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Eschmeyer's Catalog of Fishes
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Species in the genus Phoxinus: [ 46 ] records


abanticus, Phoxinus Turan [D.], Bayçelebi [E.], Özuluğ [M.], Gaygusuz [Ö.] & Aksu [İ.] 2023:4, Figs. 4, 7-10 [Journal of Fish Biology v. 102 (no. 5); ref. 40351] Outlet of Lake Abant, Bolu Province, Turkey, 40.664722N, 31.425000E. Holotype: FFR 2322. Paratypes: FFR, IUSHM. •Valid as Phoxinus abanticus Turan, Bayçelebi, Özuluğ, Gaygusuz & Aksu 2023 -- (Çiçek et al. 2023:416 [ref. 40625], Bayçelebi et al. 2024:107 [ref. 40795], Çiçek et al. 2024:267 [ref. 40789]). Current status: Valid as Phoxinus abanticus Turan, Bayçelebi, Özuluğ, Gaygusuz & Aksu 2023. Leuciscidae: Phoxininae. Distribution: Asia Minor: Lake Abant basin, Filyos River drainage (Turkey). Habitat: freshwater.

adagumicus, Phoxinus Artaev [O. A.], Turbanov [I. S.], Bolotovskiy [A. A.], Gandlin [A. A.] & Levin [B. A.] 2024:29, Figs. 3, 4 [Zoosystematics and Evolution v. 100 (no. 1); ref. 40926] Pryamaya Shchel River (Adagum River drainage) upstream Nizhnebakanskaya, Kuban River drainage, Krasnodar Krai, Russia, 44.8538°N, 37.8417°E. Holotype: IBIW FS_385. Paratypes: IBIW. Plus additional non-type material. •Valid as Phoxinus adagumicus Artaev, Turbanov, Bolotovskiy, Gandlin & Levin 2024. Current status: Valid as Phoxinus adagumicus Artaev, Turbanov, Bolotovskiy, Gandlin & Levin 2024. Leuciscidae: Phoxininae. Distribution: Southeast Europe: Adagum River drainage, Kuban River basin, northwestern Caucasus, Russia. Habitat: freshwater.

aphya, Cyprinus Linnaeus [C.] 1758:323 [Systema Naturae, Ed. X v. 1; ref. 2787] River Agger north of Lohmar, 50°50'N, 7°12'E, Nordrhein-Westfalen, Germany. Neotype: NRM 55108. Spelled aphia by Bonnaterre 1788:196 [ref. 4940] and authors. Neotype designated by Fricke 1999:25 [ref. 24101], but withdrawn in Fricke 2000:639 [ref. 24537]. Neotype validly designated by Kottelat 2007:155 [ref. 29201]; the neotype is the same as the neotype for Cyprinus phoxinus Linnaeus 1758 making them simultaneous objective synonyms. See also Kottelat 2007:155 [ref. 29201] for other nomenclatural issues. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Berg 1949:588 [ref. 20661], Bogutskaya & Naseka 1996:20 [ref. 22798], Kottelat 1997:73 [ref. 22952], Bogutskaya 1998:68 [ref. 23589], Fricke 1999:25 [ref. 24101], Sideleva 2001:51 [ref. 26181], Sideleva 2003:9 [ref. 27815], Kottelat 2007:155 [ref. 29201], Denys et al. 2020:214 [ref. 37846]). Current status: Synonym of Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae. Habitat: freshwater.

apollonicus, Phoxinus Bianco [P. G.] & De Bonis [S.] 2015:10, Figs. 13-15 [Researches on Wildlife Conservation v. 4; ref. 34040] Upper River Moraca tributary to Lake Skadar, Montenegro. Holotype: IZA 87103-1. Paratypes: IZA. First revisor action in an unavailable publication, not valid. •Synonym of Phoxinus karsticus Bianco & De Bonis 2015 -- (Palandačić et al. 2017:12 [ref. 35519]). Current status: Synonym of Phoxinus karsticus Bianco & De Bonis 2015. Leuciscidae: Phoxininae. Habitat: freshwater.

balchaschana, Phoxinus laevis var. Kessler [K. F.] 1879:283 [Bulletin de l'Académie Impériale des Sciences de St. Pétersbourg v. 25; ref. 2598] Ajagus [Ayaguz] River near Sergiopol [Ayaguz], Lake Balkhash basin, Kazakhstan. Syntypes: (90+) ZIN 4149-50 (6+, 6+). Also appeared in Kessler 1879:234 [ref. 14266]. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952]). •Questionably a synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 2006:88 [ref. 28831]). Current status: Synonym of Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae. Habitat: freshwater.

bigerri, Phoxinus Kottelat [M.] 2007:146, Fig. 2 [Ichthyological Exploration of Freshwaters v. 18 (no. 2); ref. 29201] River Adour in Tarbes (Séméac), 43°13'39"N, 0°05'27"E, Dépt. Hautes-Pyrénées, France. Holotype: MNHN 2006-1703. Paratypes: CMK 17575 (6), 17589 (5); MNHN 2006-1704 (4). Plus non-type specimens. •Valid as Phoxinus bigerri Kottelat 2007 -- (Kottelat & Freyhof 2007:225 [ref. 29996], Palandačić et al. 2017:3 [ref. 35519], Denys et al. 2020:211 [ref. 37846], Palandačić et al. 2022:[4] [ref. 39189]). Current status: Valid as Phoxinus bigerri Kottelat 2007. Leuciscidae: Phoxininae. Distribution: Southwestern Europe: Adour, Leyre and Ebro basins, France and Spain; introduced in the Aude and Garonne River drainages, France, and to Galicia and Duero River basin in Spain. Habitat: freshwater.

