Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the genus Benthophilus: [ 28 ] records

abdurahmanovi, Benthophilus magistri Ragimov [D. B.] 1978:793 [703] [Voprosy Ikhtiologii v. 18 (no. 5); ref. 8998] East coast of Tyuleniy Island, northern Caspian Sea, depth 2.7 meters. Holotype: ZISP [= ZIN] 53657 [ex IZAB 132]. Paratypes: (orig. 99) ZISP 53658 [ex IZAB 132] (15). On p. 703 of English translation. •Synonym of Benthophilus magistri Iljin 1927, but a valid subspecies as described -- (Coad 1995:30 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:129 [ref. 23591]). •Synonym of Benthophilus magistri Iljin 1927 -- (Bogutskaya et al. 2001:47 [ref. 26178]). •Valid as Benthophilus abdurahmanovi Ragimov 1978 -- (Miller & Vasil'eva in Miller 2004:169 [ref. 27853], Bogutskaya & Naseka 2004:215 [ref. 28183], Boldyrev & Bogutskaya 2004:132 [ref. 28099], Boldyrev & Bogutskaya 2007:56 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:376 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:79 [ref. 35254], Esmaeili et al. 2018:60 [ref. 36089], Parenti 2021:69 [ref. 38603], Vasil'eva et al. 2025:[8] [ref. 42434]). Current status: Valid as Benthophilus abdurahmanovi Ragimov 1978. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin (Russia, Dagestan). Habitat: freshwater, brackish.

aculeatus, Benthophilus Baer [K. E.] in Lukina 1984:168 [Caspian expedition of K. M. Baer in 1853-1857; ref. 29151] Bight at Novopetrovsk, Mangyshlak, Caspian Sea, Kazakhstan. No types known. Original not seen; see Boldyrev & Bogutskaya 2007:56 [ref. 29150]. •Synonym of Benthophilus leobergius Berg 1949 -- (Boldyrev & Bogutskaya 2007:60 [ref. 29150] with question, Neilson & Stepien 2009:96 [ref. 30487], Jouladeh-Roudbar et al. 2020:227 [ref. 37983], Parenti 2021:70 [ref. 38603], Çiçek et al. 2024:301 [ref. 40789]). Current status: Synonym of Benthophilus leobergius Berg 1949. Gobiidae: Gobiinae. Habitat: freshwater, brackish.

baeri, Benthophilus Kessler [K. F.] 1877:52, Pl. 4 (fig. 10) [The Aralo-Caspian Expedition; ref. 2597] Mangyshlak Penninsula, Kazakhstan; southern Caspian Sea, depth 49-266 feet [7-38 Russian fathoms]. Lectotype: ZISP 2239. Paralectotypes: (11) BMNH 1897.7.5.16 [ex St. Petersberg Univ.] (1), SPU 395 [463/410] (1), ZIN 53665 [ex 2239] (4). Lectotype selected y Boldyrev & Bogutskaya 2007:82 [ref. 29150]. •Valid as Benthophilus baeri Kessler 1877 -- (Berg 1949:1118 [ref. 20662], Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:752 [ref. 24702], Vasil'eva 1998:127 [ref. 23591], Coad 1998:103 [ref. 23869], Vasil'eva 2000:764 [ref. 25083], Pinchuk et al. in Miller 2004:175 [ref. 27853], Bogutskaya & Naseka 2004:215 [ref. 28183], Boldyrev & Bogutskaya 2007:81 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:376 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:79 [ref. 35254], Esmaeili et al. 2018:60 [ref. 36089], Jouladeh-Roudbar et al. 2020:226 [ref. 37983], Kovačić et al. 2021:62 [ref. 38346], Parenti 2021:69 [ref. 38603], Zarei et al. 2022:165 [ref. 39576], Çiçek et al. 2024:300 [ref. 40789], Vasil'eva et al. 2025:[8] [ref. 42434]). Current status: Valid as Benthophilus baeri Kessler 1877. Gobiidae: Gobiinae. Distribution: Eurasia: southern Caspian Sea basin. Habitat: freshwater, brackish.

casachicus, Benthophilus stellatus Ragimov [D. B.] 1978:725 [705], Fig. [Voprosy Ikhtiologii v. 18 (no. 5); ref. 8998] Kenderli Spit, eastern shore of the middle Caspian Sea, depth 36 meters. Holotype: ZISP 53661. Paratypes: (3) Caspian Biol. Station IZA 204 (2) whereabouts unknown. Description on p. 705 of English translation. •Mention -- (Kottelat 1997:176 [ref. 22952]). •Synonym of Benthophilus stellatus (Sauvage 1874), but a valid subspecies as described -- (Coad 1995:30 [ref. 23608]). •Valid as Benthophilus casachicus Ragimov 1978 -- (Reshetnikov et al. 1997:752 [ref. 24702], Vasil'eva 1998:127 [ref. 23591], Bogutskaya et al. 2001:47 [ref. 26178], Vasil'eva & Miller in Miller 2004:182 [ref. 27853], Bogutskaya & Naseka 2004:215 [ref. 28183], Boldyrev & Bogutskaya 2004:132 [ref. 28099], Boldyrev & Bogutskaya 2007:69 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:376 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:80 [ref. 35254], Esmaeili et al. 2018:60 [ref. 36089], Parenti 2021:69 [ref. 38603], Zarei et al. 2022:165 [ref. 39576], Vasil'eva et al. 2025:[8] [ref. 42434]). Current status: Valid as Benthophilus casachicus Ragimov 1978. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: freshwater, brackish.

