Online Version, Updated 15 Apr 2014


Catalog of Fishes
Select the database to search:  
Include unavailable names

Catalog of Fishes - version of 15 Apr 2014 (Continuously updated since the early 1980s.)

By default, search results will not include item 3 names -- unavailable names that detract from the main list of available names. These are mostly names mentioned in synonymy, names in lists, etc. They are still in the database, so if you want to see them, check the box labeled Include unavailable names before searching.

An important work was produced by Maurice Kottelat in late 2013: "The fishes of the inland waters of southeast Asia: a catalogue and core bibliography of the fishes known to occur in freshwaters, mangroves and estuaries." [ref. 32989] I began going through it, and I am now on page 110.

[Note for Apr. 15, 2014: I have now completed pages to 307. Christopher Scharpf has assisted with this section. He has been reviewing data entry for Kottelat (2013), improving coverage of this important work.]

The Catalog went through a similar long history. It was published in 3 volumes in 1990, and added to with updates about every six or eight weeks on the Internet. This amounts to 60,000 names of fishes and 34,000 references -- nearly all seen by Eschmeyer and/or assistants. Perhaps he did not know that Eschmeyer visited most major museums around the world (gathering information on types -- with bottles examined for smaller collections). This is why I am cited for BMNH specimens, although I checked specimens or records at other museums, and looked at nearly all major type catalogs. For localities, I just used what was found in the current literature (with modifications). Kottelat has on-the-ground experience in the localities described, and some of his localities localities are more accurate. And he was able to allocate earlier synonymous names more accurately.

We agree on nearly all primary type specimens (but I also give additional types), and we agree on the spelling of nearly all names.

Where do we differ? I am missing some information on old synonyms, and I should have paid closer attention to early works (mostly freshwater, and those appearing in journals not in a language I was familiar with). So Kottelat found a few additional synonyms and more unavailable names. We are nearly in agreement on dates of publication (except for a few Bleeker dates, which will be changed).

But here are differences:

  1. Bloch and Schneider. Kottelat treats this as Schneider in Bloch & Schneider. His statement that the text was written by Schneider has been questioned. Some names are clearly Schneider's, but others are not. Most workers treat this as Bloch and Schneider.
  2. Spelling of names. We both treat one person as an author, differing from the publication in some cases. We do differ on Rafinesque and Rafinesque-Schmaltz, both referring to the same person. This reaches an extreme for South American authors. I wrote some living ones and asked how they wanted to be cited in international journals. One person said she found her name appeared in 14 ways (including initials). But we differ on Lacepède versus La cepède -- The former was used by North American authors and the authors of the many catalogs by Bauchot and coworkers for catalogs at the MNHN in Paris. How the name is spelled in other disciplines would be helpful.

He also does not follow the Code in at least 2 cases. One allows use of common spellings, such as Engraulidae, instead of Engraulididae). The second says that names that have not been used as valid in fishes since 1900 can be eliminated. Another effort has been made by Kottelat to find other names that are not available -- I have about a third of them as type 3 names. These are names mentioned in passing, names in synonymies without a description, etc., that should be eliminated. I propose that these names be moved to a separate database. Kottelat is on the Commission, but doesn't follow the Code; I too was on the Commission, and I try to follow the code. An interpretation of the Code was published by Eschmeyer (1990) for the 1985 edition.

His work is exceptional, but he does not always agree with current usage. But I still must check references through page 110 and check the remainder of his work. This will take several months.

Feb. 3, 2014

I am now up to page 140. A few more questions: Kottelat retains the original spelling with a capital letter for species when it occurs. We automatically change species names to begin with a lower case letter (as it is known by systematists). Kottelat does not show a lower case in his synonyms. This causes confusion. Also, when the spelling has diacritical markings, we correct the spelling in remarks, but he includes this in the original spelling -- this suggesting is automatic.

The spelling of Bănărescu is shown as Bănărescu in Eschmeyer but as Banarescu in Kottelat. Some spelling errors of scientific names were found.

Kottelat goes out of his way to cite unavailable names -- names which were never available, such as names in synonymies, names appearing without a description, published manuscript names. We include only a few of these, but we exclude most of them. Eventually these unavailable names will be placed in a separate database (type 3 names). These names can be used by later authors.

The RMNH museum has two systems, a "D" series, as in D1234 and a regular overlapping series, 1234; Kottelat removes the "D" yielding two identical series without a distinction (and I usually indicate if the specimen is dry). I was told that the comma in names such as ZMA 111,345 is only a comma, so I have deleted it to agree with other systems that include 4 or more numbers; it is given as 111345; but in the ZMA series the numbering does not start until 1000 (that comma can be reinserted easily).

