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Eschmeyer's Catalog of Fishes
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Species in the genus Dipturus: [ 64 ] records


acrobelus, Dipturus Last [P. R.], White [W. T.] & Pogonoski [J. J.] 2008:10, Figs. 1-6 [CSIRO Marine and Atmospheric Research Paper No. 021; ref. 29647] East of Patricks Head, Tasmania, 42°33'S, 148°42'E, depth 905-990 meters. Holotype: CSIRO H 226-01. Paratypes: 24 at AMS, CSIRO, NMV. Plus non-type material. •Valid as Dipturus acrobelus Last, White & Pogonoski 2008 -- (Last & White 2008:109 [ref. 30618], Last & Alava 2013:214 [ref. 33041], Weigmann 2016:107 [ref. 34211], Last et al. 2016:20 [ref. 34941], Last et al. 2016:241 [ref. 35000]). Current status: Valid as Dipturus acrobelus Last, White & Pogonoski 2008. Rajidae. Distribution: Southern Australia: New South Wales to Western Australia, including Tasmania. Habitat: marine.

acus, Raja Lacepède [B. G. E.] 1803:662, 665 [Histoire naturelle des poissons (Lacepède) v. 5; ref. 4930] Coast of France or England, northeastern Atlantic. No types at MNHN. Type catalog: Séret & McEachran 1987:30 [ref. 9312]. •Synonym of Raja oxyrinchus Linnaeus 1758 -- (Stehmann 1973:63 [ref. 7168]). •Synonym of Dipturus oxyrinchus (Linnaeus 1758) -- (Dyldin 2015:70 [ref. 34524]). Current status: Synonym of Dipturus oxyrinchus (Linnaeus 1758). Rajidae. Habitat: marine.

amphispinus, Dipturus Last [P. R.] & Alava [M.] 2013:215, Figs. 1-10 [Zootaxa 3752 (no. 1); ref. 33041] Dipolog City public market, Dipolog City, Zanboanga del Norte, Philippines. Holotype: PNM 15178. Paratypes: CSIRO H 7416 (01); SUML MMLM 014, 018 (1), 018 (1) SUMM MMLM 012 (1), SUMM BRU 147 (1), Plus one non-type specimen. •Valid as Dipturus amphispinus Last & Alava 2013 -- (Weigmann 2016:107 [ref. 34211], Last et al. 2016:20 [ref. 34941], Last et al. 2016:242 [ref. 35000]). Current status: Valid as Dipturus amphispinus Last & Alava 2013. Rajidae. Distribution: Philippines Archipelago. Habitat: marine.

apricus, Dipturus Last [P. R.], White [W. T.] & Pogonoski [J. J.] 2008:19, Figs. 8-13 [CSIRO Marine and Atmospheric Research Paper No. 021; ref. 29647] South of Saumarez Reef, Queensland, Australia, 22°35'S, 153°40E, depth 314-319 meters. Holotype: CSIRO H 601-01. Paratypes: 23 at AMS, CSIRO, NMV, QM. Plus non-type specimens. •Valid as Dipturus apricus Last, White & Pogonoski 2008 -- (Last & Alava 2013:214 [ref. 33041], Weigmann 2016:107 [ref. 34211], Last et al. 2016:20 [ref. 34941], Last et al. 2016:243 [ref. 35000]). Current status: Valid as Dipturus apricus Last, White & Pogonoski 2008. Rajidae. Distribution: Off Queensland, Australia. Habitat: marine.

argentinensis, Dipturus Díaz de Astarloa [J. M.], Mabragaña [E.], Hanner [R.] & Figueroa [D. E.] 2008:37, Figs. 1-2 [Zootaxa No. 1921; ref. 29969] Off central Patagonia, continental shelf, 45°38'S, 64°08'W, depth 98 meters. Holotype: INIDEP 793. Paratypes: INIDEP 794-800, 802-803 (total 7). •Valid as Zearaja argentinensis (Díaz de Astarloa, Mabragaña, Hanner & Figueroa 2008) -- (Weigmann 2016:123 [ref. 34211] see remarks, Last et al. 2016:23 [ref. 34941], Last et al. 2016:360 [ref. 35000]). •Valid as Dipturus argentinensis Díaz de Astarloa, Mabragaña, Hanner & Figueroa 2008 -- (Mabragaña et al. 2011:7 and table S2 [ref. 31800], Vargas-Caro 2015:323 [ref. 33982], Concha et al. 2019:507 [ref. 36686]). Current status: Valid as Dipturus argentinensis Díaz de Astarloa, Mabragaña, Hanner & Figueroa 2008. Rajidae. Distribution: Southwestern Atlantic: Argentina. Habitat: marine.

batis, Raja Linnaeus [C.] 1758:231 [Systema Naturae, Ed. X v. 1; ref. 2787] European seas [original: "in Oceano Europæo"]; localities include England, northeastern Atlantic; Lesbos Island, Greece, Mediterranean Sea. No types known. Lectotype or neotype fixation needed. Originally based on multiple species and at least 22 pre-Linnaean sources including Artedi 1738:73 [ref. 30578] and Artedi 1738:102 [ref. 30349] as Raja varia, dorso medio glabro; Rondelet 1554:346 [ref. 30354] and Gesner 1558:932 [ref. 30350] as Raia vndulata siue cinerea, Jonston 1649:Pl. 9 (fig. 1) [ref. 30579] as Raia, Charleton 1668:130 [ref. 30580] as Raia undulata. Spelled batys in Risso 1810:3 [ref. 3755]. See Iglésias et al. 2010 [ref. 30586]. •Valid as Raja batis Linnaeus 1758 -- (Stehmann 1973:62 [ref. 7168], Stehmann & Bürkel in Whitehead et al. 1984:175 [ref. 13675], Séret 1989:126 [ref. 12827], Stehmann 1990:31 [ref. 19316], Stehmann 1995:105 [ref. 22055], Santos et al. 1997:119 [ref. 23531], Arruda 1997:22 [ref. 24952]). •See Iglésias et al. 2010:333 [ref. 30586] for possible splitting of the species. •Valid as Dipturus batis (Linnaeus 1758) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Bilecenoğlu et al. 2002:21 [ref. 26753], Fricke 2007:25 [ref. 30577], Fricke et al. 2007:18 [ref. 29533], Fricke 2008:11 [ref. 30182], Jeong & Nakabo 2008:189 [ref. 29612], George 2009:56 [ref. 30539], Lipej & Dulčić 2010:13 [ref. 36649], Naylor et al. 2012:94 [ref. 32267] see remarks, Parin et al. 2014:35 [ref. 33547], Dyldin 2015:69 [ref. 34524], Weigmann 2016:107 [ref. 34211] see remarks, Bañón et al. 2016:301 [ref. 34397], McEachran & Séret 2016:1396 [ref. 34551], Gordon et al. 2016:260 [ref. 34661], Last et al. 2016:20 [ref. 34941] used herein as D. cf. flossada, Last et al. 2016:244 [ref. 35000], Biscoito et al. 2018:486 [ref. 37562], Almeida & Biscoito 2019:139 [ref. 37302], Carneiro et al. 2019:43 [ref. 37250], Kovačić et al. 2020:7 [ref. 37519]). Current status: Valid as Dipturus batis (Linnaeus 1758). Rajidae. Distribution: Baltic Sea; North Sea; Mediterranean Sea; eastern Atlantic: Iceland and Norway south to Senegal, including Canary Islands. Habitat: marine.

brevicaudata, Raia Marini [T. L.] 1933:329 [4] [Physis (Buenos Aires) v. 11 (no. 39); ref. 16500] Argentina. Holotype (unique): MACN ICT-569. Type catalog: Gabbanelli et al. 2018:455 [ref. 36350]. On p. 4 of separate. •Synonym of Raja chilensis Guichenot 1848 -- (Lloris & Rucabado 1991:44 [ref. 22530]). •Synonym of Zearaja chilensis (Guichenot 1848) -- (Vargas-Caro et al. 2015:325 [ref. 33982]). •Valid as Zearaja brevicaudata (Marini 1933) -- (Gabbanelli et al. 2018:464 [ref. 36350]). •Valid as Dipturus brevicaudatus (Marini 1933) -- (Concha et al. 2019:507 [ref. 36686]). Current status: Valid as Dipturus brevicaudatus (Marini 1933). Rajidae. Distribution: Southwestern Atlantic: southern Brazil south to Beagle Channel, Argentina and Chile, Habitat: marine.

bullisi, Raja Bigelow [H. B.] & Schroeder [W. C.] 1962:168, Fig. 1 [Bulletin of the Museum of Comparative Zoology v. 128 (no. 4); ref. 5793] About 30 miles southwest of Dry Tortugas, Florida, U.S.A., 24°18'N, 83°18'W, Silver Bay station 1199, depth 200 fathoms. Holotype: USNM 196442. Paratypes: MCZ 40673 (1), plus 16 other specimens. •Valid as Raja bullisi Bigelow & Schroeder 1962 -- (Uyeno & Miyake in Uyeno et al. 1983:74 [ref. 14275], Séret 1989:126 [ref. 12827], Cervigón 1992:205 [ref. 23827], Castro-Aguirre & Espinosa Pérez 1996:36 [ref. 22793], Cervigón & Alcalá 1999:167 [ref. 24490]). •Valid as Dipturus bullisi (Bigelow & Schroeder 1961) -- (McEachran & Fechhelm 1998:154 [ref. 23897], Compagno 1999:490 [ref. 25589], Gomes & Paragó 2001:1 [ref. 25669], McEachran & Carvalho 2003:551 [ref. 26985], Nelson et al. 2004:55 [ref. 27807], Page et al. 2013:55 [ref. 32708], Vargas-Caro 2015:323 [ref. 33982], Del Moral-Flores et al. 2016:113 [ref. 34398], Weigmann 2016:107 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:245 [ref. 35000], Ehemann et al. 2018:25 [ref. 36194], Ehemann et al. 2019:[6] [ref. 36861], Jones et al. 2020:286 [ref. 37610]). Current status: Valid as Dipturus bullisi (Bigelow & Schroeder 1962). Rajidae. Distribution: Western Atlantic: including Gulf of Mexico and Caribbean Sea. Habitat: marine.

campbelli, Raja Wallace [J. H.] 1967:24, Fig. 12 [Investigational Report. Oceanographic Research Institute, Durban No. 17; ref. 7929] 24 miles east of Durban, KwaZulu-Natal, South Africa, southwestern Indian Ocean, depth 175 fathoms. Holotype: SAIAB [formerly RUSI] 992 [ex ORI B804]. Paratypes: ?SAIAB [formerly RUSI] [ex ORI B859] (1). Additional material: ORI uncat. (5). •Synonym of Raja pullopuntata Smith 1964 -- (Hulley 1986:123 [ref. 5672]). •Valid as Raja campbelli Wallace 1967 -- (Séret 1989:127 [ref. 12827], Heemstra in Smith & Heemstra 1995:vii [ref. 21953]). •Valid as Dipturus campbelli (Wallace 1967) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Heemstra & Heemstra 2004:81 [ref. 28072], Compagno & Ebert 2007:137 [ref. 30471], Last & Alava 2013:214 [ref. 33041], Weigmann 2016:107 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:246 [ref. 35000]). Current status: Valid as Dipturus campbelli (Wallace 1967). Rajidae. Distribution: Southwestern Indian Ocean: KwaZulu-Natal and southern Mozambique. Habitat: marine.

canutus, Dipturus Last [P. R.] 2008:54, Figs. 1-6 [CSIRO Marine and Atmospheric Research Paper No. 021; ref. 29648] East of Maria Island, Tasmania, Australia, 42°40'S, 148°25'E, depth 448-460 meters. Holotype: CSIRO H 153-01. Paratypes: 21 specimens at CSIRO, NMV, SAMA. Plus non-type specimens. •Valid as Dipturus canutus Last 2008 -- (Last & White 2008:109 [ref. 30618], Last & Alava 2013:214 [ref. 33041], Weigmann 2016:108 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:247 [ref. 35000]). Current status: Valid as Dipturus canutus Last 2008. Rajidae. Distribution: Southern Australia: New South Wales to Western Australia, including Tasmania. Habitat: marine.

chilensis, Raia Guichenot [A.] 1848:367 [Fauna Chilena.... in Gay.; ref. 1939] Valparaíso Bay, central Chile, southeastern Pacific Ocean, 32°53'22.94"S, 71°31'32.16"W, depth approximately 40 meters [Chile]. Neotype: MNHNCL ICT 7549. Neotype designated by Concha et al. 2019:507 [ref. 36686]. •Valid as Raja chilensis Guichenot 1848 -- (Pequeño & Lamilla 1985:249 [ref. 9593], Pequeño 1989:17 [ref. 14125], Séret 1989:127 [ref. 12827], Lloris & Rucabado 1991:44 [ref. 22530]). •Valid as Zearaja chilensis (Guichenot 1848) -- (Last & Gledhill 2007:50 [ref. 29177], Last et al. 2008:9 [ref. 29647], Vargas-Caro et al. 2015:325 [ref. 33982], Weigmann 2016:124 [ref. 34211] see remarks, Last et al. 2016:23 [ref. 34941] with remarks, Last et al. 2016:361 [ref. 35000], Nión et al. 2016:22 [ref. 35565], Gabbanelli et al. 2018:458 [ref. 36350]). •Valid as Dipturus chilensis (Guichenot 1848) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Cousseau & Perrotta 2000:46 [ref. 25942], Gomes & Paragó 2001:1 [ref. 25669], Soto & Mincarone 2001:6 [ref. 26630], Díaz de Astarloa et al. 2008:35 [ref. 29969], Mabragaña et al. 2011:7 and table S2 [ref. 31800], Concha et al. 2019:507 [ref. 36686]). Current status: Valid as Dipturus chilensis (Guichenot 1848). Rajidae. Distribution: Southeastern Pacific: central Chile to the Strait of Magellan. Habitat: marine.

chinensis, Raia Basilewsky [S.] 1855:251 [Nouveaux mémoires de la Société impériale des naturalistes de Moscou v. 10; ref. 200] "Oriental Sea", northern China. No types at ZIN. •Valid as Raja chinensis Basilewsky 1855 -- (Cheng & Zhou 1997:47 [ref. 26385], Weigmann 2016:118 [ref. 34211]). •Valid as Dipturus chinensis (Basilewsky 1855) -- (Last et al. 2016:21 [ref. 34941], Last et al. 2016:248 [ref. 35000]). Current status: Valid as Dipturus chinensis (Basilewsky 1855). Rajidae. Distribution: China to Japan including Taiwan. Habitat: marine.

crosnieri, Raja (Dipturus) Séret [B.] 1989:116, Figs. 1-9 [Cybium v. 13 (no. 2); ref. 12827] Southwestern coast of Madagascar, 22°05'S, 22°30'E, depth 300-850 meters. Holotype: MNHN 1988-1089. Paratypes: BMNH 1988.7.25.1 (1); MNHN 1988-1090 to 1108 (19); SAIAB [formerly RUSI] 28021 (1); USNM 297040 (1). •Valid as Dipturus crosnieri (Séret 1989) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Jeong & Nakabo 2008:189 [ref. 29612], Last & Alava 2013:214 [ref. 33041], Weigmann 2016:108 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:249 [ref. 35000], Fricke et al. 2018:24 [ref. 35805]). Current status: Valid as Dipturus crosnieri (Séret 1989). Rajidae. Distribution: Western Indian Ocean off Madagascar. Habitat: marine.

diehli, Dipturus Soto [J. M. R.] & Mincarone [M. M.] 2001:3, Figs. 1-3 [Mare Magnum v. 1 (no. 1); ref. 26630] Off Santa Catarina State, southern Brazil, 29°39'31"S, 47°50'32"W, depth 480 meters. Holotype (unique): MOVI 08494. Type catalog: Soto & Mincarone 2004:94 [ref. 37067]. •Valid as Dipturus diehli Soto & Mincarone 2001 -- (Gomes & Gadig in Menezes et al. 2003:28 [ref. 27192], Jeong & Nakabo 2008:189 [ref. 29612], Díaz de Astarloa et al. 2008:35 [ref. 29969], Vargas-Caro 2015:323 [ref. 33982]). •Synonym of Dipturus mennii Gomes & Paragó 2001 -- (Weigmann 2016:109 [ref. 34211], Last et al. 2016:21 [ref. 34941]). Current status: Synonym of Dipturus mennii Gomes & Paragó 2001. Rajidae. Distribution: Southwestern Atlantic (if valid). Habitat: marine.

doutrei, Raja Cadenat [J.] 1960:294, Figs. 1-11, 13, 15 [Bulletin de l'Institut Français d'Afrique Noire (Sér A) Sciences Naturelles v. 22 (no. 1); ref. 11900] Senegal, depth 450-500 meters. Lectotype: MNHN 1959-0041 [ex IFAN]. Paralectotypes: (30) IFAN 1954-679, 59-804, and R744-751 (8, jaws only), uncat. (6, jaws); MNHN 1959-0042 [ex IFAN] (1, jaws only), 1959-0043 [ex IFAN] (1, jaws only); 13 other syntypes lost. Type catalog: Séret & McEachran 1987:14-15 [ref. 9312]. Lectotype selected by Séret & McEachran 1987:14 [ref. 9312]. •Valid as Raja doutrei Cadenat 1960 -- (Hulley 1986:122 [ref. 5672], Séret 1989:127 [ref. 12827], Stehmann 1990:32 [ref. 19316], Compagno et al. 1991:95 [ref. 21083], Bianchi & Carpenter in Bianchi et al. 1993:93 [ref. 25472], Stehmann 1995:36 [ref. 22055]). •Valid as Dipturus doutrei (Cadenat 1960) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Compagno & Ebert 2007:137 [ref. 30471], Weigmann 2016:108 [ref. 34211], McEachran & Séret 2016:1389 [ref. 34551], Last et al. 2016:21 [ref. 34941], Last et al. 2016:250 [ref. 35000], Almeida & Biscoito 2019:140 [ref. 37302]). Current status: Valid as Dipturus doutrei (Cadenat 1960). Rajidae. Distribution: Eastern Atlantic, southwestern Indian Ocean: Senegal to southeastern South Africa. Habitat: marine.

