Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the genus Creagrutus: [ 95 ] records

affinis, Creagrutus Steindachner [F.] 1880:79 [27] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 42; ref. 14403] Río Cauca, northern Colombia. Lectotype: NMW 67120.1. Paralectotypes: NMW 67120.2-5 (4). On p. 27 of separate. Lectotype selected by Harold & Vari 1994:25 [ref. 21969]. •Valid as Creagrutus affinis Steindachner 1880 -- (Harold & Vari 1994:23 [ref. 21969], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:121 [ref. 27061], Román-Valencia & Cala 1997:150 [ref. 30645], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Albornoz-Garzón et al. 2018:349 [ref. 36150], Ardila Rodríguez 2020:12 [ref. 37904], Ardila Rodríguez 2021:16 [ref. 38586], Vanegas-Ríos et al. 2021:63 [ref. 38668], Ardila Rodríguez 2023:12 [ref. 40275]). Current status: Valid as Creagrutus affinis Steindachner 1880. Stevardiidae: Creagrutinae. Distribution: South America: various streams of the Pacific and Caribbean slopes, Colombia and Panama. Habitat: freshwater.

amoenus, Creagrutus Fowler [H. W.] 1943:239, Fig. 18 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 95; ref. 1443] Río Orteguasa, Florencia, Caquetá, Colombia. Holotype: ANSP 70499. Paratypes: ANSP 70500-02 (3), 70503 (1 c&s). Type catalog: Böhlke 1984:42 [ref. 13621]. •Valid as Creagrutus amoenus Fowler 1943 -- (Vari & Harold 2001:58 [ref. 25842], Powers & Mayden 2003:578 [ref. 27129], Vari & Harold in Reis et al. 2003:121 [ref. 27061], Román-Valencia & Cala 1997:145 [ref. 30645], Román-Valencia et al. 2010:94 [ref. 30781], Mirande et al. 2013:763 [ref. 33366], Barriga Salazar 2014:107 [ref. 35745], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Albornoz-Garzón et al. 2018:351 [ref. 36150], Albornoz-Garzón et al. 2020:371 [ref. 37405], Meza-Vargas et al. 2021:17 [ref. 39298]). Current status: Valid as Creagrutus amoenus Fowler 1943. Stevardiidae: Creagrutinae. Distribution: South America: upper Amazon River basin (Colombia, Ecuador and Peru). Habitat: freshwater.

anary, Creagrutus Fowler [H. W.] 1913:552, Fig. 16 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 65; ref. 1389] Rio Madeira, about 200 miles [possibly 20] east of 62°20'W, Brazil. Holotype: ANSP 39290 (missing). Paratype: ANSP 39291 (1, c&s). Type catalog: Böhlke 1984:42 [ref. 13621]. •Valid as Creagrutus anary Fowler 1913 -- (Géry & Renno 1989:5 [ref. 13420], Chang & Ortega 1995:2 [ref. 24302], Vari & Harold 2001:63 [ref. 25842], Vari & Harold in Reis et al. 2003:121-122 [ref. 27061], Mirande 2010:544 [ref. 31006], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:17 [ref. 39298], Chuctaya et al. 2022:19 [ref. 42137]). Current status: Valid as Creagrutus anary Fowler 1913. Stevardiidae: Creagrutinae. Distribution: South America: Madeira River basin (Brazil and Peru). IUCN (2022): Least Concern. Habitat: freshwater.

andaki, Creagrutus Albornoz-Garzón [J. G.], Acosta-Santos [A.], Bogotá-Gregory [J. D.] & Agudelo-Córdoba [E.] 2020:366, Figs. 1-3 [Zootaxa 4755 (no. 2); ref. 37405] Río Sucre, km 61, bridge at old road to Neiva, Río Hacha drainage, Río Caquetá basin, Florencia, Caquetá, Colombia, 01°47'43.2"N, 75°38'48.9"W, elevation 1,015 meters. Holotype: CIACOL 3462. Paratypes: CIACOL, IAVHP. •Valid as Creagrutus andaki Albornoz-Garzón, Acosta-Santos, Bogotá-Gregory & Agudelo-Córdoba 2020. Current status: Valid as Creagrutus andaki Albornoz-Garzón, Acosta-Santos, Bogotá-Gregory & Agudelo-Córdoba 2020. Stevardiidae: Creagrutinae. Distribution: Río Caquetá basin, Caquetá, Colombia. Habitat: freshwater.

ardilai, Creagrutus Ardila Rodríguez [C. A.] 2021:10, Figs. 2B, 4A, 5C, 7A, 8C, 9C [Barranquilla, Departamento del Atlántico No. 75; ref. 38586] Quebrada a 8 kilómetros del Municipio El Playón, Departamento de Santander, Colombia. Holotype: CAR 919. Paratypes: CAR 323 (8). •Valid as Creagrutus ardilai Ardila Rodríguez 2021 -- (Ardila Rodríguez 2023:12 [ref. 40275]). Current status: Valid as Creagrutus ardilai Ardila Rodríguez 2021. Stevardiidae: Creagrutinae. Distribution: South America: streams at El Playón, Departamento de Santander, Colombia. Habitat: freshwater.

argos, Creagrutus Ardila Rodríguez [C. A.] 2020:7, Figs. 1-3, 5A, 7A, 8A, 9B-B1, 10A, 11A, 12A, 13 [Barranquilla, Departamento del Atlántico No. 71; ref. 37904] Quebrada El Laurel, afluente del río Saldaña, cuenca alta del río Magdalena, municipio del Guamo, Departamento del Tolima, Colombia. Holotype: CAR 900. Paratypes: CAR, ICNMHN. •Valid as Creagrutus argos Ardila Rodríguez 2020 -- (Ardila Rodríguez 2021:15 [ref. 38586]). Current status: Valid as Creagrutus argos Ardila Rodríguez 2020. Stevardiidae: Creagrutinae. Distribution: Saldaña River drainage, Magdalena River basin, Tolima, Colombia. Habitat: freshwater.

atratus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:66, Figs. 24-25 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Caqueza, tributary To Río Negro, about 2 kilometers upstream of village of Caqueza, Río Meta basin, Cundinamarca, Colombia. Holotype: ICNMHN 4158. Paratypes: ANSP 134080 (5); NRM 16842-43 (19, 15); USNM 353866-67 (5, 5). Type catalog: Mojica & Agudelo-Zamora 2018:14 [ref. 36127]. •Valid as Creagrutus atratus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:122 [ref. 27061], Román-Valencia et al. 2010:95 [ref. 30781], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Urbano-Bonilla et al. 2018:88 [ref. 35773]). Current status: Valid as Creagrutus atratus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Orinoco River basin, Colombia. Habitat: freshwater.

atrisignum, Creagrutus Myers [G. S.] 1927:116 [Bulletin of the Museum of Comparative Zoology v. 68 (no. 3); ref. 3096] Upper Rio Maranhão, Brazil. Holotype: CAS 41339 [ex IU 17679]. Paratypes: CAS 41340 [ex IU 17680] (7), MCZ 31571 (1). •Valid as Creagrutus atrisignum Myers 1927 -- (Vari & Harold 2001:69 [ref. 25842], Vari & Harold in Reis et al. 2003:122 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Netto-Ferreira & Moreira 2018:255 [ref. 35759]). Current status: Valid as Creagrutus atrisignum Myers 1927. Stevardiidae: Creagrutinae. Distribution: Maranhão River basin in Tocantins River drainage, Brazil. Habitat: freshwater.

barrigai, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:71, Fig. 27 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Aguarico, near San Pablo de Kantesiya, 0°15'18"S, 76°25'30"W, Sucumbios, Ecuador. Holotype: MEPN 4621. Paratypes: MEPN uncat. (10), 5252 (7); USNM 340976 (8, 2 c&s), 340977-78 (4, 3). •Valid as Creagrutus barrigai Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:122 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Barriga Salazar 2014:107 [ref. 35745], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Meza-Vargas et al. 2021:17 [ref. 39298], Chuctaya et al. 2022:19 [ref. 42137]). Current status: Valid as Creagrutus barrigai Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: western Amazon River basin (Brazil, Colombia, Ecuador and Peru). IUCN (2023): Least Concern. Habitat: freshwater.

beni, Creagrutus Eigenmann [C. H.] 1911:172, Pl. 6 (fig. 2) [Annals of the Carnegie Museum v. 8 (no. 1) (art. 3); ref. 1226] Río Beni at Villa Bella, Amazon system, Bolivia. Holotype (unique): FMNH 54585 [ex CM 3216] (poor condition). Type catalog: Henn 1928:63 [ref. 12305], Ibarra & Stewart 1987:28 [ref. 12367]. No longer known from Argentina -- Koerber 2012:10 [ref. 32229]. •Valid as Creagrutus beni Eigenmann 1911 -- (Ortega & Vari 1986:8 [ref. 9708], Géry & Renno 1989:5 [ref. 13420], Taphorn 1992:167 [ref. 23654] as cf beni, Harold et al. 1994:979 [ref. 22027], Vari & Harold 2001:75 [ref. 25842], Vari & Harold in Reis et al. 2003:122 [ref. 27061], López et al. 2003:25 [ref. 27366], Menni 2004:77 [ref. 28131], Román-Valencia & Cala 1997:149 [ref. 30645], Koerber 2011:9 [ref. 31122], Mirande et al. 2013:763 [ref. 33366], Sarmiento et al. 2014:184 [ref. 35004], Thomaz et al. 2015:Add. 5 [ref. 33893], see DoNascimiento et al. 2017:[11] [ref. 35679], Chuctaya et al. 2022:19 [ref. 42137]). Current status: Valid as Creagrutus beni Eigenmann 1911. Stevardiidae: Creagrutinae. Distribution: South America: upper Amazon basin, upper Madeira River basin (Bolivia, Peru). IUCN (2023): Least Concern. Habitat: freshwater.

