Institute for Biodiversity Science and Sustainability
Online Version, Updated 14 Mar 2025

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Eschmeyer's Catalog of Fishes
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         Comments: Richard van der Laan, Ron Fricke

    Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the family or subfamily Grammatidae: [ 19 ] records


anabantoides, Lipogramma Böhlke [J. E.] 1960:8, Fig. 1 [Notulae Naturae (Philadelphia) No. 330; ref. 5941] Wood Cay, off Grand Bahama Island, Bahamas, 26°44'15"N, 79°02'37"W. Holotype: ANSP 92412. Paratypes: ANSP 92413 (1), 92414 (1, c&s), 92415 (1). Type catalog: Böhlke 1984:111 [ref. 13621]. •Valid as Lipogramma anabantoides Böhlke 1960 -- (Gilmore & Jones 1988:443 [ref. 12879], Gilmore 1997:787 [ref. 22975], Schmitter-Soto et al. 2000:158 [ref. 27754], Collette et al. 2003:107 [ref. 26784], Mooi & Gill 2003:1373 [ref. 27062], Smith et al. 2003:26 [ref. 27621], McEachran & Fechhelm 2005:197 [ref. 28815], Page et al. 2013:134 [ref. 32708], Baldwin et al. 2016:68 [ref. 34958], Baldwin et al. 2018:138, 152 [ref. 35754], Tornabene et al. 2018:91 [ref. 36382], Page et al. 2023:159 [ref. 40505], Fricke et al. 2024:382 [ref. 41622]). Current status: Valid as Lipogramma anabantoides Böhlke 1960. Grammatidae. Distribution: Western Atlantic: Bahamas south to Venezuela, including northern Gulf of Mexico (U.S.A.) and northern and western Caribbean Sea. IUCN (2015): Least Concern. Habitat: marine.

barrettorum, Lipogramma Baldwin [C. C.], Nonaka [A.] & Robertson [D. R.] in Baldwin, Tornabene, Robertson, Nonaka & Gilmore 2018:132, Figs. 1-2, 3A [ZooKeys No. 729; ref. 35754] Curaçao, west of Substation Curaçao downline, 12.083197N, 68.899058W, depth 161 meters. Holotype: USNM 440439. Paratypes: UF, USNM. •Valid as Lipogramma barrettorum Baldwin, Nonaka & Robertson 2018 -- (Tornabene et al. 2018:91 [ref. 36382]). Current status: Valid as Lipogramma barrettorum Baldwin, Nonaka & Robertson 2018. Grammatidae. Distribution: Curaçao. Habitat: marine.

brasiliensis, Gramma Sazima [I.], Gasparini [J. L.] & Moura [R. L. de] 1998:40, Figs. 1-4 [aqua, Journal of Ichthyology and Aquatic Biology v. 3 (no. 1); ref. 23432] Roncadeira, Marataízes, Espírito Santo, Brazil, 21°04'S, 40°44'W, depth 15-18 meters. Holotype: MZUSP 51089. Paratypes: MZUSP 36263 (1), 44669 (2), 46141 (1), 46320 (2), 46375 (1), 46820 (1), 51106-07 (2, 1), 51108 (2, c&s), 51109 (1), 51319 (2); USNM 343209 (1); ZUEC 2752 (1), 3052-53 (1, 1), 3072 (1), 3091-92 (2, 1), 3095 (1). •Valid as Gramma brasiliensis Sazima, Gasparini & Moura 1998 -- (Rocha & Rosa 2001:989 [ref. 25909], Mooi & Gill 2003:1370 [ref. 27062], Moura & Menezes in Menezes et al. 2003:77 [ref. 27192], Victor & Randall 2010:865 [ref. 31057], Carvalho-Filho in Marceniuk et al. 2021:252 [ref. 38945], Carvalho-Filho 2023:128 [ref. 40480]). Current status: Valid as Gramma brasiliensis Sazima, Gasparini & Moura 1998. Grammatidae. Distribution: Southwestern Atlantic: Ceará south to São Paulo (Brazil) endemic. Habitat: marine.