ecuadoriensis, Raja Beebe [W.] & Tee-Van [J.] 1941:255, Fig. 14 [Zoologica, Scientific Contributions of the New York Zoological Society v. 26 (pt 3, no. 26); ref. 16103] San Helena Bay, Ecuador. Based on Raja steindachneri of Tortonese 1939:3-5, Pl. 1 [ref. 4414]. •Valid as Dipturus ecuadoriensis (Beebe & Tee-Van 1941) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Last & Alava 2013:214 [ref. 33041], Vargas-Caro 2015:323 [ref. 33982], Weigmann 2016:108 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:251 [ref. 35000]). Current status: Valid as Dipturus ecuadoriensis (Beebe & Tee-Van 1941). Rajidae. Distribution: Ecuador. Habitat: marine.

flavirostris, Raja Philippi [R. A.] 1892:3, Pl. 1 (fig. 2) [Anales del Museo Nacional de Chile. Primera seccion, Zoología No. 3; ref. 22759] Bahia de Quinteros, Chile. Holotype (unique): whereabouts unknown. On p. 4 of German edition (Philippi 1893 [ref. 14560]). •Valid as Dipturus flavirostris (Philippi 1892) -- (Gomes & Gadig in Menezes et al. 2003:29 [ref. 27192]). •Synonym of Zearaja chilensis (Guichenot 1848) -- (Vargas-Caro et al. 2015:325 [ref. 33982]). Current status: Synonym of Dipturus chilensis (Guichenot 1848). Rajidae. Habitat: marine.

floridana, Raja Bigelow [H. B.] & Schroeder [W. C.] 1962:173, Figs. 2-3 [Bulletin of the Museum of Comparative Zoology v. 128 (no. 4); ref. 5793] Off Jacksonville, Florida, U.S.A., 29°43'N, 80°10'W, depth 195 fathoms. Holotype: USNM 196441. Paratypes: MCZ 40723-25 (1, 1, 1), plus 27 other specimens. •Valid as Raja floridana Bigelow & Schroeder 1962 -- (Séret 1989:126 [ref. 12827]). •Synonym of Raja teevani Bigelow & Schroeder 1951 -- (Jacob & McEachran 1994:444 [ref. 22422]). •Synonym of Dipturus teevani (Bigelow & Schroeder 1951) -- (authors). Current status: Synonym of Dipturus teevani (Bigelow & Schroeder 1951). Rajidae. Habitat: marine.

flossada, Raia Risso [A.] 1827:145 [Histoire naturelle des principales productions de l'Europe méridionale v. 3; ref. 3757] Nice, France, northwestern Mediterranean Sea. No types known. Type catalog: Séret & McEachran 1987:31 [ref. 9312]. •Synonym of Raja batis Linnaeus 1758 -- (Stehmann 1973:62 [ref. 7168]). •Synonym of Dipturus batis (Linnaeus 1758) -- (Iglésias et al. 2010:327 [ref. 30586] as temporary species cf. flossada, Dyldin 2015:70 [ref. 34524] with question, Weigmann 2016:107 [ref. 34211] as cf. flossada, Last et al. 2016:20 [ref. 34941] as cf. flossada). Current status: Synonym of Dipturus batis (Linnaeus 1758). Rajidae. Habitat: marine.

gaimardi, Raia Gaimard [J. P.] (ex Valenciennes) 1851:Pls. 2-3 [Voyage de la commission scientifique du Nord; ref. 1523] Coast of Iceland, North Atlantic. Holotype (unique): MNHN 0000-1794 (poor condition). Type catalog: Bertin 1939:84 [ref. 20739], Séret & McEachran 1987:15 [ref. 9312]. Original not seen; date may be incorrect. •Synonym of Raja batis Linnaeus 1758 -- (Stehmann 1973:62 [ref. 7168]). •Synonym of Dipturus batis (Linnaeus 1758) -- (Dyldin 2015:70 [ref. 34524]). Current status: Synonym of Dipturus batis (Linnaeus 1758). Rajidae. Habitat: marine.

garricki, Raja Bigelow [H. B.] & Schroeder [W. C.] 1958:213, Figs. 5-7 [Bulletin of the Museum of Comparative Zoology v. 119 (no. 2); ref. 12049] Northern Gulf of Mexico, 28°32'N, 86°20'W, Oregon station 1277, depth 260 fathoms. Holotype: USNM 156711. Paratypes: MCZ 39616 (1). •Valid as Raja garricki Bigelow & Schroeder 1958 -- (Castro-Aguirre & Espinosa Pérez 1996:38 [ref. 22793]). •Valid as Dipturus garricki (Bigelow & Schroeder 1958) -- (McEachran & Dunn 1998:285 [ref. 23312], McEachran & Fechhelm 1998:155 [ref. 23897], Compagno 1999:490 [ref. 25589], Gomes & Paragó 2001:1 [ref. 25669], McEachran & Carvalho 2003:552 [ref. 26985], Jeong & Nakabo 2008:189 [ref. 29612], Del Moral-Flores et al. 2016:113 [ref. 34398], Weigmann 2016:108 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:252 [ref. 35000], Ehemann et al. 2018:25 [ref. 36194], Ehemann et al. 2019:[6] [ref. 36861], Jones et al. 2020:288 [ref. 37610]). Current status: Valid as Dipturus garricki (Bigelow & Schroeder 1958). Rajidae. Distribution: Western Atlantic: including Gulf of Mexico and Caribbean Sea. Habitat: marine.

gigas, Raja (Tengujei) Ishiyama [R.] 1958:386 [194], Figs. 84 W, 86 [Journal of the Shimonoseki College of Fisheries v. 7 (nos 2-3); ref. 2308] Off Aichi Prefecture, Kumano-nada, Japan. Holotype: not researched. Paratypes: (20 specimens, 2 egg capsules) CAS 23173 (1), MCZ 40292 (1). •Valid as Raja gigas Ishiyama 1958 -- (Nakaya in Okamura et al. 1982:55, 314 [ref. 8256], Nakaya in Masuda et al. 1984:13 [ref. 6441], Nakaya in Okamura & Kitajima 1984:67, 308 [ref. 8057], Ishihara 1987:278 [ref. 6264], Séret 1989:128 [ref. 12827], Compagno et al. 1999:1407 [ref. 24636], Last & Compagno 1999:1454 [ref. 24639], Nakabo 2000:172 [ref. 25086], Compagno in Randall & Lim 2000:582 [ref. 25122], Shinohara et al. 2001:291 [ref. 25995]). •Valid as Dipturus gigas (Ishiyama 1958) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Nakabo 2002:172 [ref. 26001], Shinohara et al. 2005:397 [ref. 28370], Compagno et al. 2005:66 [ref. 29145], Jeong & Nakabo 2008:183 [ref. 29612], Séret & Last 2008:107 [ref. 29649], Last & Alava 2013:214 [ref. 33041], Hsu et al. 2013:253 [ref. 33043], Ebert et al. 2013:352 [ref. 33045], Motomura et al. 2015:12 [ref. 34709], Weigmann 2016:108 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:253 [ref. 35000]). Current status: Valid as Dipturus gigas (Ishiyama 1958). Rajidae. Distribution: Western Pacific: Japan to Philippines. Habitat: marine.

grahami, Dipturus Last [P. R.] 2008:84, Figs. 27-32 [CSIRO Marine and Atmospheric Research Paper No. 021; ref. 29648] East of Terrigal, New South Wales, Australia, 33°30'S, 152°04'E, depth 446-515 meters. Holotype: CSIRO H 950-01. Paratypes: 17 specimens as AMS, CSIRO and QM. Plus non-type specimens. Originally appeared as grahami, but named for two ichthyologists, Alastair Graham and Ken Graham; therefore mandatory change to grahamorum according to ICZN Art. 31.1. •Valid as Dipturus grahamorum Last 2008 -- (Last & Alava 2013:214 [ref. 33041] as grahami, Weigmann 2016:108 [ref. 34211] as grahami, Last et al. 2016:21 [ref. 34941] as grahami, Last et al. 2016:254 [ref. 35000] as grahami). Current status: Valid as Dipturus grahamorum Last 2008. Rajidae. Distribution: Eastern Australia: Queensland to New South Wales. Habitat: marine.

granulata, Raia Gill [T. N.] in Goode & Bean 1879:28 [Bulletin of the Essex Institute v. 11; ref. 5605] Le Have Bank, off Nova Scotia, Canada. Holotype: USNM 21578 (not found in 1980). •Synonym of Raja laevis Mitchill 1818 -- (Scott & Scott 1988:47 [ref. 25518]). •Synonym of Dipturus laevis (Mitchill 1818) -- (current authors). Current status: Synonym of Dipturus laevis (Mitchill 1818). Rajidae. Habitat: marine.

