Institute for Biodiversity Science and Sustainability
Online Version, Updated 1 April 2019

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Eschmeyer's Catalog of Fishes
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Říĉan, O., R. Zardoya and I. Doadria  2008 [ref. 31419]
Phylogenetic relationships of Middle America cichlids (Cichlidae, Heroini) based on combined evidence from nuclear genes, mtDNA, and morphology. Molecular Phylogenetics and Evolution v. 49: 941-957.