen, depth 475 and 839 meters. Syntypes: ZMUO J4540 (1), J4541 (1). Type catalog: Pethon 1969:12 [ref. 19268]. Also appeared as new in Collett 1880:110, Pl. 3 (figs. 26-27) [ref. 887]. •Valid as Lycodes pallidus Collett 1879 -- (Andriashev 1973:544 [ref. 9214], Andriashev in Whitehead et al. 1986:1141 [ref. 13677], Robins & Ray 1986:237 [ref. 23100], Scott & Scott 1988:409 [ref. 25518], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Saito & Okamura in Okamura et al. 1995:199 [ref. 22531], Mecklenburg et al. 2002:721 [ref. 25968], Møller 2001:982 [ref. 25812], Carl 2002:65 [ref. 26925], Anderson & Fedorov 2004:27 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Møller et al. 2010:62 [ref. 31169], Balushkin et al. 2011:1003 [ref. 32449], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Parin et al. 2014:417 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:65, 206 [ref. 36199], Mecklenburg et al. 2016:174 [ref. 34440] see remarks, Mecklenburg et al. 2018:344 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes pallidus Collett 1879. Zoarcidae: Lycodinae. Distribution: Arctic Ocean: circumpolar, and North Atlantic. Habitat: marine.

pammelas, Melanostigma Gilbert [C. H.] 1896:472, Pl. 35 (lower) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Monterey Bay, California, U.S.A., 36°46'10"N, 121°58'45"W, Albatross station 3202, depth 382 fathoms. Holotype: USNM 53034 (missing). Paratypes: MCZ 28383 [ex USNM] (1); USNM 48599 (1, nearly disintigrated), 53034 (1). Non-type: CAS-SU 4000 (1). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:20 [ref. 22953]. •Valid as Melanostigma pammelas Gilbert 1896 -- (Anderson & Hubbs 1981:341 [ref. 5476], Eschmeyer & Herald 1983:107 [ref. 9277], McAllister 1990:165 [ref. 14674], Anderson 1994:120 [ref. 21438], Mecklenburg et al. 2002:684 [ref. 25968], Anderson & Fedorov 2004:8 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Balushkin et al. 2011:970 [ref. 32449], Page et al. 2013:163 [ref. 32708], Kells et al. 2016:274 [ref. 35888], Balushkin & Moganova 2017:131 [ref. 35279], Balushkin & Orlovskaya 2019:136 [ref. 36664], Balushkin 2019:287 [ref. 36807]). Current status: Valid as Melanostigma pammelas Gilbert 1896. Zoarcidae: Gymnelinae. Distribution: Eastern North Pacific. Habitat: marine.

pammelas, Pachycara Anderson [M. E.] 1989:239, Fig. 14 [Proceedings of the California Academy of Sciences v. 46 (no. 10); ref. 13487] Off Punta Lobos, Chile, 20°50'S, 70°27'W, depth 610 meters. Holotype: CAS 62407. Paratypes: LACM 43726-1 (2), 43732-1 (2); SIO 72-183 (2). •Valid as Pachycara pammelas Anderson 1989 -- (Anderson 1991:158 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:227 [ref. 22911], Pequeño 1997:83 [ref. 23536], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson & Mincarone 2006:25 [ref. 28658], Balushkin et al. 2011:1027 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara pammelas Anderson 1989. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific. Habitat: marine.

panthera, Lycodes knipowitschi Schmidt [P. Yu.] 1950:103 [89], Pl. 8 (fig. 2) [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] Bank northeast of Iona Island, Okhotsk Sea, depth 35-75 meters. Syntypes: ?ZIN 24839 (2), 24824 (2), 24842 (1), 30000 (1). Type catalog: Balushkin et al. 2011:999 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667] with original description dated 1937. On p. 89 of English translation. •Synonym of Lycodes knipowitschi Popov 1931 -- (Toyoshima 1985:195 [ref. 5722], Sheiko & Fedorov 2000:36 [ref. 25650], Balushkin et al. 2011:999 [ref. 32449]). •Synonym of Lycodes mucosus Richardson 1855 -- (Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:27 [ref. 27603], Parin et al. 2014:416 [ref. 33547]). Current status: Synonym of Lycodes mucosus Richardson 1855. Zoarcidae: Lycodinae. Habitat: marine.

parallelus, Embryx Gilbert [C. H.] 1915:360, Pl. 19 (fig. 16) [Proceedings of the United States National Museum v. 48 (no. 2075); ref. 1632] Monterey Bay, California, U.S.A., Albatross station 4514, depth 394-406 fathoms. Holotype (unique): USNM 75818. Type catalog: Springer & Anderson 1997:20 [ref. 22953]. •Synonym of Lycenchelys crotalinus (Gilbert 1890) -- (Anderson 1994:117 [ref. 21438], Anderson 1995:69 [ref. 21928], Anderson & Fedorov 2004:16 [ref. 27603], Parin et al. 2014:398 [ref. 33547]). Current status: Synonym of Lycenchelys crotalinus (Gilbert 1890). Zoarcidae: Lycodinae. Habitat: marine.

parini, Lycenchelys Fedorov [V. V.] 1995:126, Figs. 1-2 [Voprosy Ikhtiologii v. 35 (no. 1); ref. 21957] Kuril-Kamchatka Trench, 44°39'02"N, 149°02'02"E, depth 800 meters. Holotype (unique): ZIN 45796. Type catalog: Balushkin et al. 2011:981 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. •Valid as Lycenchelys parini Fedorov 1995 -- (Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:18 [ref. 27603], Balushkin et al. 2011:981 [ref. 32449], Parin et al. 2014:401 [ref. 33547], Thiel et al. 2018:52 [ref. 35743]). Current status: Valid as Lycenchelys parini Fedorov 1995. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

parviceps, Lycodapus Gilbert [C. H.] 1896:455 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] North of Unalaska, 53°33'50"N, 167°46'50"W, Albatross station 3324, depth 0-109 fathoms. Holotype (unique): USNM 48631. Type catalog: Springer & Anderson 1997:20 [ref. 22953]. •Valid as Lycodapus parviceps Gilbert 1896 -- (Peden & Anderson 1978:1929 [ref. 8895], McAllister 1990:158 [ref. 14674], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:733 [ref. 25968], Anderson & Fedorov 2004:22 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Balushkin et al. 2011:986 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:405 [ref. 33547], Pietsch & Orr 2015:58 [ref. 34694]). Current status: Valid as Lycodapus parviceps Gilbert 1896. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific: Bering Sea to Oregon, U.S.A. Habitat: marine.

patagonica, Maynea Cunningham [R. O.] 1871:472 [The Transactions of the Linnean Society of London v. 27; ref. 992] Otter Islands, Smyth Channel, Magellan Straits. Holotype (unique): BMNH 1869.5.3.25. •Valid as Maynea patagonica Cunningham 1871 -- (Gosztonyi 1977:221 [ref. 6103], Nakamura in Nakamura et al. 1986:236 [ref. 14235], Pequeño 1989:49 [ref. 14125], López et al. 1996:33 [ref. 23681]). •Synonym of Maynea puncta (Jenyns 1842) -- (Anderson 1988:91 [ref. 6021], Anderson & Fedorov 2004:32 [ref. 27603], Balushkin et al. 2011:1019 [ref. 32449]). Current status: Synonym of Maynea puncta (Jenyns 1842). Zoarcidae: Lycodinae. Habitat: marine.

paucidens, Leurynnis Lockington [W. N.] 1879:102 [Mining and Scientific Press. An illustrated journal of mining, popular sciences and general news v. 39 (no. 7) 16 Aug.; ref. 20592] Off San Francisco, California, U.S.A. Lectotype: USNM 23502. Paralectotypes: USNM 336667 [ex USNM 23502] (1). Type catalog: Springer & Anderson 1997:20 [ref. 22953]. Original genus as Leurhynnis as above, Leurynnis in ref. 2815. Also appeared in Lockington 1880:326 [ref. 2815]. Lectotype selected by Springer & Anderson 1997:20 [ref. 22953]. •Synonym of Lycodes pacificus Collett 1879 -- (Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:704 [ref. 25968], Anderson & Fedorov 2004:27 [ref. 27603]). Current status: Synonym of Lycodes pacificus Collett 1879. Zoarcidae: Lycodinae. Habitat: marine.

paucidens, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] in Taranetz 1937:163 [Izvestiia Tikhookeanskogo nauchnogo instituta rybnogo khoziaistva [Bulletins of the Pacific Science Institute] v. 11; ref. 13384] Not available, no distinguishing features. •In the synonymy of Lycodes raridens Taranetz & Andriashev 1937 -- (Anderson 1994:119 [ref. 21438]). •In the synonymy of Lycodes brevicaudus Taranetz & Andriashev 1935 -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Nomen nudum in the family Zoarcidae, genus Lycodes -- (Anderson & Fedorov 2004:48 [ref. 27603], Parin et al. 2014:419 [ref. 33547]). Current status: Synonym of Lycodes raridens Taranetz & Andriashev 1937. Zoarcidae: Lycodinae.

paucilepidotus, Lycodes Toyoshima [M.] 1985:196, Fig. 43 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Northern Okhotsk Sea, 58°33'N, 147°25'E, depth 138 meters. Holotype: HUMZ 58117. Paratypes: HUMZ 55142 (1), 55144 (1), 55159 (1), 57658-59 (2), 57792 (1), 57862 (1), 57865 (1), 57867-69 (3), 58058 (1), 58117 (1), 58222 (1), 58224 (1), 60756 (1), 60814-20 (7), 60822-23 (2), 60844 (1), 60846 (1), 61071 (1), 61458 (1), 61480 (1), 61497 (1), 61506 (1), 61509 (1), 61512 (1), 68063 (1). •Valid as Lycodes paucilepidotus Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1038 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1038 [ref. 26193], Fedorov et al. 2003:116 [ref. 27579], Anderson & Fedorov 2004:28 [ref. 27603], Balushkin et al. 2011:1005 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:418 [ref. 33547]). Current status: Valid as Lycodes paucilepidotus Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea and Sea of Japan. Habitat: marine.

pauciporus, Gymnelus Anderson [M. E.] 1982:47, Figs. 12B, 30 [National Museum of Canada Publications in Zoology No. 17; ref. 5520] South of Cape Ozernoy, Kamchatka Peninsula, Russia, 57°31.0'N, 163°49.5'E, depth 275 meters. Holotype: USNM 92589. Paratypes: ZIN 23945 (1), 23951 (1, missing), 46507 [ex 34498] (1). Type catalog: Springer & Anderson 1997:20 [ref. 22953], Balushkin et al. 2011:965 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. •Valid as Gymnelus pauciporus Anderson 1982 -- (Anderson 1994:117 [ref. 21438], Chernova 1998:743 [ref. 24169], Chernova 2000:7 [ref. 24885], Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:6 [ref. 27603], Balushkin et al. 2011:965 [ref. 32449], Parin et al. 2014:393 [ref. 33547], Mecklenburg & Anderson 2015:263 [ref. 33742]). Current status: Valid as Gymnelus pauciporus Anderson 1982. Zoarcidae: Gymnelinae. Distribution: Western North Pacific. Habitat: marine.

pavlovskii, Krusensterniella Andriashev [A. P.] 1955:393, Fig. 1 [Trudy Zoologicheskogo Instituta, Akademii Nauk SSSR v. 21; ref. 9472] Off Cape Afrika, eastern Kamchatka, western Bering Sea. Holotype: ZIN 33748 (126.5 mm specimen). Paratypes: ZIN 33748 (2) with holotype. Type catalog: Balushkin et al. 2011:969 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. Holotype established in caption to Fig. 1. •Valid as Krusensterniella pavlovskii Andriashev 1955 -- (Anderson 1994:117 [ref. 21438], Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:7 [ref. 27603], Balushkin et al. 2011:969 [ref. 32449], Parin et al. 2014:397 [ref. 33547]). Current status: Valid as Krusensterniella pavlovskii Andriashev 1955. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Kamchatka, Bering Sea. Habitat: marine.

paxilloides, Lycodes Goode [G. B.] & Bean [T. H.] 1883:207 [Bulletin of the Museum of Comparative Zoology v. 10 (no. 5); ref. 1838] Atlantic, 39°50'45"-40°11'40"N, 68°22'00"-70°11'00"W, depth 304-466 fathoms. Syntypes: (3) MCZ 25857 (1), 25858 (1, in 2 pieces). •Synonym of Lycenchelys paxillus (Goode & Bean 1879) -- (Anderson 1994:117 [ref. 21438], Møller 1999:323 [ref. 24052], Anderson & Fedorov 2004:19 [ref. 27603]). Current status: Synonym of Lycenchelys paxillus (Goode & Bean 1879). Zoarcidae: Lycodinae. Habitat: marine.

paxillus, Lycodes Goode [G. B.] & Bean [T. H.] 1879:44 [Proceedings of the United States National Museum v. 2 (no. 64); ref. 18146] Between Le Have and Sable Island Banks, 42°48'N, 63°07'W, depth 1200-2400 feet (stomach content). Holotype (unique): USNM 22177. Type catalog: Springer & Anderson 1997:20-21 [ref. 22953]. •Valid as Lycenchelys paxillus (Goode & Bean 1879) -- (Scott & Scott 1988:405 [ref. 25518], McAllister 1990:158 [ref. 14674] as paxilla, Anderson 1994:117 [ref. 21438], Møller 1999:323 [ref. 24052], Moore et al. 2003:228 [ref. 27076], Anderson & Fedorov 2004:18 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Møller et al. 2010:59 [ref. 31169], Balushkin et al. 2011:981 [ref. 32449], Page et al. 2013:162 [ref. 32708], Afonso et al. 2016:62, 193 [ref. 36199], Mecklenburg et al. 2018:316 [ref. 36370]). Current status: Valid as Lycenchelys paxillus (Goode & Bean 1879). Zoarcidae: Lycodinae. Distribution: Western North Atlantic and adjacent Arctic region. Habitat: marine.

pearcyi, Lycenchelys Anderson [M. E.] 1995:87, Fig. 10 [Proceedings of the California Academy of Sciences v. 49 (no. 2); ref. 21928] Cascadia Abyssal Plain, 45°17'N, 126°38.8'W, depth 2753 meters. Holotype: CAS 81715. Paratypes: CAS 81693 (1), 81706 (1), 81710-11 (1, 2), 81713 (1), 81718 (1); OS 14187 (1), 14377 (1), 14379 (1); SIO 62-366 (1). •Valid as Lycenchelys pearcyi Anderson 1995 -- (Anderson & Fedorov 2004:19 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys pearcyi Anderson 1995. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

pectoralis, Lycodes Toyoshima [M.] 1985:228, Fig. 60 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Southern Okhotsk Sea, 45°37'N, 143°53'E, depth 290-480 meters. Holotype: HUMZ 49083. Paratypes: HUMZ 33951 (1), 33956 (1), 33964 (1), 49071 (1), 49075-77 (3), 49082 (1), 49084-91 (8). •Valid as Lycodes pectoralis Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1029 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:724 [ref. 25968], Nakabo 2002:1029 [ref. 26193], Fedorov et al. 2003:116 [ref. 27579], Anderson & Fedorov 2004:28 [ref. 27603], Shinohara & Shirai 2005:172 [ref. 28232], Kim et al. 2006:275 [ref. 29191], Balushkin et al. 2011:1005 [ref. 32449], Nazarkin at al. 2014:177 [ref. 33213], Saveliev et al. 2014:152 [ref. 33340], Parin et al. 2014:418 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331]). Current status: Valid as Lycodes pectoralis Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea and Kuril Islands (presence in Korea is a misidentification [ref. 33340]). Habitat: marine.

pectoralis, Seleniolycus Møller [P. R.] & Stewart [A. L.] 2006:58, Figs. 2, 3b [Zootaxa No. 1376; ref. 28888] North of Balleny and Scott Islands, 64°40.0'S to 64°42.3'S, 176°32.0'S to 176°44.6'E, Pacific-Antarctic Ridge, Antarctica, depth 1948-2594 meters by longline. Holotype: NMNZ P.40591. Paratypes: NMNZ P.40595 (2), P.42283 (1). •Valid as Seleniolycus pectoralis Møller & Stewart 2006 -- (Struthers & Møller 2009:65 [ref. 30403], Balushkin et al. 2011:971 [ref. 32449]). Current status: Valid as Seleniolycus pectoralis Møller & Stewart 2006. Zoarcidae: Gymnelinae. Distribution: Pacific-Antarctic Ridge, Antarctica. Habitat: marine.