Some corrections mentioned by Kottelet were corrected in on-line editions that appeared after he checked them.

Mar. 10, 2014

I am now up to page 282. I also went back over some pages up to 282 (disregard earlier pages to 282). There are mistakes in Kottelat (2013) that he could have solved. Some mistakes by Eschmeyer date to 2010 and even to the genera (1990); he could have marked these and taken care of them in one morning -- many were corrected in earlier editions of the online Catalog. This is what happens with a manuscript evolved over several years. I also find that he missed some synonyms that should have been in the text -- perhaps because he has overlooked them. Also, he does not give ranges for species.

Citing the Catalog of Fishes

Eschmeyer, W. N. (ed).  CATALOG OF FISHES: GENERA, SPECIES, REFERENCES. ( Electronic version accessed dd mmm 2014. [This version was edited by Bill Eschmeyer.]

Fricke, R. (ed).  REFERENCES. ( Electronic version accessed dd mmm 2014. [Ron Fricke devoted much time to improving references and journals, many of which had not been examined in 15 to 25 years. Please remember that the title of a paper often differs in three places: the title on the paper itself, the contents listed by the journal, and in the PDF provided by the journal. Ron tends to follow the title on the pdf for recent papers and the title on the original for earlier publications. Many of our dates of publication are documented for priority purposes. Some current journals have a target date, but actual publication is often delayed. For nomenclatural date problems, we will help if we can.]

van der Laan, R., Fricke, R. and W. N. Eschmeyer.  FAMILY-GROUP NAMES. ( Electronic version accessed dd mmm 2014.

Fricke, R. & Eschmeyer, W. N.  JOURNALS. ( Electronic version accessed dd mmm 2014. [Includes all journals appearing in the Catalog, including publication information and ISSN numbers.]

Fricke, R. & Eschmeyer, W. N.  GUIDE TO FISH COLLECTIONS. ( Electronic version accessed dd mmm 2014. [Arranged by museum abbreviation and by country, includes type catalogs and historical publications and www sites where available.]

Eschmeyer, W. N. & Fong, J. D.  SPECIES BY FAMILY/SUBFAMILY. ( Electronic version accessed dd mmm 2014. [Recalculated with each new version; based on current literature, this provides all available species names, valid species, and species described in the last 10 years by family/subfamily.]

Other participants are Dennis Polack (Data Analyst), Nicolas Bailly (Data Analyst), Richard van der Laan (Freshwater Fishes Specialist), Thomas Litz (Aquarium Species/Literature Specialist), and Dan Zimberlin (Data Analyst). Jon Fong prepares the online versions and does other programming tasks.

Please remember that if you got to the Catalog of Fishes from Fishbase, you are NOT in Fishbase. Fishbase is a more comprehensive database for a different audience, but they use Catalog of Fishes information in their authority file.

Some other useful sites for fishes are (1) Fishbase [Fishbase team, Manila]; (2) -- A comprehensive database on Shorefishes of the Tropical Eastern Pacific (and of the eastern Atlantic) [Ross Robertson]; (3) -- A database on elasmobranchs, with over 13,000 references, information on taxa, type specimens, etc. [Jürgen Pollerspöck]; (4) -- a comprehensive relational database of more than 99,350 scientific species name combinations and over 34,300 well-identified (mostly marine) fish pictures [Dennis Polack]; (5) Freshwater Fish List, 8th edition 2012 [via Google]; and (6) ETYFish Project (derivations of Fish Names) by Christopher Scharpf. A very useful site is [not associated with Fishbase or Worldfish]. Maintained by Michal Mikšik and Erwin Schraml. They post new fish taxa and references as they are published.


A summary paper was published in 2010 from the Catalog of Fishes database that might be of interest to users of the Catalog. "Marine fish diversity: history of knowledge and discovery (Pisces)." By William N. Eschmeyer, Ronald Fricke, Jon D. Fong & Dennis A. Polack. Zootaxa, no. 2525:19-50 (2 July 2010, open access, click title to download).

About 426 new species were described in 2013, and about 92 so far in 2014. As always, notifying us of missed original descriptions and errors in the database or providing PDFs of recent papers is most welcome.

Bill Eschmeyer edited type localities and standardized many place names. Our style is to use current place names found on an English map (or from Wikipedia and Google sites). Should you need original localities, they can be found in the original descriptions or from particular museum holdings (often online).

