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Eschmeyer's Catalog of Fishes
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Species in the genus Oxynoemacheilus: [ 91 ] records


afrenatus, Nemachilus frenatus Battalgil [F.] 1941:183 [Revue de la Faculté des Sciences de l'Université d'Instanbul, Série B: Sciences Naturelles v. 6 (nos 1-2); ref. 13194] Small creek at Diyarbakir, tributary of the Tigris River, Turkey. No types known. Original genus is correctly Nemacheilus. •Synonym of Nemacheilus frenatus (Heckel 1843), but a valid subspecies as described -- (Coad 1991:18 [ref. 15702], Coad 1995:22 [ref. 23608]). •Synonym of Oxynoemacheilus frenatus (Heckel 1843), but a valid subspecies afrenatus (Battagil 1941 [or 1942]) -- (Prokofiev 2009:880 [ref. 30604], Freyhof et al. 2012:308 [ref. 31752]). •Synonym of Oxynoemacheilus frenatus (Heckel 1843) -- (Kottelat 2012:96 [ref. 32367], Freyhof et al. 2017:557 [ref. 35290]). •Possibly valid as Oxynoemacheilus afrenatus (Battalgil 1941) -- (Sayyadzadeh & Esmaeili 2020:203 [ref. 37967]). Current status: Synonym of Oxynoemacheilus frenatus (Heckel 1843). Nemacheilidae. Habitat: freshwater.

alasanicus, Nemacheilus angorae Elanidze [R. F.] 1983 [Ichthyofauna of Georgia [Eurasia]; ref. 31784] Upper reaches of Alazani River at Alvani village, left bank tributary of the Kura River, Georgia (Eurasia). No types known. Without author; authorship attributed to the book author as this is the only taxon in the book without an author. Original description not seen. •Synonym of Oxynoemacheilus angorae (Steindachner 1897) -- (Freyhof et al. 2012:308 [ref. 31752]). •Synonym of Oxymoemacheilus lenkoranensis (Abdurakhmanov 1962) -- (Esmaeili et al. 2010:372 [ref. 33391] but may be a different species, Kottelat 2012:98 [ref. 32367]). •Synonym of Oxynoemacheilus lenkoranensis (Abdurakhmanov 1962) -- (Jouladeh-Roudbar et al. 2020:173 [ref. 37983]). •Synonym of Oxynoemacheilus bergi (Gratzianov 1907) -- (Çiçek et al. 2018:233 [ref. 36249], Freyhof et al. 2021:142 [ref. 38237]). Current status: Synonym of Oxynoemacheilus bergi (Gratzianov 1907). Nemacheilidae. Habitat: freshwater.

amanos, Oxynoemacheilus Kaya [C.], Yoğurtçuoğlu [B.] & Freyhof [J.] 2021:560, Figs. 1-3 [Zootaxa 4938 (no. 5); ref. 38174] Spring İncesu at Hassa, Hatay province, Turkey, 36.7935, 36.5135. Holotype: FFR 15582. Paratypes: FFR 1478 (45), FSJF 2273 (32). Type catalog: Kaya et al. 2021:424 [ref. 38688]. •Valid as Oxynoemacheilus amanos Kaya, Yoğurtçuoğlu & Freyhof 2021. Current status: Valid as Oxynoemacheilus amanos Kaya, Yoğurtçuoğlu & Freyhof 2021. Nemacheilidae. Distribution: Asia Minor: spring İncesu at Hassa, northern Orontes basin, Hatay province, Turkey. Habitat: freshwater.

anatolica, Oxynemacheilus Erk'akan [F.], Özeren [S. C.] & Nalbant [T. T.] 2008:117, Figs. 4-6 [Journal of Fisheries International [electronic journal] v. 3 (no. 4); ref. 29859] Input of Karamanli Dam Lake, Burdur, southwestern Turkey. Holotype: HUIC-AKD-13. Paratypes: HUIC-AKD (14 specimens). Original genus should have been Oxynoemacheilus. •Valid as Oxynoemacheilus anatolicus Erk'akan, Özeren & Nalbant 2008 -- (Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:95 [ref. 32367], Çiçek et al. 2015:148 [ref. 35151], Çiçek et al. 2018:17 [ref. 36267], Çiçek et al. 2020:244 [ref. 37644], Yoğurtçuoğlu et al. 2022:457 [ref. 39236]). Current status: Valid as Oxynoemacheilus anatolicus Erk'akan, Özeren & Nalbant 2008. Nemacheilidae. Distribution: Asia Minor: Burdur River basin and upper Dalaman River basin, Burdur Province (Turkey). Habitat: freshwater.

angorae, Nemacheilus Steindachner [F.] 1897:693 [9], Pl. 4 (fig. 4a-c) [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 64; ref. 14691] Tabakane-Sir and Tschibuk-Tschai, Turkey. Syntypes: (16) NMW. Freyhof et al. 2012:307 [ref. 31752] •Valid as Orthrias angorae (Steindachner 1897) -- (Bănărescu et al. 1978:256 [ref. 9016], Erk'akan & Kuru 1986:160 [ref. 21167]). •Valid as Nemacheilus angorae Steindachner 1897 -- (Berg 1949:873 [ref. 20661] as Nemachilus, Coad 1991:18 [ref. 15702], Coad 1995:21 [ref. 23608], Coad 1996:71 [ref. 25297], Coad 1998:102 [ref. 23869], Gabrielyan 2001:28 [ref. 25867], Fricke et al. 2007:51 [ref. 29533]). •Valid as Barbatula angorae (Steindachner 1897) -- (Stoumboudi et al. 2006:143 [ref. 28767], Erk'akan et al. 2007:75 [ref. 29026], Ninua & Japoshvili 2008:167 [ref. 31251]). •Valid as Oxynoemacheilus angorae (Steindachner 1897) -- (Prokofiev 2009:880 [ref. 30604] with subspecies, Esmaeili et al. 2010:372 [ref. 33391], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Jouladeh-Roudbar et al. 2015:882 [ref. 34062], Çiçek et al. 2015:148 [ref. 35151], Jouladeh-Roudbar et al. 2016:103 [ref. 34789], Freyhof 2016:94 [ref. 34815], Esmaeili et al. 2017:62 [ref. 35254], Sungur et al. 2017:379 [ref. 35750], Çiçek et al. 2018:18 [ref. 36267], Çiçek et al. 2019:16 [ref. 36528], Turan et al. 2019:[6] [ref. 36540], Çiçek & Sungur 2020:117 [ref. 38064], Çiçek et al. 2020:245 [ref. 37644], Yogurtçuoglu et al. 2020:[14] [ref. 37627], Kelleci et al. 2021:282 [ref. 38359], Saygun et al. 2021:46 [ref. 38525], Yoğurtçuoğlu et al. 2022:458 [ref. 39236]). Current status: Valid as Oxynoemacheilus angorae (Steindachner 1897). Nemacheilidae. Distribution: Asia Minor: western and central Anatolian Black Sea basin (Marmara, Susurluk, Sakarya and Kızılırmak River basins) (Turkey). Habitat: freshwater.

araxensis, Orthrias angorae Bănărescu [P. M.] & Nalbant [T. T.] in Bănărescu, Nalbant & Balik 1978:259, Fig. 2; Pl. 20 (figs. 1-4) [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 75; ref. 9016] Kandili Karassu, upper Karassu drainage, Euphrates-Tigris basin, eastern Turkey. Holotype: ZMH 4827. Paratypes: ISBB 2617-18 (2, 3); ZMH 4826 (5), 5951 (4). Type catalog: Wilkens & Dohse 1993:413 [ref. 21161]. •Synonym of Nemacheilus angorae Steindachner 1897, but a valid subspecies as described -- (Erk'akan & Kuru 1986:160 [ref. 21167], Coad 1995:22 [ref. 23608]). •Valid as Orthrias araxensis Bănărescu & Nalbant 1978 -- (Nalbant & Bianco 1998:111 [ref. 23810], Fricke et al. 2007:51 [ref. 29533]). •Valid as Barbatula araxensis (Bănărescu & Nalbant 1978) -- (Erk'akan et al. 2007:78 [ref. 29026]). •Synonym of Oxynoemacheilus angorae (Steindachner 1897), but a valid subspecies -- (Prokofiev 2009:880 [ref. 30604]). •Valid as Oxynoemacheilus araxensis Bănărescu & Nalbant 1978 -- (Esmaeili et al. 2010:372 [ref. 33391], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:95 [ref. 32367], Jouladeh-Roudbar et al. 2015:883 [ref. 34062], Çiçek et al. 2015:148 [ref. 35151], Freyhof 2016:94 [ref. 34815], Esmaeili et al. 2017:62 [ref. 35254], Freyhof & Geiger 2017:259 [ref. 35343], Çiçek et al. 2018:240 [ref. 36249], Çiçek et al. 2018:18 [ref. 36267], Turan et al. 2019:[6] [ref. 36540], Çiçek et al. 2020:245 [ref. 37644], Rakici et al. 2020:410 [ref. 37960], Saygun et al. 2021:46 [ref. 38525]). Current status: Valid as Oxynoemacheilus araxensis (Bănărescu & Nalbant 1978). Nemacheilidae. Distribution: Asia Minor: upper Euphrates [Firat Nehri] (Persian Gulf tributary), Erzurum Province (Turkey). Habitat: freshwater.

argyrogramma, Cobitis Heckel [J. J.] 1847:239, Pl. 18 (fig. 3) [Reisen in Europa, Asien und Africa v. 2 (pt 3); ref. 2068] Aleppo, Syria. Syntypes: NMW 48541 (8), 59913 (4). •Synonym of Nemacheilus tigris (Heckel 1843) -- (Coad 1991:18 [ref. 15702], Coad 1995:23 [ref. 23608]). •Synonym or Orthrias tigris (Heckel 1843) -- (Fricke et al. 2007:52 [ref. 29533]). •Synonym of Barbatula tigris (Heckel 1843) -- (based on status of Barbatula). •Valid as Barbatula argyrogrammus (Heckel 1847) -- (Coad 2010:206 [ref. 36620]). •Valid as Oxynoemacheilus argyrogrammus (Heckel 1847) -- (Prokofiev 2009:880 [ref. 30604]) or a synonym or subspecies of Paracobitis tigris (Heckel 1843) -- (Prokofiev 2009:888 [ref. 30604]). •Valid as Oxynoemacheilus argyrogramma (Heckel 1847) -- (Golzarianpour et al. 2011:201 [ref. 31693], Freyhof et al. 2012:307, 308 [ref. 31752], Kottelat 2012:95 [ref. 32367], Kamangar et al. 2014:39, 49 [ref. 33140], Çiçek et al. 2015:148 [ref. 35151], Freyhof 2016:99 [ref. 34815], Freyhof & Özuluğ 2017:388 [ref. 35216], Esmaeili et al. 2017:63 [ref. 35254], Freyhof & Geiger 2017:259 [ref. 35343], Esmaeili et al. 2018:43 [ref. 36089], Turan et al. 2019:[6] [ref. 36540], Çiçek et al. 2020:245 [ref. 37644], Rakici et al. 2020:410 [ref. 37960]). Current status: Valid as Oxynoemacheilus argyrogramma (Heckel 1847). Nemacheilidae. Distribution: Asia Minor and Middle East: Euphrates and Tigris river basins (Turkey, Syria, Iraq, Iran). Habitat: freshwater.

arsaniasus, Oxynoemacheilus Freyhof [J.], Kaya [C.], Turan [D.] & Geiger [M. F.] 2019:44, Figs. 12-14 [Zootaxa 4612 (no. 1); ref. 36739] Stream Kaynarca at Tepe, Muş Province, Turkey. Holotype: FFR 15530. Paratypes: FFR, FSJF. Plus additional non-type material. Type catalog: Kaya et al. 2021:424 [ref. 38688]. •Valid as Oxynoemacheilus arsaniasus Freyhof, Kaya, Turan & Geiger 2019 -- (Çiçek et al. 2020:245 [ref. 37644]). Current status: Valid as Oxynoemacheilus arsaniasus Freyhof, Kaya, Turan & Geiger 2019. Nemacheilidae. Distribution: Asia Minor: Murat and upper Karasu (Bitlis) River drainages, upper Euphrates basin (Turkey). Habitat: freshwater.

atili, Oxynoemacheilus Erk'akan [F.] 2012:99, Fig. 3-4 [Research Journal of Biological Sciences v. 7 (no. 2); ref. 32281] Eflatun pinari-Beysehir Lake, 37°49'30.95"N, 31°40'29.08"E, Turkey. Holotype: HUIC uncat. Paratypes: HUIC (33). •Valid as Oxynoemacheilus atili Erk'akan 2012 -- (Çiçek et al. 2015:148 [ref. 35151], Çiçek et al. 2018:18 [ref. 36267], Bayçelebi et al. 2020:145 [ref. 37381], Çiçek et al. 2020:245 [ref. 37644], Yoğurtçuoğlu et al. 2021:375 [ref. 38333]). •Synonym of Oxynoemacheilus eregliensis (Bănărescu & Nalbant 1978) -- (Yoğurtçuoğlu et al. 2022:460 [ref. 39236]). Current status: Synonym of Oxynoemacheilus eregliensis (Bănărescu & Nalbant 1978). Nemacheilidae. Habitat: freshwater.

axylos, Oxynoemacheilus Yoğurtçuoğlu [B.], Kaya [C.] & Freyhof [J.] 2022:472, Figs. 14-16 [Zootaxa 5133 (no. 4); ref. 39236] Spring at Baltali, 30 km east of Haymana, Konya province, Turkey, 39.2393, 32.7501. Holotype: FFR 15616. Paratypes: FFR 15602 (30), 15620 (3). Plus additional non-type material. •Valid as Oxynoemacheilus axylos Yoğurtçuoğlu, Kaya & Freyhof 2022. Current status: Valid as Oxynoemacheilus axylos Yoğurtçuoğlu, Kaya & Freyhof 2022. Nemacheilidae. Distribution: Asia Minor: springs, streams and rivers in the endorheic Lake Tuz basin, Central Anatolia, Turkey. Habitat: freshwater.