beni, Piabina Pearson [N. E.] 1924:45, Pl. 10 (fig. 5) [Indiana University Studies v. 11 (no. 64); ref. 3396] Río Popoi, Upper Beni, Amazon system, Bolivia. Lectotype: CAS 62272 [ex IU 17317]. Paralectotypes: (10) CAS 87389 [ex CAS 62272] (6); UMML 66495 (3). Subjectively invalid; secondarily preoccupied by Creagrutus beni Eigenmann 1911, replaced by Creagrutus pearsoni Mahnert & Géry 1988. Lectotype selected by Vari & Harold 2001:180 [ref. 25842]. •Objective synonym of Creagrutus pearsoni Mahnert & Géry 1988 -- (Vari & Harold 1998:256 [ref. 23845], Vari & Harold 2001:178 [ref. 25842], Vari & Harold in Reis et al. 2003:125 [ref. 27061]). 2°Homonym. Current status: Synonym of Creagrutus pearsoni Mahnert & Géry 1988. Stevardiidae: Creagrutinae. Habitat: freshwater.

boehlkei, Creagrutus Géry [J.] 1972:63, Pl. 4 (fig. 2) [Acta Humboldtiana (Series Geologica, Palaeontologica et Biologica Nr. 2) No. 2; ref. 1595] Amazon River basin, eastern Ecuador (no specific locality). Holotype: ZSM 28428a. Paratypes: (19) MTD 27515 (1) ZMH H 1757 (2), H 1758 (10, now 8), H 1868 (1); ZSM 28428b (2), 28428c (2, missing). Type catalog: Wilkens 1977:156 [ref. 22013], Zarske 2003:16 [ref. 27474], Neumann 2011:244 [ref. 31652]. •Synonym of Creagrutus amoenus Fowler 1943 -- (Vari & Harold 2001:58 [ref. 25842], Vari & Harold in Reis et al. 2003:121 [ref. 27061]). Current status: Synonym of Creagrutus amoenus Fowler 1943. Stevardiidae: Creagrutinae. Habitat: freshwater.

bolivari, Creagrutus Schultz [L. P.] 1944:334, Fig. 49 [Proceedings of the United States National Museum v. 95 (no. 3181); ref. 3960] Río Guárico and tributaries between San Sebastián and San Casimiro, State of Aragua, Venezuela. Holotype: USNM 121497. Paratypes: USNM 121498 (orig. 90, now 82), UMMZ 145040 [ex USNM 121498] (3), MCZ [ex USNM 121498] (5). Type catalog: Vari & Howe 1991:15 [ref. 19022]. •Valid as Creagrutus bolivari Schultz 1944 -- (Taphorn 1992:170 [ref. 23654], Vari & Harold 2001:63 [ref. 25842], Vari & Harold in Reis et al. 2003:122 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento 2016:529 [ref. 34727], DoNascimiento et al. 2017:57 [ref. 35633], Urbano-Bonilla et al. 2018:88 [ref. 35773], Moreno-Burbano et al. 2025:13 [ref. 42549]). Current status: Valid as Creagrutus bolivari Schultz 1944. Stevardiidae: Creagrutinae. Distribution: South America: Orinoco River basin (Venezuela and Colombia). IUCN (2021): Least Concern. Habitat: freshwater.

brevipinnis, Creagrutus Eigenmann [C. H.] 1913:10 [Indiana University Studies No. 18; ref. 1229] Piedra de Moler, Colombia. Holotype: FMNH 56095 [ex CM 4887a]. Paratypes: (105, CM and IU) CAS 41341-42 [ex IU 12728-29] (12, 5), 41375 [ex IU 12730] (8); FMNH 56096-98 (10, 5, 10), 75172 (34); USNM 79188 (4). Type catalog: Henn 1928:63 [ref. 12305], Ibarra & Stewart 1987:28 [ref. 12367], Vari & Howe 1991:15 [ref. 19022]. •Valid as Creagrutus brevipinnis Eigenmann 1913 -- (Harold & Vari 1994:18 [ref. 21969], Vari & Harold 2001:2 [ref. 25842], Román-Valencia 1998:783 [ref. 27073], Vari & Harold in Reis et al. 2003:122 [ref. 27061], Román-Valencia & Cala 1997:148 [ref. 30645], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Albornoz-Garzón et al. 2018:348 [ref. 36150], Ardila Rodríguez 2020:13 [ref. 37904], Ardila Rodríguez 2021:16 [ref. 38586], Ardila Rodríguez 2023:12 [ref. 40275]). Current status: Valid as Creagrutus brevipinnis Eigenmann 1913. Stevardiidae: Creagrutinae. Distribution: South America: upper Cauca River basin, Colombia. Habitat: freshwater.

britskii, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:83, Fig. 31 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Ribeirão Macambira, about 14°20'S, 46°45'W, near bridge on highway GO 112, Iaciara, Goiás, Brazil. Holotype: MZUSP 40537. Paratypes: MNRJ 13048 (6); MZUSP 46546 (18), 50540 (12); USNM 292214 (3), 292226 (13), 340952 (4, 2 c&s). •Valid as Creagrutus britskii Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:122 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Netto-Ferreira & Moreira 2018:255 [ref. 35759]). Current status: Valid as Creagrutus britskii Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Upper Tocantins River basin, Brazil. Habitat: freshwater.

cacique, Creagrutus Flausino Junior [N.] & Lima [F. C. T.] 2019:2, Figs. 1-2 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 166 (no. 1); ref. 36726] Rio Culuene, fish ladder of Paranatinga II hydroelectric dam, Campinápolis, Mato Grosso, Brazil, 13°50'58"S, 53°15'22"W. Holotype: ZUEC 15770. Paratypes: ANSP, INPA, LBP, ZUEC. •Valid as Creagrutus cacique Flausino Junior & Lima 2019. Current status: Valid as Creagrutus cacique Flausino Junior & Lima 2019. Stevardiidae: Creagrutinae. Distribution: Upper Rio Xingu basin, Mato Grosso, Brazil. Habitat: freshwater.

calai, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:85, Fig. 32 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Unnamed stream on Rancho El Viento, across Río Meta from Puerto Lopez, 4°08'N, 72°39'W, Meta, Colombia. Holotype: ANSP 130527. Paratypes: ANSP 139149 (18), 177718 (12); NRM 16848-49 (1, 10), 43015-16 (2, 5); USNM 353304 (4, 2 c&s), 353307 (4), 353868 (2). Type catalog: DoNascimiento et al. 2016:406 [ref. 34787]. •Valid as Creagrutus calai Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:122 [ref. 27061], Román-Valencia et al. 2010:95 [ref. 30781], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Albornoz-Garzón et al. 2018:351 [ref. 36150], Moreno-Burbano et al. 2025:14 [ref. 42549]). Current status: Valid as Creagrutus calai Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: western Meta River basin (eastern Colombia). IUCN (2021): Least Concern. Habitat: freshwater.

caucanus, Creagrutus Eigenmann [C. H.] 1913:9 [Indiana University Studies No. 18; ref. 1229] Paila, Colombia. Holotype: FMNH 56104 [ex CM 4895a]. Paratypes: (70, CM and IU) CAS 41373-74 [ex IU 12736-37] (10, 1), 69304 [ex IU 12738] (8); FMNH 56105-08 (6, 10, 1, 2). Type catalog: Henn 1928:63 [ref. 12305], Ibarra & Stewart 1987:28 [ref. 12367]. •Valid as Creagrutus caucanus Eigenmann 1913 -- (Harold & Vari 1994:5 [ref. 21969], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:122 [ref. 27061], Román-Valencia & Cala 1997:144 [ref. 30645], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Albornoz-Garzón et al. 2018:348 [ref. 36150]). Current status: Valid as Creagrutus caucanus Eigenmann 1913. Stevardiidae: Creagrutinae. Distribution: Upper Cauca River basin, Colombia. Habitat: freshwater.

cerritulus, Creagrutus Netto-Ferreira [A. L.] & Vari [R. P.] 2017:369, Figs. 1-4 [Copeia v. 105 (no. 2); ref. 35453] Río Cachiyacu, tributary of Río Huallaga, Shapaja, Tocache, San Martin, Peru, 8°17'37"S, 76°21'05"W, elevation 520 meters. Holotype: MUSM 54501 (ex-MHNG 2725.023). Paratypes: MHNG, MPEG, MUSM. •Valid as Creagrutus cerritulus Netto-Ferreira & Vari 2017. Current status: Valid as Creagrutus cerritulus Netto-Ferreira & Vari 2017. Stevardiidae: Creagrutinae. Distribution: Río Huallaga drainage, Amazon basin, eastern Peru. Habitat: freshwater.

changae, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:88, Fig. 34 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río San Alejandro, a tributary of Río Sungarayacu, just above junction with Río sungarayacu, about 9°23'S, 75°11'W, Provincia Pachitea, Huanuco, Peru. Holotype: MUSM 8858. Paratypes: FMNH 84265 (10), MUSM 8859 (16), ROM 72378 (10), USNM 285276 (14, 2 c&s). •Valid as Creagrutus changae Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:122 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:17 [ref. 39298], Chuctaya et al. 2022:19 [ref. 42137]). Current status: Valid as Creagrutus changae Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: western tributaries of Ucayali River in depts. Junin, Pasco and Huanuco (Peru). IUCN (2016): Least Concern. Habitat: freshwater.

cijerri, Creagrutus Bisbal [F. J.] & Sánchez [J.] 1997 [v. 1; ref. 25857] •Not available, name only -- (Vari & Harold 2001:3 [ref. 25842]). •Unavailable name -- (Toledo-Piza et al. 2024:443 [ref. 40817]). Nomen Nudum. Current status: Synonym of Creagrutus. Stevardiidae: Creagrutinae.

cimitarraensis, Creagrutus Ardila Rodríguez [C. A.] 2023:15, Figs. 2C, 6B, 8C, 9E, 10B [Barranquilla, Departamento del Atlántico No. 88; ref. 40588] Quebrada La Quitiana, afluente de la quabrada Guayabito, cuenca del río Carare, Municipio de Cimitarra, Departamento de Santander, Colombia. Holotype: CAR 990. Paratypes: CAR 988 (8). •Valid as Creagrutus cimitarraensis Ardila Rodríguez 2023. Current status: Valid as Creagrutus cimitarraensis Ardila Rodríguez 2023. Stevardiidae: Creagrutinae. Distribution: South America: río Carare drainage, Departamento de Santander, Colombia. Habitat: freshwater.