pelophilum, Thermarces Geistdoerfer [P.] 1999:7, Fig. 1 [Cybium v. 23 (no. 1); ref. 23832] Cold seeps of the Barbados accretionary complex, western North Atlantic, 10°19.9'N, 58°37.3'W, depth 1947 meters. Holotype (unique): MNHN 1994-0560. •Valid as Thermarces pelophilum Geistdoerfer 1999 -- (Machida & Hashimoto 2002:1 [ref. 25949], Chernova & Geistdorfer 2003:153 [ref. 26887], Anderson & Fedorov 2004:38 [ref. 27603], Balushkin et al. 2011:1027 [ref. 32449]). Current status: Valid as Thermarces pelophilum Geistdoerfer 1999. Zoarcidae: Lycodinae. Distribution: Western Atlantic. Habitat: marine.

pentactina, Lycenchelys Anderson [M. E.] 1995:90, Fig. 11 [Proceedings of the California Academy of Sciences v. 49 (no. 2); ref. 21928] Off Pta. Santa Maria, Peru, 14°44'S, 76°12'W, depth 1495-1430 meters. Holotype (unique): ZIN 48392. Type catalog: Balushkin et al. 2011:981 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. •Valid as Lycenchelys pentactina Anderson 1995 -- (Anderson & Fedorov 2004:19 [ref. 27603], Balushkin et al. 2011:981 [ref. 32449]). Current status: Valid as Lycenchelys pentactina Anderson 1995. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific: Peru. Habitat: marine.

pequenoi, Lycenchelys Anderson [M. E.] 1995:91, Fig. 12 [Proceedings of the California Academy of Sciences v. 49 (no. 2); ref. 21928] West of Lobos de Tierra Island, Peru, 6°26'S, 80°05'W, depth 1025 meters. Holotype: LACM 44132-1. Paratypes: CAS 60243 (1), SIO 72-184 (2). •Valid as Lycenchelys pequenoi Anderson 1995 -- (Pequeño 1997:83 [ref. 23536], Anderson & Fedorov 2004:19 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys pequenoi Anderson 1995. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific: Peru and Chile. Habitat: marine.

perspicillum, Lycodes Krøyer [H. N.] 1845:140 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1844 (no. 8); ref. 19477] Newfoundland, Canada. Syntypes: ZMUC 23 (1) Greenland, ZMUC (1 lost). Type catalog: Nielsen 1974:77 [ref. 9588]. •In part a synonym of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603], Parin et al. 2014:419 [ref. 33547]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Habitat: marine.

peruana, Lycenchelys Anderson [M. E.] 1995:93, Fig. 13 [Proceedings of the California Academy of Sciences v. 49 (no. 2); ref. 21928] Off Trujillo, Peru, 7°59'S, 80°37'W, depth 991-1015 meters. Holotype: LACM 45998-1. Paratypes: LACM 44134-1 (1). •Valid as Lycenchelys peruana Anderson 1995 -- (Anderson & Fedorov 2004:19 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys peruana Anderson 1995. Zoarcidae: Lycodinae. Distribution: Eastern South Pacific: Peru. Habitat: marine.

pictus, Gymnelus Günther [A.] 1862:324 [Catalogue of the fishes in the British Museum v. 4; ref. 1969] No locality. Syntypes: (2) BMNH 1855.9.19.1298 (1). •Synonym of Gymnelus viridis (Fabricius 1780) -- (Andriashev 1973:541 [ref. 9214] with question, Anderson 1982:30 [ref. 5520], Anderson & Fedorov 2004:6 [ref. 27603]). Current status: Synonym of Gymnelus viridis (Fabricius 1780). Zoarcidae: Gymnelinae. Habitat: marine.

platei, Lycodes (Phucocoetus) Steindachner [F.] 1898:320, Pl. 19 (fig. 8) [Zoologische Jahrbücher. Jena, Supplementband v. 4; ref. 4238] Cape Espíritu Santo, Tierra del Fuego, Chile. Holotype (unique): ZMB 15691. •Also a synonym of Phucocoetes latitans Jenyns 1842 -- (Anderson 1994:120 [ref. 21438]). •Synonym of Austrolycus laticinctus (Berg 1895) -- (Gosztonyi 1977:202 [ref. 6103], Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:10 [ref. 27603]). Current status: Synonym of Austrolycus laticinctus (Berg 1895). Zoarcidae: Lycodinae. Habitat: marine.

platycephalus, Gymnelus Chernova [N. V.] 1999:313, Figs. 8-9 [Voprosy Ikhtiologii v. 39 (no. 3); ref. 24868] Northern part of Bering Sea. Holotype: ZIN 51564. Paratypes: ZIN 34989 (3), 47798 (3). Plus non-type material. Type catalog: Balushkin et al. 2011:966 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. Anderson in Anderson & Fedorov 2004:6 [ref. 27603] believes this is probably a synonym of G. viridis. •Valid as Gymnelus platycephalus Chernova 1999 -- (Chernova 2000:8 [ref. 24885], Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:6 [ref. 27603], Balushkin et al. 2011:966 [ref. 32449]). •Synonym of Gymnelus hemifasciatus Andriasahev 1937 -- (Mecklenburg et al. 2011:124, 128 [ref. 31212], Parin et al. 2014:392 [ref. 33547]). •Synonym of Gymnelus viridis (Fabricius 1780) -- (Mecklenburg & Anderson 2015:278 [ref. 33742]). Current status: Synonym of Gymnelus viridis (Fabricius 1780). Zoarcidae: Gymnelinae. Distribution: North Pacific and Arctic: Bering and Chukchi seas [if valid]. Habitat: marine.

platyrhinus, Lycodes Jensen [A. S.] 1902:208 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Jan Mayen to Iceland, 67°53'N, 10°19'W, depth 1010 fathoms. Holotype: ZMUC 119. Type catalog: Nielsen 1974:78 [ref. 9588]. •Valid as Lycenchelys platyrhinus (Jensen 1902) -- (Andriashev 1973:542 [ref. 9214], Andriashev in Whitehead et al. 1986:1134 [ref. 13677]). •Valid as Lycenchelys platyrhina (Jensen 1902) -- (Anderson 1994:117 [ref. 21438], Møller 1999:327 [ref. 24052], Anderson & Fedorov 2004:19 [ref. 27603], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin et al. 2011:1026 [ref. 32449], Parin et al. 2014:401 [ref. 33547], Mecklenburg et al. 2018:318 [ref. 36370]). Current status: Valid as Lycenchelys platyrhina (Jensen 1902). Zoarcidae: Lycodinae. Distribution: Arctic. Habitat: marine.

pliciferus, Lycenchelys Andriashev [A. P.] 1955:372, Fig. 2 (no. 7), Figs. 15-16 [Trudy Instituta Zoologii / Akademiia Nauk, Azerbaidzhanskoi SSR. v. 18; ref. 12261] Northeast of Mednyi Island, southwestern part of central depression of the Bering Sea, about 56°55'N, 174°20'E, depth 3820-3830 meters. Holotype: ZIN 32961 (130 mm). Paratypes: ZIN 32961A (1). Type catalog: Balushkin et al. 2011:981 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. •Valid as Lycenchelys pliciferus Andriashev 1955 -- (Toyoshima 1985:155 [ref. 5722]). •Valid as Lycenchelys plicifera Andriashev 1955 -- (Anderson 1994:117 [ref. 21438], Anderson 1995:94 [ref. 21928], Anderson & Balanov 2000:1056 [ref. 25023], Sheiko & Fedorov 2000:35 [ref. 25650], Mecklenburg et al. 2002:696 [ref. 25968], Anderson & Imamura 2002:357 [ref. 26499], Anderson & Fedorov 2004:19 [ref. 27603], Balushkin et al. 2011:981 [ref. 32449], Parin et al. 2014:401 [ref. 33547], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys plicifera Andriashev 1955. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

poecilimon, Lycenchelys Jordan [D. S.] & Fowler [H. W.] 1902:748, Fig. 2 [Proceedings of the United States National Museum v. 25 (no. 1303); ref. 2459] Off Kinkwazan in Matsushima Bay, Japan, Albatross station 3768, depth 25-27 fathoms. Holotype: USNM 50578. Paratypes: CAS-SU 7933 [ex CAS-SU 7121] (1). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:21 [ref. 22953]. •Valid as Davidijordania poecilimon (Jordan & Fowler 1902) -- (Lindberg & Krasyukova 1975:174 [ref. 7348] as Davidojordania, Toyoshima in Masuda et al. 1984:305 [ref. 6441], Anderson 1994:116 [ref. 21438], Shinohara et al. 1996:180 [ref. 26605], Sokolovskaya et al. 1998:12 [ref. 24670] as Davidojordania, Nakabo 2000:1036 [ref. 25182], Nakabo 2002:1036 [ref. 26193], Youn 2002:375, 639 [ref. 26218], Anderson & Fedorov 2004:4 [ref. 27603], Kim et al. 2006:278 [ref. 29191], Anderson & Imamura 2008:66 [ref. 29573], Shinohara & Kim 2009:231 [ref. 31427], Shinohara et al. 2011:51 [ref. 31715] as Davidijordani, Shinohara et al. 2014:264 [ref. 33330], Parin et al. 2014:388 [ref. 33547]). Current status: Valid as Davidijordania poecilimon (Jordan & Fowler 1902). Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Japan and Okhotsk Sea. Habitat: marine.

poecilus, Lycodapus Peden [A. E.] & Anderson [M. E.] 1981:673, Fig. 6 [Canadian Journal of Zoology v. 59 (no. 4); ref. 5531] Bering Sea, 58°32'02"N, 176°06'21"W, depth 497-539 meters. Holotype: USNM 222663. Paratypes: BCPM 980-24 (1), 980-595 (22); CAS 46715 (2); HUMZ 83800 (1), 83814-15 (1, 1), 86650 (1), 86746 (1), 86764 (1); NMC 80-1037 (5); USNM 221053 (2), 221060-64 (5, 11, 5, 1, 3). Type catalog: Springer & Anderson 1997:21 [ref. 22953]. •Valid as Lycodapus poecilus Peden & Anderson 1981 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:741 [ref. 25968], Anderson & Fedorov 2004:22 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449], Parin et al. 2014:406 [ref. 33547]). Current status: Valid as Lycodapus poecilus Peden & Anderson 1981. Zoarcidae: Lycodinae. Distribution: North Pacific: Bering and Okhotsk seas. Habitat: marine.

polaris, Blennius Sabine [E.] 1824:212 [ccxii] [A supplement to the appendix of Captain Parry's voyage; ref. 18649] North Georgia, Boothia Peninsula, Northwest Territories, Canada. No types known. •Valid as Lycodes polaris (Sabine 1824) -- (Andriashev 1973:545 [ref. 9214], Andriashev in Whitehead et al. 1986:1142 [ref. 13677], Scott & Scott 1988:409 [ref. 25518], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:710 [ref. 25968], Anderson & Fedorov 2004:28 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Møller et al. 2010:62 [ref. 31169], Balushkin et al. 2011:1006 [ref. 32449], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:144, 145 [ref. 33463], Parin et al. 2014:418 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:66, 207 [ref. 36199], Kells et al. 2016:274 [ref. 35888], Mecklenburg et al. 2016:176 [ref. 34440] see remarks, Mecklenburg et al. 2018:346 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes polaris (Sabine 1824). Zoarcidae: Lycodinae. Distribution: Arctic Ocean, North Atlantic and North Pacific: nearly circumpolar. Habitat: marine.

polylepis, Ophthalmolycus Matallanas [J.] 2011:564, Fig. 3 [Journal of the Marine Biological Association of the United Kingdom v. 91 (no. 2); ref. 30839] Baited trap, 64°19'S, 61°58'W, Gerlache Strait, Southern Ocean, depth 1056 meters. Holotype: Uncat. •Valid as Ophthalmolycus polylepis Matallanas 2011. Current status: Valid as Ophthalmolycus polylepis Matallanas 2011. Zoarcidae: Lycodinae. Distribution: Gerlache Strait, Southern Ocean. Habitat: marine.

polyodon, Lycenchelys Anderson [M. E.] & Møller [P. R.] 2007:180, Figs. 3-4 [Species Diversity v. 12 (no. 3); ref. 29401] Lord Howe Rise, 34°14.81'S, 163°17.77'E, depth 1076-1083 meters. Holotype (unique): NMNZ P.39203. •Valid as Lycenchelys polyodon Anderson & Møller 2007. Current status: Valid as Lycenchelys polyodon Anderson & Møller 2007. Zoarcidae: Lycodinae. Distribution: Southwestern Pacific. Habitat: marine.

popovi, Commandorella Taranetz [A. Ya.] & Andriashev [A. P.] 1935:268, Figs. 1-2 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 1 (no. 4); ref. 4341] Medny Island, Komander Islands, western Bering Sea. Holotype: ZIN 23948. Paratypes: (4) ZIN 23950 (1), 29992 (1). Type catalog: Balushkin et al. 2011:958 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. •Valid as Commandorella popovi Taranetz & Andriashev 1935 -- (Sheiko & Fedorov 2000:34 [ref. 25650], Balushkin et al. 2011:958 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:387 [ref. 33547], Kells et al. 2016:350 [ref. 35888]). •Valid as Gymnelus popovi (Taranetz & Andriashev 1935) -- (Anderson 1982:44 [ref. 5520], Anderson 1994:117 [ref. 21438], Mecklenburg et al. 2002:688 [ref. 25968], Anderson & Fedorov 2004:6 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Mecklenburg & Anderson 2015:263 [ref. 33742]). Current status: Valid as Gymnelus popovi (Taranetz & Andriashev 1935). Zoarcidae: Gymnelinae. Distribution: North Pacific. Habitat: marine.

porifer, Lycodes Gilbert [C. H.] 1890:104 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off Baja California, Gulf of California, Mexico, 27°09'00"N, 111°42'00"W, Albatross station 3009, depth 857 fathoms. Holotype (unique): USNM 44384. Type catalog: Springer & Anderson 1997:21 [ref. 22953]. •Valid as Lycenchelys porifer (Gilbert 1890) -- (Anderson 1994:117 [ref. 21438], Anderson 1995:69 [ref. 21928], Anderson & Fedorov 2004:19 [ref. 27603], Balushkin et al. 2011:981 [ref. 32449], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys porifer (Gilbert 1890). Zoarcidae: Lycodinae. Distribution: Eastern Pacific: Gulf of California to Peru. Habitat: marine.

priedei, Pachycara Møller [P. R.] & King [N.] 2007:586, Fig. 2 [Copeia 2007 (no. 3); ref. 29259] Crozet Plateau, 45°40'S, 56°35'E, southern Indian Ocean, depth 4269-4275 meters. Holotype: ZMUC P766466. Paratypes: BMNH 2006.7.3.1 (1); ZMUC P766467-69 (1, 1, 1). •Valid as Pachycara priedei Møller & King 2007 -- (Shinohara 2012:55 [ref. 32102]). Current status: Valid as Pachycara priedei Møller & King 2007. Zoarcidae: Lycodinae. Distribution: Crozet Plateau, southern Indian Ocean. Habitat: marine.

psarostomatus, Lycodapus Peden [A. E.] & Anderson [M. E.] 1981:668, Fig. 4 [Canadian Journal of Zoology v. 59 (no. 4); ref. 5531] Bering Sea, 59°53.83'N, 178°57.12'W, depth 564-580 meters. Holotype: USNM 221057. Paratypes: USNM 221052 (1), 221056 (1), 221058 (10); UW 20804 (1). Type catalog: Springer & Anderson 1997:21 [ref. 22953]. •Valid as Lycodapus psarostomatus Peden & Anderson 1981 -- (Anderson 1989:149 [ref. 13419], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:734 [ref. 25968], Anderson & Fedorov 2004:22 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Balushkin et al. 2011:1026 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:406 [ref. 33547]). Current status: Valid as Lycodapus psarostomatus Peden & Anderson 1981. Zoarcidae: Lycodinae. Distribution: Eastern North Pacific. Habitat: marine.

