Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the genus Barbatula: [ 46 ] records

altayensis, Barbatula Zhu [S.-Q.] 1992:241 [246], Figs. 1-2 [Acta Zootaxonomica Sinica v. 17 (no. 2); ref. 21064] Kelang He River, trib of the Ertix River, near Altay City, 47°52'N, 88°06'E, Xinjiang, China. Holotype: NIGlAS 888006. Paratypes: NIGLAS 888001-05 (5), 888007 (1), 888009 (1). English description on p. 246. •Valid as Orthrias altayensis (Zhu 1992) -- (Prokofiev 2003:726 [ref. 27494], Prokofiev 2007:14 [ref. 30343]). •Valid as Barbatula altayensis Zhu 1992 -- (Zhu 1995:108 [ref. 25213], Kottelat 2006:88 [ref. 28831], Cao et al. 2012:236 [ref. 32335], Kottelat 2012:77 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Zhang et al. 2016:127 [ref. 34477], Prokofiev 2016:396 [ref. 34693], Prokofiev 2018:273 [ref. 35878] as cf. altayensis). Current status: Valid as Barbatula altayensis Zhu 1992. Nemacheilidae. Distribution: China. Habitat: freshwater.

anglicana, Nemacheilus barbatulus forma Băcescu-Meşter [L.] 1967:369 [Travaux du Muséum d'Histoire Naturelle "Grigore Antipa" v. 7; ref. 7940] Not available, infrasubspecific (a form after 1960); from Cambridge River, England. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Kottelat 2012:78 [ref. 32367]). Infrasubspecific. Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae. Habitat: freshwater.

barbatula, Cobitis Linnaeus [C.] 1758:303 [Systema Naturae, Ed. X v. 1; ref. 2787] Fresh waters of Europe and Asia. Syntypes: NRM 68 (1 or 3). Probably 3 syntypes with no geographic origin, Norén et al. 2018:64 [ref. 35742]. •Valid as Cobitis barbatula Linnaeus 1758 -- (Herrera 1896:25 [ref. 39336]). •Valid as Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:868 [ref. 20661] as Nemachilus, Lelek 1987:256 [ref. 22962] as Noemacheilus, Chereshnev 1996:601 [ref. 24659] as Noemacheilus). •Valid as Noemacheilus barbatulus (Linnaeus 1758), subspecies barbatulus -- (Mitrofanov & Dukravec 1989:58 [ref. 37748]). •Valid as Orthrias barbatulus (Linnaeus 1758) -- (Bănărescu & Nalbant 1995:449 [ref. 23187], Prokofiev 2003:726 [ref. 27494], Soric 2000:61 [ref. 28770]). •Valid as Orthrias barbatulus (Linnaeus 1758) and subspecies barbatulus (Linnaeus 1758) -- (Prokofiev 2007:15 [ref. 30343]). •Valid as Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Reshetnikov et al. 1997:743 [ref. 24702], Vasil'eva 1998:94 [ref. 23591], Novikov et al. 2000:161 [ref. 40929], Bogutskaya et al. 2001:45 [ref. 26178], Hanel 2003:55 [ref. 27894], Bogutskaya & Naseka 2004:112 [ref. 28183], Economidis 2005:67 [ref. 28213], Vassilev & Pehlivanov 2005:172 [ref. 28369], Hanel & Lusk 2005:290 [ref. 28804], Fricke et al. 2007:50 [ref. 29533], Kottelat & Freyhof 2007:329 [ref. 29996], Kottelat 2012:77 [ref. 32367], Parin et al. 2014:102 [ref. 33547], Barbieri et al. 2015:75 [ref. 35029], Çiçek et al. 2015:148 [ref. 35151], Norén et al. 2018:59 [ref. 35742], Tutman et al. 2018:220 [ref. 35876], Ćaleta et al. 2019:149 [ref. 37026], Gauliard et al. 2019:175 [ref. 36787], Lorenzoni et al. 2019:251 [ref. 37108], Pozdeev et al. 2022:6 [ref. 39511], Dyldin et al. 2023:[7] [ref. 40328], Nagy et al. 2023:356 [ref. 40348]). Current status: Valid as Barbatula barbatula (Linnaeus 1758). Nemacheilidae. Distribution: Northern and eastern Europe and western Asia. Habitat: freshwater, brackish.

blackiana, Nemacheilus barbatulus forma Băcescu-Meşter [L.] 1967:369 [Travaux du Muséum d'Histoire Naturelle "Grigore Antipa" v. 7; ref. 7940] Regarded as infrasubspecific by Kottelat 1997:93 [ref. 22952]. Not available, infrasubspecific; from Black Beck River, England. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Kottelat 2012:78 [ref. 32367]). Infrasubspecific. Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae.

caucasicus, Nemachilus barbatulus var. Berg [L. S.] 1898:67 [26] [Izvestija Kavkazskago Muzeja = Mitteilungen des Kaukasischen Museums = Bulletin du Musée du Caucase v. 1 (no. 3); ref. 20653] Argun River, Terek River basin, Russia. Syntypes: (many). Original genus is correctly Nemacheilus. On p. 26 of Russian issue. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies as described -- (Berg 1949:872 [ref. 20661] as Nemachilus, Lelek 1987:256 [ref. 22962] as Noemacheilus). •Synonym of Barbatula barbatula (Linnaeus 1758), but a valid subspecies -- (Reshetnikov et al. 1997:743 [ref. 24702], Vasil'eva 1998:95 [ref. 23591], Gabrielyan 2001:28 [ref. 25867], Ninua & Japoshvili 2008:167 [ref. 31251], Kottelat 2012:77 [ref. 32367] with question, Parin et al. 2014:103 [ref. 33547] with question). Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae. Habitat: freshwater, brackish.

