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Eschmeyer's Catalog of Fishes
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Species in the genus Engraulis: [ 41 ] records


adriaticus, Engraulis encrasicholus race Aleksandrov [A. I.] 1927:78 [Reports of the Scientific Station of Fisheries in Kertch v. 1 (2-3); ref. 21154] Ionian Sea and Adriatic Sea (Mediterranean Sea). No types known. Named for "race Méditerranéenne, group oriental" Fage 1920. Infrasubspecific. Engraulidae.

albidus, Engraulis Borsa [P.], Collet [A.] & Durand [J.-D.] 2005:1122, Fig. 3 (upper) [Comptes Rendus Biologies v. 327 (no. 12) (for 2004); ref. 28607] Cul-de-Beauduc in Camargue, southern France, northwestern Mediterranean Sea. Holotype: MNHN 2002-1716. Paratypes: MNHN 2002-1717 to 1774. •Valid as Engraulis albidus Borsa, Collet & Durand 2005. Current status: Valid as Engraulis albidus Borsa, Collet & Durand 2005. Engraulidae. Distribution: Mediterranean Sea. Habitat: marine.

amara, Engraulis Risso [A.] 1827:456 [Histoire naturelle des principales productions de l'Europe méridionale v. 3; ref. 3757] Nice, France, northwestern Mediterranean Sea. No types known. Type catalog: Whitehead & Bauchot 1986:47 [ref. 9739]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

anchoita, Engraulis Hubbs [C. L.] & Marini [T. L.] in Marini 1935:418, Figs. 2, 4 (no. 2) [Physis (Buenos Aires) v. 11; ref. 17600] Mar del Plata, Province Buenos Aires, Argentina. Holotype: UMMZ 156821 or MACN 3591. Paratypes: MACN 1070 (3); UMMZ 95471 (2, now 1), 156820 (2); USNM 116438 [ex UMMZ 95471] (1). Type catalog: Braga & Piacentino 1994:99 [ref. 23697]. •Valid as Engraulis anchoita Hubbs & Marini 1935 -- (Nelson 1984:424 [ref. 5190], Nakamura in Nakamura et al. 1986:78 [ref. 14235], Whitehead et al. 1988:312 [ref. 5725], Cousseau & Perrotta 2000:64 [ref. 25942], López et al. 2002:63 [ref. 26808], Menezes & Figueiredo in Menezes et al. 2003:40 [ref. 27192], Mabragaña et al. 2011:table S2 [ref. 31800], Catelani et al. 2014:931 [ref. 37496], Nión et al. 2016:26 [ref. 35565], Spier et al. 2018:7 [ref. 36103], Ribeiro et al. 2019:6 [ref. 36717]). Current status: Valid as Engraulis anchoita Hubbs & Marini 1935. Engraulidae. Distribution: Southwestern Atlantic: Brazil south to Argentina. Habitat: marine.

antarcticus, Engraulis Castelnau [F. L.] 1872:186 [Proceedings of the Zoological and Acclimatisation Society of Victoria, Melbourne v. 1; ref. 757] Melbourne markets, Victoria, Australia. Syntypes: MNHN 0000-3732 (8). Type catalog: Bertin 1940:298 [ref. 293], Whitehead & Bauchot 1986:31 [ref. 9739]. •Synonym of Engraulis australis (Shaw 1790) -- (Whitehead et al. 1988:314 [ref. 5725], Paxton et al. 1989:160 [ref. 12442], Wongratana et al. 1999:1718 [ref. 24719], Paxton et al. 2006:312 [ref. 28995]). Current status: Synonym of Engraulis australis (Shaw 1790). Engraulidae. Habitat: marine.