brachyurus, Phoxinus Berg [L. S.] 1912:241, Fig. 16 [v. 3 (pt 1); ref. 5874] Chilik River, Ili basin, Kazakhstan. Holotype (?): ZIN 10277 (1). Paratypes: ZIN 10277a (6). •Synonym of Phoxinus phoxinus (Linnaeus 1758), but a valid subspecies as described -- (Mitrofanov et al. 1987:141 [ref. 37731]). •Valid as Phoxinus brachyurus Berg 1912 -- (Berg 1949:586 [ref. 20661], Yang & Hwang 1964:24 [ref. 13497], Howes 1985:67 [ref. 5274], Zhu 1995:30 [ref. 25213], Luo in Chen et al. 1998:81 [ref. 23878], Strange & Mayden 2009:495 [ref. 30381], Zhang et al. 2016:54 [ref. 34477]). Current status: Valid as Phoxinus brachyurus Berg 1912. Leuciscidae: Phoxininae. Distribution: North Asia: Kazakhstan. Habitat: freshwater.

carpathicus, Phoxinus phoxinus Popescu Gorj [A.] & Dimitriu [M.] 1950:94, Fig. 6 [Buletinul Institutului de Cercetari Piscicole. v. 9; ref. 23004] Lake Roşu, Romainia. Syntypes: whereabouts unknown. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Kottelat 2007:156 [ref. 29201]). Current status: Synonym of Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae. Habitat: freshwater.

chrysoprasius, Cyprinus Pallas [P. S.] 1814:318 [Zoographia Rosso-Asiatica v. 3; ref. 3351] Upper reaches of the river Kuchuk-Karasu at Krasna Sloboda village, a tributary of Biyuk-Karasu, a tributary of Salhir, Crimea, 44°58ʹ26ʺN, 34°44ʹ19" [rocky banks of mountainous rivulets of the Crimea Peninsula]. Neotype: ZFMK 93640-59. Neotype designated by Bogutskaya et al. 2023:12 [ref. 40045]. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Kottelat 2007:156 [ref. 29201]). •Valid as Phoxinus chrysoprasius (Pallas 1814) -- (Bogutskaya et al. 2023:10 [ref. 40045], Dyldin et al. 2023:[36] [ref. 40328] with question, Artaev et al. 2024:301 [ref. 40926]). Current status: Valid as Phoxinus chrysoprasius (Pallas 1814). Leuciscidae: Phoxininae. Distribution: Southeast Europe: Salhir River drainage (Crimea). Habitat: freshwater.

colchicus, Phoxinus phoxinus Berg [L. S.] 1910:169 [Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 15; ref. 19593] Bachvis Tzchali [Bachwis-tzchali] River in Ozurgety District, Georgia, Eurasia. Holotype (unique): ZMT 141a. Based on Phoxinus laevis of Kamensky 1901:62-63 [ref. 14761]. See also Reshetnikov et al. 1997:740 [ref. 24702]. •Synonym of Phoxinus phoxinus (Linnaeus 1758), but a valid subspecies as described -- (Berg 1949:591 [ref. 20661], Lelek 1987:217 [ref. 22962], Bogutskaya 1997:174-175 [ref. 23566], Ninua & Japoshvili 2008:167 [ref. 31251]). •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Bogutskaya 1998:68 [ref. 23589], Sideleva 2001:51 [ref. 26181], Sideleva 2003:9 [ref. 27815], Vassilev & Pehlivanov 2005:170 [ref. 28369] as colhicus). •Valid as Phoxinus colchicus Berg 1910 -- (Bogutskaya & Naseka 2004:93 [ref. 28183], Kottelat 2007:156 [ref. 29201], Kottelat & Freyhof 2007:83 [ref. 29996], Kottelat & Freyhof 2007:226 [ref. 29996], Dyldin & Orlov 2016:662 [ref. 34853] mention, Çiçek et al. 2016:117 [ref. 35152], Palandačić et al. 2017:3 [ref. 35519], Çiçek et al. 2020:253 [ref. 37644], Kuljanishvili et al. 2020:[5] [ref. 37535], Çiçek et al. 2023:416 [ref. 40625], Dyldin et al. 2023:[36] [ref. 40328], Turan et al. 2023:6 [ref. 40351], Bayçelebi et al. 2024:106 [ref. 40795], Çiçek et al. 2024:267 [ref. 40789], Artaev et al. 2024:301 [ref. 40926]). Current status: Valid as Phoxinus colchicus Berg 1910. Leuciscidae: Phoxininae. Distribution: Eurasia: Caucasian Black Sea basin (Georgia, Russia and Turkey). Habitat: freshwater.