cobdonensis, Nemacheilus Gundriser [A. N.] 1973:253, Fig. 4 [Siberian aquatic resources and their prospective fishery utilization; ref. 28184] Tuva, Lakes Khintiktig-Khol and Ak-Khol Khos and River Mogen-Buren, basin of River Kobdo, Russia. Syntypes: probably lost. •Valid as Triplophysa cobdonensis (Gundriser 1973) -- (Bogutskaya & Naseka 2004:116 [ref. 28183]). •Questionably a synonym of Barbatula compressirostris (Warpachowski 1897) -- (Kottelat 2006:51 [ref. 28831]). •Questionably valid as Barbatula cobdonensis (Gundriser 1973) -- (Kottelat 2012:78 [ref. 32367], Prokofiev 2015:460 [ref. 34065], Prokofiev 2016:401 [ref. 34693], Dyldin et al. 2020:87 [ref. 37569]). •Valid as Barbatula cobdonensis (Gundriser 1973) -- (Dyldin et al. 2023:[8] [ref. 40328]). Current status: Uncertain as Barbatula cobdonensis (Gundriser 1973). Nemacheilidae. Distribution: North Asia: Upper Chovd River drainage (Russia). Habitat: freshwater.

compressirostris, Nemachilus Warpachowski [N. A.] 1897:270, Pl. 11 (figs. 1-3) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 2; ref. 4515] Lakes in northwestern Mongolia [possibly an error, probably a lake in the vicinity of Kosh-Agach]. Lectotype: ZIN 11298 (158 mm). Paralectotypes: ZIN 11298 (1). Original genus is correctly Nemacheilus. For remarks on systematics see Kottelat 2006:51 [ref. 28831]. Lectotype selected by Prokofiev 2016:392 [ref. 34693]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Barbatula toni (Dybowski 1869) -- (Vasil'eva 1998:95 [ref. 23591], Sideleva 2001:54 [ref. 26181], Sideleva 2003:11 [ref. 27815], Prokofiev 2016:392 [ref. 34693] with doubt). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Valid as Barbatula compressirostris (Warpachowski 1897) -- (Kottelat 2006:51 [ref. 28831], Cao et al. 2012:236 [ref. 32335], Kottelat 2012:78 [ref. 32367] with question, Prokofiev 2015:460 [ref. 34065] with question, Prokofiev 2018:273 [ref. 35878], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula compressirostris (Warpachowski 1897). Nemacheilidae. Distribution: East Asia: northwestern Mongolia and Tuva (Russia). Habitat: freshwater.

conilobus, Barbatula Prokofiev [A. M.] 2016:657, Figs. 1, 2a, 3a, 4, 5a [Voprosy Ikhtiologii v. 56 (no. 6); ref. 35025] Bogd River, a tributary of the Zavkhan River near Uliastai village, Mongolia, 47°43'59.2"N, 96°49'08.9"E, elevation 1749 meters. Holotype: ZMMU 23813. Paratypes: ZMMU. Plus additional non-type specimens. •Valid as Barbatula conilobus Prokofiev 2016 -- (Prokofiev 2018:273 [ref. 35878]). Current status: Valid as Barbatula conilobus Prokofiev 2016. Nemacheilidae. Distribution: Zavkhan River drainage, Mongolia. Habitat: freshwater.

dgebuadzei, Orthrias Prokofiev [A. M.] 2003:730, Fig. 2 [Voprosy Ikhtiologii v. 43 (no. 6); ref. 27494] Zag River basin of Baidrag-River, central Mongolia. Holotype: ZIN 52051. Paratypes: ZIN 52052 (18), 52052/1 (5), 52052/2 (3), 52052/3 (1). •Valid as Orthrias dgebuadzei Prokofiev 2003 -- (Prokofiev 2007:77 [ref. 30343], Cao et al. 2012:236 [ref. 32335]). •Valid as Barbatula dgebuadzei (Prokofiev 2003) -- (Kottelat 2006:52 [ref. 28831], Ocock et al. 2006:28 [ref. 28933], Kottelat 2012:78 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:396 [ref. 34693], Prokofiev 2018:273 [ref. 35878]). Current status: Valid as Barbatula dgebuadzei (Prokofiev 2003). Nemacheilidae. Distribution: Baidrag-Gol River drainage, Gobi Lakes Valley, Mongolia. Habitat: freshwater.

dsapchynensis, Barbatula Prokofiev [A. M.] 2016:661, Figs. 2b, 3b, 5b, 6 [Voprosy Ikhtiologii v. 56 (no. 6); ref. 35025] Bogd River, tributary of Zavkhan River near Uliastai village, Mongolia, 47°43'59.2"N, 96°49'08.9"E, elevation 1749 meters. Holotype: ZMMU 23816. Paratypes: ZMMU. Plus additional non-type specimens. •Valid as Barbatula dsapchynensis Prokofiev 2016 -- (Prokofiev 2018:273 [ref. 35878]). Current status: Valid as Barbatula dsapchynensis Prokofiev 2016. Nemacheilidae. Distribution: North Asia: Zavkhan River drainage (Mongolia). Habitat: freshwater.

emuensis, Barbatula Chen [H.], Zhang [H.], Chen [Y.-X.] & Freyhof [J.] 2019:19, Figs. 19-22 [Zootaxa 4565 (no. 1); ref. 36569] Zhu’Erduo River near Emu, upper Mudan River flowing into Songhua River of the middle Heilongjiang River drainage, Jilin Province, China, 43°75′N, 128°15′E. Holotype: HU 1608142. Paratypes: HU. Plus additional non-type material. •Valid as Barbatula emuensis Chen, Zhang, Chen & Freyhof 2019. Current status: Valid as Barbatula emuensis Chen, Zhang, Chen & Freyhof 2019. Nemacheilidae. Distribution: Zhu’Erduo, Erdaobai, Hun and Pushi River drainages, northern China (and possibly North Korea). Habitat: freshwater.

erythrinna, Nemacheilus barbatulus aberr. Berg [L. S.] 1933:550 [Les poissons des eaux douces de l'U.R.S.S. Part 2; ref. 279] Original not seen. Infrasubspecific, not available. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Kottelat 2012:78 [ref. 32367]). Infrasubspecific. Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae.

fowleri, Barbatula toni Nichols [J. T.] 1925:3 [American Museum Novitates No. 171; ref. 3178] Eastern Tombs, Chihli, Hopeh [Hebei] Province, China. Now: Zunhua, 40°11'N, 117°58'E, China. Holotype: AMNH 8409. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Barbatula barbatula (Linnaeus 1758), subspecies nuda (Bleeker 1868) -- (Zhu 1989:30 [ref. 17744]). •Synonym of Barbatula toni (Dybowski 1869), but a valid subspecies as described -- (Vasil'eva 1998:96 [ref. 23591]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula nuda (Bleeker 1869) with question -- (Kottelat 2012:78 [ref. 32367]). •Synonym of Barbatula toni (Dybowski 1869) -- (Kottelat 2006:53 [ref. 28831], Prokofiev 2015:461 [ref. 34065]). •Synonym of Barbatula pechiliensis (Fowler 1899) -- (Chen et al. 2019:15 [ref. 36569]). Current status: Synonym of Barbatula pechiliensis (Fowler 1899). Nemacheilidae. Habitat: freshwater.

fuerstenbergii, Cobitis Fitzinger [L. J. F. J.] in Heckel & Kner 1857:301 [Die Süsswasserfische der Österreichischen Monarchie; ref. 2078] Not available, no distinguishing features, as fürstenbergii in synonymy of Cobitis barbatula. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). Nomen Nudum. Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae.

furstenbergii, Cobitis Bonaparte [C. L.] 1846:26 [Atti della Settima Adunanza degli Scienziati Italiani Sesta Riunione, Milano 7a Adunanza, Napoli, Part 2; ref. 519] Name only, not available. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Kottelat 2012:77 [ref. 32367]). Nomen Nudum. Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae.

gibba, Barbatula Cao [L.], Causse [R.] & Zhang [E] 2012:244, Figs. 1C [Zootaxa No. 3586; ref. 32335] Dali-Nur Lake in Hexigten Banner, Inner Mongolia, China. Holotype: IHB 76X2566. Paratypes: IHB 76X2552-4, 76X2558-9, 76X2562, 76X2564, 76X2567-8, 76X2574, 76X2576, 76X2584-5 (total 13). •Synonym of Barbatula toni (Dybowski 1869) -- (Prokofiev 2015:461 [ref. 34065]). •Valid as Barbatula gibba Cao, Causse & Zhang 2012 -- (Zhang et al. 2016:127 [ref. 34477], Chen et al. 2019:10 [ref. 36569]). Current status: Valid as Barbatula gibba Cao, Causse & Zhang 2012. Nemacheilidae. Distribution: Dali-Nur Lake, Inner Mongolia, China. Habitat: freshwater.

golubtsovi, Orthrias Prokofiev [A. M.] 2003:727 or 698, Fig. 1 [Voprosy Ikhtiologii v. 43 (no. 6); ref. 27494] Chedi-Tei River, about 60 km wes of Magur-Aksk, Kobdo River drainage, Tuva Republic, Mongolia. Holotype: ZIN 52251. Paratypes: ZIN 52252 (15), 52252/1-4 (5), 52253 ( 1); ZMMGY 20413 (5). •Questionably a synonym of Barbatula compressirostris (Warpachowski 1897) -- (Kottelat 2006:51 [ref. 28831]). •Valid as Orthrias golubtsovi Prokofiev 2003 -- (Prokofiev 2007:68 [ref. 30343]). •Valid as Barbatula golubtsovi (Prokofiev 2003) -- (Cao et al. 2012:236 [ref. 32335], Kottelat 2012:78 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:396 [ref. 34693], Prokofiev 2018:273 [ref. 35878], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula golubtsovi (Prokofiev 2003). Nemacheilidae. Distribution: East Asia: Chovd River drainage (Mongolia and Russia). Habitat: freshwater.

hispanica, Nemacheilus barbatulus forma Băcescu-Meşter [L.] 1967:369 [Travaux du Muséum d'Histoire Naturelle "Grigore Antipa" v. 7; ref. 7940] Not available, infrasubspecific (forma after 1960); from Nervion R. at Durango and Tajo River, Spain. See Nemacheilus barbatulus hispanica Lelek 1987. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Kottelat 2012:79 [ref. 32367]). •In the synonymy of Barbatula quignardi (Băcescu-Meşter 1967) -- (Gauliard et al. 2019:176 [ref. 36787]). Infrasubspecific. Current status: Synonym of Barbatula quignardi (Băcescu-Meşter 1967). Nemacheilidae.

hispanica, Nemacheilus barbatulus Lelek [A.] 1987:256 [The freshwater fishes of Europe. Threatened fishes of Europe v. 9; ref. 22962] Nervion River at Durango and Tajo River, Spain. Syntypes: MGAB (11). Available by reference to infrasubspecific hispanica Bǎcescu-Meşter 1967, with Bǎcescu-Meşter specimens becoming the types. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies as described -- (Lelek 1987:256 [ref. 22962] as Noemacheilus). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Barbatula quignardi (Băcescu-Meşter1967) -- (Gauliard et al. 2019:176 [ref. 36787]). •Valid as Barbatula hispanica (Lelek 1987) -- (Denys et al. 2021:5 [ref. 38347]). Current status: Valid as Barbatula hispanica (Lelek 1987). Nemacheilidae. Distribution: Southwestern Europe: Ebro drainage and Cantabria (Spain); Adour drainage (southwestern France). Habitat: freshwater.

karabanowi, Barbatula Prokofiev [A. M.] 2018:269, Figs. 3-4 [Zootaxa 4407 (no. 2); ref. 35878] Bayan-Ölgii aimag, headwaters of the Bulgan River, Mongolia, 47°04'46.4"N, 91°02'36.1"E. Holotype: ZMMU 23829. Paratypes: ZMMU. Plus additional non-type material. •Valid as Barbatula karabanowi Prokofiev 2018:267 [ref. 35878] Current status: Valid as Barbatula karabanowi Prokofiev 2018. Nemacheilidae. Distribution: Upper Bulgan River drainage, Mongolia. Habitat: freshwater.

kirinensis, Barbatula toni Tchang [T.-L.] 1932:115, Fig. 2 [Bulletin of the Fan Memorial Institute of Biology, Peiping (Zoological Series) v. 3 (no. 8); ref. 16401] Kirin, Jilin Province, northeastern China. Holotype: ZMFMIB (now ASIZB) 7931. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:870 [ref. 20661] as Nemachilus). •Synonym of Barbatula barbatula (Linnaeus 1758), subspecies nuda (Bleeker 1868) -- (Zhu 1989:30 [ref. 17744]). •Synonym of Barbatula toni (Dybowski 1869), but a valid subspecies as described -- (Vasil'eva 1998:96 [ref. 23591]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula nuda (Bleeker 1964) -- (Kottelat 2012:79 [ref. 32367] with question). •Synonym of Barbatula toni (Dybowski 1869) -- (Kottelat 2006:53 [ref. 28831], Cao et al. 2012:241 [ref. 32335], Prokofiev 2015:461 [ref. 34065]). •Valid as Barbatula kirinensis Tchang 1932 -- (Chen et al. 2019:12 [ref. 36569], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula kirinensis Tchang 1932. Nemacheilidae. Distribution: East Asia: Jingpo Lake, Tumen River and Zhu'Erduo River (northern China). Habitat: freshwater.

leoparda, Barbatula Gauliard [C.], Dettai [A.], Persat [H.], Keith [P.] & Denys [G. P. J.] 2019:171, Figs. 1A-B, 2A, 3 [Cybium v. 43 (no. 2); ref. 36787] Têt River at Nefiach, Departement Pyrénées-Orientales, France, 42.697N, 2.663E. Holotype: MNHN 2018-0228. Paratypes: MNHN. •Valid as Barbatula leoparda Gauliard, Dettai, Persat, Keith & Denys 2019. Current status: Valid as Barbatula leoparda Gauliard, Dettai, Persat, Keith & Denys 2019. Nemacheilidae. Distribution: Têt and Tech Rivers, Pyrénées-Orientales, southern France. Habitat: freshwater.

liaoyangensis, Barbatula Chen [H.], Zhang [H.], Chen [Y.-X.] & Freyhof [J.] 2019:22, Figs. 23-25 [Zootaxa 4565 (no. 1); ref. 36569] Taizi River east of Xiaobeihezhen, Liao River drainage, Liaoning Province, China, 41°36′N, 122°85′E. Holotype: HU 1608150. Paratypes: FSJF, HU. Plus additional non-type material. •Valid as Barbatula liaoyangensis Chen, Zhang, Chen & Freyhof 2019. Current status: Valid as Barbatula liaoyangensis Chen, Zhang, Chen & Freyhof 2019. Nemacheilidae. Distribution: Taizi River, Liao River drainage, northern China. Habitat: freshwater.

linjiangensis, Barbatula Chen [H.], Zhang [H.], Chen [Y.-X.] & Freyhof [J.] 2019:25, Figs. 26-28 [Zootaxa 4565 (no. 1); ref. 36569] Yalu River south-west of Linjiang City, Jilin Province, China, 41°79′N, 126°89′E Holotype: HU 1608016. Paratypes: HU. Plus additional non-type material. •Valid as Barbatula linjiangensis Chen, Zhang, Chen & Freyhof 2019. Current status: Valid as Barbatula linjiangensis Chen, Zhang, Chen & Freyhof 2019. Nemacheilidae. Distribution: Yalu River drainage, northern China (and possibly North Korea). Habitat: freshwater.

markakulensis, Nemacheilus barbatulus Menshikov [M. I.] 1939:141, Fig. [Ucenye zapiski / Permskij Ordena Trudovogo Krasnogo Znameni Gosudarstvennyj Universitet Imeni A. M. Gor'kogo, Ministerstvo Vyssego i Srednego Special'nogo Obrazovanija RSFSR. v. 3 (no. 2); ref. 21170] Lake Marka-kul', Irtysh basin, Kazakhstan. Syntypes: ZIN 26864 (6). Author also seen as Men'shikov. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:872 [ref. 20661] as Nemachilus). •Synonym of Noemacheilus barbatulus (Linnaeus 1758), but a valid subspecies markakulensis Manshikov 1939 -- (Mitrofanov & Dukravec 1989:60 [ref. 37748]). •Synonym of Barbatula toni (Dybowski 1869) -- (Kottelat 2006:53 [ref. 28831]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (authors). •Valid as Barbatula markakulensis (Menshikov 1939) -- (Kottelat 2012:78 [ref. 32367] with question, Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:396 [ref. 34693] as makakulensis, Prokofiev 2018:273 [ref. 35878]). Current status: Valid as Barbatula markakulensis (Menshikov 1939). Nemacheilidae. Distribution: North Asia: Markakul Lake (Kazakhstan). Habitat: freshwater.

nudus, Nemacheilus Bleeker [P.] 1864:12 [Nederlandsch Tijdschrift voor de Dierkunde v. 2; ref. 17027] Mongolia [brought from China]. Apparently Hebei, Chongli county, Siwantse [Xiwanzi], 40°58'25"N, 115°16'22"E, China. [Possibly Daling or Liao Rivers, Liaoning, China] Holotype (unique): MNHN 0000-1450. Type catalog: Bertin & Estève 1948:98 [ref. 19574]. Locality information from Kottelat 2012:78 [ref. 32367]; also a lengthy discussion of status. •Mentioned -- (Ding 1994:78 [ref. 21531]). •Synonym of Barbatula barbatula (Linnaeus 1758), but a valid subspecies nuda (Bleeker 1864) -- (Zhu 1989:26 [ref. 17744], Zhu 1995:108 [ref. 25213]). •Valid as Orthrias nudus (Bleeker 1864) -- (Kim 1997:283 [ref. 24032], Kim & Park 2002:195 [ref. 25913]). •Questionably a senior synonym of Barbatula toni (Dybowski 1869) -- (Kottelat 2006:53 [ref. 28831], Bogutskaya et al. 2008:339 [ref. 30085]). •In Barbatula -- (based on status of Barbatula). •Valid as Barbatula nuda (Bleeker 1864) -- (Wang et al. 2001:168 [ref. 26566], Cao et al. 2012:237 [32335], Kottelat 2012:78 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Zhang et al. 2016:127 [ref. 34477], Prokofiev 2016:396 [ref. 34693], Prokofiev 2018:273 [ref. 35878], Chen et al. 2019:6 [ref. 36569] dated 1865, An et al. 2021:121 [ref. 38850], Du et al. 2021:4 [ref. 38600], Dyldin et al. 2023:[8] [ref. 40328] with question). Current status: Valid as Barbatula nuda (Bleeker 1864). Nemacheilidae. Distribution: East Asia: Liao and Taizi River drainages (northern China) and northern Korea. Habitat: freshwater.

oreas, Orthrias Jordan [D. S.] & Fowler [H. W.] 1903:769, Fig. 2 [Proceedings of the United States National Museum v. 26 (no. 1332); ref. 2461] Chitose in Iburi, Hokkaido, Japan. Holotype (unique): Sapporo Mus. Non-types: BMNH 1905.2.4.134 [from Jordan] (1). •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Barbatula barbatula (Linnaeus 1758), subspecies nuda (Bleeker 1868) -- (Zhu 1989:29 [ref. 17744]). •Synonym of Barbatula toni (Dybowski 1869) -- (Okada 1961:541 [ref. 6363], Vasil'eva 1998:95 [ref. 23591], Sideleva 2001:54 [ref. 26181], Sideleva 2003:11 [ref. 27815], Kottelat 2006:53 [ref. 28831]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies oreas (Jordan & Fowler 1903) -- (Prokofiev 2007:15 [ref. 30347]). •Valid as Barbatula oreas (Jordan & Fowler 1903) -- (Kottelat 2012:79 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula oreas (Jordan & Fowler 1903). Nemacheilidae. Distribution: East Asia: Hokkaido (Japan) and South Korea. Habitat: freshwater.

parisiensis, Cobitis barbatula var. De la Pylaie [A. J. M.] 1835:534 [Congrès Scientifique de France v. 2 (Sept. 1834) (art. 5); ref. 1086] Name only, not available. From Paris, France. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Kottelat 2012:77 [ref. 32367]). Nomen Nudum. Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae.

pechiliensis, Nemachilus Fowler [H. W.] 1899:181 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 51; ref. 17572] Tan lan Ho River, tributary of Shan lan Ho River, about 30 miles northeast of Lama-miau or Dolon-nor, Pechili Province, Inner Mongolia. Holotype (unique): ANSP 16394 (missing). Type catalog: Böhlke 1984:63 [ref. 13621]. Original genus is correctly Nemacheilus. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Barbatula toni (Dybowski 1869) -- (Okada 1961:541 [ref. 6363] as pichiliensis, Kottelat 2006:53 [ref. 28831]). •Synonym of Barbatula barbatula (Linnaeus 1758), subspecies nuda (Bleeker 1868) -- (Zhu 1989:29 [ref. 17744]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula toni fowleri Fowler 1925 -- (Cao et al. 2012:241 [ref. 35335]). •Synonym of Barbatula nuda (Bleeker 1964) -- (Kottelat 2012:78 [ref. 32367] with question). •Possible senior synonym of Barbatula potaninorum (Prokofiev 2007) -- (Prokofiev 2015:461 [ref. 34065]). •Valid as Barbatula pechiliensis (Fowler 1899) -- (Chen et al. 2019:15 [ref. 36569]). Current status: Valid as Barbatula pechiliensis (Fowler 1899). Nemacheilidae. Distribution: Luan, Hai, Yin and Nu'er River drainages and Dali-Nur Lake, northern China. Habitat: freshwater.

pictava, Cobitis barbatula var. De la Pylaie [A. J. M.] 1835:534 [Congrès Scientifique de France v. 2 (Sept. 1834) (art. 5); ref. 1086] Name only, not available; from France. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Kottelat 2012:77 [ref. 32367]). Nomen Nudum. Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae.

pironae, Cobitis Nardo [G. D.] 1866:143 [Atti del regio Istituto Veneto di Scienze, Littere ed Arti (Ser. 3) v. 11; ref. 23026] [Basso Friuli] Lower Friuli-Venezia Giulia, Italy. No types known. Mention as variety of Cobitis barbatula, but name appeared as Cobitis Pironae. No description given but mention that description was to be published by Ninni. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Kottelat 2012:77 [ref. 32367]). Nomen Nudum. Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae.

potaninorum, Orthrias Prokofiev [A. M.] 2007:65, Fig. 18 [Morphology, systematics and origin of the stone loach genus Orthrias (Teleostei: Balitoridae: Nemacheilinae); ref. 30343] Fansu, Edzin-Gol, 42°25'20"N, 100°40'07"E, North China. Holotype (unique): ZIN 8121. Locality from Kottelat 2012:79 [ref. 32367]. •Valid as Barbatula potaninorum (Prokofiev 2007) -- (Cao et al. 2012:236 [ref. 32335], Kottelat 2012:79 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:396 [ref. 34693], Prokofiev 2018:274 [ref. 35878], Chen et al. 2019:12 [ref. 36569]). Current status: Valid as Barbatula potaninorum (Prokofiev 2007). Nemacheilidae. Distribution: Inner Mongolia, northern China. Habitat: freshwater.

quignardi, Noemacheilus barbatulus Băcescu-Meşter [L.] 1967:359, Figs. 1 (A, B, C), 5 (C, D) [Travaux du Muséum d'Histoire Naturelle "Grigore Antipa" v. 7; ref. 7940] Le Lez River, near Montpellier, France. Holotype: MGAB 77. Paratypes: MGAB 78 (1), MGAB (16). Genus is correctly Nemacheilus. •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Barbatula barbatula (Linnaeus 1758), but a valid subspecies as described -- (Vasil'eva 1998:95 [ref. 23591]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies quignardi (Băcescu-Meşter 1967) -- (Prokofiev 2007:15 [ref. 30343]). •Valid as Barbatula quignardi (Băcescu-Meşter 1967) -- (Kottelat & Freyhof 2007:330 [ref. 29996], Kottelat 2012:79 [ref. 32367], Gauliard et al. 2019:176 [ref. 36787]). Current status: Valid as Barbatula quignardi (Băcescu-Meşter 1967). Nemacheilidae. Distribution: Southern France and northwestern Spain. Habitat: freshwater.

restricta, Barbatula Prokofiev [A. M.] 2015:458, Figs. 1-4 [Zootaxa 4052 (no. 4); ref. 34065] Saldan-Kol (= Dinnoye) Lake, Builyukem River catchment, Chuya River system, upper Ob drainage, Altai, Russia, 50°13'00"N, 89°15'30"E, elevation 2272 meters. Holotype: ZIN 52587. Paratypes: ZIN. Specific name is an adjective. •Valid as Barbatula restricta Prokofiev 2015 -- (Prokofiev 2016:399 [ref. 34693], Dyldin et al. 2020:110, 111 [ref. 37569], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula restricta Prokofiev 2015. Nemacheilidae. Distribution: North Asia: Chuya River system, Upper Ob, Altai (Russia). Habitat: freshwater.

sawadai, Orthrias Prokofiev [A. M.] 2007:84, Figs. 26-29 [Morphology, systematics and origin of the stone loach genus Orthrias (Teleostei: Balitoridae: Nemacheilinae); ref. 30343] Uram River, Ero River basin, Selenga River system, into lake Baikal, Yenisei River basin, Mongolia / Russia. Holotype: ZIN 52054. Paratypes (55): ZIN 52055 (3), 52056 (12 + 3 c&s), 52254 (9 + 2 C&S), 53145 (1), 53150 (21 + 4 c&s). •Valid as Barbatula sawadai (Prokofiev 2007) -- (Cao et al. 2012:236 [ref. 32335], Kottelat 2012:80 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:399 [ref. 34693], Prokofiev 2018:274 [ref. 35878], Dyldin et al. 2020:109 [ref. 37569], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula sawadai (Prokofiev 2007). Nemacheilidae. Distribution: North Asia: Selenga River basin (Mongolia and Russia). Habitat: freshwater.

shansi, Barbatula stoliczkai Nichols [J. T.] 1925:6 [American Museum Novitates No. 171; ref. 3178] Mai-tai-chao, Shansi (42 miles east of Paotow [Pao-t'ou] Nei Mongolia), northern China. Holotype: AMNH 84112. •Synonym of Barbatula nuda (Bleeker 1864) -- (Kottelat 2012:78 [ref. 32367], Chen et al. 2019:6 [ref. 36569]). •Uncertain -- (Prokofiev 2015:461 [ref. 34065]). Current status: Synonym of Barbatula nuda (Bleeker 1864). Nemacheilidae. Habitat: freshwater.

sibiricus, Nemacheilus Gratzianov [V. I.] 1907:164, 168 [Trudy Otdela Ichtiologii Imperatorskago Russkago Obshchestva Akklimatizacii Zivotnych' i Rastenii. v. 4; ref. 1871] Biya River near Biysk [Biysk], Altai, Russia. Lectotype: ZUMM 2531 (140 mm). Paralectotypes: ZMMU P-2532 (now 2). Type catalog: Vasil'eva et al. 2001:25 [ref. 25714]. Lectotype selected by Prokofiev 2016:394 [ref. 34693]. Former syntypes ZMMGU 2532 (3) = Barbatula compressirostris, see Prokofiev 2018:273 [ref. 35878]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:869 [ref. 20661] as Nemachilus). •Synonym of Nemacheilus toni (Dybowski 1869) -- (Bogutskaya & Naseka 1996:47 [ref. 22798]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (based on status of Barbatula). •Questionably the same as Nemacheilus tominus Ruzssky 1920 -- (Kottelat 2012:80 [ref. 32367]). •Synonym of Barbatula toni (Dybowski 1869) -- (Vasil'eva 1998:95 [ref. 23591], Sideleva 2001:54 [ref. 26181], Sideleva 2003:11 [ref. 27815], Kottelat 2006:53 [ref. 28831], Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:391 [ref. 34693] with question). Current status: Synonym of Barbatula toni (Dybowski 1869). Nemacheilidae. Habitat: freshwater.

sturanyi, Nemachilus Steindachner [F.] 1892:378 [22] [131], Pl. 2 (fig. 3) [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 59 (1. Abth.); ref. 14517] Lake Ohrid [Ochrida] at Pestani, between Ohrid City and Naum Monastery, Yugoslavia/Albania. Holotype (unique): NMW 48440. Original genus is correctly Nemacheilus. On p. 22 of separate; on p. 131 of abstract. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies -- (Lelek 1987:256 [ref. 22962] as Noemacheilus). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Barbatula barbatula (Linnaeus 1758), but a valid subspecies -- (Vasil'eva 1998:95 [ref. 23591], Economidis 2005:67 [ref. 28213]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies sturanyi (Steindachner 1892) -- (Soric 2000:64 [ref. 28770], Prokofiev 2007:15 [ref. 30343]). •Valid as Barbatula sturanyi (Steindachner 1892) -- (Kottelat 2006:89 [ref. 28831], Kottelat & Freyhof 2007:331 [ref. 29996], Kottelat 2012:80 [ref. 32367], Shumka et al. 2023:36 [ref. 40747]). Current status: Valid as Barbatula sturanyi (Steindachner 1892). Nemacheilidae. Distribution: Southeastern Europe: Lake Ohrid basin and upper Drin River (Macedonia and Albania). Habitat: freshwater.

taurica, Nemachilus barbatulus var. Kessler [K. F.] 1877:171 [The Aralo-Caspian Expedition; ref. 2597] Selgir River, Crimea, Ukraine. Syntypes: ZIN 3029 (3), 3032 (3). Original genus is correctly Nemacheilus. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:868 [ref. 20661] as Nemachilus). •Synonym of Barbatula barbatula (Linnaeus 1758). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies barbatulus (Linnaeus 1758) -- (Prokofiev 2007:54 [ref. 30343]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (based on status of Barbatula) Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae. Habitat: freshwater, brackish.

tigris, Barbatula barbatula morpha Gundriser [A. N.] 1975 [Fishes of Tucva, ASSR; ref. 28833] Original not seen; from Kottelat 2006:51 [ref. 28831]. Infrasubspecific, not available. •Questionably in the synonymy of Barbatula compressirostris (Warpachowski 1897) -- (Kottelat 2006:51 [ref. 28831]). •Not available, infrasubspecific name -- (Prokofiev 2015:461 [ref. 34065]). Infrasubspecific. Current status: Synonym of Barbatula compressirostris (Warpachowski 1897). Nemacheilidae.

tomianus, Nemacheilus barbatulus Ruzsky [M. D.] 1920:36, Figs. 1-2 [Trudy estestvenno-isthrichsk. Otdel., Tomsk No. 1; ref. 21169] Basins of the rivers Tom, Ob and Yenisei, Russia. Syntypes: not researched. See Reshetnikov et al. 1997:743 [ref. 24702]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:871 [ref. 20661] as Nemachilus). •Synonym of Barbatula toni (Dybowski 1869) -- (Vasil'eva 1998:96 [ref. 23591], Sideleva 2001:54 [ref. 26181], Sideleva 2003:11 [ref. 27815], Kottelat 2006:53 [ref. 28831]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Valid as Barbatula tomiana (Ruzsky 1920) -- (Kottelat 2012:80 [ref. 32367], Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:399 [ref. 34693], Romanov et al. 2017:823 [ref. 35687], Prokofiev 2018:274 [ref. 35878], Dyldin et al. 2023:[8] [ref. 40328]). Current status: Valid as Barbatula tomiana (Ruzsky 1920). Nemacheilidae. Distribution: East Asia: Altai Republic (Russia) and Mongolia. Habitat: freshwater.

toni, Cobitis Dybowski [B. N.] 1869:957, Pl. 18 (fig. 10) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 19; ref. 1169] Onon and Ingoda rivers, Amur River basin, Russia. Syntypes and/or Dybowski specimens: MNHN B-0738 (8), B-0739 (4); NMW 50473 (2). Type catalog: Bertin & Estève 1948:98 [ref. 19574]. For remarks on systematics see Kottelat 2006:53 [ref. 28831]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies -- (Berg 1949:869 [ref. 20661] as Nemachilus, Kawanabe & Mizuno 1989:398 [ref. 25598] as Neomacheilus, Watanabe 1998:263 [ref. 23491]). •Synonym of Noemacheilus barbatulus (Linnaeus 1758), but a valid subspecies toni -- (Chereshnev 1996:601 [ref. 24659], Mitrofanov & Dukravec 1989:58 [ref. 37748], Nakabo 2000:276 [ref. 25086], Nakabo 2002:276 [ref. 26001]). •Synonym of Barbatula barbatula, subspecies nuda (Bleeker 1865) -- (Zhu 1989:29 [ref. 17744]). •Valid as Nemacheilus toni (Dybowski 1869) -- (Wang 1984:82 [ref. 22951], Sawada in Masuda et al. 1984:59 [ref. 6441] as Noemacheilus, Bogutskaya & Naseka 1996:47 [ref. 22798], Matsuura et al. 2000:176 [ref. 26000], Choi et al. 2003:178, 517 [ref. 26218]). •Valid as Orthrias toni (Dybowski 1869) -- (Bănărescu & Nalbant 1995:449 [ref. 23187], Kim et al. 2000:224 [ref. 25434], Kim & Park 2002:193 [ref. 25913], Jang et al. 2003:119 [ref. 27681]). •Synonym of Orthrias nudus (Bleeker 1865) -- (Kim 1997:555 [ref. 24032]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies toni (Dybowski 1869) -- (Prokofiev 2003:736 [ref. 27494], Prokofiev 2007:17 [ref. 30343]). •Valid as Barbatula toni (Dybowski 1869) -- (Okada 1961:541 [ref. 6363], Nikiforov et al. 1997:330 [ref. 24694], Reshetnikov et al. 1997:743 [ref. 24702], Vasil'eva 1998:95 [ref. 23591], Sheiko & Fedorov 2000:17 [ref. 25650], Chereshnev et al. 2001:37 [ref. 26736], Tokranov 2001:268 [ref. 25407], Zemnukhov et al. 2001:420 [ref. 25412], Kottelat 2001:49 [ref. 25482], Pietsch et al. 2001:140 [ref. 25688], Bogutskaya et al. 2001:45 [ref. 26178], Sideleva 2001:54 [ref. 26181], Fedorov et al. 2003:25 [ref. 27579], Alekseyev et al. 2004:54 [ref. 27625], Sideleva 2003:11 [ref. 27815], Bogutskaya & Naseka 2004:113 [ref. 28183], Kottelat 2006:53 [ref. 28831], Ocock et al. 2006:28 [ref. 28933], Kottelat & Freyhof 2007:330 [ref. 29996], Bogutskaya et al. 2008:339 [ref. 30085], Cao et al. 2012:241 [ref. 32335], Kottelat 2012:80 [ref. 32367], Parin et al. 2014:103 [ref. 33547], Prokofiev 2015:460 [ref. 34065], Prokofiev 2016:399 [ref. 34693], Dyldin & Orlov 2016:669 [ref. 34853] with question, Prokofiev 2018:274 [ref. 35878], Chen et al. 2019:8 [ref. 36569], Dyldin et al. 2020:87 [ref. 37569], An et al. 2021:121 [ref. 38850], Dyldin et al. 2021:19 [ref. 38804] with question, Dyldin & Orlov 2021:63 [ref. 38158] with question, Dyldin et al. 2023:[8] [ref. 40328] with question). Current status: Valid as Barbatula toni (Dybowski 1869). Nemacheilidae. Distribution: East Asia: Amur (Heilongjiang) and Yin River drainages (Russia, Mongolia, northern China and northern Korea). Habitat: freshwater.

vardarensis, Nemachilus barbatulus Karaman [S.] 1928:164 [Glasnik Skopskog Nauchnog Drustva [Glasnik -- Bulletin de la Société Scientifique de Skopje]. v. 6; ref. 15903] Vardar River basin, former Yugoslavian republic of Macedonia. Original genus is correctly Nemacheilus. Also in Karaman 1929:173 [ref. 15903]. •Synonym of Nemacheilus barbatulus (Linnaeus 1758), but a valid subspecies as described -- (Lelek 1987:256 [ref. 22962] as Noemacheilus). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952]). •Synonym of Orthrias barbatulus (Linnaeus 1758), but a valid subspecies vardarensis (Karaman 1928) -- (Soric 2000:62 [ref. 28770]). •Synonym of Barbatula barbatula (Linnaeus 1758), but a valid subspecies vardarensis (Karaman 1928) -- (Vasil'eva 1998:95 [ref. 23591], Economidis 2005:67 [ref. 28213]). •Synonym of Barbatula barbatula (linnaeus 1758) -- (Kottelat 2012:78 [ref. 32367]). •Valid as Barbatula vardarensis (Karaman 1928) -- (Geiger et al. 2014:Table S2 [ref. 37408]). Current status: Valid as Barbatula vardarensis (Karaman 1928). Nemacheilidae. Distribution: Vardar / Axios River basin, Macedonia and Greece. Habitat: freshwater.

variabilis, Cobitis Günther [A.] 1868:355 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Not available, Heckel name on specimen from Moravia, Czech Republic, mentioned under Nemachilus barbatulus. •In the synonymy of Barbatula barbatula (Linnaeus 1758) -- (Kottelat 1997:93 [ref. 22952], Kottelat 2012:77 [ref. 32367]). Nomen Nudum. Current status: Synonym of Barbatula barbatula (Linnaeus 1758). Nemacheilidae.

zetensis, Orthrias barbatulus Sorić [V. M.] 2000:66, Fig. [Ichthyologia v. 32 (no. 1); ref. 28770] Near village of Vis, 5-6 kilometers west of town of Danilovgrad from the Vis brook, a tributary of the Zeta River, Montenegro. Holotype: BIK 59.4 mm SL. Paratypes: BIK (4). •Synonym of Barbatula barbatula (Linnaeus 1758) but a valid subspecies zetensis (Sorić 2000) -- (Bohlen et al. 2003 [ref. 28830]). •Valid as Barbatula zetensis (Sorić 2000) -- (Kottelat & Freyhof 2007:332 [ref. 29996], Kottelat 2012:80 [ref. 32367], Shumka et al. 2023:36 [ref. 40747]). Current status: Valid as Barbatula zetensis (Sorić 2000). Nemacheilidae. Distribution: Southeastern Europe: drainage area of Lake Skadar (Montenegro and Albania). Habitat: freshwater.

zhangwuensis, Barbatula Chen [H.], Zhang [H.], Chen [Y.-X.] & Freyhof [J.] 2019:27, Figs. 29-31 [Zootaxa 4565 (no. 1); ref. 36569] Liu River at Ertaizicun of Zhangwu, Liao River drainage, Liaoning Province, China, 42°40′N, 122°43′E. Holotype: HU 1609235. Paratypes: FSJF, HU. Plus additional non-type material. •Valid as Barbatula zhangwuensis Chen, Zhang, Chen & Freyhof 2019. Current status: Valid as Barbatula zhangwuensis Chen, Zhang, Chen & Freyhof 2019. Nemacheilidae. Distribution: Muxiga, Liu, Pushi and Erdaobai River drainages, northern China (and possibly North Korea). Habitat: freshwater.