antipodum, Engraulis encrasicholus var. Günther [A.] 1868:386 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Tasmania, Australia; New Zealand. Syntypes: (3) BMNH 2011.12.8.1 (1) Tasmania, BMNH 1855.9.19.1175 (2) New Zealand. •Synonym of Engraulis australis (Shaw 1790) -- (Whitehead et al. 1988:314 [ref. 5725], Paxton et al. 1989:160 [ref. 12442], Gomon et al. 1994:226 [ref. 22532], Wongratana et al. 1999:1718 [ref. 24719], Paxton et al. 2006:312 [ref. 28995]). Current status: Synonym of Engraulis australis (Shaw 1790). Engraulidae. Habitat: marine.

aquitanicus, Engraulis encrasicolus Aleksandrov [A. I.] 1927:77 [Reports of the Scientific Station of Fisheries in Kertch v. 1 (2-3); ref. 21154] Bay of Biscay and probably near shores of Portugal. No types known. After "race Atlantique, groupe méridional" Fage 1920. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

argyrophanus, Engraulis Valenciennes [A.] (ex Kuhl & van Hasselt) in Cuvier & Valenciennes 1848:49 [Histoire naturelle des poissons v. 21; ref. 1013] Equatorial Atlantic off western Africa. Lectotype: MNHN A-7614. Paralectotypes: MNHN B-3103 [ex MNHN A-7614] (4), RMNH 3331 (2). Type catalog: Whitehead & Bauchot 1986:31-32 [ref. 9739]. Lectotype selected by Whitehead 1967:125 [ref. 6464]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

atlantica, Engraulis encrassicholus forma de Buen [F.] 1926:49 [Resultado de las Campañas Realizadas por Acuerdos Internacionales, Madrid No. 2; ref. 5054] Gulf of Cádiz and coasts of Larache, Spain. Based on literature sources. Species should have been encrasicolus. Apparently independent of Pusanov & Zeeb 1926; earliest not reserached. •Synonym of Engraulis encrasicolus (Linnaeus 1758) Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

atlanticus, Engraulis encrasicholus Pusanov [I. I.] & Zeeb [Y.] 1926:93 [Trudy Krimskogo Naucno-Issledovatel'skogo Instituta Biologii i Biologiceskogo Fakul'teta v. 1; ref. 18576] Atlantic. After "race Atlantique" of Fage 1920. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

australis, Atherina Shaw [G.] in White 1790:296, Pl. 64 (opp. p. 296) [Journal of a voyage to New South Wales; ref. 4654] Between Broken Bay and Botany Bay, New South Wales, Australia. No types known. Authorship according to Sherborn 1891:535 [ref. 32516]. •Valid as Engraulis australis (Shaw 1790) -- (Nelson 1984:424 [ref. 5190], Whitehead et al. 1988:314 [ref. 5725], Paxton et al. 1989:160 [ref. 12442], Paulin et al. 1989:79 [ref. 24556], Kuiter 1993:36 [ref. 23929], Francis 1993:158 [ref. 25479], Gomon et al. 1994:226 [ref. 22532], Kuiter 1997:34 [ref. 25488], Wongratana et al. 1999:1718 [ref. 24719], Johnson 1999:722 [ref. 25471], Hutchins 2001:18 [ref. 25847], Paxton et al. 2006:312 [ref. 28995], Gomon 2008:194 [ref. 30616], Fricke et al. 2011:354 [ref. 31242], Duffy & Ahyong 2015:108 [ref. 36167], Stewart & Struthers 2015:300 [ref. 34231]). Current status: Valid as Engraulis australis (Shaw 1790). Engraulidae. Distribution: Southwestern Pacific: southern Australia and New Zealand to Kermadec Islands. Habitat: marine.

australis, Engraulis Steindachner [F.] 1879:31 [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Classe v. 16 (no. 4); ref. 20480] Hobson's Bay, 37°51'S, 144°56'E, Victoria, Australia. Syntypes: (original as 2) SMNS 2254 (3). Type catalog: Fricke 1995:13 [ref. 28100], Fricke 2005:37 [ref. 29864]. Species later described in more detail in Steindachner 1879:14 [ref. 4228]. Subjectively invalid; secondarily preoccupied in Engraulis by Atherina australis Shaw 1790. •Synonym of Engraulis australis (Shaw 1790) -- (Fricke 1995:13 [ref. 28100], Paxton et al. 2006:312 [ref. 28995]). 1°Homonym. Current status: Synonym of Engraulis australis (Shaw 1790). Engraulidae. Habitat: marine.