gudgeri, Zearaja Whitley [G. P.] 1940:188, Fig. 217 [The fishes of Australia. Part I; ref. 4700] Great Australian Bight, Australia. Holotype: AMS E.4284. Paratypes: AMS IB.768 (1), IB.769 (?1). •Valid as Raja gudgeri (Whitley 1940) -- (Séret 1989:128 [ref. 12827], Paxton et al. 1989:56 [ref. 12442], Gomon et al. 1994:166 [ref. 22532], Last & Stevens 1994:349 [ref. 23873], Hutchins 2001:15 [ref. 25847]). •Valid as Dipturus gudgeri (Whitley 1940) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Last & Yearsley 2002:1640 [ref. 26568], Hoese et al. 2006:182 [ref. 29001], Séret & Last 2008:107 [ref. 29649], Last & White 2008:116 [ref. 30618], Last & Alava 2013:214 [ref. 33041], Weigmann 2016:108 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:255 [ref. 35000]). Current status: Valid as Dipturus gudgeri (Whitley 1940). Rajidae. Distribution: Australia: New South Wales to southern Australia and to Western Australia. Habitat: marine.

hispanica, Raja Bloch [M. E.] & Schneider [J. G.] 1801:369 [M. E. Blochii, Systema Ichthyologiae; ref. 471] Spain, western Mediterranean Sea. Based on "Raja, Hispanorum Machuelo" of Osbeck 1770:99 [ref. 18497]. •Synonym of Dipturus oxyrinchus (Linnaeus 1758). Current status: Synonym of Dipturus oxyrinchus (Linnaeus 1758). Rajidae. Habitat: marine.

hyposticta, Propterygia Otto [A. W.] 1821:6 [Conspectus animalium quorundam maritimorum; ref. 4987] Newhaven, Edinburgh. Also described as new in Otto 1821:112 [ref. 3321], earliest not researched, and again in Otto 1824:94 [ref. 26341]. Based on an abnormal specimen of Raja batis Linnaeus 1758. •Synonym of Raja batis Linnaeus 1758 -- (authors). •Synonym of Dipturus batis (Linnaeus 1758) -- (Dyldin 2015:70 [ref. 34524]). Current status: Synonym of Dipturus batis (Linnaeus 1758). Rajidae. Habitat: marine.

innominata, Raja Garrick [J. A. F.] & Paul [L. J.] 1974:357, Figs. 6-7 [Journal of the Royal Society of New Zealand v. 4 (no. 3); ref. 7421] Cape Campbell, New Zealand. Holotype: NMNZ P.1441. Paratypes: NMNZ P.1040 (1), P.1817 (1), P.3094 (2), P.4680 (1), P.5483-87 (1, 4, 1, 2, 2), P.5534 (1), 5546 (1). Type catalog: Hardy 1990:6 [ref. 18942]. •Valid as Raja innominata Garrick & Paul 1974 -- (Séret 1989:128 [ref. 12827], Paulin et al. 1989:28 [ref. 24556], Ishihara in Amaoka et al. 1990:86 [ref. 16024]). •Valid as Dipturus innominatus (Garrick & Paul 1974) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Last & Yearsley 2002:1640 [ref. 26568] as innominata, Last & Alava 2013:214 [ref. 33041], Last & Stewart 2015:178 [ref. 34224], Weigmann 2016:108 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:256 [ref. 35000]). Current status: Valid as Dipturus innominatus (Garrick & Paul 1974). Rajidae. Distribution: New Zealand. Habitat: marine.

intermedia, Raia Parnell [R.] 1837:167, Pl. 2 [Proceedings of the Royal Society of Edinburgh v. 1; ref. 18522] Firth of Forth, Scotland, North Sea. Syntypes: (all dry) BMNH 1848.1.12.512 (1), 1848.1.12.568 (1), 1848.1.12.570 (1). Original genus spelled Raia on Pl. 2 but not spelled out in text. •Synonym of Raja batis Linnaeus 1758 -- (Stehmann 1973:62 [ref. 7168]). •Valid as Dipturus cf. intermedia (Parnell 1837) -- (Gordon et al. 2016:261 [ref. 34661]). •Synonym of Dipturus batis (Linnaeus 1758) -- (Iglésias et al. 2010:329 [ref. 30586] as temporary species cf. intermedia, Dyldin 2015:70 [ref. 34524] with question, Weigmann 2016:107 [ref. 34211] as cf. intermedia). •Valid as Dipturus intermedius (Parnell 1837) -- (Last et al. 2016:21 [ref. 34941], Last et al. 2016:257 [ref. 35000], Biscoito et al. 2018:476 [ref. 37562], Almeida & Biscoito 2019:139 [ref. 37302], Carneiro et al. 2019:43 [ref. 37250]). Current status: Valid as Dipturus intermedius (Parnell 1837). Rajidae. Distribution: Western Mediterranean Sea; northeastern Atlantic: Iceland and Norway south to Senegal, including Madeira and Canary islands. Habitat: marine.

johannisdavisi, Raja Alcock [A. W.] 1899:21 [A descriptive catalogue of the Indian deep-sea fishes in the Indian Museum; ref. 5114] Laccadive Sea, off Travancore coast, 11°14'30"N, 74°57'15"E, Investigator station 246, depth 410-520 meters. Holotype (unique): ZSI F477/1. Type catalog: Menon & Yazdani 1968:96 [ref. 20743], Menon & Rama-Rao 1975:32 [ref. 22148]. Originally as johannis-davisi. Figured in Alcock 1900: Pl. 27 (figs. 2, 2a) [ref. 20536]. •Valid as Raja johannisdavisi Alcock 1899 -- (Stehmann 1976:154 [ref. 7085], Séret 1986:258 [ref. 6753], Séret 1989:127 [ref. 12827]). •Valid as Dipturus johannisdavisi (Alcock 1899) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Manilo & Bogorodsky 2003:S93 [ref. 27377] Last & Alava 2013:214 [ref. 33041], Weigmann 2016:108 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:258 [ref. 35000], Psomadakis et al. 2020:201 [ref. 37272]). Current status: Valid as Dipturus johannisdavisi (Alcock 1899). Rajidae. Distribution: Indian Ocean. Habitat: marine.

kwangtungensis, Raja Chu [Y.-T.] 1960:148, Figs. 137-139 [Cartilaginous fishes of China; ref. 21272] Jia-Bo, China. Holotype: SFI 57-0676. Paratypes: SFI 57-0674 (1), 57-0675 (1). •Valid as Raja kwangtungensis Chu 1960 -- (Ishihara 1987:242 [ref. 6264], Séret 1989:128 [ref. 12827], Compagno 1999:490 [ref. 25589], Nakabo 2000:172 [ref. 25086], Compagno in Randall & Lim 2000:582 [ref. 25122], Shinohara et al. 2001:291 [ref. 25995], Nakabo 2002:172 [ref. 26001]). •Valid as Dipturus kwangtungensis (Chu 1960) -- (McEachran & Dunn 1998:285 [ref. 23312], Jeong 1999:202 [ref. 25171], Choi et al. 2003:66, 482 [ref. 26218], Shinohara et al. 2005:397 [ref. 28370], Jeong & Nakabo 2008:183 [ref. 29612], Last & Lim 2010:107 [ref. 30853], Last el al. 2010:170 [ref. 32461], Yamashita et al. 2012:129 [ref. 32408], Last & Alava 2013:214 [ref. 33041], Hsu et al. 2013:253 [ref. 33043], Ebert et al. 2013:352 [ref. 33045], Shinohara et al. 2014:235 [ref. 33330], Hara et al. 2016:5 [ref. 36169] as cf. kwanftungensis, Last et al. 2016:21 [ref. 34941], Last et al. 2016:259 [ref. 35000], Weigmann 2016:108 [ref. 34211], Sonoyama et al. 2020:12 [ref. 37637]). Current status: Valid as Dipturus kwangtungensis (Chu 1960). Rajidae. Distribution: Western Pacific: Borneo and China north to Japan. Habitat: marine.

laevis, Raja Mitchill [S. L.] 1818:327 [American Monthly Magazine and Critical Review v. 2 (no. 5, art. 1); ref. 30214] New York City by East River, New York, U.S.A. Holotype (unique): whereabouts unknown. •Valid as Raja laevis Mitchill 1818 -- (McEachran & Fechhelm 1982:449 [ref. 8444], Smith 1986:391 [ref. 22186], Robins & Ray 1986:39 [ref. 23100], Séret 1989:126 [ref. 12827], McAllister 1990:37 [ref. 14674], Murdy et al. 1997:38 [ref. 23144]). •Valid as Dipturus laevis (Mitchill 1818) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Musick et al. 2000:9 [ref. 24959], Gomes & Paragó 2001:1 [ref. 25669], McEachran in Collette & Klein-MacPhee 2002:65 [ref. 26158], Moore et al. 2003:171 [ref. 27076], Nelson et al. 2004:55 [ref. 27807], Page et al. 2013:55 [ref. 32708], Weigmann 2016:109 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:260 [ref. 35000]). Current status: Valid as Dipturus laevis (Mitchill 1818). Rajidae. Distribution: Western Atlantic. Habitat: marine.