Many scientific names of fish species are based on personal names; some were found to be incorrect. For example, Jordan and co-workers spelled Japanese names ending in "a" with a terminal "e" as in matsubarae; that is acceptable, with the name Latinized first. So those names do not end in "i". Remarks are provided by Eschmeyer in this version for most species based on personal names. For a review of this very technical subject, see Dubois 2007: Zootaxa No. 1550:49-68.

New treatment of synonyms: Often an author revises a genus and treats a species in detail but does not mention the species synonyms (if any). If he/she does not mention the synonyms (rather common in recent treatments) and moves the valid “parent” species to a different genus, do we move the synonyms automatically, although the revising author may not have studied or assessed the synonyms? In the past, we did not move the synonyms. Typically, this often resulted in the synonyms being in a different current genus than the valid parent species. We now move the synonyms to the genus of the valid parent species, but no “status reference” showing the new generic placement is found with the synonym. Some synonyms will be moved incorrectly, but it is considered more desirable to have the potential synonyms in the same current genus as the valid parent species.

Some conventions used in the Catalog

Author plus date and a species code: The code of Zoological Nomenclature does not require a comma between the author and date, only a suggestion. In fact, we consider that confusing as that is traditionally used to show a cited reference in journal publications and not a species authorship and date. Genus+species+author+date (with parentheses as needed) is a (nearly) unique code or formula to define every species [2 known exceptions]. So you will find no comma between the author and date. Some journals are now treating species in this way.

Figures: Under a new species heading, some journals list (a) only the major figure, some list (b) all figures (including maps, habitats and graphs), and (c) some give figures showing the full view and any illustrated parts of the species. We give all figures that show a full view and any anatomical part of the species, so our listing of figures often differs from the treatment by a journal in the original description (as listed in the species heading).

Type localities: We give a current place name that one will find on a Google search or in an ‘English’ atlas. For some species described many years ago, one may find both the original locality and in parentheses a modern name for that locality. Many collections now have their holdings on-line, and it is there one can usually find the actual original locality if needed. Current localities are often augmented by additional information, such as coordinates.

Authors: We have added initials in brackets to distinguish all individual authors of genera and species.

Other sections

Species of Fishes by Family/Subfamily -- Total valid taxa reflect new taxa added minus taxa moved into synonymy plus taxa newly moved to valid. The classification is as used in the Catalog of Fishes. Prepared by Bill Eschmeyer and Jon Fong.

Family-Group Names -- Priority applies to Family-group names. It did not in early years. As classifications are refined with use of more subfamilies and tribes, it is required to use the oldest available family-group name for these new taxa. Early authors did not provide authorships and dates for families, and it is very difficult to find them. A third preliminary list of available family-group names of fishes is included in this version. Prepared by Richard van der Laan, Ron Fricke, and Bill Eschmeyer.

Ichthyological Collections -- Presented are those specimen collections mentioned in the main Catalog of Fishes database, especially as repositories of type specimens. Besides museum names and abbreviations, this includes WWW sites, cross-referencing of abbreviations, and selection of museums by country, along with type catalogs, historical articles, name changes, transfers, and other items for individual collections. Prepared by Ron Fricke and Bill Eschmeyer.

Ichthyological Journals -- There are nearly 2600 individual journals or monograph series in the Catalog of Fishes database. We provide full citations, and other information such as changes in titles, published duration, ISSN numbers, WWW sites, etc. Prepared by Ron Fricke and Bill Eschmeyer.


Thanks to Nicolas Bailly (Fishbase) and Dennis Pollack for continuing to find errors, discrepancies, and inconsistencies, to Thomas Litz for work with aquarium literature, and to Michal Mikšik who continues to find and forward publications. Thanks for help with this version to Christopher Scharpf. Jon Fong prepared the new version for www presentation.

