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Eschmeyer's Catalog of Fishes
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Species in the family or subfamily Acheilognathidae: [ 128 ] records


albomarginatus, Rhodeus Li [F.] & Arai [R.] 2014:167, Figs. 1-8 [Zootaxa 3790 (no. 1); ref. 33267] Lvjiang River, Yangtze River system, Xuling Town, Qimen County, Anhui Province, China. Holotype: SOU 1306001. Paratypes: NSMT-P, SOU, ZUMT. •Valid as Rhodeus albomarginatus Li & Arai 2014 -- (Zhang et al. 2016:68 [ref. 34477], Li et al. 2017:84 [ref. 35680], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:337 [ref. 37574]). Current status: Valid as Rhodeus albomarginatus Li & Arai 2014. Acheilognathidae. Distribution: Lvjiang River drainage, Yangtze River basin, Anhui, China. Habitat: freshwater.

amarus, Cyprinus Bloch [M. E.] 1782:52, Pl. 8 (fig. 3) [M. Marcus Elieser Bloch's ..., ausübenden Arztes zu Berlin, Oeconomische Naturgeschichte der Fische Deutschlands v. 1; ref. 19643] Müggelsee (lake) near Köpenick, Berlin, Germany. Syntypes: ZMB 3393 (3). Type catalog: Paepke 1999:69 [ref. 24282]. On p. 45 of Bloch's Ichthyologie, v. 1 [ref. 21381]. •Synonym of Rhodeus sericeus (Pallas 1776), but a valid subspecies -- (Berg 1949:814 [ref. 20661], Coad 1995:19 [ref. 23608], Pipoyan 1996:710 [ref. 24654], Reshetnikov et al. 1997:740 [ref. 24702], Bogutskaya 1998:80 [ref. 23589], Arai et al. 2001:275 [ref. 25545], Bogutskaya & Komlev 2001:81 [ref. 26834], Vassilev & Pehlivanov 2005:170 [ref. 28369]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus sericeus (Pallas 1776) -- (Smith 1986:124 [ref. 22186], Lelek 1987:223 [ref. 22962], Naseka 1998:80 [ref. 23590], Coad 1998:102 [ref. 23869], Holčík 1999:2 [ref. 24847], Fuller et al. 1999:149 [ref. 25838], Bogutskaya et al. 2001:44 [ref. 26178]). •Valid as Rhodeus amarus (Bloch 1782) -- (Mrakovcic et al. 1995:184 [ref. 23079], Kottelat 1997:75 [ref. 22952], Gabrielyan 2001:27 [ref. 25867], Bianco & Ketmaier 2001:200 [ref. 26728], Bogutskaya & Naseka 2004:43 [ref. 28183], Hanel & Lusk 2005:256 [ref. 28804], Kottelat 2006:47 [ref. 28831], Fricke et al. 2007:45 [ref. 29533], Kottelat & Freyhof 2007:72 [ref. 29996], Yang et al. 2010:333 [ref. 30786], Esmaeili et al. 2010:371 [ref. 33391], Yang et al. 2011:158 [ref. 31154], Parin et al. 2014:87 [ref. 33547], Jouladeh-Roudbar et al. 2015:879 [ref. 34062], Barbieri et al. 2015:60 [ref. 35029], Çiçek et al. 2015:144 [ref. 35151], Dyldin & Orlov 2016:663 [ref. 34853], Esmaeili et al. 2017:55 [ref. 35254], Li et al. 2017:84 [ref. 35680], Esmaeili et al. 2018:35 [ref. 36089], Ćaleta et al. 2019:150 [ref. 37026], Lorenzoni et al. 2019:252 [ref. 37108], Özuluğ & Saç 2019:24 [ref. 36599], Çiçek et al. 2020:246 [ref. 37644], Kaya et al. 2020:330 [ref. 37582], Kuljanishvili et al. 2020:[4] [ref. 37535], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:337 [ref. 37574], Yogurtçuoglu et al. 2020:[3] [ref. 37627], Esmaeili et al. 2020:335 [ref. 37759], Jouladeh-Roudbar et al. 2020:102 [ref. 37983], Bayçelebi 2021:641 [ref. 38115]). Current status: Valid as Rhodeus amarus (Bloch 1782). Acheilognathidae. Distribution: Northern and eastern Europe, Turkey. Introduced in Italy. Habitat: freshwater, brackish.

amurensis, Acanthorhodeus asmussii Holčík [J.] 1962:160, Figs. 3-4 [Japanese Journal of Ichthyology v. 9 (nos 1-6); ref. 5934] Lake Kabar, Amur River near Jelabuga (ca 60 kilometers from town of Khabarowsk), Russia. Holotype: SNMB 386. Paratypes: SNMB 376-385 (10), 387 (1), 388 (1), 389 (1). Type catalog: Blahák 1991:204 [ref. 20023]. •In genus Acheilognathus -- (Arai & Akai 1988:206 [ref. 6999]). •Synonym of Acanthorhodeus asmussii (Dybowski 1872) -- (Naseka & Bogutskaya 2004:281 [ref. 27838], Bogutskaya & Naseka 2004:42 [ref. 28183], Bogutskaya et al. 2008:314 [ref. 30085]). •Synonym of Acheilognathus asmussii (Dybowski 1872) -- (Kottelat 2006:27 [ref. 28831]). •Valid as Acheilognathus amurensis (Holčík 1962) -- (Dyldin et al. 2020:86 [ref. 37569]). Current status: Valid as Acheilognathus amurensis (Holčík 1962). Acheilognathidae. Distribution: East Asia: Amur River system (Russia, China). Habitat: freshwater.

amurensis, Pseudoperilampus lighti Vronsky [B. B.] 1967:24 [Voprosy Ikhtiologii v. 7; ref. 12489] Lower Amur River drainage, Russia. •Synonym of Acanthorhodeus chankaensis (Dybowski 1872) -- (Bogutskaya & Naseka 1996:32 [ref. 22798]). •Synonym of Rhodeus lighti (Wu 1931) -- (Naseka 1998:80 [ref. 23590]). •Questionably a synonym of Rhodeus sinensis Günther 1868 -- (Bogutskaya et al. 2001:44 [ref. 26178]). •Valid as Rhodeus amurensis (Vronsky 1967) -- (Naseka & Bogutskaya 2004:281 [ref. 27838], Bogutskaya & Naseka 2004:43 [ref. 28183], Bogutskaya et al. 2008:316 [ref. 30085]). Current status: Valid as Rhodeus amurensis (Vronsky 1967). Acheilognathidae. Distribution: Eastern Europe. Habitat: freshwater.

argenteus, Acheilognathus Wu [H.-W.] 1939:118 [Sinensia v. 10 (nos 1-6); ref. 4805] Yangso, Li Kiang, southern China. Syntypes: NIZBN 160 (1), 404 (1). Types may be at NIZBN = National Instute of Zoology and Botany, Nanking, China. •In genus Acheilognathus (Arai & Akai 1988:206 [ref. 6999]). •Synonym of Acheilognathus barbatulus Günther 1873 -- (Zheng et al. 1989:161 [ref. 21202], Lin in Chen et al. 1998:425 [ref. 23878], Kottelat 2013:70 [ref. 32989], Zhang et al. 2016:66 [ref. 34477]). Current status: Synonym of Acheilognathus barbatulus Günther 1873. Acheilognathidae. Habitat: freshwater.

asmussii, Devario Dybowski [B. N.] 1872:212 [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 22; ref. 1170] Lake Khanka [Chanka], southeastern Russia. Neotype: ZIN 28193. Possible original types: Mus. Zool. Inst. Polish Acad. Sci. 6112 (2, poor condition), ZMB 7936 (2) Lake Khanka. Neotype designated by Holcik 1962 [ref. 5934]. See Kottelat 2006:27 [ref. 28831] for remarks on systematics. •In genus Acheilognathus -- (Arai & Akai 1988:206 [ref. 6999]). •Synonym of Acheilognathus macropterus (Bleeker 1871) -- (Kim 1997:545 [ref. 24032] as assmussi, Lin in Chen et al. 1998:419 [ref. 23878]). •Valid as Acanthorhodeus asmussii (Dybowski 1872) -- (Berg 1949:817 [ref. 20661] and Wu 1964:212 [ref. 13503] as asmussi, Reshetnikov et al. 1997:734 [ref. 24702], Naseka 1998:79 [ref. 23590], Bogutskaya & Naseka 1996:30 [ref. 22798], Bogutskaya et al. 2001:44 [ref. 26178], Naseka & Bogutskaya 2004:280 [ref. 27838], Bogutskaya & Naseka 2004:41 [ref. 28183], Bogutskaya et al. 2008:314 [ref. 30085]). •Valid as Acheilognathus asmussii (Dybowski 1872) -- (Kottelat 2006:27 [ref. 28831], Ocock et al. 2006:31 [ref. 28933], Yang et al. 2011:163 [ref. 31154], Parin et al. 2014:66 [ref. 33547]). Current status: Valid as Acheilognathus asmussii (Dybowski 1872). Acheilognathidae. Distribution: Russia, Mongolia, China and Korea. Habitat: freshwater.

atranalis, Acanthorhodeus Günther [A.] 1873:248 [Annals and Magazine of Natural History (Series 4) v. 12 (no. 69) (art. 31); ref. 2004] Shanghai, China. Syntypes: BMNH 1873.7.30.84 (1), RMNH 10384 [ex BMNH] (1). •Synonym of Acanthorhodeus chankaensis (Dybowski 1872) -- (Wu 1964:214 [ref. 13503]). •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Synonym of Acheilognathus chankaensis (Dybowski 1872) -- (Lin in Chen et al. 1998:433 [ref. 23878], Zhang et al. 2016:67 [ref. 34477]). Current status: Synonym of Acheilognathus chankaensis (Dybowski 1872). Acheilognathidae. Habitat: freshwater.

atremius, Acanthorhodeus Jordan [D. S.] & Thompson [W. F.] 1914:227, Pl. 25 (fig. 3) [Memoirs of the Carnegie Museum v. 6 (no. 4); ref. 2543] Chikugo River at Kurume, Japan. Holotype: CAS-SU 22606. Paratypes: CM (1, whereabouts unknown), USNM 74786 (1). Type catalog: Böhlke 1953:29 [ref. 12291]. •Valid as Acanthorhodeus atremius Jordan & Thompson 1914 -- (Okada 1961:406 [ref. 6363]). •Synonym of Rhodeus sinensis Günther 1868 -- (Wu 1964:202 [ref. 13503]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Valid as Rhodeus atremius (Jordan & Thompson 1914) -- (Sawada in Masuda et al. 1984:55 [ref. 6441], Kawanabe & Mizuno 1989:365 [ref. 25598], Watanabe 1998:262 [ref. 23491], Nakabo 2000:260 [ref. 25086], Arai et al. 2001:275 [ref. 25545], Li et al. 2020:[14] [ref. 37372]). •Valid as Rhodeus atremius (Jordan & Thompson 1914), subspecies atremius (Jordan & Thompson 1914) -- (Matsuura et al. 2000:116 [ref. 26000], Nakabo 2002:260 [ref. 26001]). Current status: Valid as Rhodeus atremius (Jordan & Thompson 1914). Acheilognathidae. Distribution: Japan. Habitat: freshwater.

barbatulus, Achilognathus Günther [A.] 1873:248 [Annals and Magazine of Natural History (Series 4) v. 12 (no. 69) (art. 31); ref. 2004] Shanghai, China. Holotype (unique): BMNH 1873.7.30.85. Original genus should have been Acheilognathus. •Valid as Acanthorhodeus barbatulus (Günther 1873) -- (Wu 1964:214 [ref. 13503], Wang 1984:68 [ref. 22951]). •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Valid as Acheilognathus barbatulus Günther 1873 -- (Chen & Li in Chu & Chen 1989:131 [ref. 13584], Zheng et al. 1989:161 [ref. 21202], Ye in Pan et al. 1991:130 [ref. 23876], Ding 1994:182 [ref. 21531], Zhu 1995:46 [ref. 25213], Cheng & Zhou 1997:146 [ref. 26385], Kottelat 1998:55 [ref. 23436], Lin in Chen et al. 1998:425 [ref. 23878], Doi et al. 1999:307 [ref. 24230], Kottelat 2000:83 [ref. 25486], Kottelat 2001:14 [ref. 25482], Kottelat 2001:36 [ref. 25780], Wang et al. 2001:153 [ref. 26566], Yang et al. 2010:336 [ref. 30786], Yang et al. 2011:163 [ref. 31154], Nguyen et al. 2013:20 [ref. 32689], Kottelat 2013:70 [ref. 32989], Zhang et al. 2016:66 [ref. 34477]). Current status: Valid as Acheilognathus barbatulus Günther 1873. Acheilognathidae. Distribution: Asia. Habitat: freshwater.

barbatus, Acheilognathus Nichols [J. T.] 1926:6, Fig. 6 [American Museum Novitates No. 214; ref. 15600] Ningkwo (Suancheng), Anhwei Province, China. Holotype: AMNH 8449. Paratypes: AMNH 10773 (2). Non-types: FMNH 14713 (1). Type catalog: Ibarra & Stewart 1987:5 [ref. 12367]. •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Valid as Acheilognathus barbatus Nichols 1926 -- (Wu 1964:208 [ref. 13503], Wang 1984:65 [ref. 22951], Lin in Chen et al. 1998:423 [ref. 23878], Yang et al. 2011:163 [ref. 31154], Nguyen et al. 2013:20 [ref. 32689], Zhang et al. 2016:67 [ref. 34477]). Current status: Valid as Acheilognathus barbatus Nichols 1926. Acheilognathidae. Distribution: China. Habitat: freshwater.

bergi, Acanthorhodeus Mori [T.] 1928:63 [17] [Journal of the Chosen Natural History Society No. 6; ref. 15861] Manpochin, Korea. Holotype: whereabouts unknown. On p. 17 of separate. •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Synonym of Acheilognathus macropteus (Bleeker 1871) -- (Kottelat 2013:70 [ref. 32989]). Current status: Synonym of Acheilognathus macropterus (Bleeker 1871). Acheilognathidae. Habitat: freshwater.

binidentatus, Acheilognathus Li [G.-L.] in Wang et al. 2001:145, Figs. 91 A-B [The fauna of Hebei, China 2001; ref. 26566] XiQing, District of Tianjin, China, 38°50'N, 117°15'E. Holotype: TNZ [TNHM] 79808. Paratypes: TNHM 77803 (1), 79801-03 (1, 1, 1), 79804-05 (1, 1), 79806-07 (1, 1), 79809 (1), 79901 (1), •Valid as Acheilognathus binidentatus Li 2001 -- (Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus binidentatus Li 2001. Acheilognathidae. Distribution: Tianjin, China. Habitat: freshwater.

bleekeri, Paracheilognathus Berg [L. S.] 1907:162 [Annals and Magazine of Natural History (Series 7) v. 19 (no. 110) (art. 18); ref. 269] Chang Jiang [Yangtze River], China. Holotype (unique): not researched. Replacement name for Paracheilognathus imberbis Bleeker 1871, secondarily preoccupied by Acheilognathus imberbis Günther 1868. •Synonym of Acanthorhodeus chankaensis (Dybowski 1872) -- (Wu 1964:214 [ref. 13503]). •In genus Acheilognathus (Arai & Akai 1988:207 [ref. 6999]). •Synonym of Acheilognathus chankaensis (Dybowski 1872) -- (Lin in Chen et al. 1998:434 [ref. 23878]). Replacement Name. Current status: Synonym of Acheilognathus chankaensis (Dybowski 1872). Acheilognathidae. Habitat: freshwater.

brevianalis, Acheilognathus Fowler [H. W.] 1910:479, Fig. 2 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 62; ref. 1380] Lake Biwa at Matsubara, Omi, Japan. Holotype (unique): ANSP 28580. •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Synonym of Acheilognathus cyanostigma Jordan & Fowler 1903 -- (Okada 1961:394 [ref. 6363] as brevianals, Okamura & Machida 1987:103 [ref. 22279]). Current status: Synonym of Acheilognathus cyanostigma Jordan & Fowler 1903. Acheilognathidae. Habitat: freshwater.

