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Eschmeyer's Catalog of Fishes
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Species in the genus Akysis: [ 26 ] records


bilustris, Akysis Ng [H. H.] 2011:62, Fig. 1 [Zootaxa No. 3066; ref. 31572] Xe Kong where Houai Tamopat enters it, downstream of Ban Khanmaknao, 14°36'31"N, 106°33' 08"E, Attapeu Province, Laos. Holotype: ZRC 53111. Paratypes: CMK 15692 (1), 21247 (1), 21411 (2), 21457 (1); ZRC 53112 (2). •Valid as Akysis bilustris Ng 2011 -- (Kottelat 2013:218 [ref. 32989], Taki et al. 2021:299 [ref. 39830]). Current status: Valid as Akysis bilustris Ng 2011. Akysidae: Akysinae. Distribution: Southeast Asia: Mekong River basin, southern Laos and Cambodia. Habitat: freshwater.

brachybarbatus, Akysis Chen [Y.-R.] in He & Chen 1981:210 (English p. 213), Fig. 2 [Zoological Research v. 2 (no. 3); ref. 21313] Nanrui River, Mekong River system, Menlian County, China. Holotype: KIZ 787162. Paratypes: KIZ (26). Type catalog: Ferraris 2007:13 [ref. 29155], Du et al. 2013:354 [ref. 32939] with author as He. •Valid as Akysis brachybarbatus Chen 1981 -- (Zhu 1995:158 [ref. 25213], Ng & Kottelat 1996:19 [ref. 22212], Ng 1996:3 [ref. 22484], Ng & Kottelat 1998:1059 [ref. 23903], Yue & Chen 1998:224 [ref. 26415], Ng & Tan 1999:362 [ref. 24159], Chen in Chu et al. 1999:112 [ref. 24534], Ng & Kottelat 2000:9 [ref. 25042], Ng & Freyhof 2003:315 [ref. 27399], Ng & Rainboth 2005:33 [ref. 28353], Ng 2006:20 [ref. 28586], Page et al. 2007:292 [ref. 29140], Ferraris 2007:13 [ref. 29155], Ng 2007:53 [ref. 29306], Ng 2008:260 [ref. 29979], Ng 2009:7 [ref. 30104], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989], Zhang et al. 2016:170 [ref. 34477], Fu et al. 2021:9572 [ref. 39347], Ng et al. 2022:7 [ref. 39344]). Current status: Valid as Akysis brachybarbatus Chen 1981. Akysidae: Akysinae. Distribution: Southeast Asia: Lancangjiang [Mekong] River, Yunnan (China). IUCN (2023): Vulnerable. Habitat: freshwater.

clavulus, Akysis Ng [H. H.] & Freyhof [J.] 2003:312, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 14 (no. 4); ref. 27399] Song Duc My about 10 kilometers west of Ninh Hoa, 12°32.22'N, 109°01.38'E, Khanh Hoa Province, Vietnam. Holotype: ZFMK 27159. Paratypes: UMMZ 238018 (1); ZFMK 23265 (1), 25506-16 (11), 27156-58 (3), 27160-65 (6), 27506-09 (4); ZRC 46177 (3). Type catalog: Ferraris 2007:14 [ref. 29155], Ng & Kim 2008:5 [ref. 29920], Herder et al. 2010:119 [ref. 31087]. •Valid as Akysis clavulus Ng & Freyhof 2003 -- (Ng & Rainboth 2005:33 [ref. 28353], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989], Ng et al. 2022:7 [ref. 39344]). Current status: Valid as Akysis clavulus Ng & Freyhof 2003. Akysidae: Akysinae. Distribution: Cai and Duc My rivers, draining to the central coast of Vietnam. Habitat: freshwater.

clinatus, Akysis Ng [H. H.] & Rainboth [W. J.] 2005:34, Figs. 1, 3a [Raffles Bulletin of Zoology Suppl. no. 13; ref. 28353] Stung Chhay River, Tok Kong village on road from Phnom Penh to Sihanoukville, 10°56'37.8"N, 113°47'47.4"E, Cambodia. Holotype: ZRC 47288. Paratypes: ZRC 46159 (3). Type catalog: Ferraris 2007:14 [ref. 29155], Ng & Kim 2008:5 [ref. 29920]. •Valid as Akysis clinatus Ng & Rainboth 2005 -- (Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2009:48 [ref. 30100], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989], Ng et al. 2022:7 [ref. 39344]). Current status: Valid as Akysis clinatus Ng & Rainboth 2005. Akysidae: Akysinae. Distribution: Cambodia. Habitat: freshwater.

ephippifer, Akysis Ng [H. H.] & Kottelat [M.] 1998:1078, Figs. 4h, 11, 13f [Journal of Natural History v. 32 (no. 7); ref. 23903] Stung Santréa, 40 kilometers from Pomnac, Cambodia. Holotype: MNHN 1977-0034. Paratypes: MNHN 1997-0035 to 0038 (5, 6, 5, 3); CMK 4296 (1, c&s), 11936 (6), 11938 (3); ZRC 40713 (6). Type catalog: Ferraris 2007:14 [ref. 29155], Ng & Kim 2008:7 [ref. 29920]. •Valid as Akysis ephippifer Ng & Kottelat 1998 -- (Ng & Tan 1999:362 [ref. 24159], Kottelat 2001:138 [ref. 25780] = bilustris, Ng & Freyhof 2003:315 [ref. 27399], Ng & Rainboth 2005:33 [ref. 28353], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989], Taki et al. 2021:299 [ref. 39830], Ng et al. 2022:7 [ref. 39344]). Current status: Valid as Akysis ephippifer Ng & Kottelat 1998. Akysidae: Akysinae. Distribution: Southeast Asia: lower Mekong River basin, Cambodia. Habitat: freshwater.

fontaneus, Akysis Ng [H. H.] 2009:42, Figs. 1-2, 5a, 6a [Zootaxa No. 1973; ref. 30100] Sungai Jernih at Desa Jerih, ca. 25 killometers on road to Sarolangun after turnoff at Paihm 2°00.215'S, 102°42.475'E, Jambi, Sumatra. Holotype (unique): MZB 10989. •Valid as Akysis fontaneus Ng 2009 -- (Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989]). Current status: Valid as Akysis fontaneus Ng 2009. Akysidae: Akysinae. Distribution: Central Sumatra. Habitat: freshwater.

fuliginatus, Akysis Ng [H. H.] & Rainboth [W. J.] 2005:35, Fig. 4 [Raffles Bulletin of Zoology Suppl. no. 13; ref. 28353] Stung Treng Province, Mekong River on western edge of Kaoh Han, 16 kilometers northeast of Stung Treng, 13°38'N, 106°03'E, Cambodia. Holotype: UMMZ 241338. Type catalog: Ferraris 2007:14 [ref. 29155]. •Valid as Akysis fuliginatus Ng & Rainboth 2005 -- (Ng 2006:20 [ref. 28586], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2007:53 [ref. 29306], Ng 2009:7 [ref. 30104], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis fuliginatus Ng & Rainboth 2005. Akysidae: Akysinae. Distribution: Lower Mekong River basin, Cambodia. Habitat: freshwater.

galeatus, Akysis Page [L. M.], Rachmatika [I.] & Robins [R. H.] in Page, Hadiaty, López, Rachmatika & Robins 2007:299, Figs. 3 (left column), 6 [Copeia 2007 (no. 2); ref. 29140] Way Gedongwani (Way Sekampung drainage), 5°14.499'S, 105°29.060'E, Lampung, Sumatra, Indonesia. Holotype: MZB 15192. Paratypes: ANSP 182914 (1), INHS 100121 (1), MZB 15193 (2), UF 162466 (2). •Valid as Akysis galeatus Page, Rachmatika & Robins 2007 -- (Ng 2008:260 [ref. 29979], Ng 2009:46 [ref. 30100], Ng 2011:66 [ref. 31572], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis galeatus Page, Rachmatika & Robins 2007. Akysidae: Akysinae. Distribution: Lampung Province, Sumatra, Indonesia. Habitat: freshwater.

hendricksoni, Akysis Alfred [E. R.] 1966:467, Fig. 1 [Copeia 1966 (no. 3); ref. 8002] Trengganu River at Kuala Brang, Trengganu, Malaysia. Holotype: ZRC 1414. Paratypes: BMNH 1966.2.11.1 (1), ZRC 1416 (1). Type catalog: Alfred 1970:69 [ref. 23931], Ferraris 2007:14 [ref. 29155], Ng & Kim 2008:8 [ref. 29920]. •Valid as Akysis hendricksoni Alfred 1966 -- (He & Chen 1981:212 [ref. 21313], Burgess 1989:118 [ref. 12860], Ng & Kottelat 1996:19 [ref. 22212], Ng 1996:3 [ref. 22484], Ng & Kottelat 1998:1084 [ref. 23903], Ng & Tan 1999:362 [ref. 24159], Ng 1999:547 [ref. 24609], Ng & Freyhof 2003:315 [ref. 27399], Ng & Kottelat 2004:199 [ref. 27854], Ng & Rainboth 2005:33 [ref. 28353], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989], Ng et al. 2022:8 [ref. 39344], Ahmad et al. 2023:105 [ref. 41243]). Current status: Valid as Akysis hendricksoni Alfred 1966. Akysidae: Akysinae. Distribution: Southeast Asia: Malaya (Malaysia) and Isthmus of Kra (southern Thailand). IUCN (2019): Least Concern. Habitat: freshwater.

heterurus, Akysis Ng [H. H.] 1996:4, Figs. 1-2 [Raffles Bulletin of Zoology v. 44 (no. 1); ref. 22484] Sg. Alai at 19.5 kilometers Muara Bung-Muara Tebo road, Jambi, Sumatra, Indonesia. Holotype: ZRC 34194. Paratypes: CMK 12199 (1); MZB 6209 (1); ZRC 39248 (7). Type catalog: Ferraris 2007:14 [ref. 29155], Ng & Kim 2008:9 [ref. 29920]. •Valid as Akysis heterurus Ng 1996 -- (Burgess & Finley 1996:165 [ref. 22901], Ng & Kottelat 1998:1059 [ref. 23903], Ng & Tan 1999:362 [ref. 24159], Ng 1999:547 [ref. 24609], Tan & Ng 1999:289 [ref. 24254], Ng & Freyhof 2003:315 [ref. 27399], Ng & Kottelat 2004:199 [ref. 27854], Ng & Rainboth 2005:33 [ref. 28353], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2009:46 [ref. 30100], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis heterurus Ng 1996. Akysidae: Akysinae. Distribution: Sumatra, Indonesia. Habitat: freshwater.

longifilis, Akysis Ng [H. H.] 2006:20, Fig. 1 [Zootaxa No. 1150; ref. 28586] Bago division Pyu township, Pyu stream, tributary of Sittang River, about 229 kilometers from Yangon, 18°29'N, 96°26'E, Myanmar. Holotype: UMMZ 246172. Paratypes: UF 161587 (2); UMMZ 245966 (7). Plus non-type material. •Valid as Akysis longifilis Ng 2006 -- (Page et al. 2007:292 [ref. 29140], Ng 2007:53 [ref. 29306], Ng 2008:260 [ref. 29979], Ng 2009:6 [ref. 30104], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis longifilis Ng 2006. Akysidae: Akysinae. Distribution: Sittang River drainage, southern Myanmar. Habitat: freshwater.

maculipinnis, Akysis Fowler [H. W.] 1934:97, Fig. 46 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 86; ref. 1417] Chantaboon [Chantaburi], southeastern Thailand. Holotype: ANSP 59366. Paratypes: ANSP 59367-72 (6). Type catalog: Böhlke 1984:18 [ref. 13621], Ferraris 2007:14 [ref. 29155] •Valid as Akysis maculipinnis Fowler 1934 -- (He & Chen 1981:212 [ref. 21313], Burgess 1989:118 [ref. 12860], Ng & Kottelat 1996:19 [ref. 22212], Ng 1996:3 [ref. 22484], Rainboth 1996:159 [ref. 22772], Ng & Kottelat 1998:1088 [ref. 23903], Ng & Tan 1999:362 [ref. 24159], Ng & Rainboth 2005:33 [ref. 28353], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis maculipinnis Fowler 1934. Akysidae: Akysinae. Distribution: Southeastern Thailand and southwestern Cambodia. Habitat: freshwater.

