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Eschmeyer's Catalog of Fishes
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Species in the genus Leucoraja: [ 24 ] records


caribbaea, Raja garmani McEachran [J. D.] 1977:434 [Bulletin of Marine Science v. 27 (no. 3); ref. 7268] Off Quintana Roo, 21°13'N, 86°27'W, Mexico, depth 457 meters. Holotype: UF 231707 [ex UMML 31707]. Paratypes: MCZ 42424 (1); USNM 222386 (1). •Synonym of Raja garmani Whitley 1939, but a valid subspecies as described -- (Schmitter-Soto et al. 2000:147 [ref. 27754], Weigmann 2016:112 [ref. 34211]). •Valid as Leucoraja caribbaea (McEachran 1977) -- (Nelson et al. 2004:55 [ref. 27807], Page et al. 2013:55 [ref. 32708], Del Moral-Flores et al. 2016:116 [ref. 34398]). •Synonym of Leucoraja yucatanensis (Bigelow & Schroeder 1950) -- (personal communication Ross Robertson email 6/26/2016). •Synonym of Leucoraja garmani (Whitley 1939), but a valid subspecies caribbaea (McEachran 1977) -- (Last et al. 2016:18 [ref. 34941]). Current status: Synonym of Leucoraja garmani (Whitley 1939). Rajidae. Distribution: Western Atlantic [if valid]. Habitat: marine.

chagrinea, Raja Shaw [G.] 1804:281 [General zoology or systematic natural history; ref. 4015] European Seas. Based on Pennant, 1776:87. •Synonym of Raja fullonica Linnaeus 1758 -- (Stehmann 1973:64 [ref. 7168]). •Synonym of Leucoraja fullonica (Linnaeus 1758) -- (Dyldin 2015:71 [ref. 34524]). Current status: Synonym of Leucoraja fullonica (Linnaeus 1758). Rajidae. Habitat: marine.

circularis, Raja Couch [J.] 1838:53 [A Cornish fauna, being a compendium of the natural history of the county; ref. 18049] Cornwall, England, northeastern Atlantic. No types known. Based on Sandy ray of Couch, Mag. N. Hist. Charlesworth, v. 2, 1838:71. •Valid as Raja circularis Couch 1838 -- (Stehmann 1973:64 [ref. 7168], Stehmann & Bürkel in Whitehead et al. 1984:178 [ref. 13675], Capapé 1987:22 [ref. 39326], Stehmann 1990:34 [ref. 19316], Stehmann 1995:108 [ref. 22055]). •Valid as Leucoraja circularis (Couch 1838) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], Bilecenoğlu et al. 2002:22 [ref. 26753], Fricke et al. 2007:18 [ref. 29533], Last et al. 2008:145 [ref. 29653], George 2009:56 [ref. 30539], Lipej & Dulčić 2010:13 [ref. 36649], Weigmann 2016:112 [ref. 34211], McEachran & Séret 2016:1397 [ref. 34551], Last et al. 2016:18 [ref. 34941], Last et al. 2016:280 [ref. 35000], Biscoito et al. 2018:478 [ref. 37562], Almeida & Biscoito 2019:156 [ref. 37302], Carneiro et al. 2019:44 [ref. 37250], Kovačić et al. 2020:7 [ref. 37519], Serena et al. 2020:502 [ref. 37963], Giovos et al. 2021:6 [ref. 38895], Kovačić et al. 2021:15 [ref. 39014], Bañón & Mano 2022:13 [ref. 40121], Barone et al. 2022:49 [ref. 39527], Balàka et al. 2023:6 [ref. 40410], Charalampous & Chatzispyrou in Digenis et al. 2024:104 [ref. 40958], Deval in Digenis et al. 2024:102 [ref. 40958]). Current status: Valid as Leucoraja circularis (Couch 1838). Rajidae. Distribution: Mediterranean Sea; eastern Atlantic: Iceland and central Norway south to northern Morocco, including Madeira. Habitat: marine.