pterygalces, Barbapellis Iglésias [S. P.], Dettai [A.] & Ozouf-Costaz [C.] 2012:216, Figs. 2-3 [Polar Biology v. 35; ref. 31445] Off Terre Adélie, eastern Antarctic continental slope, depth about 1200 meters. Holotype (unique): MNHN 2009-0020. Appeared first in electronic form in 2011; published in an available way in 2012. •Valid as Barbapellis pterygalces Iglésias, Dettai & Ozouf-Costaz 2012. Current status: Valid as Barbapellis pterygalces Iglésias, Dettai & Ozouf-Costaz 2012. Zoarcidae: Gymnelinae. Distribution: Off Terre Adélie, eastern Antarctic continental slope. Habitat: marine.

punctus, Conger Jenyns [L.] 1842:143 [The zoology of the voyage of H. M. S. Beagle; ref. 2344] Beagle Channel, Tierra del Fuego. Holotype (unique): BMNH 1917.7.14.12. •Valid as Maynea puncta (Jenyns 1842) -- (Anderson 1988:91 [ref. 6021], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:32 [ref. 27603], Balushkin et al. 2011:1019 [ref. 32449]). Current status: Valid as Maynea puncta (Jenyns 1842). Zoarcidae: Lycodinae. Distribution: Eastern South Pacific to western South Atlantic: southern South America. Habitat: marine.

pusilla, Maynea Bean [T. H.] 1890:39 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Alaska, U.S.A., 55°10'00"N, 160°18'00"W, Albatross station 2848, depth 110 fathoms. Holotype (unique): USNM 45360 (in pieces). Type catalog: Springer & Anderson 1997:21 [ref. 22953] with figure of holotype. •Valid as Bothrocara pusillum (Bean 1890) -- (McAllister 1990:157 [ref. 14674], Anderson 1994:116 [ref. 21438], Mecklenburg et al. 2002:730 [ref. 25968], Anderson & Fedorov 2004:11 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Stevenson & Anderson 2005:53 [ref. 28569], Anderson et al. 2009:186 [ref. 30175], Page et al. 2013:162 [ref. 32708], Kells et al. 2016:270 [ref. 35888]). •Valid as Bentartia pusilla (Bean 1890) -- (Balushkin et al. 2011:1026 [ref. 32449], Robertson et al. 2017:91 [ref. 35676]). Current status: Valid as Bentartia pusillum (Bean 1890). Zoarcidae: Lycodinae. Distribution: Eastern North Pacific: northern Bering Sea to southern British Columbia (Canada) and Gulf of Panama. Habitat: marine.

raridens, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] in Andriashev 1937:335, Figs. 13, 14 [Issled. Morey SSSR, Leningrad v. 25; ref. 20242] Bering Sea, Gulf of Anadry, 61°52'N, 173°11'W, depth 66 meters. Holotype: ZIN 25070. Paratypes: ZIN 13009 (1), 23757 (1), 24845 (6), 25071 (1), 29999 (2), 30007 (2), 30008 (1), 30010 (2). Type catalog: Balushkin et al. 2011:1008 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. •Questionably a synonym of Lycodes brevicaudus Taranetz & Andriashev 1935 -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Valid as Lycodes raridens Taranetz & Andriashev 1937 -- (Lindberg & Krasyukova 1975:161 [ref. 7348], Toyoshima 1985:198 [ref. 5722], Scott & Scott 1988:408 [ref. 25518], Anderson 1994:119 [ref. 21438], Chikilev & Kharitonov 2000:100 [ref. 24916], Mecklenburg et al. 2002:711 [ref. 25968], Fedorov et al. 2003:116 [ref. 27579], Anderson & Fedorov 2004:28 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Mecklenburg et al. 2007:182 [ref. 29448], Mecklenburg et al. 2011:128 [ref. 31212], Balushkin et al. 2011:1008 [ref. 32449], Page et al. 2013:163 [ref. 32708], Parin et al. 2014:419 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Mecklenburg et al. 2016:178 [ref. 34440] see remarks, Kells et al. 2016:274 [ref. 35888], Mecklenburg et al. 2018:348 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes raridens Taranetz & Andriashev 1937. Zoarcidae: Lycodinae. Distribution: Arctic and adjacent North Pacific. Habitat: marine.

rassi, Lycenchelys Andriashev [A. P.] 1955:359, Fig. 2 (no. 2), Figs. 5-6 [Trudy Instituta Zoologii / Akademiia Nauk, Azerbaidzhanskoi SSR. v. 18; ref. 12261] East of North Sakhalin, Okhotsk Sea, 54°28'N, 145°21.6'E, depth 1500 meters. Holotype (unique): ZIN 32962. Type catalog: Balushkin et al. 2011:981 [ref. 32449], Balushkin et al. 2012:164 [ref. 33667]. •Valid as Lycenchelys rassi Andriashev 1955 -- (Lindberg & Krasyukova 1975:164 [ref. 7348], Toyoshima in Amaoka et al. 1983:269, 332 [ref. 8379], Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:173 [ref. 5722], Anderson 1994:117 [ref. 21438], Anderson 1995:98 [ref. 21928], Nakabo 2000:1033 [ref. 25182], Sheiko & Fedorov 2000:35 [ref. 25650], Mecklenburg et al. 2002:703 [ref. 25968], Nakabo 2002:1033 [ref. 26193], Anderson & Fedorov 2004:19 [ref. 27603], Shinohara & Anderson 2007:64 [ref. 29111], Balushkin et al. 2011:981 [ref. 32449], Parin et al. 2014:402 [ref. 33547], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys rassi Andriashev 1955. Zoarcidae: Lycodinae. Distribution: North Pacific: Bering Sea to Okhotsk Sea. Habitat: marine.

ratmanovi, Lycenchelys Andriashev [A. P.] 1955:355, Fig. 2 (no. 1), Figs. 3-4 [Trudy Instituta Zoologii / Akademiia Nauk, Azerbaidzhanskoi SSR. v. 18; ref. 12261] Avachinsk Bay, southeastern Kamchatka, 52°40.8'N, 159°13'E, depth 800-1000 meters. Holotype: ZIN 32957 [ex ZIN 30012]. Paratypes: ZIN 32957a [ex ZIN 30012] (1). Type catalog: Balushkin et al. 2011:981 [ref. 32449], Balushkin et al. 2012:164 [ref. 33667]. •Valid as Lycenchelys ratmanovi Andriashev 1955 -- (Toyoshima 1985:166 [ref. 5722], Anderson 1994:117 [ref. 21438], Anderson 1995:100 [ref. 21928], Anderson & Balanov 2000:1056 [ref. 25023], Sheiko & Fedorov 2000:35 [ref. 25650], Mecklenburg et al. 2002:702 [ref. 25968], Anderson & Fedorov 2004:20 [ref. 27603], Balushkin et al. 2011:981 [ref. 32449], Parin et al. 2014:402 [ref. 33547], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys ratmanovi Andriashev 1955. Zoarcidae: Lycodinae. Distribution: North Pacific: Bering Sea to northern Kuril Islands. Habitat: marine.

regani, Lycozoarces Popov [A. M.] 1933:151, Fig. 1 [labeled 2 on plate] [Issledovaniya Morey SSSR, Leningrad No. 19; ref. 6535] Tatar Strait, 50°36.7'N, 140°53.6'E, Russia, northern Sea of Japan, depth 70 meters. Holotype (unique): ZIN 29987. Type catalog: Balushkin 2011:951 [ref. 32449], Balushkin et al. 2012:164 [ref. 33667]. Lycozoarces regani is available as above, even though the genus is unavailable until 1935 (Article 11(h)(iii)), so hubbsi should be treated as a synonym of regani. •Wrongly as unavailable and in the synonymy of Lycozoarces hubbsi Popov 1935 -- (Anderson 1994:22 [ref. 21438]). •Valid as Lycozoarces regani Popov 1933 -- (Lindberg & Krasyukova 1975:124 [ref. 7348], Toyoshima in Masuda et al. 1984:305 [ref. 6441], Nakabo 2000:1028 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1028 [ref. 26193], Fedorov et al. 2003:118 [ref. 27579], Anderson & Fedorov 2004:2 [ref. 27603], Balushkin et al. 2011:952 [ref. 32449], Parin et al. 2014:425 [ref. 33547]). Current status: Valid as Lycozoarces regani Popov 1933. Zoarcidae: Lycozoarcinae. Distribution: Western North Pacific: Japan and Okhotsk seas. Habitat: marine.

remigera, Bothrocara Gilbert [C. H.] 1915:366, Pl. 20 (fig. 19) [Proceedings of the United States National Museum v. 48 (no. 2075); ref. 1632] Monterey Bay, California, U.S.A., Albatross station 4516, depth 718-756 fathoms. Holotype: USNM 75820. Paratypes: CAS-SU 22982 (1), 22987 (1), 69828 [ex 22982] (2). Additional material: USNM 46709-10 (2, 2). Type catalog: Böhlke 1953:102 [ref. 12291], Springer & Anderson 1997:22 [ref. 22953]. Original should have been Bothrocara remigerum. All paratypes except SU 22982 (1 specimen) are B. brunneum -- (Anderson et al. 2009:184 [ref. 30175]). •Valid as Bothrocara remigera Gilbert 1915 -- (McAllister 1990:157 [ref. 14674]). •Synonym of Bothrocara molle Bean 1890 -- (Anderson 1994:116 [ref. 21438], Mecklenburg et al. 2002:732 [ref. 25968], Anderson & Fedorov 2004:11 [ref. 27603], Anderson 2006:2 [ref. 28576], Anderson et al. 2009:179 [ref. 30175], Balushkin et al.2011:973 [ref. 32499], Parin et al. 2014:385 [ref. 33547]). Current status: Synonym of Bothrocara molle Bean 1890. Zoarcidae: Lycodinae. Habitat: marine.

remissaria, Lycenchelys Fedorov [V. V.] 1995:130, Figs. 1-2 [Voprosy Ikhtiologii v. 35 (no. 1); ref. 21958] Off Japan, 36°54'04"N, 141°45'07"E, depth 1080 meters. Holotype (unique): ZIN 50586. Type catalog: Balushkin et al. 2011:982 [ref. 32449], Balushkin et al. 2012:164 [ref. 33667]. •Valid as Lycenchelys remissaria Fedorov 1995 -- (Shinohara et al. 1996:181 [ref. 26605], Nakabo 2000:1032 [ref. 25182], Nakabo 2002:1032 [ref. 26193], Anderson & Imamura 2002:357 [ref. 26499], Anderson & Fedorov 2004:20 [ref. 27603], Shinohara & Anderson 2007:63 [ref. 29111], Balushkin et al. 2011:982 [ref. 32449], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys remissaria Fedorov 1995. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Japan. Habitat: marine.

reticulatus, Lycodes Reinhardt [J. C. H.] 1835:7 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1834-35; ref. 3695] Greenland. Syntypes: ZMUC 17 (1) Umanak, ZMUC 21 (1) Fiskenaesset. Type catalog: Nielsen 1974:78 [ref. 9588]. Also appeared in Reinhardt 1837:115, 195, Pl. 6 [33, 113, Pl. 6 of separate] [ref. 3691]. •Valid as Lycodes reticulatus Reinhardt 1835 -- (Andriashev 1973:545 [ref. 9214], Andriashev in Whitehead et al. 1986:1143 [ref. 13677], Robins & Ray 1986:237 [ref. 23100], Scott & Scott 1988:410 [ref. 25518], McAllister 1990:161 [ref. 14674], Saito & Okamura in Okamura et al. 1995:198 [ref. 22531], Mecklenburg et al. 2002:712 [ref. 25968], Anderson & Fedorov 2004:28 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Møller et al. 2010:62 [ref. 31169], Mecklenburg et al. 2011:128 [ref. 31212] with comments, Balushkin et al. 2011:1009 [ref. 32449], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:419 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:66, 208 [ref. 36199], Mecklenburg et al. 2016:180 [ref. 34440] see remarks, Mecklenburg et al. 2018:350 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Distribution: North Atlantic and Arctic Ocean. Habitat: marine.

retrodorsalis, Gymnelis Le Danois [E.] 1913:258 [Bulletin de la Société Zoologique de France v. 38; ref. 14951] Denmark Strait, 65°52'N, 29°20'W, depth 335-346 meters. Neotype: NMC 79-716. Neotype designated by Anderson 1982:35 [ref. 5520]. Original genus should have been spelled Gymnelus. Also appeared as new in Le Danois 1913:429 [ref. 31503]. •Valid as Gymnelus retrodorsalis Le Danois 1913 -- (Andriashev 1973:541 [ref. 9214], McAllister et al. 1981:825 [ref. 5431], Anderson 1982:35 [ref. 5520], Andriashev in Whitehead et al. 1986:1131 [ref. 13677], Scott & Scott 1988:403 [ref. 25518], McAllister 1990:165 [ref. 14674], Anderson 1994:117 [ref. 21438], Koyanagi in Okamura et al. 1995:188 [ref. 22531], Chernova 1998:743 [ref. 24169], Chernova 1999:306 [ref. 24868], Chernova 2000:8 [ref. 24885], Anderson & Fedorov 2004:6 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Møller et al. 2010:58 [ref. 31169], Balushkin et al. 2011:966 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:393 [ref. 33547], Mecklenburg & Anderson 2015:263 [ref. 33742], Afonso et al. 2016:61, 161 [ref. 36199], Mecklenburg et al. 2018:309 [ref. 36370] see remarks). Current status: Valid as Gymnelus retrodorsalis Le Danois 1913. Zoarcidae: Gymnelinae. Distribution: North Atlantic and Arctic Ocean. Habitat: marine.

rictolata, Bothrocaropsis Garman [S.] 1899:128, Pl. 33 (fig. 1) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Off Marinato Point, Panama Bay, 7°06'15"N, 80°34'W, Albatross station 3353, depth 695 fathoms. Holotype (unique): MCZ 28693. •Status uncertain, possibly a synonym of Bothrocara alalongum (Garman 1899) -- (Anderson 1994:116 [ref. 21438]). •Synonym of Bothrocara brunneum (Bean 1890) -- (Anderson & Fedorov 2004:10 [ref. 27603], Anderson et al. 2009:173 [ref. 30175]). Current status: Synonym of Bothrocara brunneum (Bean 1890). Zoarcidae: Lycodinae. Habitat: marine.

rimae, Pachycara Anderson [M. E.] 1989:235, Figs. 11, 12 [Proceedings of the California Academy of Sciences v. 46 (no. 10); ref. 13487] Galápagos Rift hydrothermal vents, 0°47.8'N, 86°09.9'W, depth 2500 meters. Holotype (unique): LACM 44699-1. •Valid as Pachycara rimae Anderson 1989 -- (Anderson 1991:157 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:226 [ref. 22911], Møller & Anderson 2000:65 [ref. 24495], Møller 2003:364 [ref. 26908], Anderson & Fedorov 2004:35 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson & Mincarone 2006:25 [ref. 28658], Møller & King 2007:591 [ref. 29259], Shinohara 2012:55 [ref. 32102], Balushkin et al. 2011:1027 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448], Anderson et al. 2016:76 [ref. 34148]). Current status: Valid as Pachycara rimae Anderson 1989. Zoarcidae: Lycodinae. Distribution: Eastern Pacific: Galápagos Rift hydrothermal vents. Habitat: marine.

ringueleti, Piedrabuenia Gosztonyi [A. E.] 1977:236, Figs. 19-21 [Archiv für Fischereiwissenschaft v. 27 (no. 3); ref. 6103] Continental slope off Puerto Deseado, central Patagonia, Argentina, 46°54'S, 60°28'W, depth 480 meters. Holotype: ISH 279-1971. Paratypes: ISH 369-1971 (1). Type catalog: Krefft 1978:15 [ref. 9884]. •Valid as Piedrabuenia ringueleti Gosztonyi 1977 -- (Anderson & Gosztonyi 1991:13 [ref. 19348], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:36 [ref. 27603], Balushkin et al. 2011:1022 [ref. 32449]). Current status: Valid as Piedrabuenia ringueleti Gosztonyi 1977. Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

robertsi, Seleniolycus Møller [P. R.] & Stewart [A. L.] 2006:55, Figs. 1, 3a [Zootaxa No. 1376; ref. 28888] North of Balleny and Scott Islands, 63°02.0'S, 172°04.0'E, outer Ross Dependency, Pacific-Antarctic Ridge, Antarctic, depth 2290 meters by longline. Holotype: NMNZ P.37815. Paratypes: BMNH 2006.6.21.1 (1); NMNZ P.38574 (1), P.38804 (1), P.39917 (1), P.38819 (1); ZMUC P766465 (1). •Valid as Seleniolycus robertsi Møller & Stewart 2006 -- (Struthers & Møller 2009:65 [ref. 30403]). Current status: Valid as Seleniolycus robertsi Møller & Stewart 2006. Zoarcidae: Gymnelinae. Distribution: Southernmost section of the Pacific-Antarctic Ridge, Antarctica. Habitat: marine.

roseus, Lycenchelys Toyoshima [M.] 1985:152, Figs. 10-11, 31b [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Bering Sea, 52°49.13'N, 171°01.91'W, depth 750 meters. Holotype: HUMZ 88487. Paratypes: HUMZ 89341 (1). •Valid as Lycenchelys rosea Toyoshima 1985 -- (Anderson 1994:118 [ref. 21438], Anderson 1995:103 [ref. 21928], Shinohara & Matsuura 1998:154 [ref. 23382], Mecklenburg et al. 2002:698 [ref. 25968], Anderson & Fedorov 2004:20 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449], Parin et al. 2014:402 [ref. 33547], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys rosea Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Bering Sea. Habitat: marine.

rossi, Lycodes Malmgren [A. J.] 1865:516 [Öfversigt af Kongliga Vetenskaps-Akademiens förhandlingar, Kungliga Svenska Vetenskapsakademien. 1864 (no. 10) [v. 21]; ref. 17598] Treurenberg Bay, Spitzbergen, 5 fathoms and other localities. Syntypes: ?BMNH; NRM 9398 (1, juvenile), NRM not found (1). Based in part Blennius polaris of J.C. Ross, Append. to Parry's Attempt to reach the North-pole, 1827, p. 200. Sometimes as rossii but rossi is correct. •Synonym of Lycodes reticulatus Reinhardt 19835 with question -- (Meckenburg et al. 2011:127 [ref. 31212]). •Valid as Lycodes rossi Malmgren 1865 -- (Andriashev 1973:546 [ref. 9214], McAllister et al. 1981:825 [ref. 5431], Andriashev in Whitehead et al. 1986:1143 [ref. 13677], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:713 [ref. 25968], Anderson & Fedorov 2004:28 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Balushkin et al. 2011:1010 [ref. 32449], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:420 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Mecklenburg et al. 2016:182 [ref. 34440] see remarks, Mecklenburg et al. 2018:352 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948] as rossii). Current status: Valid as Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae. Distribution: Arctic and North Atlantic. Habitat: marine.

rubra, Puzanovia Fedorov [V. V.] 1975:589 [528], Figs. 1-2 [Voprosy Ikhtiologii v. 15 (no. 4); ref. 1310] Bering Sea, 61°05'N, 179°20'W, depth 460-475 meters. Holotype: ZIN 39213 (230 mm TL, 1 of 3). Paratypes: (68) BMNH 1979.8.4.8 [ex ZIN 40525] (1); ZIN 39211-12 (3, 1), 39213 (2 of 3), 39214-15 (1, 5), 40516-24 (1, 3, 6, 1, 3, 1, 1, 1, 1), 40526 (32). Type catalog: Balushkin et al. 2011:970 [ref. 32449], Balushkin et al. 2012:164 [ref. 33667]. On p. 528 of English translation. Appeared as name only in Fedorov 1973. •Valid as Puzanovia rubra Fedorov 1975 -- (Toyoshima in Amaoka et al. 1983:258, 329 [ref. 8379], Toyoshima in Masuda et al. 1984:306 [ref. 6441], Anderson 1994:120 [ref. 21438], Nakabo 2000:1042 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:687 [ref. 25968], Nakabo 2002:1042 [ref. 26193], Fedorov et al. 2003:119 [ref. 27579], Anderson & Fedorov 2004:9 [ref. 27603], Balushkin et al. 2011:970 [ref. 32449], Parin et al. 2014:428 [ref. 33547]). Current status: Valid as Puzanovia rubra Fedorov 1975. Zoarcidae: Gymnelinae. Distribution: North Pacific: Bering Sea to Okhotsk Sea and Japan. Habitat: marine.

ryukyuensis, Lycenchelys Shinohara [G.] & Anderson [M. E.] 2007:60, Figs. 1-2 [Bulletin of the National Museum of Nature and Science (Ser. A) Supplement No. 1; ref. 29111] Okinawa Trough, East China Sea, 26°11.34'N, 124°54.27'E to 26°12.65'N, 124°55.47'E, depth 1493-1533 meters. Holotype: NSMT-P 63965. Paratypes: BSKU 57850-57581 (2), NSMT-P 63966-63967 (2). •Vallid as Lycenchelys ryukyuensis Shinohara & Anderson 2007 -- (Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys ryukyuensis Shinohara & Anderson 2007. Zoarcidae: Lycodinae. Distribution: Okinawa Trough near Ryukyu Islands, East China Sea. Habitat: marine.

sadoensis, Lycodes Toyoshima [M.] & Honma [Y.] 1980:48, Fig. 1 [Japanese Journal of Ichthyology v. 27 (no. 1); ref. 8650] Off Ishikawa Prefecture, Japan, 37°33.5'N, 136°15.0'E, depth 235 meters (not from off Sado Island but off Noto Penisula [ref. 33330]). Holotype: HUMZ 65832. Paratypes: HUMZ 65828-31 (1, 1, 1, 1), 65833 (1); NMC 79-697 (1); ZUMT 54225-26 [ex HUMZ 65834-35] (1, 1). •Valid as Lycodes sadoensis Toyoshima & Honma 1980 -- (Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:189 [ref. 5722], Anderson 1994:119 [ref. 21438], Nakabo 2000:1031 [ref. 25182], Nakabo 2002:1031 [ref. 26193], Anderson & Fedorov 2004:29 [ref. 27603], Kim et al. 2006:274 [ref. 29191], Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:1026 [ref. 32449], Shinohara et al. 2014:267 [ref. 33330]). •Synonym of Petroschmidtia teraoi (Katayama 1943) -- (Nazarkin et al. 2014:171 [ref. 33213]). Current status: Synonym of Petroschmidtia teraoi (Katayama 1943). Zoarcidae: Lycodinae. Distribution: Western North Pacific: Japan and Korea. Habitat: marine.

sagamia, Ericandersonia Shinohara [G.] & Sakurai [H.] 2006:174, Figs. 1-6, 7A, 9 [Ichthyological Research v. 53 (no. 2); ref. 28693] Honshu, Sagami Bay, Kanagawa Prefecture, off Chigasaki City, 35°13.7'N, 139°21.0'E, Japan, depth 880-930 meters. Holotype: NSMT-P 59763. Paratypes: NSMT-P 60356 (1), 63685 (1, dissected); USNM 384348 (1). •Valid as Ericandersonia sagamia Shinohara & Sukurai 2006 -- (Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Ericandersonia sagamia Shinohara & Sakurai 2006. Zoarcidae: Gymnelinae. Distribution: Sagami Bay, Japan. Habitat: marine.

sagittarius, Lycodes McAllister [D. E.] 1976:9, Figs. 1-3 [Publications in Biological Oceanography, National Museum Natural Science Canada No. 9; ref. 7446] Beaufort Sea, 50 miles north-northeast of Brownlow Point, 70°51'30"N, 145°17'W, depth 357 meters. Holotype: NMC 74-282. Paratypes: BMNH 1974.10.5.1 (1); NMC 74-275 (1), 74-282A (1); ROM 30520 (1); USNM 212282 (1); ZIN 32050 (spec. 1 of 2); ZMUC P761071 (1). Type catalog: Springer & Anderson 1997:22 [ref. 22953], Balushkin et al. 2011:1010 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667]. •Valid as Lycodes sagittarius McAllister 1976 -- (Andriashev in Whitehead et al. 1986:1144 [ref. 13677], McAllister et al. 1981:828 [ref. 5431], McAllister 1990:163 [ref. 14674], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:717 [ref. 25968], Møller 2001:111 [ref. 25378], Anderson & Fedorov 2004:29 [ref. 27603], Balushkin et al. 2011:1011 [ref. 32449], Dolgov 2013:920 [ref. 33112], Mecklenburg 2014:145 [ref. 33463], Parin et al. 2014:420 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:67, 209 [ref. 36199], Mecklenburg et al. 2016:184 [ref. 34440] see remarks, Mecklenburg et al. 2018:354 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes sagittarius McAllister 1976. Zoarcidae: Lycodinae. Distribution: Arctic Ocean. Habitat: marine.

saldanhai, Pachycara Biscoito [M.] & Almeida [A. J.] 2004:562, Figs. 2-4 [Copeia 2004 (no. 3); ref. 27911] Rainbow hydrothermal vent field (PP37), north Azores segment of Mid-Atlantic Ridge, 36°13.43'N, 33°54.08'W, depth 2280 meters. Holotype: MMF 31373. Paratypes: BMNH 2003.7.24.1 (1); MMF 31374-76 (1, 1, 1); MNHN 2003-0096 (1); USNM 371905 (1). Plus non-type material. •Valid as Pachycara saldanhai Biscoito & Almeida 2004 -- (Anderson et al. 2016:76 [ref. 34148]). Current status: Valid as Pachycara saldanhai Biscoito & Almeida 2004. Zoarcidae: Lycodinae. Distribution: Rainbow hydrothermal vent field, Mid-Atlantic Ridge, Atlantic. Habitat: marine.

sarsii, Lycodes Collett [R.] 1871:62 [3], Pl. [Forhandlinger i Videnskabs-selskabet i Christiania (for 1871); ref. 18017] Utne, Hardanger Fjord, southwestern coast of Norway, depth 188-282 meters. Holotype (unique): ZMUO J4555. Type catalog: Pethon 1969:12 [ref. 19268]. On p. 3 of separate. •Valid as Lycenchelys sarsii (Collett 1871) [sometimes as sarsi] -- (Andriashev 1973:542 [ref. 9214], Andriashev in Whitehead et al. 1986:1135 [ref. 13677], Anderson 1994:118 [ref. 21438], Koyanagi in Okamura et al. 1995:193 [ref. 22531], Anderson & Balanov 2000:1060 [ref. 25023], Anderson & Fedorov 2004:20 [ref. 27603], Møller et al. 2010:60 [ref. 31169], Balushkin et al. 2011:982 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:403 [ref. 33547], Afonso et al. 2016:63, 194 [ref. 36199], Mecklenburg et al. 2018:319 [ref. 36370]). Current status: Valid as Lycenchelys sarsii (Collett 1871). Zoarcidae: Lycodinae. Distribution: North Atlantic and Arctic Ocean. Habitat: marine.

scaurus, Lycodopsis Garman [S.] 1899:132, Pl. 32 (fig. 1) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Off Bay of Panama, 7°31'30"N, 79°14'W, Albatross station 3384, depth 458 fathoms. Holotype (unique): MCZ 28689. •Valid as Lycenchelys scaurus (Garman 1899) -- (Pequeño 1989:48 [ref. 14125], Anderson 1994:118 [ref. 21438], Anderson 1995:104 [ref. 21928], Anderson & Fedorov 2004:20 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys scaurus (Garman 1899). Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

schmidti, Lycodes Gratzianov [V. I.] 1907:426, 430 [Trudy Otdela Ichtiologii Imperatorskago Russkago Obshchestva Akklimatizacii Zivotnych' i Rastenii. v. 4; ref. 1871] East coast of Sakhalin Island, Okhotsk Sea, Russia. Lectotype: ZIN 13010 (largest specimen). Paralectotypes: ZIN 13010 (1). Type catalog:Balushkin et al. 2011:1014 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667]. Based on Schmidt 1904:201 (not researched). Lectotype selected by Nazarkin 2014:188 [ref. 33213]. •Synonym of Lycodes palearis Gilbert 1896, but a valid subspecies -- (Lindberg & Krasyukova 1975:158 [ref. 7348]). •Questionably a synonym of Lycodes fasciatus (Schmidt 1904) -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Valid as Lycodes schmidti Gratzianov 1907 -- (Toyoshima 1985:214 [ref. 5722], Anderson 1994:119 [ref. 21438], Sokolovskaya et al. 1998:12 [ref. 24670], Møller 2001:111 [ref. 25378], Anderson & Fedorov 2004:29 [ref. 27603], Balushkin et al. 2011:1014 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330]). •Valid as Petroschmidtia schmidti (Gratzianov 1907) -- (Nazarkin et al. 2014:188 [ref. 33213]). Current status: Valid as Petroschmidtia schmidti (Gratzianov 1907). Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk and Japan seas. Habitat: marine.

schmidti, Lycodes Soldatov [V. K.] 1918:115, Fig. 2 [An. Mus. Zool. Acad. Sci. Petrograd v. 22; ref. 21830] Near Rymnik, eastern Sakhalin Island, Okhotsk Sea; western Sakhalin Island, 50°31'N, 141°56'E, Sea of Japan. Type catalog: Balushkin et al. 2011:1020 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667] Also appeared in Soldatov 1922:115, Fig. 2 [ref. 15429]. Objectively invalid; preoccupied by Lycodes schmidti Gratzianov 1907, replaced by Lycodes sigmatoides Lindberg & Krasyukova 1975. •Questionably a synonym of Lycodes brevicaudus Taranetz & Andriashev 1937 -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Objective synonym of Lycodes sigmatoides Lindberg & Krasyukova 1975 -- (Toyoshima 1985:204 [ref. 5722], Sheiko & Fedorov 2000:37 [ref. 25650], Anderson & Fedorov 2004:29 [ref. 27603]). •Synonym of Lycodes tanakae Jordan & Thompson 1914 -- (Balushkin et al. 2011:1014 [ref. 32449], Parin et al. 2014:422 [ref. 33547]). Current status: Synonym of Lycodes tanakae Jordan & Thompson 1914. Zoarcidae: Lycodinae. Habitat: marine.

schmidti, Lycogrammoides Soldatov [V. K.] & Lindberg [G. U.] 1928:41, Fig. [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 30 (no. 1) (for 1929); ref. 4161] Tauyskaya Bay, off Ol'skii Island, northern Okhotsk Sea, depth 108 meters. Holotype (unique): ZIN 19120. Type catalog: Balushkin et al. 2011:1018 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667]. •Valid as Lycogrammoides schmidti Soldatov & Lindberg 1928 -- (Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Fedorov et al. 2003:118 [ref. 27579] dated 1928, Anderson & Fedorov 2004:32 [ref. 27603], Balushkin et al. 2011:1018 [ref. 32449], Parin et al. 2014:424 [ref. 33547]). Current status: Valid as Lycogrammoides schmidti Soldatov & Lindberg 1928. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

schmidti, Notolycodes Gosztonyi [A. E.] 1977:224, Figs. 13-14 [Archiv für Fischereiwissenschaft v. 27 (no. 3); ref. 6103] Continental slope off Buenos Aires, Argentina, 36°00'S, 52°58'W, depth 800 meters. Holotype: ISH 1134-1966. Paratypes: ISH 1515-1966 (1), 1586-1966a-b (2), 1977-1968 (1). Type catalog: Krefft 1978:15 [ref. 9884]. •Valid as Notolycodes schmidti Gosztonyi 1977 -- (Anderson & Gosztonyi 1991:8 [ref. 19348], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:33 [ref. 27603], Balushkin et al. 2011:1027 [ref. 32449], Nión et al. 2016:51 [ref. 35565]). Current status: Valid as Notolycodes schmidti Gosztonyi 1977. Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

semenovi, Lycodes Popov [A. M.] 1931:144, Pl. 1 (fig. 2) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] 56°09'N, 144°30'E, Okhotsk Sea, off St. Iona Island, depth 307 meters. Holotype (unique): ZIN 29995 [not 2995]. Type catalog: Balushkin et al. 2011:1011 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667]. •Valid as Lycodes semenovi Popov 1931 -- (Toyoshima 1985:205 [ref. 5722], Anderson 1994:119 [ref. 21438], Fedorov et al. 2003:116 [ref. 27579], Anderson & Fedorov 2004:29 [ref. 27603], Nazakin & Fedorov 2007:701 [ref. 29424], Ikeda et al. 2008:357 [ref. 29896], Balushkin et al. 2011:1011 [ref. 32449], Parin et al. 2014:420 [ref. 33547]). Current status: Valid as Lycodes semenovi Popov 1931. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

seminuda, Lycodes reticulatus forma Smitt [F. A.] 1901:31 [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 27 (no. 4); ref. 18732] Not intended as new; subsequent treatment of Lycodes seminudus Reinhardt 1837 as a subspecific name. •Not intended as new, in the family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). •In the synonymy of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Parin et al. 2014:419 [ref. 33547]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae.

