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Eschmeyer's Catalog of Fishes
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Species in the genus Systomus: [ 44 ] records


asoka, Puntius Kottelat [M.] & Pethiyagoda [R.] 1989:472, Figs. 1, 4 [Die Aquarien- und Terrarienzeitschrift (DATZ) v. 42 (no. 8); ref. 14110] Sitawaka River, 6°55'25"N, 80°20'02"E, Kegala District, Sabaragamuwa Province, Sri Lanka. Holotype: ZSM 27188 [now ZRC 38480]. Paratypes: ZMA 120334 (1); CMK 6402 (9), 6522 (1). Type catalog: Neumann 2011:255 [ref. 31652]. The holotype is currently at ZRC. •Valid as Puntius asoka Kottelat & Pethiyagoda 1989 -- (Pethiyagoda 1991:93 [ref. 20075], Silva et al. 2008:62 [ref. 29719]). •Valid as Systomus asoka Kottelat & Pethiyagoda 1989 -- (Pethiyagoda et al. 2012:77 [ref. 32071], Sudasinghe et al. 2020:[4] [ref. 37737]). Current status: Valid as Systomus asoka (Kottelat & Pethiyagoda 1989). Cyprinidae: Smiliogastrinae. Distribution: South Asia: southwestern Sri Lanka. Habitat: freshwater.

binduchitra, Barbus (Puntius) Hora [S. L.] 1937:327, Fig. 4 [Records of the Indian Museum (Calcutta) v. 39 (pt 4); ref. 20745] Road-side drains and small streams at Sandoway, Lower Myanmar. Holotype: ZSI F 12478/1. Paratypes: ZSI F12478/1 (9). Type catalog: Menon & Yazdani 1968:116 [ref. 20743]. See Das 2003:354 [ref. 27706]. •Synonym of Puntius sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Talwar & Jhingran 1991:282 [ref. 20764]). •Synonym of Barbodes sarana (Hamilton 1822) subspecies orphoides (Valenciennes 1842) -- (Menon 1999:70 [ref. 24904]). •Synonym of Puntius sarana (Hamilton 1822). •Synonym of Systomus sarana (Hamilton 1822) based on placement of sarana. •Valid as Systomus binduchitra (Hora 1937) -- (Kottelat 2013:165 [ref. 32989]). Current status: Valid as Systomus binduchitra (Hora 1937). Cyprinidae: Smiliogastrinae. Distribution: Myanmar. Habitat: freshwater, brackish.

caudimarginatus, Barbus Blyth [E.] 1860:157 [Journal of the Asiatic Society of Bengal v. 29 (no. 2); ref. 477] Tenasserim provinces, Myanmar. No types known. •Synonym of Puntius sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Talwar & Jhingran 1991:282 [ref. 20764], Menon 1999:69 [ref. 24904] but as genus Barbodes). •Synonym of Systomus orphoides (Valenciennes 1842) -- (Rainboth 1996:104 [ref. 22772]). •Synonym of Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus rubripinnis (Valenciennes 1842). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

chryseus, Systomus Plamoottil [M.] 2014:140, Fig. 12 [International Journal of Fauna and Biological Studies v. 1 (no. 6) Part C; ref. 33650] Keezhvaipur, Manimala River, Kerala, India. Holotype: ZSI/FF 4625. Paratypes: ZSI/WGRC/IR/V. •Valid as Systomus chryseus Plamoottil 2014 -- (Plamoottil 2016:95 [ref. 34814], Arunachalam et al. 2018:45 [ref. 36280], Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Valid as Systomus chryseus Plamoottil 2014. Cyprinidae: Smiliogastrinae. Distribution: Manimala River drainage, Kerala, India. Habitat: freshwater.