compagnoi, Raja (Leucoraja) Stehmann [M. F. W.] 1995:43, Fig. 10 [Archive of Fishery and Marine Research v. 43 (no. 1); ref. 22055] Off South Africa, 30°44'S, 15°18'E, depth 550 meters. Holotype (unique): ZIN 48406. •Valid as Leucoraja compagnoi (Stehmann 1995) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], Compagno & Ebert 2007:133 [ref. 30471], Last et al. 2008:145 [ref. 29653], Weigmann 2016:112 [ref. 34211] see remarks, Last et al. 2016:18 [ref. 34941], Last et al. 2016:281 [ref. 35000], Almeida & Biscoito 2019:155 [ref. 37302], Ebert & Leslie 2019:233 [ref. 37048], Ebert et al. 2021:81 [ref. 38234], Aschliman & Ebert 2022:581 [ref. 39668]). Current status: Valid as Leucoraja compagnoi (Stehmann 1995). Rajidae. Distribution: Southeastern Atlantic, southwestern Indian Ocean: Western Cape, Eastern Cape and KwaZulu-Natal (South Africa) endemic. Habitat: marine.

elaineae, Leucoraja Ebert [D. A.] & Leslie [R. W.] 2019:226, Figs. 1-5 [Zootaxa 4691 (no. 3); ref. 37048] Off Malindi, Kenya, 03°25'S, 40°23'E, depth 484 meters. Holotype (unique): SAIAB 13742. •Valid as Leucoraja elaineae Ebert & Leslie 2019. Current status: Valid as Leucoraja elaineae Ebert & Leslie 2019. Rajidae. Distribution: Off Malindi, Kenya, western Indian Ocean. Habitat: marine.

erinaceus, Raja Mitchill [S. L.] 1825:290 [American Journal of Science and Arts v. 9; ref. 17778] Barnegat and southeast of Sandy Hook, New Jersey, U.S.A. No types known. "Erinaceus" is a noun in apposition, meaning hedgehog, and does not change its ending (Chris Scharpf, pers. comm., 2/22). •Valid as Raja erinacea Mitchill 1825 -- (McEachran & Martin 1977:121 [ref. 21983], Robins & Ray 1986:38 [ref. 23100], Scott & Scott 1988:41 [ref. 25518], McAllister 1990:37 [ref. 14674], Murdy et al. 1997:41 [ref. 23144]). •Valid as Leucoraja erinacea (Mitchill 1825) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], McEachran in Collette & Klein-MacPhee 2002:67 [ref. 26158], Nelson et al. 2004:55 [ref. 27807], Last et al. 2008:145 [ref. 29653], Page et al. 2013:55 [ref. 32708], Weigmann 2016:112 [ref. 34211], Last et al. 2016:18 [ref. 34941], Last et al. 2016:282 [ref. 35000]). •Valid as Leucoraja erinaceus (Mitchill 1825) -- (Simian et al. 2022:374 [ref. 40776], Page et al. 2023:44 [ref. 40505]). Current status: Valid as Leucoraja erinaceus (Mitchill 1825). Rajidae. Distribution: Western Atlantic: Newfoundland (Canada) south to northern North Carolina (U.S.A.). IUCN (2020): Least Concern. Habitat: marine.

falsavela, Raja Bonaparte [C. L.] 1839:fasc. 26, punt. 136, Pl. 148 (fig. 1) [Iconografia della fauna italica; ref. 4895] Italy, western Mediterranean Sea. Holotype: ANSP 406 (missing). Type catalog: Böhlke 1984:149 [ref. 13621]. Puntata written as 135 on first page, corrected to 136 in index. •Synonym of Raja circularis Couch 1838 -- (Stehmann 1973:64 [ref. 7168]). •Synonym of Leucoraja circularis (Couch 1838). Current status: Synonym of Leucoraja circularis (Couch 1838). Rajidae. Habitat: marine.

