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Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information. Species in the family or subfamily Hemiscylliidae: [ 43 ] records arabicum, Chiloscyllium Gubanov [E. P.] in Gubanov & Shleib 1980:14, Figs. 6-7 [Sharks of the Arabian Gulf; ref. 19702] Persian Gulf. No types known. Original not seen. •Valid as Chiloscyllium arabicum Gubanov 1980 -- (Compagno 1984:190 [ref. 6474], Gubanov & Timoxin 1989:221 [ref. 36138], Randall 1995:22 [ref. 22896], Compagno 1999:476 [ref. 25589], Goto 2001:82 [ref. 25912], Compagno 2001:167 [ref. 26323], Manilo & Bogorodsky 2003:S91 [ref. 27377], Moore et al. 2012:5 [ref. 31771], Psomadakis et al. 2015:75 [ref. 34104], Weigmann 2016:8 [ref. 34211], Ali et al. 2018:300 [ref. 36183], Eagderi et al. 2019:13 [ref. 37020], Zajonz et al. 2019:58 [ref. 36871], Haque et al. 2021:215 [ref. 38677] as cf. arabicum, Human 2022:455 [ref. 39664], Hsu et al. 2022:3 [ref. 39567], Borichangar et al. 2024:618 [ref. 41196]). Current status: Valid as Chiloscyllium arabicum Gubanov 1980. Hemiscylliidae. Distribution: Northwestern Indian Ocean: Persian Gulf and Oman east to western India. Habitat: marine. astigma, Chiloscyllium ornatum var. Bleeker [P.] 1863:136 [Nederlandsch Tijdschrift voor de Dierkunde v. 1; ref. 396] Amoy, China. Holotype (unique): whereabouts unknown. Not treated by Dingerkus & DeFino 1983 [ref. 5386]. Hemiscylliidae. Habitat: marine. burmensis, Chiloscyllium Dingerkus [G.] & DeFino [T. C.] 1983:9, Figs. 3, 7, 59-60 [Bulletin of the American Museum of Natural History v. 176 (art. 1); ref. 5386] Off Rangoon, Myanmar, 15°04'N, 95°51'E, depth 29-33 meters. Holotype (unique): USNM 202672. Type catalog: Howe & Springer 1993:5 [ref. 21812]. Chioscyllium is neuter, therefore change to burmense. •Valid as Chiloscyllium burmensis Dingerkus & DeFino 1983 -- (Weigmann 2016:8 [ref. 34211], Habib & Islam 2020:Supplementary table p. 1 [ref. 38321], Psomadakis et al. 2020:165 [ref. 37272]). •Valid as Chiloscyllium burmense Dingerkus & DeFino 1983 -- (Compagno 1999:476 [ref. 25589], Goto 2001:82 [ref. 25912], Compagno 2001:168 [ref. 26323]). Current status: Valid as Chiloscyllium burmense Dingerkus & DeFino 1983. Hemiscylliidae. Distribution: Northeastern Indian Ocean: southern Bangladesh and Myanmar. Habitat: marine. caeruleopunctatum, Chiloscyllium Pellegrin [J.] 1914:230 [Bulletin de la Société Zoologique de France v. 39 (no. 5); ref. 14168] Fort Dauphin, Madagascar. Holotype (unique): MNHN 1914-0009. Type catalog: Bertin 1939:68 [ref. 20739]. •Synonym of Chiloscyllium plagiosum (Anonymous [Bennett] 1830) -- (Dingerkus & DeFino 1983:23 [ref. 5386], Kharin 1987:364 [ref. 6227], Compagno 2001:174 [ref. 26323]). •Valid as Chiloscyllium caeruleopunctatum Pellegrin 1914 -- (Compagno 1984:190 [ref. 6474], Bass 1986:64 [ref. 5635], Weigmann 2016:8 [ref. 34211], Fricke et al. 2018:7 [ref. 35805]). Current status: Valid as Chiloscyllium caeruleopunctatum Pellegrin 1914. Hemiscylliidae. Distribution: Madagascar. Habitat: marine. caudatus, Squalus Gronow [L. T.] in Gray 1854:8 [Catalogue of fish collected and described by Laurence Theodore Gronow; ref. 1911] East Indies. Holotype (unique): BMNH 1853.11.12.205 [Gronovius coll.] (skin). Type catalog: Wheeler 1958:203 [ref. 13434]. •Synonym of Chiloscyllium colax (Meuschen 1781) [not available] -- (Kharin 1987:367 [ref. 6227]). •Objective synonym of Chiloscyllium indicum (Gmelin 1789) -- (Wheeler 1958:203 [ref. 13434], Compagno 1984:193 [ref. 6474], Compagno 2001:172 [ref. 26323], Kottelat 2013:18 [ref. 32989]). Current status: Synonym of Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Habitat: brackish, marine. colax, Hemiscyllium Fowler [H. W.] 1941:89 [Bulletin of the United States National Museum No. 100, v. 13; ref. 6536] China Seas and other areas. Syntypes: USNM 12623 (1), 40032 (3), 75953 (2). Available publication of Squalus colax Meuschen 1781 (unavailable work). Types would be the specimens mentioned by Fowler and specimens represented in literature cited by Fowler. Kottelat (2013) dates to Whitley 1939. •Valid as Hemiscyllium colax Fowler 1941 -- (Goren & Dor 1994:2 [ref. 25356]). •Valid as Chiloscyllium colax (Fowler 1941) -- (Kharin 1987:367 [ref. 6227] with author as Meuschen 1781 (cf. Fowler, 1941)). •Synonym of Chiloscyllium indicum (Gmelin 1789) -- (Dingerkus & DeFino 1983:21 [ref. 5386], Compagno & Niem 1998:1255 [ref. 23787], Compagno 2001:172 [ref. 26323]). Current status: Synonym of Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Habitat: brackish, marine. colax, Squalus Meuschen [F. C.] 1781:[3] [Index to Gronovius, Zoophylacium Gronovianum; ref. 19701] Not available, published in a rejected work (Opinion 261). Can date to Fowler 1941:89 [ref. 6536]. •In the synonymy of Chiloscyllium indicum (Gmelin 1789) -- (Dingerkus & DeFino 1983:21 [ref. 5386], Compagno 1984:193 [ref. 6474], Kottelat 2013:18 [ref. 32989]). •Valid as Chiloscyllium colax (Meuschen 1781) -- (Kharin 1987:367 [ref. 6227] with author as "Meuschen, 1781 (cf. Fowler, 1941)"). Suppressed. Current status: Synonym of Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Habitat: brackish, marine. confusum, Chiloscyllium Dingerkus [G.] & DeFino [T. C.] 1983:9, Figs. 2, 4-7, 51, 57, 61-64 [Bulletin of the American Museum of Natural History v. 176 (art. 1); ref. 5386] Offshore of Calicut (Kozhikode), Kerala State, India. Holotype: AMNH 44126 [ex OSU 1507]. Paratypes: AMNH 44127 (2, 1 c&s); ANSP 81662 (2); BPBM 27525 (1); BMNH 1868.10.25.22 (1), 1904.7.8.12 (1), 1912.3.1.49 (1), 1912.7.20.1 (1); CAS 50893 (1); LACM 38311-31 (3); MCZ 46564 (1); MNHN 1969-0029 (1); OSU 937 (3), 1507 (3); USNM 148105-08 (1, 1, 1, 1), 202654 (1), 202671 (2), 202674 (2), 221698 (2); ZMH 1370(1), 1371 (1), 25397 [ex ISH 1-1961] (1). Type catalog: Krefft 1987:72 [ref. 20751], Wilkens & Dohse 1993:410 [ref. 21161], Howe & Springer 1993:6 [ref. 21812], Thiel et al. 2009:12 [ref. 30918]. •Valid as Chiloscyllium confusum Dingerkus & DeFino 1983 -- (Goto 2001:82 [ref. 25912]). •Synonym of Chiloscyllium arabicum Gubanov 1980 -- (Randall 1995:22 [ref. 22896], Compagno 2001:167 [ref. 26323], Human 2022:455 [ref. 39664]). Current status: Synonym of Chiloscyllium arabicum Gubanov 1980. Hemiscylliidae. Habitat: marine. dentatus, Squalus (Scyliorhinus) Blainville [H. de] 1816:121 [Bulletin des Sciences, par la Société Philomathique de Paris v. 8; ref. 306] Not available, name only (see Dingerkus & DeFino 1983:56 [ref. 5386]). •Synonymy of Chiloscyllium indicum (Gmelin 1789) -- (Compagno 1984:193 [ref. 6474] with question, Compagno 2001:172 [ref. 26323] with question, Kottelat 2013:18 [ref. 32989]). Nomen Nudum. Current status: Synonym of Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Habitat: brackish, marine. denticulatus, Squalus Shaw [G.] 1804:351 [General zoology or systematic natural history; ref. 4015] No locality. Not treated by Dingerkus & DeFino 1983 [ref. 5386]. •Synonym of Chiloscyllium indicum (Gmelin 1789) -- (Kottelat 2013:18 [ref. 32989]). Current status: Synonym of Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Habitat: brackish, marine. dolganovi, Chiloscyllium Kharin [V. E.] 1987:366, Fig. 5 [Voprosy Ikhtiologii v. 27 (no. 3); ref. 6227] Gulf of Thailand, Vietnam, 9°12'01"N, 104°34'06"E, depth 12 meters. Holotype (unique): ZIN 46984. •Synonym of Chiloscyllium hasseltii Bleeker 1852 -- (Last el al. 2010:48 [ref. 32461]). Current status: Synonym of Chiloscyllium hasseltii Bleeker 1852. Hemiscylliidae. Habitat: marine. freycineti, Scyllium Quoy [J. R. C.] & Gaimard [J. P.] 1824:192 [Voyage autour du monde; ref. 3574] Waigeo [Waigiou]; Irian Jaya, Indonesia. Lectotype: MNHN A-7792. Paralectotypes: MNHN B-2962 [ex MNHN A-7792] (1). Type catalog: Bertin 1939:67 [ref. 20739]. Lectotype selected by Dingerkus & de Fina 1983:38 [ref. 5386]. •Valid as Hemiscyllium freycineti (Quoy & Gaimard 1824) -- (Dingerkus & DeFino 1983:38 [ref. 5386], Compagno 1984:196 [ref. 6474], Compagno & Niem 1998:1258 [ref. 23787], Compagno 1999:476 [ref. 25589], Goto 2001:81 [ref. 25912], Compagno 2001:179 [ref. 26323], Allen & Adrim 2003:21 [ref. 26830], Allen & Erdmann 2008:94 [ref. 29476], Allen & Dudgeon 2010:26 [ref. 30713], Allen & Erdmann 2012:48 [ref. 31980], Allen et al. 2013:124 [ref. 32791], Weigmann 2016:9 [ref. 34211], Allen et al. 2016:61 [ref. 35399], White et al. 2017:75 [ref. 35860], Dudgeon et al. 2020:B [ref. 37271], Gloerfelt-Tarp & Kailola 2022:7 [ref. 41223], Allen & Erdmann 2024:48 [ref. 40796]). Current status: Valid as Hemiscyllium freycineti (Quoy & Gaimard 1824). Hemiscylliidae. Distribution: Western Pacific: Raja Ampat and Kri islands (West Papua, Indonesia) endemic. Habitat: marine. galei, Hemiscyllium Allen [G. R.] & Erdmann [M. V.] 2008:96, Figs. 2B, 3-6 [aqua, International Journal of Ichthyology v. 13 (nos 3-4); ref. 29476] Reef near Rumberpon Village, 03°53.757'S, 134°06.638'E, western Cenderawasih Bay, Papua Batat Province, Indonesia. Holotype: NCIP 6324. Paratypes: WAM P.32888-001 (1). •Valid as Hemiscyllium galei Allen & Erdmann 2008 -- (Allen & Dudgeon 2010:26 [ref. 30713], Allen & Erdmann 2012:49 [ref. 31980], Allen et al. 2013:124 [ref. 32791], Weigmann 2016:9 [ref. 34211], Allen et al. 2016:66 [ref. 35399], Dudgeon et al. 2020:B [ref. 37271], Allen & Erdmann 2024:49 [ref. 40796]). Current status: Valid as Hemiscyllium galei Allen & Erdmann 2008. Hemiscylliidae. Distribution: Western Pacific: Cenderawasih Bay and western tip of Yapen Island (West Papua, Indonesia) endemic. Habitat: marine. griseum, Chiloscyllium Müller [J.] & Henle [F. G. J.] 1838:19, [Pl. 4] [Systematische Beschreibung der Plagiostomen; ref. 3069] Puducherry, India; Japan. Lectotype: MNHN 0000-1010. Paralectotypes: (7) MNHN 0000-1009 (1), 0000-1011 (1). Lectotype selected by Dingerkus & DeFino 1983:12 [ref. 5386]. Based on multiple sources including a drawing of a Japanese specimen by Heinrich Bürger. •Valid as Chiloscyllium griseum Müller & Henle 1838 -- (Dingerkus & DeFino 1983:12 [ref. 5386], Compagno 1984:191 [ref. 6474], Nakaya & Shirai in Masuda et al. 1984:8 [ref. 6441], Kharin 1987:365 [ref. 6227], Gubanov & Timoxin 1989:221 [ref. 36138], Talwar & Jhingran 1991:7 [ref. 20764], Krishnan & Mishra 1993:211 [ref. 24016], Goren & Dor 1994:2 [ref. 25356] with Fowler as author, Randall 1995:22 [ref. 22896], Compagno & Niem 1998:1253 [ref. 23787], Morón et al. 1999:144 [ref. 24253], Compagno 1999:476 [ref. 25589], Nakabo 2000:120 [ref. 25086], Compagno in Randall & Lim 2000:578 [ref. 25122], Goto 2001:82 [ref. 25912], Compagno 2001:169 [ref. 26323], Nakabo 2002:120 [ref. 26001], Manilo & Bogorodsky 2003:S91 [ref. 27377], Compagno et al. 2005:20 [ref. 29145], Kimura 2009:7 [ref. 30426], Weigmann 2012:8 [ref. 31968], Allen & Erdmann 2012:47 [ref. 31980], Naylor et al. 2012:53 [ref. 32267] see remarks, Psomadakis et al. 2015:76 [ref. 34104], Weigmann 2016:8 [ref. 34211], Ali et al. 2018:300 [ref. 36183], Kumar et al. 2018:295 [ref. 36178], Eagderi et al. 2019:13 [ref. 37020], Habib & Islam 2020:Supplementary table p. 1 [ref. 38321], Psomadakis et al. 2020:165 [ref. 37272], Haroon & Kibria 2021:43 [ref. 39241], Human 2022:455 [ref. 39664], Hsu et al. 2022:3 [ref. 39567], Gloerfelt-Tarp & Kailola 2022:6 [ref. 41223], Allen & Erdmann 2024:47 [ref. 40796]). Current status: Valid as Chiloscyllium griseum Müller & Henle 1838. Hemiscylliidae. Distribution: Indo-West Pacific: Persian Gulf and Gulf of Oman east to Philippines and Papua New Guinea, north to southern Japan. Habitat: marine. gronovianus, Squalus Shaw [G.] 1804:353 [General zoology or systematic natural history; ref. 4015] East Indies. Holotype (unique): BMNH 1853.11.12.205 [Gronovius coll.] (skin). Type catalog: Wheeler 1958:203 [ref. 13434]. Unneeded new name for (and objective synonym of) Squalus indicus Gmelin 1789. •Objective synonym of Chiloscyllium indicum (Gmelin 1789) -- (Compagno 1984:193 [ref. 6474], Compagno 2001:172 [ref. 26323], Kottelat 2013:18 [ref. 32989]). Current status: Synonym of Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Habitat: brackish, marine. hallstromi, Hemiscyllium Whitley [G. P.] 1967:178 [Australian Zoologist v. 14 (no. 2); ref. 8001] Papua New Guinea. Lectotype: AMS I.15717-001. Paralectotypes: AMS 15584-001 (1). The type specimens were still alive and in the aquarium at Taronga Zoo in June 1970 (Whitley 1970:242 [ref. 6601]). Although Whitley designated a holotype, it was impossible to determine the specimen (see Dingerkus & DeFino 1983:40 [ref. 5386], so they designated a lectotype). Not treated as valid by Goto 2001:81 [ref. 25912]. •Valid as Hemiscyllium hallstromi Whitley 1967 -- (Dingerkus & DeFino 1983:40 [ref. 5386], Compagno 1984:197 [ref. 6474], Compagno & Niem 1998:1258 [ref. 23787], Compagno 1999:476 [ref. 25589], Compagno 2001:180 [ref. 26323], Allen & Adrim 2003:21 [ref. 26830], Allen & Erdmann 2008:94 [ref. 29476], Allen & Dudgeon 2010:26 [ref. 30713], Allen & Erdmann 2012:49 [ref. 31980], Allen et al. 2013:124 [ref. 32791], Weigmann 2016:9 [ref. 34211], Allen et al. 2016:70 [ref. 35399], White et al. 2018:72 [ref. 35860], White & Ko'ou 2018:19 [ref. 35922], Dudgeon et al. 2020:B [ref. 37271], Gloerfelt-Tarp & Kailola 2022:8 [ref. 41223], Allen & Erdmann 2024:49 [ref. 40796]). Current status: Valid as Hemiscyllium hallstromi Whitley 1967. Hemiscylliidae. Distribution: Western Pacific: Gulf of Papua (southern Papua New Guinea); Torres Strait region (Queensland, Australia). Habitat: marine. halmahera, Hemiscyllium Allen [G. R.], Erdmann [M. V.] & Dudgeon [C. L.] 2013:126, Figs. 2-7 [aqua, International Journal of Ichthyology v. 19 (no. 3); ref. 32791] Northwestern Ternate, 00°50.958’N, 127°18.717’E, Halmahera, Indonesia, depth 10 meters. Holotype: MZB 21248. Paratypes: WAM P. 33784-001 (1). •Valid as Hemiscyllium halmahera Allen, Erdmann & Dudgeon 2013 -- (Weigmann 2016:9 [ref. 34211], Allen et al. 2016:74 [ref. 35399], Dudgeon et al. 2020:B [ref. 37271], Akbar et al. 2023:4 [ref. 40618], Allen & Erdmann 2024:50 [ref. 40796]). Current status: Valid as Hemiscyllium halmahera Allen, Erdmann & Dudgeon 2013. Hemiscylliidae. Distribution: Western Pacific: Halmahera (Maluku, Indonesia) endemic. Habitat: marine. hasseltii, Chiloscyllium Bleeker [P.] 1852:19 [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 24 (art. 12); ref. 333] Molucca Islands, Indonesia. Lectotype: BMNH 1867.11.28.196. Paralectotypes: (4) RMNH 7408 (2). Lectotype not regarded as established in Günther 1870:413 [ref. 1995] by use of "Adult. Moluccas ... Type of Ch. hasseltii" becuase Günther used the term to mean a type and not the type. Lectotype selected by Dingerkus & DeFino 1983:17 [ref. 5386]. •Synonym of Chiloscyllium griseum Müller & Henle 1838 -- (Compagno 1984:191 [ref. 6474], Kharin 1987:365 [ref. 6227]). •Valid as Chiloscyllium hasseltii Bleeker 1852 -- (Dingerkus & DeFino 1983:17 [ref. 5386] as hasselti, Compagno & Niem 1998:1254 [ref. 23787] as hasselti, Compagno 1999:476 [ref. 25589] as hasselti, Compagno in Randall & Lim 2000:578 [ref. 25122] as hasselti, Compagno 2001:171 [ref. 26323] as hasselti, Goto 2001:82 [ref. 25912] as hasselti, Last el al. 2010:48 [ref. 32461]. Allen & Erdmann 2012:47 [ref. 31980], Weigmann 2016:8 [ref. 34211], Kumar et al. 2018:295 [ref. 36178], Tyabji et al. 2018:16 [ref. 36598], Habib & Islam 2020:Supplementary table p. 1 [ref. 38321], Psomadakis et al. 2020:165 [ref. 37272], Haque et al. 2021:216 [ref. 38677], Gloerfelt-Tarp & Kailola 2022:6 [ref. 41223], Allen & Erdmann 2024:47 [ref. 40796]). Current status: Valid as Chiloscyllium hasseltii Bleeker 1852. Hemiscylliidae. Distribution: Eastern Indian Ocean, western Pacific: southern Bangladesh, Myanmar and Andaman Islands east to Indonesia and Philippines, north to Vietnam. Habitat: marine. henryi, Hemiscyllium Allen [G. R.] & Erdmann [M. V.] 2008:102, Figs. 2B, 6-9 [aqua, International Journal of Ichthyology v. 13 (nos 3-4); ref. 29476] Northwestern portion of Selat Iris, 3°54.544'S, 134°09.679'E, Papua Barat Province, Indonesia. Holotype: NCIP 6323. Paratypes: USNM 390771 (1), WAM P.32889-001 (1). •Valid as Hemiscyllium henryi Allen & Erdmann 2008 -- (Allen & Dudgeon 2010:26 [ref. 30713], Allen & Erdmann 2012:50 [ref. 31980], Allen et al. 2013:124 [ref. 32791], Weigmann 2016:9 [ref. 34211], Allen et al. 2016:78 [ref. 35399], Dudgeon et al. 2020:B [ref. 37271], Gloerfelt-Tarp & Kailola 2022:8 [ref. 41223], Allen & Erdmann 2024:50 [ref. 40796]). Current status: Valid as Hemiscyllium henryi Allen & Erdmann 2008. Hemiscylliidae. Distribution: Eastern Indian Ocean: Selat Iris (West Papua, Indonesia) endemic. Habitat: marine. indicus, Squalus Gmelin [J. F.] 1789:1503 [Caroli a Linné ... Systema Naturae per regna tria naturae v. 1 (pt 3); ref. 18139] East Indies. Holotype (unique): BMNH 1853.11.12.205 [Gronovius coll.] (skin). Type catalog: Wheeler 1958:203 [ref. 13434]. •Synonym of Chiloscyllium colax [not available] -- (Kharin 1987:367 [ref. 6227]). •Valid as Chiloscyllium indicum (Gmelin 1789) -- (Dingerkus & DeFino 1983:21 [ref. 5386], Compagno 1984:193 [ref. 6474], Nakaya & Shirai in Masuda et al. 1984:8 [ref. 6441], Gubanov & Timoxin 1989:221 [ref. 36138], Talwar & Jhingran 1991:8 [ref. 20764], Mohsin & Ambak 1996:54 [ref. 27969], Compagno & Niem 1998:1255 [ref. 23787], Compagno 1999:476 [ref. 25589], Nakabo 2000:120 [ref. 25086], Compagno in Randall & Lim 2000:578 [ref. 25122], Compagno 2001:172 [ref. 26323], Goto 2001:82 [ref. 25912], Shinohara et al. 2001:287 [ref. 25995], Nakabo 2002:120 [ref. 26001], Choi et al. 2003:50, 469 [ref. 26218], Manilo & Bogorodsky 2003:S91 [ref. 27377], Compagno et al. 2005:20 [ref. 29145], Last el al. 2010:50 [ref. 32461], Allen & Erdmann 2012:47 [ref. 31980], Naylor et al. 2012:52 [ref. 32267] see remarks, Kottelat 2013:18 [ref. 32989], Psomadakis et al. 2015:76 [ref. 34104], Weigmann 2016:8 [ref. 34211], Kumar et al. 2018:295 [ref. 36178], Habib & Islam 2020:Supplementary table p. 1 [ref. 38321], Psomadakis et al. 2020:166 [ref. 37272], Haroon & Kibria 2021:44 [ref. 39241], Gloerfelt-Tarp & Kailola 2022:6 [ref. 41223], Human 2022:456 [ref. 39664], Allen & Erdmann 2024:47 [ref. 40796]). Current status: Valid as Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Distribution: Indo-West Pacific: Pakistan east to western Indonesia, ?Philippines and ?Solomon Islands, north to southern China, ?Korea and ?Japan. Habitat: brackish, marine. interruptum, Chiloscyllium plagiosum var. Bleeker [P.] 1852:18 [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 24 (art. 12); ref. 333] Jakarta and Semarang, Java, Indonesia. No types known. Not treated by Dingerkus & DeFino 1983 [ref. 5386]. •Synonym of Chiloscyllium plagiosum (Anonymous [Bennett] 1830) -- (Compagno 1984:194 [ref. 6474], Compagno 2001:174 [ref. 26323] with question, Last el al. 2010:52 [ref. 32461], Endruweit 2024:31 [ref. 41040]). Current status: Synonym of Chiloscyllium plagiosum (Anonymous [Bennett] 1830). Hemiscylliidae. Habitat: marine. lambarda, Squalis (Scyliorhinus) Blainville [H. de] 1816:121 [Bulletin des Sciences, par la Société Philomathique de Paris v. 8; ref. 306] Not available, name only (see Dingerkus & DeFino 1983:56 [ref. 5386]). •Questionably in the synonymy of Chiloscyllium colax [not available] -- (Kharin 1987:367 [ref. 6227]). Nomen Nudum. Current status: Synonym of Chiloscyllium. Hemiscylliidae. malaisianum, Scyllium Lesson [R. P.] 1831:94, Poissons Pl. 6 [Voyage autour du monde Zool. v. 2 (pt 1); ref. 2776] Offack Bay, Waigeo [Waigiou], Irian Jaya, Indonesia. Lectotype: MNHN A-7767. Paralectotypes: MNHN A-7792 (1), B-2962 [ex MNHN A-7792] (1). Type catalog: Bertin 1939:67 [ref. 20739] with MNHN A.7767 as figured holotype; that specimen designated lectotype by Eschmeyer 1998:1008 [ref. 23416]. Spelled malaianum by Müller & Henle 1838:20 [ref. 3069] and malayanum by Bleeker 1852:6 (not researched). •Synonym of Hemiscyllium freycineti (Quoy & Gaimard 1824) -- (Dingerkus & DeFino 1983:38 [ref. 5386], Compagno 1984:196 [ref. 6474], Compagno 2001:179 [ref. 26323], Allen et al. 2016:61 [ref. 35399]). Current status: Synonym of Hemiscyllium freycineti (Quoy & Gaimard 1824). Hemiscylliidae. Habitat: marine. margaritiferum, Chiloscyllium Bleeker [P.] 1863:243 [Nederlandsch Tijdschrift voor de Dierkunde v. 1; ref. 17006] Obi Island, Molucca Islands, Indonesia. Holotype (unique): RMNH 7404 (260-mm male of 2). •Synonym of Chiloscyllium punctatum Müller & Henle 1838 -- (Compagno 1984:194 [ref. 6474], Kharin 1987:367 [ref. 6227], Compagno & Niem 1998:1257 [ref. 23787]). •Synonym of Chiloscyllium plagiosum (Anonymous [Bennett] 1830) -- (Dingerkus & DeFino 1983:23 [ref. 5386], Compagno 2001:174 [ref. 26323], Last el al. 2010:52 [ref. 32461], Endruweit 2024:31 [ref. 41040]). Current status: Synonym of Chiloscyllium plagiosum (Anonymous [Bennett] 1830). Hemiscylliidae. Habitat: marine. melanostigma, Chiloscyllium ornatum var. Bleeker [P.] 1863:136 [Nederlandsch Tijdschrift voor de Dierkunde v. 1; ref. 396] Singapore. Holotype (unique): whereabouts unknown. Not treated by Dingerkus & DeFino 1983 [ref. 5386]. Hemiscylliidae. Habitat: marine. michaeli, Hemiscyllium Allen [G. R.] & Dudgeon [C. L.] 2010:22, Figs. 2-4 [aqua, International Journal of Ichthyology v. 16 (no. 1); ref. 30713] Coral reef in Harvey Bay, 8°54.440'S, 148°39.306'E, east of Oro Bay, Papua New Guinea. Holotype: USNM 218602. Paratypes: AMS IA.5741-2 (2), CUMV 24992 (1), USNM 221705 (1), WAM P.32840-001 (1). •Valid as Hemiscyllium michaeli Allen & Dudgeon 2010 -- (Allen & Erdmann 2012:50 [ref. 31980], Allen et al. 2013:124 [ref. 32791], Weigmann 2016:9 [ref. 34211], Allen et al. 2016:81 [ref. 35399], White et al. 2018:74 [ref. 35860], White & Ko'ou 2018:20 [ref. 35922], Dudgeon et al. 2020:B [ref. 37271], Allen & Erdmann 2024:51 [ref. 40796]). Current status: Valid as Hemiscyllium michaeli Allen & Dudgeon 2010. Hemiscylliidae. Distribution: Western Pacific: Milne Bay (Papua New Guinea) endemic. Habitat: marine. obscura, Chiloscyllium indicum var. Günther [A.] 1870:413 [Catalogue of the fishes in the British Museum v. 8; ref. 1995] Apparently not an original description; intended for Scyllium obscurum of authors. See Dingerkus & DeFino 1983:17 [ref. 5386]. Hemiscylliidae. obscurum, Chiloscyllium Gray [J. E.] 1851:35 [List of the specimens of fish in the collection of the British Museum.; ref. 4939] Not available, no distinguishing features provided; from Moluccas Is. (based on BMNH 1845.6.22.122). •Questionably a synonym of Chiloscyllium griseum Müller & Henle 1838 -- (Compagno 1984:191 [ref. 6474]). •In the synonymy (as name only) of Chiloscyllium griseum Müller & Henle 1838 -- (Kharin 1987:365 [ref. 6227]). •In the synonymy of Chiloscyllium hasseltii Bleeker 1852 -- (Dingerkus & DeFino 1983:17 [ref. 5386], Compagno 2001:171 [ref. 26323], Last el al. 2010:48 [ref. 32461]). Nomen Nudum. Current status: Synonym of Chiloscyllium griseum Müller & Henle 1838. Hemiscylliidae. Habitat: marine. ocellatus, Squalus Walbaum [J. J.] 1792:516 [Petri Artedi sueci genera piscium Part 3; ref. 4572] No locality stated. Objectively invalid; preoccupied by Squalus ocellatus Bonnaterre 1788 and Gmelin 1789. 1°Homonym. Hemiscylliidae. Habitat: marine. ocellatus, Squalus Bonnaterre [J. P.] (ex Broussonet) 1788:8 [Tableau encyclopédique et méthodique des trois règnes de la nature... Ichthyologie; ref. 4940] Cooktown, Queensland, Australia, Coral Sea, southwestern Pacific. Holotype (unique): MNHN 0000-1003. Type catalog: Bauchot 1969:127 [ref. 20807]. Based on 'L'Œillé' of Broussonet 1784:660 [ref. 20282]. •Valid as Hemiscyllium ocellatum (Bonnaterre 1788) -- (Compagno 1984:197 [ref. 6474], Allen & Swainston 1988:20 [ref. 25477], Gubanov & Timoxin 1989:221 [ref. 36138], Paxton et al. 1989:90 [ref. 12442], Randall et al. 1990:14 [ref. 15987], Last & Stevens 1994:136 [ref. 23873], Allen 1997:40 [ref. 23977], Larson & Williams 1997:343 [ref. 23967], Randall et al. 1997:14 [ref. 25919], Compagno & Niem 1998:1258 [ref. 23787], Compagno 1999:476 [ref. 25589], Hutchins 2001:13 [ref. 25847], Goto 2001:81 [ref. 25912], Compagno 2001:181 [ref. 26323], Allen & Adrim 2003:21 [ref. 26830], Hoese et al. 2006:123 [ref. 29001], Allen & Erdmann 2008:94 [ref. 29476], Allen & Dudgeon 2010:26 [ref. 30713], Allen et al. 2013:124 [ref. 32791], Larson et al. 2013:14 [ref. 32988], Weigmann 2016:9 [ref. 34211], Allen et al. 2016:84 [ref. 35399], Dudgeon et al. 2020:B [ref. 37271]). Current status: Valid as Hemiscyllium ocellatum (Bonnaterre 1788). Hemiscylliidae. Distribution: Western Pacific: Queensland (Australia). Habitat: marine. ocellatus, Squalus Gmelin [J. F.] 1789:1494 [Caroli a Linné ... Systema Naturae per regna tria naturae v. 1 (pt 3); ref. 18139] Objectively invalid; preoccupied by Squalus ocellatus Bonnaterre 1788. 1°Homonym. Hemiscylliidae. Habitat: marine. oculatus, Squalus Banks [J.] & Solander [D. C.] in Gray 1827:436 [List of fish of Australia; ref. 18152] Cooktown, Queensland, Australia. Holotype (unique): MNHN 0000-1003. Not available from above; manuscript name mentioned in synonymy of Squalus ocellatus, subsequent validation not researched. Perhaps validated by Duméril 1853:119 (see Dingerkus & DeFino 1983:43 [ref. 5386]). •In the synonymy of Hemiscyllium ocellatum (Bonnaterre 1788) -- (Dingerkus & DeFino 1983:43 [ref. 5386], Compagno 1984:197 [ref. 6474], Paxton et al. 1989:90 [ref. 12442], Last & Stevens 1994:136 [ref. 23873], Compagno 2001:181 [ref. 26323], Hoese et al. 2006:124 [ref. 29001]). Nomen Nudum. Current status: Synonym of Hemiscyllium ocellatum (Bonnaterre 1788). Hemiscylliidae. ornatum, Scyllium Gray [J. E.] 1830:no page number, Pl. 98 (fig. 2) [Illustrations of Indian zoology; ref. 1878] China seas. Holotype (unique): BMNH 1982.2.26.1 (dry). Published about 15 July 1830. •Synonym of Chiloscyllium plagiosum (Anonymous [Bennett] 1830) -- (Dingerkus & DeFino 1983:23 [ref. 5386], Compagno 1984:194 [ref. 6474], Kharin 1987:364 [ref. 6227], Compagno & Niem 1998:1256 [ref. 23787], Compagno 2001:174 [ref. 26323], Endruweit 2024:31 [ref. 41040]). Current status: Synonym of Chiloscyllium plagiosum (Anonymous [Bennett] 1830). Hemiscylliidae. Habitat: marine. phymatodes, Chiloscyllium Bleeker [P.] 1852:21 [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 24 (art. 12); ref. 333] Semarang, Java, Indonesia. Holotype (unique): possibly RMNH 7406 (smaller specimen). Bleeker specimens: BMNH 1867.11.28.200 (1). •Synonym of Chiloscyllium colax (Meuschen 1781) [not available] -- (Kharin 1987:367 [ref. 6227]). •Synonym of Chiloscyllium indicum (Gmelin 1789) -- (Dingerkus & DeFino 1983:21 [ref. 5386], Compagno 1984:193 [ref. 6474], Compagno 2001:172 [ref. 26323], Last el al. 2010:48 [ref. 32461], Kottelat 2013:18 [ref. 32989]). Current status: Synonym of Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Habitat: brackish, marine. plagiosum, Scyllium Anonymous [Bennett [E. T.]] 1830:694 [Memoir of the life and public services of Sir Thomas Stamford Raffles; ref. 259] Java Sea, 5°58'S, 106°48'E, Sumatra, Indonesia. Neotype: CAS 36046. Neotype designated by Dingerkus & DeFino 1983:21 [ref. 5386]. On Code recommendation 51D, the anonymous author is listed in square brackets. •Valid as Chiloscyllium plagiosum (Anonymous [Bennett] 1830) -- (Dingerkus & DeFino 1983:22 [ref. 5386], Compagno 1984:194 [ref. 6474], Nakaya & Shirai in Masuda et al. 1984:8 [ref. 6441], Kharin 1987:364 [ref. 6227], Gubanov & Timoxin 1989:221 [ref. 36138], Chen et al. 1997:2 [ref. 26476], Compagno & Niem 1998:1256 [ref. 23787], Compagno 1999:476 [ref. 25589], Nakabo 2000:120 [ref. 25086], Compagno in Randall & Lim 2000:578 [ref. 25122], Goto 2001:82 [ref. 25912], Compagno 2001:173 [ref. 26323], Soto 2001:98 [ref. 26637], Nakabo 2002:120 [ref. 26001], Manilo & Bogorodsky 2003:S91 [ref. 27377], Compagno et al. 2005:21 [ref. 29145], Last el al. 2010:52 [ref. 32461], Allen & Erdmann 2012:48 [ref. 31980], Yamashita et al. 2012:120 [ref. 32408], Ebert et al. 2013:304 [ref. 33045], Fricke et al. 2014:12 [ref. 33932], Weigmann 2016:8 [ref. 34211], White et al. 2018:68 [ref. 35860], White & Ko'ou 2018:17 [ref. 35922], Gloerfelt-Tarp & Kailola 2022:7 [ref. 41223], Human 2022:456 [ref. 39664], Allen & Erdmann 2024:48 [ref. 40796], Endruweit 2024:31 [ref. 41040]). Current status: Valid as Chiloscyllium plagiosum (Anonymous [Bennett] 1830). Hemiscylliidae. Distribution: Indo-West Pacific: Madagascar; India east to Philippines and New Guinea, north to Japan and Korea, south to Solomon Islands. Habitat: marine. punctatum, Chiloscyllium Müller [J.] & Henle [F. G. J.] (ex Kuhl & van Hasselt) 1838:18, [Pl. 3] [Systematische Beschreibung der Plagiostomen; ref. 3069] Off Jakarta, Java, Indonesia. Neotype: AMNH 38153. Scyllium punctatum Kuhl & van Hasselt as name only mentioned in synonymy. Neotype designated by Dingerkus & DeFino 1983:30 [ref. 5386]. Scylia russelii Billberg 1833 is probably a senior synonym; Chiloscyllium punctatum is in prevailing usage and therefore here treated as the valid name. •Valid as Chiloscyllium punctatum Müller & Henle 1838 -- (Dingerkus & DeFino 1983:30 [ref. 5386], Compagno 1984:194 [ref. 6474], Nakaya & Shirai in Masuda et al. 1984:8 [ref. 6441], Allen & Swainston 1988:20 [ref. 25477], Gubanov & Timoxin 1989:221 [ref. 36138], Paxton et al. 1989:89 [ref. 12442], Kharin 1987:367 [ref. 6227], Randall et al. 1990:14 [ref. 15987], Last & Stevens 1994:135 [ref. 23873], Mohsin & Ambak 1996:55 [ref. 27969], Allen 1997:40 [ref. 23977], Larson & Williams 1997:343 [ref. 23967], Randall et al. 1997:14 [ref. 25919], Compagno & Niem 1998:1254 [ref. 23787], Johnson 1999:720 [ref. 25471], Compagno 1999:476 [ref. 25589], Nakabo 2000:120 [ref. 25086], Compagno in Randall & Lim 2000:578 [ref. 25122], Hutchins 2001:13 [ref. 25847], Compagno 2001:175 [ref. 26323], Goto 2001:82 [ref. 25912], Nakabo 2002:120 [ref. 26001], Allen & Adrim 2003:21 [ref. 26830], Adrim et al. 2004:118 [ref. 27715], Hoese et al. 2006:123 [ref. 29001], Compagno et al. 2005:21 [ref. 29145], Kimura 2009:7 [ref. 30426], Motomura et al. 2010:68 [ref. 31256], Last el al. 2010:53 [ref. 32461], Weigmann 2012:11 [ref. 31968], Allen & Erdmann 2012:48 [ref. 31980], Naylor et al. 2012:52 [ref. 32267] see remarks, Larson et al. 2013:14 [ref. 32988], Ebert et al. 2013:305 [ref. 33045], Yoshida et al. 2013:8 [ref. 34464], Fricke et al. 2014:13 [ref. 33932], Weigmann 2016:9 [ref. 34211], Koeda et al. 2016:4 [ref. 35333], Kumar et al. 2018:295 [ref. 36178], White et al. 2018:70 [ref. 35860], White & Ko'ou 2018:18 [ref. 35922], Habib & Islam 2020:Supplementary table p. 1 [ref. 38321], Psomadakis et al. 2020:166 [ref. 37272], Sonoyama et al. 2020:8 [ref. 37637], Haroon & Kibria 2021:44 [ref. 39241], Gloerfelt-Tarp & Kailola 2022:7 [ref. 41223], Human 2022:457 [ref. 39664], Hsu et al. 2022:3 [ref. 39567], Vilasri et al. 2023:71 [ref. 40764], Allen & Erdmann 2024:48 [ref. 40796]). Current status: Valid as Chiloscyllium punctatum Müller & Henle 1838. Hemiscylliidae. Distribution: Indo-West Pacific: southwestern India east to Philippines and Papua New Guinea, north to Japan, south to northern Australia. Habitat: marine. russelii, Scylia Billberg [G. J.] 1833:51 [Linnéska Samfundets Handlingar v. 1 (for 1832); ref. 34146] Vizagapatnam, India. No types known. Based on the "Boku Sorsa" [sic, should have been "Bokee sorrah"] of Russell 1803, v. 1:10, Pl.16 [ref. 13862]; see Kullander 2016:112 [ref. 34147]. Not used as valid after 1899; therefore reversal of priority in favor of Chiloscyllium punctatum Müller & Henle 1838 is followed (ICZN Art. 23.9). Named for Russell (misspelled as Russel), an inadvertent error that must be corrected, Code Art. 32.5.1. •Possibly a synonym of Chiloscyllium punctatum Müller & Henle 1838 -- (Kullander 2016:112 [ref. 34147]). Current status: Synonym of Chiloscyllium punctatum Müller & Henle 1838. Hemiscylliidae. Habitat: marine. russellianus, Squalus (Scyliorhinus) Blainville [H. de] 1816:121 [Bulletin des Sciences, par la Société Philomathique de Paris v. 8; ref. 306] Not available, name only (see Dingerkus & DeFino 1983:54 [ref. 5386]). •Questionably in the synonymy of Chiloscyllium punctatum Müller & Henle 1838 -- (Compagno 1984:194 [ref. 6474]). Nomen Nudum. Current status: Synonym of Chiloscyllium punctatum Müller & Henle 1838. Hemiscylliidae. strahani, Hemiscyllium Whitley [G. P.] 1967:176, Fig. 1 [Australian Zoologist v. 14 (no. 2); ref. 8001] Vicinity of Port Moresby [in error], Papua New Guinea. Holotype (unique): AMS IB.7938. •Valid as Hemiscyllium strahani Whitley 1967 -- (Dingerkus & DeFino 1983:46 [ref. 5386], Compagno 1984:198 [ref. 6474], Compagno & Niem 1998:1259 [ref. 23787], Compagno 1999:476 [ref. 25589], Goto 2001:81 [ref. 25912], Compagno 2001:182 [ref. 26323], Allen & Adrim 2003:21 [ref. 26830], Allen & Erdmann 2008:94 [ref. 29476], Allen & Dudgeon 2010:26 [ref. 30713], Allen & Erdmann 2012:51 [ref. 31980], Allen et al. 2013:124 [ref. 32791], Fricke et al. 2014:13 [ref. 33932], Weigmann 2016:9 [ref. 34211], Allen et al. 2016:89 [ref. 35399], White et al. 2018:76 [ref. 35860], White & Ko'ou 2018:20 [ref. 35922], Dudgeon et al. 2020:B [ref. 37271], Gloerfelt-Tarp & Kailola 2022:8 [ref. 41223], Allen & Erdmann 2024:51 [ref. 40796]). Current status: Valid as Hemiscyllium strahani Whitley 1967. Hemiscylliidae. Distribution: Western Pacific: northern New Guinea from eastern West Papua (Indonesia) east to Madang (Papua New Guinea). Habitat: marine. trispeculare, Hemiscyllium Richardson [J.] 1843:5, Pl. 1 (fig. 2) [Icones piscium, or plates of rare fishes; ref. 6537] North West Australia. Neotype: BMNH 1953.5.10.1. Based on a drawing of specimen which was not kept. Neotype designated by Dingerkus & DeFino 1983:51 [ref. 5386]. •Valid as Hemiscyllium trispeculare Richardson 1843 -- (Compagno 1984:199 [ref. 6474], Allen & Swainston 1988:20 [ref. 25477], Paxton et al. 1989:90 [ref. 12442], Randall et al. 1990:14 [ref. 15987], Last & Stevens 1994:137 [ref. 23873], Allen 1997:40 [ref. 23977], Larson & Williams 1997:343 [ref. 23967], Randall et al. 1997:14 [ref. 25919], Compagno & Niem 1998:1253 [ref. 23787], Compagno 1999:476 [ref. 25589], Hutchins 2001:13 [ref. 25847], Goto 2001:81 [ref. 25912], Compagno 2001:183 [ref. 26323], Allen & Adrim 2003:21 [ref. 26830], Hoese et al. 2006:124 [ref. 29001], Allen & Erdmann 2008:94 [ref. 29476], Allen & Dudgeon 2010:29 [ref. 30713], Allen et al. 2013:124 [ref. 32791], Larson et al. 2013:14 [ref. 32988], Weigmann 2016:9 [ref. 34211], Allen et al. 2016:92 [ref. 35399], Dudgeon et al. 2020:B [ref. 37271], Gloerfelt-Tarp & Kailola 2022:8 [ref. 41223], Allen & Erdmann 2024:52 [ref. 40796]). Current status: Valid as Hemiscyllium trispeculare Richardson 1843. Hemiscylliidae. Distribution: Eastern Indian Ocean: Point Quobba northward (Western Australia); Northern Territory (Australia); Aru Islands (Maluku, Indonesia). Habitat: marine. tuberculatus, Squalus Bloch [M. E.] & Schneider [J. G.] 1801:137 [M. E. Blochii, Systema Ichthyologiae; ref. 471] No locality. Non-types: BMNH 1853.11.12.205 (1, dried skin), ZMB 4443 (2). Type catalog: Paepke & Schmidt 1988:162 [ref. 21041]. •Synonym of Chiloscyllium colax (Meuschen 1781) [not available] -- (Kharin 1987:367 [ref. 6227]). •Synonym of Chiloscyllium indicum (Gmelin 1789) -- (Dingerkus & DeFino 1983:21 [ref. 5386], Compagno 1984:193 [ref. 6474], Compagno 2001:172 [ref. 26323], Kottelat 2013:18 [ref. 32989]). Current status: Synonym of Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Habitat: brackish, marine. unicolor, Squalis (Scyliorhinus) Blainville [H. de] 1816:121 [Bulletin des Sciences, par la Société Philomathique de Paris v. 8; ref. 306] Not available, name only (see Dingerkus & DeFino 1983:56 [ref. 5386]). •In the synonymy of Chiloscyllium griseum Müller & Henle 1838 -- (Kharin 1987:365 [ref. 6227]). Nomen Nudum. Current status: Synonym of Chiloscyllium griseum Müller & Henle 1838. Hemiscylliidae. Habitat: marine. variegatus, Squalus (Scyliorhinus) Blainville [H. de] 1816:121 [Bulletin des Sciences, par la Société Philomathique de Paris v. 8; ref. 306] Not available, name only (see Dingerkus & DeFino 1983:56 [ref. 5386]). •Questionably in the synonymy of Chiloscyllium colax Gmelin 1789 -- (Kharin 1987:367 [ref. 6227] with author as Meuschen). •In the synonymy of Chiloscyllium indicum (Gmelin 1789). Nomen Nudum. Current status: Synonym of Chiloscyllium indicum (Gmelin 1789). Hemiscylliidae. Habitat: brackish, marine. |