manipurensis, Laguvia Arunkumar [L.] 2000:194, Fig. 1 [Indian Journal of Fisheries v. 47 (no. 3); ref. 25658] Lairaok Maru stream near Moreh, 110 kilometers from Imphal City, Yu River System, Manipur, India. Holotype: MUMF 3001/1A. Paratypes: MUMF 3001/4A (4). Type catalog: Ferraris 2007:382 [ref. 29155]. •Valid, but generic placement insertae sedis -- (Ng & Kottelat 2005:20 [ref. 28207], Thomson & Page 2006:23 [ref. 28859]). •Valid as Akysis manipurensis (Arunkumar 2000) -- (Jayaram 2006:161 [ref. 28762], Vishwanath et al. 2007:2675 [ref. 29124], Ng 2008:261 [ref. 29979], Ng 2009:6 [ref. 30104], Ng 2011:66 [ref. 31572], Kottelat 2013:218 [ref. 32989]). •Incertae sedis in Sisoridae -- (Ferraris 2007:382 [ref. 29155]). Current status: Valid as Akysis manipurensis (Arunkumar 2000). Akysidae: Akysinae. Distribution: Manipur, India. Habitat: freshwater.

microps, Akysis Ng [H. H.] & Tan [H. H.] 1999:359, Figs. 5-7 [Zoological Studies v. 38 (no. 3); ref. 24159] Sungai Kahang and tributary, about 44.4 kilometers from Mersing turnoff to Kluang just before side road to Endau Rompin Taman Negara, kilometer 96 from Mersing to Batu Pahat (2°3.93'N, 103°31.58'E), Johor, Malaysia. Holotype: ZRC 42596. Paratypes: BMNH 1998.7.19.1-2 (2); CAS 99371 (1); CMK 14803 (2); MZUSP 53470 (4, 1 c&s); ZRC 42597 (6), 42599 (5). Type catalog: Ferraris 2007:14 [ref. 29155], Ng & Kim 2008:11 [ref. 29920]. •Valid as Akysis microps Ng & Tan 1999 -- (Ng & Freyhof 2003:315 [ref. 27399], Ng & Rainboth 2005:33 [ref. 28353], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis microps Ng & Tan 1999. Akysidae: Akysinae. Distribution: Malay Peninsula. Habitat: freshwater.

patrator, Akysis Ng [H. H.], Pawangkhanant [P.] & Suwannapoom [C.] 2022:3, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 31 (no. 3): IEF-1186; ref. 39344] Huay Saung Ga Ria, Khwae Noi subdrainage, Mae Klong drainage, Kanchanaburi Province, Thailand, 15°13'42"N, 98°26'42"E. Holotype: AUP 02025. Paratypes: AUP 02026-02030 (5). •Valid as Akysis patrator Ng, Pawangkhanant & Suwannapoom 2022. Current status: Valid as Akysis patrator Ng, Pawangkhanant & Suwannapoom 2022. Akysidae: Akysinae. Distribution: Southeast Asia: Mae Klong drainage, Kanchanaburi Province, western Thailand. Habitat: freshwater.

pictus, Akysis Günther [A.] 1883:138 [Annals and Magazine of Natural History (Series 5) v. 11 (no. 62); ref. 14339] Tenasserim, Myanmar. Syntypes: BMNH 1880.12.1.25-26 (2). Type catalog: Ferraris 2007:14 [ref. 29155] •Valid as Akysis pictus Günther 1883 -- (He & Chen 1981:211 [ref. 21313], Burgess 1989:118 [ref. 12860], Talwar & Jhingran 1991:617 [ref. 20764], Ng & Kottelat 1996:19 [ref. 22212], Ng 1996:3 [ref. 22484], Ng & Kottelat 1998:1059 [ref. 23903], Ng & Tan 1999:362 [ref. 24159], Ng 1999:542 [ref. 24609], Ng & Kottelat 2000:9 [ref. 25042], Ng & Freyhof 2003:315 [ref. 27399], Ng & Kottelat 2004:194 [ref. 27854], Ng & Rainboth 2005:33 [ref. 28353], Ng 2006:20 [ref. 28586], Jayaram 2006:162 [ref. 28762], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2007:53 [ref. 29306], Ng 2008:260 [ref. 29979], Ng 2009:6 [ref. 30104], Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis pictus Günther 1883. Akysidae: Akysinae. Distribution: Myanmar. Habitat: freshwater.

portellus, Akysis Ng [H. H.] 2009:3, Figs. 1-5 [Journal of Threatened Taxa v. 1 (no. 1); ref. 30104] Shwe Kyinm 17°55'N, 96°53'E, Than Kyaw Toe, Bago division, Myanmar. Holotype: ZRC 51138. Paratypes: NRM 57228 (5), UMMZ 248469 (12), ZRC 51139 (9). Plus non-type material. •Valid as Akysis portellus Ng 2009 -- (Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis portellus Ng 2009. Akysidae: Akysinae. Distribution: Sittang River drainage, southern Myanmar. Habitat: freshwater.

prashadi, Akysis Hora [S. L.] 1936:200, Figs. 1, 2(b-c) [Records of the Indian Museum (Calcutta) v. 38 (pt 2); ref. 13703] Indawgyi Lake and round about Kamaing in the Myitkyina District, upper Myanmar. Holotype (unique): ZSI F10873/1. Type catalog: Ferraris 2007:14 [ref. 29155] See Das 2003:378 [ref. 27706]. •Valid as Akysis prashadi Hora 1936 -- (He & Chen 1981:211 [ref. 21313], Burgess 1989:118 [ref. 12860], Talwar & Jhingran 1991:617 [ref. 20764], Ng & Kottelat 1996:19 [ref. 22212], Ng 1996:3 [ref. 22484], Ng & Kottelat 1998:1059 [ref. 23903], Ng & Tan 1999:362 [ref. 24159], Ng 1999:545 [ref. 24609], Ng & Kottelat 2000:9 [ref. 25042], Ng & Freyhof 2003:315 [ref. 27399], Ng & Kottelat 2004:194 [ref. 27854], Ng & Rainboth 2005:33 [ref. 28353], Ng 2006:20 [ref. 28586], Jayaram 2006:163 [ref. 28762], Vishwanath et al. 2007:2676 [ref. 29124], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2007:53 [ref. 29306], Ng 2008:260 [ref. 29979], Ng 2009:6 [ref. 30104], Ng 2011:66 [ref. 31572], Ahmed et al. 2013:286 [ref. 32872], Kottelat 2013:219 [ref. 32989], Ng et al. 2022:8 [ref. 39344], Chen et al. 2024:115 [ref. 41792]). Current status: Valid as Akysis prashadi Hora 1936. Akysidae: Akysinae. Distribution: Southeast Asia: Manipur, India, Bangladesh and Irrawaddy River basin, Myanmar. Habitat: freshwater.

pulvinatus, Akysis Ng [H. H.] 2007:52, Figs. 1, 2, 5a [Zootaxa No. 1608; ref. 29306] Stream draining into Andaman Sea upstream of Kapoe, 9°34'14.0"N, 98°41'40.4"E, Ranong Province, Thailand. Holotype: UMMZ 248249. Paratypes: UMMZ 245696 (1), 248250 (2); ZRC 51009 (3). Type catalog: Ng & Kim 2008:12 [ref. 29920]. •Valid as Akysis pulvinatus Ng 2007 -- (Ng 2008:261 [ref. 29979], Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis pulvinatus Ng 2007. Akysidae: Akysinae. Distribution: Kra Isthmus, upper Tapi River drainage and hillstreams flowing into the Andaman Sea, southern Thailand. Habitat: freshwater.