brevicaudatus, Acheilognathus elongatus Chen [Y.-R.] & Li [Z.-Y.] 1987:62 [English p. 66], Fig. 2B [Zoological Research v. 8 (no. 1); ref. 8970] Yangzonhai Lake, flowing into the Nanpan River, Yunnan Province, China, 24°51'-24°58'N, 102°59'-103°02'E. Holotype: KIZ 843056. Paratypes: KIZ (27). Type catalog: Du et al. 2013:347 as breviacauatus. •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acheilognathus elongatus (Regan 1908), but a valid subspecies as described -- (Yang et al. 1990:69 [ref. 16186], Yang & Chen 1995:58 [ref. 23555], Zhu 1995:45 [ref. 25213], Lin in Chen et al. 1998:437 [ref. 23878], Zhang et al. 2016:67 [ref. 34477], Zhang et al. 2019:214 [ref. 36699]). •Valid as Acheilognathus brevicaudatus Chen & Li 1987 -- (Kottelat 1998:55 [ref. 23436], Yang et al. 2010:336 [ref. 30786], Yang et al. 2011:163 [ref. 31154], Yang et al. 2011:163 [ref. 31154], Nguyen et al. 2013:20 [ref. 32689]). Current status: Valid as Acheilognathus brevicaudatus Chen & Li 1987. Acheilognathidae. Distribution: Upper Yangtze River basin, Yunnan Province, China. Habitat: freshwater.

caspius, Rhodeus Esmaeili [H. R.], Sayyadzadeh [G.], Japoshvili [B.], Eagderi [S.], Abbasi [K.] & Mousavi-Sabet [H.] 2020:322, Figs. 2-11 [Zootaxa 4851 (no. 2); ref. 37759] Anzali Lagoon drainage of the Shakhraz (= Shakhazar or Siahdarvishan) River, Caspian Sea basin, Tolamshahr, Guilan province, Iran, 37°16'3.98"N, 49°21'53.82"E. Holotype: ZM-CBSU H1005. Paratypes: VMFC, ZM-CBSU. Plus additional non-type material. •Valid as Rhodeus caspius Esmaeili, Sayyadzadeh, Japoshvili, Eagderi, Abbasi & Mousavi-Sabet 2020. Current status: Valid as Rhodeus caspius Esmaeili, Sayyadzadeh, Japoshvili, Eagderi, Abbasi & Mousavi-Sabet 2020. Acheilognathidae. Distribution: Southern Caspian Sea basin, Iran; translocated to the Tigris River drainage and Urmia Lake basin. Habitat: freshwater.

changtingensis, Acheilognathus Yang [Q.], Zhu [Y.], Xiong [B.] & Liu [H.] 2011:158, Fig. 2 [Zoological Science v. 28 (no. 2); ref. 31154] Hanjiang River, Changting County, 25°85'N, 116°37'E, southeastern China. Holotype: IHASW 06027. Paratypes: IHASW 06028 (1), 060221(1), 060222 (1), 060223 (1), 060224 (1). •Valid as Acheilognathus changtingensis Yang, Zhu, Xiong & Liu 2011 -- (Zhang et al. 2016:67 [ref. 34477]). Current status: Valid as Acheilognathus changtingensis Yang, Zhu, Xiong & Liu 2011. Acheilognathidae. Distribution: Hanjiang River, Fujian Province, southeastern China. Habitat: freshwater.

chankaensis, Devario Dybowski [B. N.] 1872:212 [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 22; ref. 1170] Lake Khanka, southeastern Russia. No types known. Original as chankaënsis. •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Valid as Acanthorhodeus chankaensis (Dybowski 1872) -- (Berg 1949:819 [ref. 20661], Wu 1964:214 [ref. 13503], Wang 1984:70 [ref. 22951], Ye in Pan et al. 1991:135 [ref. 23876], Bogutskaya & Naseka 1996:31 [ref. 22798], Reshetnikov et al. 1997:734 [ref. 24702], Naseka 1998:79 [ref. 23590], Bogutskaya et al. 2001:44 [ref. 26178], Bogutskaya & Naseka 2004:42 [ref. 28183], Bogutskaya et al. 2008:314 [ref. 30085]). •Valid as Acheilognathus chankaensis (Dybowski 1872) -- (Chen & Li in Chu & Chen 1989:133 [ref. 13584], Zheng et al. 1989:162 [ref. 21202], Ding 1994:187 [ref. 21531], Zhu 1995:45 [ref. 25213], Cheng & Zhou 1997:149 [ref. 26385], Kottelat 1998:55 [ref. 23436], Lin in Chen et al. 1998:433 [ref. 23878], Doi et al. 1999:307 [ref. 24230], Wang et al. 2001:150 [ref. 26566], Yang et al. 2011:163 [ref. 31154], Parin et al. 2014:66 [ref. 33547], Zhang et al. 2016:67 [ref. 34477], Zhang et al. 2019:216 [ref. 36699]). Current status: Valid as Acheilognathus chankaensis (Dybowski 1872). Acheilognathidae. Distribution: Eastern Asia: Russia and China. Habitat: freshwater.

chii, Acheilognathus Miao [C.-P.] 1934:182, Fig. 33 [Contributions from the Biological Laboratory of the Science Society of China. (Zoological Series) v. 10 (no. 3); ref. 13903] Lanchie, Upper Tsien Tang River, Chinkiang, Jiangsu Province, China. Holotype: SSCN 13182. Spelled chi in title, chii elsewhere. •Synonym of Paracheilognathus himantegus (Günther 1868) -- (Wu 1964:210 [ref. 13503] as himategus, Lin in Chen et al. 1998:439 [ref. 23878]). •In genus Tanakia -- (Arai & Akai 1988:205 [ref. 6999]). •Synonym of Tanaka himantegus (Günther 1868) but a valid subspecies chii Miao 1934 -- (Chang et al. 2009:824 [ref. 31127]). •Valid as Tanakia chii (Miao 1934) -- (Zhang et al. 2016:69 [ref. 34477]). •Valid as Paratanakia chii (Miao 1934) -- (Li et al. 2017:84 [ref. 35680]). Current status: Valid as Paratanakia chii (Miao 1934). Acheilognathidae. Distribution: Taiwan and China. Habitat: freshwater.

colchicus, Rhodeus Bogutskaya [N. G.] & Komlev [A. M.] 2001:90, Fig. 3 [Proceedings of the Zoological Institute, Academy of Sciences, St.-Petersburg v. 287; ref. 26834] River Notanebi, West Transcaucasia, Georgia. Holotype: ZIN 15164a. Paratypes: ZIN 15164 (18), ZMMGU P-17589 (15). •Valid as Rhodeus colchicus Bogutskaya & Komlev 2001 -- (Bogutskaya et al. 2001:44 [ref. 26178], Bogutskaya & Naseka 2004:44 [ref. 28183], Ninua & Japoshvili 2008:165 [ref. 31251], Yang et al. 2010:333 [ref. 30786], Yang et al. 2011:158 [ref. 31154], Dyldin & Orlov 2016:663 [ref. 34853], Dyldin et al. 2020:105 [ref. 37569], Kuljanishvili et al. 2020:[4] [ref. 37535], Li et al. 2020:[14] [ref. 37372], Esmaeili et al. 2020:335 [ref. 37759]). Current status: Valid as Rhodeus colchicus Bogutskaya & Komlev 2001. Acheilognathidae. Distribution: Eurasia: west Transcaucasia, south and east of the Black Sea, Georgia. Habitat: freshwater.

coreanus, Achilognathus Steindachner [F.] 1892:366 [10] [133] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 59 (1. Abth.); ref. 14517] Seoul, Korea. Syntypes: (2) NMW? On p. 10 of separate; on p. 133 of abstract. The spelling coreanus is probably acceptable. Original genus should have been Acheilognathus. •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Valid as Acheilognathus koreanus Steindachner 1892 -- (Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus coreanus Steindachner 1892. Acheilognathidae. Distribution: Korea. Habitat: freshwater.

cyanorostris, Rhodeus Li [F.], Liao [T.-Y.] & Arai [R.] 2020:[3], Figs. 1-7, 15C [Journal of Vertebrate Biology v. 69 (no. 1); ref. 37372] Bai-Tiao River, tributary of the Yangtze River, Pidu District, Chengdu City, Sichuan Province, China. Holotype: SOU 1801001. Paratypes: DOS, NSMT, SOU. Plus additional non-type material. •Valid as Rhodeus cyanorostris Li, Liao & Arai 2020 -- (Li et al. 2020:337 [ref. 37574]). Current status: Valid as Rhodeus cyanorostris Li, Liao & Arai 2020. Acheilognathidae. Distribution: Bai-Tiao River drainage, Yangtze River basin, Sichuan Province, China. Habitat: freshwater.

cyanostigma, Acheilognathus Jordan [D. S.] & Fowler [H. W.] 1903:820, Fig. 2 [Proceedings of the United States National Museum v. 26 (no. 1334); ref. 2463] Lake Biwa at Matsubara, Omi, Japan. Holotype: CAS-SU 7724. Paratypes: CAS-SU 7143 (27). Additional material: CAS-SU 22563 (5). Type catalog: Böhlke 1953:29 [ref. 12291]. Holotype was bottled with two specimens thought to be paratypes after 1906 earthquake; these were removed and catalogued as SU 69111 and are probably not A. cyanostigma. •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Valid as Acheilognathus cyanostigma Jordan & Fowler 1903 -- (Okada 1961:394 [ref. 6363], Sawada in Masuda et al. 1984:55 [ref. 6441], Okamura & Machida 1987:103 [ref. 22279], Kawanabe & Mizuno 1989:372 [ref. 25598], Sato 1995:49 [ref. 25531], Watanabe 1998:262 [ref. 23491], Nakabo 2000:257 [ref. 25086], Matsuura et al. 2000:45 [ref. 26000], Nakabo 2002:257 [ref. 26001], Hagiwara 2003:35 [ref. 27188], Arai et al. 2007:24 [ref. 29106], Yang et al. 2010:336 [ref. 30786], Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus cyanostigma Jordan & Fowler 1903. Acheilognathidae. Distribution: Western Japan. Habitat: freshwater.

danubicus, Rhodeus sericeus amarus natio Holčík [J.] 1959:50 [Voprosy Ikhtiologii v. 13; ref. 23006] Not available, infrasubspecific; from Danube R. basin, Slovakia. •In the synonymy of Rhodeus sericeus (Pallas 1776) -- (Holčík 1999:2 [ref. 24847], Bogutskaya & Komlev 2001:81 [ref. 26834]). Infrasubspecific. Current status: Synonym of Rhodeus sericeus (Pallas 1776). Acheilognathidae.

dayeus, Acanthorhodeus Mai [D. Y.] 1978:187, Fig. 85 [Identification of the fresh-water fishes of North Viet Nam v. 1; ref. 13511] Dáy River, Hòa-Binh, northern Vietnam. Syntypes: DVZUT. •Synonym of Acanthorhodeus polyspinus Holčík 1972 -- (Kottelat 1998:55 [ref. 23436]). •Tentative synonym of Acheilognathus polyspinus (Holčík 1972) -- (Kottelat 2001:15 [ref. 25482]). Current status: Synonym of Acheilognathus polyspinus (Holčík 1972). Acheilognathidae. Habitat: freshwater.

deignani, Acanthorhodeus Smith [H. M.] 1945:219, Fig. 39 [Bulletin of the United States National Museum No. 188; ref. 4056] Huey Nam Puat, tributary of Mekong River at Ban Nam Puat, Laos. Holotype: USNM 107938. Paratypes: USNM 107892 (3). Type catalog: Monkolprasit 1969:3 [ref. 25088]. In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). See Kottelat 2001:13 [ref. 25482]. •Valid as Acheilognathus deignani (Smith 1945) -- (Doi 1997:3 [ref. 22832], Kottelat 1998:55 [ref. 23436], Kottelat 2001:36 [ref. 25780], Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus deignani (Smith 1945). Acheilognathidae. Distribution: Laos and Vietnam. Habitat: freshwater.

dicaeus, Acanthorhodeus Rutter [C. L.] 1897:58 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 49; ref. 12329] Swatow, Shantou, coast of southeastern China. Syntypes: CAS-SU 4997 (1), 69687 [ex CAS-SU 4997] (2). Type catalog: Böhlke 1953:29 [ref. 12291]. •Synonym of Acanthorhodeus macropterus Bleeker 1871 -- (Wu 1964:212 [ref. 13503], Ye in Pan et al. 1991:132 [ref. 23876], Yang et al. 2011:165 [ref. 31154]). •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Synonym of Acheilognathus macropterus (Bleeker 1871) -- (Lin in Chen et al. 1998:420 [ref. 23878], Kottelat 2013:70 [ref. 32989]). •Valid as Acanthorhodeus dicaeus Rutter 1897 -- (Yang et al. 2011:165 [ref. 31154]). Current status: Synonym of Acheilognathus macropterus (Bleeker 1871). Acheilognathidae. Distribution: Guangdong Province, China. Habitat: freshwater.

elongatoides, Acheilognathus Kottelat [M.] 2001:14 [Freshwater fishes of northern Vietnam. A preliminary check-list; ref. 25482] Thuong River, Lang Son, northern Vietnam. Syntypes: SVZUT. Replacement name for Pararhodeus elongatus Mai 1978, secondarily preoccupied in Acheilognathus by Acanthothodeus elongatus Regan 1908. •Valid as Acheilognathus elongatoides Kottelat 2001 -- (Yang et al. 2011:163 [ref. 31154], Kottelat 2013:70 [ref. 32989]). Replacement Name. Current status: Valid as Acheilognathus elongatoides Kottelat 2001. Acheilognathidae. Distribution: Vietnam. Habitat: freshwater.

elongatus, Acanthorhodeus Regan [C. T.] 1908:356 [Annals and Magazine of Natural History (Series 8) v. 2 (no. 10); ref. 14350] Yunnan Fu, China. Syntypes: (several) BMNH 1908.2.27.21-22 (2), MNHN 1923-0001 and 0002 (2), RMNH 10385 [ex BMNH] (1). Type catalog: Bertin & Estève 1948:76 [ref. 19574]. •Valid as Acanthorhodeus elongatus Regan 1908 -- (Wu 1964:217 [ref. 13503]). •In genus Acheilognathus -- (Arai & Akai 1988:207 [ref. 6999]). •Valid as Acheilognathus elongatus (Regan 1908) -- (Chen & Li in Chu & Chen 1989:134 [ref. 13584], Yang et al. 1990:69 [ref. 16186], Yang & Chen 1995:59 [ref. 23555], Zhu 1995:46 [ref. 25213], Kottelat 1998:55 [ref. 23436], Chen in Chen 1998:130 [ref. 23556], Lin in Chen et al. 1998:435 [ref. 23878], Yang et al. 2010:336 [ref. 30786], Yang et al. 2011:163 [ref. 31154], Zhang et al. 2016:67 [ref. 34477], Zhang et al. 2019:214 [ref. 36699]). Current status: Valid as Acheilognathus elongatus (Regan 1908). Acheilognathidae. Distribution: Yangtze River basin, China. Habitat: freshwater.

elongatus, Pararhodeus Mai [D. Y.] 1978:181, Fig. 82 [Identification of the fresh-water fishes of North Viet Nam v. 1; ref. 13511] Thuong River, Lang Son, northern Vietnam. Syntypes: DVZUT. Subjectively invalid; secondarily preoccupied in Acheilognathus by Acanthorhodeus elongatus Regan 1908, replaced by Acheilognathus elongatoides Kottelat 2001. •Synonym of Pararhodeus elongatus Mai 1978 -- (Kottelat 1998:54 [ref. 23436], Arai et al. 2001:281 [ref. 25545]). •Synonym of Acheilognathus elongatoides Kottelat 2001 -- (Kottelat 2013:70 [ref. 32989]). 2°Homonym. Current status: Synonym of Acheilognathus elongatoides Kottelat 2001. Acheilognathidae. Habitat: freshwater.