csikii, Phoxinus Hankó [B.] 1922:1, Pl. 1 (figs. 1-3) [A Magyar Tudományos Akadémia Balkán-Kutatásainak tudományos eredményei. v. 1; ref. 16373] Rožaje [Rozaj], on the Ibar River, tributary to the Zapadna Morava river, a tributary of the Danube, Montenegro, 42°50′39″N, 20°10′00″E. Neotype: NMW 98673 [ex 51266:2]. Original types MNH (2), destroyed by fire in 1956. Neotype designated by Palandačić et al. 2017:3 [ref. 35648]. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Kottelat 2007:156 [ref. 29201]). •Valid as Phoxinus csikii Hankó 1922 -- (Palandačić et al. 2017:14 [ref. 35519], Palandačić et al. 2017:3 [ref. 35648], Denys & Manne 2019:199 [ref. 36790], Bogutskaya et al. 2019:8 [ref. 37234], Denys et al. 2020:213 [ref. 37846], Palandačić et al. 2022:[4] [ref. 39189]). Current status: Valid as Phoxinus csikii Hankó 1922. Leuciscidae: Phoxininae. Distribution: Europe: Danube River system: from the Balkans to Germany, and the upper Rhine basin west to Lake Geneva; introduced in the lower Rhine, Germany, The Netherlands, France, and northern Italy. Habitat: freshwater.

dragarum, Phoxinus Denys [G. P. J.], Dettaï [A.], Persat [H.], Daszkiewicz [P.], Hautecoeur [M.] & Keith [P.] 2020:220, Fig. 8 [Cybium v. 44 (no. 3); ref. 37846] Arrat-Devant Stream, Manent-Montané, France, 43°20.7'N, 00°37.5'E, elevation 275 meters. Holotype: MNHN 2019-0262. Paratypes: MNHN. Plus additional non-type material. •Valid as Phoxinus dragarum Denys, Dettaï, Persat, Daszkiewicz, Hautecoeur & Keith 2020 -- (Palandačić et al. 2022:[4] [ref. 39189]). Current status: Valid as Phoxinus dragarum Denys, Dettaï, Persat, Daszkiewicz, Hautecoeur & Keith 2020. Leuciscidae: Phoxininae. Distribution: Western Europe: Garonne River basin, France; introduced in the Ebro and Guadalquivir drainages (Spain) and the Rhone Basin (France). Habitat: freshwater.

fayollarum, Phoxinus Denys [G. P. J.], Dettaï [A.], Persat [H.], Daszkiewicz [P.], Hautecoeur [M.] & Keith [P.] 2020:223, Fig. 9 [Cybium v. 44 (no. 3); ref. 37846] Boron Stream, Pionsat, France, 46°06.4'N, 02°41.1'E, elevation 522 meters. Holotype: MNHN 2019-0263. Paratypes: MNHN. Plus additional non-type material. •Valid as Phoxinus fayollarum Denys, Dettaï, Persat, Daszkiewicz, Hautecoeur & Keith 2020 -- (Palandačić et al. 2022:[4] [ref. 39189]). Current status: Valid as Phoxinus fayollarum Denys, Dettaï, Persat, Daszkiewicz, Hautecoeur & Keith 2020. Leuciscidae: Phoxininae. Distribution: Western Europe: Loire River basin and Sèvre Niortaise catchment (France); introduced in Garonne and Rhône River basins. Habitat: freshwater.

galian, Cyprinus Gmelin [J. F.] 1789:1421 [Caroli a Linné ... Systema Naturae per regna tria naturae v. 1 (pt 3); ref. 18139] "Circa Catharinopolin Sibiriae." •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 2006:45 [ref. 28831] as cf. phoxinus). Current status: Synonym of Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae. Habitat: freshwater.

grumi, Phoxinus Berg [L. S.] (ex Herzenstein) 1907:211 [16] [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 11 (for 1906); ref. 15129] Turfan Lake, Xinjiang Province, China. Syntypes: ZIN 10537-38 (1, 5). Apparently appeared first as name only as Herzenstein in Grum-Grzhimailo 1899:414 of v. 2. •Valid, tentatively in genus Lagowskiella -- (Howes 1985:63 [ref. 5274]). •Valid as Phoxinus grumi Berg 1907 -- (Zhu 1995:30 [ref. 25213], Luo in Chen et al. 1998:82 [ref. 23878], Zhang et al. 2016:54 [ref. 34477]). Current status: Valid as Phoxinus grumi Berg 1907. Leuciscidae: Phoxininae. Distribution: East Asia: Turfan Lake, Xinjiang Province, China. Habitat: freshwater.

isetensis, Cyprinus Georgi [J. G.] 1775:621 [Bemerkungen einer Reise im russischen Reich in 1772; ref. 18104] Cartharinopolis, Siberia, Russia. Syntypes: whereabouts unknown. Based on literature sources. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Kottelat 2006:45 [ref. 28831] as cf. phoxinus). •Valid as Phoxinus isetensis Georgi 1775 -- (Dyldin et al. 2023:[36] [ref. 40328]). Current status: Valid as Phoxinus isetensis (Georgi 1775). Leuciscidae: Phoxininae. Distribution: North Asia: Arctic Ocean basin, from Murman coast to East Siberian Sea basin (Kolyma drainage); rivers of northern and western Sea of Okhotsk drainage (Ola and Uda rivers), rivers of Peter the Great Bay drainage, probably Amur River basin and northwestern Sakhalin Island (Russia). Habitat: freshwater.

issykkulensis, Phoxinus Berg [L. S.] 1912:243, Fig. 17 [v. 3 (pt 1); ref. 5874] Lake Issyk-Kul, Kyrgyzstan. Syntypes: ZIN 9696 (5, not found), 13511 (5), 14536 (6), 15704 (3). •Valid as Phoxinus issykkulensis Berg 1912 -- (Berg 1949:587 [ref. 20661], Howes 1985:67 [ref. 5274] as issykylensis, Strange & Mayden 2009:495 [ref. 30381]). Current status: Valid as Phoxinus issykkulensis Berg 1912. Leuciscidae: Phoxininae. Distribution: Central Asia: Kyrgyzstan. Habitat: freshwater.