capensis, Engraulis Gilchrist [J. D. F.] 1913:62, Fig. [Marine biological report: for the year ended ... and for the half year ending ...; to be presented to the Provincial Council; Union of South Africa, Province of the Cape of Good Hope. No. 1 (art. 2); ref. 14154] South Africa. No types known. •Synonym of Engraulis japonicus Temminck & Schlegel 1846 -- (Whitehead & Wongratana 1986:205 [ref. 6284]). •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead 1990:228 [ref. 14039]). •Valid as Engraulis capensis Gilchrist 1913 -- (Whitehead et al. 1988:315 [ref. 5725], Bianchi & Carpenter in Bianchi et al. 1993:138 [ref. 25472], Fricke et al. 2018:52 [ref. 35805]). Current status: Valid as Engraulis capensis Gilchrist 1913. Engraulidae. Distribution: Southeastern Atlantic, Western Indian Ocean: South and East Africa to Madagascar and Mascarenes. Habitat: marine.

celebicus, Stolephorus Hardenberg [J. D. F.] 1933:262 [Natuurkundig Tijdschrift voor Nederlandsch Indië v. 93 (no. 2); ref. 16492] Manado, Sulawesi, Indonesia. Appeared first in, and probably dates to Hardenberg 1933:249 [ref. 18244] in table; full description as above. Also in Hardenberg 1934:327 [ref. 16426]. •Synonym of Engraulis japonicus Temminck & Schlegel 1846 -- (Whitehead et al. 1988:318 [ref. 5725], Wongratana et al. 1999:1719 [ref. 24719]). Current status: Synonym of Engraulis japonicus Temminck & Schlegel 1846. Engraulidae. Habitat: marine.

encrasicolus, Clupea Linnaeus [C.] 1758:318 [Systema Naturae, Ed. X v. 1; ref. 2787] 18 nautical miles off Sète, Gulf of Lyon, France, northwestern Mediterranean Sea. MNHN 2002-1775. Neotype designated by Fricke 1999:20 [ref. 24101], but withdrawn in Fricke 2000:639 [ref. 24537]. Neotype designated by Borsa et al. 2004:1113 [ref. 28607]. Spelled encrasiolus in Brünnich 1768:83 [ref. 17290]. Sometimes spelled encrasicholus or encrassicolus.. •Valid as Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Dor 1984:43 [ref. 29757], Nelson 1984:424 [ref. 5190], Whitehead in Whitehead et al. 1984:282 [ref. 13675], Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039], Goren & Dor 1994:11 [ref. 25356], Coad 1995:9 [ref. 23608], Coad 1996:84 [ref. 25304], Santos et al. 1997:121 [ref. 23531], Arruda 1997:30 [ref. 24952] as encrasicholus, Fricke 1999:20 [ref. 24101], Bilecenoğlu et al. 2002:39 [ref. 26753], Manilo & Bogorodsky 2003:S97 [ref. 27377], Bogutskaya & Naseka 2004:30 [ref. 28183], Borsa et al. 2005:1121 [ref. 28607], Golani 2005:20 [ref. 37112], Fricke 2007:26 [ref. 30577], Fricke et al. 2007:25 [ref. 29533], Vasil'eva 2007:44 [ref. 30517], Fricke 2008:15 [ref. 30182], Lipej & Dulčić 2010:20 [ref. 36649], Wirtz et al. 2013:118 [ref. 32972], Parin et al. 2014:51 [ref. 33547], Nizinski & Munroe 2016:1717 [ref. 34559], Bogorodsky & Randall 2018:[22] 260 [ref. 36492], Golani & Fricke 2018:32 [ref. 36273], Martins & Carneiro 2018:29 [ref. 36550], Artüz & Fricke 2019:553 [ref. 36612], Carneiro et al. 2019:74 [ref. 37250], Elbaraasi et al. 2019:92 [ref. 36864], Reiner 2019:79 [ref. 38139], Bariche & Fricke 2020:45 [ref. 37515], Kovačić et al. 2020:12 [ref. 37519], Golani 2021:20 [ref. 38303]). Current status: Valid as Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Distribution: Western Baltic Sea; North Sea; Mediterranean Sea; Sea of Marmara; Black Sea; Sea of Azov; eastern Atlantic: Norway south to South Africa, including Azores, Madeira, Canary Islands, and São Tomé and Principe; Red Sea (Mediterranean Sea immigrant); southwestern Indian Ocean. Habitat: brackish, marine.