cochui, Creagrutus Géry [J.] 1964:56, Fig. 12 [Tropical Fish Hobbyist v. 13 (no. 4, Dec.); ref. 2264] Upper Amazon, region surrounding Iquitos, Peru. Holotype: USNM 200426 [ex Géry coll. 400.1]. Paratypes: (1). Type catalog: Vari & Howe 1991:15 [ref. 19022]. •Valid as Creagrutus cochui Géry 1964 -- (Ortega & Vari 1986:8 [ref. 9708], Géry & Renno 1989:5 [ref. 13420], Vari & Harold 2001:91 [ref. 25842], Vari & Harold in Reis et al. 2003:122 [ref. 27061], Román-Valencia & Cala 1997:148 [ref. 30645], Mirande et al. 2013:763 [ref. 33366], Barriga Salazar 2014:107 [ref. 35745], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Netto-Ferreira & Moreira 2018:255 [ref. 35759], Flausino & Lima 2019:7 [ref. 36726], Albornoz-Garzón et al. 2020:372 [ref. 37405], Meza-Vargas et al. 2021:17 [ref. 39298], Chuctaya et al. 2022:19 [ref. 42137]). Current status: Valid as Creagrutus cochui Géry 1964. Stevardiidae: Creagrutinae. Distribution: South America: Amazon River basin (Brazil, Ecuador, Colombia and Peru). IUCN (2023): Least Concern. Habitat: freshwater.

convencionensis, Creagrutus Ardila Rodríguez [C. A.] 2023:7, Figs. 1, 4A, 5A, 6A, 7A [Barranquilla, Departamento del Atlántico No. 83; ref. 40275] Quebrada Guamal, afluente de la quebrada Búrbura, alto Río Catatumbo, Municipio de Convención, Departamento de Norte de Santander, Colombia. Holotype: CAR 928. Paratypes: CAR 927 (12). •Valid as Creagrutus convencionensis Ardila Rodríguez 2023. Current status: Valid as Creagrutus convencionensis Ardila Rodríguez 2023. Stevardiidae: Creagrutinae. Distribution: South America: Río Catatumbo drainage, Departamento de Norte de Santander, Colombia. Habitat: freshwater.

corredori, Creagrutus Ardila Rodríguez [C. A.] 2023:7, Figs. 1, 5A, 7A, 9A [Barranquilla, Departamento del Atlántico No. 88; ref. 40588] Río Tucurinca, parte baja, Municipio de Tucurinca, Departamento del Magdalena, Colombia. Holotype: CAR 980. Paratypes: CAR 915 (15). •Valid as Creagrutus corredori Ardila Rodríguez 2023 -- (Ardila Rodríguez 2024:3 [ref. 41561]). Current status: Valid as Creagrutus corredori Ardila Rodríguez 2023. Stevardiidae: Creagrutinae. Distribution: South America: Río Tucurinca drainage, Departamento del Magdalena, Colombia. Habitat: freshwater.

cracentis, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:93, Fig. 26 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Rio Tapajós, sandy beaches on an island located 5 kilometers south of Itiatuba, Pará, Brazil. Holotype: MCP 15213. Paratypes: MCP 22018 (3), USNM 353862 (2, 1 c&s). •Valid as Creagrutus cracentis Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:122 [ref. 27061], Dagosta & Pastana 2014:578 [ref. 33181], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Flausino & Lima 2019:7 [ref. 36726]). Current status: Valid as Creagrutus cracentis Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Lower Tapajós River basin, Brazil. Habitat: freshwater.

crenatus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:95, Fig. 37 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Quebrada Sanare in Yay, 9°45'N, 69°43'W, Lara, Venezuela. Holotype: MHNLS 505. Paratypes: INHS 28851 (5), MHNLS 12900 (6, 1c&s). •Valid as Creagrutus crenatus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:122 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Ardila Rodríguez 2021:16 [ref. 38586], Ardila Rodríguez 2023:11 [ref. 40275]). Current status: Valid as Creagrutus crenatus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: upper Tocuyo, Claro and Turbio River drainages, Caribbean versant, Lara, north-central Venezuela. Habitat: freshwater.

dulima, Creagrutus Albornoz-Garzón [J. G.], Conde-Saldaña [C. C.], García-Melo [J. E.], Taphorn [D. C.] & Villa-Navarro [F. A.] 2018:342, Figs. 1, 2A, 3 [Zootaxa 4482 (no. 2); ref. 36150] Quebrada Chumba, upper Río Magdalena drainage, Ibagué, Departamento Tolima, Colombia, 4°29'06"N, 75°05'49"W, elevation 973 meters. Holotype: CZUT-IC 17663. Paratypes: CP-UCO, CZUT-IC, IAVHP, ICNMHN. Plus additional non-type material. •Valid as Creagrutus dulima Albornoz-Garzón, Conde-Saldaña, García-Melo, Taphorn & Villa-Navarro 2018 -- (Ardila Rodríguez 2020:12 [ref. 37904], Ardila Rodríguez 2021:15 [ref. 38586], Ardila Rodríguez 2023:12 [ref. 40275]). Current status: Valid as Creagrutus dulima Albornoz-Garzón, Conde-Saldaña, García-Melo, Taphorn & Villa-Navarro 2018. Stevardiidae: Creagrutinae. Distribution: South America: upper Río Magdalena basin, Departamento Tolima, Colombia. Habitat: freshwater.

embera, Creagrutus Ardila Rodríguez [C. A.] 2023:10, Figs. 1, 3, 5B, 7B, 9B [Barranquilla, Departamento del Atlántico No. 88; ref. 40588] Río Sinú, cerca de la desembocadura al río Manso, Municipio de Tierralta, Departamento de Córdoba, Colombia. Holotype: CAR 995. Paratypes: CAR. •Valid as Creagrutus embera Ardila Rodríguez 2023. Current status: Valid as Creagrutus embera Ardila Rodríguez 2023. Stevardiidae: Creagrutinae. Distribution: South America: Río Sinú drainage, Departamento de Córdoba, Colombia. Habitat: freshwater.

ephippiatus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:98, Fig. 39 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Upper Río Siapa, Campamento Siapa (Siapa Base Camp), Amazonas, Venezuela. Holotype: MBUCV V-29068. Paratypes: MBUCV V-18623 (3), V-29069 (24); USNM 355116 (7, c&s). •Valid as Creagrutus ephippiatus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:122-123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus ephippiatus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Siapa River, southern tributary of Casiquiare River, upper Negro River basin, Venezuela. Habitat: freshwater.

figueiredoi, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:101, Fig. 40 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Rio Maranhão, upper Rio Tocantins basin, about 35 air kilometers north of Brasilia, about 15°32'S, 47°49'W, Distrito Federal, Brazil. Holotype: MZUSP 50542. Paratypes: MNRJ 14482 (2), 17334 (2), 17339 (3); MZUSP 50543 (9); USNM 292221 (11). •Valid as Creagrutus figueiredoi Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus figueiredoi Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Tocantins River drainage in upper Tocantins and Araguaia River basins, Brazil. Habitat: freshwater.

flavescens, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:104, Fig. 41 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Toguino basin, RíoTiguino No. 3, unnamed tributary of Río Tiguino, 1°07'35"S, 76°56'52"W, Pastaza, Ecuador. Holotype: MEPN 4622. Paratypes: MEPN 4642 (7), 6380 (7); USNM 340974-75 (8, 4). •Valid as Creagrutus flavescens Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Barriga Salazar 2014:107 [ref. 35745], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Albornoz-Garzón et al. 2020:372 [ref. 37405], Meza-Vargas et al. 2021:18 [ref. 39298]). Current status: Valid as Creagrutus flavescens Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: upper Amazon River basin (Colombia, Ecuador and Peru). Habitat: freshwater.

florianensis, Creagrutus Ardila Rodríguez [C. A.] 2023:14, Figs. 2B, 6A, 8B, 9D, 10A [Barranquilla, Departamento del Atlántico No. 88; ref. 40588] Quebrada Huaz, afluente del río Minero, Municipio de Florián, Departamento de Santander, Colombia. Holotype: CAR 975. Paratypes: CAR 904 (6). •Valid as Creagrutus florianensis Ardila Rodríguez 2023. Current status: Valid as Creagrutus florianensis Ardila Rodríguez 2023. Stevardiidae: Creagrutinae. Distribution: South America: río Minero drainage, Departamento de Santander, Colombia. Habitat: freshwater.

gephyrus, Creagrutus Böhlke [J. E.] & Saul [W. G.] 1975:25, Figs. 1-4 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 127 (no. 3); ref. 7109] Río Aguarico at Santa Cecilia, Napo Province, Ecuador. Holotype: ANSP 130516. Paratypes: ANSP 130517 (1), 130519-22 (4); KU 13468 (2). Type catalog: Böhlke 1984:47 [ref. 13621]. •Valid as Creagrutus gephyrus Böhlke & Saul 1975 -- (Vari & Harold 1998:256 [ref. 23845], Vari & Harold 2001:108 [ref. 25842], Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Barriga Salazar 2014:107 [ref. 35745], Dagosta & Pastana 2014:578 [ref. 33181], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298]). Current status: Valid as Creagrutus gephyrus Böhlke & Saul 1975. Stevardiidae: Creagrutinae. Distribution: South America: Andean piedmont rivers (eastern Ecuador, northeastern Peru). Habitat: freshwater.

gracilis, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:111, Fig. 44 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Santiago, at La Poza, 4°01'S, 77°47'W, Provincia Condorcanqui, Amazonas, Peru. Holotype: LACM 41724-27. Paratypes: LACM 41724-32 (10); MUSM 8864 (10), 10760 (6); USNM 341370 (10, 2 c&s), 344534 (4). •Valid as Creagrutus gracilis Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Barriga Salazar 2014:107 [ref. 35745], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298]). Current status: Valid as Creagrutus gracilis Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: Amazonas and Loreto States (Peru), and various localities along eastern slobe of Andean Cordilleras in Ecuador (Ecuador and Peru). Habitat: freshwater.