karsticus, Phoxinus Bianco [P. G.] & De Bonis [S.] 2015:8, Figs. 11-12 [Researches on Wildlife Conservation v. 4; ref. 34040] Endorheic River Trebinje, near Jasenica Village, Bosnia-Herzegovina. Holotype: IZA 00314-1. Paratypes: IZA. •Valid as Phoxinus karsticus Bianco & De Bonis 2015 -- (Palandačić et al. 2017:3 [ref. 35519], Shumka et al. 2023:39 [ref. 40747]). Current status: Valid as Phoxinus karsticus Bianco & De Bonis 2015. Leuciscidae: Phoxininae. Distribution: Southeastern Europe: Skadar Lake drainage (Bosnia-Herzegovina and Albania). Habitat: freshwater.

ketmaieri, Phoxinus Bianco [P. G.] & De Bonis [S.] 2015:6, Figs. 8-10 [Researches on Wildlife Conservation v. 4; ref. 34040] Krk Island, Baska brook at Baska draga, Croatia. Holotype: IZA 8823-1. Paratypes: IZA. •Synonym of Phoxinus lumaireul (Schinz 1840) -- (Palandačić et al. 2017:13 [ref. 35519]). Current status: Synonym of Phoxinus lumaireul (Schinz 1840). Leuciscidae: Phoxininae. Habitat: freshwater.

krkae, Phoxinus Bogutskaya [N. G.], Jelić [D.], Vucić [M.], Jelić [M,], Diripasko [O. A.], Stefanov [T.] & Klobučar [G.] 2019:4, Figs. 3-4 [Journal of Fish Biology v. 96 (no. 2); ref. 37234] Radljevac stream at Radljevac, Plavno, Krka drainage, Croatia, 44°06'22.6"N, 16°10'47.6"E, elevation about 200 meters. Holotype: MNW 99380 [ex HDBI 824]. Paratypes: HDBI, NMW. •Valid as Phoxinus krkae Bogutskaya, Jelić, Vucić, Jelić, Diripasko, Stefanov & Klobučar 2019. Current status: Valid as Phoxinus krkae Bogutskaya, Jelić, Vucić, Jelić, Diripasko, Stefanov & Klobučar 2019. Leuciscidae: Phoxininae. Distribution: Southeastern Europe: upper Krka River drainage, Knin Valley, Croatia. Habitat: freshwater.

kubanicum, Phoxinus phoxinus Emtyl [M. K.] & Ivanenko [A. M.] 2002:90, Fig. 59 [ ref. 40927] Trans-Kuban Rivers, Russia. No types known. No types mentioned, not available [ICZN Art. 16.4.1]. •Unavailable name -- (Artaev et al. 2024:300 [ref. 40926]). Current status: Synonym of Phoxinus. Leuciscidae: Phoxininae.

kuldschiensis, Phoxinus Warpachowski [N. A.] 1887:690 [Mélanges biologiques v. 12; ref. 13838] Lake Balkhash, Kazakhstan. Syntypes: ZIN 7785 (1). •Synonym of Phoxinus poljakowii Kessler 1879 -- (Berg 1905:141 [ref. 15338]). Current status: Synonym of Phoxinus poljakowii Kessler 1879. Leuciscidae: Leuciscinae. Habitat: freshwater.

laevis, Phoxinus Fitzinger [L. J. F. J.] (ex Agassiz) 1832:337 [Beiträge zur Landeskunde Österreichs unter der Enns, Wien v. 1; ref. 5019] Unneeded new name for Cyprinus phoxinus Linnaeus 1758. •Objective synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Bogutskaya 1998:68 [ref. 23589], Sideleva 2001:51 [ref. 26181], Sideleva 2003:9 [ref. 27815], Vassilev & Pehlivanov 2005:170 [ref. 28369], Parin et al. 2014:86 [ref. 33547], Denys et al. 2020:214 [ref. 37846]). Current status: Synonym of Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae. Habitat: freshwater.

likai, Phoxinus Bianco [P. G.] & De Bonis [S.] 2015:12, Figs. 16-17 [Researches on Wildlife Conservation v. 4; ref. 34040] River Oruča [Otuča] system, near the village of Gračac, Croatia. Holotype: IZA 02263-1. Paratypes: IZA. •Synonym of Phoxinus lumaireul (Schinz 1840) -- (Palandačić et al. 2017:13 [ref. 35519]). Current status: Synonym of Phoxinus lumaireul (Schinz 1840). Leuciscidae: Phoxininae. Habitat: freshwater.

lumaireul, Cyprinus Schinz [H. R.] 1840:331 [Europaïsche Fauna; ref. 18695] Po River, Italy. No types known. Appeared first as name only, not available, from Pô, Italy as Bonelli in Agassiz 1835:37 [ref. 22]. Also appeared in Nordmann 1840:482 [ref. 3209] as name in synonymy. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Bianco & Delmastro 2011:5 [ref. 31648], Bianco 2013:9 [ref. 32897], Bianco & De Bonis 2015:3 [ref. 34044]). •Valid as Phoxinus lumaireul (Schinz 1840) -- (Kottelat 2007:156 [ref. 29201], Kottelat & Freyhof 2007:227 [ref. 29996], Palandačić et al. 2017:13 [ref. 35519], Ćaleta et al. 2019:162 [ref. 37026], Lorenzoni et al. 2019:252 [ref. 37108], Bogutskaya et al. 2019:10 [ref. 37234], Palandačić et al. 2022:[4] [ref. 39189]). Current status: Valid as Phoxinus lumaireul (Schinz 1840). Leuciscidae: Phoxininae. Distribution: Southeastern Europe: North Adriatic Sea basin and middle Danube drainage, Italy, Slovenia, Croatia and Bosnia and Herzegovina. Habitat: freshwater.