fullonica, Raja Linnaeus [C.] 1758:231 [Systema Naturae, Ed. X v. 1; ref. 2787] European seas [original: "in M. Europeo"]; localities include England, Northeastern Atlantic; southern France, Mediterranean Sea. No types known. Lectotype or neotype fixation needed. Originally based on multiple species and at least 14 pre-Linnaean sources including Artedi 1738:72 [ref. 30578] and Artedi 1738:101 [ref. 30349] as Raja toto dorso aculeato; Rondelet 1554:356 [ref. 30354] as Raia fullonica, Gesner 1558:938 [ref. 30350] as Raia fvllonica, Jonston 1649:Pl. 11 (fig. 4) [ref. 30579] as Raia fullonica, Charleton 1668:130 [ref. 30580] as Raia fullonica, Willughby 1686:78 [ref. 30164] and Ray 1713:26 [ref. 30231] as Raia aspera nostras, an fullonica. Spelled fullonia by Bloch & Schneider 1801:367 [ref. 471]. Name spelled Raya fallonica by Berthelot 1840:115 [ref. 30346]. •Valid as Raja fullonica Linnaeus 1758 -- (Stehmann 1973:64 [ref. 7168], Stehmann & Bürkel in Whitehead et al. 1984:179 [ref. 13675], Stehmann 1990:35 [ref. 19316], Stehmann 1995:108 [ref. 22055], Santos et al. 1997:14 [ref. 23531], Arruda 1997:22 [ref. 24952]). •Valid as Leucoraja fullonica (Linnaeus 1758) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], Bilecenoğlu et al. 2002:22 [ref. 26753], Fricke et al. 2007:18 [ref. 29533], Last et al. 2008:145 [ref. 29653], George 2009:57 [ref. 30539], Lipej & Dulčić 2010:13 [ref. 36649], Parin et al. 2014:36 [ref. 33547], Dyldin 2015:70 [ref. 34524], Weigmann 2016:112 [ref. 34211], Last et al. 2016:18 [ref. 34941], Last et al. 2016:283 [ref. 35000], Biscoito et al. 2018:477 [ref. 37562], Almeida & Biscoito 2019:142 [ref. 37302], Carneiro et al. 2019:44 [ref. 37250], Saad & Alkusairy 2019:304 [ref. 36710], Kovačić et al. 2020:7 [ref. 37519], Serena et al. 2020:502 [ref. 37963], Giovos et al. 2021:6 [ref. 38895], Kovačić et al. 2021:15 [ref. 39014], Bañón & Mano 2022:14 [ref. 40121], Barone et al. 2022:49 [ref. 39527], Balàka et al. 2023:6 [ref. 40410], Shakman et al. 2023:464 [ref. 40810]). Current status: Valid as Leucoraja fullonica (Linnaeus 1758). Rajidae. Distribution: Arctic Ocean; White Sea; western Baltic Sea; North Sea; Mediterranean Sea; eastern Atlantic: Murmansk (Russia) south to northern Morocco, including Iceland, Faroe Islands, and Madeira. Habitat: marine.

fullonica, Raja Stephan [C. F.] 1779:23 [De rajis, schediasma primum; ref. 18818] •Synonym of Leucoraja fullonica (Linnaeus 1758) -- (Kottelat 2010:65 [ref. 30995], Dyldin 2015:71 [ref. 34524]). Current status: Synonym of Leucoraja fullonica (Linnaeus 1758). Rajidae. Habitat: marine.

gallica, Raja Walbaum [J. J.] 1792:533 [Petri Artedi sueci genera piscium Part 3; ref. 4572] British Isles. No types known. •Synonym of Leucoraja fullonica (Linnaeus 1758) -- (Dyldin 2015:71 [ref. 34524]). Current status: Synonym of Leucoraja fullonica (Linnaeus 1758). Rajidae. Habitat: marine.