Persons assisting in 2013: Arturo Acero, Abu Ahmed, Harald Ahnelt, K. V. Akhilesh, Gerald Allen, Frederico Almada, Daniel Almeida, Tilaian Alpermann, M. Eric Anderson, William Anderson, Jr., Arturo Angulo, D. V. Antonenko, Carlos Ardila, Jonathan Armbruster, Rachel J. Arnold, Gloria Arratia, Nicolas Bailly, Andrey Balanov, Carole Baldwin, Andred Balonic, Arcady Balushkin, Evgeny Barabanshchikov, Philippe Béarez, Ricardo Betancur-R., Murat Bilecenoglu, Roger Bills, Anthony J. Booth, John Briggs, Paulo Buckup, Kai Burkenhagen, Noel Burkhead, Rudolf Caers, Emily Capuli, Miguel Carneiro, Giorgia Carnevale, Kent Carpenter, Fernando Carvalho, Dave Catania, Albert Chakona, Bruce Collette, Kevin Conway, Rongfeng Cui, Regina Cunha, Neelesh Dahanukar, Paul L. Davis, Rafael Baron Diaz, Michael Doosey, Jean-Domonique Durand, Yu Dydyldin, Joseph Eastman, Dave Ebert, Mark Erdmann, Sergei Evseenko, Vicente Faria, Bo Fernholm, Carl Ferraris, Pierre Feutry, Jose de Lima Figueiredo, Lloyd Findley, Sergio Floeter, Matt Ford, Thomas Fraser, Katia Freire, John Friel, Katie Friere, Fulvio Garibaldi, Tom Geerinckx, Ali Gholamifard, Anthony Gill, Pier Giorgia, Dick Glodinski, Daniel Golani, Martin Gomon, Steve Grant, David Greenfield, Jessica Grosso, Lubomer Hanel, Akos Harka, Karsten Hartel, Phil Hastings, Malorie Hayes, Elaine Heemstra, Kai Heller, Dean Hendrickson, Eric Hilton, Hans Ho, Mysi Hoang, Doug Hoese, Wouter Holleman, Byron Hosking, Jean Huber, Jean-Claude Hureau, Markus Hussen, Tine Huyse, Samuel Iglésias, Leonardo Ingenito, Yukio Iwatsuki, Laith Jawad, Jean-Lou Justine, Yuichi Kano, Toshio Kawai, Heller Kei, Philippe Keith, Jin-Koo Kim, Seishi Kimura, Andrew Kinziger, Marcus Knight, Stas Kobyliansky, Keita Koeda, Steven Koerber, Alexander Kotlyar, Maurice Kottelat, Marcelo Kovacic, Bernd Kramer, Rudie Kuiter, Sven Kullander, Dinesh Kumar, Samuel Lalronunga, Daniel Lumbantobing, Francisco Langeani, Helen Larson, Peter Last, Robert Lea, Jeff Leis, Thomas Litz, Liu Jing, Liu Min, Marina Loeb, Yumnam Lokeshwor, Hernan Lopez-Fernandez, M. de L. Lozano Vilano, Carlos Alberto Santos de Lucena, Nathan Lujan, John Lundberg, Sherwyn Mack, James Maclaine, Luiz R. Malabarba, Javier Maldonado, Manoela Ferrera Marinho, Maniola Marinthe, Fernando Martins, Wilfredo Matamoros, Mizuki Matsunuma, George Mattox, Dishma Mayanglambam, Richard Mayden, Maricio De la Maza, John McCosker, Mark McGrouther, Catherine Mecklenburg, Canilo Melo, Naercio Menezes, Michael Mincarone, Alessandro Minelli, George Minos, Marcos Mirande, Susan Mochel, Anil Mohapatra, David Morgan, Jon Moore, Hiroyuki Motomura, Prachya Musikasinthorn, Brett Nagle, Rekha Nair, Tetsuji Nakabo, Douglas Nelson, André Netto-Ferreira, Dirk Neumann, Heok Hee Ng, Nguyen Huu Duc, Dalton Nielsen, Jørgen Nielsen, Dirk Nolf, Chiraher Nonpayon, Shinpei Ohashi, Makoto Okamoto, Alex Orlov, Guillermo Orti, Gerhard Ott, Larry Page, Mats Petersson, Ted Pietsch, Hudson Pinheiro, Diane Pitassy, Matthews Plamoottil, Gianluca Polgar, William Poly, Dominique Ponton, Artem Prokofiev, Graham Proudlove, Marti Pujolar, Joe Quattro, Olga Radchenko, Rajeev Raghavan, Mohammad Rahman, John Randall, Zachery Randall, David Raubenheimer, Tony Rees, Hector Reyes Bonilla, Adela Roa-Varon, Ross Robertson, Rob Robins, Ross Robinson, Luiz Rocha, Evgeny Romanov, Randall de Ruiter, Barry Russell, Mark Sabaj Perez, K. K. Satpathy, Christopher Scharpf, Ingo Schindler, Ulrich Schliewen, Erwin Schraml, Werner Schwarzhans, Bernard Séret, Ricardo Serrao Santos, Richard Sexton, Ilia Shakhovskoy, Bungdon Shangningam, Boris Sheiko, Koichi Shibukawa, Veronica Slobodian, Gabriel de Souza da Costa e Silva, David Smith, Gerald Smith, Bill Smith-Vaniz, Rainer Sonnenberg, Victor Springer, Hank Stam, Mathias Stehmann, Carl Struthers, John Sullivan, Lakpa Tamang, Donald Taphorn, James van Tassell, Christine Thacker, Armi Torres, Franz Uiblein, Josiane Vachon, Stefano Valdesalici, Richard van der Laan, J. A. Vanegas-Ríos, Henrique Varella, Katerina Vasil'eva, Diego Vaz, Molly Vergaese, Eric Verheyen, Frans Vermeulen, Leo Vernerus, Benjamin Victor, Ma de Lourdes Lazuno Vilano, Waikhom Vishwanath, Lei Wang, Simon Weigmann, David de Weirdt, Mark Westneat, Dan White, William White, Jeff Williams, Peter Wirtz, Tomoyuki Yamanaka, Claudio Henrique Zawadzki, Vladimir Zemnokhov, E. Zhang, Dan Zimberlin. Thanks to others we may have missed, including those mentioned elsewhere.