marsilii, Phoxinus Heckel [J. J.] 1836:232 [Annalen des Wiener Museums der Naturgeschichte v. 1 (no. 10); ref. 18254] Clear brooks of the environs of Vienna and beyond, Austria. Lectotype: NMW 51225:2. Paralectotypes: NMW-51225 (5). Lectotype selected by Palandačić et al. 2017:2 [ref. 35648]. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Kottelat 2007:156 [ref. 29201]). •Valid as Phoxinus marsilii Heckel 1836 -- (Palandačić et al. 2017:13 [ref. 35519], Palandačić et al. 2017:2 [ref. 35648], Ćaleta et al. 2019:163 [ref. 37026], Bogutskaya et al. 2019:15 [ref. 37234], Palandačić et al. 2022:[4] [ref. 39189]). Current status: Valid as Phoxinus marsilii Heckel 1836. Leuciscidae: Phoxininae. Distribution: Europe: middle and lower Danube drainage, Austria, Hungary and Croatia. Habitat: freshwater.

mikrosquamatus, Phoxinus laevis Kaschenko [N.] 1899:145 [Results of the Altai Zoological Expedition; ref. 19630] Lake Karalachinskoye in Argut River basin, Altai, Siberia, Russia. Holotype (unique): whereabouts unknown. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Berg 1949:591 [ref. 20661], Kottelat 1997:74 [ref. 22952], Bogutskaya 1998:68 [ref. 23589], Sideleva 2001:51 [ref. 26181], Bogutskaya 1997:175 [ref. 23566], Bogutskaya & Naseka 2004:96 [ref. 28183]). •Synonym of Phoxinus ujmonensis Kaschenko 1899 -- (Kottelat 2006:46 [ref. 28831]). Current status: Synonym of Phoxinus ujmonensis Kaschenko 1899. Leuciscidae: Phoxininae. Habitat: freshwater.

montanus, Phoxynus Ogérien [L. F.] 1863:362 [Histoire naturelle du Jura et des départements voisins; ref. 23003] Ain River near Champagnole; Queue de Cheval waterfall, near Saint-Claude, France. No types known. Original genus should have been spelled Phoxinus. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952]). •Synonym of Phoxinus septimaniae Kottelat 2007 -- (Denys et al. 2020:220 [ref. 37846] as nomen oblitum). Nomen Oblitum. Current status: Synonym of Phoxinus septimaniae Kottelat 2007. Leuciscidae: Phoxininae. Habitat: freshwater.

morella, Cyprinus Leske [N. G.] 1774:47 [Ichthyologiae Lipsiensis specimen.; ref. 17505] Bode Creek near Rübeland, 103 kilometers northwest of Leipzig, Germany. Neotype: ZMB 32661. Neotype designated by Kottelat 1997:74 [ref. 22952]. •Possibly valid as Phoxinus morella (Leske 1774) -- (Palandačić et al. 2017:14 [ref. 35519], Palandačić et al. 2022:[4] [ref. 39189] as cf. morella). •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:73 [ref. 22952]). Current status: Synonym of Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae. Distribution: Central Europe: Elbe River drainage, Czech Republic and Germany (if valid). Habitat: freshwater.

obtusus, Leuciscus De la Pylaie [A. J. M.] 1835:533 [Congrès Scientifique de France v. 2 (Sept. 1834) (art. 5); ref. 1086] Name only, not available; from Vendée, France. •In the synonymy of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952]). Nomen Nudum. Current status: Synonym of Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae.

oxyrhynchus, Moroco Mori [T.] 1930:45 [7], Pl. 3 (fig. 2) [Journal of the Chosen Natural History Society No. 11; ref. 16505] Mo-san, Korea. Holotype: not researched. Additional specimens: (3). On p. 7 of separate. •Synonym of Phoxinus lagowskii Dybowski 1869 -- (Berg 1949:584 [ref. 20661]). •Valid as Phoxinus oxyrhynchus (Mori 1930) -- (Shedko & Shedko 2003:322 [ref. 27966], Bogutskaya & Naseka 2004:94 [ref. 28183]). Current status: Valid as Phoxinus oxyrhynchus (Mori 1930). Leuciscidae: Phoxininae. Distribution: Korea. Habitat: freshwater.