garmani, Raja Whitley [G. P.] 1939:248 [Australian Zoologist v. 9 (pt 3); ref. 4695] Off South Carolina, 32°24'N, 78°44'W, depth 142 fathoms. Lectotype: MCZ 915-S. Paralectotypes: MCZ 585-S (1), MCZ 51759 (1); USNM 43727 (1, or now 0). Type catalog: Hartel & Dingerkus 1997:xlvii [ref. 23119]. New name for Raja ackleyi var. ornata Garman 1881. Lectotype established by McEachran for garmani which established the lectotype for ornata as well, see R. ornata. •Valid as Raja garmani Whitley 1939 -- (McEachran 1977:425 [ref. 7268], Robins & Ray 1986:39 [ref. 23100], Séret 1989:126 [ref. 12827], Boschung 1992:26 [ref. 23239], Castro-Aguirre & Espinosa Pérez 1996:38 [ref. 22793], Schmitter-Soto et al. 2000:146 [ref. 27754]). •Valid as Leucoraja garmani (Whitley 1939) -- (McEachran & Dunn 1998:286 [ref. 23312], McEachran & Fechhelm 1998:162 [ref. 23897], Compagno 1999:491 [ref. 25589], McEachran & Carvalho 2003:540 [ref. 26985], Nelson et al. 2004:55 [ref. 27807], Last et al. 2008:145 [ref. 29653], Page et al. 2013:55 [ref. 32708], Del Moral-Flores et al. 2016:116 [ref. 34398], Weigmann 2016:112 [ref. 34211], Last et al. 2016:18 [ref. 34941], Last et al. 2016:284 [ref. 35000], Ehemann et al. 2018:25 [ref. 36194], Ehemann et al. 2019:[6] [ref. 36861], Blanco-Parra & Niño-Torres 2022:Correction, p. 5 [ref. 38986], Page et al. 2023:44 [ref. 40505]). Current status: Valid as Leucoraja garmani (Whitley 1939). Rajidae. Distribution: Western Atlantic: Massachusetts south to Florida (U.S.A.); western and southern Caribbean Sea. Habitat: marine.

intermedia, Raja falsavela var. Doderlein [P.] 1884:202 [Manuale Ittiologico del Mediterraneo ossia sinossi metodica delle varie specie di pesci riscontrate sin qui nel Mediterraneo ossia senossi metodica delle di Sicilia. 5 parts.; ref. 17416] Palermo, Sicily, Italy, Mediterranean Sea. •Apparently a synonym of Raja circularis Couch 1838. •Synonym of Leucoraja circularis (Couch 1838). Current status: Synonym of Leucoraja circularis (Couch 1838). Rajidae. Habitat: marine.

lentiginosa, Raja Bigelow [H. B.] & Schroeder [W. C.] 1951:383, Fig. 1 [Journal of the Washington Academy of Sciences v. 41 (no. 12); ref. 12883] Campeche Bank, Gulf of Mexico, 22°32'N, 88°47'W, depth 29 fathoms. Holotype: USNM 153552. Paratypes or additional material: (7) MCZ 37188 (1); USNM 153553-55 (1, 2, 1). •Valid as Raja lentiginosa Bigelow & Schroeder 1951 -- (McEachran 1977:430 [ref. 7268], Robins & Ray 1986:39 [ref. 23100]). •Synonym of Raja garmani Whitley 1939 -- (Castro-Aguirre & Espinosa Pérez 1996:38 [ref. 22793], Schmitter-Soto et al. 2000:147 [ref. 27754]). •Valid as Leucoraja lentiginosa (Bigelow & Schroeder 1951) -- (McEachran & Dunn 1998:286 [ref. 23312], McEachran & Fechhelm 1998:163 [ref. 23897], Compagno 1999:491 [ref. 25589], McEachran & Carvalho 2003:541 [ref. 26985], Nelson et al. 2004:55 [ref. 27807], Last et al. 2008:145 [ref. 29653], Page et al. 2013:55 [ref. 32708], Del Moral-Flores et al. 2016:116 [ref. 34398], Weigmann 2016:113 [ref. 34211], Last et al. 2016:18 [ref. 34941], Last et al. 2016:285 [ref. 35000], Ehemann et al. 2018:25 [ref. 36194], Jones et al. 2020:290 [ref. 37610], Page et al. 2023:44 [ref. 40505]). Current status: Valid as Leucoraja lentiginosa (Bigelow & Schroeder 1951). Rajidae. Distribution: Western Atlantic: Gulf of Mexico. Habitat: marine.

leucosticta, Raja (Leucoraja) Stehmann [M. F. W.] 1971:3, Figs. 3-9, 12 [Archiv für Fischereiwissenschaft v. 22 (no. 1); ref. 7474] West Africa, 4°58'S, 11°20'30"E, eastern Atlantic, depth 200-300 meters. Holotype: IRSNB 522 [ex IRSNB 8545]. Paratypes: IRSNB 523 [ex IRSNB 8495] (1), IRSNB 524 [ex IRSNB 8535] (1); ZMH 25255 [ ex ISH 273-1963] (1). Type catalog: Krefft 1978:6 [ref. 9884], Walschaerts 1987:5 [ref. 20755], Thiel et al. 2009:25 [ref. 30918]. •Valid as Raja leucosticta Stehmann 1971 -- (Stehmann 1990:35 [ref. 19316], Stehmann 1995:109 [ref. 22055]). •Valid as Leucoraja leucosticta (Stehmann 1971) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], Last et al. 2008:145 [ref. 29653], Weigmann 2016:113 [ref. 34211], McEachran & Séret 2016:1390 [ref. 34551], Last et al. 2016:18 [ref. 34941], Last et al. 2016:286 [ref. 35000], Almeida & Biscoito 2019:155 [ref. 37302], Reiner 2019:44 [ref. 38139], Fermon et al. 2022:104 [ref. 39448]). Current status: Valid as Leucoraja leucosticta (Stehmann 1971). Rajidae. Distribution: Eastern Atlantic: Mauritania south to Gabon, including São Tomé and Principe. Habitat: marine.

maggiore, Raja falsavela var. Doderlein [P.] 1884:202 [Manuale Ittiologico del Mediterraneo ossia sinossi metodica delle varie specie di pesci riscontrate sin qui nel Mediterraneo ossia senossi metodica delle di Sicilia. 5 parts.; ref. 17416] Palermo, Sicily, Italy, Mediterranean Sea. •Apparently a synonym of Raja circularis Couch 1838. •Synonym of Leucoraja circularis (Couch 1838). Current status: Synonym of Leucoraja circularis (Couch 1838). Rajidae. Habitat: marine.

melitensis, Raia Clark [R. S.] 1926:43, Pl. 21 (fig. b) [Fishery Board for Scotland, Scientific Investigations 1926 No. 1; ref. 15582] Malta, Mediterranean Sea. Syntypes: (2) BMNH 1926.12.20.2 (1). •Valid as Raja melitensis Clark 1926 -- (Stehmann 1973:67 [ref. 7168], Capapé 1980:114 [ref. 9546], Stehmann & Bürkel in Whitehead et al. 1984:180 [ref. 13675], Capapé 1987:22 [ref. 39326]). •Valid as Leucoraja melitensis (Clark 1926) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], Last et al. 2008:145 [ref. 29653], Weigmann 2016:113 [ref. 34211], Last et al. 2016:18 [ref. 34941], Last et al. 2016:287 [ref. 35000], Ben Amor et al. 2018:10 [ref. 36367], Almeida & Biscoito 2019:153 [ref. 37302], Serena et al. 2020:502 [ref. 37963], Kovačić et al. 2021:15 [ref. 39014], Barone et al. 2022:49 [ref. 39527], Shakman et al. 2023:464 [ref. 40810]). Current status: Valid as Leucoraja melitensis (Clark 1926). Rajidae. Distribution: Western Mediterranean Sea: Algeria,Tunisia, Malta. Habitat: marine.