guanes, Creagrutus Torres-Mejia [M.] & Vari [R. P.] 2005:812, Fig. 1 [Copeia 2005 (no. 4); ref. 28477] Río Fonce, beside road from San Gil to Charalá, 6°28'39.4"N, 73°07'3.0"W, Río Magdalena system, Velle de San José, Santander, Colombia. Holotype: ICNMHN 8520. Paratypes: ICNMHN 8521 (25), 8522 (1), 9893 (20, 1 c&s), 9894 (9), 9895 (1), 9896 (3), 9897 (2, 2 c&s), 9898 (2, 2 c&s). Plus non-type material. Type catalog: Mojica & Agudelo-Zamora 2018:14 [ref. 36127]. The specific name is a noun in apposition. •Valid as Creagrutus guanes Torres-Mejia & Vari 2005 -- (Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Albornoz-Garzón et al. 2018:348 [ref. 36150], Ardila Rodríguez 2020:13 [ref. 37904], Ardila Rodríguez 2021:15 [ref. 38586], Ardila Rodríguez 2023:11 [ref. 40275]). Current status: Valid as Creagrutus guanes Torres-Mejia & Vari 2005. Stevardiidae: Creagrutinae. Distribution: South America: middle Magdalena River basin, Colombia. Habitat: freshwater.

gyrospilus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:113, Fig. 45 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Saguaz, Río Guanare-Apure drainage, bridge near park on road to Chabasquén (=Paraíso de Chabasquén, 9°23'30"N, 70°00'30"W), Portuguesa, Venezuela. Holotype: INHS 69479. Paratypes: INHS 51282 (32), USNM 359486 (8). •Valid as Creagrutus gyrospilus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Bogotá-Gregory et al. 2020:1399 [ref. 37878]). Current status: Valid as Creagrutus gyrospilus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South Ameica: western Orinoco River basin (Venezuela and Colombia). Habitat: freshwater.

hildebrandi, Creagrutus Schultz [L. P.] 1944:330, Figs. 47-48 [Proceedings of the United States National Museum v. 95 (no. 3181); ref. 3960] Río Táchira, 7 kilometers northern of San Antonio, Catatumbo system, Venezuela. Holotype: USNM 121482. Paratypes: MCZ [ex USNM 121486] (10); MHNG [ex USNM 121496 ex Géry] (6); MNHN 1980-1433 [ex USNM 121496] (1); UMMZ 145032 [ex USNM 121496] (10); USNM 121483-85 (1, 20, 97), 121486 (orig. 334, now 314), 121487 (orig. 229, ?now 219), 121488-95 (159, 154, 148, 10, 66, 11, 261, 16), USNM ? (now 387, orig. 403); ZMA 102113 (7) and 113826 (1) [ex USNM 121486] (orig. 10) and ZMA [ex USNM 121487] (10); MHNG [ex USNM 121496 through Géry] (6). Type catalog: Nijssen et al. 1982:17 [ref. 19236], Vari & Howe 1991:15-16 [ref. 19022]. •Valid as Creagrutus hildebrandi Schultz 1944 -- (Harold & Vari 1994:20 [ref. 21969], Galvis et al. 1997:46 [ref. 23651], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:123 [ref. 27061], Román-Valencia & Cala 1997:147 [ref. 30645], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Albornoz-Garzón et al. 2018:349 [ref. 36150]). Current status: Valid as Creagrutus hildebrandi Schultz 1944. Stevardiidae: Creagrutinae. Distribution: Lake Maracaibo basin, Venezuela and Colombia. Habitat: freshwater.

holmi, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:116, Fig. 46 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Provincia Utcubamba, Bagua Grande, San Antonio, Quebrada Jaimito, about 5°47'S, 78°23'W, Amazonas, Peru. Holotype: MUSM 5670. Paratypes: MUSM 8865-66 (7, 5); ROM 55331 (5), 72379 (10); USNM 341368 (6, 1 c&s), 341369 (5). •Valid as Creagrutus holmi Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Barriga Salazar 2014:107 [ref. 35745], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298], Chuctaya et al. 2022:19 [ref. 42137]). Current status: Valid as Creagrutus holmi Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: Marañón River basin (northeastern Peru and Ecuador). IUCN (2016): Least Concern. Habitat: freshwater.

hysginus, Creagrutus Harold [A. S.], Vari [R. P.], Machado-Allison [A.] & Provenzano [F.] 1994:975, Fig. 1 [Copeia 1994 (no. 4); ref. 22027] Río Güiria, near La Toma, Estado Sucre, Venezuela. Holotype: MBUCV-V-20310. Paratypes: CAS 79623 (9, 1 c&s), 80270 (10); MBUCV-V-20304 (12, 1 c&s), V-24002 (20); MCNG 19691 (148); USNM 326035 (10), 326055 (10, 2 c&s). Type catalog: Provenzano et al. 1998:6 [ref. 24196]. •Valid as Creagrutus hysginus Harold, Vari, Machado-Allison & Provenzano 1994 -- (Vari & Harold 2001:119 [ref. 25842], Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366] as hysiginus, Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus hysginus Harold, Vari, Machado-Allison & Provenzano 1994. Stevardiidae: Creagrutinae. Distribution: Rivers draining into Gulf of Paria, Estados Sucre and Monagas, Venezuela. Habitat: freshwater.

ignotus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:122, Fig. 48 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Riacho (small stream) 1, tributary To Rio Preto at road to San Francisco, Município de Diamantino (Rio Arinos basin), Mato Grosso, Brazil. Holotype: MZUSP 45310. Paratypes: MZUSP 42456 (3), 45349 (3), 50165 (2); USNM 326672 (3, 1 c&s), 340960 (3). •Valid as Creagrutus ignotus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:123 [ref. 27061], Dagosta & Pastana 2014:580 [ref. 33181], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Netto-Ferreira & Moreira 2018:255 [ref. 35759]). Current status: Valid as Creagrutus ignotus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Upper Tapajós River and Xingu River basin, Brazil. Habitat: freshwater.

jordanensis, Creagrutus Ardila Rodríguez [C. A.] 2021:9, Figs. 2A, 3B, 5B, 6B, 8B, 9B [Barranquilla, Departamento del Atlántico No. 75; ref. 38586] Río Chicamocha, cuenca del río Magdalena, municipio de Jordán, Departamento de Santander, Colombia. Holotype: CAR 890. Paratypes: CAR 416 (33). •Valid as Creagrutus jordanensis Ardila Rodríguez 2021 -- (Ardila Rodríguez 2023:12 [ref. 40275]). Current status: Valid as Creagrutus jordanensis Ardila Rodríguez 2021. Stevardiidae: Creagrutinae. Distribution: South America: Río Chicamocha drainage, upper Magdalena basin, Departamento de Santander, Colombia. Habitat: freshwater.

kunturus, Creagrutus Vari [R. P.], Harold [A. S.] & Ortega [H.] 1995:290, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 6 (no. 4); ref. 21942] Upper Río Comainas, 20 meters upriver of Puesto de Vigilancia No.22, Cordillera del Condor, Peru, 3°56'30"S, 78°24'20"W. Holotype: MUSM 5667. Paratypes: MUSM 5663 (1), 5665 (2), 5668-69 (2, 33), 6593 (1); USNM 335146-48 (1, 2, 2). •Valid as Creagrutus kunturus Vari, Harold & Ortega 1995 -- (Vari & Harold 2001:125 [ref. 25842], Powers & Mayden 2003:578 [ref. 27129], Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Barriga S. 2042:107 [ref. 35745], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298], Chuctaya et al. 2022:19 [ref. 42137]). Current status: Valid as Creagrutus kunturus Vari, Harold & Ortega 1995. Stevardiidae: Creagrutinae. Distribution: South America: upper Marañón River in northeastern Peru, upper Pastaza River and southwestern portion of Napo River in southeastern Ecuador (Ecuador and Peru). IUCN (2016): Least Concern. Habitat: freshwater.

lassoi, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:128, Fig. 52 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Tupe, Río Aroa drainage, about 12 kilometers north or Aroa, on highway 3, 10°30'19"N, 68°52'33"W, Yaracuy, Venezuela. Holotype: INHS 60094. Paratypes: INHS 28890 (1), 34917 (2), 51283 (22); MCNG 24685 (29); MHNLS 4143 (18); USNM 359487 (5). •Valid as Creagrutus lassoi Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Ardila Rodríguez 2021:16 [ref. 38586], Ardila Rodríguez 2023:11 [ref. 40275]). Current status: Valid as Creagrutus lassoi Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: Aroa and Yaracuy River basins, Caribbean versant drainages of north-central Venezuela. Habitat: freshwater.

lepidus, Creagrutus Vari [R. P.], Harold [A. S.], Lasso [C. A.] & Machado-Allison [A.] 1993:352, Fig. 1 [Ichthyological Exploration of Freshwaters v. 4 (no. 4); ref. 21107] Río Aroa basin, Quebrada El Charal, Finca El Jaguar, Sierra de Aroa, Yaracuy State, Venezuela, about 10°32'N, 68°32'W. Holotype: MHNLS 9659. Paratypes: CAS 79520 (7); MBUCV-V-23560 (12); MHNLS 4144 (12), 4715 (5, ?now 4); USNM 325045 (10). Type catalog: Lasso et al. 1997:41-42 [ref. 23639], Provenzano et al. 1998:6 [ref. 24196]. •Valid as Creagrutus lepidus Vari, Harold, Lasso & Machado-Allison 1993 -- (Vari & Harold 2001:131 [ref. 25842], Vari & Harold in Reis et al. 2003:123 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus lepidus Vari, Harold, Lasso & Machado-Allison 1993. Stevardiidae: Creagrutinae. Distribution: Aroa and Urama River basins, Caribbean versant drainages of Venezuela. Habitat: freshwater.

leuciscus, Creagrutus Regan [C. T.] 1913:463 [Annals and Magazine of Natural History (Series 8) v. 12 (no. 71); ref. 3656] Río Condoto, Río San Juan basin, southwestern Colombia. Lectotype: BMNH 1913.10.1.6. Paralectotypes: BMNH 1910.1.11.170 (1), 1913.10.1.7 (1). Lectotype selected by Harold & Vari 1994:26 [ref. 21969]. •Synonym of Creagrutus affinis Steindachner 1880 -- (Harold & Vari 1994:23 [ref. 21969], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:121 [ref. 27061]). •Valid as Creagrutus leuciscus Regan 1913 -- (Román-Valencia & Cala 1997:145 [ref. 30645], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Albornoz-Garzón et al. 2018:352 [ref. 36150], Ardila Rodríguez 2020:12 [ref. 37904], Ardila Rodríguez 2021:16 [ref. 38586], Ardila Rodríguez 2023:12 [ref. 40275]). Current status: Valid as Creagrutus leuciscus Regan 1913. Stevardiidae: Creagrutinae. Distribution: South America: Río San Juan basin, Chocó, Colombia. Habitat: freshwater.