phoxinus, Cyprinus Linnaeus [C.] 1758:322 [Systema Naturae, Ed. X v. 1; ref. 2787] River Agger north of Lohmar, 50°50'N, 7°12'E, Nordrhein-Westfalen, Germany. Neotype: NRM 55108. Neotype designated by Fricke 1999:25 [ref. 24101], but withdrawn in Fricke 2000:639 [ref. 24537]. Neotype validly established by Kottelat 2007:153 [ref. 29201]. For remarks on systematics see Kottelat 2006:45-46 [ref. 28831]. •Valid as Cyprinus phoxinus Linnaeus 1758 -- (Herrera 1896:21 [ref. 39336]). •Valid as Phoxinus phoxinus (Linnaeus 1758), as a valid subspecies -- (Mitrofanov et al. 1987:132 [ref. 37731]). •Valid as Phoxinus phoxinus (Linnaeus 1758) -- (Berg 1949:588 [ref. 20661], Yang & Hwang 1964:23 [ref. 13497], Howes 1985:67 [ref. 5274], Mrakovcic et al. 1995:184 [ref. 23079], Lelek 1987:217 [ref. 22962], Zhu 1995:30 [ref. 25213], Bogutskaya & Naseka 1996:19 [ref. 22798], Chereshnev 1996:600 [ref. 24659], Bianco et al. 1996:52 [ref. 26724], Kottelat 1997:73 [ref. 22952], Bogutskaya 1997:174 [ref. 23566], Kim 1997:256 [ref. 24032], Reshetnikov et al. 1997:740 [ref. 24702], Bogutskaya 1998:68 [ref. 23589], Luo in Chen et al. 1998:75 [ref. 23878], Briolay et al. 1998:101 [ref. 25586], Fricke 1999:25 [ref. 24101], Novikov et al. 2000:161 [ref. 40929], Sheiko & Fedorov 2000:17 [ref. 25650], Bogutskaya et al. 2001:43 [ref. 26178], Chereshnev et al. 2001:46 [ref. 26736], Pietsch et al. 2001:139 [ref. 25688], Sideleva 2001:51 [ref. 26181], Kim & Park 2002:164 [ref. 25913], Choi et al. 2003:158, 517 [ref. 26218], Fedorov et al. 2003:25 [ref. 27579], Sideleva 2003:9 [ref. 27815], Hanel 2003:49 [ref. 27894], Bogutskaya & Naseka 2004:96 [ref. 28183], Vassilev & Pehlivanov 2005:170 [ref. 28369], Hanel & Lusk 2005:200 [ref. 28804], Kottelat 2006:88 [ref. 28831], Ocock et al. 2006:42 [ref. 28933], Kottelat 2007:145 [ref. 29201], Fricke 2007:28 [ref. 30577], Fricke et al. 2007:44 [ref. 29533], Kottelat & Freyhof 2007:228 [ref. 29996], Bianco & Delmastro 2011:5 [ref. 31648], Strange & Mayden 2009:495 [ref. 30381], Bianco 2013:9 [ref. 32897], Parin et al. 2014:85 [ref. 33547], Bianco & De Bonis 2015:3 [ref. 34040], Çiçek et al. 2015:146 [ref. 35151], Zhang et al. 2016:54 [ref. 34477], Dyldin & Orlov 2016:662 [ref. 34853], Palandačić et al. 2017:3 [ref. 35519], Lorenzoni et al. 2019:252 [ref. 37108], Denys et al. 2020:214 [ref. 37846], Dyldin et al. 2021:26 [ref. 38804] with question, Dyldin & Orlov 2021:67 [ref. 38158] with question, Palandačić et al. 2022:[4] [ref. 39189], Nagy et al. 2023:357 [ref. 40348]). Current status: Valid as Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae. Distribution: Western Europe: Rhine, Meuse and Seine River drainages, coastal catchments of Normandy and English Channel and Thames River drainage. Introduced in Italy, Corsica and Ireland. Habitat: freshwater.

poljakowii, Phoxinus Kessler [K. F.] 1879:283 [Bulletin de l'Académie Impériale des Sciences de St. Pétersbourg v. 25; ref. 2598] Ajagus [Ayaguz] near Sergiopol, Lake Balkhash, Kazakhstan. Syntypes: (ca. 20) ZIN 4151 (15). Also appeared as Phoxinus poljakowii in Kessler 1879:235 [ref. 14266]. •Synonym of Phoxinus phoxinus (Linnaeus 1758), but a valid subspecies as described -- (Mitrofanov et al. 1987:138 [ref. 37731]). •Valid as Phoxinus poljakowii Kessler 1879 -- (Berg 1949:585 [ref. 20661]). •Valid, tentatively in genus Lagowskiella -- (Howes 1985:63 [ref. 5274]). Current status: Valid as Phoxinus poljakowii Kessler 1879. Leuciscidae: Phoxininae. Distribution: Central Asia: Kazakhstan. Habitat: freshwater.

radeki, Phoxinus Bayçelebi [E.], Aksu [İ.] & Turan [D.] 2024:103, Figs. 2-4 [Zoosystematics and Evolution v. 100 (no. 1); ref. 40795] Stream Büyükdere about 6 km west of Pınarhisar, Kırklareli Province, Türkiye, 41.6337, 27.5994. Holotype: FFR 2322. Paratypes: FFR. •Valid as Phoxinus radeki Bayçelebi, Aksu & Turan 2024. Current status: Valid as Phoxinus radeki Bayçelebi, Aksu & Turan 2024. Leuciscidae: Phoxininae. Distribution: Asia Minor: Ergene River drainage, Ageaen Sea watershed, Turkey. Habitat: freshwater.

