quadrispinosa, Gasterosteus Crespon [J.] 1844:283 [Faune méridionale, ou description de tous les animaux vertébrés vivans et fosses v. 2; ref. 18056] Vistre River and ditches around Nîmes, France. Syntypes: whereabouts unknown. •Synonym of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:397 [ref. 40085]). Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.