londonoi, Creagrutus Fowler [H. W.] 1945:3, Figs. 1-3 [Notulae Naturae (Philadelphia) No. 158; ref. 13140] Honda, Río Magdalena basin, Colombia. Holotype: ANSP 71610. Paratypes: ANSP 71611-18 (8), 71619 (1, c&s), 71620 (1). Type catalog: Böhlke 1984:50 [ref. 13621]. •Synonym of Creagrutus affinis Steindachner 1880 -- (Harold & Vari 1994:23 [ref. 21969], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:121 [ref. 27061], Román-Valencia & Cala 1997:150 [ref. 30645]). Current status: Synonym of Creagrutus affinis Steindachner 1880. Stevardiidae: Creagrutinae. Habitat: freshwater.

machadoi, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:134, Fig. 54 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Upper Río Caura, Caño Yumucukenã, within 4 kilometers its mouth, Departmento Cedeno, Bolivar, Venezuela. Holotype: MCNG 18852. Paratypes: MCNG 18896 (3), 41717 (6); USNM 359641 (2). •Valid as Creagrutus machadoi Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:123-124 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:57 [ref. 35633], Taphorn et al. 2022:14 [ref. 41297] as cf. machadoi). Current status: Valid as Creagrutus machadoi Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: Orinoco River basin (Venezuela and Colombia). Habitat: freshwater.

maculosus, Creagrutus Román-Valencia [C.], García-Alzate [C. A.], Ruiz-C. [R. I.] & Taphorn [D. C.] 2010:88, Figs. 1-8 [Ichthyological Exploration of Freshwaters v. 21 (no. 1); ref. 30781] Guadualito Creek drainage, 3°05'34"N, 73°51'53"W, Buenos Aires village, Vista Hermosa municípality, Güejar River, Meta State, Colombia. Holotype: IUQ 2506. Paratypes: many at AUM, IUQ. •Valid as Creagrutus maculosus Román-Valencia, García-Alzate, Ruiz-C. & Taphorn 2010 -- (Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Albornoz-Garzón et al. 2018:352 [ref. 36150]). Current status: Valid as Creagrutus maculosus Román-Valencia, García-Alzate, Ruiz-C. & Taphorn 2010. Stevardiidae: Creagrutinae. Distribution: Orinoco River basin, Colombia. Habitat: freshwater.

magdalenae, Creagrutus Eigenmann [C. H.] 1913:8 [Indiana University Studies No. 18; ref. 1229] Girardot, Colombia. Holotype: FMNH 56088 [ex CM 4880]. Paratypes: (over 100, CM and IU) CAS 41641 [ex IU 12722] (18), 60056-57 [ex IU 12725-26] (6, 8); FMNH 56089 (20), 56092-93 (7, 9), 69730 (72). •Valid as Creagrutus magdalenae Eigenmann 1913 -- (Harold & Vari 1994:16 [ref. 21969], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:124 [ref. 27061], Román-Valencia & Cala 1997:148 [ref. 30645], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Albornoz-Garzón et al. 2018:348 [ref. 36150], Ardila Rodríguez 2020:12 [ref. 37904], Ardila Rodríguez 2021:15 [ref. 38586], Ardila Rodríguez 2023:11 [ref. 40275]). Current status: Valid as Creagrutus magdalenae Eigenmann 1913. Stevardiidae: Creagrutinae. Distribution: South America: Magdalena and lower Cauca River basins, Colombia. Habitat: freshwater.

magoi, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:137, Fig. 55 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Chaviripa, where crossed by bridge on road from Caicara to San Fernado de Atabapo, about 7°11'N, 66°18'W, Bolivar, Venezuela. Holotype: UF 80477. Paratypes: ANSP 159834-35 (10, 10), 161405 (1); MBUCV V-17787 (8); UF 11079 (6), 80423 (5); USNM 353305-06 (2, 2), 353863 (1), 353864 (2, 1 c&s), 355118 (2). •Valid as Creagrutus magoi Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:124 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], see DoNascimiento et al. 2017:[12] [ref. 35679]). Current status: Valid as Creagrutus magoi Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Chaviripa and Parguaza River basins, right bank tributaries to the Orinoco River, Venezuela. Habitat: freshwater.

manu, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:140, Fig. 56 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Manu basin, Parque Nacional Manu, Quebrada Agua Clara, first major quebrada on trial 1 leading from Pakitza, about 11°57'S, 71°17'W, Provincia Manu, Madre de Dios, Peru. Holotype: MUSM 8867. Paratypes: MUSM 8868 (1); USNM 303067 (2), 313476 (1), 326903 (1, c&s). •Valid as Creagrutus manu Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:124 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus manu Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Upper Manu River basin in southeastern Peru. Habitat: freshwater.

maracaiboensis, Creagrutops Schultz [L. P.] 1944:327, Figs. 45-46 [Proceedings of the United States National Museum v. 95 (no. 3181); ref. 3960] Río Negro below mouth of Río Yasa, Lake Maracaibo basin, Venezuela. Holotype: USNM 121531. Paratypes: USNM 121532 (orig. 48, now 42, 1 c&s), UMMZ 145042 [ex USNM 121532] (2), MCZ 37280 [ex USNM 121532] (2), MHNG [ex USNM 121532] (1). Type catalog: Vari & Howe 1991:15 [ref. 19022]. •Valid as Creagrutus maracaiboensis (Schultz 1944) -- (Harold & Vari 1994:10 [ref. 21969], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:124 [ref. 27061], Román-Valencia & Cala 1997:145 [ref. 30645], Harold & Salcedo 2010:382 [ref. 30692], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Albornoz-Garzón et al. 2018:348 [ref. 36150]). Current status: Valid as Creagrutus maracaiboensis (Schultz 1944). Stevardiidae: Creagrutinae. Distribution: Lake Maracaibo and Río Rancheria basins, Venezuela and Colombia. Habitat: freshwater.

mariposa, Creagrutus Ardila Rodríguez [C. A.] 2021:5, Figs. 1, 3A, 5A, 6A, 8A, 9A [Barranquilla, Departamento del Atlántico No. 75; ref. 38586] Quebrada La Ruitoca (En Pozo Azul), Vereda del Valle de Ruitoque, afluente del río de Oro, cuenca alta del río Lebrija, cuenca del río Magdalena, municipio de Floridablanca, Departamento de Santander, Colombia. Holotype: CAR 877. Paratypes: CAR. •Valid as Creagrutus mariposa Ardila Rodríguez 2021 -- (Ardila Rodríguez 2023:12 [ref. 40275]). Current status: Valid as Creagrutus mariposa Ardila Rodríguez 2021. Stevardiidae: Creagrutinae. Distribution: South America: Río de Oro drainage, upper Magdalena basin, Departamento de Santander, Colombia. Habitat: freshwater.

martinezi, Creagrutus Ardila Rodríguez [C. A.] 2021:12, Figs. 2C, 4B, 5D, 7B, 8D, 9D [Barranquilla, Departamento del Atlántico No. 75; ref. 38586] Quebrada La Salitre, cuenca del río Magdalena, Municipio de San Vicente de Chucurí, Departamento de Santander, Colombia. Holotype: CAR 905. Paratypes: CAR 13 (4), 885 (9). •Valid as Creagrutus martinezi Ardila Rodríguez 2021 -- (Ardila Rodríguez 2023:12 [ref. 40275]). Current status: Valid as Creagrutus martinezi Ardila Rodríguez 2021. Stevardiidae: Creagrutinae. Distribution: South America: La Salitre stream, upper Magdalena basin, Departamento de Santander, Colombia. Habitat: freshwater.

maxillaris, Creagrudite Myers [G. S.] 1927:118 [Bulletin of the Museum of Comparative Zoology v. 68 (no. 3); ref. 3096] Sandbank on the Colombian border at Cucuhy, Rio Negro, Brazil. Holotype: CAS 30419 [ex IU 17682]. Paratypes: CAS 30420 [ex IU 17684] (2), CAS-SU 18071 [ex IU 17683] (1), MCZ 31572 (1). Type catalog: Böhlke 1953:23 [ref. 12291]. •Valid as Creagrutus maxillaris (Myers 1927) -- (Vari & Harold 1998:256 [ref. 23845], Vari & Harold 2001:142 [ref. 25842], Vari & Harold in Reis et al. 2003:124 [ref. 27061], Dagosta & Pastana 2014:578 [ref. 33181], Mirande et al. 2013:763 [ref. 33366], Sarmiento et al. 2014:184 [ref. 35004], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Albornoz-Garzón et al. 2018:352 [ref. 36150], Flausino & Lima 2019:7 [ref. 36726], Urbano-Bonilla et al. 2024:137 [ref. 42338]). Current status: Valid as Creagrutus maxillaris (Myers 1927). Stevardiidae: Creagrutinae. Distribution: South America: Orinoco River basin and rivers draining the Guiana Shield, upper Negro River in Venezuela, and other localities (Brazil, Colombia, Bolivia and Venezuela). Habitat: freshwater.

melanzonus, Creagrutus Eigenmann [C. H.] 1909:30 [Annals of the Carnegie Museum v. 6 (no. 1); ref. 1222] Crab Falls, Essequibo River, Guyana. Holotype: FMNH 52705 [ex CM 1067]. Paratypes: (5, CM and IU) CAS 30421-22 [ex IU 11753-54] (1, 2), FMNH 51706-07 (1, 1). Type catalog: Henn 1928:63 [ref. 12305], Ibarra & Stewart 1987:28 [ref. 12367]. •Valid as Creagrutus melanzonus Eigenmann 1909 -- (Planquette et al. 1996:324 [ref. 22818], Vari & Harold 2001:146 [ref. 25842], Vari & Harold in Reis et al. 2003:124 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], see DoNascimiento et al. 2017:[12] [ref. 35679], Albornoz-Garzón et al. 2018:352 [ref. 36150], Borstein et al. 2022:16 [ref. 39507], Taphorn et al. 2022:14 [ref. 41297]). Current status: Valid as Creagrutus melanzonus Eigenmann 1909. Stevardiidae: Creagrutinae. Distribution: South America: Cuyani River of eastern Venezuela to Sinnamary River basin French Guiana (French Guiana, Guyana, Suriname and Venezuela). Habitat: freshwater.