lamillai, Dipturus Concha [F. J.], Caira [J. N.], Ebert [D. A.] & Pompert [J. H. W.] 2019:513, Figs. 7-11, 12C-D [Zootaxa 4590 (no. 5); ref. 36686] Off the Falkland Islands, southwestern Atlantic Ocean, Station 2019, 49.61°S, 61.19°W, depth 162 meters. Holotype: MNHNCL ICT 7531. Paratypes: CAS 242403 (1), MNHNCL ICT 7532 (1). Plus additional non-type material. •Valid as Dipturus lamillai Concha, Caira, Ebert & Pompert 2019. Current status: Valid as Dipturus lamillai Concha, Caira, Ebert & Pompert 2019. Rajidae. Distribution: Southwestern Atlantic adjacent to the Falkland Islands Habitat: marine.

lanceorostrata, Raja Wallace [J. H.] 1967:15, Fig. 8 [Investigational Report. Oceanographic Research Institute, Durban No. 17; ref. 7929] Off Limpopo River mouth, South Africa, depth 235-240 fathoms. Holotype: SAIAB [formerly RUSI] 986 [ex ORI B869]. Paratypes: SAIAB [formerly RUSI] 987 [ex ORI B868] (1). Additional material: ORI uncat. (1). •Valid as Raja lanceorostrata Wallace 1967 -- (McEachran & Fechhelm 1982:449 [ref. 8444], Hulley 1986:122 [ref. 5672], Séret 1989:127 [ref. 12827]). •Valid as Dipturus lanceorostratus (Wallace 1967) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Compagno & Ebert 2007:139 [ref. 30471], Shao et al. 2008:237 [ref. 29952], Last & Alava 2013:214 [ref. 33041], Weigmann 2016:109 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:261 [ref. 35000]). Current status: Valid as Dipturus lanceorostratus (Wallace 1967). Rajidae. Distribution: Mozambique and ?Taiwan. Habitat: brackish, marine.

latastei, Raja Delfin [F. T.] 1902:264, Pl. 11 [Revista Chilena de Historia Natural (Santiago) v. 6; ref. 17397] Chile. Syntypes: (2). •Synonym of Raja chilensis Guichenot 1848 -- (Lloris & Rucabado 1991:44 [ref. 22530]). •Valid as Zearaja chilensis (Guichenot 1848) -- (Vargas-Caro et al. 2015:325 [ref. 33982]). Current status: Synonym of Dipturus chilensis (Guichenot 1848). Rajidae. Habitat: marine.

leptocauda, Raja (Dipturus) Krefft [G.] & Stehmann [M. F. W.] 1975:79, Figs. 1-2 [Archiv für Fischereiwissenschaft v. 26 (no. 3); ref. 7435] Off southern Brazil, 24°21'S, 43°54'W, depth 500 meters. Holotype: ZMH 25254 [ex ISH 1913-1968]. Type catalog: Krefft 1978:6 [ref. 9884]. •Valid as Raja leptocauda Krefft & Stehmann 1975 -- (Séret 1989:127 [ref. 12827]). •Valid as Dipturus leptocauda (Krefft & Stehmann 1975) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Menni & Stehmann 2000:80 [ref. 24909], Gomes & Paragó 2001:1 [ref. 25669], Soto & Mincarone 2001:6 [ref. 26630], Gomes & Gadig in Menezes et al. 2003:29 [ref. 27192], Jeong & Nakabo 2008:189 [ref. 29612], Díaz de Astarloa et al. 2008:35 [ref. 29969], Vargas-Caro 2015:323 [ref. 33982], Weigmann 2016:109 [ref. 34211] as leptocaudus, Last et al. 2016:21 [ref. 34941] as leptocaudus, Last et al. 2016:262 [ref. 35000] as leptocaudus, Nión et al. 2016:22 [ref. 35565]). Current status: Valid as Dipturus leptocauda (Krefft & Stehmann 1975). Rajidae. Distribution: Southwestern Atlantic. Habitat: marine.

macrocauda, Raja Ishiyama [R.] 1955:43, Figs. 1-2 [Journal of the Shimonoseki College of Fisheries v. 4 (no. 1); ref. 12276] Miya fish market, Japan. Holotype: not researched. Paratypes: (23) CAS 23169 (1), 23172 (1). •Valid as Raja macrocauda Ishiyama 1955 -- (Nakaya in Masuda et al. 1984:13 [ref. 6441], Nakaya in Okamura & Kitajima 1984:65, 307 [ref. 8057], Ishihara 1987:278 [ref. 6264], Séret 1989:128 [ref. 12827], Compagno in Randall & Lim 2000:582 [ref. 25122], Shinohara et al. 2001:291 [ref. 25995]). •Valid as Dipturus macrocauda (Ishiyama 1955) -- (McEachran & Dunn 1998:285 [ref. 23312], Jeong 1999:202 [ref. 25171], Compagno 1999:490 [ref. 25589], Nakabo 2000:173 [ref. 25086], Nakabo 2002:173 [ref. 26001], Choi et al. 2003:66, 482 [ref. 26218], Shinohara et al. 2005:398 [ref. 28370], Jeong & Nakabo 2008:189 [ref. 29612], Last & Alava 2013:214 [ref. 33041] as macrocaudus, Hsu et al. 2013:253 [ref. 33043], Ebert et al. 2013:353 [ref. 33045], Shinohara et al. 2014:235 [ref. 33330], Weigmann 2016:109 [ref. 34211] as macrocaudus, Last et al. 2016:21 [ref. 34941] as macrocaudus, Last et al. 2016:263 [ref. 35000] as macrocaudus). Current status: Valid as Dipturus macrocauda (Ishiyama 1955). Rajidae. Distribution: Western North Pacific: Japan, Taiwan and China. Habitat: marine.

macrorynchus, Raja Rafinesque [C. S.] 1810:15 [Caratteri di alcuni nuovi generi e nuove specie di animali e piante della Sicilia; ref. 3594] Sicily, Italy, Mediterranean Sea. No types known. Appeared first in Rafinesque 1810:47 [ref. 3595] with citation to Rafinesque 1810 [ref. 3594] [3594 technically published later]. •Valid as Raja macrorynchus Rafinesque 1810 -- (Stehmann 1990:33 [ref. 19316], Biscoito et al. 2018:488 [ref. 37562]). •Synonym of Dipturus batis (Linnaeus 1758) -- (Lipej & Dulčić 2010:13 [ref. 36649] as macrorhynchus, Parenti 2019:103 [ref. 37117]). Current status: Synonym of Dipturus batis (Linnaeus 1758). Rajidae. Habitat: marine.

maugeana, Zearaja Last [P. R.] & Gledhill [D. C.] 2007:52, Figs. 4-13 [Zootaxa No. 1494; ref. 29177] Macquarie Harbour, 42°21'S, 145°32'E, Australia. Holotype: CSIRO H 3975-01. Paratypes: AMS I 40748-001 (1); CSIRO H 1987-01 (1), H 3747-01 (1), H 3976-01 (1), H 3976-02 (1), H 4486-01 (1), H 4487-01 (1), H 5544-01 (1). •Valid as Zearaja maugeana Last & Gledhill 2007 -- (Last et al. 2008:9 [ref. 29647], Weigmann 2016:124 [ref. 34211], Last et al. 2016:23 [ref. 34941], Last et al. 2016:362 [ref. 35000]). •Valid as Dipturus maugeanus (Last & Gledhill 2007) -- (Concha et al. 2019:507 [ref. 36686]). Current status: Valid as Dipturus maugeanus (Last & Gledhill 2007). Rajidae. Distribution: Southwestern Tasmania, Australia. Habitat: brackish.

melanospilus, Dipturus Last [P. R.], White [W. T.] & Pogonoski [J. J.] 2008:34, Figs. 20-25 [CSIRO Marine and Atmospheric Research Paper No. 021; ref. 29647] South of Saumarez Reef, Saumarez Plateau, Queensland, Australia, 22°49'S, 154°10'E, depth 445-450 meters. Holotype: CSIRO H 715-05. Paratypes: 21 specimens at AMS, CSIRO, QM. Plus non-type specimens. •Valid as Dipturus melanospilus Last, White & Pogonoski 2008 -- (Last & Alava 2013:214 [ref. 33041], Weigmann 2016:109 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:264 [ref. 35000]). Current status: Valid as Dipturus melanospilus Last, White & Pogonoski 2008. Rajidae. Distribution: Eastern Australia: Queensland to New South Wales. Habitat: marine.

mennii, Dipturus Gomes [U. L.] & Paragó [C.] 2001:2, Figs. 1-3 [Boletim do Museu Nacional do Rio de Janeiro No. 448; ref. 25669] Santa Catarina, 26°44'15"S, 47°09'57", western Brazil. Holotype: MZUSP 51600. Paratypes: UERJ 1893 (1). •Valid as Dipturus mennii Gomes & Paragó 2001 -- (Gomes & Gadig in Menezes et al. 2003:29 [ref. 27192] as menni, Díaz de Astarloa et al. 2008:35 [ref. 29969], Vargas-Caro 2015:323 [ref. 33982], Weigmann 2016:109 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:265 [ref. 35000]). Current status: Valid as Dipturus mennii Gomes & Paragó 2001. Rajidae. Distribution: Brazil. Habitat: marine.