erythropterus, Acheilognathus tabira Arai [R.], Fujikawa [H.] & Nagata [Y.] 2007:11, Figs. 3B-D, 8 [Bulletin of the National Museum of Nature and Science (Ser. A) Suppement No. 1; ref. 29106] Lake Kasumigaura, Hama, Tamatsukuri, Ibaraki Prefecture, Japan. Holotype: NSMT-P 72656. Paratypes: NSMT-P 74702 (49), 74703 (35), 27043 (2); ZUMT 61512 (2). •Synonym of Acheilognathus tabira Jordan & Thompson 1914, but a valid subspecies erythropterus Arai, Fujikawa & Nagata 2007 as described. Current status: Synonym of Acheilognathus tabira Jordan & Thompson 1914. Acheilognathidae. Distribution: Pacific side of eastern Honshu [subspecies]. Habitat: freshwater.

fangi, Pararhodeus Miao [C.-P.] 1934:180, Fig. 31 [Contributions from the Biological Laboratory of the Science Society of China. (Zoological Series) v. 10 (no. 3); ref. 13903] Chinkiang, China. Holotype: SSCN 3216. •Valid as Pararhodeus fangi Miao 1934 -- (Wu 1964:206 [ref. 13503]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Valid as Rhodeus fangi (Miao 1934) -- (Zhu 1995:47 [ref. 25213], Lin in Chen et al. 1998:451 [ref. 23878], Arai et al. 2001:280 [ref. 25545], Naseka & Bogutskaya 2004:282 [ref. 27838], Bogutskaya & Naseka 2004:44 [ref. 28183], Chen et al. 2005:90 [ref. 28516], Li & Arai 2010:303 [ref. 31188], Li & Arai 2011:303 [ref. 31188], Li & Arai 2014:175 [ref. 33267], Zhang et al. 2016:68 [ref. 34477], Li et al. 2017:84 [ref. 35680], Dyldin et al. 2020:98 [ref. 37569], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:337 [ref. 37574]). Current status: Valid as Rhodeus fangi (Miao 1934). Acheilognathidae. Distribution: East Asia: Yangtze River basin (China). Habitat: freshwater.

fasciodorsalis, Acheilognathus Nguyen [V. H.] in Nguyen & Ngô 2001:243, Fig. 114 [Freshwater fishes of Viet Nam. Volume I. Family Cyprinidae v. 1; ref. 28301] Sông Bàng, Cao Bàng, Cao Bàng Province, Viêt Nam. Holotype: VNCNTTS I: H.01.47.08.01. Paratypes: VNCNTTS I: H.01.47.08.02 (1). •Valid as Acheilognathus fasciodorsalis Nguyen 2001 -- (Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus fasciodorsalis Nguyen 2001. Acheilognathidae. Distribution: Bàng River, Cao Bàng Province, Vietnam. Habitat: freshwater.

flaviventris, Rhodeus Li [F.], Arai [R.] & Liao [T.-Y.] 2020:330, Figs. 1-8 [Zootaxa 4790 (no. 2); ref. 37574] Le-An River, a tributary flowing into Poyang Lake, lower Yangtze River basin, Xitou Township, Wuyuan County, Jiangxi Province, China. Holotype: SOU 1909001. Paratypes: DOS, NSMT, SOU. Plus additional non-type material. •Valid as Rhodeus flaviventris Li, Arai & Liao 2020. Current status: Valid as Rhodeus flaviventris Li, Arai & Liao 2020. Acheilognathidae. Distribution: Le-An River drainage, lower Yangtze River basin, Jiangxi, China. Habitat: freshwater.

fowleri, Acanthorhodeus Holčík [J.] & Nalbant [T. T.] 1964:3, Figs. 1-3 [Annotationes Zoologicae et Botanicae No. 2; ref. 2282] Chang Jiang [Yangtze River] at Uhan [Wuhan], Hupeh Province, China. Holotype (unique): SNMB Ry 473. Type catalog: Blahák 1991:204 [ref. 20023]. •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acanthorhodeus gracilis Nichols 1926 -- (Blahák 1991:204 [ref. 20023], Lin in Chen et al. 1998:418 [ref. 23878]). Current status: Synonym of Acheilognathus gracilis Nichols 1926. Acheilognathidae. Habitat: freshwater.

genitalis, Rhodeus Walecki [A.] 1863:364 [Wiadomosci Z Nauk [Science News] v. 4; ref. 22954] Apparently an unneeded replacement name for Cyprinus amarus Bloch 1782. •Synonym of Rhodeus amarus (Bloch 1782) -- (Kottelat 1997:75 [ref. 22952]). Current status: Synonym of Rhodeus amarus (Bloch 1782). Acheilognathidae. Habitat: freshwater, brackish.

gracilis, Acanthorhodeus Regan [C. T.] 1908:60, Pl. 2 (fig. 1) [Proceedings of the Zoological Society of London 1908 (pt 1) (art. 3) (for 4 Feb. 1908); ref. 14334] Chong-ju, South Korea. Lectotype: BMNH 1907.12.10.51. Paralectotypes: BMNH 1907.12.10.52 (1). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acanthorhodeus gracilis Regan 1908 -- (Kim 1997:190 [ref. 24032], Kim & Park 2002:92 [ref. 25913], Choi et al. 2003:156, 504 [ref. 26218]). •Synonym of Acanthorhodeus chankaensis (Dybowski 1872) -- (Bogutskaya & Naseka 2004:42 [ref. 28183]). Current status: Synonym of Acheilognathus chankaensis (Dybowski 1872). Acheilognathidae. Distribution: South Korea. Habitat: freshwater.

gracilis, Acheilognathus Nichols [J. T.] 1926:5, Fig. 5 [American Museum Novitates No. 214; ref. 15600] Huping, Tungting Lake, Hunan Province, China. Holotype: AMNH 8448. Paratypes: AMNH 10923 (3), ?FMNH 14668 (1). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus gracilis Nichols 1926 -- (Wu 1964:207 [ref. 13503], Ding 1994:185 [ref. 21531], Zhu 1995:46 [ref. 25213], Cheng & Zhou 1997:148 [ref. 26385], Lin in Chen et al. 1998:417 [ref. 23878], Doi et al. 1999:307 [ref. 24230], Kim & Lee 2001:46 [ref. 25334], Yang et al. 2011:163 [ref. 31154], Zhang et al. 2016:67 [ref. 34477]). Current status: Valid as Acheilognathus gracilis Nichols 1926. Acheilognathidae. Distribution: China. Habitat: freshwater.

grahami, Acanthorhodeus Nichols [J. T.] 1918:18 [Proceedings of the Biological Society of Washington v. 31; ref. 3171] Yunnan-fu, Yunnan Province, China. Holotype: AMNH 7029. Paratypes: AMNH 7027 (7). •Synonym of Acanthorhodeus elongatus Regan 1908 -- (Wu 1964:217 [ref. 13503]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acheilognathus elongatus (Regan 1908) -- (Lin in Chen et al. 1998:435 [ref. 23878]). Current status: Synonym of Acheilognathus elongatus (Regan 1908). Acheilognathidae. Habitat: freshwater.

guichenoti, Acanthorhodeus Bleeker [P.] 1871:41, Pl. 13 (fig. 2) [Verhandelingen der Koninklijke Akademie van Wetenschappen (Amsterdam) v. 12 (art. 2); ref. 6421] Chang Jiang [Yangtze River], China. Holotype (unique): MNHN 0000-5043. Also in Bleeker 1871:73 [ref. 17074]. •Synonym of Acanthorhodeus macropterus Bleeker 1871 -- (Wu 1964:212 [ref. 13503], Mai 1978:177 [ref. 13511]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acheilognathus macropterus (Bleeker 1871) -- (Lin in Chen et al. 1998:419 [ref. 23878]). •Valid as Acanthorhodeus guichenoti Bleeker 1871 -- (Yang et al. 2011:165 [ref. 31154], Kottelat 2013:70 [ref. 32989]). Current status: Synonym of Acheilognathus macropterus (Bleeker 1871). Acheilognathidae. Habitat: freshwater.

hainanensis, Pseudoperilampus Nichols [J. T.] & Pope [C. H.] 1927:379, Fig. 42 [Bulletin of the American Museum of Natural History v. 54 (art. 2); ref. 5747] Nodoa, Hainan Island, China. Holotype: AMNH 8386. Paratypes: ?MCZ 32663 [ex AMNH] (2). •Valid as Pseudoperilampus hainanensis Nichols & Pope 1927 -- (Wu 1964:206 [ref. 13503], Mai 1978:179 [ref. 13511], Ye in Pan et al. 1991:129 [ref. 23876]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus spinalis Oshima 1926 -- (Arai et al. 1990:143 [ref. 16800], Kottelat 1998:53 [ref. 23436], Lin in Chen et al. 1998:448 [ref. 23878], Kottelat 2013:157 [ref. 32989], Zhang et al. 2016:69 [ref. 34477]). Current status: Synonym of Rhodeus spinalis Oshima 1926. Acheilognathidae. Habitat: freshwater.

haradai, Rhodeus Arai [R.], Suzuki [N.] & Shen [S.-C.] 1990:151, Fig. 8 [Bulletin of the National Science Museum Series A (Zoology) v. 16 (no. 3); ref. 16800] Longtang, Hainan Island, China. Holotype: NTUM 7600. Paratypes: NSMT-P 30930-31 (1, 1), NTUM 2203 (10). •Questionably a synonym of Rhodeus spinalis Oshima 1926 -- (Lin in Chen et al. 1998:448 [ref. 23878]). •Valid as Rhodeus haradai Arai, Suzuki & Shen 1990 -- (Kottelat 1998:53 [ref. 23436], Arai et al. 2001:280 [ref. 25545], Li & Arai 2010:303 [ref. 31188], Li & Arai 2011:303 [ref. 31188], Kottelat 2013:157 [ref. 32989], Li & Arai 2014:175 [ref. 33267], Zhang et al. 2016:68 [ref. 34477], Li et al. 2017:85 [ref. 35680], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:337 [ref. 37574]). Current status: Valid as Rhodeus haradai Arai, Suzuki & Shen 1990. Acheilognathidae. Distribution: Hainan Island, China. Habitat: freshwater.

himantegus, Achilognathus Günther [A.] 1868:277 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Taiwan. Syntypes: (at least 4) BMNH 1865.5.2.54-58 (5). Original genus should have been Acheilognathus. •Valid as Paracheilognathus himantegus (Günther 1868) -- (Wu 1964:210 [ref. 13503] as himategus, Zhu 1995:45 [ref. 25213] as himantagus, Cheng & Zhou 1997:151 [ref. 26385], Lin in Chen et al. 1998:439 [ref. 23878]). •In genus Tanakia -- (Arai & Akai 1988:205 [ref. 6999]). •Valid as Tanakia himantegus subspecies himantegus (Günther 1868) -- (Chang et al. 2009:824 [ref. 31127]). •Valid as Tanakia himantegus (Günther 1868) -- (Arai et al. 1995 [ref. 22109], Ho & Shao 2011:25 [ref. 31432], Zhang et al. 2016:69 [ref. 34477]). •Valid as Paratanakia himantegus (Günther 1868) -- (Chang et al. 2014:191 [ref. 33502], Li et al. 2017:84 [ref. 35680]). Current status: Valid as Paratanakia himantegus (Günther 1868). Acheilognathidae. Distribution: Taiwan and China. Habitat: freshwater.

hondae, Pseudoperilampus Jordan [D. S.] & Metz [C. W.] 1913:20, Pl. 2 (fig. 3) [Memoirs of the Carnegie Museum v. 6 (no. 1); ref. 2490] Suigen, Korea. Holotype (unique): FMNH 55803 [ex CM 4566]. Type catalog: Henn 1928:73 [ref. 12305], Ibarra & Stewart 1987:71 [ref. 12367]. •Valid as Rhodeus hondae (Jordan & Metz 1913) -- (Kim 1997:171 [ref. 24032], Choi et al. 2003:158, 501 [ref. 26218]). •Valid as Acheilognathus hondae (Jordan & Metz 1913) -- (Arai et al. 2001:281 [ref. 25545], Yang et al. 2011:163 [ref. 31154], Li et al. 2017:83 [ref. 35680]). Current status: Valid as Rhodeus hondae (Jordan & Metz 1913). Acheilognathidae. Distribution: Korea. Habitat: freshwater.

hwanghoensis, Rhodeus Mori [T.] 1928:68 [Japanese Journal of Zoology v. 2 (no. 1) (art. 4); ref. 15862] Tsi-nan, Shantung, northern China. Holotype: whereabouts unknown. Additional material: (2). •Synonym of Rhodeus sinensis Günther 1868 -- (Wu 1964:202 [ref. 13503]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus ocellatus (Kner 1866) -- (Lin in Chen et al. 1998:445 [ref. 23878], Kottelat 2013:157 [ref. 32989]). Current status: Synonym of Rhodeus ocellatus (Kner 1866). Acheilognathidae. Habitat: freshwater.

hypselonotus, Acanthorhodeus Bleeker [P.] 1871:43, Pl. 11 (fig. 2) [Verhandelingen der Koninklijke Akademie van Wetenschappen (Amsterdam) v. 12 (art. 2); ref. 6421] Chang Jiang [Yangtze River], China. Holotype (unique): MNHN 0000-5934 (poor condition). Type catalog: Bertin & Estève 1948:76 [ref. 19574]. Also in Bleeker 1871:71 [ref. 17074]. •Valid as Acanthorhodeus hypselonotus Bleeker 1871 -- (Wu 1964:218 [ref. 13503]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus hypselonotus (Bleeker 1871) -- (Ding 1994:184 [ref. 21531], Zhu 1995:45 [ref. 25213], Lin in Chen et al. 1998:432 [ref. 23878], Doi et al. 1999:307 [ref. 24230], Yang et al. 2011:163 [ref. 31154], Zhang et al. 2016:67 [ref. 34477]). Current status: Valid as Acheilognathus hypselonotus (Bleeker 1871). Acheilognathidae. Distribution: China. Habitat: freshwater.

imberbis, Achilognathus Günther [A.] 1868:278 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] China. Holotype (unique): BMNH 1858.10.19.148. Original genus should have been Acheilognathus. •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Paracheilognathus imberbis (Günther 1868) -- (Wu 1964:209 [ref. 13503], Ding 1994:188 [ref. 21531], Zhu 1995:45 [ref. 25213], Cheng & Zhou 1997:150 [ref. 26385], Lin in Chen et al. 1998:442 [ref. 23878], Wang et al. 2001:144 [ref. 26566]). •Valid as Acheilognathus imberbis Günther 1868 -- (Doi et al. 1999:307 [ref. 24230], Yang et al. 2011:163 [ref. 31154], Zhang et al. 2016:67 [ref. 34477]). Current status: Valid as Acheilognathus imberbis Günther 1868. Acheilognathidae. Distribution: China. Habitat: freshwater.

imberbis, Parachilognathus Bleeker [P.] 1871:37, Pl. 4 (fig. 1) [Verhandelingen der Koninklijke Akademie van Wetenschappen (Amsterdam) v. 12 (art. 2); ref. 6421] Chang Jiang [Yangtze River], China. Holotype (unique): not researched. Original genus should have been Paracheilognathus. Regarded as an original description by some; Günther's Achilognathus imberbis included with question by Bleeker (p. 38) as the same species; probably can consider imberbis Bleeker as a subsequent identification and not an original description. Subjectively invalid; secondarily preoccupied by Acheilognathus imberbis Günther 1868, replaced by Paracheilognathus bleekeri Berg 1907. •Synonym of Acheilognathus chankaensis (Dybowski 1872) -- (Lin in Chen et al. 1998:434 [ref. 23878], Yang et al. 2010:336 [ref. 30786]). 2°Homonym. Current status: Synonym of Acheilognathus chankaensis (Dybowski 1872). Acheilognathidae. Habitat: freshwater.

imfasciodorsalis, Acheilognathus Nguyen [V. H.] in Nguyen & Ngô 2001:242, Fig. 113 [Freshwater fishes of Viet Nam. Volume I. Family Cyprinidae v. 1; ref. 28301] Cao Bàng, Cao Bàng Province, Viêt Nam. Holotype: VNCNTTS I: H.01.47.07.01. Paratypes: VNCNTTS I: H.01.47.07.02 (1). •Valid as Acheilognathus imfasciodorsalis Nguyen 2001 -- (Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus imfasciodorsalis Nguyen 2001. Acheilognathidae. Distribution: Bàng River, Cao Bàng Province, Vietnam. Habitat: freshwater.

intermedia, Capoeta Temminck [C. J.] & Schlegel [H.] 1846:203, Pl. 100 (figs. 4, 4a-b) [Fauna Japonica Parts 10-14; ref. 4374] Near Nagasaki, Japan. Lectotype: RMNH 2500. Paralectotypes: RMNH 32441 [ex 2500] (1). Lectotype selected by Boeseman 1947:158 [ref. 12876]. •Synonym of Acheilognathus lanceolatus (Temminck & Schlegel 1846) -- (Okada 1961:384 [ref. 6363]). •In genus Tanakia -- (Arai & Akai 1988:205 [ref. 6999]). •Synonym of Tanakia lanceolata (Temminck & Schlegel 1846). •Valid as Acheilognathus intermedia (Temminck & Schlegel 1846) -- (Yang et al. 2011:163 [ref. 31154] as imtermebia). Current status: Valid as Acheilognathus intermedia (Temminck & Schlegel 1846). Acheilognathidae. Distribution: Japan. Habitat: freshwater.

jeholicus, Acanthorhodeus Mori [T.] 1934:33, Pl. 1 (figs. 1-2) [Report of the first scientific expedition to Manchoukuo, Tokyo, Sec. 5, Zoology Pt 1; ref. 3042] Summer Palace, Jehol, China. Syntypes: (13) NSMT. •Synonym of Acanthorhodeus macropterus Bleeker 1871 -- (Wu 1964:212 [ref. 13503]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acheilognathus macropterus (Bleeker 1871) -- (Lin in Chen et al. 1998:420 [ref. 23878], Kottelat 2013:70 [ref. 32989]). Current status: Synonym of Acheilognathus macropterus (Bleeker 1871). Acheilognathidae. Habitat: freshwater.

jordani, Acheilognathus tabira Arai [R.], Fujikawa [H.] & Nagata [Y.] 2007:17, Figs. 4A-B, 5C-E, 10 [Bulletin of the National Museum of Nature and Science (Ser. A) Suppement No. 1; ref. 29106] Lake Kibagata, Komatsu, Ishikawa Prefecture, Japan. Holotype: NSMT-P 72658. Paratypes: at NSMT, OKU, ZMUT. Plus non-type specimens. •Synonym of Acheilognathus tabira Jordan & Thompson 1914, but a valid subspecies jordani Arai, Fujikawa & Nagata 2007 as described -- (Ito et al. 2021:47 [ref. 38540]). Current status: Synonym of Acheilognathus tabira Jordan & Thompson 1914. Acheilognathidae. Distribution: East Asia: Japan Sea side of western Honshu [subspecies]; introduced in Mie Prefecture (Japan). Habitat: freshwater.

koreensis, Acheilognathus Kim [I.-S.] & Kim [C. H.] 1990:47, Figs. 1-2 [Korean Journal of Ichthyology v. 2 (no. 1); ref. 20304] Chon-chon stream, tributary of the Kum River at Chonchon-myon, Changsu-gun, Chollabuk-do, Korea. Holotype: CUBK 15546. Paratypes: CUBK 15547-61 (15). •Valid as Tanakia koreensis (Kim & Kim 1990) -- (Chang et al. 2009:824 [ref. 31127]). •Valid as Acheilognathus koreensis Kim & Kim 1990 -- (Kim 1997:178 [ref. 24032], Kim & Yang 1998:29 [ref. 24125], Kim & Lee 2001:46 [ref. 25334], Kim & Park 2002:78 [ref. 25913], Choi et al. 2003:156, 504 [ref. 26218], Yang et al. 2011:163 [ref. 31154], Lim et al. 2014:67 [ref. 33508]). Current status: Valid as Tanakia koreensis (Kim & Kim 1990). Acheilognathidae. Distribution: South Korea. Habitat: freshwater.

kristinae, Acanthorhodeus Holčík [J.] 1971:270, Figs. 1-2 [Věstnik Československé společnosti zoologické v. 35 (no. 4); ref. 7682] Szechuan Province, China. Holotype: USNM 87600. Paratypes: USNM 206045 (1). Genus appeared as Acenthorhodeus in title. •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Questionably a synonym of Rhodeus fangi (Miao 1934) -- (Lin in Chen et al. 1998:451 [ref. 23878]). •Possibly not in Rhodeus -- (Li et al. 2017:83 [ref. 35680]). Acheilognathidae. Habitat: freshwater.

kurumeus, Rhodeus Jordan [D. S.] & Thompson [W. F.] 1914:229, Pl. 26 (fig. 2) [Memoirs of the Carnegie Museum v. 6 (no. 4); ref. 2543] Chikugo River at Kurume, Kiusiu, Japan. Holotype: CAS-SU 22605. Paratypes: FMNH 57074 (1), ?CAS-SU 22644 (18), USNM 74785 (1). Additional original material: CAS-SU 22571 (1), 22576 (22), 22580 (1). Type catalog: Böhlke 1953:38 [ref. 12291], Ibarra & Stewart 1987:75 [ref. 12367]. •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus ocellatus (Kner 1866), but a valid subspecies kurumeus Jordan & Thompson 1914 -- (Kimura & Nagata 1992:425 [ref. 19826], Watanabe 1998:262 [ref. 23491], Nakabo 2000:259 [ref. 25086], Matsuura et al. 2000:118 [ref. 26000], Arai et al. 2001:281 [ref. 25545], Kawamura et al. 2001:369 [ref. 25794], Nakabo 2002:258 [ref. 26001], Li et al. 2020:[14] [ref. 37372]). •Synonym of Rhodeus ocellatus (Kner 1866) -- (Wu 1964:203 [ref. 13503], Kottelat 2013:157 [ref. 32989], Zhang et al. 2016:69 [ref. 34477]). Current status: Synonym of Rhodeus ocellatus (Kner 1866). Acheilognathidae. Distribution: Japan and ? China (if valid). Habitat: freshwater.

kyphus, Pararhodeus Mai [D. Y.] 1978:182, Fig. 83 [Identification of the fresh-water fishes of North Viet Nam v. 1; ref. 13511] Creeks in northern Vietnam. Syntypes: DVZUT. •Valid as Parathodeus kyphus Mai 1978 -- (Kottelat 1998:54 [ref. 23436], Arai et al. 2001:281 [ref. 25545]). •Valid as Acheilognathus kyphus (Mai 1978) -- (Kottelat 2001:15 [ref. 25482], Yang et al. 2011:163 [ref. 31154], Kottelat 2013:70 [ref. 32989]). Current status: Valid as Acheilognathus kyphus (Mai 1978). Acheilognathidae. Distribution: Northern Vietnam. Habitat: freshwater.

lamus, Acanthorhodeus Nguyen [T. T.], Nguyen [X. K.] & Le [V. T.] 1999 [ ref. 25485] •A nomen nudum -- see Kottelat 2001:15 [ref. 25482]. Nomen Nudum. Acheilognathidae.

lanceolata, Capoeta Temminck [C. J.] & Schlegel [H.] 1846:202, Pl. 100 (figs. 3, 3a-b) [Fauna Japonica Parts 10-14; ref. 4374] Streams about Nagasaki, Japan. Holotype (unique): RMNH 2501. Type information: Boeseman 1947:157 [ref. 12876]. •Valid as Acheilognathus lanceolatus (Temminck & Schlegel 1846) -- (Okada 1961:384 [ref. 6363], Sawada in Masuda et al. 1984:54 [ref. 6441], Kim 1997:173 [ref. 24032], Kim & Yang 1998:29 [ref. 24125], Kim & Park 2002:72 [ref. 25913], Choi et al. 2003:156, 503 [ref. 26218]). •In genus Tanakia -- (Arai & Akai 1988:205 [ref. 6999]). •Valid as Tanakia lanceolata (Temminck & Schlegel 1846) -- (Kawanabe & Mizuno 1989:354 [ref. 25598], Arai et al. 1995:196 [ref. 22109], Sato 1995:48 [ref. 25531], Watanabe 1998:262 [ref. 23491], Sato et al. 1998:30 [ref. 24972], Nakabo 2000:256 [ref. 25086], Matsuura et al. 2000:129 [ref. 26000], Nakabo 2002:256 [ref. 26001], Chang et al. 2009:824 [ref. 31127], Lim et al. 2014:67 [ref. 33508], Li et al. 2017:85 [ref. 35680]). Current status: Valid as Tanakia lanceolata (Temminck & Schlegel 1846). Acheilognathidae. Distribution: Japan and Korea. Habitat: freshwater.

lanchiensis, Acanthorhodeus Herre [A. W. C. T.] & Lin [S.-Y.] 1936:17, Fig. 5 [Bulletin of the Chekiang Provincial Fisheries Experiment Station, Tinghai, Chekiang, China v. 2 (no. 7); ref. 10995] Pool in rapids at Lanchie, upper Tsien Tang River, Chekiang Province, China. Syntypes: (4) CAS-SU 32555 (1). Type catalog: Böhlke 1953:29 [ref. 12291]. •Synonym of Paracheilognathus himantegus (Günther 1868) -- (Wu 1964:210 [ref. 13503] as himategus, Lin in Chen et al. 1998:439 [ref. 23878]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus lanchiensis (Herre & Lin 1936) -- (Yang et al. 2010:336 [ref. 30786]). Current status: Valid as Acheilognathus lanchiensis (Herre & Lin 1936). Acheilognathidae. Distribution: Upper Tsien Tang River, Chekiang Province, China. Habitat: freshwater.

laoensis, Rhodeus Kottelat [M.], Doi [A.] & Musikasinthorn [P.] in Kottelat 1998:52, Fig. 81 [Ichthyological Exploration of Freshwaters v. 9 (no. 1); ref. 23436] Nam Theun, immediately upriver of confluence with Nam Ong, 17°43'N, 105°16'20"E, Laos. Holotype: ZRC 41786. Paratypes: CMK 12564 (3), 12637 (8); DFV uncat. (1), DFV uncat. (1); ZRC 41787 (1). •Valid as Rhodeus laoensis Kottelat, Doi & Musikasinthorn 1998 -- (Arai et al. 2001:281 [ref. 25545], Kottelat 2001:79 [ref. 25780], Kottelat 2015:157 [ref. 32989], Kottelat 2015:43 [ref. 33971], Li et al. 2020:[14] [ref. 37372]). Current status: Valid as Rhodeus laoensis Kottelat, Doi & Musikasinthorn 1998. Acheilognathidae. Distribution: Nam Theun, Laos. Habitat: freshwater.

latimarginata, Tanakia Kim [D.], Jeon [H.-B.] & Suk [H. Y.] 2014:63, Figs. 3-5 (a, c, e) [Ichthyological Exploration of Freshwaters v. 25 (no. 1); ref. 33508] Gyeongsangbuk Province, Yeongcheon County, Jogyo-dong, Keumho River, 35°57'55"N, 128°57'32"E, South Korea. Holotype: NIBR P0000023407. Paratypes: NIBR P0000023408 (10), -23409 (8); SLIC D4 (6), D5 (2), D6 (4), D7 (3), D8 (3), D9 (3). •Valid as Tanakia latimarginata Kim, Jeon & Suk 2014. Current status: Valid as Tanakia latimarginata Kim, Jeon & Suk 2014. Acheilognathidae. Distribution: Dakdong River drainage, South Korea. Habitat: freshwater.

lighti, Pseudoperilampus Wu [H.-W.] 1931:25, Fig. 4 [Contributions from the Biological Laboratory of the Science Society of China. (Zoological Series) v. 7 (no. 1); ref. 16348] Fukien, China. Holotype (unique): whereabouts unknown. •Synonym of Rhodeus ocellatus (Kner 1867) -- (Okada 1961:401 [ref. 6363]). •Valid as Pseudoperilampus lighti Wu 1931 -- (Wu 1964:205 [ref. 13503] as light, Wang 1984:64 [ref. 22951], Jin in Kuang et al. 1986:64 [ref. 26639], Ye in Pan et al. 1991:128 [ref. 23876]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus sinensis Günther 1868 -- (Akai & Arai 1998:105 [ref. 23354], Zhang et al. 2016:69 [ref. 34477]). •Valid as Rhodeus lighti (Wu 1931) -- (Zheng et al. 1989:158 [ref. 21202], Ding 1994:176 [ref. 21531], Zhu 1995:47 [ref. 25213], Cheng & Zhou 1997:142 [ref. 26385], Reshetnikov et al. 1997:740 [ref. 24702], Kottelat 1998:55 [ref. 23436], Naseka 1998:80 [ref. 23590], Lin in Chen et al. 1998:449 [ref. 23878], Tang et al. 2001:12 [ref. 25373], Wang et al. 2001:156 [ref. 26566], Naseka & Bogutskaya 2004:282 [ref. 27838], Bogutskaya & Naseka 2004:43 [ref. 28183], Kottelat 2013:157 [ref. 32989]). Current status: Valid as Rhodeus lighti (Wu 1931). Acheilognathidae. Distribution: Former USSR and Asia. Habitat: freshwater.

limbata, Capoeta Temminck [C. J.] & Schlegel [H.] 1846:203, Pl. 100 (figs. 5, 5a-b) [Fauna Japonica Parts 10-14; ref. 4374] Near Nagasaki, Japan. Lectotype: RMNH 2497. Paralectotypes: RMNH 26608 [ex RMNH 2497] (3). Lectotype selected by Boeseman 1947:158 [ref. 12876]. •Valid as Acheilognathus limbatus (Temminck & Schlegel 1846) -- (Okada 1961:388 [ref. 6363], Sawada in Masuda et al. 1984:54 [ref. 6441]). •Valid as Tanakia limbata (Temminck & Schlegel 1846) -- (Arai & Akai 1988:200 [ref. 6999], Kawanabe & Mizuno 1989:356 [ref. 25598], Watanabe 1998:262 [ref. 23491], Nakabo 2000:256 [ref. 25086], Matsuura et al. 2000:133 [ref. 26000], Nakabo 2002:256 [ref. 26001], Chang et al. 2009:824 [ref. 31127], Li et al. 2017:85 [ref. 35680]). Current status: Valid as Tanakia limbata (Temminck & Schlegel 1846). Acheilognathidae. Distribution: Japan. Habitat: freshwater.

longibarbatus, Acanthorhodeus Mai [D. Y.] 1978:186, Fig. 84 [Identification of the fresh-water fishes of North Viet Nam v. 1; ref. 13511] Na-Ri River and Cao Bang River, northern Vietnam. Syntypes: DVZUT. •Valid as Acanthorhodeus longibarbatus Mai 1978 -- (Kottelat 1998:54 [ref. 23436]). •Valid as Acheilognathus longibarbatus (Mai 1978) -- (Kottelat 2001:15 [ref. 25482], Kottelat 2001:36 [ref. 25780], Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus longibarbatus (Mai 1978). Acheilognathidae. Distribution: Vietnam. Habitat: freshwater.

longipinnis, Achilognathus Regan [C. T.] 1905:364 [Annals and Magazine of Natural History (Series 7) v. 16 (no. 93) (art. 42); ref. 15382] Yamasabu River, Lake Biwa, Japan. Syntypes: BMNH 1905.6.7.57-60 (4). Original genus should have been Acheilognathus. •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus longipinnis Regan 1905 -- (Okada 1961:370 [ref. 6363], Sawada in Masuda et al. 1984:55 [ref. 6441], Kawanabe & Mizuno 1989:370 [ref. 25598], Nakabo 2000:257 [ref. 25086], Matsuura et al. 2000:46 [ref. 26000], Nakabo 2002:257 [ref. 26001], Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus longipinnis Regan 1905. Acheilognathidae. Distribution: Japan. Habitat: freshwater.

longispinnis, Acanthorhodeus Oshima [M.] 1926:14 [Annotationes Zoologicae Japonenses v. 11 (no. 1); ref. 3314] Kachek River near Kachek, Hainan Island, China. Syntypes: (7, one described). •Synonym of Acanthorhodeus macropterus Bleeker 1871 -- (Wu 1964:212 [ref. 13503], Jin in Kuang et al. 1986:67 [ref. 26639], Ye in Pan et al. 1991:130 [ref. 23876]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acheilognathus macropterus (Bleeker 1871) -- (Lin in Chen et al. 1998:420 [ref. 23878], Kottelat 2013:70 [ref. 32989]). •Valid as Acheilognathus longispinnis (Oshima 1926) -- (Watanabe 1998:262 [ref. 23491], Arai et al. 2007:24 [ref. 29106]). Current status: Valid as Acheilognathus longispinnis (Oshima 1926). Acheilognathidae. Distribution: China. Habitat: freshwater.