recavus, Akysis Ng [H. H.] & Kottelat [M.] 1998:1090, Figs. 4l, 13i, 16, 17a [Journal of Natural History v. 32 (no. 7); ref. 23903] Western Chao Phraya basin, Thailand. Holotype: ZRC 40716. Paratypes: CMK 10740 (3), 11020 (1); UMMZ 233005 (2); ZRC 40717 (2). Type catalog: Ferraris 2007:14 [ref. 29155], Ng & Kim 2008:12 [ref. 29920]. •Valid as Akysis recavus Ng & Kottelat 1998 -- (Ng & Tan 1999:362 [ref. 24159], Ng 1999:547 [ref. 24609], Ng & Freyhof 2003:315 [ref. 27399], Ng & Rainboth 2005:33 [ref. 28353], Page et al. 2007:292 [ref. 29140], Ferraris 2007:14 [ref. 29155], Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis recavus Ng & Kottelat 1998. Akysidae: Akysinae. Distribution: Chao Phraya and Mae Klong drainages, Thailand. Habitat: freshwater.

scorteus, Akysis Page [L. M.], Hadiaty [R. K.] & López [J. A.] in Page, Hadiaty, López, Rachmatika & Robins 2007:296, Figs. 2, 3 (right column) [Copeia 2007 (no. 2); ref. 29140] Way Seputih, 5°04.310'S, 104°53.048'E, Lampung, Sumatra, Indonesia. Holotype: MZB 15191. Paratypes: UF 162465 (1). •Valid as Akysis scorteus Page, Hadiaty & López 2007 -- (Ng 2008:261 [ref. 29979], Ng 2009:46 [ref. 30100], Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis scorteus Page, Hadiaty & López 2007. Akysidae: Akysinae. Distribution: Lampung, Sumatra. Habitat: freshwater.

variegatus, Akysis variegatus Prashad [B.] & Mukerji [D. D.] 1929:180, Pl. 8 (figs. 1-2) [Records of the Indian Museum (Calcutta) v. 31 (pt 3); ref. 3558] South end of Indawgyi Lake, along southwestern shore, Myitkyina District, Upper Myanmar. Holotype (unique): ZSI F10873/1. Type catalog: Menon & Yazdani 1968:126 [ref. 20743], Ferraris 2007:14 [ref. 29155] Subjectively invalid; preoccupied in Akysis by Pimelodus variegatus Bleeker 1846; the authors even state that this new subspecies is closely allied both to Akysis variegatus Bleeker and Akysis pictus Günther. See Das 2003:378 [ref. 27706] with date as 1930. •Synonym of Akysis prashadi Hora 1936 -- (Ng 1999:545 [ref. 24609], Ng & Freyhof 2003:315 [ref. 27399], Jayaram 2006:163 [ref. 28762], Vishwanath et al. 2007:2676 [ref. 29124], Ferraris 2007:14 [ref. 29155], Kottelat 2013:219 [ref. 32989]). 2°Homonym. Current status: Synonym of Akysis prashadi Hora 1936. Akysidae: Akysinae. Habitat: freshwater.

variegatus, Pimelodus Bleeker [P.] 1846:53 [177] [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 21 (art. 5); ref. 313] Jakarta, Java, Indonesia. Syntypes: BMNH 1863.12.4.187 (1) Java, ?NMV 46004 (1), RMNH 6881 (16). Type catalog: Ferraris 2007:15 [ref. 29155]. Also in Bleeker 1846:53 [ref. 313]. See van Oijen et al. 2009:205 [ref. 30423]. •Valid as Akysis variegatus (Bleeker 1846) -- (He & Chen 1981:212 [ref. 21313], Kottelat et al. 1993:75 [ref. 23448], Roberts 1993:31 [ref. 21799], Ng & Kottelat 1996:19 [ref. 22212], Ng 1996:3 [ref. 22484], Rainboth 1996:159 [ref. 22772], Ng & Kottelat 1998:1059 [ref. 23903], Ng & Tan 1999:362 [ref. 24159], Ng 1999:547 [ref. 24609], Ng & Kottelat 2000:9 [ref. 25042], Ng & Kottelat 2004:194 [ref. 27854], Ng & Rainboth 2005:33 [ref. 28353], Ng 2006:20 [ref. 28586], Page et al. 2007:292 [ref. 29140], Ferraris 2007:15 [ref. 29155], Ng 2008:261 [ref. 29979], Ng 2009:48 [ref. 30100], Ng 2009:7 [ref. 30104], Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Hasan et al. 2022:6035 [ref. 39888], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis variegatus (Bleeker 1846). Akysidae: Akysinae. Distribution: Southeast Asia: western Java (Indonesia). Habitat: freshwater.

