Institute for Biodiversity Science and Sustainability
Online Version, Updated 1 July 2019

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Eschmeyer's Catalog of Fishes
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Squalius (subgenus of Leuciscus) Bonaparte [C. L.] 1837:fasc. 19 Masc. Leuciscus squalus Bonaparte 1837. Type by subsequent designation. Occurred in puntata 96, fasc. 19 under Leuciscus squalus and two additional species; also in later fascicles. Type evidently designated first by Jordan 1919:187 [ref. 2410]. •Valid as Squalius Bonaparte 1837 -- (Sanjur et al. 2003:20 [ref. 28742], Bogutskaya & Naseka 2004:88 [ref. 28183], Doadrio & Carmona 2006:199 [ref. 28975], Doadrio et al. 2007:89 [ref. 29250], Doadrio et al. 2007:247 [ref. 29342], Kottelat & Freyhof 2007:154, 257 [ref. 29996], Turan et al. 2009:53 [ref. 30540], Bogutskaya & Zupančič 2010:53 [ref. 30905], Coad 2010:193 [ref. 36620], Perea et al. 2010:3, 14 [ref. 31802], Zupančič et al. 2010:155 [ref. 31002], Özuluğ & Freyhof 2011:107 [ref. 31360], Shinohara et al. 2011:38 [ref. 31715], Parin et al. 2014:94 [ref. 33547], Jouladeh-Roudbar et al. 2015:880 [ref. 34062], Khaefi et al. 2016:145 [ref. 34748], Esmaeili et al. 2016:118 [ref. 34985], Turan et al. 2017:1 [ref. 35136], Esmaeili et al. 2017:57 [ref. 35254], Çiçek et al. 2018:14 [ref. 36267], Esmaeili et al. 2018:38 [ref. 36089]). Current status: Valid as Squalius Bonaparte 1837. Leuciscidae: Leuciscinae.