chrysopoma, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:165, Pl. 466 [Histoire naturelle des poissons v. 16; ref. 1009] Coast of Malabar, India. Syntypes: MNHN 0000-5776 (2). Type catalog: Bertin & Estève 1948:24 [ref. 19574]. •Synonym of Barbodes sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Menon 1999:72 [ref. 24904]). •Synonym of Puntius sarana (Hamilton 1822). •Valid as Systomus chrysopoma (Valenciennes 1842) -- (Plamoottil 2014:145 [ref. 33650], Bleher 2018:788 [ref. 36647]). •Synonym of Systomus subnasutus (Valenciennes 1842) -- (Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989], Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

chrysosomus, Systomus McClelland [J.] 1839:284 [Asiatic Researches v. 19 (pt 2); ref. 2923] Bengal, India. No types known. •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904] as chrysopomus). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

clavatus, Barbus McClelland [J.] 1845:280, Pl. 21 (fig. 2) [Calcutta Journal of Natural History v. 5 (no. 18); ref. 2929] Rivers at foot of Sikkim Mountains, northern frontier of Bengal. Holotype (unique): whereabouts unknown. •Valid as Poropuntius clavatus (McClelland 1845) -- (Roberts 1998:110 [ref. 23549], Menon 1999:83 [ref. 24904], Vishwanath & Kosygin 2001:35 [ref. 25682]). •Valid as Puntius clavatus (McClelland 1845) -- (Talwar & Jhingran 1991:265 [ref. 20764], Sen 1995:513 [ref. 23778], Kundu 2000:96 [ref. 25663], Karmakar 2000:28 [ref. 25662], Arunkumar & Tombe Singh 2003:481 [ref. 26926], Shrestha 2008:77 [ref. 29923]). •Valid as Systomus clavatus (McClelland 1845) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Bleher 2018:790 [ref. 36647]). Current status: Valid as Systomus clavatus (McClelland 1845). Cyprinidae: Smiliogastrinae. Distribution: Ganges and Brahmaputra River basins, Nepal, India Bangladesh and Myanmar. Habitat: freshwater.

compressiformis, Barbus Cockerell [T. D. A.] 1913:133 [Bulletin of the Bureau of Fisheries v. 32 (Doc. 779) (for 1912); ref. 870] Fort Stedman, southern Shan States, Indochina, elevation 3000 feet. Syntypes: BMNH 1893.6.30.43-52 (10). Replacement name for Barbus compressus Boulenger 1893, preoccupied by Barbus (Barboides) compressus Day 1870. •Synonym of Puntius clavatus (McClelland 1845) -- (Talwar & Jhingran 1991:265 [ref. 20764]). •Valid as Puntius compressiformis (Cockerell 1913) -- (Doi 1997:11 [ref. 22832]). •Valid as Systomus compressiformis (Cockerell 1913) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Kottelat 2013:165 [ref. 32989], Fuke et al. 2021:[3] [ref. 38221]). Replacement Name. Current status: Valid as Systomus compressiformis (Cockerell 1913). Cyprinidae: Smiliogastrinae. Distribution: South Asia: Myanmar. Habitat: freshwater.

compressus, Barbus Boulenger [G. A.] 1893:202 [Annals and Magazine of Natural History (Series 6) v. 12 (no. 69); ref. 14522] Fort Stedman, southern Shan States, Indochina, elevation 3000 feet. Syntypes: BMNH 1893.6.30.43-52 (10). Objectively invalid; preoccupied by Barbus (Barbodes) compressus Day 1870, replaced by Barbus compressiformis Cockerell 1913. Also replaced by Barbus liui Fowler 1958. •In the synonymy Puntius clavatus (McClelland 1845) -- (Talwar & Jhingran 1991:265 [ref. 20764]). •In the synonym of Systomus compresiformis (Cockerell 1913) -- (Kottelat 2013:165 [ref. 32989]). 1°Homonym. Current status: Synonym of Systomus compressiformis (Cockerell 1913). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

diliciosus, Barbus McClelland [J.] 1839:272, 342, Pl. 39 (fig. 3) [Asiatic Researches v. 19 (pt 2); ref. 2923] Upper Assam, India. No types known. Spelled diliciosus on pp. 272, 343 and plate caption (p. 272); spelled deliciosus in index. •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904] as deliciosus). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

duvaucelii, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:167 [Histoire naturelle des poissons v. 16; ref. 1009] Bengal, India. Holotype (unique): MNHN 0000-3817. Type catalog: Bertin & Estève 1948:23 [ref. 19574]. •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904] as duvancelli). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