banarescui, Orthrias brandti Delmastro [G. B.] 1982:53, Fig. 1 [Rivista Piemontese di Storia Naturale No. 3; ref. 8988] Devrekani creek near Devrekani, Anatolia, Turkey. Holotype: MCSNC. Paratypes: ISBB (4), IZA 8171 (4), MCSNC (80), MSNVR (4), MRSNT (4). •Synonym of Oxynoemacheilus brandtii (Kessler 1877), but a valid subspecies -- (Prokofiev 2009:880 [ref. 30604]). •Valid as Oxynoemacheilus banarescui (Delmastro 1982) -- (Stoumboudi et al. 2006:143 [ref. 28767], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:95 [ref. 32367], Çiçek et al. 2015:148 [ref. 35151], Çiçek et al. 2018:18 [ref. 36267], Turan et al. 2019:[6] [ref. 36540], Çiçek et al. 2020:245 [ref. 37644], Yogurtçuoglu et al. 2020:[14] [ref. 37627], Saygun et al. 2021:46 [ref. 38525], Freyhof et al. 2022:167 [ref. 39240]). Current status: Valid as Oxynoemacheilus banarescui (Delmastro 1982). Nemacheilidae. Distribution: Asia Minor: Black Sea tributaries, Kastamonu Province (Turkey). Habitat: freshwater.

bergamensis, Barbatula Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:71, Fig. 3 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Bergama, Kozak (Madra) Creek, 39°40'N, 27°28'E, Turkey. Holotype: HUIC KE-1. Paratypes: HUIC KE-1 (16). •Valid as Barbatula bergamensis Erk'akan, Nalbant & Özeren 2007 -- (Erk'akan et al. 2008:115 [ref. 29859]). •Valid as Oxynoemacheilus bergamensis (Erk'akan, Nalbant & Özeren 2007) -- (Prokofiev 2009:880 [ref. 30604]). •Synonym of Oxynoemacheilus theophilii (Stoumboudi, Kottelat & Barbieri 2006) -- (Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:99 [ref. 32367], Yoğurtçuoğlu et al. 2022:471 [ref. 39236]). Current status: Synonym of Oxynoemacheilus theophilii Stoumboudi, Kottelat & Barbieri 2006. Nemacheilidae. Habitat: freshwater.

bergi, Nemacheilus Gratzianov [V. I.] 1907:163, 167 [Trudy Otdela Ichtiologii Imperatorskago Russkago Obshchestva Akklimatizacii Zivotnych' i Rastenii. v. 4; ref. 1871] Akstapha [Akstafa] River, right tributary of Kura River, Azerbaijan. Syntypes: (2) ZMMU (not found in 2002). Based on 'Nemachilus spec.' of Berg 1898:30, 72 [ref. 20653]. •Synonym of Nemacheilus angorae Steindachner 1897 -- (Coad 1995:21 [ref. 23608], Prokofiev 2009:880 [ref. 30604]). •Synonym of Oxynoemacheilus angorae (Steindachner 1897) -- (Freyhof et al. 2012:308 [ref. 31752]). •Valid as Oxynoemacheilus bergi (Gratzianov 1907) -- (Çiçek et al. 2018:233 [ref. 36249], Çiçek et al. 2020:245 [ref. 37644], Freyhof et al. 2021:142 [ref. 38237]). Current status: Valid as Oxynoemacheilus bergi (Gratzianov 1907). Nemacheilidae. Distribution: Eurasia: Caspian Sea basin, Azerbaijan and possibly Iran. IUCN (2014): LC. Habitat: freshwater.

bergianus, Nemachilus Derjavin [A. N.] 1934:109, Fig. 8 [Trudy Azerbaidzhanskogo otdela Zakavkazskogo filiala Akademii Nauk SSSR, Sektor Zoologii v. 7; ref. 19591] Kisum village, Shah-rud River, Safid Rud basin, Iran. Holotype (unique): ZIN 25433. Original genus is correctly Nemacheilus. •Valid as Nemacheilus bergianus Derjavin 1934 -- (Berg 1949:873 [ref. 20661] as Nemachilus, Coad 1996:71 [ref. 25297], Coad 1998:102 [ref. 23869]). •Questionably valid or questionably a synonym of Nemacheilus angorae Steindachner 1897 -- (Coad 1995:21, 22 [ref. 23608]). •Valid as Orthrias bergianus (Derzhavin 1934) -- (Nalbant & Bianco 1998:111 [ref. 23810]). •Valid as Barbatula bergianus (Derzhavin 1934) -- (based on status of Barbatula). •Synonym of Oxynoemacheilus brandtii (Kessler 1877), but a valid subspecies -- (Prokofiev 2009:880 [ref. 30604]). •Valid as Oxynoemacheilus bergianus (Derjavin 1934) -- (Esmaeili et al. 2010:372 [ref. 33391], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:95 [ref. 32367], Kamangar et al. 2014:43 [ref. 33140], Jouladeh-Roudbar et al. 2015:883 [ref. 34062], Teimori et al. 2015:90 [ref. 34095], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:99 [ref. 34815], Sayyadzadeh et al. 2016:247 [ref. 34976], Freyhof & Abdullah 2017:84 [ref. 35169], Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:64 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Eagderi et al. 2018:458 [ref. 35940], Esmaeili et al. 2018:43 [ref. 36089], Çiçek et al. 2018:240 [ref. 36249], Çiçek et al. 2020:245 [ref. 37644], Jouladeh-Roudbar et al. 2020:162 [ref. 37983], Kuljanishvili et al. 2020:[6] [ref. 37535], Rakici et al. 2020:410 [ref. 37960], Çiçek et al. 2021:173 [ref. 38560], Freyhof et al. 2021:142 [ref. 38237], Saygun et al. 2021:47 [ref. 38525], Freyhof et al. 2022:168 [ref. 39240]). Current status: Valid as Oxynoemacheilus bergianus (Derjavin 1934). Nemacheilidae. Distribution: Eurasia: Caspian Sea, Urmia and Namak Lake basins. Habitat: freshwater.

brandtii, Nemachilus Kessler [K. F.] 1877:174, Pl. 6 (fig. 23) [The Aralo-Caspian Expedition; ref. 2597] Upper Kura River at Tbilis, Georgia, Eurasia. Syntypes: (5) BMNH 1897.7.5.39 [ex ZIN] (1) Tiflis, ZIN 2923 (4, lost in 1924). Original genus is correctly Nemacheilus. •Valid as Nemacheilus brandtii Kessler 1877 -- (Berg 1949:876 [ref. 20661] as Nemachilus brandti, Coad 1995:22 [ref. 23608] as brandti, Coad 1998:102 [ref. 23869] as brandti). •Valid as Orthrias brandtii (Kessler 1877) -- (Bănărescu et al. 1978:260 [ref. 9016] as brandti, Nalbant & Bianco 1998:111 [ref. 23810] as brandti, Fricke et al. 2007:51 [ref. 29533]). •Valid as Barbatula brandtii (Kessler 1877) -- (Erk'akan et al. 2007:78 [ref. 29026]). •Valid as Oxynoemacheilus brandtii (Kessler 1877) -- (Ninua & Japoshvili 2008:168 [ref. 31251], Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kamangar et al. 2014:43 [ref. 33140], Jouladeh-Roudbar et al. 2015:883 [ref. 34062], Çiçek et al. 2015:148 [ref. 35151], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:94 [ref. 34815], Sayyadzadeh et al. 2016:248 [ref. 34976], Esmaeili et al. 2017:64 [ref. 35254], Eagderi et al. 2018:458 [ref. 35940], Esmaeili et al. 2018:44 [ref. 36089], Çiçek et al. 2018:240 [ref. 36249], Turan et al. 2019:[10] [ref. 36540], Çiçek et al. 2020:245 [ref. 37644], Jouladeh-Roudbar et al. 2020:163 [ref. 37983], Kaya et al. 2020:339 [ref. 37582], Kuljanishvili et al. 2020:[6] [ref. 37535], Çiçek et al. 2021:173 [ref. 38560], Freyhof et al. 2021:135 [ref. 38237], Saygun et al. 2021:47 [ref. 38525]). Current status: Valid as Oxynoemacheilus brandtii (Kessler 1877). Nemacheilidae. Distribution: Eurasia: Kura and Aras River drainages, Caspian Sea basin (Georgia, Turkey and Iran). Habitat: freshwater.

bureschi, Nemachilus Drensky [P.] 1928:160, Fig. 1 [Izvestija na Carskite Prirodonaucni Instituti va Sofija = Mitteilungen aus den Königlichen Naturwissenschaftlichen Instituten in Sofia, Bulgraien = Bulletin des Institutions Royales d'Histoire Naturelle à Sofia, Bulgarie v. 1; ref. 15891] Bezirk Radomir and a tributary of the Struma River in Bezirk Küstendil and Dupniza, Bulgaria. Syntypes: NMNHS (2). Original genus should have been Nemacheilus •Synonym of Orthrias brandtii (Kessler 1877), but a valid subspecies -- (Bănărescu et al. 1978:261 [ref. 9016], Bănărescu & Nalbant 1995:449 [ref. 23187]). •Valid as Barbatula bureschi (Drensky 1928) -- (Kottelat 1997:94 [ref. 22952], Vassilev & Pehlivanov 2005:172 [ref. 28369], Fricke et al. 2007:50 [ref. 29533]). •Valid as Oxynoemacheilus bureschi (Drensky 1928) -- (Stoumboudi et al. 2006:143 [ref. 28767], Kottelat & Freyhof 2007:333 [ref. 29996], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:95 [ref. 32367], Barbieri et al. 2015:75 [ref. 35029], Çiçek et al. 2015:148 [ref. 35151], Esmaeili et al. 2017:63 [ref. 35254], Çiçek et al. 2020:245 [ref. 37644]). Current status: Valid as Oxynoemacheilus bureschi (Drensky 1928). Nemacheilidae. Distribution: Eurasia: Aegean Sea watershed (Macedonia, Greece, Bulgaria and Turkey). Habitat: freshwater.

cemali, Oxynoemacheilus Turan [D.], Kaya [C.], Kalayci [G.], Bayçelebi [E.] & Aksu [I.] 2019:[2], Figs.1-4 [Journal of Fish Biology v.94 (no. 3); ref. 36540] Stream Yaglı, about 2 km southeast of lncesu, Erzurum Province, Turkey, 40°18'30.49"N, 41°00'24.36"E. Holotype: FFR 15504. Paratypes: FFR. Plus additional non-type material. Type catalog: Kaya et al. 2021:425 [ref. 38688]. •Valid as Oxynoemacheilus cemali Turan, Kaya, Kalayci, Bayçelebi & Aksu 2019 -- (Kuljanishvili et al. 2020:[6] [ref. 37535], Çiçek et al. 2020:245 [ref. 37644], Saygun et al. 2021:47 [ref. 38525]). Current status: Valid as Oxynoemacheilus cemali Turan, Kaya, Kalayci, Bayçelebi & Aksu 2019. Nemacheilidae. Distribution: Eurasia: Çoruh and upper Yesilırmak River drainages, Black Sea basin (Turkey and probably Georgia). Habitat: freshwater.

ceyhanensis, Schistura Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:80, Fig. 10 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Yalak Village, Elbistan, Kahramanmaras Province, 38°39'N, 36°37'E, Turkey. Holotype: HUIC CEY-1. Paratypes: HUIC CEY-1 (3). •Valid as Oxynoemacheilus ceyhanensis (Erk'akan, Nalbant & Özeren 2007) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:95 [ref. 32367], Çiçek et al. 2015:148 [ref. 35151], Çiçek et al. 2018:18 [ref. 36267], Bayçelebi 2020:758 [ref. 37964], Çiçek et al. 2020:245 [ref. 37644], Kaya et al. 2020:289 [ref. 37622]). Current status: Valid as Oxynoemacheilus ceyhanensis (Erk'akan, Nalbant & Özeren 2007). Nemacheilidae. Distribution: Asia Minor: Ceyhan River basin (Mediterranean tributary), Kahramanmaraş Province (Turkey). Habitat: freshwater.

chomanicus, Oxynoemacheilus Kamangar [B. B.], Prokofiev [A. M.], Ghaderi [E.] & Nalbant [T. T.] 2014:46, Fig. 5 [Zootaxa 3755 (no. 1); ref. 33140] Baneh River, (Korhe-Pazi sta. 8), Baneh, Kurdistan, Iran, 36°01'03"N, 45°55'20"E. Holotype: FCFUK 176. Paratypes: FCFUK 177-209 (33), ZMUU 23066 (4). Plus non-type specimens. •Valid as Oxynoemacheilus chomanicus Kamangar, Prokofiev, Ghaderi & Nalbant 2014 -- (Jouladeh-Roudbar et al. 2015:883 [ref. 34062], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:94 [ref. 34815], Sayyadzadeh et al. 2016:248 [ref. 34976], Freyhof & Abdullah 2017:84 [ref. 35169], Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:64 [ref. 35254], Freyhof et al. 2017:559 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Esmaeili et al. 2018:44 [ref. 36089], Çiçek et al. 2020:245 [ref. 37644], Jouladeh-Roudbar et al. 2020:164 [ref. 37983]). Current status: Valid as Oxynoemacheilus chomanicus Kamangar, Prokofiev, Ghaderi & Nalbant 2014. Nemacheilidae. Distribution: Asia Minor and Middle East: Tigris River basin (Turkey, Iraq, Iran). Habitat: freshwater.

ciceki, Oxynoemacheilus Sungur [S.], Jalili [P.] & Eagderi [S.] 2017:376, Figs. 1-5 [Iranian Journal of Ichthyology v. 4 (no. 4); ref. 35750] Sultan Marsh, Kayseri Province, Turkey, 38°23'23.53"N, 35°21'54.52"E. Holotype: NHVUIC 2017-03-15-h. Paratypes: IMNRF-UT, NHVUIC. •Valid as Oxynoemacheilus ciceki Sungur, Jalili & Eagderi 2017 -- (Çiçek et al. 2018:18 [ref. 36267], Çiçek & Sungur 2020:117 [ref. 38064], Çiçek et al. 2020:245 [ref. 37644], Yoğurtçuoğlu et al. 2021:376 [ref. 38333]). Current status: Valid as Oxynoemacheilus ciceki Sungur, Jalili & Eagderi 2017. Nemacheilidae. Distribution: Asia Minor: Sultan Marsh, eastern Central Anatolia, Kayseri Province (Turkey). Habitat: freshwater.

cilicicus, Oxynoemacheilus Kaya [C.], Turan [D.], Bayçelebi [E.], Kalayci [G.] & Freyhof [J.] 2020:289, Figs. 2-5 [Zootaxa 4808 (no. 2); ref. 37622] Irrigation canal 6 km southeast of Silifke, Mersin Province, Turkey, 36.3465, 33.9806. Holotype: FFR 15579. Paratypes: FFR 15560 (18); FSJF 4098 (5). Plus additional non-type material. Type catalog: Kaya et al. 2021:425 [ref. 38688]. •Valid as Oxynoemacheilus cilicicus Kaya, Turan, Bayçelebi, Kalayci & Freyhof 2020 -- (Bayçelebi 2020:758 [ref. 37964], Kaya et al. 2021:564 [ref. 38174]). Current status: Valid as Oxynoemacheilus cilicicus Kaya, Turan, Bayçelebi, Kalayci & Freyhof 2020. Nemacheilidae. Distribution: Asia Minor: Lower Göksu, Seyhan and Ceyhan River basins, southern Anatolia (Turkey). Habitat: freshwater.

cinica, Barbatula Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:73, Fig. 4 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Road of Kütahya to Denizli, Cin Stream, upper Emir drainage, 37°40'41.07"N, 29°30'13.09"E [corrected], Turkey. Holotype: HUIC BM-3. Paratypes: HUIC BM-3 (23). •Valid as Barbatula cinica Erk'akan, Nalbant & Özeren 2007 -- (Erk'akan et al. 2008:117 [ref. 29859]). •Valid as Oxynoemacheilus cinicus (Erk'akan, Nalbant & Özeren 2007) -- (Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:96 [ref. 32367], Çiçek et al. 2015:148 [ref. 35151], Çiçek et al. 2018:18 [ref. 36267], Çiçek et al. 2020:245 [ref. 37644]). •Synonym of Oxynoemacheilus germencicus (Erk'akan, Nalbant & Özeren 2007) -- (Yoğurtçuoğlu et al. 2022:463 [ref. 39236]). Current status: Synonym of Oxynoemacheilus germencicus (Erk'akan, Nalbant & Özeren 2007). Nemacheilidae. Habitat: freshwater.

cyri, Nemacheilus tigris Berg [L. S.] 1910:170 [Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 15; ref. 19593] Upper reaches of the Kura River (Göle depression), near Okam village, Ardahan Province, Turkey. Syntypes: (several) ZIN 13291 (6+), 16885 (2). Also in Berg 1910:128 [ref. 14429], earliest not researched. •Synonym of Nemacheilus tigris (Heckel 1843), but a valid subspecies as described -- (Berg 1949:878 [ref. 20661] as nemachilus, Coad 1995:23 [ref. 23608]). •Valid as Orthrias cyri (Berg 1910) -- (Fricke et al. 2007:51 [ref. 29533]). •Valid as Barbatula cyri (Berg 1910) -- (authors). •Synonym of Paracobitis tigris (Heckel 1843), but a valid subspecies -- (Prokofiev 2009:888 [ref. 30604]). •Valid as Oxynoemacheilus cyri (Berg 1910) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:96 [ref. 32367], Çiçek et al. 2015:148 [ref. 35151], Esmaeili et al. 2017:63 [ref. 35254], Çiçek et al. 2018:18 [ref. 36267], Eagderi et al. 2018:458 [ref. 35940], Çiçek et al. 2018:240 [ref. 36249], Turan et al. 2019:[9] [ref. 36540], Çiçek et al. 2020:245 [ref. 37644], Kaya et al. 2020:339 [ref. 37582], Çiçek et al. 2021:173 [ref. 38560], Saygun et al. 2021:47 [ref. 38525]). Current status: Valid as Oxynoemacheilus cyri (Berg 1910). Nemacheilidae. Distribution: Asia Minor: Kura Nehri basin (Caspian Sea tributary), Ardahan Province (Turkey). Habitat: freshwater.

dori, Orthrias Goren [M.] & Bănărescu [P. M.] in Bănărescu, Nalbant & Goren 1982:12, Figs. 9-13 [Israel Journal of Zoology v. 31 (nos 1-2); ref. 174] Northern Malqoah [Nahal Malkoah], Bet-She'an Valley, Israel, about 32°25'N, 35°30'E, Jordan River system. Holotype: TAU 8278 [= SMNHTAU]. Paratypes: TAU [= SMNHTAU] 8270 and ISBB 3186 (6), TAU [= SMNHTAU] 7063 (25). •Synonym of Nemacheilus insignis (Heckel 1843) -- (Krupp & Schneider 1989:380 [ref. 13651]). •Valid as Oxynoemacheilus dori Goren & Bănărescu 1982 -- (Prokofiev 2009:880 [ref. 30604]). •Synonym of Oxynoemacheilus insignis (Heckel 1843) -- (Freyhof et al. 2012:309 [ref. 31752], Kottelat 2012:97 [ref. 32367]). Current status: Synonym of Oxynoemacheilus insignis (Heckel 1843). Nemacheilidae. Habitat: freshwater.

eliasi, Oxynoemacheilus Yoğurtçuoğlu [B.], Kaya [C.] & Freyhof [J.] 2022:478, Figs. 18-20 [Zootaxa 5133 (no. 4); ref. 39236] Inlet of Tahtalı Reservoir, under Şaşal bridge, Izmir province, Turkey, 38.1990, 27.1705. Holotype: FFR 15619. Paratypes: FFR 15581 (21), FSJF 1878 (6). Plus additional non-type material. •Valid as Oxynoemacheilus eliasi Yoğurtçuoğlu, Kaya & Freyhof 2022. Current status: Valid as Oxynoemacheilus eliasi Yoğurtçuoğlu, Kaya & Freyhof 2022. Nemacheilidae. Distribution: Asia Minor: Gediz, Küçük Menderes and Tahtalı River drainages, western Anatolia, Turkey. Habitat: freshwater.

elsae, Oxynoemacheilus Eagderi [S.], Jalili [P.] & Çiçek [E.] 2018:454, Figs. 1-4 [FishTaxa v. 3 (no. 2); ref. 35940] Zarineh River, near Shahin-Dej city, Lake Urmia basin, west Azerbaijan Province, Iran, 36°37'40"N, 46°43'30"E. Holotype: IMNRF-UT-1404-H. Paratypes: IMNRF-UT. •Valid as Oxynoemacheilus elsae Eagderi, Jalili & Çiçek 2018 -- (Esmaeili et al. 2018:44 [ref. 36089], Jouladeh-Roudbar et al. 2020:164 [ref. 37983], Kaya 2020:327 [ref. 39101], Freyhof et al. 2021:135 [ref. 38237], Çiçek et al. 2021:173 [ref. 38560] as cf. elsae, Freyhof et al. 2021:144 [ref. 38237]). Current status: Valid as Oxynoemacheilus elsae Eagderi, Jalili & Çiçek 2018. Nemacheilidae. Distribution: Middle East: Lake Urmia basin (Iran; Turkey). Habitat: freshwater.

ercisianus, Orthrias angorae Erk'akan [F.] & Kuru [M.] 1986:161, Fig. 1a [Doga Turk Biyoloji Dergisi. v. 10 (no. 2); ref. 21167] Ercis stream, Lake Van basin, Turkey. Holotype: HUIC. Paratypes: HUIC (4+9+3). •Synonym of Oxynoemacheilus angorae (Steindachner 1897), but a valid subspecies -- (Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693]). •Valid as Oxynoemacheilus ercisianus (Erk'akan & Kuru 1986) -- (Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:96 [ref. 32367], Kamangar et al. 2014:45 [ref. 33140], Çiçek et al. 2015:148 [ref. 35151], Freyhof & Geiger 2017:259 [ref. 35343], Çiçek et al. 2018:18 [ref. 36267], Şen et al. 2019:67 [ref. 36474], Turan et al. 2019:[9] [ref. 36540], Freyhof et al. 2019:34 [ref. 36739], Çiçek et al. 2020:245 [ref. 37644]). Current status: Valid as Oxynoemacheilus ercisianus (Erk'akan & Kuru 1986). Nemacheilidae. Distribution: Asia Minor: Lake Van basin endemic, Van Province (Turkey). Habitat: freshwater.

erdali, Barbatula Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:78, Fig. 8 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Murat River-Agri, 39°40'N, 43°44'E, Turkey. Holotype: HUIC F12. Paratypes: HUIC F12 (7). •Valid as Oxynoemacheilus erdali (Erk'akan, Nalbant & Özeren 2007) -- (Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kamangar et al. 2014:43 [ref. 33140], Çiçek et al. 2015:148 [ref. 35151], Freyhof & Özuluğ 2017:379 [ref. 35216], Freyhof & Geiger 2017:259 [ref. 35343], Çiçek et al. 2018:19 [ref. 36267], Turan et al. 2019:[9] [ref. 36540]). •Synonym of Oxynoemacheilus bergianus (Derjavin 1934) -- (Freyhof et al. 2019:33 [ref. 36739], Jouladeh-Roudbar et al. 2020:162 [ref. 37983], Freyhof et al. 2022:168 [ref. 39240]). Current status: Synonym of Oxynoemacheilus bergianus (Derjavin 1934). Nemacheilidae. Habitat: freshwater.

eregliensis, Orthrias angorae Bănărescu [P. M.] & Nalbant [T. T.] in Bănărescu, Nalbant & Balik 1978:258, Fig. 1 [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 75; ref. 9016] Eregli, southwestern central Turkey. Holotype: ZMH 1921. Paratypes: ZMH 5637 (4). Type catalog: Wilkens & Dohse 1993:413 [ref. 21161]. •Synonym of Orthrias angorae (Steindachner 1897), but a valid subspecies as described -- (Erk'akan & Kuru 1986:160 [ref. 21167]). •Valid as Barbatula eregliensis (Bănărescu & Nalbant 1978) -- (Erk'akan et al. 2007:74 [ref. 29026]). •Synonym of Nemacheilus angorae Steindachner 1897 -- (Fricke et al. 2007:51 [ref. 29533]). •Synonym of Oxynoemacheilus angorae (Steindachner 1897), but a valid subspecies -- (Prokofiev 2009:880 [ref. 30604]). •Valid as Oxynoemacheilus eregliensis (Bănărescu & Nalbant 1978) -- (Stoumboudi et al. 2006:143 [ref. 28767], Erk'akan et al. 2008:117 [ref. 29859] as Oxynemacheilus, Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:96 [ref. 32367], Çiçek et al. 2015:148 [ref. 35151], Çiçek et al. 2018:19 [ref. 36267], Çiçek et al. 2020:245 [ref. 37644], Kelleci et al. 2021:282 [ref. 38359], Yoğurtçuoğlu et al. 2022:460 [ref. 39236]). Current status: Valid as Oxynoemacheilus eregliensis (Bănărescu & Nalbant 1978). Nemacheilidae. Distribution: Asia Minor: Manavgat River basin and Central Anatolian lake tributaries (Turkey). Habitat: freshwater.

euphraticus, Noemacheilus insignis Bănărescu [P. M.] & Nalbant [T. T.] 1964:175, Pl. 7 (figs. 11-12) [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 61; ref. 217] Euphrates (Firat Nehri) Basin, Malatya, eastern Anatolia, Turkey. Holotype: ZMH 1889. Paratypes: ZMH H1890 (36). Also 10 additional non-type specimens in ZMH. Type catalog: Wilkens 1977:158 [ref. 22013]. Original genus is correctly Nemacheilus. •Synonym of Orthrias insignis (Heckel 1847), but a valid subspecies euphratica (Bănărescu & Nalbant 1964) -- (Bănărescu et al. 1978:264 [ref. 9016]). •Valid as Nemacheilus euphraticus Bănărescu & Nalbant 1964 -- (Nalbant & Bianco 1998:110 [ref. 23810]). •Valid as Orthrias euphraticus (Bănărescu & Nalbant 1964) -- (Nalbant 1998:371 [ref. 26011], Fricke et al. 2007:51 [ref. 29533]). •Valid as Barbatula euphratica (Bănărescu & Nalbant 1964) -- (authors). •Synonym of Oxynoemacheilus insignis (Heckel 1847), but a valid subspecies -- (Prokofiev 2009:880 [ref. 30604]). •Synonym of Oxynoemacheilus argyrogramma (Heckel 1847) -- (Freyhof et al. 2012:308 [ref. 31752], Kottelat 2012:95 [ref. 32367], Kamangar et al. 2014:39 [ref. 33140]). •Valid as Oxynoemacheilus euphraticus (Bănărescu & Nalbant 1964) -- (Freyhof 2016:99 [ref. 34815], Freyhof & Abdullah 2017:84 [ref. 35169], Freyhof & Özuluğ 2017:385 [ref. 35216], Esmaeili et al. 2017:64 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Esmaeili et al. 2018:44 [ref. 36089], Turan et al. 2019:[9] [ref. 36540], Çiçek et al. 2020:245 [ref. 37644], Jouladeh-Roudbar et al. 2020:165 [ref. 37983], Rakici et al. 2020:410 [ref. 37960], Saygun et al. 2021:47 [ref. 38525]). Current status: Valid as Oxynoemacheilus euphraticus (Bănărescu & Nalbant 1964). Nemacheilidae. Distribution: Asia Minor and Middle East: Euphrates River and Tigris River drainages (Turkey, Syria, Iraq and Iran). Habitat: freshwater.

evreni, Schistura Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:82, Fig. 12 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Tekir Stream, Göksu Basin, 38°39'N, 36°37'E, Turkey. Holotype: SEY-3. Paratypes: HUIC CEY-1 (2). •Valid as Oxynoemacheilus evreni (Erk'akan, Nalbant & Özeren 2007) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:96 [ref. 32367], Çiçek et al. 2015:148 [ref. 35151], Çiçek et al. 2018:19 [ref. 36267], Bayçelebi 2020:758 [ref. 37964], Çiçek et al. 2020:245 [ref. 37644], Kaya et al. 2020:289 [ref. 37622], Freyhof et al. 2021:138 [ref. 38277]). Current status: Valid as Oxynoemacheilus evreni (Erk'akan, Nalbant & Özeren 2007). Nemacheilidae. Distribution: Asia Minor: Ceyhan River basin (Mediterranean tributary), Kayseri Province (Turkey). Habitat: freshwater.

farsicus, Orthrias Nalbant [T. T.] & Bianco [P. G.] 1998:111, Fig. 6 [Italian Journal of Zoology v. 65, Suppl.; ref. 23810] River Kor near Persepolis, Iran. Holotype: IZA 7823. Paratypes: ISBB 3442 (7), 3446 (7), 3451 (7); IZA 7824-26 (64, 61, 31), 7845 (7). •Valid as Barbatula farsica (Nalbant & Bianco 1998) -- (based on status of Barbatula). •Valid as Oxynoemacheilus farsicus (Nalbant & Bianco 1998) -- (Prokofiev 2009:880 [ref. 30604], Esmaeili et al. 2010:372 [ref. 33391]). •Synonym of Oxynoemacheilus persa (Heckel 1847) -- (Freyhof et al. 2012:309 [ref. 31752], Kottelat 2012:98 [ref. 32367], Freyhof 2016:100 [ref. 34815], Esmaeili et al. 2017:67 [ref. 35254], Sayyadzadeh et al. 2018:45 [ref. 36026], Jouladeh-Roudbar et al. 2020:175 [ref. 37983]). Current status: Synonym of Oxynoemacheilus persa (Heckel 1847). Nemacheilidae. Habitat: freshwater.

fatsaensis, Oxynoemacheilus Saygun [S.], Ağdamar [S.] & Özuluğ [M.] 2021:40, Figs. 2-6 [Zoologischer Anzeiger v. 294; ref. 38525] Elekçi stream, Ordu Province, northern Anatolia, Turkey, 40.53200N, 37.23390E. Holotype: IUSHM 2021-1449. Paratypes: IUSHM 2021-1450 (6), 2021-1451 (8). Plus additional non-type material. •Valid as Oxynoemacheilus fatsaensis Saygun, Ağdamar & Özuluğ 2021 -- (Freyhof et al. 2022:176 [ref. 39240]). Current status: Valid as Oxynoemacheilus fatsaensis Saygun, Ağdamar & Özuluğ 2021. Nemacheilidae. Distribution: Asia Minor: Elekçi stream drainage, northern Anatolia, Turkey. Habitat: freshwater.

flavus, Nemacheilus Berg [L. S.] 1905:193, Pl. 6 (figs. 3-5) [Izviestii Turkestanskago otdiela Russkago geograficheskago obschchestva v. 4; ref. 15338] Lower Amu Darya River and Upper Syr Darya River, Turkmenistan. Lectotype: ZIN 13282. Paralectotypes: MLSU 297 (1); ZIN 10306 (2), 13282-83 (10, 2). Lectotype established in caption to Fig. 625 in Berg 1949:880 [ref. 20661] if figured specimen is identifiable. •Synonym of Nemacheilus oxianus Kessler 1877 -- (Berg 1949:879 [ref. 20661] as Nemachilus). •Synonym of 'Oxynoemacheilus' oxianus (Kessler 1877) -- (Kottelat 2012:98 [ref. 32367]). Current status: Synonym of Oxynoemacheilus oxianus (Kessler 1877). Nemacheilidae. Habitat: freshwater.

frenata, Cobitis Heckel [J. J.] 1843:1086 [96] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Tigris River, Mosul, Iraq. Syntypes: NMW 48552 (5), NRM 15477 (1). Illustrated by Heckel 1843, Pl. 12 (fig. 1) [ref. 2067]. On p. 96 of separate [ref. 2067]. •Valid as Nemacheilus frenatus (Heckel 1843) -- (Coad 1991:18 [ref. 15702], Coad 1995:22 [ref. 23608], Coad 1998:103 [ref. 23869]). •Valid as Orthrias frenatus (Heckel 1843) -- (Nalbant & Bianco 1998:112 [ref. 23810], Fricke et al. 2007:52 [ref. 29533]). •Valid as Barbatula frenata (Heckel 1843) -- (Coad 2010:207 [ref. 36620]). •Valid as Oxynoemacheilus frenatus (Heckel 1843) -- (Prokofiev 2009:880 [ref. 30604], Esmaeili et al. 2010:372 [ref. 33391], Golzarianpour et al. 2011:205 [ref. 31693], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:96 [ref. 32367], Kamangar et al. 2014:45 [ref. 33140], Jouladeh-Roudbar et al. 2015:883 [ref. 34062], Çiçek et al. 2015:148 [ref. 35151], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:94 [ref. 34815], Freyhof & Abdullah 2017:86 [ref. 35169], Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:65 [ref. 35254], Freyhof et al. 2017:559 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Esmaeili et al. 2018:44 [ref. 36089], Çiçek et al. 2020:245 [ref. 37644], Sayyadzadeh & Esmaeili 2020:201 [ref. 37967]). Current status: Valid as Oxynoemacheilus frenatus (Heckel 1843). Nemacheilidae. Distribution: Asia Minor and Middle East: Tigris River basin and Lesser Zab drainage (Turkey, Iraq, Iran). Habitat: freshwater.

freyhofi, Oxynoemacheilus Jouladeh-Roudbar [A.], Eagderi [S.] & Hosseinpour [T.] 2016:95, Figs. 1-7 [FishTaxa v. 1 (no. 2); ref. 34789] Roudbar River, tributary of Bakhtiyari River, near Kazem Abad village, Lorstan province, Iran, 33°08'17.8"N, 49°40'43.9"E. Holotype: IMNRF-UT-1036-1. Paratypes: IMNRF-UT. •Valid as Oxynoemacheilus freyhofi Jouladeh-Roudbar, Eagderi & Hosseinpour 2016 -- (Esmaeili et al. 2017:65 [ref. 35254], Esmaeili et al. 2018:45 [ref. 36089] with question, Jouladeh-Roudbar et al. 2020:166 [ref. 37983]). •Synonym of Oxynoemacheilus euphraticus (Bănărescu & Nalbant 1964) -- (Freyhof 2016:99 [ref. 34815], Freyhof & Özuluğ 2017:385 [ref. 35216]). Current status: Synonym of Oxynoemacheilus euphraticus (Bănărescu & Nalbant 1964). Nemacheilidae. Habitat: freshwater.

germencica, Barbatula Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:70, Fig. 2 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Aydin, Germencik, 15th kilometer, 37°38'N, 27°18'E [not correct, probably north of Germencik], Turkey. Holotype: HUIC BM-1. Paratypes: HUIC BM-1 (7). •Valid as Oxynoemacheilus germencicus (Erk'akan, Nalbant & Özeren 2007) -- (Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:96 [ref. 32367], Çiçek et al. 2015:148 [ref. 35151], Çiçek et al. 2018:19 [ref. 36267], Çiçek et al. 2020:245 [ref. 37644], Yoğurtçuoğlu et al. 2022:463 [ref. 39236]). Current status: Valid as Oxynoemacheilus germencicus (Erk'akan, Nalbant & Özeren 2007). Nemacheilidae. Distribution: Asia Minor: Büyük Menderes River basin (Aegean Sea tributary), Aydιn Province (Turkey). Habitat: freshwater.

gibbusnazus, Nemacheilus brandtii Elanidze [R. F.] 1983 [Ichthyofauna of Georgia [Eurasia]; ref. 31784] Upper reaches of Alazani River at Alvani Village, Left bank tributary of Kura River basin, Georgia [Eurasia]. Without author; authorship attributed to the book author. Original description not seen. Possibly described in an earlier paper -- Coad, 2012, Freshwater Fishes of Iran, online database. •Synonym of Oxynoemacheilus brandtii (Kessler 1877) -- (Freyhof et al. 2012:308 [ref. 31752], Kottelat 2012:95 [ref. 32367], Çiçek et al. 2018:233 [ref. 36249], Jouladeh-Roudbar et al. 2020:163 [ref. 37983], Freyhof et al. 2021:142 [ref. 38237]). Current status: Synonym of Oxynoemacheilus brandtii (Kessler 1877). Nemacheilidae. Habitat: freshwater.

gyndes, Oxynoemacheilus Freyhof [J.] & Abdullah [Y. S.] 2017:74, Figs. 1-5 [Zootaxa 4238 (no. 1); ref. 35169] Stream Zalm south of Taparezina, Iraq, 35°18'23"N, 45°58'14"E. Holotype: ZFMK 103019. Paratypes: FSJF. •Valid as Oxynoemacheilus gyndes Freyhof & Abdullah 2017 -- (Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:63 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Jouladeh-Roudbar et al. 2020:168 [ref. 37983]). Current status: Valid as Oxynoemacheilus gyndes Freyhof & Abdullah 2017. Nemacheilidae. Distribution: Middle East: upper Sirwan drainage, Tigris basin, Kurdistan (Iraq and Iran). Habitat: freshwater.

hamwii, Nemacheilus Krupp [F.] & Schneider [W.] 1991:24, Figs. 1-5 [Senckenbergiana Biologica v. 71 (nos 1/3); ref. 18977] Nahr Afrin in Afrin, Syria, 36°31'N, 36°52'E. Holotype: SMF 17398. Paratypes: SMF 23463-65 (22, 2, 5), 23468 (11), plus non-type material. •Valid as Oxynoemacheilus hamwii (Krupp & Schneider 1991) -- (Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:97 [ref. 32367], Kamangar et al. 2014:43, 47 [ref. 33140], Çiçek et al. 2016:116 [ref. 35152], Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:63 [ref. 35254], Bayçelebi 2020:758 [ref. 37964], Çiçek et al. 2020:245 [ref. 37644], Kaya et al. 2020:289 [ref. 37622], Freyhof & Geiger 2021:580 [ref. 38072]). Current status: Valid as Oxynoemacheilus hamwii (Krupp & Schneider 1991). Nemacheilidae. Distribution: Asia Minor and Middle East: Orontes River basin (Syria and Turkey). Habitat: freshwater.

hanae, Oxynoemacheilus Freyhof [J.] & Abdullah [Y. S.] 2017:79, Figs. 6-10 [Zootaxa 4238 (no. 1); ref. 35169] Stream Zalm south of Taparezina, Iraq, 35°18'23"N, 45°58'14"E. Holotype: ZFMK 103020. Paratypes: FSJF. •Valid as Oxynoemacheilus hanae Freyhof & Abdullah 2017 -- (Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:63 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Jouladeh-Roudbar et al. 2020:169 [ref. 37983]). Current status: Valid as Oxynoemacheilus hanae Freyhof & Abdullah 2017. Nemacheilidae. Distribution: Middle East: upper Sirwan drainage, Tigris basin, Kurdistan (Iraq and Iran). Habitat: freshwater.

hazarensis, Oxynoemacheilus Freyhof [J.] & Özuluğ [M.] 2017:379, Figs. 1-6 [Zootaxa 4247 (no. 4); ref. 35216] North-eastern shore of Lake Hazar, Elazığ province, Turkey, 38°28.398'N, 39°18.093'E. Holotype: IUSHM 2017-1171. Paratypes: FSJF, IUSHM. •Valid as Oxynoemacheilus hazarensis Freyhof & Özuluğ 2017 -- (Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Çiçek et al. 2018:19 [ref. 36267], Freyhof et al. 2019:37 [ref. 36739], Çiçek et al. 2020:245 [ref. 37644]). Current status: Valid as Oxynoemacheilus hazarensis Freyhof & Özuluğ 2017. Nemacheilidae. Distribution: Asia Minor: Lake Hazar basin endemic, upper Tigris basin (Turkey). Habitat: freshwater.

insignis, Cobitis Heckel [J. J.] 1843:1087 [97] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Damascus, Syria. Syntypes: NMW (4, not found), SMF 166 [ex NMW] (2, poor condition). Illustrated by Heckel 1843, Pl. 12 (fig. 3) [ref. 2067]. On p. 97 of separate [ref. 2067]. •Valid as Nemacheilus insignis (Heckel 1843) -- (Krupp & Schneider 1989:380 [ref. 13651], Coad 1991:24 [ref. 15702], Hamidan 2004:389 [ref. 37679], Fricke et al. 2007:51 [ref. 29533]). •Valid as Oxynoemacheilus insignis (Heckel 1843) -- (Prokofiev 2009:880 [ref. 30604] with subspecies, Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307, 309 [ref. 31752], Kottelat 2012:97 [ref. 32367], Kamangar et al. 2014:37 [ref. 33140], Çiçek et al. 2015:149 [ref. 35151], Çiçek et al. 2020:245 [ref. 37644]). Current status: Valid as Oxynoemacheilus insignis (Heckel 1843). Nemacheilidae. Distribution: Asia Minor and Middle East: Turkey, Syria, Israel and Jordan. Habitat: freshwater.

isauricus, Oxynoemacheilus Yoğurtçuoğlu [B.], Kaya [C.], Özuluğ [M.] & Freyhof [J.] 2021:372, Figs. 2-4, 6 [Zootaxa 4975 (no. 2); ref. 38333] Stream Çeltek at Çeltek, south of Şarkikaraağaç, Isparat Province, Turkey, 38.0124N, 31.3152E. Holotype: IUSHM 2021-1425. Paratypes: IUSHM 2021-1463 (3). Plus additional non-type material. •Valid as Oxynoemacheilus isauricus Yoğurtçuoğlu, Kaya, Özuluğ & Freyhof 2021 -- (Yoğurtçuoğlu et al. 2022:467 [ref. 39236]). Current status: Valid as Oxynoemacheilus isauricus Yoğurtçuoğlu, Kaya, Özuluğ & Freyhof 2021. Nemacheilidae. Distribution: Asia Minor: Lake Beyşehir and Suğla basins, Central Anatolia, Turkey. Habitat: freshwater.

israeliticus, Orthrias Goren [M.] & Nalbant [T. T.] in Bănărescu, Nalbant & Goren 1982:19, Figs. 19-23 [Israel Journal of Zoology v. 31 (nos 1-2); ref. 174] Ein Rewaya, Bet-She'an Valley, Israel, 32°28'N, 35°28'E, Jordan River system. Holotype: TAU 3073 [= SMNHTAU]. Paratypes: ISBB 3670 (1); TAU [= SMNHTAU] 8293-94 (1, 1). •Synonym of Nemacheilus insignis (Heckel 1843) -- (Krupp & Schneider 1989:380 [ref. 13651]). •Valid as Oxynoemacheilus israeliticus (Goren & Nalbant 1982) -- (Prokofiev 2009:880 [ref. 30604]), •Synonym of Oxynoemacheilus insignis (Heckel 1843) -- (Freyhof et al. 2012:309 [ref. 31752], Kottelat 2012:97 [ref. 32367]). Current status: Synonym of Oxynoemacheilus insignis (Heckel 1843). Nemacheilidae. Habitat: freshwater.

jordanicus, Noemacheilus angorae Bănărescu [P. M.] & Nalbant [T. T.] 1966:329, Fig. 1; Pl. 5 (fig. 1) [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 63; ref. 8005] Wadi Kufrinja, east shore of Jordan, about 1.5 kilometers before river mouth, 32°16'N, 35°34'E. Holotype: ZMH H2738. Paratypes: ZMH H2739 (orig. 16, now 14), 2740-41 (9, 5). Genus is correctly Nemacheilus. •Synonym of Nemacheilus angorae Steindachner 1897, but a valid subspecies as described -- (Goren 1974:94 [ref. 7502] as Noemacheilus). •Valid as Orthrias jordanicus (Bănărescu & Nalbant 1966) -- (Bănărescu et al. 1982:4 [ref. 174]). •Synonym of Nemacheilus insignis (Heckel 1843) -- (Krupp & Schneider 1989:380 [ref. 13651]). •Valid as Oxynoemacheilus jordanicus (Bănărescu & Nalbant 1966) -- (Prokofiev 2009:880 [ref. 30604]). •Synonym of Oxynemacheilus insignis (Heckel 1843) -- (Freyhof et al. 2012:309 [ref. 31752], Kottelat 2012:97 [ref. 32367]). Current status: Synonym of Oxynoemacheilus insignis (Heckel 1843). Nemacheilidae. Habitat: freshwater.

karunensis, Oxynoemacheilus Freyhof [J.] 2016:94, Figs. 1-5 [Zootaxa 4175 (no. 1); ref. 34815] Gamasiab River at Do Ab, Hamadan province, Iran, 34°22'20.76"N, 47°55'00.1"E. Holotype: ZFMK-ICH 102205. Paratypes: FSJF, SMF. •Valid as Oxynoemacheilus karunensis Freyhof 2016 -- (Freyhof & Abdullah 2017:86 [ref. 35169], Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:65 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Esmaeili et al. 2018:45 [ref. 36089], Sayyadzadeh & Esmaeili 2020:201 [ref. 37967], Jouladeh-Roudbar et al. 2020:170 [ref. 37983]). Current status: Valid as Oxynoemacheilus karunensis Freyhof 2016. Nemacheilidae. Distribution: Middle East: Jarrahi and Karun river drainages, Tigris River basin (Iran). Habitat: freshwater.

kaynaki, Oxynemacheilus Erk'akan [F.], Özeren [S. C.] & Nalbant [T. T.] 2008:115, Figs. 1-3 [Journal of Fisheries International [electronic journal] v. 3 (no. 4); ref. 29859] Goksu River, Nurhak, Elbistan, Firat basin, 37°53'22.82''N, 37°22'19.99''E, Turkey. Holotype: HUIC-F-20. Paratypes: HUCI-F-20 (14). Original genus should have been Oxynoemacheilus. •Valid as Oxynoemacheilus kaynaki Erk'akan, Özeren & Nalbant 2008 -- (Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:97 [ref. 32367], Kamangar et al. 2014:43, 48 [ref. 33140], Çiçek et al. 2015:149 [ref. 35151], Freyhof & Özuluğ 2017:389 [ref. 35216], Freyhof & Geiger 2017:259 [ref. 35343], Çiçek et al. 2018:19 [ref. 36267], Turan et al. 2019:[9] [ref. 36540], Freyhof et al. 2019:39 [ref. 36739], Çiçek et al. 2020:245 [ref. 37644], Rakici et al. 2020:410 [ref. 37960]). Current status: Valid as Oxynoemacheilus kaynaki Erk'akan, Özeren & Nalbant 2008. Nemacheilidae. Distribution: Asia Minor: upper Göksu River drainage, upper Euphrates River [Firat Nehri] basin, Kahramanmaraş Province (Turkey). Habitat: freshwater.

kentritensis, Oxynoemacheilus Freyhof [J.], Kaya [C.] & Turan [D.] 2017:552, Figs. 1-4 [Zootaxa 4258 (no. 6); ref. 35290] Stream Kesan about 1 km south of Güntepe, Bitlis province, Turkey, 38°21'24''N, 42°37'39''E. Holotype: FFR 01566. Paratypes: FFR, FSJF. Plus additional non-type material. Type catalog: Kaya et al. 2021:425 [ref. 38688]. •Valid as Oxynoemacheilus kentritensis Freyhof, Kaya & Turan 2017 -- (Freyhof & Geiger 2017:259 [ref. 35343] as kentritesensis, Çiçek et al. 2020:245 [ref. 37644]). Current status: Valid as Oxynoemacheilus kentritensis Freyhof, Kaya & Turan 2017. Nemacheilidae. Distribution: Asia Minor and Middle East: upper Tigris River basin (Turkey, Iraq and Syria). Habitat: freshwater.

kiabii, Oxynoemacheilus Golzarianpour [K.], Abdoli [A.] & Freyhof [J.] 2011:202, Figs. 1-3 [Ichthyological Exploration of Freshwaters v. 22 (no. 3) [for Sept.]; ref. 31693] Stream Dehnoo, 3 km west of Nahavand on road from Nahavand to Sarab-e-Gamasiab, 34°10'N, 48°24'E, Hamadan Province, Iran. Holotype: ZFMK 41847. Paratypes: FCKG 8658 (8), 8659 (7); FSJF 3003 (7), 3004 (2). Plus additional not-type material. •Valid as Oxynoemacheilus kiabii Golzarianpour, Abdoli & Freyhof 2011 -- (Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:97 [ref. 32367], Kamangar et al. 2014:45 [ref. 33140], Jouladeh-Roudbar et al. 2015:884 [ref. 34062], Teimori et al. 2015:90 [ref. 34095], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:94 [ref. 34815], Sayyadzadeh et al. 2016:248 [ref. 34976], Freyhof & Abdullah 2017:86 [ref. 35169], Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:65 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Esmaeili et al. 2018:45 [ref. 36089], Jouladeh-Roudbar et al. 2020:170 [ref. 37983]). Current status: Valid as Oxynoemacheilus kiabii Golzarianpour, Abdoli & Freyhof 2011. Nemacheilidae. Distribution: Middle East: Karkheh River drainage, Tigris River basin (Iran). Habitat: freshwater.

kosswigi, Orthrias angorae Erk'akan [F.] & Kuru [M.] 1986:160, Fig. 1a [Doga Turk Biyoloji Dergisi. v. 10 (no. 2); ref. 21167] Yildiz çayi, Yildizeli, Sivas, Turkey. Holotype: HUIC. Paratypes: HUIC (9). •Apparently valid in Barbatula -- (Stoumboudi et al. 2006:143 [ref. 28767]). •Synonym of Oxynoemacheilus angorae (Steindachner 1897), but a valid subspecies -- (Prokofiev 2009:880 [ref. 30604]). •Valid as Oxynoemacheilus kosswigi (Erk'akan & Kuru 1986) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:97 [ref. 32367], Çiçek et al. 2015:149 [ref. 35151], Sungur et al. 2017:379 [ref. 35750], Çiçek et al. 2018:19 [ref. 36267], Turan et al. 2019:[9] [ref. 36540]). •Synonym of Oxynoemacheilus seyhanensis (Bănărescu 1968) -- (Seçer et al. 2020:261 [ref. 37933], Sungur 2020:[1] [ref. 37465]). Current status: Synonym of Oxynoemacheilus seyhanensis (Bănărescu 1968). Nemacheilidae. Distribution: Asia Minor: Kızılırmak River basin (Black Sea tributary), Sivas Province (Turkey) [if valid]. Habitat: freshwater.

kurdistanicus, Oxynoemacheilus Kamangar [B. B.], Prokofiev [A. M.], Ghaderi [E.] & Nalbant [T. T.] 2014:38, Fig. 3 [Zootaxa 3755 (no. 1); ref. 33140] Choman River (Tajaban, sta. 6) Baneh, Kurdistan, Iran, 35°56'53"N, 45°41'40"E. Holotype: FCFUK 146. Paratypes: FCFUK 146-170 (35), ZMUU 23084 (4). Plus non-type specimens. •Possibly a synonym of Oxynoemacheilus euphraticus (Bănărescu & Nalbant 1964) -- (Freyhof & Özuluğ 2017:388 [ref. 35216]). •Valid as Oxynoemacheilus kurdistanicus Kamangar, Prokofiev, Ghaderi & Nalbant 2014 -- (Jouladeh-Roudbar et al. 2015:884 [ref. 34062], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:100 [ref. 34815], Freyhof & Abdullah 2017:86 [ref. 35169], Esmaeili et al. 2017:65 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Esmaeili et al. 2018:45 [ref. 36089], Çiçek et al. 2020:245 [ref. 37644], Jouladeh-Roudbar et al. 2020:172 [ref. 37983]). Current status: Valid as Oxynoemacheilus kurdistanicus Kamangar, Prokofiev, Ghaderi & Nalbant 2014. Nemacheilidae. Distribution: Asia Minor and Middle East: Tigris River drainage (Turkey and Iran). Habitat: freshwater.

lenkoranensis, Nemacheilus angorae Abdurakhmanov [Y. A.] 1962:285, Fig. 50 [Freshwater fishes off Azerbaijan; ref. 21165] Lenkoran, Azerbaijan. No types known. •Synonym of Nemacheilus angorae Steindachner 1897, but a valid subspecies as described -- (Coad 1995:22 [ref. 23608]). •Synonym of Oxynoemacheilus angorae (Steindachner 1897), but a valid subspecies -- (Prokofiev 2009:880 [ref. 30604]). •Valid as Oxynoemacheilus lenkoranensis (Abdurakhmanov 1962) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:98 [ref. 32367], Esmaeili et al. 2017:62 [ref. 35254], Esmaeili et al. 2017:66 [ref. 35254], Esmaeili et al. 2018:46 [ref. 36089], Jouladeh-Roudbar et al. 2020:173 [ref. 37983], Kuljanishvili et al. 2020:[6] [ref. 37535]). •Synonym of Oxynoemacheilus bergi (Gratzianov 1907) -- (Çiçek et al. 2018:233 [ref. 36249], Freyhof et al. 2022:168 [ref. 39240]). Current status: Synonym of Oxynoemacheilus bergi (Gratzianov 1907). Nemacheilidae. Distribution: Eurasia: Azerbaijan (if valid). Habitat: freshwater.

leontinae, Nemachilus Lortet [L.] 1883:171, Pl. 18 (fig. 1) [Archives du Muséum d'Histoire Naturelle de Lyon v. 3; ref. 14341] Lake Tiberias [Galilée], Israel. Lectotype: MGHN 3665 (64 mm SL, poor condition). Paralectotypes: MGHN 3655 (1). Original genus is correctly Nemacheilus. Lectotype selected by Krupp & Schneider 1989:383 [ref. 13651]. •Synonym of Orthrias panthera (Heckel 1843) -- (Bănărescu et al. 1982:14 [ref. 174]). •Valid as Nemacheilus leontinae Lortet 1883 -- (Krupp & Schneider 1989:383 [ref. 13651]). •Synonym of Oxynoemacheilus panthera (Heckel 1843) -- (Prokofiev 2009:880 [ref. 30604]). •Valid as Oxynoemacheilus leontinae (Lortet 1883) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:98 [ref. 32367], Kamangar et al. 2014:45 [ref. 33140]). Current status: Valid as Oxynoemacheilus leontinae (Lortet 1883). Nemacheilidae. Distribution: Israel. Habitat: freshwater.

leopardus, Cobitis Heckel [J. J.] 1843:1089 [99] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Barada River, Damascus, Syria. Holotype (unique): NMW 48559. On p. 99 of separate [ref. 2067]. First reviser Heckel 1848:241 gives precedence to panthera over leopardus. •Synonym of Orthrias panthera (Heckel 1843) -- (Bănărescu et al. 1982:14 [ref. 174]). •In the synonymy of Barbatula panthera (Heckel 1843) -- (based on status of Barbatula). •Synonym of Oxynoemacheilus panthera (Heckel 1843) -- (Prokofiev 2009:880 [ref. 30604], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:98 [ref. 32367], Kamangar et al. 2014:45 [ref. 33140]). Current status: Synonym of Oxynoemacheilus panthera (Heckel 1843). Nemacheilidae. Habitat: freshwater.

longipinnis, Ilamnemacheilus Coad [B. W.] & Nalbant [T. T.] 2005:304, Figs. 1, 3 [Travaux du Muséum d'Histoire Naturelle "Grigore Antipa" v. 48; ref. 28672] Meymeh River (formerly tributary of Tigris River), Tigris-Euphrates drainage, 17 kilometers west of Dehloran City, about 21 kilometers east of Iraqi border, 32°45'30"N, 47°05'30"E, Iran. Holotype (unique): CMNFI 1979-0366 (ex NMC 79-966). •Valid as Ilamnemacheilus longipinnis Coad & Nalbant 2005 -- (Golzarianpour et al. 2011:201 [ref. 31693], Esmaeili et al. 2010:372 [ref. 33391], Kottelat 2012:85 [ref. 32367], Jouladeh-Roudbar et al. 2015:882 [ref. 34062], Teimori et al. 2015:90 [ref. 34095]). •Synonym of Oxynoemacheilus bergianus (Derjavin 1934) -- (Freyhof et al. 2022:168 [ref. 39240]). •Valid as Oxynoemacheilus longipinnis (Coad & Nalbant 2005) -- (Freyhof et al. 2016:76 [ref. 34414], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:100 [ref. 34815], Sayyadzadeh et al. 2016:248 [ref. 34976], Freyhof & Abdullah 2017:87 [ref. 35169], Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:66 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Sayyadzadeh et al. 2017:[3] 230 [ref. 36413], Esmaeili et al. 2018:46 [ref. 36089], Jouladeh-Roudbar et al. 2020:173 [ref. 37983]). Current status: Valid as Oxynoemacheilus longipinnis (Coad & Nalbant 2005). Nemacheilidae. Distribution: Middle East: Meymeh River drainage, Tigris River basin (Ilam and Khuzestan provinces, Iran). Habitat: freshwater.

macedonicus, Orthrias brandti Sorić [V. M.] 1999:83, Fig. 1 [Ichthyologia v. 31 (no. 1); ref. 28225] Doŝnica Brook, near Demir Kapija, Vardar River System, Republic of Macedonia. Holotype (unique): BIK uncat. •Synonym of Barbatula bureschi (Drensky 1928) -- (Economidis 2005:67 [ref. 28213]). •Synonym of Oxynoemacheilus bureschi (Drensky 1928) -- (Freyhof et al. 2012:306 [ref. 31752] as bureschii, Kottelat 2012:95 [ref. 32367]). Current status: Synonym of Oxynoemacheilus bureschi (Drensky 1928). Nemacheilidae. Habitat: freshwater.

marunensis, Oxynoemacheilus Sayyadzadeh [G.] & Esmaeili [H. R.] 2020:195, Figs. 2-7 [Zootaxa 4885 (no. 2); ref. 37967] Marun/Maroon River at Kharestan, Khuzestan province, Iran, 30°39'22"N, 50°12'31"E. Holotype: ZM-CBSU H2509. Paratypes: ZM-CBSU H2510 (36), H1998 (9). Plus additional non-type material. •Valid as Oxynoemacheilus marunensis Sayyadzadeh & Esmaeili 2020. Current status: Valid as Oxynoemacheilus marunensis Sayyadzadeh & Esmaeili 2020. Nemacheilidae. Distribution: Marun River drainage, Jarrahi River basin, flowing to the Persian Gulf, Khuzestan, Iran. Habitat: freshwater.

mediterraneus, Barbatula Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:74, Fig. 5 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Eğirdir, Çandir, Aksu Stream, 37°38'N, 30°31'E, Turkey. Holotype: HUIC AAKD-2a. HUIC AKD-2a (7), AKD-3 (27). •Valid as Oxynoemacheilus mediterraneus (Erk'akan, Nalbant & Özeren 2007) -- (Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:98 [ref. 32367], Çiçek et al. 2015:149 [ref. 35151], Çiçek et al. 2018:19 [ref. 36267], Çiçek et al. 2020:245 [ref. 37644], Yogurtçuoglu et al. 2021:135 [ref. 38325], Yoğurtçuoğlu et al. 2022:468 [ref. 39236]). Current status: Valid as Oxynoemacheilus mediterraneus (Erk'akan, Nalbant & Özeren 2007). Nemacheilidae. Distribution: Asia Minor: Aksu and Köprüçay drainages, Gulf of Antalya, Burdur Province (Turkey). Habitat: freshwater.

merga, Cobitis Krynicki [J.] in Nordmann 1840:470 [Voyage dans la Russie méridionale et la Crimée; ref. 3209] Rivière Podcoumok, près de Paitigorsk [Podkumok ve Pyatigorska, north Caucases]. nurga is a misspelling in Cuvier & Valenciennes 1846:24 [ref. 1011]. •Valid as Nemacheilus merga Krynicki 1840 -- (Berg 1949:873 [ref. 20661] as Nemachilus). •Valid as Orthrias merga (Krynicki 1840) -- (Bănărescu & Nalbant 1995:449 [ref. 23187]). •Valid as Barbatula merga (Krynicki 1840) -- (Reshetnikov et al. 1997:743 [ref. 24702], Vasil'eva 1998:95 [ref. 23591], Bogutskaya et al. 2001:45 [ref. 26178], Bogutskaya & Naseka 2004:114 [ref. 28183]). •Valid as Oxynoemacheilus merga (Krynicki 1840) -- (Kottelat & Freyhof 2007:333 [ref. 29996], Prokofiev 2009:881 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:98 [ref. 32367], Kuljanishvili et al. 2020:[6] [ref. 37535]). Current status: Valid as Oxynoemacheilus merga (Krynicki 1840). Nemacheilidae. Distribution: Western Caspian Sea basin. Habitat: freshwater.

mesudae, Oxynoemacheilus Erk'akan [F.] 2012:98, Fig. 1-2 [Research Journal of Biological Sciences v. 7 (no. 2); ref. 32281] Büyük Menderes river, Dinar, Civril, Denizli Province, 38°07.366'N, 30°05.723'E, western Turkey. Holotype: HUIC. Paratypes: HUIC (15). •Valid as Oxynoemacheilus mesudae Erk'akan 2012 -- (Çiçek et al. 2015:149 [ref. 35151], Çiçek et al. 2018:19 [ref. 36267], Çiçek et al. 2020:245 [ref. 37644]). •Synonym of Oxynoemacheilus germencicus (Erk'akan, Nalbant & Özeren 2007) -- (Yoğurtçuoğlu et al. 2022:463 [ref. 39236]). Current status: Synonym of Oxynoemacheilus germencicus (Erk'akan, Nalbant & Özeren 2007). Nemacheilidae. Habitat: freshwater.

muefiti, Oxynoemacheilus Freyhof [J.], Kaya [C.], Turan [D.] & Geiger [M. F.] 2019:48, Figs. 15-17 [Zootaxa 4612 (no. 1); ref. 36739] Murat River at Ballıbostan, Ağrı Province, Turkey. Holotype: FFR 15532. Paratypes: FFR 15507 (2). Plus additional non-type material. Type catalog: Kaya et al. 2021:425 [ref. 38688]. •Valid as Oxynoemacheilus muefiti Freyhof, Kaya, Turan & Geiger 2019 -- (Çiçek et al. 2020:245 [ref. 37644]). Current status: Valid as Oxynoemacheilus muefiti Freyhof, Kaya, Turan & Geiger 2019. Nemacheilidae. Distribution: Asia Minor: upper Murat River drainage and the Eğri, a tributary to Atatürk reservoir (Turkey). Habitat: freshwater.

namiri, Nemacheilus Krupp [F.] & Schneider [W.] 1991:28, Figs. 7-13 [Senckenbergiana Biologica v. 71 (nos 1/3); ref. 18977] Orontes at Jisr ash-Shughur, Syria, 35°48'N, 36°19'E. Holotype: SMF 17387. Paratypes: SMF 17388 (15), 23448-57 (6, 7, 14, 14, 50, 11, 4, 2, 6, 22), 23458-60 (4, 14, 8), 23461-62 (13, 11); NMC 90-0147 (47); plus non-type material. •Valid as Schistura namiri (Krupp & Schneider 1991) -- (Nalbant 1998:371 [ref. 26011], Erk'akan et al. 2007:80 [ref. 29026]). •Questionably a synonym of or subspecies of Paracobitis tigris (Heckle 1843) -- (Prokofiev 2009:881, 888 [ref. 30604]). •Valid as Oxynoemacheilus namiri (Krupp & Schneider 1991) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kamangar et al. 2014:45 [ref. 33140], Çiçek et al. 2015:149 [ref. 35151], Bayçelebi 2020:758 [ref. 37964], Çiçek et al. 2020:246 [ref. 37644], Kaya et al. 2020:289 [ref. 37622], Freyhof & Geiger 2021:580 [ref. 38072], Kaya et al. 2021:569 [ref. 38174]). Current status: Valid as Oxynoemacheilus namiri (Krupp & Schneider 1991). Nemacheilidae. Distribution: Asia Minor and Middle East: Orontes River basin (Syria and Turkey). Habitat: freshwater.

nasreddini, Oxynoemacheilus Yoğurtçuoğlu [B.], Kaya [C.] & Freyhof [J.] 2021:138, Figs. 2-5 [Zootaxa 4974 (no. 1); ref. 38325] Stream Aksu at Ayvali, 6 km north of Sincanli, Afyon Province, Turkey, 38.8101, 30.2560. Holotype: FFR 15588. Paratypes: FFR 15589 (8), FSJF 3205 (5). Plus additional non-type material. Type catalog: Kaya et al. 2021:426 [ref. 38688]. •Valid as Oxynoemacheilus nasreddini Yoğurtçuoğlu, Kaya & Freyhof 2021 -- (Yoğurtçuoğlu et al. 2022:470 [ref. 39236]). Current status: Valid as Oxynoemacheilus nasreddini Yoğurtçuoğlu, Kaya & Freyhof 2021. Nemacheilidae. Distribution: Asia Minor: Lakes Akşehir, Eber and Eğirdir, and Ilgın basins, Central Anatolia, Turkey. Habitat: freshwater.

oxiana, Nemacheilus brandtii var. Kessler [K. F.] 1877:177 [The Aralo-Caspian Expedition; ref. 2597] Amu Darya River, Uzbekistan. Holotype (unique): ZIN (not found in 1996). •Valid as Nemacheilus oxianus Kessler 1877 -- (Berg 1949:879 [ref. 20661] as Nemachilus, Coad 1981:15 [ref. 22348] as Noemacheilus). •Valid as 'Oxynoemacheilus' oxiana (Kessler 1877) -- (Kottelat 2012:98 [ref. 32367]). Current status: Valid as Oxynoemacheilus oxianus (Kessler 1877). Nemacheilidae. Distribution: Uzbekistan. Habitat: freshwater.

panthera, Cobitis Heckel [J. J.] 1843:1087 [97] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Damascus, Syria. Syntypes: NMW 48565 (3), NRM 15478 [ex NMW] (1). Type catalog: Nijssen et al. 1982:31-32 [ref. 19236], Menon 1987:46 [ref. 14149]). Illustrated by Heckel 1843, Pl. 12 (fig. 2) [ref. 2067]. On p. 97 of separate [ref. 2067]. •Valid as Nemacheilus panthera (Heckel 1843) -- (Goren 1974:94 [ref. 7502] as Noemacheilus, Coad 1991:24 [ref. 15702]). •Valid as Orthrias panthera (Heckel 1843) -- (Bănărescu et al. 1978:264 [ref. 9016], Bănărescu et al. 1982:14 [ref. 174]). •Valid as Barbatula panthera (Heckel 1843) -- (based on status of Barbatula). •Valid as Oxynoemacheilus panthera (Heckel 1843) -- (Prokofiev 2009:880 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:98 [ref. 32367], Khalaf 2013:4 [ref. 37897], Çiçek et al. 2015:149 [ref. 35151]). Current status: Valid as Oxynoemacheilus panthera (Heckel 1843). Nemacheilidae. Distribution: Asia Minor and Middle East: Turkey, Syria, Israel. Habitat: freshwater.

pantheroides, Orthrias Goren [M.] & Nalbant [T. T.] in Bănărescu, Nalbant & Goren 1982:16, Fig. 14-18; Pl. 2 (figs. 9-10) [Israel Journal of Zoology v. 31 (nos 1-2); ref. 174] Tanuria Springs, Golan Heights, Israel, 32°58'N, 35°48'E, Jordan River system. Holotype: TAU 8292 [= SMNHTAU]. Paratypes: ISBB 3437 (1), 3438 [ex TAU [= SMNHTAU] 2916] (?1), 3455 (5); SMF 17125 [ex ISBB 3455] (3); TAU [= SMNHTAU] 2916 (?16), 3083 (1), 3250 (1), 8280 (9); USNM 227927 (1). •Synonym of Nemacheilus insignis (Heckel 1843) -- (Krupp & Schneider 1989:380 [ref. 13651]). •Valid as Oxynoemacheilus pantheroides (Goren & Nalbant 1982) -- (Prokofiev 2009:881 [ref. 30604]). •Synonym of Oxynoemacheilus insignis (Heckel 1843) -- (Freyhof et al. 2012:309 [ref. 31752], Kottelat 2012:97 [ref. 32367]). Current status: Synonym of Oxynoemacheilus insignis (Heckel 1843). Nemacheilidae. Habitat: freshwater.

parvinae, Oxynoemacheilus Sayyadzadeh [G.], Eagderi [S.] & Esmaeili [H. R.] 2016:238, Figs. 2-10 [Iranian Journal of Ichthyology v. 3 (no. 4); ref. 34976] Javanrud city, at Sharvineh village, Leilehrud (Leileh River), a tributary of Sirvan River drainage, Tigris basin, Kermanshah Province, Iran, 34°49'37.9"N, 46°21'30.0"E. Holotype: ZM-CBSU H1987. Paratypes: ZM-CBSU. Plus additional non-type material. •Synonym of Oxynoemacheilus bergianus (Derjavin 1934) -- (Freyhof et al. 2022:168 [ref. 39240]). •Valid as Oxynoemacheilus parvinae Sayyadzadeh, Eagderi & Esmaeili 2016 -- (Freyhof & Abdullah 2017:87 [ref. 35169], Freyhof & Özuluğ 2017:389 [ref. 35216], Esmaeili et al. 2017:66 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Esmaeili et al. 2018:46 [ref. 36089], Jouladeh-Roudbar et al. 2020:174 [ref. 37983]). Current status: Valid as Oxynoemacheilus parvinae Sayyadzadeh, Eagderi & Esmaeili 2016. Nemacheilidae. Distribution: Middle East: Sirvan River drainage, Tigris river basin (Iran). Habitat: freshwater.

paucilepis, Barbatula Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:79, Fig. 9 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Sivas, Mancilik Stream, Gürün, 38°39'N, 37°38'E, Turkey. Holotype: HUIC F2. Paratypes: HUIC F2 (7). •Valid as Paracobitis paucilepis (Erk'akan, Nalbant & Özeren 2007) -- (Prokofiev 2009:881 [ref. 30604], Prokofiev 2009:888 [ref. 30604]). •Valid as Oxynoemacheilus paucilepis (Erk'akan, Nalbant & Özeren 2007) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:98 [ref. 32367], Çiçek et al. 2015:149 [ref. 35151], Freyhof & Özuluğ 2017:389 [ref. 35216], Freyhof & Geiger 2017:259 [ref. 35343], Çiçek et al. 2018:20 [ref. 36267], Turan et al. 2019:[9] [ref. 36540], Çiçek et al. 2020:246 [ref. 37644], Rakici et al. 2020:410 [ref. 37960]). Current status: Valid as Oxynoemacheilus paucilepis (Erk'akan, Nalbant & Özeren 2007). Nemacheilidae. Distribution: Asia Minor: tributaries in Kangal and Gürün, Sivas Province (Turkey). Habitat: freshwater.

persa, Cobitis Heckel [J. J.] 1847:266 [Reisen in Europa, Asien und Africa v. 2 (pt 3); ref. 2068] Springs at Persepolis, Fars Province, Iran. Holotype (unique): NMW 48567. If persa is a noun, then persa is the correct spelling when in Oxynoemacheilus. •Questionably a synonym of Nemacheilus angorae Steindachner 1897 -- (Coad 1995:21, 22 [ref. 23608]). •Valid as Nemacheilus persa (Heckel 1847) -- (Coad 1996:71 [ref. 25297], Coad 1998:103 [ref. 23869]). •Valid as Orthrias persus (Heckel 1847) -- (Nalbant & Bianco 1998:110 [ref. 23810]). •Valid as Barbatula persa (Heckel 1847). •Valid as Oxynoemacheilus persa (Heckel 1947) -- (Prokofiev 2009:881 [ref. 30604] dated 1846 as persus, Esmaeili et al. 2010:372 [ref. 33391] as persus, Freyhof et al. 2012:307, 309 [ref. 31752], Kottelat 2012:98 [ref. 32367], Mousavi-Sabet & Eagderi 2015:299 [ref. 34027], Jouladeh-Roudbar et al. 2015:884 [ref. 34062], Teimori et al. 2015:90 [ref. 34095], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:100 [ref. 34815], Sayyadzadeh et al. 2016:248 [ref. 34976], Esmaeili et al. 2017:66 [ref. 35254], Esmaeili et al. 2018:46 [ref. 36089], Sayyadzadeh et al. 2018:45 [ref. 36026], Jouladeh-Roudbar et al. 2020:175 [ref. 37983]). Current status: Valid as Oxynoemacheilus persa (Heckel 1847). Nemacheilidae. Distribution: Middle East: Mond and Kor River and Lake Maharlu drainages (Iran). Habitat: freshwater.

phasicus, Oxynoemacheilus Freyhof [J.], Kaya [C.], Epitashvili [G.] & Geiger [M. F.] 2021:139, Figs. 2-5 [Zootaxa 4952 (no. 1); ref. 38237] Dzirula River at road from Tbilisi to Kutaisi 1 km upstream from Khunevi, Georgia, 42.0975 43.3900. Holotype: ZFMK ICH-124180. Paratypes: FSJF. Plus additional non-type material. •Valid as Oxynoemacheilus phasicus Freyhof, Kaya, Epitashvili & Geiger 2021. Current status: Valid as Oxynoemacheilus phasicus Freyhof, Kaya, Epitashvili & Geiger 2021. Nemacheilidae. Distribution: Rioni and Enguri River drainages, Georgia. Habitat: freshwater.

phoxinoides, Barbatula Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:75, Fig. 6 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Iznik, Bursa Province, 40°41'N, 29°30'E, Turkey. Holotype: HUC MAR-7g. Paratypes: HUIC D12 (10). •Valid as Oxynoemacheilus phoxinoides (Erk'akan, Nalbant & Özeren 2007) -- (Prokofiev 2009:881 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:98 [ref. 32367], Çiçek et al. 2015:149 [ref. 35151], Çiçek et al. 2018:20 [ref. 36267]). •Synonym of Oxynoemacheilus angorae (Steindachner 1897) -- (Çiçek et al. 2019:16 [ref. 36528], Yoğurtçuoğlu et al. 2022:458 [ref. 39236]). Current status: Synonym of Oxynoemacheilus angorae (Steindachner 1897). Nemacheilidae. Habitat: freshwater.

pindus, Barbatula Economidis [P. S.] 2005:67, Figs. 1-3 [Ichthyological Exploration of Freshwaters v. 16 (no. 1); ref. 28213] Stream Sarantaporos near village Drosopygi, Aoos drainage, Epiros, Greece. Holotype: LZUT 1977-215a. Paratypes: CMK 13003 (2), 16926 (27), 16931 (2); LZUT 1977-215 (12), 1980-13 (13), 1985-15 (2), 1985-16 (6), 1987-13 (25), 1987-14 (1), 1987-31 (28), 1987-32 (6). •Valid as Oxynoemacheilus pindus (Economidis 2005) -- (Stoumboudi et al. 2006:143 [ref. 28767], Kottelat & Freyhof 2007:334 [ref. 29996], Prokofiev 2009:881 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:99 [ref. 32367], Barbieri et al. 2015:76 [ref. 35029]). Current status: Valid as Oxynoemacheilus pindus (Economidis 2005). Nemacheilidae. Distribution: Albania and Greece. Habitat: freshwater.

pindus, Orthrias Economidis [P. S.] 1991:26, 42, (fig. 4) [Check list of freshwater fishes of Greece; ref. 23018] Not available; name only, no description. Also appeared in Economidis 1992:49 [ref. 23021]. Later made available by Economidis 2005. •Treated as valid as Barbatula sp. -- (Kottelat 1997:94 [ref. 22952]). Nomen Nudum. Current status: Synonym of Oxynoemacheilus pindus (Economidis 2005). Nemacheilidae.

pulsiz, Nemacheilus Krupp [F.] 1992:291, Fig. 1 [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 89; ref. 20207] Lake Van drainage basin, road from Erçis to Patnos, Turkey. Holotype: ZMH 7995. Paratypes: ZMH 3606 (13), 3636 (66, now 64). Type catalog: Wilkens & Dohse 1993:413 [ref. 21161]. •Valid as Barbatula pulsiz (Krupp 1992) -- (Fricke et al. 2007:51 [ref. 29533]). •Valid as Paracobitis pulsiz (Krupp 1992) -- (Prokofiev 2009:888 [ref. 30604]). •Synonym of Oxynoemacheilus ercisianus (Erk'akan & Kuru 1986) -- (Freyhof et al. 2012:308 [ref. 31752], Kottelat 2012:96 [ref. 32367], Kamangar et al. 2014:45 [ref. 33140], Freyhof et al. 2019:34 [ref. 36739]). Current status: Synonym of Oxynoemacheilus ercisianus (Erk'akan & Kuru 1986). Nemacheilidae. Habitat: freshwater, brackish.

samantica, Orthrias brandti Bănărescu [P. M.] & Nalbant [T. T.] in Bănărescu, Nalbant & Balik 1978:263, Fig. 4 [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 75; ref. 9016] Tributary to Samanti, between Pinarbsi and Sarizi, Seyhan basin, southern Turkey. Holotype: ZMH 3633. Paratypes: ISBB 2978 (1), ZMH 5638 [not 5637] (8). Type catalog: Wilkens & Dohse 1993:413 [ref. 21161]. •Valid as Schistura samantica (Bănărescu & Nalbant 1978) -- (Erk'akan et al. 2007:81 [ref. 29026]). •Synonym of Oxynoemacheilus brandtii (Kessler 1877), but a valid subspecies samanticus -- (Prokofiev 2009:880 [ref. 30604]). •Synonym of Oxynoemacheilus bergianus (Derjavin 1934) -- (Freyhof et al. 2022:168 [ref. 39240]). •Valid as Oxynoemacheilus samanticus (Bănărescu & Nalbant 1978) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:99 [ref. 32367], Çiçek et al. 2015:149 [ref. 35151], Freyhof & Geiger 2017:259 [ref. 35343], Sungur et al. 2017:379 [ref. 35750], Çiçek et al. 2018:20 [ref. 36267], Turan et al. 2019:[9] [ref. 36540], Bayçelebi 2020:758 [ref. 37964], Çiçek et al. 2020:246 [ref. 37644], Kaya et al. 2020:289 [ref. 37622], Yogurtçuoglu et al. 2020:[14] [ref. 37627], Saygun et al. 2021:47 [ref. 38525]). Current status: Valid as Oxynoemacheilus samanticus (Bănărescu & Nalbant 1978). Nemacheilidae. Distribution: Asia Minor: Seyhan River basin, Mediterranean Sea tributaries (Turkey). Habitat: freshwater.

sarus, Oxynoemacheilus Freyhof [J.], Yoğurtçuoğlu [B.] & Kaya [C.] 2021:128, Figs. 2-5 [Zootaxa 4964 (no. 1); ref. 38277] Lower stream Çakıt, south of Salbaş, Adana Province, Turkey, 37.1031N, 35.1094E. Holotype: FFR 15585. Paratypes: FFR, FSJF. Plus additional non-type material. Type catalog: Kaya et al. 2021:427 [ref. 38688]. •Valid as Oxynoemacheilus sarus Freyhof, Yoğurtçuoğlu & Kaya 2021. Current status: Valid as Oxynoemacheilus sarus Freyhof, Yoğurtçuoğlu & Kaya 2021. Nemacheilidae. Distribution: Asia Minor: lower Ceyhan and Seyhan drainages, Turkey. Habitat: freshwater.

seyhanensis, Noemacheilus (Paracobitis) tigris Bănărescu [P. M.] 1968:355, Pl. 3 (figs. 2-3) [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 65; ref. 16001] Between Viransehir and Kazancik, Turkey. Holotype: ZMH H4014. Paratypes: ZMH H4015 (3). Type catalog: Wilkens 1977:159 [ref. 22013]. Genus is correctly Nemacheilus. •Synonym of Paracobitis tigris (Heckel 1843), but a valid subspecies -- (Prokofiev 2009:888 [ref. 30604]). •Valid as Oxynoemacheilus seyhanensis (Bănărescu 1968) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:99 [ref. 32367], Kamangar et al. 2014:45 [ref. 33140], Çiçek et al. 2015:149 [ref. 35151], Çiçek et al. 2018:20 [ref. 36267], Bayçelebi 2020:759 [ref. 37964], Çiçek & Sungur 2020:117 [ref. 38064], Çiçek et al. 2020:246 [ref. 37644], Kaya et al. 2020:289 [ref. 37622], Seçer et al. 2020:261 [ref. 37933], Sungur 2020:[1] [ref. 37465], Yogurtçuoglu et al. 2020:[14] [ref. 37627], Saygun et al. 2021:47 [ref. 38525]). Current status: Valid as Oxynoemacheilus seyhanensis (Bănărescu 1968). Nemacheilidae. Distribution: Asia Minor: Seyhan River basin, Mediterranean Sea tributaries (Turkey). Habitat: freshwater.

seyhanicola, Schistura Erk'akan [F.], Nalbant [T. T.] & Özeren [S. C.] 2007:81, Fig. 11 [Journal of Fisheries International [electronic journal] v. 2 (no. 1); ref. 29026] Dam Bridge, near Adana, Seyhan River basin, 38°39'N, 36°37'E, Turkey. Holotype (unique): HUIC SEY-1. Also spelled Nanoschistura seyhanicolo in Fig. 11, an obvious error. •Valid as Schistura seyhanicola Erk'akan, Nalbant & Özeren 2007 -- (Golzarianpour et al. 2011:205 [ref. 31693]). •Valid as Oxynoemacheilus seyhanicola (Erk'akan, Nalbant & Özeren 2007) -- (Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:99 [ref. 32367], Çiçek et al. 2015:149 [ref. 35151], Çiçek et al. 2018:20 [ref. 36267], Bayçelebi 2020:759 [ref. 37964], Çiçek et al. 2020:246 [ref. 37644], Kaya et al. 2020:289 [ref. 37622], Freyhof & Geiger 2021:580 [ref. 38072] as "seyhanicola", Freyhof et al. 2021:133 [ref. 38277]). Current status: Valid as Oxynoemacheilus seyhanicola (Erk'akan, Nalbant & Özeren 2007). Nemacheilidae. Distribution: Asia Minor: Seyhan River basin (Mediterranean tributary), Adana Province (Turkey). Habitat: freshwater.

shehabi, Oxynoemacheilus Freyhof [J.] & Geiger [M. F.] 2021:573, Figs. 2-6 [Zootaxa 4908 (4); ref. 38072] Orontes at Al Qusayr, Syria, 34.5086N, 36.5389E. Holotype: ZFMK ICH-124181. Paratypes: ZFMK ICH-125126-28 (3) [ex FSJF 2739]. •Valid as Oxynoemacheilus shehabi Freyhof & Geiger 2021. Current status: Valid as Oxynoemacheilus shehabi Freyhof & Geiger 2021. Nemacheilidae. Distribution: Western Asia: upper Orontes basin (Syria). Habitat: freshwater.

simavica, Orthrias brandti Balik [S.] & Bănărescu [P. M.] in Bănărescu, Nalbant & Balik 1978:261, Fig. 3 [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 75; ref. 9016] Simav stream, Balikesir. Turkey. Holotype: ISBB 2976. Paratypes: ISBB 2977 (2). See Stoumboudi et al. 2006:143 [ref. 28767] for information on the type locality. •Valid as Barbatula simavica (Balik & Bănărescu 1978) -- (Erk'akan et al. 2007:70 [ref. 29026]). •Synonym of Oxynoemacheilus brandtii (Kessler 1877), but a valid subspecies simavicus -- (Prokofiev 2009:880 [ref. 30604]). •Valid as Oxynoemacheilus simavicus (Balik & Bănărescu 1978) -- (Stoumboudi et al. 2006:143 [ref. 28767], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:99 [ref. 32367], Çiçek et al. 2015:149 [ref. 35151], Çiçek et al. 2018:20 [ref. 36267], Turan et al. 2019:[10] [ref. 36540], Çiçek et al. 2020:246 [ref. 37644], Saygun et al. 2021:47 [ref. 38525], Freyhof et al. 2022:179 [ref. 39240]). Current status: Valid as Oxynoemacheilus simavicus (Balik & Bănărescu 1978). Nemacheilidae. Distribution: Asia Minor: Marmara Sea tributary, Balιkesir Province (Turkey). Habitat: freshwater.

terekensis, Nemacheilus Gratzianov [V. I.] 1907:163, 167 [Trudy Otdela Ichtiologii Imperatorskago Russkago Obshchestva Akklimatizacii Zivotnych' i Rastenii. v. 4; ref. 1871] Terek River at Vladikavkas, North Osetia-Alanya, southern Russia. Syntypes: ZMMU P-3318 (24). Type catalog: Vasil'eva et al. 2001:24-25 [ref. 25714]. •Synonym of Nemacheilus marga [or merga] Krynicki 1840 -- (Berg 1949:873 [ref. 20661] as Nemachilus). •Synonym of Barbatula merga (Krynicki 1840) -- (Vasil'eva 1998:95 [ref. 23591]). •Synonym of Oxynoemacheilus merga (Krynicki 1840) -- (Freyhof et al. 2012:306 [ref. 31752], Kottelat 2012:98 [ref. 32367]). Current status: Synonym of Oxynoemacheilus merga (Krynicki 1840). Nemacheilidae. Habitat: freshwater.

theophilii, Oxynoemacheilus Stoumboudi [M. Th.], Kottelat [M.] & Barbieri [R.] 2006:140, Figs. 8-9 [Ichthyological Exploration of Freshwaters v. 17 (no. 2); ref. 28767] Lesbos Island, Tsingou springs, in Evergetoulas drainage, Greece. Holotype: MHNG 2679.009. Paratypes: HCMR 1469 (2). •Valid as Oxynoemacheilus theophili [sic] or theophilii Stoumboudi, Kottelat & Barbieri 2006 -- (Erk'akan et al. 2008:115 [ref. 29859], Kottelat & Freyhof 2007:334 [ref. 29996], Prokofiev 2009:881 [ref. 30604], Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Erk'akan 2012:97 [ref. 32281], Kottelat 2012:99 [ref. 32367], Barbieri et al. 2015:76 [ref. 35029], Çiçek et al. 2015:149 [ref. 35151], Çiçek et al. 2020:246 [ref. 37644], Yoğurtçuoğlu et al. 2022:471 [ref. 39236]). Current status: Valid as Oxynoemacheilus theophilii Stoumboudi, Kottelat & Barbieri 2006. Nemacheilidae. Distribution: Eurasia: Lesbos Island, Greece and Bakir River drainage, western Anatolia, Turkey. Habitat: freshwater.

tigris, Cobitis Heckel [J. J.] 1843:1088 [98] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Kuiek River near Aleppo, Syria. Syntypes: NMW 48441 (2), 49444-46 (2, 4, 2); SMF 405 (3). Illustrated by Heckel 1843, Pl. 12 (fig. 4) [ref. 2067]. On p. 98 of separate [ref. 2067]. •Valid as Noemacheilus tigris (Heckel 1843) -- (Goren 1974:93 [ref. 7502]). •Valid as Nemacheilus tigris (Heckel 1843) -- (Berg 1949:879 [ref. 20661] as Nemachilus, Coad 1991:18 [ref. 15702], Coad 1995:23 [ref. 23608], Coad 1998:103 [ref. 23869]). •Valid as Orthrias tigris (Heckel 1843) -- (Bănărescu et al. 1982:21 [ref. 174], Bănărescu & Nalbant 1995:443 [ref. 23187], Nalbant & Bianco 1998:111 [ref. 23810], Fricke et al. 2007:52 [ref. 29533]). •Valid as Paracobitis tigris (Heckel 1843) -- (Erk'akan et al. 2007:79 [ref. 29026], Prokofiev 2009:881, 888 [ref. 30604]). •Valid as Barbatula tigris (Heckel 1843) -- (Khalaf 2013:4 [ref. 37897]). •Valid as Oxynoemacheilus tigris (Heckel 1843) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307 [ref. 31752], Kottelat 2012:99 [ref. 32367], Kamangar et al. 2014:45 [ref. 33140], Çiçek et al. 2015:149 [ref. 35151], Freyhof et al. 2019:42 [ref. 36739], Çiçek et al. 2020:246 [ref. 37644]). Current status: Valid as Oxynoemacheilus tigris (Heckel 1843). Nemacheilidae. Distribution: Asia Minor and Middle East: upper Euphrates River basin (Syria and Turkey). Habitat: freshwater.

tongiorgii, Seminemacheilus Nalbant [T. T.] & Bianco [P. G.] 1998:113, Figs. 11 A-D [Italian Journal of Zoology v. 65, Suppl.; ref. 23810] Large water spring near Darab town, Kul River basin, Iran. Holotype: IZA 801. Paratypes: ISBB uncat. (1). •Valid as Seminemacheilus tongiorgii Nalbant & Bianco 1998 -- (Esmaeili et al. 2010:373 [ref. 33391]). •Valid as Oxynoemacheilus tongiorgii (Nalbant & Bianco 1998) -- (Golzarianpour et al. 2011:205 [ref. 31693], Freyhof et al. 2012:307, 309 [ref. 31752], Kottelat 2012:99 [ref. 32367], Jouladeh-Roudbar et al. 2015:884 [ref. 34062], Teimori et al. 2015:90 [ref. 34095], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:94 [ref. 34815], Sayyadzadeh et al. 2016:248 [ref. 34976], Esmaeili et al. 2017:68 [ref. 35254], Esmaeili et al. 2018:47 [ref. 36089], Jouladeh-Roudbar et al. 2020:176 [ref. 37983]). Current status: Valid as Oxynoemacheilus tongiorgii (Nalbant & Bianco 1998). Nemacheilidae. Distribution: Middle East: Kor River basin (Iran). Habitat: freshwater.

tortonesei, Noemacheilus insignis Bănărescu [P. M.] & Nalbant [T. T.] 1966:331, Figs. 2-3; Pl. 4 (figs. 2-4) [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 63; ref. 8005] Wadi-Umm ad-Dananir Spring, tributary of Zerqua River, about 25 kilometers north of Amman, Jordan, about 32°04'N, 35°50'E. Holotype: ZMH 2742. Paratypes: ZMH 2743 (orig. 11, now 8). Type catalog: Wilkens 1977:158 [ref. 22013]. Genus is correctly Nemacheilus. •Valid as Nemacheilus insignis (Heckel 1843), but a valid subspecies as described -- (Goren 1974:93 [ref. 7502]). •Synonym of Nemacheilus insignis (Heckel 1843) -- (Krupp & Schneider 1989:380 [ref. 13651]). •Synonym of Oxynoemacheilus insignis (Heckel 1843), but a valid subspecies or a synonym of Oxynoemacheilus jordanicus -- (Prokofiev 2009:880 [ref. 30604]). •Synonym of Oxynoemacheilus insignis (Heckel 1843) -- (Freyhof et al. 2012:309 [ref. 31752], Kottelat 2012:97 [ref. 32367] with question). Current status: Synonym of Oxynoemacheilus insignis (Heckel 1843). Nemacheilidae. Habitat: freshwater.

tschaiyssuensis, Noemacheilus Bănărescu [P. M.] & Nalbant [T. T.] 1964:177, Pl. 8 (fig. 13) [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 61; ref. 217] Çayı Su [Tchaiy-su], between Keyson (37°30'N, 37°50'E) and Gaziantep (37°00'N, 37°30'E), Euphrates basin, Turkey. Holotype: BMNH 1935.9.12.57. Paratypes: BMNH 1935.9.12.58-60 (3) Tchaiy-su. Genus is correctly Nemacheilus. •Valid as Orthrias tschaiyssuensis (Bănărescu & Nalbant 1964) -- (Bănărescu et al. 1978:264 [ref. 9016], Fricke et al. 2007:52 [ref. 29533]). •Valid as Oxynoemacheilus tschaiyssuensis (Bănărescu & Nalbant 1964) -- (Prokofiev 2009:881 [ref. 30604]). •Synonym of Oxynoemacheilus argyrogramma (Heckel 1847) -- (Freyhof et al. 2012:308 [ref. 31752], Kottelat 2012:95 [ref. 32367], Freyhof 2016:99 [ref. 34815]). Current status: Synonym of Oxynoemacheilus argyrogramma (Heckel 1847). Nemacheilidae. Habitat: freshwater.

veyseli, Oxynoemacheilus Çiçek [E.], Eagderi [S.] & Sungur [S.] 2018:233, Figs. 1A, 2-6 [Iranian Journal of Ichthyology v. 5 (no. 3); ref. 36249] Bozkuş River, a tributary of the Aras River at Kars, Kars Province, Turkey, 40°37'03.7"N, 42°47'04.9"E. Holotype: NHVUIC 14005-H. Paratypes: NHVUIC. Named after father and son Veysel, so mandatorily changed to veyselorum. •Valid as Oxynoemacheilus veyselorum Çiçek, Eagderi & Sungur 2018 -- (Çiçek et al. 2018:20 [ref. 36267] as veyseli, Çiçek et al. 2020:246 [ref. 37644], Kaya et al. 2020:339 [ref. 37582], Kuljanishvili et al. 2020:[6] [ref. 37535], Jouladeh-Roudbar et al. 2020:177 [ref. 37983], Çiçek et al. 2021:173 [ref. 38560], Saygun et al. 2021:47 [ref. 38525]). Current status: Valid as Oxynoemacheilus veyselorum Çiçek, Eagderi & Sungur 2018. Nemacheilidae. Distribution: Asia Minor and Middle East: upper Aras River drainage, Kars Province (Turkey and northwest Iran). Habitat: freshwater.

zagrosensis, Oxynoemacheilus Kamangar [B. B.], Prokofiev [A. M.], Ghaderi [E.] & Nalbant [T. T.] 2014:43, Fig. 4 [Zootaxa 3755 (no. 1); ref. 33140] Shooei River (Jemli, sta. 9), Baneh, Kurdistan, Iran, 35°58'01"N, 45°42'43"E. Holotype: FCFUK 101. Paratypes: FCFUK 136-136 (31), ZMUU 23085 (4). •Valid as Oxynoemacheilus zagrosensis Kamangar, Prokofiev, Ghaderi & Nalbant 2014 -- (Jouladeh-Roudbar et al. 2015:884 [ref. 34062], Jouladeh-Roudbar et al. 2016:105 [ref. 34789], Freyhof 2016:94 [ref. 34815], Sayyadzadeh et al. 2016:248 [ref. 34976], Freyhof & Özuluğ 2017:379 [ref. 35216], Esmaeili et al. 2017:68 [ref. 35254], Freyhof et al. 2017:552 [ref. 35290], Freyhof & Geiger 2017:259 [ref. 35343], Esmaeili et al. 2018:47 [ref. 36089], Jouladeh-Roudbar et al. 2020:178 [ref. 37983]). Current status: Valid as Oxynoemacheilus zagrosensis Kamangar, Prokofiev, Ghaderi & Nalbant 2014. Nemacheilidae. Distribution: Middle East: Tigris River basin (Iran). Habitat: freshwater.

zarzianus, Oxynoemacheilus Freyhof [J.] & Geiger [M. F.] 2017:261, Figs. 3-7 [Zootaxa 4273 (no. 2); ref. 35343] Stream Kunamasi in Sevanja, Iraq, 35°47.35'N, 45°24.18'E. Holotype: ZFMK Ich-103667. Paratypes: FSJF. Plus additional non-type material. •Valid as Oxynoemacheilus zarzianus Freyhof & Geiger 2017 -- (Eagderi et al. 2022:12 [ref. 39252]). Current status: Valid as Oxynoemacheilus zarzianus Freyhof & Geiger 2017. Nemacheilidae. Distribution: Middle East: Lesser Zab drainage (Iraq), and Marivan (Iran). Habitat: freshwater.

zeravschani, Nemachilus oxianus natio Turdakov [F. A.] 1936:199 [Trudy Uzbekskogo Gosudarstvennogo Universiteta v. 7; ref. 21172] Original genus is correctly Nemacheilus. Not available, infrasubspecific; from Zeravschan River, Ak-Darya. •In the synonymy of Nemachilus oxianus Kessler 1877 -- (Berg 1949:879 [ref. 20661]). •In the synonymy of 'Oxynoemacheilus'oxianus (Kessler 1877) -- (Kottelat 2012:98 [ref. 32367]). Infrasubspecific. Current status: Synonym of Oxynoemacheilus oxianus (Kessler 1877). Nemacheilidae.


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