rivularis, Salmo Pallas [P. S.] 1773:617, 717 [Reise durch verschiedene Provinzen des russischen Reiches; ref. 18509] Basin of the river Altaï, Siberia, Russia. No types known. Spelled riuularis on p. 717, rivularis once on p. 617; first reviser not researched but riuularis probably can be regarded as a spelling error. Placed in genus Cyprinus by Bonnaterre 1788:195 [ref. 4940]. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Berg 1949:591 [ref. 20661], Bogutskaya & Naseka 1996:20 [ref. 22798], Kottelat 1997:73 [ref. 22952], Bogutskaya 1998:68 [ref. 23589], Sideleva 2003:9 [ref. 27815]). •Questionably a synonym of Phoxinus ujmonensis Kaschenko 1899 -- (Kottelat 2006:46 [ref. 28831]). Current status: Synonym of Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae. Habitat: freshwater.

saposchnikowi, Phoxinus Kaschenko [N.] 1899:146 [Results of the Altai Zoological Expedition; ref. 19630] Lake on plateau of Ukök [or Ukëk] , source of river Kalguty, tributary of River Argut, Argut River basin, Altai, Siberia, Russia. Holotype (unique): whereabouts unknown. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Berg 1949:591 [ref. 20661], Kottelat 1997:74 [ref. 22952], Bogutskaya 1998:68 [ref. 23589], Sideleva 2001:51 [ref. 26181], Sideleva 2003:9 [ref. 27815], Bogutskaya & Naseka 2004:96 [ref. 28183]). •Synonym of Phoxinus ujmonensis Kaschenko 1899 -- (Kottelat 2006:46 [ref. 28831]). Current status: Synonym of Phoxinus ujmonensis Kaschenko 1899. Leuciscidae: Phoxininae. Habitat: freshwater.

sedelnikowi, Phoxinus czekanowskii Berg [L. S.] 1908:226 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 13 (no. 2); ref. 14302] Saissan [Zaisan Lake], upper Irtysch basin, Kazakhstan. Lectotype: ZIN 14361. Paralectotypes: ZIN 14361a (6). Lectotype from ZSIP catalog, selected by Berg but publication not reserched [not designated here]. •Synonym of Phoxinus phoxinus (Linnaeus 1758), but a valid subspecies as described -- (Mitrofanov et al. 1987:135 [ref. 37731]). •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Bogutskaya & Naseka 2004:96 [ref. 28183] as sedelnikovi). •Synonym of Phoxinus ujmonensis Kaschenko 1899 -- (Kottelat 2006:46 [ref. 28831]). •Valid as Phoxinus sedelnikowi Berg 1908 -- (Berg 1949:585 [ref. 20661], Howes 1985:67 [ref. 5274]). Current status: Synonym of Phoxinus ujmonensis Kaschenko 1899. Leuciscidae: Phoxininae. Distribution: Central Asia: upper Irtysh River basin, Kazakhstan [if valid]. Habitat: freshwater.

septimaniae, Phoxinus Kottelat [M.] 2007:148, Fig. 3 [Ichthyological Exploration of Freshwaters v. 18 (no. 2); ref. 29201] River Agly at bridge downstream of Latour-de-France, 42°46'09"N, 2°39'39"E, Dépt. Pyrénées-Orientales, France. Holotype: MNHN 2006-1701. Paratypes: CMK 13006 (5), 17565 (4), 17603 (12), 18348 (8), 18508 (2); MNHN 2006-1702 (7). Plus non-type material. •Valid as Phoxinus septimaniae Kottelat 2007 -- (Kottelat & Freyhof 2007:229 [ref. 29996], Palandačić et al. 2017:12 [ref. 35519] as septimanae, Denys et al. 2020:216 [ref. 37846] as nomen protectum, Palandačić et al. 2022:[4] [ref. 39189]). Nomen Protectum. Current status: Valid as Phoxinus septimaniae Kottelat 2007. Leuciscidae: Phoxininae. Distribution: Western Europe: Rhône River basin and the French coastal Mediterranean catchments, France; introduced to the Roya and Rhine River drainages and Lakes Geneva and Ceresole (Italy). Habitat: freshwater.

strandjae, Phoxinus phoxinus Drensky [P.] 1926:137 [Trudove na Bulgarskogo Prirodoizpitatelno Druzhestvo = Travaux de la Société Bulgare des Sciences Naturelles v. 12; ref. 15583] Strandscha [Istrandzhha] range, Bulgaria. Syntypes: NMNHS. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Vassilev & Pehlivanov 2005:170 [ref. 28369], Kottelat 2007:156 [ref. 29201]). •Valid as Phoxinus strandjae Drensky 1926 -- (Kottelat & Freyhof 2007:229 [ref. 29996], Çiçek et al. 2015:146 [ref. 35151], Palandačić et al. 2017:14 [ref. 35519] with question, Özuluğ & Saç 2019:30 [ref. 36599], Çiçek et al. 2020:253 [ref. 37644], Çiçek et al. 2023:417 [ref. 40625], Turan et al. 2023:7 [ref. 40351], Bayçelebi et al. 2024:106 [ref. 40795], Çiçek et al. 2024:268 [ref. 40789]). Current status: Valid as Phoxinus strandjae Drensky 1926. Leuciscidae: Phoxininae. Distribution: Southeastern Europe: southwest Black Sea watersheds, Bulgaria and Istanbul and Kirklareli Provinces (Turkey). Habitat: freshwater.