luchowensis, Acheilognathus gracilis Wu [H.-W.] 1930:79, Fig. 6 [Sinensia v. 1 (no. 6); ref. 16240] Fu-tsi-Chang, Luchow, upper Chang Jiang [Yangtze River] valley, Szechwan, China. Holotype: not researched. •Synonym of Acheilognathus gracilis Nichols 1926 -- (Wu 1964:207 [ref. 13503], Lin in Chen et al. 1998:418 [ref. 23878], Zhang et al. 2016:67 [ref. 34477]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). Current status: Synonym of Acheilognathus gracilis Nichols 1926. Acheilognathidae. Habitat: freshwater.

lucinae, Rhodeus Walecki [A.] 1863:364 [Wiadomosci Z Nauk [Science News] v. 4; ref. 22954] Apparently an unneeded replacement name for Cyprinus amarus Bloch 1782. •Synonym of Rhodeus amarus (Bloch 1782) -- (Kottelat 1997:75 [ref. 22952]). Current status: Synonym of Rhodeus amarus (Bloch 1782). Acheilognathidae. Habitat: freshwater, brackish.

macrodorsalis, Acheilognathus Hsia [W. P.] 1949:200, Fig. [Contributions from the Institute of Zoology, National Academy of Peiping v. 5 (no. 5); ref. 20561] Paiyang Tien, Hopei Province, China. Holotype: ASZIB [now NSMC] 43644. Paratypes: ASZIB [now NSMC] 42252 and 43645-43648. Type catalog: Zhang 1996:499 [ref. 24036], Ye et al. 2015:15 [ref. 33808]. Original not seen. •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Paracheilognathus imberbis (Günther 1868) -- (Wu 1964:209 [ref. 13503], Ye et al. 2015:15 [ref. 33808]). Current status: Synonym of Acheilognathus imberbis Günther 1868. Acheilognathidae. Distribution: Hopei Province, China. Habitat: freshwater.

macromandibularis, Acheilognathus Doi [A.], Arai [R.] & Liu [H.-Z.] 1999:305, Fig. 1 [Ichthyological Exploration of Freshwaters v. 10 (no. 4); ref. 24230] Zong Yang, Anhui Province, China. Holotype: IHASW 59VII0308. Paratypes: IHASW 590147 (1). •Valid as Acheilognathus macromandibularis Doi, Arai & Liu 1999 -- (Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus macromandibularis Doi, Arai & Liu 1999. Acheilognathidae. Distribution: China. Habitat: freshwater.

macropterus, Acanthorhodeus Bleeker [P.] 1871:40, Pl. 2 (fig. 2) [Verhandelingen der Koninklijke Akademie van Wetenschappen (Amsterdam) v. 12 (art. 2); ref. 6421] Chang Jiang [Yangtze River], China. Holotype (unique): MNHN 0000-5045. Type catalog: Bertin & Estève 1948:75 [ref. 19574]. Also in Bleeker 1871:71 [ref. 17074]. •Valid as Acanthorhodeus macropterus Bleeker 1871 -- (Wu 1964:212 [ref. 13503], Mai 1978:177 [ref. 13511], Wang 1984:66 [ref. 22951], Jin in Kuang et al. 1986:67 [ref. 26639] dated 1870, Ye in Pan et al. 1991:132 [ref. 23876], Kim 1997:188 [ref. 24032], Kottelat 1998:54 [ref. 23436], Kim & Park 2002:90 [ref. 25913], Choi et al. 2003:156, 504 [ref. 26218]). •In the genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Questionably the same as Acanthorhodeus asmussii (Dybowski 1872) -- (Bogutskaya et al. 2001:44 [ref. 26178]). •Synonym of Acanthorhodeus asmussii (Dybowski 1872) -- (Bogutskaya & Naseka 2004:42 [ref. 28183]). •Valid as Acheilognathus macropterus (Bleeker 1871) -- (Chen & Li in Chu & Chen 1989:129 [ref. 13584], Zheng et al. 1989:159 [ref. 21202], Ding 1994:179 [ref. 21531], Zhu 1995:46 [ref. 25213], Cheng & Zhou 1997:144 [ref. 26385], Kottelat 1998:55 [ref. 23436], Lin in Chen et al. 1998:419 [ref. 23878], Doi et al. 1999:307 [ref. 24230], Kottelat 2001:15 [ref. 25482], Tang et al. 2001:12 [ref. 25373], Wang et al. 2001:152 [ref. 26566], Arai et al. 2007:24 [ref. 29106], Bogutskaya et al. 2008:314 [ref. 30085], Yang et al. 2011:163 [ref. 31154], Kottelat 2013:70 [ref. 32989], Zhang et al. 2016:67 [ref. 34477], Zhang et al. 2019:212 [ref. 36699]). Current status: Valid as Acheilognathus macropterus (Bleeker 1871). Acheilognathidae. Distribution: China, Korea and northern Vietnam. Habitat: freshwater, brackish.

maculatus, Rhodeus Fowler [H. W.] 1910:476, Fig. 1 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 62; ref. 1380] Pei-Ho River, Tien-Tsin, China. Holotype: ANSP 29472. Paratypes: ANSP 29473 (1). Type catalog: Böhlke 1984:82 [ref. 13621]. •Synonym of Rhodeus sinensis Günther 1868 -- (Wu 1964:202 [ref. 13503]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus ocellatus (Kner 1866) -- (Lin in Chen et al. 1998:445 [ref. 23878], Kottelat 2013:157 [ref. 32989]). Current status: Synonym of Rhodeus ocellatus (Kner 1866). Acheilognathidae. Distribution: China (if valid). Habitat: freshwater.

majusculus, Acheilognathus Kim [I.-S.] & Yang [H.] 1998:27, Figs. 1-2 [Korean Journal of Biological Sciences v. 2 (no. 1); ref. 24125] Somjin River, Dae-ri, Kwanchon-myon, Imsil-gun, North Cholla, Chollabuk-do Province, South Korea. Holotype: CNUC 21810. Paratypes: CNUC 21807 (1), 21823 (1), 21825 (1), 23452 (1), 23454 (1), 23458 (1), 23461 (1), 23620-22 (1, 1, 1), 23633 (1), 23699-23702 (4), NSMT-P54556-54559 (4). •Valid as Acheilognathus majusculus Kim & Yang 1998 -- (Kim & Park 2002:86 [ref. 25913], Choi et al. 2003:156 [ref. 26218] as Acheilohnathus, Yang et al. 2010:336 [ref. 30786], Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus majusculus Kim & Yang 1998. Acheilognathidae. Distribution: Korea. Habitat: freshwater.

mantschuricus, Rhodeus Mori [T.] 1934:57 [1], Fig. 1 [Journal of the Chosen Natural History Society No. 17; ref. 16439] Koga, near Kitsurin, upper Sungari River, Manchuria. Holotype: not researched. Additional specimens: (2). On p. 1 of separate. •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus sericeus (Pallas 1776) -- (Berg 1949:816 [ref. 20661], Wu 1964:201 [ref. 13503], Lin in Chen et al. 1998:453 [ref. 23878], Holčík 1999:2 [ref. 24847], Bogutskaya & Naseka 2004:45 [ref. 28183]). •Valid as Rhodeus mantschuricus Mori 1934 -- (Arai et al. 2001:280 [ref. 25545], Bogutskaya et al. 2008:316 [ref. 30085], Dyldin & Orlov 2016:663 [ref. 34853]). Current status: Valid as Rhodeus mantschuricus Mori 1934. Acheilognathidae. Distribution: Amur River basin. Habitat: freshwater.

melanogaster, Acheilognathus Bleeker [P.] 1860:92, Pl. 2 (fig. 1) [Acta Societatis Regiae Scientiarum Indo-Neêrlandicae v. 8 (art. 1); ref. 374] Tokyo, Japan. Lectotype: BMNH 1866.5.2.81. One type not found at RMNH; lectotype selected by Arai & Akai 1988:205 [ref. 1988]. •Synonym of Acheilognathus lanceolatus (Temminck & Schlegel 1846) -- (Okada 1961:384 [ref. 6363]). •Valid as Acheilognathus melanogaster Bleeker 1860 -- (Arai & Akai 1988:199 [ref. 6999], Kawanabe & Mizuno 1989:371 [ref. 25598], Watanabe 1998:262 [ref. 23491], Nakabo 2000:257 [ref. 25086], Matsuura et al. 2000:47 [ref. 26000], Nakabo 2002:257 [ref. 26001], Hagiwara 2003:36 [ref. 27188], Arai et al. 2007:24 [ref. 29106], Yang et al. 2011:163 [ref. 31154], Nguyen et al. 2013:20 [ref. 32689]). Current status: Valid as Acheilognathus melanogaster Bleeker 1860. Acheilognathidae. Distribution: Eastern Honshu, Japan. Habitat: freshwater, brackish.

meridianus, Paracheilognathus Wu [H.-W.] 1939:117 [Sinensia v. 10 (nos 1-6); ref. 4805] Yangao, Li Kiang, southern China. Syntypes: 146 (1), 148 (1), 159 (1), 161-163 (3), 165 (1), 556-557 (2), 559 (1). •Valid as Paracheilognathus meridianus Wu 1939 -- (Wu 1964:210 [ref. 13503], Zheng et al. 1989:163 [ref. 21202], Zhu 1995:45 [ref. 25213], Lin in Chen et al. 1998:441 [ref. 23878], Tang et al. 2001:12 [ref. 25373]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus meridianus (Wu 1939) -- (Kottelat 1998:55 [ref. 23436], Kottelat 2001:15 [ref. 25482] as aff. meridianus, Yang et al. 2010:336 [ref. 30786], Yang et al. 2011:163 [ref. 31154], Nguyan et al. 2013:20 [ref. 32689], Zhang et al. 2016:68 [ref. 34477]). Current status: Valid as Acheilognathus meridianus (Wu 1939). Acheilognathidae. Distribution: Southern China. Habitat: freshwater.

meridionalis, Rhodeus amarus var. Karaman [S.] 1924:41 [Pisces Macedoniae; ref. 15902] Vardar River at Skoplje [Skopje] and Lake Dojran, Republic of Macedonia. Syntypes: IEMBT (several); MNHN 1935-0358 to 0360 (3) Vardar R. Type catalog: Bertin & Estève 1948:75 [ref. 19574]. •Synonym of Rhodeus sericeus (Pallas 1776) -- (Berg 1949:814 [ref. 20661], Holčík 1999:2 [ref. 24847]). •Synonym of Rhodeus amarus (Bloch 1782) -- (Kottelat 1997:75 [ref. 22952]). •Valid as Rhodeus meridionalis Karaman 1924 -- (Kottelat & Freyhof 2007:84 [ref. 29996], Barbieri et al. 2015:61 [ref. 35029], Dyldin & Orlov 2016:663 [ref. 34853], Li et al. 2020:[14] [ref. 37372]). Current status: Valid as Rhodeus meridionalis Karaman 1924. Acheilognathidae. Distribution: Aegean Sea basin, Balkan, southern Europe. Habitat: freshwater.

microlepis, Sinorhodeus Li [F.], Liao [T.-Y.] & Arai [R.] in Li, Liao, Arai & Zhao 2017:73, Figs. 2-9 [Zootaxa 4353 (no. 1); ref. 35680] Tributary of Yangtze River, Banan District, Chongqing City, China, elevation 625 meters. Holotype: SOU 1604001. Paratypes: NSMT, SOU. Plus additional non-type material. •Valid as Sinorhodeus microlepis Li, Liao & Arai 2017. Current status: Valid as Sinorhodeus microlepis Li, Liao & Arai 2017. Acheilognathidae. Distribution: Yangtze River basin, Sichuan Province, China. Habitat: freshwater.

microphysa, Acheilognathus elongatus Yang [J.-X.], Chu [X.-L.] & Chen [Y.-R.] 1990:73, Fig. 2c [Ichthyological Exploration of Freshwaters v. 1 (no. 1); ref. 16186] Haikou, Mingxing and Xijia, 24°21'-24°38'N, 102°49'-102°57'E, Fuxian Lake, Yunnan Province, China. Holotype: KIZ 862894. Paratypes: KIZ 862893 (1), 893200-12 (13). Type catalog: Du et al. 2013:347[ref. 32939]. •Synonym of Acheilognathus elongatus (Regan 1908), but a valid subspecies as described -- (Yang 1991:194 [ref. 13585], Yang & Chen 1995:55 [ref. 23555]). •Valid as Acheilognathus microphysa Yang, Chu & Chen 1990 -- (Kottelat 1998:55 [ref. 23436], Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus microphysa Yang, Chu & Chen 1990. Acheilognathidae. Distribution: China. Habitat: freshwater.

miobuta, Rhodeus Tanaka [S.] 1909:12 [Journal of the College of Science. Imperial University, Tokyo v. 27 (art. 8); ref. 4322] Obama, Kadsusa Province, Japan. Holotype: ZUMT 1860. Paratypes: (4) ZUMT 2006 (1), 2157 (2, lost). •In genus Tanakia -- (Arai & Akai 1988:205 [ref. 6999]). •Synonym of Tanakia tanago (Tanaka 1909) -- (Okada 1961:399 [ref. 6363]). •Synonym of Pseudorhodeus tanago (Tanaka 1909). Current status: Synonym of Pseudorhodeus tanago (Tanaka 1909). Acheilognathidae. Habitat: freshwater.

monguonensis, Pseudoperilampus Li [G.-L.] 1989:3, Fig. 1 [Tianjin Natural History Museum Thesis Coll. v. 6; ref. 25327] Er-dao He-tze, Ku-lun Qi, Inner Mongolia Autonomous Region, China. Holotype: TNZ 02515 (North 503), Tianjin Nat. Hist. Mus. •Valid as Pseudoperilampus monguonensis Li 1989 -- (Zhu 1995:47 [ref. 25213]). •No longer in Pseudoperilampus -- (authors). Current status: Valid as Rhodeus monguonensis (Li 1989). Acheilognathidae. Distribution: China. Habitat: freshwater.

moriokae, Acheilognathus Jordan [D. S.] & Thompson [W. F.] 1914:225, Pl. 25 (fig. 2) [Memoirs of the Carnegie Museum v. 6 (no. 4); ref. 2543] Morioka, Lake Biwa, Funayado, Japan. Holotype: CAS-SU 22643. Paratypes: CAS-SU 22591 [Sex U 7360] (5), USNM 74784 (1). Additional original material: CAS-SU 22584-85 (1, 12). Type catalog: Böhlke 1953:29 [ref. 12291]. •Valid as Acheilognathus moriokae Jordan & Thompson 1914 -- (Okada 1961:377 [ref. 6363], Sawada in Masuda et al. 1984:54 [ref. 6441]). •Synonym of Acheilognathus melanogaster Bleeker 1860 -- (Arai & Akai 1988:199 [ref. 6999]). Current status: Synonym of Acheilognathus melanogaster Bleeker 1860. Acheilognathidae. Habitat: freshwater, brackish.

nakamurae, Acheilognathus tabira Arai [R.], Fujikawa [H.] & Nagata [Y.] 2007:19, Figs. 4D, 5F, 11 [Bulletin of the National Museum of Nature and Science (Ser. A) Suppement No. 1; ref. 29106] Futatsukawa River, Yanagawa, Fukuoka Prefecture, Japan. Holotype: NSMT-P 72659. Paratypes: at NSMT. Plus non-type material. The spelling nakamure is acceptable; early workers almost certainly Latinized the name first, and then added an 'e' (Art. 31.1.1) [WNE, 12 Mar. 2012] -- we suggest using nakamurae when the species is treated as valid. •Synonym of Acheilognathus tabira Jordan & Thompson 1914, but a valid subspecies nakamurae Arai, Fujikawa & Nagata 2007 as described. Current status: Synonym of Acheilognathus tabira Jordan & Thompson 1914. Acheilognathidae. Distribution: Kyushu area, Nagasaki Prefecture, Japan [subspecies]. Habitat: freshwater.