estauquae, Anchoviella Hildebrand [S. F.] 1943:115, Fig. 48 [Bulletin of the Bingham Oceanographic Collection Yale University v. 8 (art. 2); ref. 13209] Sandy beach in Estauques Bay, Gulf of Venezuela. Holotype: USNM 119795. Paratypes: USNM 119796 (3). •Synonym of Engraulis eurystole (Swain & Meek 1884) -- (Whitehead et al. 1988:317 [ref. 5725], Nizinski & Munroe 2003:789 [ref. 26995]). Current status: Synonym of Engraulis eurystole (Swain & Meek 1884). Engraulidae. Habitat: marine.

eurystole, Stolephorus Swain [J.] & Meek [S. E.] 1884:35 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 36; ref. 18867] Woods Hole, Massachusetts, U.S.A. Syntypes: USNM 19003 to 19017. Based on Stolephorus perfasciatus of Jordan & Gilbert 1883:273 [ref. 2476] and Swain 1883:55 [ref. 38302] (not perfasciatus of Poey 1860) on one specimen that was lost; but Swain & Meek list additional specimens [syntypes] as, "Specimens of this species, perhaps mixed with others, have been distributed by the U.S. National Museum under the following number, 19,003 to 19,017. •Valid as Engraulis eurystole (Swain & Meek 1884) -- (Nelson 1984:424 [ref. 5190], Robins & Ray 1986:75 [ref. 23100], Whitehead et al. 1988:317 [ref. 5725], Scott & Scott 1988:119 [ref. 25518], McAllister 1990:54 [ref. 14674], Cervigón 1991:121 [ref. 24472], Boschung 1992:44 [ref. 23239], Cervigón 1992:315 [ref. 23827], McEachran & Fechhelm 1998:358 [ref. 23897], Aguilera 1998:46 [ref. 24221], Schmitter-Soto et al. 2000:150 [ref. 27754], Menezes & Figueiredo in Menezes et al. 2003:40 [ref. 27192], Nizinski & Munroe 2003:789 [ref. 26995], García-Hernández et al. 2009:91 [ref. 37265], Page et al. 2013:67 [ref. 32708]). Current status: Valid as Engraulis eurystole (Swain & Meek 1884). Engraulidae. Distribution: Western Atlantic: Maine (U.S.A.) south to Florida (U.S.A.); eastern Colombia to mouth of Amazon River (Brazil); including Gulf of Mexico and southern Caribbean Sea. IUCN (2015): Least concern. Habitat: marine.

fraseri, Engraulis australis Blackburn [M.] 1950:60, 67 [Australian Journal of Marine and Freshwater Research v. 1 (no. 1); ref. 12938] Western Australia. Syntypes: none designated; many specimens are listed on p. 84, Appendix G. •Synonym of Engraulis australis (Shaw 1790) -- (Whitehead et al. 1988:314 [ref. 5725], Wongratana et al. 1999:1718 [ref. 24719]). Current status: Synonym of Engraulis australis (Shaw 1790). Engraulidae. Habitat: marine.