melasma, Creagrutus Vari [R. P.], Harold [A. S.] & Taphorn [D. C.] 1994:91, Fig. 1 [Proceedings of the Biological Society of Washington v. 107 (no. 1); ref. 21268] Río Orituco, first bridge along road from Santa Teresa to Altagracia, Parque Nacional Guatopo, Guarico, Venezuela. Holotype: MBUCV-V-22198. Paratypes: ANSP 134171 (48); CAS 79622 (10); INHS 60021 (10); MBUCV-V-21257 (74 not 56), V-24020 (2); MCNG 15354 (56), 24622 (34); MHNLS 503 (6); USNM 219615-6 (3, 1). Type catalog: Sabaj et al. 1997:269 [ref. 23225], Lasso et al. 1997:42 [ref. 23639], Provenzano et al. 1998:6 [ref. 24196]. •Valid as Creagrutus melasma Vari, Harold & Taphorn 1994 -- (Vari & Harold 2001:149 [ref. 25842], Vari & Harold in Reis et al. 2003:124 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento 2016:529 [ref. 34727], DoNascimiento et al. 2017:58 [ref. 35633]). Current status: Valid as Creagrutus melasma Vari, Harold & Taphorn 1994. Stevardiidae: Creagrutinae. Distribution: Orinoco River basin and Tuy and Neveri rivers of Caribbean Sea versant, Venezuela and Colombia. Habitat: freshwater.

menezesi, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:153, Figs. 63-64 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Rio Tocantins basin, Ribeirão Paranoa do Meio, at road crossing 11 air kilometers north northeast of Formosa, about 15°25'S, 47°18'W, Goiás, Brazil. Holotype: MZUSP 50544. Paratypes: MNRJ 12614 (8), 17333 (10), 17338 (11); MZUSP 50545 (8); USNM 292229 (11, 2 c&s), 350456 [in part ex MNRJ 16566] (10), 350456 [in part ex MNRJ 16562] (5). •Valid as Creagrutus menezesi Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:124 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Koerber et al. 2022:41 [ref. 39235]). Current status: Valid as Creagrutus menezesi Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: Tocantis River basin and tentatively Bronco and Negro River near mouth of Branco (northeastern Brazil). Habitat: freshwater.

meridionalis, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:157, Fig. 65 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Ribeirão Chiqueirão, tributary of Rio Jangada that drains into the Rio Cuiabá, about 21 kilometers west of Jangada on road to Barra do Bugres, Município de Jangada, Mato Brosso, Brazil. Holotype: MZUSP 50546. Paratypes: FML 2527 (1), MCP 18738 (13), MNHNP 2999 (13), MZUSP 50547 (12, 1 c&s), NRM 22619 (20), USNM 341363 (12, 2 c&s). •Valid as Creagrutus meridionalis Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:124 [ref. 27061], Baicere-Silva et al. 2011:381 [ref. 31395], Dagosta & Pastana 2014:581 [ref. 33181], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Koerber et al. 2017:33 [ref. 35267]). Current status: Valid as Creagrutus meridionalis Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Upper Paraguay River basin in Mato Grosso State and eastern tributaries to Paraguay River: Brazil and Paraguay. Habitat: freshwater.

metropolitana, Creagrutus Ardila Rodríguez [C. A.] 2024:8, Figs. 1, 2, 5A, 6A, 7A, 8A, 9A, 10 [Barranquilla, Departamento del Atlántico No. 99; ref. 41561] Río Sevilla, Sierra Nevada de Santa Marta, Departamento del Magdalena, Colombia. Holotype: CAR 1024. Paratypes: CAR 1007 (15), CAR 1077 (77, 5 at Museo de la Universidad del Atlántico). •Valid as Creagrutus metropolitana Ardila Rodríguez 2024. Current status: Valid as Creagrutus metropolitana Ardila Rodríguez 2024. Stevardiidae: Creagrutinae. Distribution: South America: Río Sevilla drainage, Departamento del Magdalena, Colombia. Habitat: freshwater.

molinus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:159, Fig. 66 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Rio Araguaia basin, riacho (small stream) tributary to Ribeirão Lajeadinho, Município de Alto Araguaia, about 17°19'S, 53°12'W, mato Grosso, Brazil. Holotype (unique): MZUSP 41461. •Valid as Creagrutus molinus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:124 [ref. 27061], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus molinus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Upper Araguaia River basin, Brazil. Habitat: freshwater.

mucipu, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:162, Fig. 68 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Rio Tocantins, at port of Rubao, 13°44'31"S, 48°08'29"W, Município de Minaçu, Goiás, Brazil. Holotype: MCP 19511. Paratypes: MNRJ 17337 (6), USNM 350449 (2). •Valid as Creagrutus mucipu Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:124 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Netto-Ferreira & Moreira 2018:255 [ref. 35759], Flausino & Lima 2019:6 [ref. 36726]). Current status: Valid as Creagrutus mucipu Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Upper Tocantins and Xingu River basins, Brazil. Habitat: freshwater.

muelleri, Leporinus Günther [A.] 1859:92 [Proceedings of the Zoological Society of London 1859 (pt 1); ref. 18209] Andes of western Ecuador. Lectotype: BMNH 1858.7.25.42 (82.5 mm SL). Paralectotypes: BMNH 1858.7.25.43-45 (4). Originally as mülleri, change mandatory. Lectotype selected by Vari & Harold 2001:165 [ref. 25842]. •Valid as Creagrutus muelleri (Günther 1859) -- (Ovchynnyk 1967:28 [ref. 37559] as mulleri, Vari & Harold 2001:165 [ref. 25842], Vari & Harold in Reis et al. 2003:124 [ref. 27061], Harold & Salcedo 2010:382 [ref. 30692], Mirande et al. 2013:763 [ref. 33366], Barriga Salazar 2014:107 [ref. 35745], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298], Chuctaya et al. 2022:19 [ref. 42137]). Current status: Valid as Creagrutus muelleri (Günther 1859). Stevardiidae: Creagrutinae. Distribution: South America: rivers of eastern slope of the Andean piedmont (Ecuador and Peru). IUCN (2016): Least Concern. Habitat: freshwater.

narvaezi, Creagrutus Ardila Rodríguez [C. A.] 2023:12, Figs. 2A, 5C, 8A, 9C, 10B [Barranquilla, Departamento del Atlántico No. 88; ref. 40588] Río Fundación, Sierra Nevada de Santa Marta, Municipio de Fundación, Departamento del Magdalena, Colombia. Holotype: CAR 970. Paratypes: CAR. •Valid as Creagrutus narvaezi Ardila Rodríguez 2023. Current status: Valid as Creagrutus narvaezi Ardila Rodríguez 2023. Stevardiidae: Creagrutinae. Distribution: South America: Río Fundación drainage, Departamento del Magdalena, Colombia. Habitat: freshwater.

nasutus, Creagrutus Günther [A.] 1876:400 [Annals and Magazine of Natural History (Series 4) v. 17 (no. 101) (art. 43); ref. 2007] Monterico, Peru. Lectotype: BMNH 1875.10.14.26 (78.9 mm SL). Paralectotypes: BMNH 1875.10.14.27-33 (?7). Lectotype selected by Vari & Harold 2001:42 [ref. 25842]. •Synonym of Creagrutus peruanus (Steindachner 1876) -- (Vari & Harold 2001:181 [ref. 25842]). Current status: Synonym of Creagrutus peruanus (Steindachner 1876). Stevardiidae: Creagrutinae. Habitat: freshwater.

nigrostigmatus, Creagrutus Dahl [G.] 1960:353 [Caldasia v. 8 (no. 38); ref. 9309] Arroyo (Caño) Pechilín, a small stream draining into the Caribbean Sea west of the Río Magdalena, northern Colombia. Neotype: ICNMHN 989. Possible paratypes: CAS-SU 49491 (2). Type catalog: Mojica & Agudelo-Zamora 2018:15 [ref. 36127]. Original type series (44 specimens) lost. Neotype designated by Harold & Vari 1994:10 [ref. 21969]. •Valid as Creagrutus nigrostigmatus Dahl 1960 -- (Harold & Vari 1994:7 [ref. 21969], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:124-125 [ref. 27061], Román-Valencia & Cala 1997:145 [ref. 30645], Harold & Salcedo 2010:382 [ref. 30692], Román-Valencia et al. 2010:95 [ref. 30781], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Albornoz-Garzón et al. 2018:348 [ref. 36150]). Current status: Valid as Creagrutus nigrostigmatus Dahl 1960. Stevardiidae: Creagrutinae. Distribution: Caño Pechilín, Caño Zaragocilla and other Caribbean slope rivers, Colombia. Habitat: freshwater.

nigrotaeniatus, Creagrutus Dagosta [F. C. P.] & Pastana [M. N. L.] 2014:572, Figs. 1- 5, 8 [Zootaxa 3765 (no. 6); ref. 33181] Mata Grosso State, Sepezal Municipality, rio Juruena, Bacia do rio Tapajós, 12°50'59"S, 58°55'36"W. Brazil. Holotype: LIRP 8967. Paratypes: LIRP 8966 (1 c&s), 8968 (1); MZUSP 113740 (1). •Valid as Creagrutus nigrotaeniatus Dagosta & Pastana 2014 -- (Thomaz et al. 2015:Add. 5 [ref. 33893], Netto-Ferreira & Vari 2017:368 [ref. 35453], Netto-Ferreira & Moreira 2018:255 [ref. 35759]). Current status: Valid as Creagrutus nigrotaeniatus Dagosta & Pastana 2014. Stevardiidae: Creagrutinae. Distribution: Rio Juruena, Bacia do rio Tapajós, Brazil. Habitat: freshwater.

notropoides, Creagrutus Meek [S. E.] & Hildebrand [S. F.] 1912:68 [Field Museum of Natural History, Publications, Zoölogical Series v. 10 (no. 6); ref. 10934] Río Indio, upper Chagres River basin, Panama. Holotype: FMNH 7574. Paratypes: CAS 44208 [IU 13996] (1), FMNH 12539-40 (39, 1), 12541-54 (14), 12555-58 (5, 4, 1, 1), 32021-22 (1, 1). Type catalog: Ibarra & Stewart 1987:28 [ref. 12367]. •Synonym of Creagrutus affinis Steindachner 1880 -- (Harold & Vari 1994:23 [ref. 21969], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:121 [ref. 27061], Román-Valencia & Cala 1997:150 [ref. 30645]). Current status: Synonym of Creagrutus affinis Steindachner 1880. Stevardiidae: Creagrutinae. Habitat: freshwater.

occidaneus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:167, Fig. 70 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Beach along Río Manu close to mouth of Quebrada Pachija, about 11°57'S, 71°17'W, Parque Nacional Manu, Pakitza, Provincia Manu, Madre do Dios, Peru. Holotype: MUSM 8869. Paratypes: ANSP 143765 (1); MUSM 8870-71 (4, 5); USNM 295621 (9), 303065 (1), 326920 (7, 2 c&s). •Valid as Creagrutus occidaneus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:125 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298], Chuctaya et al. 2022:20 [ref. 42137]). Current status: Valid as Creagrutus occidaneus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: eastern Peru, and Department of Ucayali, Peru, and upper Purus River (Brazil and Peru). IUCN (2023): Least Concern. Habitat: freshwater.

ortegai, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:170, Fig. 71 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Banda de Shilcayo, upper portions of Quebrada Choclin, tributary to Río Huallaga, near town of Shapaja, Provincia San Martin, San Martin, Peru. Holotype: MUSM 7498. Paratypes: MUSM 2948 (1), 5678 (4), 6552 (8), 6571 (2), 7499 (2), 7538-39 (2, 3); USNM 340969-71 (2, 3, 4), 340972 (8, 2 c&s). •Valid as Creagrutus ortegai Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:125 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298], Chuctaya et al. 2022:20 [ref. 42137]). Current status: Valid as Creagrutus ortegai Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: Huallaga River basin and headwaters of the Aguaytia River (Peru). IUCN (2016): Least Concern. Habitat: freshwater.

ouranonastes, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:173, Fig. 72 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Chalhuanca, about 14°01'S, 73°11'W, Provincia Aymaraes, Apurimac, Peru. Holotype: MUSM 8872. Paratypes: FMNH 41801 (14); MUSM 6574 (5); USNM 340988 (4, 1 c&s), 340989 (1). •Valid as Creagrutus ouranonastes Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:125 [ref. 27061], Harold & Salcedo 2010:382 [ref. 30692], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Chuctaya et al. 2022:20 [ref. 42137]). Current status: Valid as Creagrutus ouranonastes Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: upper Apurimac River basin (Peru). IUCN (2016): Least Concern. Habitat: freshwater.

paraguayensis, Creagrutus Mahnert [V.] & Géry [J.] 1988:5, Figs. 4-5; Pl. 3 [Revue française d'Aquariologie Herpétologie v. 15 (no. 1); ref. 6407] Río Azotey at Cororo, near Río Ypané mouth, Concepción Province, Paraguay. Holotype: MHNG 2386.01. Paratypes: MHNG 2386-2-3 (3, 1 stained). Type catalog: Weber 1998:7 [ref. 32707]. •Valid as Creagrutus paraguayensis Mahnert & Géry 1988 -- (Vari & Harold 2001:175 [ref. 25842], Vari & Harold in Reis et al. 2003:125 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Koerber et al. 2017:33 [ref. 35267]). Current status: Valid as Creagrutus paraguayensis Mahnert & Géry 1988. Stevardiidae: Creagrutinae. Distribution: Paraguay River basin, central and northern Paraguay. Habitat: freshwater.

paralacus, Creagrutus Harold [A. S.] & Vari [R. P.] 1994:13, Fig. 8 [Smithsonian Contributions to Zoology No. 551; ref. 21969] Río Táchira, Río Catatumbo basin, 7 kilometers north of San Antonio, Estado Táchira, Venezuela. Holotype: USNM 121504. Paratypes: MBUCV-V-23300 to 01 (2, 10); USNM 121499 (10), 324379 (3). Type catalog: Provenzano et al. 1998:6-7 [ref. 24196]. •Valid as Creagrutus paralacus Harold & Vari 1994 -- (Galvis et al. 1997:45 [ref. 23651], Vari & Harold 2001:2 [ref. 25842], Vari & Harold in Reis et al. 2003:125 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Albornoz-Garzón et al. 2018:348 [ref. 36150], Ardila Rodríguez 2023:11 [ref. 40275]). Current status: Valid as Creagrutus paralacus Harold & Vari 1994. Stevardiidae: Creagrutinae. Distribution: South America: southern and eastern tributaries to Lake Maracaibo, Venezuela and Colombia. Habitat: freshwater.

pearsoni, Creagrutus Mahnert [V.] & Géry [J.] 1988:5 [Revue française d'Aquariologie Herpétologie v. 15 (no. 1); ref. 6407] Río Beni at Villa Bella, Amazon system, Bolivia. Lectotype: CAS 62272 [ex IU 17317]. Paralectotypes: (10) CAS 87389 [ex CAS 62272] (6); UMML 66495 (3). Replacement name for Piabina beni Pearson 1924, secondarily preoccupied by Creagrutus beni Eignemann 1911. Lectotype selected by Vari & Harold 2001:180 [ref. 25842]. •Valid as Creagrutus pearsoni Mahnert & Géry 1988 -- (Vari & Harold 1998:256 [ref. 23845], Vari & Harold 2001:178 [ref. 25842], Vari & Harold in Reis et al. 2003:125 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Sarmiento et al. 2014:184 [ref. 35004], Thomaz et al. 2015:Add. 5 [ref. 33893], Careaga et al. 2020:75 [ref. 37774]). Replacement Name. Current status: Valid as Creagrutus pearsoni Mahnert & Géry 1988. Stevardiidae: Creagrutinae. Distribution: South America: upper Madeira River (northeastern Bolivia, southeastern Peru). Habitat: freshwater.

peruana, Piabina Steindachner [F.] 1876:596 [46] [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 72 (1. Abth.); ref. 18772] Monterico, Peru. Lectotype: NMW 19852 (87.0 mm SL). Paralectotypes: NMW 19853-79 (29). On p. 46 of separate. Lectotype selected by Vari & Harold 2001:182 [ref. 25842]. •Valid as Creagrutus peruanus (Steindachner 1876) -- (Ortega & Vari 1986:8 [ref. 9708], Vari & Harold 2001:181 [ref. 25842], Vari & Harold in Reis et al. 2003:125 [ref. 27061], Harold & Salcedo 2010:382 [ref. 30692], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298], Chuctaya et al. 2022:20 [ref. 42137]). Current status: Valid as Creagrutus peruanus (Steindachner 1876). Stevardiidae: Creagrutinae. Distribution: South America: Apurimac and Urubamba River basins (Peru). IUCN (2016): Least Concern. Habitat: freshwater.

petilus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:184, Fig. 78 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Rio Marco Rondon, Pimenta Bueno, about 11°29'S, 61°12'W, Rondônia, Brazil. Holotype: MNRJ 14807. Paratypes: MNRJ 13368 (1), 13370 (12); USNM 340957 (8, 2 c&s). •Valid as Creagrutus petilus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:125 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus petilus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Upper Machado River, right bank tributary of Madeira River, Brazil. Habitat: freshwater.

phasma, Creagrutus Myers [G. S.] 1927:117 [Bulletin of the Museum of Comparative Zoology v. 68 (no. 3); ref. 3096] Mouth of Curamuni, Río Cassiquiare, Venezuela. Lectotype: CAS 44210 [ex IU 17681]. Paralectotypes: CAS 209191 [ex CAS 44210 and IU 17681] (1). Lectotype selected by Vari & Harold 2001:188 [ref. 25842]. •Valid as Creagrutus phasma Myers 1927 -- (Vari & Harold 2001:186 [ref. 25842], Vari & Harold in Reis et al. 2003:125 [ref. 27061], Román-Valencia & Cala 1997:149 [ref. 30645], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Albornoz-Garzón et al. 2018:352 [ref. 36150]). Current status: Valid as Creagrutus phasma Myers 1927. Stevardiidae: Creagrutinae. Distribution: Upper Negro River basin north to Orinoco River basin: Brazil, Colombia and Venezuela. Habitat: freshwater.

pila, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:189, Fig. 80 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Huacamayo, kilometer 155 on highway (Carratera Federico Basadre) from Pucallpa to Tingo Maria, Provincia Padre Abad, Ucayali, Peru. Holotype: MUSM 8874. Paratypes: MUSM 2956 (3), 5240 (3), 8875-77 (6, 1, 4); USNM 341364-66 (3, 5, 3), 341367 (4, 2 c&s). •Valid as Creagrutus pila Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:125 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298], Chuctaya et al. 2022:20 [ref. 42137]). Current status: Valid as Creagrutus pila Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: smaller rivers draining into Aguaytia River, left bank tributary of Ucayali River (Peru). IUCN (2016): Least Concern. Habitat: freshwater.

planquettei, Creagrutus Géry [J.] & Renno [J.-F.] 1989:1, Figs. 1-5 [Revue française d'Aquariologie Herpétologie v. 16 (no. 1); ref. 13420] Japigny Creek, Arataye, tributary of Approuague, upstream of Pierrette, French Guiana. Holotype: MNHN 1989-0031. Paratypes: (8) MNHN 1989-0036 (4), MTD F 27519 [es Géry coll.] (1). Type catalog: Zarske 2003:17 [ref. 27474]. •Valid as Creagrutus planquettei Géry & Renno 1989 -- (Géry et al. 1991:45 [ref. 21021], Planquette et al. 1996:326 [ref. 22818], Vari & Harold 2001:191 [ref. 25842], Vari & Harold in Reis et al. 2003:125 [ref. 27061], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus planquettei Géry & Renno 1989. Stevardiidae: Creagrutinae. Distribution: Middle and upper Approuague River basin, French Guiana. Habitat: freshwater.