nanchongensis, Acheilognathus Deng [Q.-X.] 1996:19? [Journal of Sichuan Teachers College (Natural Science) v. 17 (no. 4); ref. 30152] Xi-Chong river, Nanchong, Sichuan, China. Holotype: Sichuan Teachers College. Full article not yet seen. •Valid as Acheilognathus nanchongensis Deng 1996 -- (Zhang et al. 2016:68 [ref. 34477]). Current status: Valid as Acheilognathus nanchongensis Deng 1996. Acheilognathidae. Distribution: Nanchong, Sichuan, China. Habitat: freshwater.

ngowyangi, Acanthorhodeus Tchang [T.-L.] 1930:115, Pl. 3 (fig. 2) [Thèses: présentées à la Faculté des Sciences de Paris pour obtenir le grade de docteur des sciences naturelles, Sér. A. No. 209 (order no. 233); ref. 16212] Chang Jiang [Yangtze River], China. Syntypes: (2) MNHN 1934-0097 (1). Type catalog: Bertin & Estève 1948:76 [ref. 19574] with MNHN 1934-97 as holotype. •Synonym of Acanthorhodeus taenianalis Günther 1873 -- (Wu 1964:217 [ref. 13503]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acheilognathus macropterus (Bleeker 1871) -- (Lin in Chen et al. 1998:420 [ref. 23878], Kottelat 2013:70 [ref. 32989], Zhang et al. 2016:67 [ref. 34477]). Current status: Synonym of Acheilognathus macropterus (Bleeker 1871). Acheilognathidae. Habitat: freshwater.

nguyenvanhaoi, Acheilognathus Nguyen [H. D.], Tran [D. H.] & Ta [T. T.] 2013:18, Figs. 1-2 [Tap Chí Sinh Hoc (Journal of Biology) v. 35 (no. 1); ref. 32689] Quing Ninh Province, Tien Yen River at Binh Lieu town, Vietnam. Holotype: MB-HNUE VI.1- FO1. Paratypes: MB-HNUE VI.1-FO2 (1). •Valid as Acheilognathus nguyenvanhaoi Nguyen, Tran & Ta 2013 -- (Kottelat 2013:71 [ref. 32989]). Current status: Valid as Acheilognathus nguyenvanhaoi Nguyen, Tran & Ta 2013. Acheilognathidae. Distribution: Tien Yen River, Vietnam. Habitat: freshwater.

nigrodorsalis, Rhodeus Li [F.], Liao [T.-Y.] & Arai [R.] 2020:[6], Figs. 9-14, 15A-B, 16 [Journal of Vertebrate Biology v. 69 (no. 1); ref. 37372] Le-An River, debouching into Lake Poyang, Yangtze River basin, Ziyang Town, Wuyuan County, Jiangxi Province, China. Holotype: SOU 1712001. Paratypes: DOS, NSMT, SOU. •Valid as Rhodeus nigrodorsalis Li, Liao & Arai 2020 -- (Li et al. 2020:337 [ref. 37574], Li et al. 2020:337 [ref. 37574]). Current status: Valid as Rhodeus nigrodorsalis Li, Liao & Arai 2020. Acheilognathidae. Distribution: Le-An River drainage, Yangtze River basin, Jiangxi and Anhui Provinces, China. Habitat: freshwater.

notatus, Rhodeus Nichols [J. T.] 1929:6, Fig. 4 [American Museum Novitates No. 377; ref. 15912] Tsinan, Shantung Province, China. Holotype: AMNH 9654. Paratypes: AMNH 10812 (8). •Synonym of Rhodeus sinensis Günther 1868 -- (Wu 1964:202 [ref. 13503]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus ocellatus (Kner 1866) -- (Lin in Chen et al. 1998:445 [ref. 23878]). •Valid as Rhodeus notatus Nichols 1929 -- (Kim 1997:168 [ref. 24032], Arai et al. 2001:275 [ref. 25545], Arai et al. 2001:280 [ref. 25545], Kim & Park 2002:68 [ref. 25913], Choi et al. 2003:158, 502 [ref. 26218], Li & Arai 2010:303 [ref. 31188], Li & Arai 2011:303 [ref. 31188], Li & Arai 2014:175 [ref. 33267], Li et al. 2017:85 [ref. 35680], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:337 [ref. 37574]). Current status: Valid as Rhodeus notatus Nichols 1929. Acheilognathidae. Distribution: Northeast China and Korean Peninsula. Habitat: freshwater.

ocellatus, Pseudoperilampus Kner [R.] 1866:548 [6] [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 53; ref. 2636] Shanghai, China. Holotype (unique): NMW 10837. On p. 6 of separate. Also appeared in Kner 1867:365, Pl. 15 (fig. 6) [ref. 18426]. •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Valid as Rhodeus ocellatus (Kner 1866) -- (Wu 1964:203 [ref. 13503], Mai 1978:177 [ref. 13511], Sawada in Masuda et al. 1984:55 [ref. 6441], Jin in Kuang et al. 1986:61 [ref. 26639] dated 1867, Okamura & Machida 1987:102 [ref. 22279], Chen & Li in Chu & Chen 1989:126 [ref. 13584], Zheng et al. 1989:157 [ref. 21202], Ye in Pan et al. 1991:125 [ref. 23876], Kimura & Nagata 1992 [ref. 19826], Ding 1994:174 [ref. 21531], Zhu 1995:46 [ref. 25213], Cheng & Zhou 1997:141 [ref. 26385] dated 1867, Kim 1997:162 [ref. 24032], Kottelat 1998:55 [ref. 23436], Lin in Chen et al. 1998:445 [ref. 23878], Sato et al. 1998:31 [ref. 24972], Tang et al. 2001:12 [ref. 25373], Wang et al. 2001:159 [ref. 26566] dated 1867, Kim & Park 2002:61 [ref. 25913], Bogutskaya et al. 2001:44 [ref. 26178], Choi et al. 2003:158, 502 [ref. 26218], Chen et al. 2005:90 [ref. 28516], Kottelat & Freyhof 2007:84 [ref. 29996], Zhang et al. 2016:68 [ref. 34477], Li et al. 2017:85 [ref. 35680], Zhang et al. 2019:209 [ref. 36699]). •Valid as Rhodeus ocellatus (Kner 1866), subspecies ocellatus (Kner 1866) -- (Okada 1961:400 [ref. 6363], Kawanabe & Mizuno 1989:360 [ref. 25598], Sato 1995:49 [ref. 25531], Nakabo 2000:259 [ref. 25086], Matsuura et al. 2000:119 [ref. 26000], Arai et al. 2001:275 [ref. 25545], Kawamura et al. 2001:369 [ref. 25794], Nakabo 2002:259 [ref. 26001], Bogutskaya & Naseka 2004:44 [ref. 28183], Li & Arai 2010:303 [ref. 31188], Li & Arai 2011:303 [ref. 31188], Kottelat 2013:157 [ref. 32989], Li & Arai 2014:175 [ref. 33267], Dyldin et al. 2020:98 [ref. 37569], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:338 [ref. 37574]). Current status: Valid as Rhodeus ocellatus (Kner 1866). Acheilognathidae. Distribution: East Asia: Yangtze River basin (China); introduced in Russia. Habitat: freshwater.

omeiensis, Acanthorhodeus Shih [H.-J.] & Tchang [T.-L.] 1934:8, Fig. 1 [Contributions of the Biology Department, Scientific Institite West China, Chungking No. 2; ref. 20562] Omei, Szechwan, China. Holotype (unique): whereabouts unknown. •Valid as Acanthorhodeus omeiensis Shih & Tchang 1934 -- (Wu 1964:216 [ref. 13503]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus omeiensis (Shih & Tchang 1934) -- (Ding 1994:181 [ref. 21531], Zhu 1995:46 [ref. 25213], Lin in Chen et al. 1998:422 [ref. 23878], Doi et al. 1999:307 [ref. 24230], Yang et al. 2011:163 [ref. 31154], Zhang et al. 2016:68 [ref. 34477]). Current status: Valid as Acheilognathus omeiensis (Shih & Tchang 1934). Acheilognathidae. Distribution: Yangtze River basin, China. Habitat: freshwater.

oryzae, Rhodeus Jordan [D. S.] & Seale [A.] 1906:144, Fig. 2 [Proceedings of the United States National Museum v. 30 (no. 1445); ref. 2496] Kawatana, northwest of Nagasaki, Kiusiu Island, Japan. Holotype: USNM 53412. Paratypes: CAS-SU 9258 (4), USNM 57798 (2). Type catalog: Böhlke 1953:38 [ref. 12291]. •Valid as Tanakia oryzae (Jordan & Seale 1906) -- (Okada 1961:398 [ref. 6363] as orizae). •Synonym of Tanakia limbata (Temminck & Schlegel 1846) -- (Arai & Akai 1988:201 [ref. 6999]). •Possibly in the genus Acheilognathus -- (Li et al. 2017:83 [ref. 35680]). Current status: Synonym of Tanakia limbata (Temminck & Schlegel 1846). Acheilognathidae. Habitat: freshwater.

peihoensis, Paracheilognathus Fowler [H. W.] 1910:481, Fig. 3 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 62; ref. 1380] Pei-Ho River, Tien-Tsin, China. Holotype: ANSP 29468. Paratypes: ANSP 29469-71 (3, in jar with holotype). Type catalog: Böhlke 1984:86 [ref. 13621]. •Valid as Acanthorhodeus peihoensis (Fowler 1910) -- (Wu 1964:215 [ref. 13503], Wang 1984:71 [ref. 22951]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus peihoensis (Fowler 1910) -- (Zhu 1995:45 [ref. 25213], Kottelat 1998:55 [ref. 23436], Wang et al. 2001:149 [ref. 26566]). •Synonym of Acheilognathus barbatulus Günther 1873 -- (Lin in Chen et al. 1998:425 [ref. 23878], Kottelat 2013:70 [ref. 32989], Zhang et al. 2016:66 [ref. 34477]). Current status: Synonym of Acheilognathus barbatulus Günther 1873. Acheilognathidae. Distribution: China (if valid). Habitat: freshwater.

pingi, Rhodeus Miao [C.-P.] 1934:176, Fig. 30 [Contributions from the Biological Laboratory of the Science Society of China. (Zoological Series) v. 10 (no. 3); ref. 13903] Chinkiang, China. Holotype: SSCN 13200. Other specimens: SSCN 3229 (1), 13188 (1). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus ocellatus (Kner 1866) -- (Wu 1964:203 [ref. 13503], Lin in Chen et al. 1998:446 [ref. 23878], Kottelat 2013:157 [ref. 32989], Zhang et al. 2016:69 [ref. 34477]). Current status: Synonym of Rhodeus ocellatus (Kner 1866). Acheilognathidae. Habitat: freshwater.

polylepis, Acanthorhodeus Wu [H.-W.] in Wu & et al. 1964:219, Fig. 5-22 [The cyprinid fishes of China v. 1; ref. 4806] Hunan and Zhejiang, China. Syntypes: (9). Authorship also seen as Woo. •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus polylepis (Wu 1964) -- (Zhu 1995:46 [ref. 25213], Lin in Chen et al. 1998:427 [ref. 23878], Doi et al. 1999:307 [ref. 24230], Yang et al. 2011:163 [ref. 31154], Zhang et al. 2016:68 [ref. 34477] with author as Wu). Current status: Valid as Acheilognathus polylepis (Wu 1964). Acheilognathidae. Distribution: Yangtze River basin, China. Habitat: freshwater.

polyspinus, Acanthorhodeus Holčík [J.] 1972:183, Fig. 5 [Věstnik Československé společnosti zoologické v. 36 (no. 3); ref. 7615] Red River, Hanoi, northern Vietnam. Holotype: MNHN 1934-0205. Paratypes: MNHN 1934-0206 (1). Other material: MNHN 1907-0287 (1), 1907-0288 (1). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as 'Acanthorhodeus polyspinus (Holčík 1972) -- (Kottelat 1998:54 [ref. 23436]). •Valid as Acheilognathus polyspinus (Holčík 1972) -- (Kottelat 2001:15 [ref. 25482], Yang et al. 2011:163 [ref. 31154], Kottelat 2013:71 [ref. 32989]). Current status: Valid as Acheilognathus polyspinus (Holčík 1972). Acheilognathidae. Distribution: Vietnam. Habitat: freshwater.

pseudorhombea, Paracheilognathus Mori [T.] 1935:565, 574, Fig. 3 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 47; ref. 13904] Rakuto River, Chosen, Korea. Syntypes: a primary specimen plus others. In English on p. 574. •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). Acheilognathidae. Habitat: freshwater.

pseudosericeus, Rhodeus Arai [R.], Jeon [S.-R.] & Ueda [T.] 2001:276, Figs. 1-6 [Ichthyological Research v. 48 (no. 3); ref. 25545] Gumgye River, tributary of Som River, Namhan River system, Hakdam-ri, Gonggun-myon, Hoengsong-gun, Gangwon-do, South Korea. Holotype: SMU 211. Paratypes: SMU 206-210 (5), 212-13 (2, 2), 7229 (2); ZUMT 61146-48 (4, 3, 2), 61149-51 (3). •Valid as Rhodeus pseudosericeus Arai, Jeon & Ueda 2001 -- (Kim & Park 2002:64 [ref. 25913], Li & Arai 2014:175 [ref. 33267], Dyldin & Orlov 2016:663 [ref. 34853], Li et al. 2017:85 [ref. 35680], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:338 [ref. 37574]). Current status: Valid as Rhodeus pseudosericeus Arai, Jeon & Ueda 2001. Acheilognathidae. Distribution: Korea. Habitat: freshwater.

rheinardti, Danio Tirant [G.] 1883:[27] [Bulletin de la Société des Études Indo-chinoises. Saigon. 1883; ref. 17920] Hué, Vietnam. Lectotype: MHNL 4200-0039 [3260]. Paralectotypes: MHNL 4200-0040 (9). Type catalog: Chevey 1937:249 [ref. 31779]. On p. 27 of reprint in Chevey 1929 [ref. 16215]. Also in Tirant 1885: [139] [ref. 17923]. Lectotype selected by Kottelat 1987:13 [ref. 11434]. •Valid as Rhodeus rheinardti (Tirant 1883) -- (Kottelat 1987:14 [ref. 11434], Doi 1997:13 [ref. 22832], Kottelat 1998:53 [ref. 23436], Fang 2001:18 [ref. 25285], Arai et al. 2001:281 [ref. 25545], Kottelat 2013:157 [ref. 32989], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:338 [ref. 37574]). Current status: Valid as Rhodeus rheinardti (Tirant 1883). Acheilognathidae. Distribution: Vietnam. Habitat: freshwater.

rhombea, Capoeta Temminck [C. J.] & Schlegel [H.] 1846:204, Pl. 100 (figs. 6, 6a-b) [Fauna Japonica Parts 10-14; ref. 4374] Near Nagasaki, Japan. Lectotype: RMNH 2490. Paralectotypes: (9 of the following) BMNH 1864.2.16.141 [ex RMNH] (1); RMNH 2491a-c (3), 2492 (1), 2493a-c (3), 2494a-b (2), 2495 (1), 2505a-c (3). Lectotype selected by Boeseman 1947:159 [ref. 12876]. •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus rhombeus (Temminck & Schlegel 1846) -- (Okada 1961:373 [ref. 6363], Sawada in Masuda et al. 1984:54 [ref. 6441], Kawanabe & Mizuno 1989:368 [ref. 25598], Sato 1995:49 [ref. 25531], Kim 1997:186 [ref. 24032], Watanabe 1998:262 [ref. 23491], Kim & Yang 1998:29 [ref. 24125], Nakabo 2000:256 [ref. 25086], Matsuura et al. 2000:49 [ref. 26000], Kim & Park 2002:88 [ref. 25913], Nakabo 2002:256 [ref. 26001], Choi et al. 2003:156, 503 [ref. 26218], Arai et al. 2007:24 [ref. 29106], •Valid as Acanthorhodeus chankaensis (Dybowski 1872) -- (Berg 1949:819 [ref. 20661], Wu 1964:214 [ref. 13503], Wang 1984:70 [ref. 22951], Ye in Pan et al. 1991:135 [ref. 23876], Bogutskaya & Naseka 1996:31 [ref. 22798], Reshetnikov et al. 1997:734 [ref. 24702], Naseka 1998:79 [ref. 23590], Bogutskaya et al. 2001:44 [ref. 26178], Bogutskaya & Naseka 2004:42 [ref. 28183], Bogutskaya et al. 2008:314 [ref. 30085]).). Current status: Valid as Acheilognathus rhombeus (Temminck & Schlegel 1846). Acheilognathidae. Distribution: Japan and Korea. Habitat: freshwater.