gardonides, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:156, Pl. 465 [Histoire naturelle des poissons v. 16; ref. 1009] Calcutta and Mumbai, India; Java, Indonesia. Lectotype: MNHN 0000-3819. Syntypes: MNHN 0000-3820 to 0000-3821 (1, 2), B-1214 (1). Type catalog: Bertin & Estève 1948:24-25 [ref. 19574] with MNHN 3819 as holotype. Lectotype selected by Kottelat 2013:166 [ref. 32989]. •Java specimens are questionably a synonym of Puntius orphoides (Valenciennes 1842) -- (Roberts 1993:21 [ref. 21799]). •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904] as gardonoides). •Synonym of Puntius orphoides (Valenciennes 1842) -- (Zhang et al. 2016:97 [ref. 34477]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

gibbosus, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:155 [Histoire naturelle des poissons v. 16; ref. 1009] Aleppey, Kerala, India. Holotype (unique): MNHN 0000-3816. Type catalog: Bertin & Estève 1948:24 [ref. 19574]. •Synonym of Barbodes sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Menon 1999:72 [ref. 24904]). •Synonym of Systomus subnasutus (Valenciennes 1842) -- (Plamoottil & Maji 2020:1036 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

gracilus, Systomus Plamoottil [M.] & Maji [D.] 2020:1034, Figs. 1-2 [Journal of Experimental Zoology of India v. 23 (no. 2); ref. 37708] Ganges River near Naihati, West Bengal, India, 22.8895°N, 88.422°E. Holotype: ZSI/ANRC M-23593. Paratypes: ZSI PUNE P-5540 (3). •Valid as Systomus gracilus Plamoottil & Maji 2020. Current status: Valid as Systomus gracilus Plamoottil & Maji 2020. Cyprinidae: Smiliogastrinae. Distribution: Lower Ganges River basin, West Bengal, India. Habitat: freshwater.

immaculatus, Systomus McClelland [J.] 1839:284, 380, Pl. 44 (fig. 5) [Asiatic Researches v. 19 (pt 2); ref. 2923] Assam, India [a Day specimen]. Neotype: AMS B.7920. Neotype selected by Pethiyagoda et al. 2012:77 [ref. 32071]. •Synonym of Puntius sarana (Hamilton 1822) -- (Coad 1981:11 [ref. 22348]). •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904]). •Valid as Systomus immaculatus McClelland 1839 -- (Bleher 2018:792 [ref. 36647]). •Synonym of Systomus sarana (Hamilton 1822) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

kadoon, Barbus Bleeker [P.] 1853:60 [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 25 (art. 8); ref. 341] Coromandel, India. No types known. Appeared as "Barbus ? kadon Blkr., Kadoon Russ. no. 206" which is Russell 1803: Pl. 206 [ref. 13862]. •Synonymy of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [re. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

kadoon, Cyprinus Heckel [J. J.] 1843:1018 [28] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Vizagapatam, India. No types known. On p. 28 of separate. Based on Russell 1803, Pl. 206 [ref. 13862]. •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

kakus, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:153 [Histoire naturelle des poissons v. 16; ref. 1009] Tank near Tartoor, Vizagapatam, India. No types known. Bases on Russell, Coromondal 2 1803, 83, no. 205. •Synonym of Barbodes sarana (Hamilton 1822), subspecies sarana -- (Menon 1999:70 [ref. 24904]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

kunama, Cyprinus Cuvier [G.] in Russell 1829:273 (footnote 1) [Le Règne Animal (Edition 2) v. 2; ref. 995] Vizagapatham, India. As "C. kunama Russel, 204", in reference to Russel 1803:fig. 204. •Synonym of Systomus sarana (Hamilton 1922) with question -- (Kottelat 2013:166 [ref. 32989]). Current status: Uncertain as Systomus kunama (Cuvier 1829). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

kunamo, Cyprinus McClelland [J.] in Russell 1839:272, 340 [Asiatic Researches v. 19 (pt 2); ref. 2923] Not available, name mentioned in synonymy. •In the synonymy of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Nomen Nudum. Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae.