seminudus, Lycodes Reinhardt [J. C. H.] 1837:223 [117] [Det Kongelige Danske videnskabernes selskabs naturvidenskabelige og mathematiske afhandlinger v. 7; ref. 13398] Omenak, Greenland, 71°N. Holotype: ZMUC 15 [or 9]. Type catalog: Nielsen 1974:78 [ref. 9588]. On p. 117 of separate. •Valid as Lycodes seminudus Reinhardt 1837 -- (Andriashev 1973:546 [ref. 9214], McAllister et al. 1981:833 [ref. 5431], Andriashev in Whitehead et al. 1986:1145 [ref. 13677], McAllister 1990:163 [ref. 14674], Anderson 1994:119 [ref. 21438], Saito & Okamura in Okamura et al. 1995:200 [ref. 22531], Mecklenburg et al. 2002:707 [ref. 25968], Anderson & Fedorov 2004:29 [ref. 27603], Møller et al. 2010:62 [ref. 31169], Balushkin et al. 2011:1011 [ref. 32449], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:141 [ref. 33463], Parin et al. 2014:421 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:67, 210 [ref. 36199], Mecklenburg et al. 2016:186 [ref. 34440] see remarks, Mecklenburg et al. 2018:356 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes seminudus Reinhardt 1837. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic and North Pacific. Habitat: marine.

semisquamata, Hadropareia Andriashev [A. P.] & Matyushin [V. M.] in Matyushin 1989:525 [Voprosy Ikhtiologii v. 29 (no. 4); ref. 17696] Western Pacific, southern Kuril Islands, Shikotan Island, Krabovaya Bay. Holotype: ZIN 35815. Paratypes: ZIN 35815A (1), 46783 (1), 47951-52 (1, 3). Type catalog: Balushkin et al. 2011:968 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. First author is A. N. Andriashev. Appeared first as name only in Andriashev in Pinchuk 1974. •Valid as Hadropareia semisquamata Andriashev & Matyushin 1989 -- (Anderson 1994:117 [ref. 21438], Sheiko & Fedorov 2000:35 [ref. 25650], Anderson & Fedorov 2004:7 [ref. 27603], Balushkin et al. 2011:968 [ref. 32449], Parin et al. 2014:395 [ref. 33547]). Current status: Valid as Hadropareia semisquamata Andriashev & Matyushin 1989. Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Kuril Islands. Habitat: marine.

septentrionalis, Lycenchelys sarsii var. Knipowitsch [N. M.] 1906:4, Figs. 1-3 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Western coast of Kola Peninsula, 69°23'N, 32°55'E, Russia, depth 271 meters. Lectotype: ZIN 13360. Paralectotypes: ZIN 12128 (1), 13357-59 (1, 4, 2), 13361-72 (1 each). Type catalog: Balushkin et al. 2011:982 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667]. •Synonym of Lycenchelys sarsii (Collett 1871) -- (Andriashev 1973:542 [ref. 9214], Anderson & Fedorov 2004:20 [ref. 27603], Balushkin et al. 2011:982 [ref. 32449], Parin et al. 2014:403 [ref. 33547]). Current status: Synonym of Lycenchelys sarsii (Collett 1871). Zoarcidae: Lycodinae. Habitat: marine.

septentrionalis, Lycodes vahli var. Knipowitsch [N. M.] 1906:95 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Barents Sea, Russia. Syntypes: ZIN 13437-71 (1 lost, 1 lost, 1, 3, 3, 1, 2, 1 lost, 3, 1, 1, 1, 1 lost, 2, 3, 1, 1, 1 lost, 1, 1 lost, 2 lost, 3, 2, 1 lost, 1, 1 lost, 1 lost, 1, 1, 1, 2, 1, 1, 1, 4), 14248 (1). Type catalog: Balushkin et al. 2011:995 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. Spelled sententrionalis by Knipowitsch 1926:111 [ref. 22166]. •Synonym of Lycodes vahlii Reinhardt 1831 -- (Andriashev 1973:543 [ref. 9214], Anderson 1994:119 [ref. 21438]). •Synonym of Lycodes gracilis Sars 1867 -- (Carl 2002:75 [ref. 26925], Anderson & Fedorov 2004:25 [ref. 27603], Balushkin et al. 2011:995 [ref. 32449], Parin et al. 2014:411 [ref. 33547]). Current status: Synonym of Lycodes gracilis Sars 1867. Zoarcidae: Lycodinae. Habitat: marine.

serpens, Lycodes Garman [S.] 1899:134 [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Off Gulf of California, 27°34'N, 110°53'40"W, Albatross station 3436, depth 905 fathoms. Holotype: MCZ 28686. Paratypes: MCZ 117364 (1) destroyed. •Synonym of Lycenchelys porifer (Gilbert 1890) -- (Anderson 1994:117 [ref. 21438], Anderson 1995:96 [ref. 21928], Anderson & Fedorov 2004:19 [ref. 27603]). Current status: Synonym of Lycenchelys porifer (Gilbert 1890). Zoarcidae: Lycodinae. Habitat: marine.

shcherbachevi, Pachycara Anderson [M. E.] 1989:233, Fig. 10 [Proceedings of the California Academy of Sciences v. 46 (no. 10); ref. 13487] Bay of Bengal, 15°54'N, 90°17'W, depth 2600 meters. Holotype (unique): ZMUC P761147. •Valid as Pachycara shcherbachevi Anderson 1989 -- (Anderson 1991:157 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:226 [ref. 22911], Møller 2003:363 [ref. 26908], Anderson & Fedorov 2004:36 [ref. 27603], Biscoito & Almeida 2004:566 [ref. 27911], Anderson & Mincarone 2006:24 [ref. 28658], Shinohara 2012:55 [ref. 32102], Balushkin et al. 2011:1027 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara shcherbachevi Anderson 1989. Zoarcidae: Lycodinae. Distribution: Northern Indian Ocean. Habitat: marine.

sigmatoides, Lycodes Lindberg [G. U.] & Krasyukova [Z. V.] 1975:161, Fig. 127 [ ref. 7348] Near Rymnik, eastern Sakhalin Island, Okhotsk Sea; western Sakhalin Island, 50°31'N, 141°56'E, Sea of Japan. Syntypes: (5) ZIN 13009 (3). Non-types: ZIN 19165-66 (3, 1). Type catalog: Balushkin et al. 2011:1014 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667] with ZIN 19165 as syntypes. Replacement name for Lycodes schmidti Soldatov 1918, preoccupied by Lycodes schmidti Gratzianov 1907. Appeared in error as sigmatus in the index. •Valid as Lycodes sigmatoides Lindberg & Krasyukova 1975 -- (Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:204 [ref. 5722], Anderson 1994:119 [ref. 21438], Sokolovskaya et al. 1998:12 [ref. 24670] as sygmatoides, Nakabo 2000:1038 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1038 [ref. 26193], Fedorov et al. 2003:117 [ref. 27579], Anderson & Fedorov 2004:29 [ref. 27603], Kim et al. 2006:276 [ref. 29191], Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:1014 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:421 [ref. 33547]). Current status: Valid as Lycodes sigmatoides Lindberg & Krasyukova 1975. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea and Sea of Japan, Kamchata and Kuril islands. Habitat: marine.

similis, Lycodes Jensen [A. S.] 1902:205 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Jan Mayen, northeastern Atlantic, depth 371 fathoms. Syntypes: ZMUC 220-234 (25) (but see Møller 2001:985 [ref. 25812], who gives different catalog numbers). Type catalog: Nielsen 1974:78 [ref. 9588]. Appeared in Jensen 1904:39, 46-48, Pl. 5 (figs.2a-d), Pl. 6 (figs. 3a-d) [ref. 15186] as Lycodes pallidus var. similis according to Møller 2001:983 [ref. 25812]. •Synonym of Lycodes pallidus Collett 1879 -- (Andriashev 1973:544 [ref. 9214], Anderson 1994:119 [ref. 21438], Møller 2001:983 [ref. 25812], Anderson & Fedorov 2004:27 [ref. 27603], Parin et al. 2014:417 [ref. 33547]). Current status: Synonym of Lycodes pallidus Collett 1879. Zoarcidae: Lycodinae. Habitat: marine.

skopetsi, Magadania Shinohara [G.], Nazarkin [M. V.] & Chereshnev [I. A.] 2004:138, Figs. 1-9 [Ichthyological Research v. 51 (no. 2); ref. 27581] Nagayeva Bay near Magadan, Sea of Okhotsk, about 59°30'N, 151°00'E, Magadan, Russia. Holotype: NSMT-P 55537. Paratypes: HUMZ 178695-96 (2), 178697-99 (3); NSMT-P 55538 (1, dissected), 55539 (1), 60857 (4); ZIN 52519 (2), 52520 (3). Type catalog: Balushkin et al. 2011:969 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. •Valid as Magadania skopetsi Shinohara, Nazarkin & Chereshnev 2004. •Valid as Magadania skopetzi Shinohara, Nazarkin & Chereshnev unpubl. -- (Fedorov et al. 2003:119 [ref. 27579] as a nomen nudum). •Valid as Magadanichthys skopetsi (Shinohara, Nazarkin & Chereshnev 2004) -- (Shinohara, Nazaekin, Yabe & Chereshnev 2006:94 [ref. 28853], Balushkin et al. 2011:969 [ref. 32449], Parin et al. 2014:425 [ref. 33547], Radchenko et al. 2018:315 [ref. 36243]). Current status: Valid as Magadanichthys skopetsi (Shinohara, Nazarkin & Chereshnev 2004). Zoarcidae: Gymnelinae. Distribution: Western North Pacific: Sea of Okhotsk. Habitat: marine.

sobrali, Crossostomus Lloris [D.] & Rucabado [J. A.] 1989:264, Fig. 1 [Miscellània Zoològica v. 11 (1987); ref. 15956] Piedra Redonda, Ushuaia Peninsula, Tierra del Fuego, 54°50.8'S, 68°19.2'W, Argentina. Holotype (unique): IIPB 14/1987. Type catalog: Allué et al. 2000:75 [ref. 30499]. •Valid as Crossostomus sobrali Lloris & Rucabado 1989 -- (López et al. 1996:31 [ref. 23681], Pequeño 1997:83 [ref. 23536]). •Synonym of Crossostomus chilensis (Regan 1913) -- (Anderson & Gosztonyi 1991:5 [ref. 19348], Lloris & Rucabado 1991:68 [ref. 22530], Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:12 [ref. 27603]). Current status: Synonym of Crossostomus chilensis (Regan 1913). Zoarcidae: Lycodinae. Distribution: Southern South America. Habitat: marine.

soldatovi, Gymnelus Chernova [N. V.] 2000:8 [4], Figs. 1, 2a [Voprosy Ikhtiologii v. 40 (no. 1); ref. 24885] Luzhina Strait, between Paramushir and Antsiferovo Islands, Kuril Islands, depth 100 meters. Holotype: ZIN 51558. Paratypes: ZIN 20166 (2), ?23951 (1), ?34999 (1). Type catalog: Balushkin et al. 2011:966 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. Anderson in Anderson & Fedorov 2004:6 [ref. 27603] believes this is probably a synonym of G. hemifasciatus. On p. 4 of English translation. •Valid as Gymnelus soldatovi Chernova 2000 -- (Fedorov et al. 2003:111 [ref. 27579], Anderson & Fedorov 2004:6 [ref. 27603], Balushkin et al. 2011:966 [ref. 32449], Parin et al. 2014:394 [ref. 33547]). •Synonym of Gymnelus hemifasciatus Andriashev 1937 -- (Mecklenburg & Anderson 2015:277 [ref. 33742], Mecklenburg et al. 2016:154 [ref. 34440]). Current status: Synonym of Gymnelus hemifasciatus Andriashev 1937. Zoarcidae: Gymnelinae. Distribution: Western North Pacific (if valid). Habitat: marine.

soldatovi, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] 1935:246, Fig. 3 [Zoologischer Anzeiger v. 112 (nos 9-10); ref. 15744] Okhotsk Sea, east of Cape Terpenija, 49°09.5'N, 145°17.5'E, depth 440 meters. Syntypes: ZIN 24846 (1), 25190 (2). Type catalog: Balushkin et al. 2011:1012 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. •Valid as Lycodes soldatovi Taranetz & Andriashev 1935 -- (Lindberg & Krasyukova 1975:153 [ref. 7348], Toyoshima in Amaoka et al. 1983:267, 331 [ref. 8379], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:207 [ref. 5722], Anderson 1994:119 [ref. 21438], Nakabo 2000:1035 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:718 [ref. 25968], Møller 2001:111 [ref. 25378], Nakabo 2002:1035 [ref. 26193], Fedorov et al. 2003:117 [ref. 27579], Anderson & Fedorov 2004:29 [ref. 27603], Balushkin et al. 2011:1012 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:422 [ref. 33547]). Current status: Valid as Lycodes soldatovi Taranetz & Andriashev 1935. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea, Sea of Japan and Bering seas (doubtful on 33547). Habitat: marine.

soldatovi, Lycogramma Schmidt [P. Yu.] 1950:116 [103], Pl. 11 (fig. 1) [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] East of Cape Terpenie, 49°12'N, 145°21'E, Okhotsk Sea, depth 440 meters. Holotype (unique): ZIN 30963. Type catalog: Balushkin et al. 2011:974 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. On p. 103 of English translation. •Status uncertain, probably a synonym of Bothrocara brunneum (Bean 1890) -- (Anderson 1994:116 [ref. 21438]). •Synonym of Bothrocara zestum (Jordan & Fowler 1902) -- (Anderson et al. 2009:190 [ref. 30175] but original description not cited). •Valid as Bothrocara soldatovi (Schmidt 1950) -- (Sheiko & Fedorov 2000:34 [ref. 25650], Fedorov et al. 2003:109 [ref. 27579], Anderson & Fedorov 2004:11 [ref. 27603], Balushkin et al. 2011:974 [ref. 32449], Parin et al. 2014:385 [ref. 33547]). Current status: Valid as Bothrocara soldatovi (Schmidt 1950). Zoarcidae: Lycodinae. Distribution: Northwestern Pacific. Habitat: marine.

spilotus, Lycenchelys Fowler [H. W.] 1943:89, Fig. 24 [Bulletin of the United States National Museum No. 100, v. 14 (pt 2); ref. 1441] Off Niigata Light, Sea of Japan, 38°12'N, 138°52'E, Albatross station 4817, depth 61 fathoms. Holotype: USNM 99511. Paratypes: USNM 122362 (1). Type catalog: Springer & Anderson 1997:22 [ref. 22953]. •Valid as Davidojordania [sic] spilota (Fowler 1943) -- (Lindberg & Krasyukova 1975:177 [ref. 7348]). •Valid as Davidijordania spilota (Fowler 1943) -- (Toyoshima in Masuda et al. 1984:305 [ref. 6441]). •Synonym of Davidijordania poecilimon (Jordan & Fowler 1902) -- (Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:4 [ref. 27603], Parin et al. 2014:388 [ref. 33547]). Current status: Synonym of Davidijordania poecilimon (Jordan & Fowler 1902). Zoarcidae: Gymnelinae. Habitat: marine.