dejongi, Gramma Victor [B. C.] & Randall [J. E.] 2010:866, Figs. 1-3 [Zoological Studies v. 49 (no. 6); ref. 31057] Sancti Spiritus, off Trinidad, 21.7896°N, 80.0579°W, Cuba, depth about 20 meters. Holotype: USNM 398656. Paratypes: UF 178386 (1); USNM 398657 (1). Plus aquarium photo. •Valid as Gramma dejongi Victor & Randall 2010. Current status: Valid as Gramma dejongi Victor & Randall 2010. Grammatidae. Distribution: Cuba. Habitat: marine.

evides, Lipogramma Robins [C. R.] & Colin [P. L.] 1979:41, Figs. 1-2, 6a [Bulletin of Marine Science v. 29 (no. 1); ref. 6986] Arrowsmith Bank, western Caribbean Sea, 21°05'N, 86°23'W, depth 146-265 meters. Holotype: ANSP 134329. Paratypes: ANSP 134330-33 (2, 1, 1); FMNH 82583 (1). Type catalog: Böhlke 1984:111 [ref. 13621]. •Valid as Lipogramma evides Robins & Colin 1979 -- (Robins & Ray 1986:147 [ref. 23100], Gilmore & Jones 1988:441 [ref. 12879], Gilmore 1997:782 [ref. 22975], Claro et al. 2000:22 [ref. 26458], Schmitter-Soto et al. 2000:158 [ref. 27754], Mooi & Gill 2003:1373 [ref. 27062], Nelson et al. 2004:130 [ref. 27807], Page et al. 2013:134 [ref. 32708], Baldwin et al. 2016:63 [ref. 34958], Baldwin et al. 2018:138, 152 [ref. 35754], Tornabene et al. 2018:91 [ref. 36382], Robertson et al. 2020:155 [ref. 38098], Page et al. 2023:159 [ref. 40505], Fricke et al. 2024:382 [ref. 41622]). Current status: Valid as Lipogramma evides Robins & Colin 1979. Grammatidae. Distribution: Western Atlantic: Bahamas south to Venezuela, including Caribbean Sea. IUCN (2015): Least Concern. Habitat: marine.

flavescens, Lipogramma Gilmore [R. G.] & Jones [R. S.] 1988:436, Figs. 1 (upper), 2 [Bulletin of Marine Science v. 42 (no. 3); ref. 12879] French Bay, 0.74 kilometers west of Low Cay, San Salvador, Bahamas, 23°36.3'N, 74°29.4'W, depth 285 fathoms. Holotype (unique): USNM 276559. •Valid as Lipogramma flavescens Gilmore & Jones 1988 -- (Gilmore 1997:787 [ref. 22975], Mooi & Gill 2003:1373 [ref. 27062], Baldwin et al. 2018:151 [ref. 35754], Tornabene et al. 2018:91 [ref. 36382]). Current status: Valid as Lipogramma flavescens Gilmore & Jones 1988. Grammatidae. Distribution: Western Atlantic. Habitat: marine.

haberorum, Lipogramma Baldwin [C. C.], Nonaka [A.] & Robertson [D. R.] in Baldwin, Robertson, Nonaka & Tornabene 2016:59, Figs. 5B, 7 [ZooKeys No. 638; ref. 34958] Curaçao, southwest tip of Klein Curaçao, southern Caribbean, 11.975783N, 68.646192W, depth 152 meters. Holotype: USNM 422679. Paratypes: USNM. Incorrect original spelling haberi emended to haberorum, as the species name was based on more than one person, Art. 31.1.2. •Valid as Lipogramma haberorum Baldwin, Nonaka & Robertson 2016 -- (Baldwin et al. 2018:138, 152 [ref. 35754] as haberi, Tornabene et al. 2018:91 [ref. 36382]). Current status: Valid as Lipogramma haberorum Baldwin, Nonaka & Robertson 2016. Grammatidae. Distribution: Curaçao, southern Caribbean. Habitat: marine.

hemichrysos, Gramma Mowbray [L. L.] in Breder 1927:42, Fig. 21 [Bulletin of the Bingham Oceanographic Collection Yale University v. 1 (art. 1); ref. 635] Royal Island harbor, Bahamas. Holotype: YPM 526. Paratypes: YPM 215 (1, missing), 216 (1), 218 (1), 219 (1), 220 (13). Additional material: YPM 217 (1). Type catalog: Moore & Boardman 1991:13 [ref. 16741]. Current status: Uncertain. Grammatidae. Habitat: marine.