provenzanoi, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:195, Fig. 84 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Upper Río Cataniapo basin, small caño above Saramä Sota, Amazonas, Venezuela. Holotype: MBUCV V-14392. Paratypes: ANSP 165530 (1); MBUCV V-21657 (1), V-29070-71 (5, 12); USNM 207229 (3), 355117 (2), 355119 (5, 2 c&s). •Valid as Creagrutus provenzanoi Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:125 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus provenzanoi Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Cataniapo River basin, right bank tributary of Orinoco River, Venezuela. Habitat: freshwater.

runa, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:198, Fig. 85 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Rio Negro, Paraná do Jacaré, about 0°30's, 66°30'W, amazonas, Brazil. Holotype: MZUSP 29888. Paratypes: MZUSP 55473 (30); USNM 359622 (5), 270338 (2). •Valid as Creagrutus runa Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633]). Current status: Valid as Creagrutus runa Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Amazon River basin: Brazil, Colombia and Venezuela. Habitat: freshwater.

saxatilis, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:200, Fig. 86 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Rio Indaial, left bank tributary Of rio Maranhão, upper Rio Tocantins basin, 14°12'S, 48°37'W, Niquelândia, Goiás, Brazil. Holotype: MNRJ 14544. Paratypes: MCP 15977 (2); MNRJ 12590 (3), 12641 (3), 17335 (3); USNM (6, 1 c&s). •Valid as Creagrutus saxatilis Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus saxatilis Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Upper Tocantins River basin, Brazil. Habitat: freshwater.

seductus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:202, Fig. 87 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Upper Rio Araguaia basin, Córrego Fundo, Município de Barra do Garças, about 15°53'S, 52°15'W, Mato Grosso, Brazil. Holotype: MZUSP 51026 [in part ex ICLMA 229]. Paratypes: MZUSP 51027 [in part ex ICLMA 229] (1), 51028 [in part ex ICLMA 190] (3), 51029 (9); USNM 342231 (9, 1 c&s), 342232 [in part ex ICLMA 190] (3), 342233 [in part ex ICLMA 229] (1). •Valid as Creagrutus seductus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Dagosta & Pastana 2014:581 [ref. 33181], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus seductus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Upper Araguaia River basin, Brazil. Habitat: freshwater.

simus, Creagrutus Meek [S. E.] & Hildebrand [S. F.] 1913:85 [Field Museum of Natural History, Publications, Zoölogical Series v. 10 (no. 8); ref. 12295] Río Cupe, Cituro, Panama. Holotype: FMNH 7591. Paratypes: CAS 44212 [ex IU 13997] (1), 44213 [ex IU 13998] (2); FMNH 12562-603 (42), 12604-09 (13, 18, 12, 18, 38, 85), 32023-28 (6); USNM 78570 (orig. 42, now 41). Type catalog: Ibarra & Stewart 1987:28 [ref. 12367], Vari & Howe 1991:16 [ref. 19022]. Only holotype cited, but description based on more than one specimen. •Synonym of Creagrutus affinis Steindachner 1880 -- (Harold & Vari 1994:23 [ref. 21969], Vari & Harold 2001:3 [ref. 25842], Vari & Harold in Reis et al. 2003:121 [ref. 27061], Román-Valencia & Cala 1997:150 [ref. 30645]). Current status: Synonym of Creagrutus affinis Steindachner 1880. Stevardiidae: Creagrutinae. Habitat: freshwater.

taphorni, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:205, Fig. 88 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Orituco, Parque Nacional Guatopo, first bridge along road from Santa Teresa to Altagracia, Guarico, Venezuela. Holotype: MBUCV V-29288. Paratypes: MBUCV V-21032 (10), V-21244 (18), V-29289 (5); MCNG 18736 (36), 28752 (4); USNM 357730 (2, 1 c&s), 357784 (6). •Valid as Creagrutus taphorni Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Mirande 2010:544 [ref. 31006] as cf. taphorni, Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Urbano-Bonilla et al. 2018:88 [ref. 35773] as cf. taphorni). Current status: Valid as Creagrutus taphorni Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Largely in Orinoco River and Tuy River basins of Caribbean versant, Venezuela and Colombia. Habitat: freshwater.

tuyuka, Creagrutus Vari [R. P.] & Lima [F. C. T.] 2003:583, Fig. 1 [Copeia 2003 (no. 3); ref. 27130] Rio Tiquié, comunidade de Cachoeira Comprida, beach below the cachoeira, 0°15'44.0"N, 70°01'05.2"W, Amazonas, Brazil. Holotype: MZUSP 65679. Paratypes: INPA 18658 (2); MZUSP 64689 (2), 78519 (6, 1 c&s); USNM 369731 (2). •Valid as Creagrutus tuyuka Vari & Lima 2003 -- (Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Flausino & Lima 2019:7 [ref. 36726]). Current status: Valid as Creagrutus tuyuka Vari & Lima 2003. Stevardiidae: Creagrutinae. Distribution: Amazon River basin: Brazil and Colombia. Habitat: freshwater.

ungulus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:209, Fig. 90 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Quebrada Soga, tributary Of Río Madre de Dios, 1 kilometer upstream from Erika, opposite Salvacion, about 12°53'S, 71°12'W, Provincia Manu, Madre de Dios, Peru. Holotype: MUSM 8878. Paratypes: MNRJ 14481 (6); MUSM 8879-80 (7, 4); USNM 302788 (3), 303066 (10, 3 c&s). •Valid as Creagrutus ungulus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], Meza-Vargas et al. 2021:18 [ref. 39298]). Current status: Valid as Creagrutus ungulus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: Madre de Dios River basin (Peru). Habitat: freshwater.

varii, Creagrutus Ribeiro [A. C.], Benine [R. C.] & Figueiredo [C. A.] 2004:598, Figs. 1-10 [Journal of Fish Biology v. 64 (no. 3); ref. 27648] Córrego da Prata, 17°47'26"S, 47°35'48"W, Rio São Marcos basin, tributary of Rio Paranaíba basin, Munic. de Catalão, Goiás, Brazil. Holotype: LIRP 4342. Paratypes: DZSJRP 5424 (8); LIRP 3358 (32, 2 c&s), 3359 (7), 3625 (14); MCP 31989 (20); MZUSP 81069 (20); ZUEC 6165 (8). Type catalog: Andrade et al. 2017:438 [ref. 35588]. •Valid as Creagrutus varii Ribeiro, Benine & Figueiredo 2004 -- (Harold & Salcedo 2010:382 [ref. 30692], Dagosta & Pastana 2014:581 [ref. 33181], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus varii Ribeiro, Benine & Figueiredo 2004. Stevardiidae: Creagrutinae. Distribution: Upper Paranaíba River basin, central Brazil. Habitat: freshwater.

veruina, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:213, Fig. 91 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Cataniapo, 200 meters above Las Pavas, Amazonas, Venezuela. Holotype: MBUCV V-29072. Paratypes: MBUCV V-14638 (2), USNM 355120 (2). •Valid as Creagrutus veruina Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus veruina Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Cataniapo River basin, east bank tributary of middle Oirnoco River, Venezuela. Habitat: freshwater.

vexillapinnus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:215, Fig. 92 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Rio Negro, Cachoeira de São Gabriel, 0°08'S, 67°05'W, Amazonas, Brazil. Holotype: MZUSP 29894. Paratypes: MZUSP 29893 (1), 29896 (1), 31220 (1); USNM 341360 (1), 341361 (1 c&s). •Valid as Creagrutus vexillapinnus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893], DoNascimiento et al. 2017:58 [ref. 35633], Flausino & Lima 2019:7 [ref. 36726], Urbano-Bonilla et al. 2024:137 [ref. 42338]). Current status: Valid as Creagrutus vexillapinnus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: South America: upper Negro, Vaupés and upper Orinoco River basins (Brazil, Colombia and Venezuela). Habitat: freshwater.

xiphos, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:218, Fig. 93 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Sand bank along Río Mato, 7°02'N, 65°13'W, Bolivar, Venezuela. Holotype: ANSP 165815. Paratypes: ANSP 177717 (3). •Valid as Creagrutus xiphos Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus xiphos Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Mato River in Caura River basin, Orinoco River drainage, Venezuela. Habitat: freshwater.

yanatili, Creagrutus Harold [A. S.] & Salcedo [N. J.] 2010:378, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 20 (no. 4) [Dec. 2009]; ref. 30692] Río Urubamba (Ucayali River drainage), vicinity of Chaullay, 13°00'48"S 072°38'31"W, elevation 1155 m, Cuzco Department, Peru. Holotype: MUSM 24082. Paratypes: ANSP 180423 (3), MUSM 24083 (2). •Valid as Creagrutus yanatili Harold & Salcedo 2010 -- (Thomaz et al. 2015:Add. 5 [ref. 33893], Chuctaya et al. 2022:20 [ref. 42137]). Current status: Valid as Creagrutus yanatili Harold & Salcedo 2010. Stevardiidae: Creagrutinae. Distribution: South America: Río Ucayali drainage (southeastern Peru). IUCN (2016): Data Deficient. Habitat: freshwater.

yudja, Creagrutus Netto-Ferreira [A. L.] & Moreira [C. R.] 2018:251, FIg. 1 [Zootaxa 4375 (no. 2); ref. 35759] Praia do Levi, Rio Iriri, Rio Xingu basin, Altamira, Pará, Brazil, 3°50'37"S, 52°44'03"W. Holotype: MNRJ 48662. Paratypes: INPA, MNRJ, MPEG, MZUSP. •Valid as Creagrutus yudja Netto-Ferreira & Moreira 2018. Current status: Valid as Creagrutus yudja Netto-Ferreira & Moreira 2018. Stevardiidae: Creagrutinae. Distribution: South America: lower Rio Xingu basin, Pará State, Brazil. Habitat: freshwater.

zephyrus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:220, Fig. 94 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Río Casiquiare, playa and backwater about 2 kilometers downstream from mouth of Río Pamoni, 2°48'N, 65°57'W, Amazonas, Venezuela. Holotype: ANSP 161238. Paratypes: ANSP 161236-37 (10, 6), 177829 (2). •Valid as Creagrutus zephyrus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Barriga Salazar 2014:107 [ref. 35745] as cf. zephyrus, Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus zephyrus Vari & Harold 2001. Stevardiidae: Creagrutinae. Distribution: Central and upper Negro River basins: Brazil, Ecuador? and Venezuela. Habitat: freshwater.