spitzbergensis, Lycodes celatus var. Jensen [A. S.] 1902:209 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Spitsbergen, Arctic regions, depth 39-75 fathoms. No types known. •Synonym of Lycodes rossi Malmgren 1865 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603], Parin et al. 2014:420 [ref. 33547]). Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae. Habitat: marine.

squamiventer, Lycodes pallidus var. Jensen [A. S.] 1904:39, 46, Pl. 4 (fig. 2) [Danish Ingolf-Expedition v. 2 (no. 4); ref. 15186] Iceland. Lectotype: ZMUC 235. Paralectotypes: ZMUC 7-8 (2), 237239 (3); NMC 74-0395 (1), ZIN 40718 (1), ZMUB 1811 (2), ZMUO J4546 (2). Type catalog: Nielsen 1974:77 [ref. 9588]; Balushkin et al. 2011:1013 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. •Valid as Lycodes squamiventer Jensen 1904 -- (Andriashev 1973:546 [ref. 9214], Andriashev in Whitehead et al. 1986:1145 [ref. 13677], McAllister et al. 1981:832 [ref. 5431], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:722 [ref. 25968], Møller 2001:985 [ref. 25812], Anderson & Fedorov 2004:29 [ref. 27603], Møller et al. 2010:63 [ref. 31169], Balushkin et al. 2011:1013 [ref. 32449], Dolgov 2013:920 [ref. 33112], Mecklenburg 2014:145 [ref. 33463], Parin et al. 2014:422 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:67, 211 [ref. 36199], Mecklenburg et al. 2016:188 [ref. 34440], Mecklenburg et al. 2018:358 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes squamiventer Jensen 1904. Zoarcidae: Lycodinae. Distribution: North Atlantic and Arctic Ocean. Habitat: marine.

squamosa, Lycodes turneri atlanticus morpha Vladykov [V. D.] & Tremblay [J.-L.] 1936:39 [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Described as a morpha, regarded as infrasubspecific and unavailable. •Infrasubspecific and unavailable, in family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Zoarcidae: Lycodinae.

squamosus, Lycenchelys Toyoshima [M.] in Amaoka et al. 1983:145, Pl. 93; Figs. 20-22 [Fishes from the north-eastern Sea of Japan and the Okhotsk Sea off Hokkaido; ref. 8379] Off Miyagi Prefecture, Sea of Japan, 37°55.4'N, 142°24.5'E, depth 1005 meters. Holotype: HUMZ 78464. Paratypes: HUMZ 72563 (1), 78930 (1). •Valid as Lycenchelys squamosus Toyoshima 1983 -- (Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:156 [ref. 5722]). •Valid as Lycenchelys squamosa Toyoshima 1983 -- (Anderson 1994:118 [ref. 21438], Fedorov 1995:130 [ref. 21958], Shinohara et al. 1996:182 [ref. 26605], Shinohara & Matsuura 1998:154 [ref. 23382], Nakabo 2000:1033 [ref. 25182], Sheiko & Fedorov 2000:35 [ref. 25650], Nakabo 2002:1033 [ref. 26193], Anderson & Fedorov 2004:20 [ref. 27603], Shinohara & Anderson 2007:64 [ref. 29111], Balushkin et al. 2011:983 [ref. 32449], Parin et al. 2014:403 [ref. 33547], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys squamosa Toyoshima 1983. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Japan. Habitat: marine.

stehmanni, Ophthalmolycus Gosztonyi [A. E.] 1977:230, Fig. 16 [Archiv für Fischereiwissenschaft v. 27 (no. 3); ref. 6103] North of the Malvinas Islands (Falkland Islands), 49°13'S, 58°45'W, depth 485-506 meters. Holotype (unique): ISH 317-1971. Type catalog: Krefft 1978:15 [ref. 9884]. •Valid as Plesienchelys stehmanni (Gosztonyi 1977) -- (Anderson 1988:268 [ref. 6334], Anderson 1994:120 [ref. 21438], Anderson & Fedorov 2004:36 [ref. 27603], Balushkin et al. 2011:1022 [ref. 32449]). Current status: Valid as Plesienchelys stehmanni (Gosztonyi 1977). Zoarcidae: Lycodinae. Distribution: Western South Atlantic. Habitat: marine.

stigma, Ophidium Lay [G. T.] & Bennett [E. T.] 1839:67, Pl. 20 (fig. 1) [The zoology of Captain Beechey's voyage; ref. 2730] Kotzebue Sound, Alaska, U.S.A. Holotype (unique): not preserved. Based on notes and a sketch by Mr. Lay, and notes by Mr. Collie. •Synonym of Gymnelus viridis (Fabricius 1780) -- (Anderson 1982:30 [ref. 5520], Mecklenburg et al. 2002:690 [ref. 25968], Anderson & Fedorov 2004:6 [ref. 27603], Balushkin et al.2011:966 [ref. 32499], Parin et al. 2014:394 [ref. 33547]). Current status: Synonym of Gymnelus viridis (Fabricius 1780). Zoarcidae: Gymnelinae. Habitat: marine.

subarcticus, Lycodes rossi forma Knipowitsch [N. M.] 1906:27 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Syntypes: ZIN 13408 (1), 13409 (1), 13424 (2), 13425 (1), 13426 (2), 13427 (2). Type catalog: Balushkin et al. 2011:1010 [ref. 32449], Balushkin et al. 2012:164 [ref. 33667]. Not available, infrasubspecific; publication in an available way as a species or subspecies not investigated [Knipowitsch used v. [variety] and f. [forma] -- in his work, the former is considered subspecific the latter infrasubspecific; see Article 45(g).]. From Barents Sea, Russia. •In the synonymy of Lycodes rossii Malmgren 1865 -- (Anderson 1994:119 [ref. 21438] as rossi, Anderson & Fedorov 2004:48 [ref. 27603], Balushkin et al. 2011:1010 [ref. 32449], Parin et al. 2014:420 [ref. 33547]). Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae.

sulaki, Pachycara Anderson [M. E.] 1989:231, Fig. 9 [Proceedings of the California Academy of Sciences v. 46 (no. 10); ref. 13487] Mona Passage off Puerto Rico, 18°28.7'N, 67°20.6'W, depth 2000 meters. Holotype: USNM 292811. Paratypes: SIO 87-33 (1), USNM 233627 (1). Type catalog: Springer & Anderson 1997:22 [ref. 22953]. •Valid as Pachycara sulaki Anderson 1989 -- (Anderson 1991:158 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:226 [ref. 22911], Møller 2003:359 [ref. 26908], Anderson 2003:1741 [ref. 27106], Anderson & Fedorov 2004:36 [ref. 27603], Biscoito & Almeida 2004:566 [ref. 27911], McEachran & Fechhelm 2005:549 [ref. 28815], Anderson 2006:12 [ref. 28576], Møller & King 2007:588 [ref. 29259], Balushkin et al. 2011:1027 [ref. 32449], Anderson et al. 2016:76 [ref. 34148]). Current status: Valid as Pachycara sulaki Anderson 1989. Zoarcidae: Lycodinae. Distribution: Western Atlantic. Habitat: marine.

suspectus, Phucocoetes Garman [S.] 1899:137, Pl. 30 (figs. 3-3a) [Memoirs of the Museum of Comparative Zoology v. 24; ref. 1540] Off Acapulco, Mexico, 16°33'N, 99°52'30"W, Albatross station 3418, depth 660 fathoms. Holotype (unique): MCZ 28683. •Valid as Pachycara suspectum (Garman 1899) -- (Anderson & Peden 1988:84 [ref. 7215], Anderson 1989:235 [ref. 13487], Anderson 1991:157 [ref. 20019], Anderson 1994:120 [ref. 21438], Anderson & Bluhm 1997:226 [ref. 22911], Pequeño 1997:83 [ref. 23536], Møller 2003:359 [ref. 26908], Anderson & Fedorov 2004:36 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Anderson 2012:41 [ref. 32270], Balushkin et al. 2011:1027 [ref. 32449], Corbella & Møller 2014:185 [ref. 32448]). Current status: Valid as Pachycara suspectum (Garman 1899). Zoarcidae: Lycodinae. Distribution: Eastern Pacific. Habitat: marine.

taeniatus, Gymnelus Chernova [N. V.] 1999:309, Figs. 3b, 4 [Voprosy Ikhtiologii v. 39 (no. 3); ref. 24868] earing Seda, Kuna Island, Franz Josef Island, depth 25-40 meters. Holotype (unique): ZIN 51567. Type catalog: Balushkin et al. 2011:966 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. Anderson in Anderson & Fedorov 2004:6 [ref. 27603] believes this is probably a synonym of G. retrodorsalis. •Valid as Gymnelus taeniatus Chernova 1999 -- (Chernova 2000:8 [ref. 24885], Anderson & Fedorov 2004:6 [ref. 27603], Balushkin et al. 2011:966 [ref. 32449].). •Synonym of Gymnelus retrodorsalis Le Danois 1913 with question -- (Mecklenburg et al. 2011:124, 128 [ref. 31212], Mecklenburg & Anderson 2015:284 [ref. 33742]), Current status: Synonym of Gymnelus retrodorsalis Le Danois 1913. Zoarcidae: Gymnelinae. Distribution: Arctic Ocean: Franz Josef Island [if valid]. Habitat: marine.

tanakae, Lycodes Jordan [D. S.] & Thompson [W. F.] 1914:299, Pl. 37 (fig. 2) [Memoirs of the Carnegie Museum v. 6 (no. 4); ref. 2543] Noto, Hondo, Japan. Holotype (unique): FMNH 57068 [ex CM 6004]. Type catalog: Henn 1928:98 [ref. 12305], Ibarra & Stewart 1987:54 [ref. 12367]. The spelling tanakae is acceptable; early workers almost certainly Latinized the name first, and then added an 'e' (Art. 31.1.1) [WNE, 12 Mar. 2012] -- we now suggest using tanakae.. •Valid as Lycodes tanakae Jordan & Thompson 1914 -- (Lindberg & Krasyukova 1975:160 [ref. 7348] as tanakae, Toyoshima in Amaoka et al. 1983:261, 330 [ref. 8379] as tanakae, Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:193 [ref. 5722], Anderson 1994:119 [ref. 21438] as tanakae, Sokolovskaya et al. 1998:12 [ref. 24670], Nakabo 2000:1038 [ref. 25182] as tanakai, Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1038 [ref. 26193] as tanakai, Youn 2002:375, 640 [ref. 26218] as tanakai, Anderson & Fedorov 2004:30 [ref. 27603], Kim et al. 2006:278 [ref. 29191], Nazarkin 2010:603 [ref. 31019], Shinohara et al. 2011:52 [ref. 31715], Balushkin et al. 2011:1013 [ref. 32449], Nazarkin et al. 2014:177 [ref. 33213], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:422 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331]). Current status: Valid as Lycodes tanakae Jordan & Thompson 1914. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea and Sea of Japan. Habitat: marine.

tanakae, Zestichthys Jordan [D. S.] & Hubbs [C. L.] 1925:321, Pl. 12 (fig. 1) [Memoirs of the Carnegie Museum v. 10 (no. 2); ref. 2486] Kushiro, Hokkaido, Japan. Holotype (unique): FMNH 58841 [ex CM 7951]. Type catalog: Henn 1928:98 [ref. 12305], Ibarra & Stewart 1987:85 [ref. 12367]. Correction to tanakai is not necessary. The spelling tanakae is acceptable as early workers almost certainly Latinized the name first, and then added an "e' (Art. 31.1.1) [WNE, 12 Mar. 2012] -- suggesting tanakae •Status uncertain, in Bothrocara -- (Anderson & Fedorov 2004:48 [ref. 27603]). •Valid as Bothrocara tanakae (Jordan & Hubbs 1925) -- (Anderson 1994:116 [ref. 21438], Nakabo 2000:1041 [ref. 25182], Nakabo 2002:1041 [ref. 26193], Anderson et al. 2009:188 [ref. 30175], Shinohara et al. 2011:50 [ref. 31715], Shinohara et al. 2014:264 [ref. 33330]). •Valid as Zestichthys tanakae Jordan & Hubbs 1925 -- (Lindberg & Krasyukova 1975:185 [ref. 7348], Toyoshima in Amaoka et al. 1983:149, 210 [ref. 8379], Toyoshima in Masuda et al. 1984:309 [ref. 6441] as tanakai, Shinohara et al. 1996:182 [ref. 26605] as tanakai, Balushkin et al. 2011:1022 [ref. 32449], Parin et al. 2014:429 [ref. 33547]). Current status: Valid as Zestichthys tanakae Jordan & Hubbs 1925. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

tangwangi, Zoarces Wu [H.-W.] 1930:60, fig. [Contributions from the Biological Laboratory of the Science Society of China. (Zoological Series) v. 6 (no. 6); ref. 16347] Foochow, China. Holotype (unique): whereabouts unknown. •Synonym of Zoarces gillii Jordan & Starks 1905 -- (Lindberg & Krasyukova 1975:133 [ref. 7348], Anderson & Fedorov 2004:2 [ref. 27603], Parin et al. 2014:430 [ref. 33547]). Current status: Synonym of Zoarces gillii Jordan & Starks 1905. Zoarcidae: Zoarcinae. Habitat: marine.

taranetzi, Lycodes (Furcimanus) Andriashev [A. P.] in Lindberg & Krasyukova 1975:152, Fig. 119-A [ ref. 7348] Pacific side of Iturup Island, depth 415 meters. Holotype: ZIN 42247. Paratypes: ZIN 42248 (7), 42262 (1). Type catalog: Balushkin et al. 2011:977 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. Author is A. N. Andriashev. On p. 185 of English translation. Appeared first as name only in Lindberg 1959. •Synonym of Lycodes hubbsi Matsubara 1955 -- (Toyoshima 1985:226 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Anderson & Fedorov 2004:25 [ref. 27603], Balushkin et al. 2011:997 [ref. 32449], Parin et al. 2014:412 [ref. 33547]). Current status: Synonym of Lycodes hubbsi Matsubara 1955. Zoarcidae: Lycodinae. Habitat: marine.

teraoi, Lycodes Katayama [M.] 1943:103, Fig. 2 [Annotationes Zoologicae Japonenses v. 22 (no. 2); ref. 2567] Off Tsuiyama, near Moroyose, Hyogo-ken, Japan, depth 100 fathoms. Lectotype: NSMT-P 78785 [163 mm TL]. Paralectotypes: HUMZ 81282-85 (4). Holotype is figured specimen (if used); designated lectotype by Nazarkin et al. 2014:173 [ref. 33213]. •Valid as Lycodes teraoi Katayama 1943 -- (Lindberg & Krasyukova 1975:147 [ref. 7348], Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:188 [ref. 5722], Anderson 1994:119 [ref. 21438], Nakabo 2000:1034 [ref. 25182], Nakabo 2002:1034 [ref. 26193], Anderson & Fedorov 2004:30 [ref. 27603], Shinohara et al. 2011:52 [ref. 31715], Balushkin et al. 2011:1026 [ref. 32449]). •Valid as Petroschmidtia teraoi (Katayama 1943) -- (Nazarkin et al. 2014:171, 179 [ref. 33213], Tohkairin et al. 2015:12 [ref. 35331]). Current status: Valid as Petroschmidtia teraoi (Katayama 1943). Zoarcidae: Lycodinae. Distribution: Western North Pacific: Sea of Okhotsk, Japan Sea (Japan and Korea), and Sakhalin Island, Russia. Habitat: marine.

terraenovae, Lycodes Collett [R.] 1896:54 [Poissons provenant des campagnes du Yacht "L'Hirondelle" (1885-1888) Fasc. 10; ref. 890] Newfoundland Banks, northwestern Atlantic, 46°04'40"N, 49°02'30"W, probably station 161 [not 162], depth 1267 meters. Lectotype: MOM-POI 295 [ex syntype b]. Paralectotypes: (3) MOM-POI 295 (1, former syntype a), 4651 (1, former syntype c, in 3 pieces), 3005 (former syntype d, stuffed, poor condition). Type catalog: Belloc 1949:14 [ref. 21544]. Originally as Terrae Novae; as a name "donnes ici provisoirement..." Lectotype selected by Møller 2000:847 [ref. 24585]. See Møller 2000:849 [ref. 24585] for information on the type locality. •Status uncertain -- (Anderson 1994:119 [ref. 21438]). •Valid as Lycodes terraenovae Collett 1896 -- (McAllister 1990:163 [ref. 14674], Møller 1997:47 [ref. 22879], Møller 2000:846 [ref. 24585], Carl 2002:65 [ref. 26925], Moore et al. 2003:228 [ref. 27076], Anderson 2003:1741 [ref. 27106], Anderson & Fedorov 2004:30 [ref. 27603], Møller et al. 2010:63 [ref. 31169], Balushkin et al. 2011:1015 [ref. 32449], Afonso et al. 2016:67, 212 [ref. 36199], Anderson in Carpenter & De Angelis 2016:2760 [ref. 34618]). Current status: Valid as Lycodes terraenovae Collett 1896. Zoarcidae: Lycodinae. Distribution: Northwestern Atlantic, northeastern Atlantic, South Atlantic. Habitat: marine.

thalassium, Melanostigma Orlovskaya [M. V.] & Balushkin [A. V.] 2019:500 [634], Figs. 1-3 [Voprosy Ikhtiologii v. 59 (no. 5); ref. 37050] Walvis Ridge, southeastern Atlantic, 33°16'2"S, 02°17'4"E, depth 960 - 1080 meters. Holotype: ZIN 56395. Paratypes: ZIN. •Valid as Melanostigma thalassium Orlovskaya & Balushkin 2019. Current status: Valid as Melanostigma thalassium Orlovskaya & Balushkin 2019. Zoarcidae: Gymnelinae. Distribution: Walvis Ridge, southeastern Atlantic. Habitat: marine.

thermophilum, Pachycara Geistdoerfer [P.] 1994:110, Fig. 1 [Cybium v. 18 (no. 2); ref. 21428] Mid-Atlantic Ridge at Snake Pit hydrothermal field, 23°22.94'N, 44°56.09'W, depth 3480 meters. Holotype: MNHN 1993-0255. Paratypes: MNHN 1993-0256 (1). •Valid as Pachycara thermophilum Geistdoerfer 1994 -- (Anderson & Bluhm 1997:223 [ref. 22911], Machida & Hashimoto 2002:1 [ref. 25949], Møller 2003:359 [ref. 26908] as termophilum, Chernova & Geistdorfer 2003:153 [ref. 26887] as thermophylum, Anderson & Fedorov 2004:36 [ref. 27603], Biscoito & Almeida 2004:564 [ref. 27911], Balushkin et al. 2011:1020 [ref. 32449] as Thermophila, Anderson et al. 2016:76 [ref. 34148]). Current status: Valid as Pachycara thermophilum Geistdoerfer 1994. Zoarcidae: Lycodinae. Distribution: Mid-Atlantic Ridge. Habitat: marine.

tohokuensis, Lycenchelys Anderson [M. E.] & Imamura [H.] 2002:355, Figs. 1-2 [Ichthyological Research v. 49 (no. 4); ref. 26499] Off Fukushima Prefecture, Japan, 37°31.6'N, 142°12.4'E to 37°32.9'N, 142°13.3' E, depth 697-709 meters. Holotype: HUMZ 156757. Paratypes: HUMZ 152369 (1), 171668 (1), 178280 (1); NSMT P-64042 [ex HUMZ 171671] (1). •Valid as Lycenchelys tohokuensis Anderson & Imamura 2002 -- (Anderson & Fedorov 2004:20 [ref. 27603], Shinohara & Anderson 2007:63 [ref. 29111], Balushkin et al. 2011:1026 [ref. 32449], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys tohokuensis Anderson & Imamura 2002. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Japan. Habitat: marine.

toyamensis, Petroschmidtia Katayama [M.] 1941:593, Fig. [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 53 (no. 12); ref. 13018] Toyama Bay, Japan. Holotype (unique): whereabouts unknown. •Valid as Lycodes toyamensis (Katayama 1941) -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:30 [ref. 27603], Kim et al. 2006:278 [ref. 29191], Shinohara et al. 2011:52 [ref. 31715]). •Valid as Petroschmidtia toyamensis Katayama 1941 -- (Lindberg & Krasyukova 1975:165 [ref. 7348], Toyoshima in Amaoka et al. 1983:257, 329 [ref. 8379], Toyoshima in Masuda et al. 1984:306 [ref. 6441], Toyoshima 1985:138 [ref. 5722], Nakabo 2000:1030 [ref. 25182], Nakabo 2002:1030 [ref. 26193], Youn 2002:375, 640 [ref. 26218], Antonenko et al. 2004:283 [ref. 27734], Balushkin et al. 2011:1021 [ref. 32449], Nazarkin et al. 2014:177 [ref. 33213], Shinohara et al. 2014:268 [ref. 33330], Parin et al. 2014:427 [ref. 33547]). Current status: Valid as Petroschmidtia toyamensis Katayama 1941. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk and Japan seas. Habitat: marine.

tristichodon, Lycenchelys DeWitt [H. H.] & Hureau [J.-C.] 1980:816, Fig. 10 [Bulletin du Museum National d'Histoire Naturelle, Série 4: Section A: Zoologie, Biologie et Écologie Animales v. 1 (no. 3) (for 1979); ref. 1125] Bellingshausen Sea, west of Adelaide Island, 67°15.6'S, 70°12.0'W, depth 630-650 meters. Holotype (unique): MNHN 1974-0087. Appeared first as name only in Gosztonyi 1977:196 [ref. 6103]. •Valid as Lycenchelys tristichodon DeWitt & Hureau 1980 -- (Anderson 1988:99 [ref. 7304], Pequeño 1989:49 [ref. 14125], Anderson 1990:264 [ref. 21695], Miller 1993:670 [ref. 21297], Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:20 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys tristichodon DeWitt & Hureau 1980. Zoarcidae: Lycodinae. Distribution: Southern Ocean: Bellingshausen Sea, Antarctica. Habitat: marine.

turneri, Lycodes Bean [T. H.] 1879:463 [Proceedings of the United States National Museum v. 1 (no. 59); ref. 16020] Saint Michaels Island, Alaska, U.S.A. Holotype (unique): USNM 21529. Type catalog: Springer & Anderson 1997:22 [ref. 22953]. •Valid as Lycodes turneri Bean 1879 -- (Toyoshima 1985:202 [ref. 5722], Robins & Ray 1986:237 [ref. 23100], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:709 [ref. 25968], Anderson & Fedorov 2004:30 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Mecklenburg et al. 2007:181 [ref. 29448], Balushkin et al. 2011:1015 [ref. 32449], Page et al. 2013:163 [ref. 32708], Parin et al. 2014:423 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Mecklenburg et al. 2016:190 [ref. 34440], Kells et al. 2016:351 [ref. 35888], Mecklenburg et al. 2018:360 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes turneri Bean 1879. Zoarcidae: Lycodinae. Distribution: Arctic and adjacent North Pacific. Habitat: marine.

typica, Lycodes rossi forma Knipowitsch [N. M.] 1906:27 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Not available, infrasubspecific; publication in an available way as a species or subspecies not investigated [Knipowitsch used v. [variety] and f. [forma] -- in his work, the former is considered subspecific, the latter infrasubspecific; see Article 45(g)]. Not treated by Anderson 1994 [ref. 21438]. Also, typica probably means nominal subspecies, "rossi rossi". •Infrasubspecific, in subfamily Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Zoarcidae: Lycodinae.

typica, Lycodes vahlii forma Smitt [F. A.] 1901:26 [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 27 (no. 4); ref. 18732] Not intended as new; treatment of Lycodes vahlii Reinhardt 1831 as a subspecific name vahlii (i.e., typical) -- see Anderson & Fedorov 2004:48 [ref. 27603]. •In the synonymy of Lycodes pallidus Collett 1879 -- (Møller 2001:983 [ref. 25812]). •In the synonymy of Lycodes vahlii Reinhardt 1831 -- (Carl 2002:69 [ref. 26925] as vahli). •Not available, in family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Current status: Synonym of Lycodes vahlii Reinhardt 1831. Zoarcidae: Lycodinae.

unernak, Ophidium Lacepède [B. G. E.] 1800:278, 280 [Histoire naturelle des poissons (Lacepède) v. 2; ref. 2709] Greenland. No types known. Unnecessary replacement name for Gymnelus viride Fabricius 1786. Also appeared in Sonnini 1803:145 [ref. 30473]. •Synonym of Gymnelus viridis (Fabricius 1780) -- (Andriashev 1973:540 [ref. 9214], Anderson 1982:29 [ref. 5520], Anderson & Fedorov 2004:6 [ref. 27603], Parin et al. 2014:394 [ref. 33547]). Current status: Synonym of Gymnelus viridis (Fabricius 1780). Zoarcidae: Gymnelinae. Habitat: marine.

unicolor, Blennius viviparus var. Walbaum [J. J.] 1792:185 [Petri Artedi sueci genera piscium Part 3; ref. 4572] American Ocean. No types known. •Synonym of Zoarces americanus (Bloch & Schneider 1801) -- (Anderson & Fedorov 2004:2 [ref. 27603]). Current status: Synonym of Zoarces americanus (Bloch & Schneider 1801). Zoarcidae: Zoarcinae. Habitat: marine.

unimaculatus, Gymnelis viridis var. Richardson [J.] 1855:371, Pl. 30 [The last of the Arctic voyages in search of Sir J. Franklin; ref. 18631] Northumberland Sound, western Greenland. Holotype (unique): BMNH?. •Nomen dubium in family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Zoarcidae.

uschakovi, Lycenchelys Andriashev [A. P.] 1958:176, Figs. 1 (no.2), 3 [Voprosy Ikhtiologii v. 11; ref. 12042] Northern part of Kurie-Kamchatka Trench, 49°49'N, 157°45'E, depth 3960-4070 meters. Holotype (unique): ZIN 34670. Type catalog: Balushkin et al. 2011:984 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. •Valid as Lycenchelys uschakovi Andriashev 1958 -- (Toyoshima 1985:155 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Anderson & Fedorov 2004:20 [ref. 27603], Balushkin et al. 2011:984 [ref. 32449], Parin et al. 2014:403 [ref. 33547], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys uschakovi Andriashev 1958. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Kuris-Kamchatka Trench. Habitat: marine.

uschakovi, Lycodes Popov [A. M.] 1931:141, Pl. 2 (fig. 7) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] Okhotsk Sea, 58°11.5'N, 148°19.5'N, depth 141 meters. Holotype (unique): ZIN 29986. Type catalog: Balushkin et al. 2011:1015 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. •Valid as Lycodes uschakovi Popov 1931 -- (Lindberg & Krasyukova 1975:147 [ref. 7348], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:205 [ref. 5722], Anderson 1994:119 [ref. 21438], Sokolovskaya et al. 1998:12 [ref. 24670], Nakabo 2000:1037 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1037 [ref. 26193], Fedorov et al. 2003:117 [ref. 27579], Anderson & Fedorov 2004:30 [ref. 27603], Balushkin et al. 2011:1015 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:423 [ref. 33547]). Current status: Valid as Lycodes uschakovi Popov 1931. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk and Japan seas. Habitat: marine.

vachonii, Lycodes Vladykov [V. D.] & Tremblay [J.-L.] 1936:23, Pls. 1 (fig. 3, 4), 2 (fig. 5) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] St. Lawrence River estuary, Canada. Syntypes: (5) whereabouts unknown. •Synonym of Lycodes esmarki [sic] Collett 1875, but probably a valid subspecies -- (Andriashev in Whitehead et al. 1986:1138 [ref. 13677]). •Synonym of Lycodes esmarkii Collett 1875 -- (Scott & Scott 1988:407 [ref. 25518] as esmarki, Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:24 [ref. 27603], Parin et al. 2014:410 [ref. 33547]). Current status: Synonym of Lycodes esmarkii Collett 1875. Zoarcidae: Lycodinae. Habitat: marine.

vahlii, Lycodes Reinhardt [J. C. H.] 1831:18 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1830-31; ref. 6533] Julianehåb, Greenland. Holotype: ZMUC 9 (skeleton). Type catalog: Nielsen 1974:78 [ref. 9588]. Also appeared in Reinhardt 1837:114, 123, 195, Pl. 5 [32, 41, 113, Pl. 5 of separate] [ref. 3691]. •Valid as Lycodes vahlii Reinhardt 1831 -- (Andriashev 1973:542 [ref. 9214] as vahli, Andriashev in Whitehead et al. 1986:1142 [ref. 13677] as vahli, Scott & Scott 1988:411 [ref. 25518], McAllister 1990:163 [ref. 14674] as vahli, Anderson 1994:119 [ref. 21438], Miki in Okamura et al. 1995:210 [ref. 22531], Møller 2000:847 [ref. 24585], Carl 2002:69 [ref. 26925] as vahli, Anderson & Fedorov 2004:31 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Møller et al. 2010:63 [ref. 31169] as vahli on p. 61, Balushkin et al. 2011:1016 [ref. 32449], Page et al. 2013:163 [ref. 32708], Afonso et al. 2016:67, 213 [ref. 36199], Mecklenburg et al. 2018:362 [ref. 36370]). Current status: Valid as Lycodes vahlii Reinhardt 1831. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and northwestern Atlantic. Habitat: marine.

variegatus, Lycodes Günther [A.] 1862:322 [Catalogue of the fishes in the British Museum v. 4; ref. 1969] Falkland Islands. Syntypes: BMNH 1842.2.12.1 (2). •Synonym of Iluocoetes fimbriatus Jenyns 1842 -- (Gosztonyi 1977:213 [ref. 6103], Anderson 1994:117 [ref. 21438], Anderson & Fedorov 2004:14 [ref. 27603]). Current status: Synonym of Iluocoetes fimbriatus Jenyns 1842. Zoarcidae: Lycodinae. Habitat: marine.

vermiformis, Lycodonus Barnard [K. H.] 1927:74 [Annals and Magazine of Natural History (Series 9) v. 20 (no. 115) (art. 8); ref. 193] Off Cape Point, South Africa, depth 460-630 fathoms [841-1152 meters]. Syntypes: BMNH 1927.12.6.7 [ex SAM] (1), SAM 12794 (2) depth 500-550 fathoms. Also appeared as new in Barnard 1927:873, Pl. 35 (fig. 3) [ref. 194]. •Valid as Lycodonus vermiformis Barnard 1927 -- (Gosztonyi 1981:156 [ref. 8596], Toyoshima 1985:143 [ref. 5722], Anderson 1986:343 [ref. 5633], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:32 [ref. 27603], Balushkin et al. 2011:1018 [ref. 32449]). Current status: Valid as Lycodonus vermiformis Barnard 1927. Zoarcidae: Lycodinae. Distribution: Southeastern Atlantic: South Africa. Habitat: marine.

verrillii, Lycodes Goode [G. B.] & Bean [T. H.] 1877:474 [American Journal of Science and Arts (Series 3) v. 14; ref. 18128] 30 miles south of Cape Negro, Nova Scotia, Canada, depth 90 fathoms; 27 miles south of Chebucto Head, Halifax, Canada, depth 101 fathoms. Lectotype: USNM 21013. Paralectotypes: USNM 336784 [ex USNM 21013] (orig. 5, now 2), 21015 (1); YPM 3633 (1, not found in 1991). Type catalog: Moore & Boardman 1991:27 [ref. 16741] as verrilli, Springer & Anderson 1997:22-23 [ref. 22953] with extensive comments. Lectotype selected by Springer & Anderson 1997:23 [ref. 22953]. •Valid as Lycenchelys verrillii (Goode & Bean 1877) -- (Robins & Ray 1986:236 [ref. 23100] as verrilli, Scott & Scott 1988:405 [ref. 25518] as verrilli, McAllister 1990:158 [ref. 14674] as verrilli, Anderson 1994:118 [ref. 21438], Møller 1999:327 [ref. 24052], Klein-MacPhee & Collette in Collette & Klein-MacPhee 2002:467 [ref. 26158], Moore et al. 2003:228 [ref. 27076], Anderson & Fedorov 2004:21 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Balushkin et al. 2011:984 [ref. 32449], Page et al. 2013:162 [ref. 32708]). Current status: Valid as Lycenchelys verrillii (Goode & Bean 1877). Zoarcidae: Lycodinae. Distribution: Western North Atlantic. Habitat: marine.