naevus, Raja Müller [J.] & Henle [F. G. J.] 1841:138, 194 [Systematische Beschreibung der Plagiostomen; ref. 3069] Abbeville, France; Toulon, France, western Mediterranean Sea; Atlantic coasts of Europe; Mediterranean Sea. Syntypes: MNHN 0000-1306 (1) Abbeville, 0000-1332 (1) Toulon, ?RMNH 4237 (1). Type catalog: Bertin 1939:83 [ref. 20739], Séret & McEachran 1987:15 [ref. 9312]. •Valid as Raja naevus Müller & Henle 1841 -- (Stehmann 1973:65 [ref. 7168], Stehmann & Bürkel in Whitehead et al. 1984:181 [ref. 13675], Capapé 1987:22 [ref. 39326], Stehmann 1990:36 [ref. 19316], Stehmann 1995:39 [ref. 22055]). •Valid as Leucoraja naevus (Müller & Henle 1841) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], Bilecenoğlu et al. 2002:22 [ref. 26753], Fricke et al. 2007:18 [ref. 29533], Fricke et al. 2007:18 [ref. 29533], Last et al. 2008:145 [ref. 29653], George 2009:57 [ref. 30539], Weigmann 2016:113 [ref. 34211], McEachran & Séret 2016:1398 [ref. 34551], Gordon et al. 2016:262 [ref. 34661], Last et al. 2016:19 [ref. 34941], Last et al. 2016:288 [ref. 35000], Almeida & Biscoito 2019:153 [ref. 37302], Carneiro et al. 2019:45 [ref. 37250], Serena et al. 2020:502 [ref. 37963], Arroyo Martínez 2021:88 [ref. 38962], Giovos et al. 2021:6 [ref. 38895], Kovačić et al. 2021:15 [ref. 39014], Bañón & Mano 2022:14 [ref. 40121], Barone et al. 2022:49 [ref. 39527], Capapé et al. 2023:5 [ref. 40801], Shakman et al. 2023:464 [ref. 40810]). Current status: Valid as Leucoraja naevus (Müller & Henle 1841). Rajidae. Distribution: North Sea; Mediterranean Sea; eastern Atlantic: British Isles south to Senegal. Habitat: marine.

ocellata, Raja Mitchill [S. L.] 1815:477 [Transactions of the Literary and Philosophical Society of New-York v. 1 (art. 5) (for 1814); ref. 13292] New York, U.S.A. No types known. •Valid as Raja ocellata Mitchill 1815 -- (McEachran & Martin 1977:121 [ref. 21983], Robins & Ray 1986:38 [ref. 23100], Scott & Scott 1988:49 [ref. 25518], McAllister 1990:37 [ref. 14674], Murdy et al. 1997:40 [ref. 23144]). •Valid as Leucoraja ocellata (Mitchill 1815) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], McEachran in Collette & Klein-MacPhee 2002:70 [ref. 26158], Nelson et al. 2004:55 [ref. 27807], Last et al. 2008:145 [ref. 29653], Page et al. 2013:55 [ref. 32708], Weigmann 2016:113 [ref. 34211], Last et al. 2016:19 [ref. 34941], Last et al. 2016:289 [ref. 35000], Simian et al. 2022:374 [ref. 40776], Page et al. 2023:44 [ref. 40505]). Current status: Valid as Leucoraja ocellata (Mitchill 1815). Rajidae. Distribution: Western Atlantic: Newfoundland (Canada) to North Carolina (U.S.A.). Habitat: marine.