guineensis, Anchoviella Rossignol [M.] & Blache [J.] 1961:285, Fig. 1 [Bulletin du Muséum National d'Histoire Naturelle (Série 2) v. 33 (no. 3); ref. 9375] Off Cape Lopez, Gabon, depth 20 meters. Syntypes: MNHN 1961-0310 (10). Additional material: ?MNHN [ex ORSTOM Dakar] uncat. (70). Type catalog: Whitehead & Bauchot 1986:30 [ref. 9739]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Nelson 1984:424 [ref. 5190], Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:229 [ref. 14039], Nizinski & Munroe 2016:1717 [ref. 34559]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

japonica, Argentina Forster [J. R.] 1795:16 [Zoologia Indica, sistens descriptiones animalium selectorum; ref. 18091] India. Name only. Based on Chaetodon Atherina japonica Houttuyn 1782. •Synonym of Engraulis japonicus Temminck & Schlegel 1846. Current status: Synonym of Engraulis japonicus Temminck & Schlegel 1846. Engraulidae. Habitat: marine.

japonica, Atherina Houttuyn [M.] 1782:340 [Verhandelingen der Hollandsche Maatschappij der Wetenschappen, Haarlem v. 20 (pt 2); ref. 2220] Name suppressed by ICZN (Opinion 749) for priority, but not for homonymy; see also Boeseman 1995:6 [ref. 21811]; from Japan. •In the synonymy of Engraulis japonicus Temminck & Schlegel 1846 -- (Whitehead et al. 1988:318 [ref. 5725], Mizuno et al. 1999:5 [ref. 25311], Nakabo 2000:248 [ref. 25086]). •Treated as valid as Engraulis japonicus (Houttuyn 1782) -- (Nakabo 2002:248 [ref. 26001], Matsunuma et al. 2010:82 [ref. 31558]). Suppressed. Current status: Synonym of Engraulis japonicus Temminck & Schlegel 1846. Engraulidae. Habitat: marine.

japonicus, Engraulis Temminck [C. J.] & Schlegel [H.] 1846:239, Pl. 108 (fig. 3) [Fauna Japonica Parts 10-14; ref. 4374] Japan. No types known. Based on figure and manuscript of Burger (Boeseman 1947:176 [ref. 12876]). •Valid as Engraulis japonicus Temminck & Schlegel 1846 -- (Uyeno & Sato in Masuda et al. 1984:20 [ref. 6441], Nelson 1984:424 [ref. 5190], Whitehead & Wongratana 1986:205 [ref. 6284], Whitehead et al. 1988:318 [ref. 5725], Youn & Kim 1996:37 [ref. 25727], Cheng & Zhou 1997:75 [ref. 26385], Sokolovskaya et al. 1998:8 [ref. 24670] with author as Houttuyn, Fricke 1999:77 [ref. 24106], Wongratana et al. 1999:1719 [ref. 24719], Munroe & Nizinski in Randall & Lim 2000:588 [ref. 25122], Sheiko & Fedorov 2000:17 [ref. 25650], Shimizu 2001:21 [ref. 25799], Wang et al. 2001:55 [ref. 26566], Zhang 2001:120 [ref. 26586], Mecklenburg et al. 2002:139 [ref. 25968], Choi et al. 2003:149, 495 [ref. 26218], Fedorov et al. 2003:24 [ref. 27579], Bogutskaya & Naseka 2004:31 [ref. 28183], Heemstra & Heemstra 2004:110 [ref. 28072], Love et al. 2005:25 [ref. 37547], Parin et al. 2014:52 [ref. 33547], Koeda et al. 2016:10 [ref. 35333], Kwun et al. 2017:141 [ref. 37287], Motomura et al. 2017:44 [ref. 35490] as japonica, Sonoyama et al. 2020:21 [ref. 37637] as japonica, Dyldin & Orlov 2021:62 [ref. 38158]). Current status: Valid as Engraulis japonicus Temminck & Schlegel 1846. Engraulidae. Distribution: Northwestern Pacific: Indonesia north to Korea, Japan and Kamchatka (Russia), straying to western Aleutian Islands (U.S.A.). Western Indian Ocean populations may belong to a separate species. Habitat: marine.

maeoticus, Engraulis encrasicholus Pusanov [I. I.] & Zeeb [Y.] 1926:94 [Trudy Krimskogo Naucno-Issledovatel'skogo Instituta Biologii i Biologiceskogo Fakul'teta v. 1; ref. 18576] Sea of Azov, Russia/Ukraine. Syntypes: whereabouts unknown. •Synonym of Engraulis encrasicolus (Linnaeus 1758), but a valid subspecies as described -- (Ninua & Japoshvili 2008:164 [ref. 31251]). •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725], Bogutskaya & Naseka 2004:30 [ref. 28183], Parin et al. 2014:51 [ref. 33547]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

mauii, Anchoviella Fowler [H. W.] & Bean [B. A.] 1923:4 [Proceedings of the United States National Museum v. 63 (no. 2488); ref. 1474] Maui Island, Hawaiian Islands [probably in error for eastern Pacific]. Holotype: USNM 82904. Paratypes: (1). Additional material: (9). •Synonym of Engraulis mordax Girard 1854 -- (Whitehead et al. 1988:320 [ref. 5725]). •Synonym of Engraulis australis (White 1790) -- (Mundy 2005:147 [ref. 28379]). Current status: Synonym of Engraulis mordax Girard 1854. Engraulidae. Habitat: marine.

mediterraneus, Engraulis encrasicolus Pusanov [I. I.] & Zeeb [Y.] 1926:93 [Trudy Krimskogo Naucno-Issledovatel'skogo Instituta Biologii i Biologiceskogo Fakul'teta v. 1; ref. 18576] Mediterranean Sea. After "race Méditerranéenne" Fage 1920. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725], Bogutskaya & Naseka 2004:30 [ref. 28183]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

meletta, Engraulis Cuvier [G.] 1829:323 [Le Règne Animal (Edition 2) v. 2; ref. 995] Mediterranean Sea. No types known. Type catalog: Whitehead & Bauchot 1986:49 [ref. 9739]. Based on brief description and citation to "Duham. sect. VI, pl. iii, f. 5" [Duhamel 1777:157-158, Pl. 3 (fig. 5)]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

mordax, Engraulis Girard [C. F.] 1854:138 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] San Francisco, California, U.S.A. No types known. •Valid as Engraulis mordax Girard 1854 -- (Eschmeyer & Herald 1983:74 [ref. 9277], Nelson 1984:424 [ref. 5190], Whitehead et al. 1988:320 [ref. 5725], Hammann & Cisneros-Mata 1989:49 [ref. 23109], Matarese et al. 1989:48 [ref. 26885], Swift et al. 1993:130 [ref. 22379], Whitehead & Rodriguez-Sánchez in Fischer et al. 1995:1082 [ref. 22829], De La Cruz Agüero et al. 1997:135 [ref. 24545], Fuller et al. 1999:41 [ref. 25838], Mecklenburg et al. 2002:138 [ref. 25968], Nelson et al. 2004:66 [ref. 27807], Love et al. 2005:25 [ref. 37547], Page et al. 2013:67 [ref. 32708], Pietsch & Orr 2015:19 [ref. 34694], Kells et al. 2016:90 [ref. 35888], Burton & Lea 2019:36 [ref. 37205]). Current status: Valid as Engraulis mordax Girard 1854. Engraulidae. Distribution: Eastern Pacific: eastern Gulf of Alaska (U.S.A.) south to Pacific coast of Baja California Sur (Mexico), including Gulf of California (Mexico). Habitat: marine.

nanus, Engraulis Girard [C. F.] 1858:335 [General report upon zoology of the several Pacific railroad routes, 1857; ref. 4911] San Francisco, California, U.S.A. Holotype: USNM 946. •Synonym of Engraulis mordax Girard 1854 -- (Whitehead et al. 1988:320 [ref. 5725]). Current status: Synonym of Engraulis mordax Girard 1854. Engraulidae. Habitat: marine.

nattereri, Engraulis Steindachner [F.] 1879:174 [56] [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 80 (nos 1-2); ref. 4229] Pará, Brazil. Holotype (unique): NMW (not found). Type information: Whitehead 1970:24 [ref. 11605]. On p. 56 of separate. •Valid as Anchoviella nattereri (Steindachner 1879) -- (Whitehead et al. 1988:336 [ref. 5725], Kullander & Ferraris in Reis et al. 2003:40 [ref. 27061], Menezes & Figueiredo in Menezes et al. 2003:39 [ref. 27192]). •Nomen nudum -- (Loeb 2012:14 [ref. 31849] dated 1880). •Nomen dubium as Engraulis nattereri Steindachner 1879 -- (Loeb & Menezes 2015:299 [ref. 33727] dated 1880). Nomen Dubium. Current status: Uncertain as Engraulis nattereri Steindachner 1879. Engraulidae. Distribution: Amazon River mouth, Brazil. Habitat: freshwater, brackish.

occidentalis, Engraulis encrasicholus ponticus natio Majorowa [A. A.] 1934:18 [Traudy Biologiceskogo Sektora / Akademija Nauk SSSR, Zakavkazskij Filial, Gruzinskoe Otdelenie = Biologiuro Sek'toris sromebi / SSRK Mec'nierebat'a Akademia, Amier-Kavkasiis P'iliali, Sak'art'velos Ganqop'ileba = Travaux biologiques / Académie des + v. 1 (no. 1); ref. 21155] Not available, infrasubspecific. •In the synonymy of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Infrasubspecific. Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae.

oliveri, Engraulis russoi var. Scuderi [I.] 1957:242 [Bollettino di Pesca, Piscicoltura e Idrobiologia, Roma (n. s.) v. 12 (no. 2); ref. 18715] Lago Verdi, brackish marine pond along the Tyrrhenian coast of Sicily near Oliveri, Italy. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

orientalis, Engraulis encrasicholus ponticus natio Majorowa [A. A.] 1934:19 [Traudy Biologiceskogo Sektora / Akademija Nauk SSSR, Zakavkazskij Filial, Gruzinskoe Otdelenie = Biologiuro Sek'toris sromebi / SSRK Mec'nierebat'a Akademia, Amier-Kavkasiis P'iliali, Sak'art'velos Ganqop'ileba = Travaux biologiques / Académie des + v. 1 (no. 1); ref. 21155] Apparently not available, infrasubspecific; from "Toto spatio partis orientalis Ponti." •In the synonymy of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Infrasubspecific. Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae.

pontica, Engraulis encrasicolus mediterraneus natio Pusanov [I. I.] & Zeeb [Y.] 1926:93 [Trudy Krimskogo Naucno-Issledovatel'skogo Instituta Biologii i Biologiceskogo Fakul'teta v. 1; ref. 18576] Sevastopol, Crimea, Ukraine, Black Sea. No types known. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Bogutskaya & Naseka 2004:30 [ref. 28183]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

ponticus, Engraulis encrasicholus Aleksandrov [A. I.] 1927:78 [Reports of the Scientific Station of Fisheries in Kertch v. 1 (2-3); ref. 21154] Western part of Black Sea. No types known. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Bogutskaya & Naseka 2004:31 [ref. 28183], Parin et al. 2014:51 [ref. 33547]). •Synonym of Engraulis encrasicolus (Linnaeus 1758), but a valid subspecies as described -- (Ninua & Japoshvili 2008:164 [ref. 31251]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Distribution: The subspecies occurs in the Black Sea basin. Habitat: brackish, marine.

pulchellus, Engraulis Girard [C. F.] 1855:199 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1819] Caldera Bay, Chile. No types known. •Synonym of Engraulis ringens Jenyns 1842 -- (Whitehead et al. 1988:322 [ref. 5725]). Current status: Synonym of Engraulis ringens Jenyns 1842. Engraulidae. Habitat: marine.

ringens, Engraulis Jenyns [L.] 1842:136 [The zoology of the voyage of H. M. S. Beagle; ref. 2344] Iquique, northern Chile. Syntypes: BMNH 1843.2.8.48 (1), 1917.7.14.1 (1). •Valid as Engraulis ringens Jenyns 1842 -- (Nelson 1984:424 [ref. 5190], Whitehead et al. 1988:322 [ref. 5725], Pequeño 1989:28 [ref. 14125], Grove & Lavenberg 1997:195 [ref. 24023], Chirichigno F. & Vélez D. 1998:183 [ref. 24555], McCosker & Rosenblatt 2010:189 [ref. 30957]). Current status: Valid as Engraulis ringens Jenyns 1842. Engraulidae. Distribution: Southeastern Pacific: Ecuador to Chile, including the Galápagos Islands. Habitat: marine.

russoi, Engraulis Dulzetto [F.] 1947:183, Figs. 1-6 [Memorie di matematica e di scienze fisiche e naturali della Società Italiana delle Scienze. v. 26; ref. 17424] Messina, Sicily, Italy, Mediterranean Sea. Neotype: MSNG 39522. Neotype designated by Tortonese 1967 [ref. 10366]. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:229 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

symaetensis, Engraulis encrasicholus Dulzetto [F.] 1938:397, Pls. 1 (fig. 2), 2 (fig. 4) [Memorie della Classe di Scienze Fisiche, Matematiche e Naturali, Reale Accademia d'Italia Roma v. 11 (no. 8); ref. 17423] Catania, eastern Sicily, Italy, Mediterranean Sea. Neotype: MSNG 39522. Neotype designated by Tortonese 1967. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Svetovidov 1973:111 [ref. 7169], Whitehead et al. 1988:316 [ref. 5725]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.

tapirulus, Engraulis Cope [E. D.] 1877:45 [29] [Proceedings of the American Philosophical Society v. 17; ref. 18033] Probably Bay of Pacasmayo, Peru. Syntypes: ANSP 21385-86 (1, 1). Type catalog: Böhlke 1984:101 [ref. 13621] as holotype (21385) and paratype. On p. 29 of separate. •Synonym of Engraulis ringens Jenyns 1842 -- (Whitehead et al. 1988:322 [ref. 5725]). Current status: Synonym of Engraulis ringens Jenyns 1842. Engraulidae. Habitat: marine.

vittargentea, Clupea Lacepède [B. G. E.] 1803:424, 458 [Histoire naturelle des poissons (Lacepède) v. 5; ref. 4930] Mauritius, Mascarenes, southwestern Indian Ocean. No types known. Type catalog: Whitehead & Bauchot 1986:47 [ref. 9739]. Based on Commerson manuscript. Status uncertain, Stolephorus or Engraulis (Whitehead & Bauchot 1985:46 [ref. 9739]). •Questionably the same as Engraulis capensis Gilchrist 1913 -- (Whitehead et al. 1988:315 [ref. 5725]). •Mention as incertae sedis -- (Fricke 1999:78 [ref. 24106]). Engraulidae. Habitat: freshwater.

vulgaris, Engraulis Nilsson [S.] 1832:25 [Prodromus ichthyologiae Scandinavicae; ref. 3204] Scandinavia. No types known. •Synonym of Engraulis encrasicolus (Linnaeus 1758) -- (Whitehead et al. 1988:316 [ref. 5725], Whitehead 1990:228 [ref. 14039]). Current status: Synonym of Engraulis encrasicolus (Linnaeus 1758). Engraulidae. Habitat: brackish, marine.