virgata, Puzanovia Fedorov [V. V.] 1982:77, Fig. [Trudy Zoologicheskogo Instituta, Akademii Nauk SSSR v. 114; ref. 5252] Western Pacific, off northern Kuril Islands, 48°24'30"N, 153°49'12"E, depth 480-500 meters. Holotype: ZIN 44079. Paratypes: ZIN 44080 (1). Type catalog: Balushkin et al. 2011:971 [ref. 32449], Balushkin et al. 2012:167 [ref. 33667]. •Valid as Puzanovia virgata Fedorov 1982 -- (Anderson 1994:120 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Anderson & Fedorov 2004:9 [ref. 27603], Balushkin et al. 2011:971 [ref. 32449], Parin et al. 2014:428 [ref. 33547]). Current status: Valid as Puzanovia virgata Fedorov 1982. Zoarcidae: Gymnelinae. Distribution: Western North Pacific. Habitat: marine.

viride, Ophidium Fabricius [O.] 1780:141 [Fauna groenlandica, systematice sistens animalia Groenlandiae occidentalis, etc.; ref. 17464] Southwestern Greenland. Neotype: ZMUC P761096. Neotype designated by Anderson 1982:30 [ref. 5520]. •Valid as Gymnelis viridis (Fabricius 1780) -- (Robins & Ray 1986:236 [ref. 23100]). •Valid as Gymnelus viridis (Fabricius 1780) -- (Andriashev 1972:540 [ref. 7214], Anderson 1982:29 [ref. 5520], Toyoshima in Masuda et al. 1984:306 [ref. 6441], Andriashev in Whitehead et al. 1986:1131 [ref. 13677], Scott & Scott 1988:404 [ref. 25518], McAllister 1990:165 [ref. 14674], Anderson 1994:117 [ref. 21438], Koyanagi in Okamura et al. 1995:189 [ref. 22531], Chernova 1998:186 [ref. 24672], Chernova 1999:9 [ref. 24863], Chernova 1999:306 [ref. 24868], Chernova 2000:8 [ref. 24885], Nakabo 2000:1040 [ref. 25182], Mecklenburg et al. 2002:690 [ref. 25968], Nakabo 2002:1040 [ref. 26193], Anderson & Fedorov 2004:6 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Møller et al. 2010:59 [ref. 31169], Balushkin et al.2011:966 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:394 [ref. 33547], Mecklenburg & Anderson 2015:263 [ref. 33742], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:61, 162 [ref. 36199], Kells et al. 2016:270 [ref. 35888], Mecklenburg et al. 2016:155, 157 [ref. 34440], Mecklenburg et al. 2018:311 [ref. 36370]). Current status: Valid as Gymnelus viridis (Fabricius 1780). Zoarcidae: Gymnelinae. Distribution: Arctic Ocean including Bering Sea, North Atlantic and western North Pacific. Habitat: marine.

vitiazi, Lycenchelys Andriashev [A. P.] 1955:371 [Trudy Instituta Zoologii / Akademiia Nauk, Azerbaidzhanskoi SSR. v. 18; ref. 12261] Kuril-Kamchatka Trench near Paramushiro Island, 50°00'N, 157°40'E, depth 2450 meters. Holotype (unique): ZIN 33747. Type catalog: Balushkin et al. 2011:984 [ref. 32449], Balushkin et al. 2012:167 [ref. 33667]. •Valid as Lycenchelys vitiazi Andriashev 1955 -- (Toyoshima 1985:164 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Anderson & Fedorov 2004:21 [ref. 27603], Balushkin et al. 2011:984 [ref. 32449], Parin et al. 2014:404 [ref. 33547], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys vitiazi Andriashev 1955. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

vitiazi, Melanostigma (Bandichthys) Parin [N. V.] 1980:167 [150], Figs. 1-2 [Voprosy Ikhtiologii v. 19 (no. 1), 1980; ref. 3363] Banda Sea, 6°39'S, 126°37'E, depth 0-890 meters. Holotype: ZIN 44000. Type catalog: Balushkin et al. 2011:970 [ref. 32449], Balushkin et al. 2012:167 [ref. 33667]. On p. 150 of English translation. •Valid as Melanostigma vitiazi Parin 1980 -- (Anderson 1988:66 [ref. 7304], Paulin et al. 1989:211 [ref. 24556], Anderson 1990:5 [ref. 16143], Anderson 1990:271 [ref. 21695], Anderson 1994:120 [ref. 21438], Anderson 2001:3494 [ref. 26279], Anderson & Fedorov 2004:8 [ref. 27603], Duhamel et al. 2005:324[ref. 32059], Anderson & Møller 2007:183 [ref. 29401], Balushkin et al. 2011:970 [ref. 32449], Møller & Anderson 2015:1430 [ref. 34289], Balushkin & Moganova 2017:131 [ref. 35279], Balushkin & Moganova 2018:136 [ref. 35877], Balushkin & Orlovskaya 2019:136 [ref. 36664], Balushkin 2019:286 [ref. 36807]). Current status: Valid as Melanostigma vitiazi Parin 1980. Zoarcidae: Gymnelinae. Distribution: Banda Sea, Indo-West Pacific. Habitat: marine.

viviparus, Blennius Linnaeus [C.] 1758:258 [Systema Naturae, Ed. X v. 1; ref. 2787] European seas. Possible syntypes: BMNH 1853.11.12.122 [Gronovius coll.] (1, skin), LS 28 (left half-skin). Type catalog: Wheeler 1958:235 [ref. 13434], Wheeler 1985:39 [ref. 20705]. •Valid as Zoarces viviparus (Linnaeus 1758) -- (Berg 1949:1053 [ref. 20662], Andriashev 1973:540 [ref. 9214], Andriashev in Whitehead et al. 1986:1149 [ref. 13677], Anderson 1994:120 [ref. 21438], Reshetnikov et al. 1997:751 [ref. 24702], Reshetnikov 1998:124 [ref. 23570], Anderson & Fedorov 2004:2 [ref. 27603], Bogutskaya & Naseka 2004:208 [ref. 28183], Chereshev et al. 2007:555 [ref. 29371], Fricke 2007:28 [ref. 30577], Voskoboinikova et al. 2010:704 [ref. 31095], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin 2011:956 [ref. 32449], Parin et al. 2014:430 [ref. 33547], Mecklenburg et al. 2018:368 [ref. 36370]). Current status: Valid as Zoarces viviparus (Linnaeus 1758). Zoarcidae: Zoarcinae. Distribution: Eastern North Atlantic and adjacent Arctic Region. Habitat: marine.

volki, Lycenchelys Andriashev [A. P.] 1955:369, Fig. 2 (no. 6), Figs. 13-14 [Trudy Instituta Zoologii / Akademiia Nauk, Azerbaidzhanskoi SSR. v. 18; ref. 12261] About 130 miles north of Mednyi Island, southwestern Bering Sea, 57°03'N, 168°29'E, depth 3940 meters. Holotype (unique): ZIN 32964. Type catalog: Balushkin et al. 2011:984 [ref. 32449], Balushkin et al. 2012:167 [ref. 33667]. •Valid as Lycenchelys volki Andriashev 1955 -- (Toyoshima 1985:152 [ref. 5722], Anderson 1994:118 [ref. 21438], Anderson 1995:106 [ref. 21928], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:697 [ref. 25968], Anderson & Fedorov 2004:21 [ref. 27603], Balushkin et al. 2011:984 [ref. 32449], Parin et al. 2014:404 [ref. 33547], Thiel et al. 2018:53 [ref. 35743]). Current status: Valid as Lycenchelys volki Andriashev 1955. Zoarcidae: Lycodinae. Distribution: North Pacific: Bering Sea. Habitat: marine.

wilkesi, Lycenchelys Anderson [M. E.] 1988:100, Fig. 37 [Antarctic Research Series, Washington DC. v. 47; ref. 7304] Bransfield Strait, Antarctica, 62°07'S, 55°58'W, depth 1113-1153 meters. Holotype (unique): LACM 10476-2. •Valid as Lycenchelys wilkesi Anderson 1988 -- (Anderson 1990:265 [ref. 21695], Miller 1993:671 [ref. 21297], Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:21 [ref. 27603], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys wilkesi Anderson 1988. Zoarcidae: Lycodinae. Distribution: Southern Ocean: Antarctic Peninsula. Habitat: marine.

xanthoptera, Lycenchelys Anderson [M. E.] 1991:152, Figs. 1-2 [Cybium v. 15 (no. 2); ref. 20019] Weddell Sea, off Kapp Norvegia, Queen Maud Land, Antarctica, 71°06.2'S, 12°53.8'W to 71°05.7'S, 12°58.4'W, depth 771-793 meters. Holotype: MNHN 1990-1023. Paratypes: ISH 1-1991 (1); LACM 45053-1 (1); MNHN 1999-1024 (1), 1990-1025 (1), 1990-1026 (1). •Valid as Lycenchelys xanthoptera Anderson 1991 -- (Anderson & Balanov 2000:1056 [ref. 25023], Anderson & Fedorov 2004:21 [ref. 27603], Struthers & Møller 2009:65 [ref. 30403], Balushkin et al. 2011:1026 [ref. 32449]). Current status: Valid as Lycenchelys xanthoptera Anderson 1991. Zoarcidae: Lycodinae. Distribution: Southern Ocean: Weddell Sea and Ross Sea, Antarctica. Habitat: marine.

yabei, Davidijordania Anderson [M. E.] & Imamura [H.] 2008:63, Figs. 5-6 [Bulletin of the National Museum of Nature and Science (Ser. A) Supplement No. 2; ref. 29573] Off Usujiri, Hokkaido, Japan. Holotype: HUMZ 101869. Paratypes: HUMZ 46074-78 (5), 95339 (1), 97358 (1), 133724 (1); NSMT-P 76023 [ex HUMZ 46072] (1), 76024 [ex HUMZ 46073] (1). •Valid as Davidijordania yabei Anderson & Imamura 2008 -- (Shinohara & Kim 2009:231 [ref. 31427]). Current status: Valid as Davidijordania yabei Anderson & Imamura 2008. Zoarcidae: Gymnelinae. Distribution: Off Usujiri, Hokkaido, Japan, and Olga Bay, Russia (Sea of Japan). Habitat: marine.

yamatoi, Lycodes Toyoshima [M.] 1985:208, Fig. 49 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Sea of Japan off Kasumi, Hyogo Prefecture, Japan. Holotype: HUMZ 41094. Paratypes: HUMZ 42480 (1), 53191 (1), 53597-98 (2), 53610-11 (2), 53618 (1), 53620 (1), 53630 (1), 53634 (1), 53658 (1), 53668 (1), 53672 (1), 53682 (1), 53684 (1), 53726 (1), 53734-35 (2), 53737-38 (2), 53740-42 (3), 53798 (1), 53805 (1), 53817 (1), 53856 (1), 53866 (1), 53911 (1), 53917-18 (2), 53921 (1), 53923 (1). The species matsubarai and yamatoi were described in the same journal article; the first name was kept as the valid one in Parin et al. 2014:415 [ref. 33547], but see matsubarai. •Valid as Lycodes yamatoi Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Møller 2001:111 [ref. 25378], Nakabo 2002:1039 [ref. 26193], Anderson & Fedorov 2004:31 [ref. 27603], Nazarkin 2010:603 [ref. 31019], Shinohara et al. 2011:52 [ref. 31715], Balushkin et al. 2011:1016 [ref. 32449]). •Synonym of Lycodes matsubarai Toyoshima 1985 -- (Nazarkin & Shinohara 2012:33 [ref. 32106], Parin et al. 2014:415 [ref. 33547]). Current status: Synonym of Lycodes matsubarai Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk and Japan seas (if valid). Habitat: marine.

ygreknotatus, Lycodes Schmidt [P. Yu.] 1950:92 [79] [Transactions of the Pacific Committee of the Academy of Sciences of the U.S.S.R. v. 6; ref. 12471] Shantar Islands, north of Cape Aleksandir, northwestern part of Okhotsk Sea, Russia, depth 135 meters. Holotype (unique): ZIN 24843. Type catalog: Balushkin et al. 2011:1017 [ref. 32449], Balushkin et al. 2012:167 [ref. 33667]. On p. 79 of English translation. Originally ygrek-notatus. Appeared first without description as Schmidt in Taranetz 1937:163 [ref. 13384]. •Valid as Lycodes ygreknotatus Schmidt 1950 -- (Lindberg & Krasyukova 1975:155 [ref. 7348], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:210 [ref. 5722], Anderson 1994:119 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Møller 2001:111 [ref. 25378], Nakabo 2002:1039 [ref. 26193], Fedorov et al. 2003:117 [ref. 27579] dated 1937, Anderson & Fedorov 2004:31 [ref. 27603], Nazarkin 2010:9 [ref. 30733], Balushkin et al. 2011:1017 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:424 [ref. 33547]). Current status: Valid as Lycodes ygreknotatus Schmidt 1950. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea and Sea of Japan. Habitat: marine.

zesta, Bothrocara Jordan [D. S.] & Fowler [H. W.] 1902:749, Fig. 3 [Proceedings of the United States National Museum v. 25 (no. 1303); ref. 2459] Sagami Bay, Japan, Albatross station 3696, depth 501-749 fathoms. Holotype: USNM 50576 (19 in., 482 mm). Paratypes: CAS-SU 7122 (1), 7256 (1); USNM 331717 [ex USNM 50576] (1). Type catalog: Böhlke 1953:102 [ref. 12291], Springer & Anderson 1997:23 [ref. 22953]. Types: Shinohara & Williams 2006:561 [ref. 31656]. Original should have been Bothrocara zestum. Paratype incorrectly listed as SU 7256 in Böhlke 1953 [ref. 12291]. •Valid as Lycogramma zesta (Jordan` & Fowler 1902) -- (Lindberg & Krasyukova 1975:184 [ref. 7348]). •In genus Bothrocara but status uncertain -- (Anderson 1994:116 [ref. 21438]). •Valid as Bothrocara zestum Jordan & Fowler 1902 -- (Balushkin et al. 2011:974 [ref. 32449], Shinohara et al. 2014:264 [ref. 33330], Parin et al. 2014:386 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331]). Current status: Valid as Bothrocara zestum Jordan & Fowler 1902. Zoarcidae: Lycodinae. Distribution: North Pacific: Izu Islands (Japan) and Sea of Okhotsk east to Gulf of Alaska. Habitat: marine.

zinorum, Aiakas Anderson [M. E.] & Gosztonyi [A. E.] 1991:2, Figs. 1-2 [Ichthyological Bulletin of the J. L. B. Smith Institute of Ichthyology No. 55; ref. 19348] Argentine slope, 49°16'00"S, 57°01'09"W, depth 630-650 meters. Holotype (unique): ZIN 39893. Type catalog: Balushkin et al. 2011:972 [ref. 32449], Balushkin et al. 2012:167 [ref. 33667]. •Valid as Aiakas zinorum Anderson & Gosztonyi 1991 -- (Anderson 1994:116 [ref. 21438], Anderson & Fedorov 2004:9 [ref. 27603], Balushkin et al. 2011:972 [ref. 32449]). Current status: Valid as Aiakas zinorum Anderson & Gosztonyi 1991. Zoarcidae: Lycodinae. Habitat: marine.

zoarchus, Lycodes Goode [G. B.] & Bean [T. H.] 1896:308, Pls. 79, 81 (figs. 283) [Special Bulletin U. S. National Museum No. 2; ref. 1848] Off Nova Scotia, Canada, 44°46'30"N, 59°55'45"W, Albatross [station 2499], depth 130 fathoms. Holotype: USNM 39298 (366 mm). Paratypes: USNM 336785 [ex USNM 39298] (3, poor condition), 39299 (1, poor condition). Type catalog: Springer & Anderson 1997:23-24 [ref. 22953]. •Synonym of Lycodes vahlii Reinhardt 1831 -- (Anderson 1994:119 [ref. 21438], Carl 2002:69 [ref. 26925] as vahli, Anderson & Fedorov 2004:31 [ref. 27603]). Current status: Synonym of Lycodes vahlii Reinhardt 1831. Zoarcidae: Lycodinae. Habitat: marine.