We thank the following for help with this edition:
K. Akhilesh, Gerry Allen, William Anderson, Jr., Nicolas Bailly, Vinicius Bertaco, Roger Bills, John Briggs, Paulo Buckup, Rudolf Caers, Jim Carlton, David Catania, Gustavo Chiarmonte, Kendel Clements, Kevin Conway, Andreas Dang, Andreas Dunz, Dave Ebert, Hector Espinosa, Herbert Federhen, Tom Fraser, Rui Freitas, Ron Fricke, Anthony Gill, Christofer Girel, Steve Grant, David Greenfield, Amanda Hay, Elaine Heemstra, Hans Ho, Mysi Hoang, Bryan Hosking, Laith Jawad, Max Kasparek, Seishi Kimura, Steen Knudsen, Bernd Kramer, Sebastien Lavaoé, Peter Last, Robert Lea, Flavio Lima, John Lundberg, James Maclaine, Ken Maeda, Fernanda Martins, John McCosker, Ernst Mikschi, Michal Miksik, Michael Mincarone, Rarina Mondonca, Rebecca Morin, Hiroyuki Motomura, Thomas Munroe, Prachya Musikasinthorn, Dirk Neumann, Maria Palomares, Ron Peters, Stuart Poss, Jurgen Pollerspock, William Poly, Heather Prestridge, Sally Reader, R. R. dos Reis, Bill Richards, Tyson Roberts, Luiz Rocha, Ronald de Ruiter, Barry Russell, Pasakorn Saenjeun, Keiichi Sakai, Ingo Schindler, Erwin Schraml, Janet Scott, Randall Singer, Bill Smith-Vaniz, Hank Stam, David Stein, Carl Struthers, Matthias Svojtka, Stefanio Valdeslici, James van Tassell, Armi Torres, Peter Unmack, Franz Uiblein, Stefanio Valdesalici, Richard Vari, Leo Venerus, Ben Victor, Steven Walsh, Simon Weighmann, William White, Erica Wilber, Rick Winterbottom, Peter Wirtz, Axel Zarske. Thanks to others whom we may have missed.

We are grateful for assistance in catching errors to Dan Zimberlin, Nicolas Bailly and the staff of Fishbase, and especially Christopher Scharpf.

Bill Eschmeyer
California Academy of Sciences

Search Instructions:

A search expression can be specified as

  • A single search term
    • a word
    • a phrase (multiple words) enclosed in double quotes – ["Jordan & Evermann"]
    • a partial word or phrase enclosed in double quotes (") and ending with an asterisk (*) – ["Hawai*"]
  • Two (or more) search terms joined by one of three logical operators: { and | or | and not }
    (terms can be grouped by using parentheses - see examples below).

Some examples of search expressions and the number of hits when that query is issued against the indicated database.

Example of
Search Expression
single word
"valid as Albula vulpes"
"synonym of Albula vulpes"
"Comoro Is"
x and y
Cuvier and Valenciennes
x or y
Cuvier or Valenciennes
x and not y
Salmo and not Salmonidae
Salmo and not Salmonidae and not Osmeridae
x and not (y or z)
Salmo and not (Salmonidae or Osmeridae)
wild card (*)
"Boo*" and not book

Technical feedback: