Institute for Biodiversity Science and Sustainability
Online Version, Updated 1 Nov 2022

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Eschmeyer's Catalog of Fishes
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tymensis, Gasterosteus Nikolskii [A. M.] 1889:293 [Sakhalin Island and its fauna of vertebarate animals; ref. 14493] Tym River, Sakhalin Island, Russia. Syntypes: (many) BMNH 1892.4.28.1 [ex ZIN]; ZIN 6593-95 (6+, 6+, 6+). Authorship sometimes as Nikolsky. •Synonym of Pungitius pungitius (Linnaeus 1758), but a valid subspecies -- (Okada 1961:619 [ref. 6363], Monod 1973:285 [ref. 7193], Ida in Masuda et al. 1984:83 [ref. 6441], Nikiforov et al. 1997:330 [ref. 24694], Keivany & Nelson 2000:119 [ref. 24720], Chereshnev 2002:704 [ref. 26468], Kolpakov 2003:847 [ref. 27537], Pichugin et al. 2004:15 [ref. 27623]). •Valid as Pungitius tymensis (Nikolskii 1889) -- (Berg 1949:968 [ref. 20662], Kawanabe & Mizuno 1989:442 [ref. 25598], Kim 1997:390 [ref. 24032], Reshetnikov et al. 1997:748 [ref. 24702], Reshetnikov & Savvaitove 1998:112 [ref. 23593], Nakabo 2000:514 [ref. 25086], Matsuura et al. 2000:199 [ref. 26000], McLennan & Mattern 2001:11 [ref. 25222], Pietsch et al. 2001:144 [ref. 25688], Kim & Park 2002:312 [ref. 25913], Bogutskaya et al. 2001:46 [ref. 26178], Nakabo 2002:514 [ref. 26001], Choi et al. 2003:231, 554 [ref. 26218], Bogutskaya & Naseka 2004:182 [ref. 28183], Takahashi et al. 2016:49 [ref. 34337], Dyldin & Orlov 2017:58 [ref. 35138], Dyldin et al. 2021:59 [ref. 38804], Matsumoto et al. 2021:15 [ref. 38508], Wang et al. 2021:[3] [ref. 38800], Dyldin & Orlov 2022:36 [ref. 38972]). Current status: Valid as Pungitius tymensis (Nikolskii 1889). Gasterosteidae. Distribution: East Asia: Russia and Japan. Habitat: freshwater.


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