idabeli, Lipogramma Tornabene [L.], Robertson [D. R.] & Baldwin [C. C.] 2018:83, Figs. 2-4 [ZooKeys No. 809; ref. 36382] Reef slope off Halfmoon Bay, West End, Roatan, Honduras, 16.305557, -86.597669, depth 165 meters. Holotype: USNM 444940. Paratypes: MUVS-V, UF, UW. •Valid as Lipogramma idabeli Tornabene, Robertson & Baldwin 2018. Current status: Valid as Lipogramma idabeli Tornabene, Robertson & Baldwin 2018. Grammatidae. Distribution: Off Roatan, Honduras. Habitat: marine.

klayi, Lipogramma Randall [J. E.] 1963:85, Pl. 1 (fig. B) [Studies on the Fauna of Curacao and other Caribbean Islands v. 19 (no. 80); ref. 5209] South of Hotel Avila, Willemstad, Curaçao, depth 240 feet. Holotype: USNM 179000. Paratypes: RMNH 24607 (1), UPRM 1129 (1). •Valid as Lipogramma klayi Randall 1963 -- (Robins & Ray 1986:148 [ref. 23100], Gilmore & Jones 1988:439 [ref. 12879], Mooi & Gill 2003:1373 [ref. 27062], Baldwin et al. 2016:68 [ref. 34958], Baldwin et al. 2018:138, 153 [ref. 35754], Tornabene et al. 2018:92 [ref. 36382], Robertson et al. 2020:155 [ref. 38098]). Current status: Valid as Lipogramma klayi Randall 1963. Grammatidae. Distribution: Western Atlantic. Habitat: marine.

levinsoni, Lipogramma Baldwin [C. C.], Nonaka [A.] & Robertson [D. R.] in Baldwin, Robertson, Nonaka & Tornabene 2016:50, Figs. 1B, 1C, 2, 4 (right) 5A, 6A [ZooKeys No. 638; ref. 34958] Curaçao, 12.083197N, 68.899058W, depth 137-146 meters. Holotype: USNM 406139. Paratypes: ANSP, USNM. Plus additional non-type specimens. •Valid as Lipogramma levinsoni Baldwin, Nonaka & Robertson 2016 -- (Baldwin et al. 2018:138, 151 [ref. 35754], Tornabene et al. 2018:91 [ref. 36382], Robertson et al. 2020:155 [ref. 38098]). Current status: Valid as Lipogramma levinsoni Baldwin, Nonaka & Robertson 2016. Grammatidae. Distribution: Western Atlantic: Saint Eustatius, Jamaica, Dominica, Curaçao and Bonaire. Habitat: marine.

linki, Gramma Starck [W. A., II] & Colin [P. L.] 1978:147, Figs. 1-3 [Bulletin of Marine Science v. 28 (no. 1); ref. 8876] Dropoff west of airport, Cozumel Island, Mexico, Caribbean Sea, depth 70-80 meters. Holotype: ANSP 117314. Paratypes: ANSP 117312-13 (13, 1), 117315 (2); UF 229140 [ex UMML 29140] (1); UPR 32828 (2); USNM 205946 (1). Additional material: ANSP 13446 (1); UF 232933 [ex UMML 32933] (1); UPR 3417-18 (3, 5). Type catalog: Böhlke 1984:111 [ref. 13621]. •Valid as Gramma linki Starck & Colin 1978 -- (Robins & Ray 1986:147 [ref. 23100], Schmitter-Soto et al. 2000:158 [ref. 27754], Mooi & Gill 2003:1373 [ref. 27062], Nelson et al. 2004:130 [ref. 27807], Victor & Randall 2010:865 [ref. 31057], Page et al. 2013:133 [ref. 32708], Smith-Vaniz & Jelks 2014:43 [ref. 33341], Robertson et al. 2020:155 [ref. 38098], Page et al. 2023:159 [ref. 40505], Fricke et al. 2024:382 [ref. 41622]). Current status: Valid as Gramma linki Starck & Colin 1978. Grammatidae. Distribution: Western Atlantic: Bahamas south to Trinidad and Tobago, including Caribbean Sea. IUCN (2015): Least Concern. Habitat: marine.

loreto, Gramma Poey [F.] 1868:296 [Repertorio Fisico-Natural de la Isla de Cuba v. 2; ref. 3505] Matanzas, Cuba. Holotype (unique): MCZ 10031. •Valid as Gramma loreto Poey 1868 -- (Robins & Ray 1986:147 [ref. 23100], Cervigón 1991:381 [ref. 24472], Sazima et al. 1998:41 [ref. 23432], Smith-Vaniz et al. 1999:220 [ref. 25013], Schmitter-Soto et al. 2000:158 [ref. 27754], Collette et al. 2003:107 [ref. 26784], Mooi & Gill 2003:1373 [ref. 27062], Smith et al. 2003:26 [ref. 27621], Nelson et al. 2004:130 [ref. 27807], McEachran & Fechhelm 2005:196 [ref. 28815], Victor & Randall 2010:865 [ref. 31057], Page et al. 2013:133 [ref. 32708], Smith-Vaniz & Jelks 2014:43 [ref. 33341], Robertson et al. 2016:145 [ref. 34961], Robertson et al. 2020:155 [ref. 38098], Escobar-Sierra et al. 2021:79 [ref. 38844], Bennema & van Moorsel 2022:71 [ref. 39585], Page et al. 2023:159 [ref. 40505], Fricke et al. 2024:382 [ref. 41622]). Current status: Valid as Gramma loreto Poey 1868. Grammatidae. Distribution: Western Atlantic: Bahamas south to Trinidad and Tobago, including Gulf of Mexico and Caribbean Sea. IUCN (2015): Least Concern. Habitat: marine.

melacara, Gramma Böhlke [J. E.] & Randall [J. E.] 1963:43, Fig. 3 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 115 (no. 2); ref. 4245] About 1/2 mile north of the western end of Treasure Island (Salt Cay), New Providence area, Bahamas, depth 165 feet. Holotype: ANSP 94687. Paratypes: ANSP 94686 (1), 94688-90 (10, 5, 1); SMF 6830 [ex UMML 9428] (1); UF 204948 [ex UMML 4948] (1), ?209420 [ex UMML 9420] (4), 209427-28 [ex UMML 9427-28] (15, 2); UPRM 1040 (1); USNM [ex ANSP 94691] 199486 (3). Type catalog: Böhlke 1984:111 [ref. 13621]. •Valid as Gramma melacara Böhlke & Randall 1963 -- (Robins & Ray 1986:147 [ref. 23100], Sazima et al. 1998:41 [ref. 23432], Schmitter-Soto et al. 2000:158 [ref. 27754], Mooi & Gill 2003:1373 [ref. 27062], Smith et al. 2003:26 [ref. 27621], Nelson et al. 2004:130 [ref. 27807], Victor & Randall 2010:865 [ref. 31057], Page et al. 2013:134 [ref. 32708], Escobar-Sierra et al. 2021:79 [ref. 38844], Page et al. 2023:159 [ref. 40505], Fricke et al. 2024:382 [ref. 41622]). Current status: Valid as Gramma melacara Böhlke & Randall 1963. Grammatidae. Distribution: Western Atlantic: Bahamas south to Venezuela, including northern and western Caribbean Sea. IUCN (2015): Least Concern. Habitat: marine.

regia, Lipogramma Robins [C. R.] & Colin [P. L.] 1979:44, Fig. 3-4, 6b [Bulletin of Marine Science v. 29 (no. 1); ref. 6986] 6 miles south of La Parguera at edge of island shelf, Puerto Rico, depth 75 meters. Holotype: ANSP 138115. Paratypes: ANSP 134333 (1). Type catalog: Böhlke 1984:111 [ref. 13621]. Lipogramma is feminine, so correct species spelling is regia as originally proposed. •Valid as Lipogramma regia Robins & Colin 1979 -- (Robins & Ray 1986:148 [ref. 23100], Gilmore & Jones 1988:439 [ref. 12879] as regium, Mooi & Gill 2003:1373 [ref. 27062] as regium, Page et al. 2013:134 [ref. 32708] as regium, Baldwin et al. 2018:138, 152 [ref. 35754], Tornabene et al. 2018:91 [ref. 36382], Robertson et al. 2020:155 [ref. 38098], Page et al. 2023:159 [ref. 40505]). Current status: Valid as Lipogramma regia Robins & Colin 1979. Grammatidae. Distribution: Western Atlantic. Habitat: marine.

robinsi, Lipogramma Gilmore [R. G.] 1997:783, Fig. 1 [Bulletin of Marine Science v. 60 (no. 3); ref. 22975] East coast Glover Reef, Belize, 16°49.35'N, 87°44.04'W, depth 202 meters. Holotype: USNM 430015. Paratypes: UF 101375 (1). •Valid as Lipogramma robinsi Gilmore 1997 -- (Claro et al. 2000:22 [ref. 26458], Mooi & Gill 2003:1373 [ref. 27062], Baldwin et al. 2016:68 [ref. 34958] as 'robinsi', Baldwin et al. 2018:152 [ref. 35754], Tornabene et al. 2018:91 [ref. 36382]). Current status: Valid as Lipogramma robinsi Gilmore 1997. Grammatidae. Distribution: Western Atlantic. Habitat: marine.

rosea, Lipogramma Gilbert [C. R.] in Robins & Colin 1979:48, Figs. 5, 6d [Bulletin of Marine Science v. 29 (no. 1); ref. 6986] Northwestern coast of Isla Providencia, northwest of Santa Catalina Island, Caribbean sea, Colombia, depth 32-33 meters. Holotype (unique): UF 19636. Author is C. R. Gilbert. Lipogramma is feminine, based on line as in lateral line or lines on the body; the correct species spelling is rosea as originally proposed. •Valid as Lipogramma roseum Gilbert 1979 -- (Gilmore & Jones 1988:439 [ref. 12879], Gilmore 1997:782 [ref. 22975], Mooi & Gill 2003:1373 [ref. 27062], Dennis et al. 2004:74 [ref. 31601]). •Valid as Lipogramma rosea Gilbert 1979 -- (Baldwin et al. 2018:152 [ref. 35754]). Current status: Valid as Lipogramma rosea Gilbert 1979. Grammatidae. Distribution: Western Atlantic. Habitat: marine.

schrieri, Lipogramma Baldwin [C. C.], Nonaka [A.] & Robertson [D. R.] in Baldwin, Tornabene, Robertson, Nonaka & Gilmore 2018:138, Figs. 3B, 5-6 [ZooKeys No. 729; ref. 35754] Curaçao, Jan Thiel Bay, 12.0746N, 68.8825W, depth 197 meters. Holotype: USNM 431722. Paratypes: USNM. •Valid as Lipogramma schrieri Baldwin, Nonaka & Robertson 2018 -- (Tornabene et al. 2018:91 [ref. 36382], Page et al. 2023:159 [ref. 40505]). Current status: Valid as Lipogramma schrieri Baldwin, Nonaka & Robertson 2018. Grammatidae. Distribution: Western Atlantic: Curaçao. Habitat: marine.

trilineata, Lipogramma Randall [J. E.] 1963:81, Pl. 1 (fig. A) [Studies on the Fauna of Curacao and other Caribbean Islands v. 19 (no. 80); ref. 5209] Off the jail at Willemstad, Curaçao, depth 160 feet. Holotype: USNM 179004. Paratypes: ANSP 98681 (1), RMNH 24608 (1), UF 211141 [ex UMML 11141] (1), UPRM 1127-28 (1, 1), USNM 179003 (1). Type catalog: Böhlke 1984:111 [ref. 13621]. Lipogramma is feminine, based on line as in lateral line or lines on the body; the correct species spelling is trilineata as originally proposed. •Valid as Lipogramma trilineatum Randall 1963 -- (Gilmore & Jones 1988:439 [ref. 12879], Gilmore 1997:782 [ref. 22975], Schmitter-Soto et al. 2000:158 [ref. 27754], Mooi & Gill 2003:1373 [ref. 27062], Smith et al. 2003:26 [ref. 27621], Nelson et al. 2004:130 [ref. 27807], McEachran & Fechhelm 2005:198 [ref. 28815], Page et al. 2013:134 [ref. 32708]). •Valid as Lipogramma trilineata Randall 1963 -- (Robins & Ray 1986:148 [ref. 23100], Baldwin et al. 2016:68 [ref. 34958], Baldwin et al. 2018:138, 153 [ref. 35754], Tornabene et al. 2018:92 [ref. 36382], Robertson et al. 2020:155 [ref. 38098], Page et al. 2023:159 [ref. 40505], Fricke et al. 2024:382 [ref. 41622]). Current status: Valid as Lipogramma trilineata Randall 1963. Grammatidae. Distribution: Western Atlantic: eastern-central Florida and Bahamas south to Venezuela, including Gulf of Mexico and northern and western Caribbean Sea. IUCN (2015): Least Concern. Habitat: marine.


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