ornata, Raja ackleyi var. Garman [S.] 1881:235 [Bulletin of the Museum of Comparative Zoology v. 8 (no. 11); ref. 14294] Off South Carolina, 32°24'N, 78°44'W, depth 142 fathoms. Lectotype: MCZ 915-S. Paralectotypes: MCZ 585-S (1), MCZ 51759 (1); USNM 43727 (1, or now 0). Appeared after Raja ackleyi sp. nov. as Raja ornata var. nov. R. ackleyi which authors have misinterpreted as describing a new species ornata, but Garman states (p. 236), "...that these young skates represent a variety of R. Ackleyi." Objectively invalid; Raja ackleyi ornata is preoccupied by Raia ornata Agassiz 1843 in fossil fishes, replaced by Raja garmani Whitley 1939. Lectotype established by McEachran for garmani, which also establishes a lectotype for ornata. •Synonym of Raja garmani Whitley 1939 -- (Castro-Aguirre & Espinosa Pérez 1996:38 [ref. 22793]). •Synonym of Leucoraja garmani (Whitley 1939). 1°Homonym. Current status: Synonym of Leucoraja garmani (Whitley 1939). Rajidae. Habitat: marine.

pristispina, Leucoraja Last [P. R.], Stehmann [M. F. W.] & Séret [B.] 2008:145, Figs. 1-6 [CSIRO Marine and Atmospheric Research Paper No. 021; ref. 29653] Southwest of Imperieuse Reef (Rowley Shoals), Western Australia, 18°07'S, 118°09'E, depth 400-404 meters. Holotype: CSIRO CA 3905. Paratypes: 27 specimens at CSIROand WAM. Plus non-type specimens. •Valid as Leucoraja pristispina Last, Stehmann & Séret 2008 -- (Weigmann 2016:113 [ref. 34211], Last et al. 2016:19 [ref. 34941], Last et al. 2016:290 [ref. 35000], Ebert & Leslie 2019:233 [ref. 37048]). Current status: Valid as Leucoraja pristispina Last, Stehmann & Séret 2008. Rajidae. Distribution: Western Australia and eastern Indonesia. Habitat: marine.

rondeleti, Raja Bougis [P.] 1959:108, Figs. 1-4 [Vie et Milieu v. 10 (fasc. 1); ref. 11978] Northwestern Mediterranean Sea. Syntypes: MNHN 1898-1249 (1) Sète; uncat. (1) Banyuls; uncat. (1) Sète; uncat. (1, dry) Livourne; MZUF 6132 [ex MZUF 1075] (1). Type catalog: Séret & McEachran 1987:15 [ref. 9312], Vanni 1993:105 [ref. 21837]. •Valid as Raja rondeleti Bougis 1959 -- (Stehmann 1973:68 [ref. 7168], Stehmann & Bürkel in Whitehead et al. 1984:195 [ref. 13675], Compagno 1999:492 [ref. 25589], Weigmann 2016:120 [ref. 34211] see remarks). •Synonym of Raja fullonica (Linnaeus 1758) -- (Psomadakis et al. 2012:15 [ref. 33457]). •Synonym of Leucoraja fullonica (Linnaeus 1758) -- (Dyldin 2015:71 [ref. 34524]). •Probably synonymous with Raja clavata (Stehmann & Bürkel 1984) or Leucoraja fullonica (Serena 2005) -- (Last et al. 2016:17 [ref. 34941]). Current status: Synonym of Leucoraja fullonica (Linnaeus 1758). Rajidae. Habitat: marine.

virginica, Raja garmani McEachran [J. D.] 1977:434 [Bulletin of Marine Science v. 27 (no. 3); ref. 7268] Off North Carolina, U.S.A., 36°46'N, 74°41'W, U.S.A., depth 104-117 meters. Holotype: MCZ 34762. Paratypes: MCZ 36315 (1), VIMS 1370 (1). •Valid as Leucoraja garmani virginica (McEachran 1977) -- (McEachran in Collette & Klein-MacPhee 2002:69 [ref. 26158], Moore et al. 2003:171 [ref. 27076], Weigmann 2016:112 [ref. 34211]). •Valid as Leucoraja virginica (McEachran 1977) -- (McEachran & Fechhelm 1998:162 [ref. 23897], Nelson et al. 2004:55 [ref. 27807], Page et al. 2013:55 [ref. 32708]). •Synonym of Leucoraja garmani (Whitley 1939), but a valid subspecies virginica (McEachran 1977) -- (Last et al. 2016:18 [ref. 34941]). Current status: Synonym of Leucoraja garmani (Whitley 1939). Rajidae. Distribution: Western Atlantic [if valid]. Habitat: marine.

wallacei, Raja Hulley [P. A.] 1970:210, Pl. 12 (fig. b); Fig. 19 (a-b) [Annals of the South African Museum v. 55 (pt 4); ref. 7750] Off Cape Town, 34°10'S, 17°45'E, South Africa, southeastern Atlantic, depth 292 meters. Holotype: ORI B.155. Paratypes: ORI B.126 (1). •Valid as Raja wallacei Hulley 1970 -- (Hulley 1986:127 [ref. 5672], Compagno et al. 1991:96 [ref. 21083], Bianchi & Carpenter in Bianchi et al. 1993:96 [ref. 25472], Walmsley-Hart et al. 1999:165 [ref. 24222]). •Valid as Leucoraja wallacei (Hulley 1970) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], Compagno & Ebert 2007:133 [ref. 30471], Last et al. 2008:145 [ref. 29653], Weigmann 2016:113 [ref. 34211] see remarks, Last et al. 2016:19 [ref. 34941], Last et al. 2016:291 [ref. 35000], Almeida & Biscoito 2019:154 [ref. 37302], Ebert & Leslie 2019:233 [ref. 37048], Ebert et al. 2021:82 [ref. 38234], Aschliman & Ebert 2022:582 [ref. 39668], Leeney 2024:73 [ref. 40985]). Current status: Valid as Leucoraja wallacei (Hulley 1970). Rajidae. Distribution: Southeastern Atlantic, southwestern Indian Ocean: Lüderitz (Namibia) south to Cape of Good Hope and Agulhas Bank (South Africa), then north to southern Mozambique. Habitat: marine.

yucatanensis, Breviraja Bigelow [H. B.] & Schroeder [W. C.] 1950:402, Pl. 7 [Bulletin of the Museum of Comparative Zoology v. 103 (no. 7); ref. 302] Off northeastern Yucatán, Mexico, 20°19'10"N, 87°03'30"W, Albatross station 2359, depth 231 fathoms. Holotype (unique): USNM 148273. •Valid as Raja yucatanensis (Bigelow & Schroeder 1950) -- (McEachran & Compagno 1982:415 [ref. 2952], Castro-Aguirre & Espinosa Pérez 1996:41 [ref. 22793], Schmitter-Soto et al. 2000:147 [ref. 27754]). •Valid as Leucoraja yucatanensis (Bigelow & Schroeder 1950) -- (McEachran & Dunn 1998:286 [ref. 23312], Compagno 1999:491 [ref. 25589], McEachran & Carvalho 2003:557 [ref. 26985], Last et al. 2008:145 [ref. 29653], Del Moral-Flores et al. 2016:117 [ref. 34398], Weigmann 2016:114 [ref. 34211], Last et al. 2016:19 [ref. 34941], Last et al. 2016:292 [ref. 35000], Ehemann et al. 2018:25 [ref. 36194], Blanco-Parra & Niño-Torres 2022:Correction, p. 5 [ref. 38986], Page et al. 2023:44 [ref. 40505]). Current status: Valid as Leucoraja yucatanensis (Bigelow & Schroeder 1950). Rajidae. Distribution: Western Atlantic: Yucatan (Mexico) south to Nicaragua (southeastern Gulf of Mexico, western Caribbean Sea). Habitat: marine.


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