laticeps, Systomus Plamoottil [M.] 2016:92, Figs. 1-4 [International Journal of Fauna and Biological Studies v. 3 (no. 5) Part B; ref. 34814] Freshwater stream at Thiruvalla, Pathanamthitta district, Kerala, India. Holotype (unique): FBRC/ZSI F2316. •Valid as Systomus laticeps Plamoottil 2016 -- (Arunachalam et al. 2018:45 [ref. 36280], Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Valid as Systomus laticeps Plamoottil 2016. Cyprinidae: Smiliogastrinae. Distribution: Kerala, India. Habitat: freshwater.

liui, Barbus Fowler [H. W.] 1958:12 [Notulae Naturae (Philadelphia) No. 310; ref. 1470] Fort Stedman, southern Shan States, Myanmar, elevation 3000 feet. Syntypes: BMNH 1893.6.30.43-52. Unneeded replacement name for Barbus compressus Boulenger 1898, preoccupied by Barbus (Barbodes) compressus Day 1869, but replaced earlier by Barbus compressiformis Cockerell 1913. •Synonym of Puntius clavatus (McClelland 1845) -- (Talwar & Jhingran 1991:265 [ref. 20764] as lini). •Synonym of Systomus compressformis (Cockerell 1913) -- (Kottelat 2013:165 [ref. 32989]). Current status: Synonym of Systomus compressiformis (Cockerell 1913). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

macclellandi, Cyprinus Valenciennes [A.] in Cuvier & Valenciennes 1842:390 [Histoire naturelle des poissons v. 16; ref. 1009] Assam streams and probably occasionally in the large rivers, India. No type known. Original as M'clellandi. New name for Systomus immaculatus McClelland 1839. •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

martenstyni, Puntius Kottelat [M.] & Pethiyagoda [R.] 1991:303, Figs. 6-8 [Freshwater fishes of Sri Lanka; ref. 16752] Kalu River about 2 kilometers north of Pallegama, 7°33'N, 80°50'E, Mahaweli River basin, Marale District, Sri Lanka. Holotype: ZSM 27966 [now ZRC 38482]. Paratypes: CMK 6539 (2), 7078 (2), 7168 (11). Type catalog: Neumann 2011:255 [ref. 31652]. The holotype is at ZRC. •Valid as Systomus martenstyni (Kottelat & Pethiyagoda 1991) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Ren et al. 2020:[5] [ref. 37805], Sudasinghe et al. 2020:[4] [ref. 37737]). Current status: Valid as Systomus martenstyni (Kottelat & Pethiyagoda 1991). Cyprinidae: Smiliogastrinae. Distribution: South Asia: central highlands of Sri Lanka. Habitat: freshwater.

morehensis, Puntius Arunkumar [L.] & Tombi Singh [H.] 1998:254, Fig.3 [Journal of Nature Conservation: an International journal devoted to natural resource conservation and environment. v. 10 (no. 2); ref. 24931] Lairok Maru stream of Lokchao River near Moreh, 2 kilometers from Moreh Bazar, Manipur, India. Holotype: MUMF 240/1A. Paratypes: MUMF 240/3A (19). •Valid as Puntius morehensis Arunkumar & Tombi Singh 1998 -- (Arunkumar & Tombe Singh 2003:481 [ref. 26926]). •Synonym of Systomus sewelli (Prashad & Mukerji 1929) -- (Kotteat 2013:167 [ref. 32989] with question). Current status: Synonym of Systomus sewelli (Prashad & Mukerji 1929). Cyprinidae: Smiliogastrinae. Distribution: India (if valid). Habitat: freshwater.

orphoides, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:193 [Histoire naturelle des poissons v. 16; ref. 1009] Java, Indonesia. No types known. Type information: Roberts 1993:21-22 [ref. 21799]. Simultaneous synonym of Barbus rubripinnis Valenciennes 1842; Bleeker 1860:338 [ref. 380] gave preference to rubripinnis, with question; see Kottelat 2013:166 [ref. 32989]. Due to prevailing usage, the name orphoides is here retained. •Synonym of Puntius sarana (Hamilton 1822), but a valid subspecies -- (Talwar & Jhingran 1991:282 [ref. 20764], Arunkumar & Tombi Singh 1998:256 [ref. 24931], Arunachalam & Sankaranarayanan 1999:336 [ref. 24216], Karmakar 2000:29 [ref. 25662]). •Synonym of Barbodes sarana (Hamilton 1822), but a valid subspecies -- (Menon 1999:69 [ref. 24904]). •Valid as Puntius orphoides (Valenciennes 1842) -- (Roberts 1993:21 [ref. 21799], Kottelat et al. 1993:43 [ref. 23448], Kottelat & Lim 1995:233 [ref. 23210], Doi 1997:11 [ref. 22832], Doi 2000:396 [ref. 25097], Kottelat 2001:74 [ref. 25780], Vishwanath & Laisram 2004:136 [ref. 27863], Shantakumar & Vishwanath 2006:2279 [ref. 28676], Zhang et al. 2016:97 [ref. 34477]). •Synonym of Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989]). •Valid as Systomus orphoides (Valenciennes 1842) -- (Rainboth 1996:104 [ref. 22772], Parenti & Lim 2005:186 [ref. 28285], Pethiyagoda et al. 2012:77 [ref. 32071], Plamoottil 2014:145 [ref. 33650], Plamoottil 2016:96 [ref. 34814], Bleher 2018:794 [ref. 36647], Ren et al. 2020:[5] [ref. 37805]). Current status: Valid as Systomus orphoides (Valenciennes 1842). Cyprinidae: Smiliogastrinae. Distribution: Southeast Asia; introduced elsewhere. Habitat: freshwater.

pinnauratus, Cyclocheilichthys Day [F.] 1865:300 [Proceedings of the Zoological Society of London 1865 (pt 1); ref. 5295] Small pond, Cochin, India. Holotype (unique): ?ZSI 2728. Day specimens: BMNH 1889.2.1.4332-33 (2), FMNH 2309 (2), NMW 54468-70 (3), RMNH 2582 (1), ZIN 8216 (2). Type catalog: Whitehead & Talwar 1976:156 [ref. 23762]. Also appeared in Day 1865:209, Pl. 15, fig. 2 [ref. 1074]. •Mention under Puntius sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Talwar & Jhingran 1991:285 [ref. 20764], Indra 1993:184 [ref. 24009], Indra 1994:416 [ref. 24015]). •Synonym of Barbodes sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Menon 1999:72 [ref. 24904]). •Valid as Systomus pinnauratus (Day 1865) -- (Plamoottil 2014:142 [ref. 33650], Plamoottil 2016:92 [ref. 34814]). •Synonym of Systomus subnasutus (Valenciennes 1842) -- (Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989], Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

pleurotaenia, Puntius (Barbodes) Bleeker [P.] 1863:242 [Verslagen en Mededeelingen der Koninklijke Akademie van Wetenschappen. Afdeling Natuurkunde. v. 15; ref. 17011] Sri Lanka. Holotype (unique): whereabouts unknown. Also appeared in Bleeker 1864:13, Pl. 3 (fig. 2) [ref. 13925]. •Valid as Puntius pleurotaenia Bleeker 1863 -- (Pethiyagoda 1991:111 [ref. 20075], Talwar & Jhingran 1991:279 [ref. 20764], Silva et al. 2008:62 [ref. 29719]). •Valid as Systomus pleurotaenia (Bleeker 1863) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Ren et al. 2020:[5] [ref. 37805], Sudasinghe et al. 2020:[4] [ref. 37737]). Current status: Valid as Systomus pleurotaenia (Bleeker 1863). Cyprinidae: Smiliogastrinae. Distribution: South Asia: southwestern Sri Lanka. Habitat: freshwater.

polydori, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:170 [Histoire naturelle des poissons v. 16; ref. 1009] Mumbai, India. Holotype (unique): MNHN 0000-3827. Type catalog: Bertin & Estève 1948:25 [ref. 19574]. •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904]). •Uncertain in genus Puntius -- (Plamoottil & Johnson 2020:99 [ref. 37958]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

roseipinnis, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:169 [Histoire naturelle des poissons v. 16; ref. 1009] Puducherry [Pondicherry], India. Syntypes: MNHN 1405 (2). •Valid as Puntius roseipinnis (Valenciennes 1842) -- (Talwar & Jhingran 1991:281 [ref. 20764]). •Synonym of Barbodes sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Menon 1999:72 [ref. 24904]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 3289]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

rubripinna, Barbus van Hasselt [J. C.] 1823:132 [Algemeene Konst- en Letter-bode voor het Jaar II Deel (no. 35); ref. 5963] Not available, name only; from Batavia, Java (see Alfred 1961:85 [ref. 20553], Kottelat 1987:370 [ref. 5962]). Also in van Hasselt 1824:375 [ref. 5964]. •In the synonymy of Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989]). Nomen Nudum. Current status: Synonym of Systomus rubripinnis (Valenciennes 1842). Cyprinidae: Smiliogastrinae.

rubripinnis, Barbus Valenciennes [A.] (ex Kuhl & van Hasselt) in Cuvier & Valenciennes 1842:194 [Histoire naturelle des poissons v. 16; ref. 1009] Java, Indonesia. Syntypes: whereabouts unknown. See also remarks under Barbus orphoides Valenciennes 1842. •Synonym of Barbodes sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Menon 1999:69 [ref. 24904]). •Synonym of Puntius sarana (Hamilton 1822). •Valid as Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989], Ng et al. 2019:525 [ref. 36606]). •In the synonymy of Systomus orphoides (Valenciennes 1842). Current status: Synonym of Systomus orphoides (Valenciennes 1842). Cyprinidae: Smiliogastrinae. Distribution: Southeast Asia: Myanmar, Malaya (Malaysia) and Indonesia [if valid]. Habitat: freshwater.

rufus, Systomus Plamoottil [M.] 2014:142, Fig. 17 [International Journal of Fauna and Biological Studies v. 1 (no. 6) Part C; ref. 33650] Venpala, Manimala River, Kerala, India. Holotype: ZSI/WGRC/IR/2367. Paratypes: ZSI/WGRC/IR/2368 (1). •Valid as Systomus rufus Plamoottil 2014 -- (Plamoottil 2016:95 [ref. 34814], Arunachalam et al. 2018:45 [ref. 36280], Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Valid as Systomus rufus Plamoottil 2014. Cyprinidae: Smiliogastrinae. Distribution: Manimala River drainage, Kerala, India. Habitat: freshwater.

russellii, Barbus Günther [A.] 1868:121 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Indus River, Sabzilkot, India. Syntypes: BMNH 1860.3.19.732 (1), 1860.3.19.856 (1). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

saberi, Puntius Datta [A. K.] & Karmakar [A. K.] 1981:179, Fig. 1 [Bulletin of the Zoological Survey of India v. 3 (no. 3); ref. 8616] Cheroguru Talao, Pujaripal village, 23 kilometers from Sukma, Bastar District, Madhya Pradesh, India. Holotype: ZSI FF1398. Paratypes: ZSI FF1399 (12). See Das 2003:360 [ref. 27706]. •Synonym of Puntius sarana (Hamilton 1822) -- (Talwar & Jhingran 1991:283 [ref. 20764]). •Synonym of Barbodes sarana (Hamilton 1822), subspecies sarana -- (Menon 1999:72 [ref. 24904]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

sarana, Cyprinus Hamilton [F.] 1822:307, 388 [An account of the fishes found in the river Ganges; ref. 2031] Ponds and rivers of Bengal. No types known. Hamilton's original, unpublished illustration reproduced by Britz 2019:Pl. 180 [ref. 37532]. •Valid as Puntius sarana (Hamilton 1822), subspecies sarana -- (Karmakar 2000:29 [ref. 25662], Shrestha 2008:80 [ref. 29923], Gurung et al. 2013:4882 [ref. 32969], Arunkumar & Manimekalan 2018:1739 [ref. 36172]). •Valid as Barbodes sarana (Hamilton 1822) -- (Rema Devi & Raghunathan 1999:150 [ref. 24759], Rema Devi & Raghunathan 1999:167 [ref. 24760], Rema Devi et al. 1999:155 [ref. 24758], Menon 1999:70 [ref. 24904], Rafique 2000:323 [ref. 25220], Yadav 2003:13 [ref. 27711], Ataur Rahman 2003:145 [ref. 31338]). •Valid as Puntius sarana (Hamilton 1822) -- (Shrestha 1978:36 [ref. 15970], Ataur Rahman 1989:126 [ref. 24860], Coad 1981:11 [ref. 22348], Pethiyagoda 1991:113 [ref. 20075], Talwar & Jhingran 1991:283 [ref. 20764], Indra 1991:239 [ref. 24005], Sen 1995:515 [ref. 23778], Doi 1997:11 [ref. 22832], Vu & Nguyen 1997:26 [ref. 26438], Arunkumar & Tombi Singh 1998:256 [ref. 24931], Manimekalan 1998:436 [ref. 24989], Arunachalam & Sankaranarayanan 1999:336 [ref. 24216], Bijukumar & Sushama 2000:185 [ref. 25703], Arunkumar & Tombe Singh 2003:481 [ref. 26926], Rafique et al. 2003:97 [ref. 27173], Vishwanath & Laisram 2004:136 [ref. 27863], Shantakumar & Vishwanath 2006:2279 [ref. 28676]). •Valid as Systomus sarana (Hamilton 1822) -- (Mirza 2003:9 [ref. 27265], Pethiyagoda et al. 2012:77 [ref. 32071], Kottelat 2013:166 [ref. 32989], Plamoottil 2014:145 [ref. 33650], Plamoottil 2016:95 [ref. 34814], Arunachalam et al. 2018:45 [ref. 36280], Suresh et al. 2018:113 [ref. 36495], Bleher 2018:798 [ref. 36647], Plamoottil & Maji 2020:1035 [ref. 37708], Ren et al. 2020:[5] [ref. 37805], Sudasinghe et al. 2020:[4] [ref. 37737]). Current status: Valid as Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Distribution: South Asia: Nepal, Bhutan, India, Sri Lanka, Myanmar, Pakistan and Bangladesh. Habitat: freshwater, brackish.

sarananella, Barbus Bleeker [P.] 1849:16 [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 23 (no. 12); ref. 16883] Serayu River, Banjumas; Gombong; Bogowonto River, Purworedjo, Java, Indonesia. •Synonym of Barbodes sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Menon 1999:69 [ref. 24904] as saranella). •Synonym of Puntius sarana (Hamilton 1822). •Synonym of Systomus sarana (Hamilton 1822) based on placement of sarana. •Synonym of Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus rubripinnis (Valenciennes 1842). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

sewelli, Barbus Prashad [B.] & Mukerji [D. D.] 1929:197, Pl. 9 (figs. 1, 1a-b) [Records of the Indian Museum (Calcutta) v. 31 (pt 3); ref. 3558] Indawgyi Lake, Myitkyina District, Upper Myanmar. Holotype (unique): ZSI F10910/1. Type catalog: Menon & Yazdani 1968:106 [ref. 20743]. See Das 2003:354 [ref. 27706] with date as 1930. •Synonym of Puntius sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Talwar & Jhingran 1991:282 [ref. 20764]). •Synonym of Barbodes sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Menon 1999:70 [ref. 24904]). •Synonym of Puntius sarana (Hamilton 1822). •Valid as Systomus sewelli (Prashad & Mujkerji 1929) -- (Kottelat 2013:167 [ref. 32989]). Current status: Valid as Systomus sewelli (Prashad & Mukerji 1929). Cyprinidae: Smiliogastrinae. Distribution: Myanmar. Habitat: freshwater, brackish.

spilurus, Barbus Günther [A.] 1868:114 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Sri Lanka. Syntypes: BMNH 1852.2.19.124 (1), 1858.10.19.86-91 (6), 1859.5.31.47 (1). •Synonym of Puntius sarana (Hamilton 1822), but a valid subspecies -- (Talwar & Jhingran 1991:284 [ref. 20764], Arunkumar & Manimekalan 2018:1739 [ref. 36172] as spirulus). •Valid as Systomus spilurus (Günther 1868) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Plamoottil 2014:145 [ref. 33650], Plamoottil 2016:96 [ref. 34814], Arunachalam et al. 2018:46 [ref. 36280], Ren et al. 2020:[5] [ref. 37805]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989], Sudasinghe et al. 2020:[16] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

stedmanensis, Barbus Annandale [N.] (ex Boulenger) 1918:47 [Records of the Indian Museum (Calcutta) v. 14; ref. 127] Unneeded replacement name for Barbus compressus Boulenger 1893, preoccupied by Barbus compressus Day 1870; replaced earlier by Barbus compressiformis Cockerell 1913. •Synonym of Puntius clavatus (McClelland 1845) -- (Talwar & Jhingran 1991:265 [ref. 20764]). •Synonym of Systomus compresssiformis (Cockerell 1913) -- (Kottelat 2013:165 [ref. 32989], Fuke et al. 2021:[3] [ref. 38221]). Replacement Name. Current status: Synonym of Systomus compressiformis (Cockerell 1913). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

subnasutus, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:154 [Histoire naturelle des poissons v. 16; ref. 1009] Puducherry [Ponducherry], India. Holotype (unique): MNHN 0000-3823. Type catalog: Bertin & Estève 1948:24 [ref. 19574]. •Synonym of Puntius sarana (Hamilton 1822), but a valid subspecies -- (Talwar & Jhingran 1991:285 [ref. 20764], Indra 1993:184 [ref. 24009], Rema Devi 1993:198 [ref. 24010], Indra 1994:416 [ref. 24015], Manimekalan 1998:436 [ref. 24989], Arunachalam et al. 2001:128 [ref. 25692], Bijukumar & Sushama 2000:185 [ref. 25703]). •Synonym of Barbodes sarana (Hamilton 1822), but a valid subspecies -- (Rema Devi et al. 1996:132 [ref. 24021], Rema Devi & Raghunathan 1999:150 [ref. 24759], Rema Devi & Raghunathan 1999:167 [ref. 24760], Rema Devi et al. 1999:155 [ref. 24758], Menon 1999:70 [ref. 24904], Raju Thomas et al. 2000:444 [ref. 25319], Raju Thomas et al. 2002:49 [ref. 26359], Yadav 2003:13 [ref. 27711]). •Valid as Systomus subnasutus (Valenciennes 1842) -- (Plamoottil 2014:145 [ref. 33650], Plamoottil 2016:95 [ref. 34814], Arunachalam et al. 2018:45 [ref. 36280], Bleher 2018:800 [ref. 36647], Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989], Sudasinghe et al. 2020:[12] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

takhoaensis, Puntius Mai [D. Y.] (ex Hao & Hoa) 1978:110 [Identification of the fresh-water fishes of North Viet Nam v. 1; ref. 13511] Northern Vietnam. Syntypes: DVZUT. Apparently unavailable from Hao & Hoa 1969 according to Kottelat 1989:1102 [ref. 14361], but Kottelat 2001 [ref. 25482] reversed this interpretation and dated to Nguyen & Doan [= Hao & Hoa] 1969. •See the account of Puntius takhoaensis Nguyen & Doan 1969. •In the synonymy of Puntius takhoaensis Nguyen & Doan 1969 -- (Kottelat 2001:37 [ref. 25482]). •Should be referred to the genus Systomus -- (Roberts & Catania 2008:94 [ref. 29794]). 1°Homonym. Current status: Synonym of Systomus takhoaensis (Nguyen & Doan 1969). Cyprinidae: Smiliogastrinae.

timbiri, Puntius Deraniyagala [P. E. P.] 1963:63, Pl. 1 (figs. 2-3) [Spolia Zeylanica v. 30 (pt 1); ref. 4605] Valavé River at Timbirigas-mankada, Sri Lanka, depth about 5 feet. Holotype (unique): NMSL FF801. Type information: Pethiyagoda 1991:336 [ref. 20075]. •Synonym of Puntius sarana (Hamilton 1822), subspecies spilurus (Günther 1868) -- (Talwar & Jhingran 1991:285 [ref. 20764]). •Valid as Systomus timbiri (Deraniyagala 1963) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Ren et al. 2020:[5] [ref. 37805]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989] with question, Sudasinghe et al. 2020:[16] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

tripunctatus, Systomus Jerdon [T. C.] 1849:316 [Madras Journal of Literature and Science v. 15 (pt 2); ref. 4902] Near the coast in Canara [Karnataka], southern India. No types known. •Status uncertain -- (Kawate et al. 2014:242 [ref. 33433]). Current status: Uncertain as Systomus tripuinctatus Jerdon 1849. Cyprinidae: Smiliogastrinae. Habitat: freshwater.