robustus, Acanthorhodeus Holčík [J.] 1972:181, Fig. 1 [Věstnik Československé společnosti zoologické v. 36 (no. 3); ref. 7615] Chang Jiang [Yangtze River], Yi-Hing, China. Holotype: MNHN 1934-0088. Paratypes: MNHN 1934-0096 (1). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acheilognathus tonkinensis (Vaillant 1892) -- (Lin in Chen et al. 1998:428 [ref. 23878], Kottelat 2013:71 [ref. 32989], Zhang et al. 2016:68 [ref. 34477]). Current status: Synonym of Acheilognathus tonkinensis (Vaillant 1892). Acheilognathidae. Habitat: freshwater.

sciosemus, Acanthorhodeus Jordan [D. S.] & Thompson [W. F.] 1914:228, Pl. 26 (fig. 1) [Memoirs of the Carnegie Museum v. 6 (no. 4); ref. 2543] Chikugo River, Kurume, Japan. Holotype: CAS-SU 22608. Paratypes: CAS-SU 22609 (6), USNM 74787 (2). Type catalog: Böhlke 1953:29 [ref. 12291]. •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Valid as Acanthorhodeus sciosemus Jordan & Thompson 1914 -- (Okada 1961:407 [ref. 6363]). Current status: Valid as Rhodeus sciosemus (Jordan & Thompson 1914). Acheilognathidae. Distribution: Japan. Habitat: freshwater.

sericeus, Cyprinus Pallas [P. S.] 1776:208, 704 [Reise durch verschiedene Provinzen des russischen Reiches; ref. 18509] Onon River, Dauriya, Russia. No types known. For remarks on systematics see Kottelat 2006:47 [ref. 28831]. •Valid as Rhodeus sericeus (Pallas 1776) -- (Berg 1949:816 [ref. 20661], Wu 1964:201 [ref. 13503], Lee et al. 1980:357 [ref. 22416], Smith 1986:123 [ref. 22186], Lelek 1987:223 [ref. 22962], Arai & Akai 1988:209 [ref. 6999], Page & Burr 1991:66 [ref. 18983], Coad 1995:19 [ref. 23608], Zhu 1995:47 [ref. 25213], Bogutskaya & Naseka 1996:32 [ref. 22798], Cheng & Zhou 1997:139 [ref. 26385], Kim 1997:164 [ref. 24032], Nikiforov et al. 1997:330 [ref. 24694], Reshetnikov et al. 1997:740 [ref. 24702], Naseka 1998:80 [ref. 23590], Lin in Chen et al. 1998:453 [ref. 23878], Briolay et al. 1998:101 [ref. 25586], Holčík 1999:2 [ref. 24847], Fuller et al. 1999:149 [ref. 25838], Zemnukhov et al. 2001:420 [ref. 25412], Arai et al. 2001:275 [ref. 25545], Pietsch et al. 2001:140 [ref. 25688], Bogutskaya et al. 2001:44 [ref. 26178], Bogutskaya & Komlev 2001:84 [ref. 26834] but a valid subspecies, Choi et al. 2003:158, 502 [ref. 26218], Hanel 2003:53 [ref. 27894], Nelson et al. 2004:77 [ref. 27807], Bogutskaya & Naseka 2004:45 [ref. 28183], Kottelat 2006:47 [ref. 28831], Ocock et al. 2006:44 [ref. 28933], Scharpf 2005:40 [ref. 28940], Bogutskaya et al. 2008:316 [ref. 30085], Ninua & Japoshvili 2008:165 [ref. 31251], Li & Arai 2010:303 [ref. 31188], Page & Burr 2011:155 [ref. 31215], Li & Arai 2011:303 [ref. 31188], Page et al. 2013:78 [ref. 32708], Li & Arai 2014:175 [ref. 33267], Parin et al. 2014:88 [ref. 33547], Zhang et al. 2016:69 [ref. 34477], Dyldin & Orlov 2016:663 [ref. 34853], Li et al. 2017:85 [ref. 35680], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:338 [ref. 37574], Dyldin & Orlov 2021:64 [ref. 38158]). Current status: Valid as Rhodeus sericeus (Pallas 1776). Acheilognathidae. Distribution: East Asia: eastern Russia and China; introduced elsewhere. Habitat: freshwater.

shibatae, Acheilognathus Mori [T.] 1928:67 [Japanese Journal of Zoology v. 2 (no. 1) (art. 4); ref. 15862] Tsi-nan, Shantung, northern China. Holotype: not researched. •Synonym of Acanthorhodeus barbatulus (Günther 1873) -- (Wu 1964:214 [ref. 13503]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acheilognathus barbatulus Günther 1873 -- (Lin in Chen et al. 1998:425 [ref. 23878], Kottelat 2013:70 [ref. 32989]). Current status: Synonym of Acheilognathus barbatulus Günther 1873. Acheilognathidae. Habitat: freshwater.

shimazui, Acheilognathus Tanaka [S.] 1908:234, Fig. 1 [v. 20 (no. 237); ref. 33911] Tokiwa, Lake Biwa, Japan. Syntypes: ZUMT (1, lost), 1833 (1, described spec., lost). Also appeared in Tanaka 1908:3, Fig. 1 [ref. 14422]. •In genus Tanakia -- (Arai & Akai 1988:205 [ref. 6999]). •Valid as Tanakia shimazui (Tanaka 1908). Current status: Valid as Tanakia shimazui (Tanaka 1908). Acheilognathidae. Distribution: Japan. Habitat: freshwater.

shitaiensis, Rhodeus Li [F.] & Arai [R.] 2011:305, Figs 1-5, 6a-b, 7 [Ichthyological Exploration of Freshwaters v. 21 (no. 4) [Dec. 2010]; ref. 31188] Qiupu River, Chang Jiang [Yangtze River] system, 30°11'N, 117°30'E, Shitai county, Anhui Province, China. Holotype: SOU 0811001. Paratypes: NSMT-P 90863 (1), 98064 (1); ZMUT 61947 (4); SOU 0811002-0811005 (4), 0905001-0905006 (6), 0905007-0905010 (4). Plus non-type specimens. •Valid as Rhodeus shitaiensis Li & Arai 2011 -- (Li & Arai 2014:175 [ref. 33267], Zhang et al. 2016:69 [ref. 34477], Dyldin & Orlov 2016:663 [ref. 34853], Li et al. 2017:85 [ref. 35680], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:337 [ref. 37574]). Current status: Valid as Rhodeus shitaiensis Li & Arai 2011. Acheilognathidae. Distribution: Qiupu River drainage, lower Yangtze basin, Anhui Province, China. Habitat: freshwater.

signifer, Acheilognathus Berg [L. S.] 1907:159 [Annals and Magazine of Natural History (Series 7) v. 19 (no. 110) (art. 18); ref. 269] Pungtung, Korea. Holotype (unique): ZIN 10265. •In genus Tanakia -- (Arai & Akai 1988:205 [ref. 6999]). •Valid as Acheilognathus signifer Berg 1907 -- (Kim 1997:176 [ref. 24032], Kim & Yang 1998:29 [ref. 24125], Kim & Park 2002:76 [ref. 25913], Choi et al. 2003:156, 503 [ref. 26218], Yang et al. 2011:163 [ref. 31154]). •Valid as Tanaka signifer (Berg 1907) -- (Chang et al. 2009:824 [ref. 31127], Lim et al. 2014:67 [ref. 33508]). Current status: Valid as Tanakia signifer (Berg 1907). Acheilognathidae. Distribution: Korea. Habitat: freshwater.

sinensis, Rhodeus Günther [A.] 1868:280 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Zhejiang, China. Lectotype: BMNH 1868.10.19.150. Paralectotypes: (2) BMNH 1858.10.19.149 (1). Lectotype selected by Akai & Arai 1998:105 [ref. 23354]. •Synonym of Rhodeus ocellatus (Kner 1866) -- (Lin in Chen et al. 1998:445 [ref. 23878]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Valid as Rhodeus sinensis Günther 1868 -- (Wu 1964:202 [ref. 13503], Coad 1981:11 [ref. 22348], Wang 1984:63 [ref. 22951], Lelek 1987:223 [ref. 22962], Chen & Li in Chu & Chen 1989:125 [ref. 13584], Zheng et al. 1989:156 [ref. 21202], Ye in Pan et al. 1991:124 [ref. 23876], Ding 1994:172 [ref. 21531], Yang & Chen 1995:53 [ref. 23555], Zhu 1995:47 [ref. 25213], Coad 1996:84 [ref. 25304], Cheng & Zhou 1997:140 [ref. 26385], Akai & Arai 1998:105 [ref. 23354], Kottelat 1998:55 [ref. 23436], Arai et al. 2001:280 [ref. 25545], Tang et al. 2001:12 [ref. 25373], Wang et al. 2001:157 [ref. 26566], Bogutskaya & Komlev 2001:82 [ref. 26834], Naseka & Bogutskaya 2004:282 [ref. 27838], Bogutskaya & Naseka 2004:43 [ref. 28183], Chen et al. 2005:90 [ref. 28516], Li & Arai 2010:303 [ref. 31188], Li & Arai 2011:303 [ref. 31188], Kottelat 2013:157 [ref. 32989], Li & Arai 2014:175 [ref. 33267], Zhang et al. 2016:69 [ref. 34477], Li et al. 2017:85 [ref. 35680], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:338 [ref. 37574]). Current status: Valid as Rhodeus sinensis Günther 1868. Acheilognathidae. Distribution: Yangtze River basin, China. Habitat: freshwater.

smithii, Achilognathus Regan [C. T.] 1908:150 [Annals and Magazine of Natural History (Series 8) v. 1 (no. 2) (art. 24); ref. 14338] Nodogawa River, Kyoto, Japan. Holotype (unique): BMNH 1907.12.23.125. Original genus should have been Acheilognathus. •Synonym of Acheilognathus rhombeus (Temminck & Schlegel 1846) -- (Okada 1961:373 [ref. 6363]). •Synonym of Rhodeus ocellatus (Kner 1866), but a valid subspecies -- (Sawada in Masuda et al. 1984:55 [ref. 6441], Kawanabe & Mizuno 1989:364 [ref. 25598]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •See Kimura & Nagata 1992:425 [ref. 19826]. •Valid as Rhodeus smithii (Regan 1908) -- (Arai et al. 2001:280 [ref. 25545], Li et al. 2020:[14] [ref. 37372]). Current status: Valid as Rhodeus smithii (Regan 1908). Acheilognathidae. Distribution: Japan. Habitat: freshwater.

somjinensis, Acheilognathus Kim [I.-S.] & Kim [C. H.] 1991:189, Fig. 1 [Korean Journal of Systematic Zoology v. 7 (no. 2); ref. 20050] Owon strait, off the Somjin River, at Sinpyong-myon, Imsil-gun, Chollabuk-do, South Korea. Holotype: CUBK 16419. Paratypes: CUBK 16420-31 (12). Name misspelled as somjinesis once on p. 189. •Valid as Acheilognathus somjinensis Kim & Kim 1991 -- (Kim 1997:180 [ref. 24032], Kim & Yang 1998:29 [ref. 24125], Kim & Park 2002:61 [ref. 25913], Choi et al. 2003:156, 503 [ref. 26218], Yang et al. 2011:163 [ref. 31154]). •Valid as Tanakia somjinensis (Kim & Kim 1991) -- (Chang et al. 2009:824 [ref. 31127], Lim et al. 2014:67 [ref. 33508]). Current status: Valid as Tanakia somjinensis (Kim & Kim 1991). Acheilognathidae. Distribution: South Korea. Habitat: freshwater.

spinalis, Rhodeus Oshima [M.] 1926:16 [Annotationes Zoologicae Japonenses v. 11 (no. 1); ref. 3314] Ding'an, Hainan Island, China. Neotype: NSMT-P 31906. Original 5 specimens from Kachek R. lost; neotype designated by Arai et al. 1990:143 [ref. 16800]. •Valid as Pseudoperilampus spinalis (Oshima 1926) -- (Li 1989:4 [ref. 25327], Zhu 1995:47 [ref. 25213]). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus ocellatus (Kner 1866) -- (Mai 1978:177 [ref. 13511]). •Synonym of Rhodeus rheinardti (Tirant 1883) -- (Kottelat 1987:14, 21 [ref. 11434]). •Valid as Rhodeus spinalis Oshima 1926 -- (Wu 1964:204 [ref. 13503], Jin in Kuang et al. 1986:62 [ref. 26639], Chen & Li in Chu & Chen 1989:127 [ref. 13584], Zheng et al. 1989:158 [ref. 21202], Ye in Pan et al. 1991:126 [ref. 23876], Kottelat 1998:53 [ref. 23436], Lin in Chen et al. 1998:448 [ref. 23878], Kottelat 2001:39 [ref. 25482], Arai et al. 2001:280 [ref. 25545], Li & Arai 2010:303 [ref. 31188], Li & Arai 2011:303 [ref. 31188], Kottelat 2013:157 [ref. 32989], Li & Arai 2014:175 [ref. 33267], Zhang et al. 2016:69 [ref. 34477], Li et al. 2017:85 [ref. 35680], Li et al. 2020:[14] [ref. 37372], Li et al. 2020:338 [ref. 37574]). Current status: Valid as Rhodeus spinalis Oshima 1926. Acheilognathidae. Distribution: Hainan Island, China. Habitat: freshwater, brackish.

steenackeri, Acheilognathus Sauvage [H.-E.] 1883:146 [3] [Bulletin de la Société philomathique de Paris (7th Série) v. 7; ref. 3896] Lake Biwa, Japan. Syntypes: MNHN A-6471 (3), A-6526 (3). Type catalog: Bertin & Estève 1948:75 [ref. 19574]. On p. 3 of separate. •Synonym of Acheilognathus rhombeus (Temminck & Schlegel 1846) -- (Okada 1961:373 [ref. 6363]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). Current status: Synonym of Acheilognathus rhombeus (Temminck & Schlegel 1846). Acheilognathidae. Habitat: freshwater.

striatus, Acheilognathus Yang [Q.], Xiong [B.-X.], Tang [Q.-G.] & Liu [H.-Z.] 2010:334, Figs.1-5 [Environmental Biology of Fishes v. 88 (no. 4); ref. 30786] Lower Chang Jiang [Yangtze River] basin, Wuyuan County, Jiangxi Province, China, 29°25'N, 117°83'E. Holotype: IHASW 04116. Paratypes: many at IHASW. •Valid as Acheilognathus striatus Yang, Xiong, Tang & Liu 2010 -- (Yang et al. 2011:163 [ref. 31154], Zhang et al. 2016:68 [ref. 34477], Li et al. 2017:83 [ref. 35680]). Current status: Valid as Acheilognathus striatus Yang, Xiong, Tang & Liu 2010. Acheilognathidae. Distribution: Le'an River, lower Yangtze River basin, China. Habitat: freshwater.

strumicae, Rhodeus sericeus sericeus forma Karaman [S. L.] 1955:183, Fig. 2 [Acta Musei Macedonici Scientiarum Naturalium v. 3 (no. 7); ref. 12279] Nor available, infrasubspecific. •In the synonymy of Rhodeus amarus (Bloch 1782) -- (Kottelat 1997:75 [ref. 22952]). •In the synonymy of Rhodeus sericeus (Pallas 1776) -- (Holčík 1999:2 [ref. 24847], Bogutskaya & Komlev 2001:82 [ref. 26834]). Infrasubspecific. Current status: Synonym of Rhodeus sericeus (Pallas 1776). Acheilognathidae.

suigensis, Pseudoperilampus Mori [T.] 1935:563, 573, Fig. 2 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 47; ref. 13904] Chosen, Korea. Syntypes: a primary specimen plus others. In English on p. 573. •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus sinensis Günther 1868, but a valid subspecies -- (Kawanabe & Mizuno 1989:366 [ref. 25598]). •Synonym of Rhodeus atremius (Jordan & Thompson 1914), but a valid subspecies -- (Watanabe 1998:262 [ref. 23491], Nakabo 2000:260 [ref. 25086], Matsuura et al. 2000:117 [ref. 26000], Nakabo 2002:260 [ref. 26001]). •Valid as Rhodeus suigensis (Mori 1935) -- (Sawada in Masuda et al. 1984:55 [ref. 6441] as suigenis, Arai et al. 2001:275 [ref. 25545], Li et al. 2020:[14] [ref. 37372]). Current status: Valid as Rhodeus suigensis (Mori 1935). Acheilognathidae. Distribution: Korea. Habitat: freshwater.

sungariensis, Acanthorhodeus asmussi Berg [L. S.] 1931:218, Fig. 6 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 32; ref. 18893] Sungari [Songhua] River, Harbin, China. Syntypes: ZIN 22256 (10). Additional material: ZIN 14470 (5). •In genus Acheilognathus (Arai & Akai 1988:207 [ref. 6999]). •Synonym of Acanthorhodeus chankaensis (Dybowski 1872) -- (Berg 1949:819 [ref. 20661], Bogutskaya & Naseka 1996:31 [ref. 22798], Naseka 1998:79 [ref. 23590], Bogutskaya & Naseka 2004:42 [ref. 28183]). Current status: Synonym of Acheilognathus chankaensis (Dybowski 1872). Acheilognathidae. Habitat: freshwater.

svetovidovi, Rhodeus sericeus amarus natio Holčík [J.] 1959:50 [Voprosy Ikhtiologii v. 13; ref. 23006] Not available, infrasubspecific; from Dnieper and Bug rivers, Ukraine. •In the synonymy of Rhodeus sericeus (Pallas 1776) -- (Holčík 1999:2 [ref. 24847], Bogutskaya & Komlev 2001:82 [ref. 26834]). Infrasubspecific. Current status: Synonym of Rhodeus sericeus (Pallas 1776). Acheilognathidae.

tabira, Acheilognathus Jordan [D. S.] & Thompson [W. F.] 1914:220, Pl. 25 (fig. 1) [Memoirs of the Carnegie Museum v. 6 (no. 4); ref. 2543] Lake Biwa at Matsubara, Japan. Holotype: FMNH 57071 [ex CM 6007a]. Paratypes: CAS 62880 [ex IU 10734] (6), 11116 [ex IU 10743] (3); FMNH 57079 (41); CAS-SU 7363 (75), 22645 (20); USNM 74783 (1). Additional material: CAS-SU 22616 (2), 22620 (1), 22639 (29), 23216 (65). Type catalog: Henn 1928:71 [ref. 12305], Böhlke 1953:29 [ref. 12291], Ibarra & Stewart 1987:5 [ref. 12367]. •Valid as Acanthorhodeus tabira (Jordan & Thompson 1914) -- (Okada 1961:383 [ref. 6363], Wu 1964:218 [ref. 13503] but as tabiro). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus tabira Jordan & Thompson 1914 -- (Sawada in Masuda et al. 1984:54 [ref. 6441], Zhu 1995:46 [ref. 25213] as tabiro, Watanabe 1998:262 [ref. 23491], Lin in Chen et al. 1998:430 [ref. 23878], Doi et al. 1999:306 [ref. 24230], Zhang et al. 2016:68 [ref. 34477]). •Valid as Acheilognathus tabira Jordan & Thompson 1914, subspecies tabira Jordan & Thompson 1914 -- (Kawanabe & Mizuno 1989:373 [ref. 25598], Nakabo 2000:258 [ref. 25086], Matsuura et al. 2000:50 [ref. 26000], Nakabo 2002:258 [ref. 26001], Arai et al. 2007:4 [ref. 29106], Yang et al. 2011:163 [ref. 31154], Ito et al. 2021:47 [ref. 38540]). Current status: Valid as Acheilognathus tabira Jordan & Thompson 1914. Acheilognathidae. Distribution: East Asia: Japan and China; introduced in Mie Prefecture (Japan). Habitat: freshwater.

taenianalis, Acanthorhodeus Günther [A.] 1873:247 [Annals and Magazine of Natural History (Series 4) v. 12 (no. 69) (art. 31); ref. 2004] Shanghai, China. Syntypes: BMNH 1873.7.30.80-83 (4). •Valid as Acanthorhodeus taenianalis Günther 1873 -- (Wu 1964:217 [ref. 13503], Wang 1984:69 [ref. 22951]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acanthorhodeus asmussii (Dubowski 1872) -- (Bogutskaya & Naseka 1996:30 [ref. 22798], Naseka 1998:79 [ref. 23590]). •Valid as Acheilognathus taenianalis (Günther 1873) -- (Chen & Li in Chu & Chen 1989:132 [ref. 13584], Zhu 1995:45 [ref. 25213], Cheng & Zhou 1997:147 [ref. 26385], Kottelat 1998:55 [ref. 23436], Tang et al. 2001:12 [ref. 25373], Wang et al. 2001:148 [ref. 26566]). •Synonym of Acheilognathus macropterus (Bleeker 1871) -- (Lin in Chen et al. 1998:419 [ref. 23878], Kottelat 2001:15 [ref. 25482] with question, Kottelat 2013:70 [ref. 32989], Zhang et al. 2016:67 [ref. 34477]). Current status: Synonym of Acheilognathus macropterus (Bleeker 1871). Acheilognathidae. Distribution: China (if valid). Habitat: freshwater.

tanago, Rhodeus Tanaka [S.] 1909:10 [Journal of the College of Science. Imperial University, Tokyo v. 27 (art. 8); ref. 4322] Pond in Botanical Garden of Imperial University, Tokyo, Japan. Holotype: ZUMT 1710 (lost). Paratypes: ZUMT 2156 (1, lost). •In genus Tanakia -- (Arai & Akai 1988:205 [ref. 6999]). •Valid as Tanakia tanago (Tanaka 1909) -- (Okada 1961:399 [ref. 6363], Sawada in Masuda et al. 1984:55 [ref. 6441], Kawanabe & Mizuno 1989:358 [ref. 25598], Watanabe 1998:262 [ref. 23491], Nakabo 2000:255 [ref. 25086], Matsuura et al. 2000:135 [ref. 26000], Nakabo 2002:255 [ref. 26001], Chang et al. 2009:824 [ref. 31127]). •Valid as Pseudorhodeus tanago (Tanaka 1909) -- (Chang et al. 2014:190 [ref. 33502], Li et al. 2017:84 [ref. 35680]). Current status: Valid as Pseudorhodeus tanago (Tanaka 1909). Acheilognathidae. Distribution: Japan. Habitat: freshwater.

tohokuensis, Acheilognathus tabira Arai [R.], Fujikawa [H.] & Nagata [Y.] 2007:13, Figs. 3A, 5A-B, 9 [Bulletin of the National Museum of Nature and Science (Ser. A) Suppement No. 1; ref. 29106] Lake Nishinuma, Yokote, Akita Prefecture, Japan. Holotype: NSMT-P 72657. Paratypes: AKPM-P 435 (5), 436 (3), 437 (5), 438 (2); NSMT-P 27553 (2), 74701 (65), 74699 (10), 74700 (40); ZMUT 61513 (1), Plus non-type material. •Synonym of Acheilognathus tabira Jordan & Thompson 1914, but a valid subspecies tohokuensis Arai, Fujikawa & Nagata 2007 as described. Current status: Synonym of Acheilognathus tabira Jordan & Thompson 1914. Acheilognathidae. Distribution: Japan Sea side of eastern Honshu, Japan [subspecies]. Habitat: freshwater.

tokunagai, Acanthorhodeus Mori [T.] 1934:35, Pl. 13 (fig. 2) [Report of the first scientific expedition to Manchoukuo, Tokyo, Sec. 5, Zoology Pt 1; ref. 3042] Summer Palace, Jehol, China. Syntypes: NSMT-P 3193 (2). •Synonym of Acanthorhodeus chankaensis (Dybowsky 1872) -- (Wu 1964:214 [ref. 13503]). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Synonym of Acheilognathus chankaensis (Dybowski 1872) -- (Lin in Chen et al. 1998:434 [ref. 23878]). Current status: Synonym of Acheilognathus chankaensis (Dybowski 1872). Acheilognathidae. Habitat: freshwater.

tonkinensis, Acanthorhodeus Vaillant [L. L.] 1892:127 [3] [Bulletin de la Société philomathique de Paris (8th Série) v. 4 (no. 3); ref. 14518] Nam-kia River, affluent to River Noire, Upper Tonkin, northern Vietnam. Holotype (unique): MNHN 1892-0046. Type catalog: Bertin & Estève 1948:76 [ref. 19574]. On p. 3 of separate. See also Kottelat 1998:54 [ref. 23436]. •Valid as Acanthorhodeus tonkinensis Vaillant 1892 -- (Wu 1964:213 [ref. 13503], Mai 1978:184 [ref. 13511], Wang 1984:67 [ref. 22951], Jin in Kuang et al. 1986:68 [ref. 26639], Ye in Pan et al. 1991:133 [ref. 23876] as tonkiennsis and authorship and date as Fang 1942). •In genus Acheilognathus -- (Arai & Akai 1988:208 [ref. 6999]). •Valid as Acheilognathus tonkinensis (Vaillant 1892) -- (Chen & Li in Chu & Chen 1989:130 [ref. 13584], Zheng et al. 1989:160 [ref. 21202], Cheng & Zhou 1997:145 [ref. 26385], Lin in Chen et al. 1998:428 [ref. 23878], Doi et al. 1999:307 [ref. 24230], Kottelat 2001:16 [ref. 25482], Kottelat 2001:38 [ref. 25780], Tang et al. 2001:12 [ref. 25373], Wang et al. 2001:155 [ref. 26566], Yang et al. 2011:163 [ref. 31154], Kottelat 2013:71 [ref. 32989], Zhang et al. 2016:68 [ref. 34477]). Current status: Valid as Acheilognathus tonkinensis (Vaillant 1892). Acheilognathidae. Distribution: China and Vietnam. Habitat: freshwater.

typus, Pseudoperilampus Bleeker [P.] 1863:258, Fig. [Verslagen en Mededeelingen der Koninklijke Akademie van Wetenschappen. Afdeling Natuurkunde. v. 15; ref. 17013] Japan. Holotype (unique): ?BMNH 1866.5.2.43. •Valid as Pseudoperilampus typus Bleeker 1863 -- (Okada 1961:409 [ref. 6363], Sawada in Masuda et al. 1984:55 [ref. 6441]). •In genus Acheilognathus -- (Arai & Akai 1988:209 [ref. 6999]). •Valid as Rhodeus typus (Bleeker 1863) -- (Zhu 1995:46 [ref. 25213]). •Valid as Acheilognathus typus (Bleeker 1863) -- (Kawanabe & Mizuno 1989:367 [ref. 25598], Watanabe 1998:262 [ref. 23491], Nakabo 2000:259 [ref. 25086], Matsuura et al. 2000:55 [ref. 26000], Nakabo 2002:259 [ref. 26001], Arai et al. 2007:24 [ref. 29106], Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus typus (Bleeker 1863). Acheilognathidae. Distribution: Eastern Honshu, Japan. Habitat: freshwater.

uyekii, Pseudoperilampus Mori [T.] 1935:562, 573, Fig. 1 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 47; ref. 13904] Chosen, China. Syntypes: a primary specimen plus others (whereabouts unknown). In English on p. 573. •Synonym of Pseudoperilampus lighti Wu 1931 -- (Wu 1964:205 [ref. 13503] as light). •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus sinensis Günther 1868 -- (Akai & Arai 1998:105 [ref. 23354], Arai et al. 2001:275 [ref. 25545], Kottelat 2013:157 [ref. 32989]). •Valid as Rhodeus uyekii (Mori 1935) -- (Kim 1997:165 [ref. 24032], Kim & Lee 2001:46 [ref. 25334], Kim & Park 2002:66 [ref. 25913], Choi et al. 2003:158, 502 [ref. 26218], Naseka & Bogutskaya 2004:282 [ref. 27838], Bogutskaya & Naseka 2004:44 [ref. 28183]). Current status: Valid as Rhodeus uyekii (Mori 1935). Acheilognathidae. Distribution: Korea and north China. Habitat: freshwater.

vietnamensis, Rhodeus ocellatus Mai [D. Y.] 1978:179 [Identification of the fresh-water fishes of North Viet Nam v. 1; ref. 13511] Northern Vietnam. Syntypes: DVZUT. •Synonym of Rhodeus rheinhardti (Tirant 1883) -- (Kottelat 1987:14, 21 [ref. 11434]). •Status uncertain, as Rhodeus vietnamensis Mai 1978 -- (Kottelat 2001:39 [ref. 25482]). •Synonym of Rhodeus ocellatus (Kner 1866), but a valid subspecies vietnamensis Mai 1978 as described -- (Arai et al. 2001:281 [ref. 25545]). •Synonym of Rhodeus spinalis Oshima 1926 -- (Kottelat 2013:157 [ref. 32989]). Current status: Synonym of Rhodeus spinalis Oshima 1926. Acheilognathidae. Habitat: freshwater.

wangi, Acanthorhodeus Tchang [T.-L.] 1930:115, Pl. 3 (fig. 2) [Thèses: présentées à la Faculté des Sciences de Paris pour obtenir le grade de docteur des sciences naturelles, Sér. A. No. 209 (order no. 233); ref. 16212] Chang Jiang [Yangtze River], China. Holotype (unique): MNHN 1934-0098. Type catalog: Bertin & Estève 1948:76 [ref. 19574]. •Synonym of Acanthorhodeus chankaensis (Dybowsky 1872) -- (Wu 1964:214 [ref. 13503]). •In genus Acheilognathus (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Acheilognathus chankaensis (Dybowski 1872) -- (Lin in Chen et al. 1998:434 [ref. 23878], Zhang et al. 2016:67 [ref. 34477], Zhang et al. 2019:216 [ref. 36699]). Current status: Synonym of Acheilognathus chankaensis (Dybowski 1872). Acheilognathidae. Habitat: freshwater.

wangkinfui, Rhodeus Wu [H.-W.] 1930:77, Fig. 5 [Sinensia v. 1 (no. 6); ref. 16240] Luchow, Chang Jiang [Yangtze River] valley, Szechwan, China. Holotype (unique): not researched. •In genus Rhodeus -- (Arai & Akai 1988:209 [ref. 6999]). •Synonym of Rhodeus ocellatus (Kner 1866) -- (Okada 1961:400 [ref. 6363], Wu 1964:203 [ref. 13503], Okamura & Machida 1987:102 [ref. 22279] as wangkingui, Lin in Chen et al. 1998:446 [ref. 23878], Kottelat 2013:157 [ref. 32989], Zhang et al. 2016:69 [ref. 34477], Zhang et al. 2019:209 [ref. 36699]). Current status: Synonym of Rhodeus ocellatus (Kner 1866). Acheilognathidae. Habitat: freshwater.

yamatsutae, Acheilognathus Mori [T.] 1928:62 [16] [Journal of the Chosen Natural History Society No. 6; ref. 15861] Fen-huang-cheng, Korea. Holotype: not researched. Additional specimens: (3). On p. 16 of separate. The spelling yamatsutae is acceptable; early workers almost certainly Latinized the name first, and then added an 'e' (Art. 31.1.1) [WNE, 12 Mar. 2012] -- we suggest using yamatsutae. •In genus Acheilognathus -- (Arai & Akai 1988:209 [ref. 6999]). •Valid as Acheilognathus yamatsutae Mori 1928 -- (Kim 1997:183 [ref. 24032], Kim & Yang 1998:29 [ref. 24125], Kim & Park 2002:83 [ref. 25913], Choi et al. 2003:156, 504 [ref. 26218], Yang et al. 2010:336 [ref. 30786], Yang et al. 2011:163 [ref. 31154]). Current status: Valid as Acheilognathus yamatsutae Mori 1928. Acheilognathidae. Distribution: Korea. Habitat: freshwater.