strymonicus, Phoxinus Kottelat [M.] 2007:151, Fig. 4 [Ichthyological Exploration of Freshwaters v. 18 (no. 2); ref. 29201] Strymon drainage, Angitis system, canals near Kalambaki, about 14 kilometers south of Drama, Macedonia, Greece. Holotype: MHNG 2680.072. Paratypes: CMK 12912 (2), 12918 (38); HCMR 1553 (10); MHNG 2680.073 (10). •Valid as Phoxinus strymonicus Kottelat 2007 -- (Kottelat & Freyhof 2007:230 [ref. 29996], Barbieri et al. 2015:59 [ref. 35029], Palandačić et al. 2017:14 [ref. 35519] with question, Bayçelebi et al. 2024:107 [ref. 40795]). Current status: Valid as Phoxinus strymonicus Kottelat 2007. Leuciscidae: Phoxininae. Distribution: Southeastern Europe: Struma River basin, Macedonia, Greece and Bulgaria. Habitat: freshwater.

tchangi, Phoxinus Chen [X.-Y.] 1988:35 [English p. 37], Fig. 2 [Sinozoologia v. 6; ref. 16227] Weifen River, China. Holotype: ASIZB S01012. Paratypes: ASIZB S01001-05, 01006 (dissected), 01007-11, 01013-19 (dissected). •Valid as Phoxinus tchangi Chen 1988. Current status: Valid as Phoxinus tchangi Chen 1988. Leuciscidae: Phoxininae. Distribution: East Asia: northern China. Habitat: freshwater.

tschuensis, Phoxinus brachyurus Pivnev [I. A.] 1985:55 [8 pls.; ref. 8966] Chu River, Kyrgyzstan. Syntypes: (many). Leuciscidae: Phoxininae. Habitat: freshwater.

tumensis, Phoxinus phoxinus Luo [Y.-L.] 1996:291, Fig. 1 [Acta Hydrobiologica Sinica v. 20 (no. 3); ref. 23189] Jilin, China. Holotype: Inst. Hydrobiol., Chinese Acad. Sci. Wuhan 790237. Paratypes: (?10, not 9) Inst. Hydrobiol. Chinese Acad. Sci. Wuhan 790239-41 (3), 790245-47 (3), 790253 (1), 773454-55 (2), 773457 (1). •Synonym of Phoxinus phoxinus (Linnaeus 1758), but a valid subspecies as described -- (Luo in Chen et al. 1998:79 [ref. 23878], Zhang et al. 2016:54 [ref. 34477], Dyldin & Orlov 2016:662 [ref. 34853]). Current status: Valid as Phoxinus tumensis Luo 1996. Leuciscidae: Phoxininae. Distribution: East Asia: northern China. Habitat: freshwater.

ujmonensis, Phoxinus laevis Kaschenko [N.] 1899:144 [Results of the Altai Zoological Expedition; ref. 19630] Katun River, near Uimon village, Altai, Ob River basin, Siberia, Russia. Syntypes: (4) whereabouts unknown. Lectotype established by Berg 1949:592, Fig. 349 if specimen is identifiable. Author also seen as Kashchenko. For remarks on systematics see Kottelat 2006:46 [ref. 28831]. •Synonym of Phoxinus phoxinus (Linnaeus 1758), but a valid subspecies -- (Berg 1949:591 [ref. 20661], Bogutskaya 1997:175 [ref. 23566] as uimonensis, Luo in Chen et al. 1998:78 [ref. 23878]). •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], see Bogutskaya 1998:68 [ref. 23589], Sideleva 2001:51 [ref. 26181], Sideleva 2003:9 [ref. 27815], Bogutskaya & Naseka 2004:96 [ref. 28183]). •See also (Reshetnikov et al. 1997:740 [ref. 24702], Ocock et al. 2006:43 [ref. 28933]). •Valid as Phoxinus phoxinus (Linnaeus 1758) subspecies ujmonensis Luo 1996 -- (Zhang et al. 2016:54 [ref. 34477] as Phoxinus laevis and author as Kaschtschenko). •Valid as Phoxinus ujmonensis Kaschenko 1889 -- (Kottelat 2006:46 [ref. 28831], Kottelat 2007:157 [ref. 29201], Dyldin & Orlov 2016:662 [ref. 34853], Romanov et al. 2017:822 [ref. 35687], Dyldin et al. 2023:[37] [ref. 40328] with question). Current status: Valid as Phoxinus ujmonensis Kaschenko 1899. Leuciscidae: Phoxininae. Distribution: Central and east Asia: Mongolia, China, Kazakhstan and Russia. Habitat: freshwater.

varius, Phoxinus Perty [J. A. M.] 1832:719 [Isis (Oken) v. 25 (no. 7); ref. 18534] Unneeded new name for Cyprinus phoxinus Linnaeus 1758. •Synonym of Phoxinus phoxinus (Linnaeus 1758) -- (Kottelat 1997:74 [ref. 22952], Denys et al. 2020:215 [ref. 37846]). Current status: Synonym of Phoxinus phoxinus (Linnaeus 1758). Leuciscidae: Phoxininae. Habitat: freshwater.

varius, Pisciculus Dralet [É.-F.] 1821:35 [Considérations sur l'histoire naturelle des poissons, sur la pêche et les lois qui la régissent.; ref. 17418] Luech Stream, lower Rhône drainage, Chamborigaud, France [no locality]. Neotype: MNHN 2019-0264. Based on Rondelet. Neotype designated by Denys et al. 2020:219 [ref. 37846]. •Synonym of Phoxinus septimaniae Kottelat 2007 -- (Denys et al. 2020:216 [ref. 37846] as nomen oblitum). Nomen Oblitum. Current status: Synonym of Phoxinus septimaniae Kottelat 2007. Leuciscidae: Leuciscinae. Habitat: freshwater.


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