varius, Akysis Ng [H. H.] & Kottelat [M.] 1998:1093, Figs. 4m, 13k, 17b, 18 [Journal of Natural History v. 32 (no. 7); ref. 23903] Xe Bangfai, about 3 kilometers upriver of Pan Pakphanang, Khammouan Province, Laos. Holotype: ZRC 41015. Paratypes: CMK 12433 (6), 12609 (1); UMMZ 214913 (2); USNM 232930 (7). Type catalog: Ferraris 2007:15 [ref. 29155], Ng & Kim 2008:14 [ref. 29920]. •Valid as Akysis varius Ng & Kottelat 1998 -- (Ng 1999:547 [ref. 24609], Ng & Kottelat 2000:9 [ref. 25042], Kottelat 2001:140 [ref. 25780], Ng & Freyhof 2003:315 [ref. 27399], Ng & Kottelat 2004:194 [ref. 27854], Ng & Rainboth 2005:33 [ref. 28353], Ng 2006:20 [ref. 28586], Ferraris 2007:15 [ref. 29155], Ng 2008:261 [ref. 29979], Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Kottelat 2015:47 [ref. 33971], Taki et al. 2021:298 [ref. 39830], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis varius Ng & Kottelat 1998. Akysidae: Akysinae. Distribution: Southeast Asia: Mekong River basin, Cambodia, Laos and Thailand. Habitat: freshwater.

vespa, Akysis Ng [H. H.] & Kottelat [M.] 2004:194, Figs. 1-3, 5 [Ichthyological Exploration of Freshwaters v. 15 (no. 3); ref. 27854] Stream "Chon Son" between Kyondaw and Phadaw, about 20 kilometers northwest of Payathouzu (at border with Thailand), 15°25'N, 98°15'E, Kayin State, Myanmar. Holotype: ZRC 46423. Paratypes: CMK 17788 (8), 17953 (5), 17977 (5); ZRC 49155 (4). Type catalog: Ferraris 2007:15 [ref. 29155], Ng & Kim 2008:15 [ref. 29920]. •Valid as Akysis vespa Ng & Kottelat 2004 -- (Ng & Rainboth 2005:33 [ref. 28353], Ng 2006:20 [ref. 28586], Jayaram 2006:164 [ref. 28762], Page et al. 2007:292 [ref. 29140], Ferraris 2007:15 [ref. 29155], Ng 2007:53 [ref. 29306], Ng 2008:261 [ref. 29979], Ng 2009:6 [ref. 30104], Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis vespa Ng & Kottelat 2004. Akysidae: Akysinae. Distribution: Ataran River drainage, Kayin State, Myanmar. Habitat: freshwater.

vespertinus, Akysis Ng [H. H.] 2008:256, Fig. 1, 4a, 5a [Ichthyological Exploration of Freshwaters v. 19 (no. 3); ref. 29979] Headwaters of Ann Chaung drainage, ca. 19 kilometers east by south of Ann, 19°43'N, 94°11'E, Rakhine state, Myanmar. Holotype: UMMZ 248755. Paratypes: CMK 20752 (2); UMMZ 248756 (4), 248757 (2); ZRC 51345 (2). •Valid as Akysis vespertinus Ng 2008 -- (Ng 2009:6 [ref. 30104], Ng 2011:66 [ref. 31572], Kottelat 2013:219 [ref. 32989], Ng et al. 2022:8 [ref. 39344]). Current status: Valid as Akysis vespertinus Ng 2008. Akysidae: Akysinae. Distribution: Ann Chaung drainage, Rakhine, western Myanmar. Habitat: freshwater.


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