ctenolepidus, Benthophilus Kessler [K. F.] 1877:48, Pl. 4 (fig. 11) [The Aralo-Caspian Expedition; ref. 2597] Caspian Sea, 40°08'N, 0°26'E of Baku, Azerbaijan. Lectotype: ZIN 10897. Paralectotypes: (4) BMNH 1897.7.5.13 [ex ZIN] (1), ZIN (lost). Lectotype selected by Ragimov [Rahimov] 1982:48 [ref. 8985]. •Valid as Benthophilus ctenolepidus Kessler 1877 -- (Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:128 [ref. 23591], Coad 1998:103 [ref. 23869], Pinchuk & Miller in Miller 2004:187 [ref. 27853], Bogutskaya & Naseka 2004:216 [ref. 28183], Boldyrev & Bogutskaya 2004:134 [ref. 28099], Boldyrev & Bogutskaya 2007:70 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:80 [ref. 35254], Esmaeili et al. 2018:61 [ref. 36089], Jouladeh-Roudbar et al. 2020:226 [ref. 37983], Parenti 2021:69 [ref. 38603], Zarei et al. 2022:165 [ref. 39576], Çiçek et al. 2024:300 [ref. 40789], Vasil'eva et al. 2025:[8] [ref. 42434]). Current status: Valid as Benthophilus ctenolepidus Kessler 1877. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: freshwater, brackish.

durrelli, Benthophilus Boldyrev [V. S.] & Bogutskaya [N. G.] 2004:130, Fig. 1 [Zoosystematica Rossica v. 13 (no. 1); ref. 28099] Tsimlyansk Reservoir, Lower Don River, Russia. Holotype: ZISP [= ZIN 53211]. Paratypes: ZIN 9972 (7), 10167 (7), 44445 (5), 44449 (1), 53212 (4), 53213 (4), 53214 (23), 53215 (25). •Synonym of Benthophilus stellatus (Sauvage 1874) -- (Parin et al. 2015:478 [ref. 33547]). •Valid as Benthophilus durrelli Boldyrev & Bogutskaya 2004 -- (Bogutskaya & Naseka 2004:216 [ref. 28183], Boldyrev & Bogutskaya 2007:53 [ref. 29150], Kottelat & Freyhof 2007:557 [ref. 29996], Neilson & Stepien 2009:96 [ref. 30497], Freyhof 2011:282 [ref. 31742], Kovačić et al. 2021:57 [ref. 38346], Parenti 2021:70 [ref. 38603]). Current status: Valid as Benthophilus durrelli Boldyrev & Bogutskaya 2004. Gobiidae: Gobiinae. Distribution: Eurasia: Sea of Azov and Don River basin; introduced into the Volga River. Habitat: freshwater, brackish.

granulosus, Benthophilus Kessler [K. F.] 1877:57, Pl. 5 (fig. 14) [The Aralo-Caspian Expedition; ref. 2597] Baku Bay, Caspian Sea, Azerbaijan. Lectotype: ZISP [= ZIN] 2242. Syntypes: (7 + 1) BMNH 1897.7.5.14 [ex ZIN] (1), ZIN 10898 (1, damaged). Lectotype selected by Boldyrev & Bogutskaya 2007:51 [ref. 29150]. •Valid as Benthophilus granulosus Kessler 1877 -- (Berg 1949:1119 [ref. 20662], Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:128 [ref. 23591], Bogutskaya et al. 2001:47 [ref. 26178], Pinchuk & Miller in Miller 2004:192 [ref. 27853], Bogutskaya & Naseka 2004:216 [ref. 28183], Boldyrev & Bogutskaya 2007:49 [ref. 29150], Kottelat & Freyhof 2007:558 [ref. 29996], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:376 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:80 [ref. 35254], Esmaeili et al. 2018:61 [ref. 36089], Jouladeh-Roudbar et al. 2020:227 [ref. 37983], Kovačić et al. 2021:62 [ref. 38346], Parenti 2021:70 [ref. 38603], Zarei et al. 2022:166 [ref. 39576], Çiçek et al. 2024:301 [ref. 40789], Vasil'eva et al. 2025:[8] [ref. 42434]). Current status: Valid as Benthophilus granulosus Kessler 1877. Gobiidae: Gobiinae. Distribution: Eurasia: southern Caspian Sea basin. Habitat: freshwater, brackish.

grimmi, Benthophilus Kessler [K. F.] 1877:59, Pl. 5 (fig. 13) [The Aralo-Caspian Expedition; ref. 2597] Middle and southern Caspian Sea, depth 245-756 feet [35-108 Russian fathoms]. Lectotype: BMNH 1897.7.5.15. Paralectotypes: (7) ?NMW 76987 (1), SPU 394 [465/412] (1), ZIN 10893 (1). Lectotype selected by Boldyrev & Bogutskaya 2007:45 [ref. 29150]. Boldyrev & Bogutskaya 2007:45 [ref. 29150] discuss types but do not mention NMW 76987. •Valid as Benthophilus grimmi Kessler 1877 -- (Berg 1949:1121 [ref. 20662], Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:128 [ref. 23591], Pinchuk & Miller in Miller 2004:198 [ref. 27853], Bogutskaya & Naseka 2004:216 [ref. 28183], Boldyrev & Bogutskaya 2007:42 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:376 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:80 [ref. 35254], Esmaeili et al. 2018:61 [ref. 36089], Parenti 2021:70 [ref. 38603], Zarei et al. 2022:166 [ref. 39576], Vasil'eva et al. 2025:[8] [ref. 42434]). Current status: Valid as Benthophilus grimmi Kessler 1877. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: brackish.

kessleri, Benthophilus grimmi var. Berg [L. S.] 1927:343 [Collection of papers in honor of Prof. Knipowitsch; ref. 15840] 41°51'N, 52°15'E, depth 75 meters, Caspian Sea. Lectotype: ZISP [= ZIN] 53670 [ex 24349]. Paralectotypes: (10) ZIN 24349 (now 6). Boldyrev & Bogutskaya 2007:41 [ref. 29150] identify 5 paralectotypes of kessleri as svetovidovi. Lectotype selected by Boldyrev & Bogutskaya 2007:49 [ref. 29150]. •Synonym of Benthophilus grimmi Kessler 1877, but a valid subspecies as described -- (Berg 1949:1122 [ref. 20662], Coad 1995:29 [ref. 23608]). •Valid as Benthophilus kessleri Berg 1927 -- (Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:128 [ref. 23591], Pinchuk & Miller in Miller 2004:201 [ref. 27853], Bogutskaya & Naseka 2004:217 [ref. 28183], Boldyrev & Bogutskaya 2007:47 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30497], Esmaeili et al. 2010:376 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:80 [ref. 35254], Esmaeili et al. 2018:61 [ref. 36089], Parenti 2021:70 [ref. 38603], Zarei et al. 2022:167 [ref. 39576], Vasil'eva et al. 2025:[9] [ref. 42434]). Current status: Valid as Benthophilus kessleri Berg 1927. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: freshwater, brackish.

lencoranicus, Benthophilus magistri Ragimov [D. B.] 1982:53 [Zoologichesky Zhurnal v. 61 (no. 1); ref. 8985] Off Zelenyy Bugor, 38°10'N, southeastern Caspian Sea, Turkmenistan, depth 42 meters. Holotype: whereabouts unknown. Paratypes: CBSIZA 546 (20). Additional material: ZIN 23131 and 41912 (9). •Synonym of Benthophilus magistri Iljin 1927, but a valid subspecies as described -- (Coad 1995:30 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:130 [ref. 23591]). •Synonym of Benthophilus magistri Iljin 1927 -- (Pinchuk et al. in Miller 2004:235 [ref. 27853]). •Synonym of Benthophilus ctenolepidus Kesssler 1877 -- (Boldyrev & Bogutskaya 2007:70 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Jouladeh-Roudbar et al. 2020:226 [ref. 37983], Parenti 2021:70 [ref. 38603], Çiçek et al. 2024:300 [ref. 40789]). Current status: Synonym of Benthophilus ctenolepidus Kessler 1877. Gobiidae: Gobiinae. Habitat: freshwater.

leobergius, Benthophilus stellatus Berg [L. S.] 1949:1116, Figs. 858-859 [Freshwater fishes of the U.S.S.R. and adjacent countries No. 30; ref. 20662] Astrabadskiy Bay, Caspian Sea, Iran. Lectotype: ZISP 23128. Syntypes: ZISP 10891 (1). Appeared first as name only in Iljin 1949:28 [ref. 18323] -- not available, no distinguishing features provided. The original spelling was leobergius, and that spelling probalbly should be maintained [Art. 31]. Lectotype selected by Boldyrev & Bogutskaya 2007:62 [ref. 29150] but they selected a 48-mm male on p. 60 and an 85-mm male on p. 63 and Fig. 28a -- I [WNE] regard the 83-mm male as lectotype. •Synonym of Benthophilus stellatus (Sauvage 1874), but a valid subspecies as described -- (Coad 1995:30 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:131 [ref. 23591]). •Mention, with name emended to leobergi and dated to Ragimov [Rahimov] 1978:707 [ref. 8998] -- (Kottelat 1997:176 [ref. 22952]). •Synonym of Benthophilus stellatus (Sauvage 1874) -- (Bogutskaya et al. 2001:47 [ref. 26178] as leobergi). •Valid as Benthophilus leobergi Berg 1949 -- (Pinchuk et al. in Miller 2004:206 [ref. 27853]). •Not available -- (Parenti 2021:445 [ref. 38603]). •Valid as Benthophilus leobergius Berg 1949 -- (Bogutskaya & Naseka 2004:217 [ref. 28183], Boldyrev & Bogutskaya 2004:132 [ref. 28099], Boldyrev & Bogutskaya 2007:60 [ref. 29150], Kottelat & Freyhof 2007:559 [ref. 29996], Neilson & Stepien 2009:96 [ref. 30497], Esmaeili et al. 2010:376 [ref. 33391], Parin et al. 2015:479 [ref. 33547] with question, Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:80 [ref. 35254], Esmaeili et al. 2018:61 [ref. 36089], Jouladeh-Roudbar et al. 2020:227 [ref. 37983], Zarei et al. 2021:[7] [ref. 38262], Kovačić et al. 2021:62 [ref. 38346], Parenti 2021:70 [ref. 38603], Zarei et al. 2022:167 [ref. 39576], Çiçek et al. 2024:301 [ref. 40789], Vasil'eva et al. 2025:[9] [ref. 42434]). Current status: Valid as Benthophilus leobergius Berg 1949. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin and Volga River Delta. Habitat: freshwater, brackish.

leptocephalus, Benthophilus Kessler [K. F.] 1877:45 [The Aralo-Caspian Expedition; ref. 2597] Southern Caspian Sea, depth 756 feet [108 Russian fathoms, 230 meters]. Lectotype: ZIN 10894. Paralectotypes: (1). Lectotype selected by Ragimov 1982:51 as the 32-mm specimen in ZISP 10894. Comments by Boldyrev & Bogutskaya 2007:75 [ref. 29150] on Berg's lectotype establishment (see Eschmeyer 1998:893) subject to interpretation. •Valid as Benthophilus leptocephalus Kessler 1877 -- (Berg 1949:1117 [ref. 20662], Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:129 [ref. 23591], Pinchuk & Miller in Miller 2004:216 [ref. 27853], Bogutskaya & Naseka 2004:217 [ref. 28183], Boldyrev & Bogutskaya 2004:134 [ref. 28099], Boldyrev & Bogutskaya 2007:73 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:376 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:81 [ref. 35254], Esmaeili et al. 2018:61 [ref. 36089], Jouladeh-Roudbar et al. 2020:228 [ref. 37983], Parenti 2021:70 [ref. 38603], Zarei et al. 2022:167 [ref. 39576], Vasil'eva et al. 2025:[9] [ref. 42434]). Current status: Valid as Benthophilus leptocephalus Kessler 1877. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: freshwater, brackish.

leptorhynchus, Benthophilus Kessler [K. F.] 1877:56, Pl. 5 (figs. 12-12a) [The Aralo-Caspian Expedition; ref. 2597] Middle Caspian Sea, depth 490 feet [70 Russian fathoms]. lectotype: ZISP [= ZIN] 10895 (damaged). Paralectotypes: (2) whereabouts unknown. Lectotype selected by Boldyrev & Bogutskaya 2007:42 [ref. 29150]. •Valid as Benthophilus leptorhynchus Kessler 1877 -- (Berg 1949:1120 [ref. 20662], Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:129 [ref. 23591], Pinchuk & Miller in Miller 2004:220 [ref. 27853], Bogutskaya & Naseka 2004:217 [ref. 28183], Boldyrev & Bogutskaya 2007:41 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:376 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:81 [ref. 35254], Esmaeili et al. 2018:62 [ref. 36089], Parenti 2021:71 [ref. 38603], Zarei et al. 2022:168 [ref. 39576], Vasil'eva et al. 2025:[9] [ref. 42434]). Current status: Valid as Benthophilus leptorhynchus Kessler 1877. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: brackish.

macrocephalus, Gobius Pallas [P. S.] 1787:352, Pl. 10 (figs. 4-6) [Nova Acta Academiae Scientiarum Imperialis Petropolitanae v. 1 (Physica) (for 1783); ref. 18507] Mare Caspicum [Caspian Sea]. No types known. Page as "(52)" between pp. 351 and 353. •Valid as Benthophilus macrocephalus (Pallas 1787) -- (Berg 1949:1109 [ref. 20662], Mitrofanov & Dukravec 1989:280 [ref. 37748], Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:129 [ref. 23591], Coad 1998:103 [ref. 23869], Vasil'eva 2000:764 [ref. 25083], Bogutskaya et al. 2001:47 [ref. 26178], Pinchuk et al. in Miller 2004:225 [ref. 27853], Bogutskaya & Naseka 2004:218 [ref. 28183], Boldyrev & Bogutskaya 2004:132 [ref. 28099], Boldyrev & Bogutskaya 2007:67 [ref. 29150], Kottelat & Freyhof 2007:560 [ref. 29996], Neilson & Stepien 2009:96 [ref. 30497], Esmaeili et al. 2010:377 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:81 [ref. 35254], Esmaeili et al. 2018:62 [ref. 36089], Jouladeh-Roudbar et al. 2020:228 [ref. 37983], Parenti 2021:71 [ref. 38603], Zarei et al. 2022:168 [ref. 39576], Çiçek et al. 2024:302 [ref. 40789], Vasil'eva et al. 2025:[9] [ref. 42434]). Current status: Valid as Benthophilus macrocephalus (Pallas 1787). Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: freshwater, brackish.

maeotica, Benthophilus macrocephalus var. Kuznetsov [I. D.] 1888:207 [Trudy St.-Peterburgskago Obshchestva Estestvoispytateley. Otdel Zoologii i Fiziologii / Travaux de la Société des Naturalistes de St. Pétersbourg v. 19; ref. 18441] Arrabatskaya Strelka, Sea Of Azov, Ukraine. Lectotype: ZISP 10890. Paralectotypes: (3 + 2) None found. Lectotype selected by Boldyrev & Bogutskaya 2007:65 [ref. 29150]. •Synonym of Benthophilus stellatus (Sauvage 1874) -- (Berg 1949:1114 [ref. 20662], Miller 1973:490 [ref. 6888], Kottelat 1997:175 [ref. 22952], Vasil'eva 2003:S43 [ref. 28444], Vasil'eva 1998:130 [ref. 23591], Pinchuk & Miller in Miller 2004:257 [ref. 27853], Bogutskaya & Naseka 2004:219 [ref. 28183], Boldyrev & Bogutskaya 2007:63 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Parin et al. 2015:478 [ref. 33547], Parenti 2021:72 [ref. 38603]). Current status: Synonym of Benthophilus stellatus (Sauvage 1874). Gobiidae: Gobiinae. Habitat: freshwater.

magistri, Benthophilus Iljin [B. S.] 1927:131, 137, Figs. ?, 5, 8, 9. [Trudy Azovsko-Chernomorskoy Nauchno-Promyslovoy Ekspeditsii v. 2; ref. 5613] Sea of Azov at Krivaya Kosa Spit, Ukraine. Lectotype: ZISP 23138. Paralectotypes: ZISP 53667 [ex 23138] (1). Lectotype selected by Boldyrev & Bogutskaya 2007:60 [ref. 29150]. •Valid as Benthophilus magistri Iljin 1927 -- (Berg 1949:1112 [ref. 20662], Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:129 [ref. 23591], Coad 1998:103 [ref. 23869], Vasil'eva 2000:764 [ref. 25083], Bogutskaya et al. 2001:47 [ref. 26178], Vasil'eva 2003:S42 [ref. 28444], Pinchuk et al. in Miller 2004:235 [ref. 27853], Bogutskaya & Naseka 2004:218 [ref. 28183], Boldyrev & Bogutskaya 2004:132 [ref. 28099], Boldyrev & Bogutskaya 2007:58 [ref. 29150], Kottelat & Freyhof 2007:560 [ref. 29996], Vasil'eva 2007:150 [ref. 30517], Neilson & Stepien 2009:96 [ref. 30487], Kovačić & Patzner 2011:184 [ref. 31737], Parin et al. 2015:477 [ref. 33547], Manilo 2020:67 [ref. 37795], Kovačić et al. 2021:57 [ref. 38346], Parenti 2021:71 [ref. 38603], Vasil'eva et al. 2025:[9] [ref. 42434]). Current status: Valid as Benthophilus magistri Iljin 1927. Gobiidae: Gobiinae. Distribution: Eurasia: eastern part of Sea of Azov and Caspian Sea. Habitat: freshwater, brackish.

mahmudbejovi, Benthophilus Ragimov [D. B.] 1976:1196, Fig. [Zoologichesky Zhurnal v. 55 (no. 8); ref. 7386] Off Cape Peschanyy, middle Caspian Sea, Kazakhstan, depth 40 meters. Holotype: ZISP 53655 [ex IZAB 027]. Paratypes: ZISP 53656 [ex IZAB 012] (20). •Valid as Benthophilus mahmudbejovi Ragimov 1976 -- (Coad 1995:30 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:130 [ref. 23591], Bogutskaya et al. 2001:47 [ref. 26178], Vasil'eva 2003:S43 [ref. 28444], Pinchuk & Miller in Miller 2004:241 [ref. 27853], Bogutskaya & Naseka 2004:218 [ref. 28183], Boldyrev & Bogutskaya 2004:130 [ref. 28099], Boldyrev & Bogutskaya 2007:51 [ref. 29150], Kottelat & Freyhof 2007:561 [ref. 29996], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:377 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:81 [ref. 35254], Esmaeili et al. 2018:62 [ref. 36089], Parenti 2021:71 [ref. 38603], Zarei et al. 2022:168 [ref. 39576], Vasil'eva et al. 2025:[9] [ref. 42434]). Current status: Valid as Benthophilus mahmudbejovi Ragimov 1976. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea and upper Volga delta [not Sea of Azov]. Habitat: freshwater, brackish.

monstrosus, Benthophilus Kuznetsov [I. D.] 1888:208 [Trudy St.-Peterburgskago Obshchestva Estestvoispytateley. Otdel Zoologii i Fiziologii / Travaux de la Société des Naturalistes de St. Pétersbourg v. 19; ref. 18441] Berdyanskiy Liman, Sea of Azov, Ukraine. Holotype (unique): lost. •Synonym of Benthophilus stellatus (Sauvage 1874) -- (Berg 1949:1114 [ref. 20662], Miller 1973:490 [ref. 6888], Kottelat 1997:176 [ref. 22952], Vasil'eva 1998:130 [ref. 23591], Vasil'eva 2003:S43 [ref. 28444] as mapstresus, Pinchuk & Miller in Miller 2004:257 [ref. 27853], Boldyrev & Bogutskaya 2007:63 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Parin et al. 2015:478 [ref. 33547], Parenti 2021:72 [ref. 38603]). Current status: Synonym of Benthophilus stellatus (Sauvage 1874). Gobiidae: Gobiinae. Habitat: freshwater, brackish.

nudus, Benthophilus macrocephalus var. Berg [L. S.] 1898:334 [Dnevnik Zoologicheskogo Otdeleniya Obshchestva Lyubiteley Estestvoznaniya, Antropologii i Etnografii. Moskva v. 2 (no. 8); ref. 19641] Dniester River at Bendery [Benderi], Moldova. Lectotype: ZMMU P-1431. Paralectotypes: ZMUU P-4206 (7), P-4222 (2). Lectotype selected by Boldyrev & Bogutskaya 2007:67 [ref. 29150]. •Synonym of Benthophilus stellatus (Sauvage 1874) -- (Berg 1949:1114 [ref. 20662], Miller 1973:490 [ref. 6888], Kottelat 1997:176 [ref. 22952], Vasil'eva 1998:130 [ref. 23591], Vasil'eva 2003:S43 [ref. 28444], Pinchuk & Miller in Miller 2004:257 [ref. 27853], Parin et al. 2015:478 [ref. 33547]). •Valid as Benthophilus nudus Berg 1898 -- (Boldyrev & Bogutskaya 2007:65 [ref. 29150], Kottelat & Freyhof 2007:562 [ref. 29996], Neilson & Stepien 2009:96 [ref. 30497], Kovačić & Patzner 2011:184 [ref. 31737], Freyhof 2011:282 [ref. 31742], Çiçek et al. 2015:152 [ref. 35151], Çiçek et al. 2020:256 [ref. 37644], Manilo 2020:67 [ref. 37795], Kovačić et al. 2021:57 [ref. 38346], Parenti 2021:71 [ref. 38603], Çiçek et al. 2023:428 [ref. 40625], Iftime & Oţel 2025:23 [ref. 42600]). Current status: Valid as Benthophilus nudus Berg 1898. Gobiidae: Gobiinae. Distribution: Eurasia: Black Sea watersheds. IUCN (2024): Least Concern. Habitat: freshwater, brackish.

persicus, Benthophilus Kovačić [M.], Esmaeili [H. R.], Zarei [F.], Abbasi [K.] & Schliewen [U. K.] 2021:47, Figs. 2-7 [Zootaxa 4980 (no. 1); ref. 38346] Anzali, southern Caspian Sea, Gilan Province, Iran, 37°29'N, 49°29'E. Holotype: ZSM 47595. Paratypes: PMR, ZM-CBSU, ZSM. Plus additional non-type material. •Valid as Benthophilus persicus Kovačić, Esmaeili, Zarei, Abbasi & Schliewen 2021 -- (Zarei et al. 2022:168 [ref. 39576], Çiçek et al. 2024:302 [ref. 40789]). Current status: Valid as Benthophilus persicus Kovačić, Esmaeili, Zarei, Abbasi & Schliewen 2021. Gobiidae: Gobiinae. Distribution: Eurasia: southwestern Caspian Sea. Habitat: brackish.

pinchuki, Benthophilus ctenolepidus Ragimov [D. B.] 1982:49 [Zoologichesky Zhurnal v. 61 (no. 1); ref. 8985] Off Belyy Bugor, 37°40'N, southeastern Caspian Sea, Turkmenistan, depth 30 meters. Holotype: ZISP 53569 [ex IZA]. Paratypes: IZA 304 (8), ZISP 53660 [ex IZA 304] (6). Additional material: ZIN 33143 (6), 44346 (1). •Synonym of Benthophilus ctenolepidus Kessler 1877, but a valid subspecies as described -- (Coad 1995:29 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:128 [ref. 23591]). •Valid as Benthophilus pinchuki Ragimov 1982 -- (Pinchuk & Miller in Miller 2004:246 [ref. 27853], Bogutskaya & Naseka 2004:218 [ref. 28183], Boldyrev & Bogutskaya 2004:134 [ref. 28099], Boldyrev & Bogutskaya 2007:77 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:377 [ref. 33391], Jouladeh-Roudbar et al. 2015:897 [ref. 34062], Esmaeili et al. 2017:81 [ref. 35254], Esmaeili et al. 2018:62 [ref. 36089], Jouladeh-Roudbar et al. 2020:230 [ref. 37983], Kovačić et al. 2021:62 [ref. 38346], Parenti 2021:71 [ref. 38603], Zarei et al. 2022:169 [ref. 39576], Çiçek et al. 2024:303 [ref. 40789], Vasil'eva et al. 2025:[10] [ref. 42434]). Current status: Valid as Benthophilus pinchuki Ragimov 1982. Gobiidae: Gobiinae. Distribution: Eurasia: southern Caspian Sea basin. Habitat: freshwater, brackish.

ponticus, Benthophilus macrocephalus Berg [L. S.] 1916:427 [Les Poissons des eaux douces de la Russie; ref. 277] Northwestern Black Sea. No types known. •Synonym of Benthophilus nudus Berg 1898 -- (Boldyrev & Bogutskaya 2007:65 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487]). •Synonym of Benthophilus stellatus (Sauvage 1874) -- (Berg 1949:1114 [ref. 20662], Miller 1973:490 [ref. 6888], Vasil'eva 1998:131 [ref. 23591], Vasil'eva 2003:S43 [ref. 28444], Pinchuk & Miller in Miller 2004:257 [ref. 27853], Parin et al. 2015:478 [ref. 33547], Parenti 2021:72 [ref. 38603]). Current status: Synonym of Benthophilus stellatus (Sauvage 1874). Gobiidae: Gobiinae. Habitat: freshwater, brackish.

ragimovi, Benthophilus Boldyrev [V. S.] & Bogutskaya [N. G.] 2004:132, Fig. 2 [Zoosystematica Rossica v. 13 (no. 1); ref. 28099] Western coast of Caspian Sea, off Yamma-Kilyazi, Azerbaijan, depth 50 meters. Holotype: ZIN 53216. Paratypes: ZIAZ 93 (8). 467 (6); ZIN 23126 (3), 53217 (1), 53218 (7), 53219 (4); plus additional non-type material. •Valid as Benthophilus ragimovi Boldyrev & Bogutskaya 2004 -- (Bogutskaya & Naseka 2004:218 [ref. 28183], Boldyrev & Bogutskaya 2007:75 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30497], Esmaeili et al. 2010:377 [ref. 33391], Jouladeh-Roudbar et al. 2015:898 [ref. 34062], Esmaeili et al. 2017:81 [ref. 35254], Esmaeili et al. 2018:62 [ref. 36089], Jouladeh-Roudbar et al. 2020:230 [ref. 37983], Parenti 2021:71 [ref. 38603], Zarei et al. 2022:170 [ref. 39576], Vasil'eva et al. 2025:[10] [ref. 42434]). Current status: Valid as Benthophilus ragimovi Boldyrev & Bogutskaya 2004. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: freshwater, brackish.

rassi, Benthophilus macrocephalus Ragimov [D. B.] 1988 Appeared in 1988 and 1991, but not in an available publication. •Not available -- see Boldyrev & Bogutskaya 2007:68-69 [ref. 29150]. Current status: Synonym of Benthophilus macrocephalus (Pallas 1787). Gobiidae: Gobiinae.

spinosus, Benthophilus Kessler [K. F.] 1877:50 [The Aralo-Caspian Expedition; ref. 2597] Middle Caspian Sea, depth 140 feet [20 Russian fathoms]. Lectotype: ZISP 10896. Syntypes: (5) BMNH 1897.7.5.17 [ex MLSU] (1), others lost. Lectotype selected by Boldyrev & Bogutskaya 2007:81 [ref. 29150]. •Valid as Benthophilus spinosus Kessler 1877 -- (Berg 1949:1117 [ref. 20662], Coad 1995:30 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:130 [ref. 23591], Pinchuk & Miller in Miller 2004:251 [ref. 27853], Bogutskaya & Naseka 2004:219 [ref. 28183], Boldyrev & Bogutskaya 2007:79 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:377 [ref. 33391], Jouladeh-Roudbar et al. 2015:898 [ref. 34062], Esmaeili et al. 2017:81 [ref. 35254], Esmaeili et al. 2018:63 [ref. 36089], Parenti 2021:71 [ref. 38603], Zarei et al. 2022:170 [ref. 39576], Vasil'eva et al. 2025:[10] [ref. 42434]). Current status: Valid as Benthophilus spinosus Kessler 1877. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: brackish.

squamatus, Benthophilus Baer [K. M.] in Lukina 1984:168 [Caspian expedition of K. M. Baer in 1853-1857; ref. 29151] Bight at Novopetrovsk, Mangyshlak. No types known. Original not seen; see Boldyrev & Bogutskaya 2007:51 [ref. 29150]. •Synonym of Benthophilus granulosus Kessler 1877 -- (Boldyrev & Bogutskaya 2007:49, 51 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30497], Jouladeh-Roudbar et al. 2020:227 [ref. 37983], Parenti 2021:70 [ref. 38603]). Current status: Synonym of Benthophilus granulosus Kessler 1877. Gobiidae: Gobiinae. Habitat: freshwater, brackish.

stellatus, Doliichthys Sauvage [H.-E.] 1874:336 [Revue et Magasin de Zoologie (Sér. 3) v. 2 [1874]; ref. 3873] Don River at Voronezh [Woronesch], Russia. Syntypes: MNHN 0000-7341 (2). Type catalog: Bauchot et al. 1991:21 [ref. 19958]. •Valid as Benthophilus stellatus (Sauvage 1874) -- (Berg 1949:1114 [ref. 20662], Miller in Whitehead et al. 1986:1025 [ref. 13677], Coad 1995:30 [ref. 23608], Kottelat 1997:175 [ref. 22952], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:130 [ref. 23591], Coad 1998:103 [ref. 23869], Bogutskaya et al. 2001:47 [ref. 26178], Vasil'eva 2003:S43 [ref. 28444], Pinchuk & Miller in Miller 2004:257 [ref. 27853], Bogutskaya & Naseka 2004:219 [ref. 28183], Boldyrev & Bogutskaya 2004:130 [ref. 28099], Vassilev & Pehlivanov 2005:182 [ref. 28369], Boldyrev & Bogutskaya 2007:63 [ref. 29150], Kottelat & Freyhof 2007:563 [ref. 29996], Vasil'eva 2007:150 [ref. 30517], Neilson & Stepien 2009:96 [ref. 30487], Kovačić & Patzner 2011:184 [ref. 31737], Parin et al. 2015:478 [ref. 33547], Manilo 2020:68 [ref. 37795], Kovačić et al. 2021:61 [ref. 38346], Kovačić et al. 2021:87 [ref. 39014], Parenti 2021:72 [ref. 38603], Vasil'eva et al. 2025:[10] [ref. 42434]). Current status: Valid as Benthophilus stellatus (Sauvage 1874). Gobiidae: Gobiinae. Distribution: Eurasia: Sea of Azov and Don River basin. Habitat: freshwater, brackish.

svetovidovi, Benthophilus Pinchuk [V. I.] & Ragimov [D. B.] 1979:515, fig'd [Zoologichesky Zhurnal v. 58 (no. 4); ref. 8797] Caspian Sea, depth 86 meters. Holotype: ZIN 43672. Paratypes: ZIN 43673 (1, not on shelves in 1996). •Valid as Benthophilus svetovidovi Pinchuk & Ragimov 1979 -- (Coad 1995:30 [ref. 23608], Reshetnikov et al. 1997:753 [ref. 24702], Vasil'eva 1998:131 [ref. 23591], Pinchuk & Miller in Miller 2004:268 [ref. 27853], Boldyrev & Bogutskaya 2007:45 [ref. 29150], Neilson & Stepien 2009:96 [ref. 30487], Esmaeili et al. 2010:377 [ref. 33391], Jouladeh-Roudbar et al. 2015:898 [ref. 34062], Esmaeili et al. 2017:82 [ref. 35254], Esmaeili et al. 2018:63 [ref. 36089], Parenti 2021:72 [ref. 38603], Zarei et al. 2022:170 [ref. 39576], Vasil'eva et al. 2025:[10] [ref. 42434]). Current status: Valid as Benthophilus svetovidovi Pinchuk & Ragimov 1979. Gobiidae: Gobiinae. Distribution: Eurasia: Caspian Sea basin. Habitat: brackish.