Institute for Biodiversity Science and Sustainability
Online Version, Updated 9 Dec 2025

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Eschmeyer's Catalog of Fishes
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Papyrocephalus (subgenus of Hymenocephalus) Gilbert [C. H.] & Hubbs [C. L.] 1920:539 Masc. Hymenocephalus aterrimus Gilbert 1905. Type by original designation. •Synonym of Hymenocephalus Giglioli 1884, but a valid subgenus as described -- (Iwamoto et al. 2011:515 [ref. 31434], Schwarzhans 2014:26 [ref. 33586], Nakayama 2020:207 [ref. 37888]). Current status: Synonym of Hymenocephalus Giglioli 1884. Macrouridae.


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