nasuta, Raja Müller [J.] & Henle [F. G. J.] (ex Banks) 1841:150 [Systematische Beschreibung der Plagiostomen; ref. 3069] Australian sea and near Totaranni, New Zealand. No types known. •Valid as Raja nasuta Müller & Henle 1841 -- (Garrick & Paul 1974:353 [ref. 7421], Séret 1989:128 [ref. 12827], Paxton et al. 1989:56 [ref. 12442], Paulin et al. 1989:28 [ref. 24556], Ishihara in Amaoka et al. 1990:87 [ref. 16024]). •Valid as Dipturus nasutus (Müller & Henle 1841) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], Last & Yearsley 2002:1640 [ref. 26568] as nasuta with author as Banks). •Valid as Zearaja nasuta (Müller & Henle 1841) -- (Last & Gledhill 2007:50 [ref. 29177], Last et al. 2008:9 [ref. 29647], Last & Stewart 2015:179 [ref. 34224], Weigmann 2016:124 [ref. 34211], Last et al. 2016:23 [ref. 34941], Last et al. 2016:363 [ref. 35000], Concha et al. 2019:506 [ref. 36686]). Current status: Valid as Dipturus nasutus (Müller & Henle 1841). Rajidae. Distribution: New Zealand and nearby plateaus and the Chatham Rise. Habitat: marine.

nidarosiensis, Raja Storm [V.] (ex Collett) 1881:80 [Det Kongelige Norske videnskabers selskabs skrifter, Trondheim 1880; ref. 19334] Munkholmen, Trondheim Fjord, Norway, eastern North Atlantic. Syntypes: ZMUO J6339 (1, stuffed male in poor condition), ZMUO (2, lost). Type catalog: Pethon 1969:3 [ref. 19268]. Also appeared as new in Collett 1882:1, Pl. (figs. 1-5) [ref. 18020] [date from Pethon] as Raja nidrosiensis. •Valid as Raja nidarosiensis Storm 1881 -- (Stehmann 1973:63 [ref. 7168], Stehmann & Bürkel in Whitehead et al. 1984:176 [ref. 13675], Séret 1989:126 [ref. 12827], Stehmann 1990:32 [ref. 19316], Stehmann 1995:105 [ref. 22055]). •Valid as Dipturus nidarosiensis (Storm 1881) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:490 [ref. 25589], George 2009:56 [ref. 30539], Iglésias et al. 2010:329 [ref. 30586], Cannas et al. 2010:341 [ref. 30787], Weigmann 2016:109 [ref. 34211] see remarks, McEachran & Séret 2016:1396 [ref. 34551], Last et al. 2016:21 [ref. 34941] with remarks, Last et al. 2016:266 [ref. 35000], Almeida & Biscoito 2019:137 [ref. 37302], Carbonara et al. 2019:4 [ref. 36762], Carneiro et al. 2019:43 [ref. 37250], Geraci et al. 2019:204 [ref. 36776], Kovačić et al. 2020:7 [ref. 37519]). Current status: Valid as Dipturus nidarosiensis (Storm 1881). Rajidae. Distribution: Western and central Mediterranean Sea; eastern Atlantic: Iceland south to South Africa. Habitat: marine.

obscura, Raia Cabrera y Corro [A.], Pérez [L.] & Hänseler [F.] 1817:12 [Lista de los peces del mar de Andalucía; ref. 17319] Western Andalusia, near Cádiz, Spain, eastern Atlantic. Genus should have been Raja. Name only, not available. Possibly an unpublished work. See also Raia obscura Cabrera y Corro, Pérez & Hänseler in Graells 1887:186 [ref. 30438]. •In the synonymy of Dipturus batis (Linnaeus 1758) -- (Dyldin 2015:70 [ref. 34524]). Current status: Synonym of Dipturus batis (Linnaeus 1758). Rajidae.

obscura, Raia Cabrera y Corro [A.], Pérez [L.] & Hänseler [F.] in Graells 1887:186 [Revista de los Progresos de las Ciencias Exactas, Fisicas y Naturales, Madrid v. 22 (no. 3); ref. 30438] Western Andalusia, near Cádiz, Spain, eastern Atlantic. Genus should have been Raja. Name only; not available. Name from Cabrera y Corro, Pérez & Hänseler 1817:12 [ref. 17319], possibly an unpublished work. •In the synonymy of Dipturus batis (Linnaeus 1758). Current status: Synonym of Dipturus batis (Linnaeus 1758). Rajidae.

olseni, Raja Bigelow [H. B.] & Schroeder [W. C.] 1951:386, Fig. 2 [Journal of the Washington Academy of Sciences v. 41 (no. 12); ref. 12883] Gulf of Mexico, 27°25'N, 96°13'W, Oregon station 157, depth 76 fathoms. Holotype: USNM 153556. Paratypes: MCZ 37176 (1). •Valid as Raja olseni Bigelow & Schroeder 1951 -- (McEachran & Fechhelm 1982:447 [ref. 8444], Robins & Ray 1986:39 [ref. 23100], Séret 1989:126 [ref. 12827], Boschung 1992:26 [ref. 23239], Castro-Aguirre & Espinosa Pérez 1996:39 [ref. 22793]). •Valid as Dipturus olseni (Bigelow & Schroeder 1951) -- (McEachran & Dunn 1998:285 [ref. 23312], McEachran & Fechhelm 1998:156 [ref. 23897], Compagno 1999:490 [ref. 25589], Gomes & Paragó 2001:1 [ref. 25669], McEachran & Carvalho 2003:552 [ref. 26985], Nelson et al. 2004:55 [ref. 27807], Page et al. 2013:55 [ref. 32708], Del Moral-Flores et al. 2016:113 [ref. 34398], Weigmann 2016:109 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:267 [ref. 35000], Ehemann et al. 2018:25 [ref. 36194], Jones et al. 2020:288 [ref. 37610]). Current status: Valid as Dipturus olseni (Bigelow & Schroeder 1951). Rajidae. Distribution: Western Atlantic: Gulf of Mexico and ?Caribbean Sea. Habitat: marine.

oregoni, Raja Bigelow [H. B.] & Schroeder [W. C.] 1958:201, Figs. 1-2 [Bulletin of the Museum of Comparative Zoology v. 119 (no. 2); ref. 12049] Northern Gulf of Mexico, 29°10'N, 88°13'W, Oregon station 1247, depth 205 fathoms. Holotype: USNM 156710. Paratypes: MCZ 39617 (1). Plus non-type material. •Valid as Raja oregoni Bigelow & Schroeder 1958 -- (Séret 1989:126 [ref. 12827], Castro-Aguirre & Espinosa Pérez 1996:39 [ref. 22793]). •Valid as Dipturus oregoni (Bigelow & Schroeder 1958) -- (McEachran & Dunn 1998:285 [ref. 23312], McEachran & Fechhelm 1998:157 [ref. 23897], Compagno 1999:491 [ref. 25589], Gomes & Paragó 2001:1 [ref. 25669], McEachran & Carvalho 2003:553 [ref. 26985], Del Moral-Flores et al. 2016:115 [ref. 34398], Weigmann 2016:109 [ref. 34211], Last et al. 2016:21 [ref. 34941], Last et al. 2016:268 [ref. 35000], Ehemann et al. 2018:25 [ref. 36194], Jones et al. 2020:289 [ref. 37610]). Current status: Valid as Dipturus oregoni (Bigelow & Schroeder 1958). Rajidae. Distribution: Western Atlantic: eastern Florida (U.S.A.) and Gulf of Mexico. Habitat: marine.

oxyptera, Raja Philippi [R. A.] 1892:4, Pl. 2 (fig. 1) [Anales del Museo Nacional de Chile. Primera seccion, Zoología No. 3; ref. 22759] Chile. Holotype (unique): whereabouts unknown. On p. 5 of German edition (Philippi 1893 [ref. 14560]). •Synonym of Raja chilensis Guichenot 1848 -- (Lloris & Rucabado 1991:44 [ref. 22530]). •Synonym of Zearaja chilensis (Guichenot 1848) -- (Vargas-Caro et al. 2015:325 [ref. 33982]). Current status: Synonym of Dipturus chilensis (Guichenot 1848). Rajidae. Habitat: marine.

oxyrinchus, Raja Linnaeus [C.] 1758:231 [Systema Naturae, Ed. X v. 1; ref. 2787] Mediterranean Sea [original: "in M. Mediterraneo & O. Europæo"]; localities include Marseille, France; Liguria and Rome, Italy; Syria; Lesbos Island, Greece. No types known. Lectotype or neotype fixation needed. Originally based on multiple species and at least 25 pre-Linnaean sources including Artedi 1738:72 [ref. 30578] and Artedi 1738:101 [ref. 30349] as Raja varia, tuberculi decem aculeatis in medio dorsi; Rondelet 1554:348 [ref. 30354] as alia Raja, Gesner 1558:938 [ref. 30350] as Raia oxyrhynchos maior, Salviani 1558:149, Pl. 51 [ref. 30229] as Raia, Jonston 1649:Pl. 9 (fig. 8) [ref. 30579] as Raia oxyrhynchos maior, Ray 1713:26 [ref. 30231] as Raia oxyrhynchos major. Spelled oxyrinchos in Brünnich 1768:2 [ref. 17290], and oxirincus in Cabrera y Corro, Pérez & Hänseler 1817:12 [ref. 17319], see Graells 1887:186 [ref. 30438]. •Valid as Raja oxyrinchus Linnaeus 1758 -- (Stehmann 1973:63 [ref. 7168], Stehmann & Bürkel in Whitehead et al. 1984:177 [ref. 13675], Séret 1989:126 [ref. 12827], Stehmann 1990:33 [ref. 19316], Stehmann 1995:105 [ref. 22055], Santos et al. 1997:119 [ref. 23531], Arruda 1997:22 [ref. 24952], Uiblein et al. 1999:60 [ref. 24045], Golani 2005:14 [ref. 37112]). •Valid as Dipturus oxyrinchus (Linnaeus 1758) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:491 [ref. 25589] as oxyrhynchus, Bilecenoğlu et al. 2002:21 [ref. 26753], George 2009:56 [ref. 30539], Iglésias et al. 2010:329 [ref. 30586], Lipej & Dulčić 2010:13 [ref. 36649], Naylor et al. 2012:95 [ref. 32267] see remarks, Dyldin 2015:70 [ref. 34524], Weigmann 2016:109 [ref. 34211], McEachran & Séret 2016:1397 [ref. 34551], Last et al. 2016:22 [ref. 34941] with remarks, Last et al. 2016:269 [ref. 35000], Biscoito et al. 2018:477 [ref. 37562], Almeida & Biscoito 2019:138 [ref. 37302], Carneiro et al. 2019:43 [ref. 37250], Bariche & Fricke 2020:26 [ref. 37515], Kovačić et al. 2020:7 [ref. 37519]). Current status: Valid as Dipturus oxyrinchus (Linnaeus 1758). Rajidae. Distribution: North Sea; Mediterranean Sea; eastern Atlantic: Norway south to Senegal, including Madeira (rare) and Canary Islands. Habitat: marine.

oxyrynchus, Raia Stephan [C. F.] 1779:22 [De rajis, schediasma primum; ref. 18818] •Synonym of Dipturus oxyrinchus (Linnaeus 1758) -- (Kottelat 2010:64 [ref. 30995]). Current status: Synonym of Dipturus oxyrinchus (Linnaeus 1758). Rajidae. Habitat: marine.

pullopunctata, Raia Smith [J. L. B.] 1964:285, Pl. 25 (fig. A) [Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, Kjøbenhavn. v. 126; ref. 3444] Algoa Bay, 31°7'E, 29°59'S, off coast of South Africa, western Indian Ocean, depth 100 fathoms. Holotype: SAIAB [formerly RUSI] 37. Paratypes: (1) not researched. •Valid as Raja pullopunctata Smith 1964 -- (Hulley 1986:123 [ref. 5672], Séret 1989:127 [ref. 12827], Compagno et al. 1991:95 [ref. 21083], Bianchi & Carpenter in Bianchi et al. 1993:94 [ref. 25472], Stehmann 1995:106 [ref. 22055], Walmsley-Hart et al. 1999:165 [ref. 24222]). •Valid as Dipturus pullopunctatus (Smith 1964) (often as pullopunctata) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:491 [ref. 25589], Heemstra & Heemstra 2004:81 [ref. 28072], Compagno & Ebert 2007:139 [ref. 30471], Last & Alava 2013:214 [ref. 33041], Weigmann 2016:110 [ref. 34211], Last et al. 2016:22 [ref. 34941], Last et al. 2016:270 [ref. 35000], Almeida & Biscoito 2019:138 [ref. 37302]). Current status: Valid as Dipturus pullopunctatus (Smith 1964). Rajidae. Distribution: Southeastern Atlantic, southwestern Indian Ocean: Namibia and South Africa. Habitat: marine.

queenslandicus, Dipturus Last [P. R.], White [W. T.] & Pogonoski [J. J.] 2008:42, Figs. 26-31 [CSIRO Marine and Atmospheric Research Paper No. 021; ref. 29647] South of Saumarez Reef, Saumarez Plateau, Queensland, Australia, 22°49'S, 154°10'E, depth 445-450 meters. Holotype: CSIRO H 715-07. Paratypes: 11specimens at CSIRO and QM. Plus one non-type specimen. •Valid as Dipturus queenslandicus Last, White & Pogonoski 2008 -- (Last & Alava 2013:214 [ref. 33041], Weigmann 2016:110 [ref. 34211], Last et al. 2016:22 [ref. 34941], Last et al. 2016:271 [ref. 35000]). Current status: Valid as Dipturus queenslandicus Last, White & Pogonoski 2008. Rajidae. Distribution: Queensland, Australia. Habitat: marine.

salviani, Raja Müller [J.] & Henle [F. G. J.] 1841:143, 195 [Systematische Beschreibung der Plagiostomen; ref. 3069] Mediterranean Sea; Atlantic. Syntypes or possible syntypes: MNHN 0000-1570 (1) Nice, 0000-1791 (1) Nice, 0000-3479 (1, stuffed) Nice; RMNH 4240 (1) or D2511 (1, dry), D2509 (1); ZMB 4599 (1) Sicily. Type catalog: Bertin 1939:84 [ref. 20739], Séret & McEachran 1987:15 [ref. 9312], Paepke & Schmidt 1988:174 [ref. 21041]. Based on Laeviraja of Salviani 1558:149, Pl. 52 [ref. 30372] and other sources. •Synonym of Raja oxyrinchus Linnaeus 1758 -- (Stehmann 1973:63 [ref. 7168]). •Synonym of Dipturus oxyrinchus (Linnaeus 1758) -- (Dyldin 2015:70 [ref. 34524]). Current status: Synonym of Dipturus oxyrinchus (Linnaeus 1758). Rajidae. Habitat: marine.

springeri, Raja Wallace [J. H.] 1967:18, Figs. 9-10 [Investigational Report. Oceanographic Research Institute, Durban No. 17; ref. 7929] 48 kilometers east-southeast of Durban, KwaZulu-Natal, South Africa, western Indian Ocean, depth 549 meters. Holotype: SAIAB [formerly RUSI] 989 [ex ORI B909]. Paratypes: SAIAB [formerly RUSI] 990 [ex ORI B185] (1). Additional material: ORI uncat. (7). •Valid as Raja springeri Wallace 1967 -- (Hulley 1986:125 [ref. 5672], Séret 1989:127 [ref. 12827], Compagno et al. 1991:96 [ref. 21083], Bianchi & Carpenter in Bianchi et al. 1993:95 [ref. 25472], Stehmann 1995:105 [ref. 22055]). •Valid as Dipturus springeri (Wallace 1967) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:491 [ref. 25589], Compagno et al. 2005:66 [ref. 29145], Compagno & Ebert 2007:139 [ref. 30471], Last & Alava 2013:214 [ref. 33041], Weigmann 2016:110 [ref. 34211], Last et al. 2016:22 [ref. 34941], Last et al. 2016:272 [ref. 35000], Fricke et al. 2018:24 [ref. 35805], Almeida & Biscoito 2019:140 [ref. 37302]). Current status: Valid as Dipturus springeri (Wallace 1967). Rajidae. Distribution: Southeastern Atlantic, western Indian Ocean: Namibia, South Africa and Madagascar to Oman and western India. Habitat: marine.

stabuliforis, Raia Garman [S.] 1913:341, Pls. 22 (fig. 2), 44 (4-6) [Memoirs of the Museum of Comparative Zoology v. 36; ref. 1545] New England, U.S.A. Syntypes: MCZ 1138-S (1), 1294-S (1). Type catalog: Hartel & Dingerkus 1997:xlvi [ref. 23119]. •Synonym of Dipturus laevis (Mitchill 1818) -- (Scott & Scott 1988:47 [ref. 25518] as stabuliformis). •Synonym of Raja chilensis Guichenot 1848 -- (Lloris & Rucabado 1991:44 [ref. 22530]). •In genus Dipturus and not Zearaja and possibly synonym of D. laevis (Mitchill) -- (Last & Gledhill 2007:50 [ref. 29177]). Current status: Synonym of Dipturus laevis (Mitchill 1818). Rajidae. Habitat: marine.

stenorhynchus, Raja Wallace [J. H.] 1967:23, Fig. 11 [Investigational Report. Oceanographic Research Institute, Durban No. 17; ref. 7929] East of False Bay, 22°25'S, 35°54'E, South Africa, western Indian Ocean, Anton Bruun station 398B, depth 405 fathoms. Holotype (unique): ORI B186. •Valid as Raja stenorhynchus Wallace 1967 -- (Hulley 1986:126 [ref. 5672], Séret 1989:127 [ref. 12827]). •Valid as Dipturus stenorhynchus (Wallace 1967) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:491 [ref. 25589], Compagno & Ebert 2007:139 [ref. 30471], Last & Alava 2013:214 [ref. 33041], Weigmann 2016:110 [ref. 34211], Last et al. 2016:22 [ref. 34941], Last et al. 2016:273 [ref. 35000]). Current status: Valid as Dipturus stenorhynchus (Wallace 1967). Rajidae. Distribution: Western Indian Ocean: South Africa. Habitat: marine.

teevani, Raja Bigelow [H. B.] & Schroeder [W. C.] 1951:388, Fig. 3 [Journal of the Washington Academy of Sciences v. 41 (no. 12); ref. 12883] Off Pensacola, Florida, U.S.A., 29°11'N, 86°52'30"W, Oregon station 279, depth 305 fathoms. Holotype: USNM 153557. Paratypes: MCZ 37189 (1). •Valid as Raja teevani Bigelow & Schroeder 1951 -- (Uyeno & Miyake in Uyeno et al. 1983:77 [ref. 14275], Séret 1989:126 [ref. 12827], Cervigón 1992:206 [ref. 23827], Jacob & McEachran 1994:444 [ref. 22422], Castro-Aguirre & Espinosa Pérez 1996:40 [ref. 22793]). •Valid as Dipturus teevani (Bigelow & Schroeder 1951) -- (McEachran & Dunn 1998:285 [ref. 23312], McEachran & Fechhelm 1998:158 [ref. 23897], Compagno 1999:491 [ref. 25589], Gomes & Paragó 2001:1 [ref. 25669], Gomes & Gadig in Menezes et al. 2003:29 [ref. 27192], McEachran & Carvalho 2003:553 [ref. 26985], Jeong & Nakabo 2008:189 [ref. 29612], Díaz de Astarloa et al. 2008:35 [ref. 29969], Vargas-Caro 2015:323 [ref. 33982], Del Moral-Flores et al. 2016:115 [ref. 34398], Weigmann 2016:110 [ref. 34211], Last et al. 2016:22 [ref. 34941], Last et al. 2016:274 [ref. 35000], Ehemann et al. 2018:25 [ref. 36194], Ehemann et al. 2019:[6] [ref. 36861], Jones et al. 2020:290 [ref. 37610]). Current status: Valid as Dipturus teevani (Bigelow & Schroeder 1951). Rajidae. Distribution: Western Atlantic: Florida (U.S.A.) south to Colombia and Bahia (Brazil), including Gulf of Mexico and Caribbean Sea. Habitat: marine.

tengu, Raja Jordan [D. S.] & Fowler [H. W.] 1903:654, Fig. 8 [Proceedings of the United States National Museum v. 26 (no. 1324); ref. 2460] Matsushima Bay, Sendai, Japan. Holotype: CAS-SU 12912 (specimen dissected) [not 7138]. Paratypes: USNM. Type catalog: Böhlke 1953:10 [ref. 12291]. •Valid as Raja tengu Jordan & Fowler 1903 -- (Nakaya in Masuda et al. 1984:13 [ref. 6441], Ishihara 1987:278 [ref. 6264], Séret 1989:128 [ref. 12827], Sokolovskaya et al. 1998:7 [ref. 24670], Compagno et al. 1999:1407 [ref. 24636], Last & Compagno 1999:1454 [ref. 24639], Dolganov 1999:429 [ref. 24873], Compagno 1999:491 [ref. 25589], Compagno in Randall & Lim 2000:582 [ref. 25122], Shinohara et al. 2001:291 [ref. 25995], Compagno et al. 2005:66 [ref. 29145]). •Valid as Dipturus tengu (Jordan & Fowler 1903) -- (McEachran & Dunn 1998:285 [ref. 23312], Jeong 1999:202 [ref. 25171], Nakabo 2000:173 [ref. 25086], Nakabo 2002:173 [ref. 26001], Choi et al. 2003:66, 482 [ref. 26218], Shinohara et al. 2005:398 [ref. 28370], Jeong & Nakabo 2008:189 [ref. 29612], Yamashita et al. 2012:130 [ref. 32408], Last & Alava 2013:214 [ref. 33041], Hsu et al. 2013:253 [ref. 33043], Ebert et al. 2013:353 [ref. 33045], Parin et al. 2014:36 [ref. 33547], Dyldin 2015:70 [ref. 34524], Motomura et al. 2015:12 [ref. 34709], Weigmann 2016:110 [ref. 34211], Last et al. 2016:22 [ref. 34941], Last et al. 2016:275 [ref. 35000], Dyldin & Orlov 2018:166 [ref. 35920], Sonoyama et al. 2020:12 [ref. 37637]). Current status: Valid as Dipturus tengu (Jordan & Fowler 1903). Rajidae. Distribution: Northwestern Pacific: Philippines and Taiwan north to Japan. Habitat: marine.

trachyderma, Raja (Dipturus) Krefft [G.] & Stehmann [M. F. W.] 1975:86, Figs. 7-8 [Archiv für Fischereiwissenschaft v. 26 (no. 3); ref. 7435] Off southern Argentina, southwestern Atlantic, 49°00'S, 60°52'W, depth 200-195 meters. Holotype (unique): ZMH 25402 [ex ISH 130-1971]. Type catalog: Krefft 1978:7 [ref. 9884], Thiel et al. 2009:26 [ref. 30918]. Specific name is an undeclinable noun. •Valid as Raja trachyderma Krefft & Stehmann 1975 -- (Pequeño 1989:17 [ref. 14125], Séret 1989:127 [ref. 12827], Lloris & Rucabado 1991:47 [ref. 22530], López et al. 1996:13 [ref. 23681]). •Valid as Dipturus trachyderma (Krefft & Stehmann 1975) -- (McEachran & Dunn 1998:285 [ref. 23312], Compagno 1999:491 [ref. 25589], Menni & Stehmann 2000:80 [ref. 24909], Gomes & Paragó 2001:1 [ref. 25669], Soto & Mincarone 2001:6 [ref. 26630], López et al. 2002:62 [ref. 26808] as trachydermus, Jeong & Nakabo 2008:189 [ref. 29612], Mabragaña et al. 2011:7 [ref. 31800], Vargas-Caro et al. 2015:328 [ref. 33982], Weigmann 2016:110 [ref. 34211] as trachydermus, Last et al. 2016:22 [ref. 34941] as trachydermus, Last et al. 2016:276 [ref. 35000] as trachydermus, Nión et al. 2016:22 [ref. 35565]). Current status: Valid as Dipturus trachyderma (Krefft & Stehmann 1975). Rajidae. Distribution: Southeastern Pacific and southwestern Atlantic: Uruguay to Chile, possibly southern Brazil. Habitat: marine.

vomer, Raja Fries [B. F.] 1838:161 [36] [Kongliga Vetenskaps-Academiens Handlingar, Stockholm 1838; ref. 18097] Sweden, Skagerrak, North Sea. Holotype (?): NRM 209e (stuffed). On p. 36 of separate. •Synonym of Raja oxyrinchus Linnaeus 1758 -- (Stehmann 1973:63 [ref. 7168]). •Synonym of Dipturus oxyrinchus (Linnaeus 1758) -- (Dyldin 2015:70 [ref. 34524]). Current status: Synonym of Dipturus oxyrinchus (Linnaeus 1758). Rajidae. Habitat: marine.

vulgaris, Batis Couch [J.] 1862:87, Pl. 18 [The history of the fishes of the British islands v. 1; ref. 989] British Isles. No types known. Apparently an unneeded substitute for Raja batis. •Apparently a synonym of Raja batis of earlier authors. •Synonym of Dipturus batis (Linnaeus 1758) -- (Dyldin 2015:70 [ref. 34524]). Current status: Synonym of Dipturus batis (Linnaeus 1758). Rajidae. Habitat: marine.

vulgaris, Raia Stephan [C. F.] 1779:23 [De rajis, schediasma primum; ref. 18818] •Synonym of Dipturus batis (Linnaeus 1758) -- (Kottelat 2010:65 [ref. 30995], Dyldin 2015:70 [ref. 34524]). Current status: Synonym of Dipturus batis (Linnaeus 1758). Rajidae. Habitat: marine.

wengi, Dipturus Séret [B.] & Last [P. R.] 2008:99, Figs. 1-6 [CSIRO Marine and Atmospheric Research Paper No. 021; ref. 29649] East of Sydney, New South Wales, Australia, 33°43'S, 151°54'E, depth 486-509 meters. Holotype: CSIRO H 891-01. Paratypes: (15 specimens) at AMS and CSIRO. Plus non-type specimens. •Valid as Dipturus wengi Séret & Last 2008 -- (Last & White 2008:109 [ref. 30618], Weigmann 2016:110 [ref. 34211], Last et al. 2016:22 [ref. 34941], Last et al. 2016:277 [ref. 35000]). Current status: Valid as Dipturus wengi Séret & Last 2008. Rajidae. Distribution: Eastern Australia: Queensland, New South Wales, and Tasmania. Habitat: marine.

wuhanlingi, Dipturus Jeong [C.-H.] & Nakabo [T.] 2008:184, Figs. 1-7 [Ichthyological Research v. 55 (no. 2); ref. 29612] Open waters of the South China Sea. Holotype: SFC D 01113. Paratypes: SFC S 07672 (1). •Valid as Dipturus wuhanlingi Jeong & Nakabo 2008 -- (Last & Alava 2013:214 [ref. 33041], Ebert et al. 2013:354 [ref. 33045], Weigmann 2016:110 [ref. 34211], Last et al. 2016:22 [ref. 34941], Last et al. 2016:278 [ref. 35000]). Current status: Valid as Dipturus wuhanlingi Jeong & Nakabo 2008. Rajidae. Distribution: South China Sea, East China Sea. Habitat: marine.