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Eschmeyer's Catalog of Fishes
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Species in the genus Cyprinella: [ 65 ] records


alvarezdelvillari, Cyprinella Contreras-Balderas [S.] & Lozano-Vilano [M. de L.] 1994:898, Fig. 1 [Copeia 1994 (no. 4); ref. 22061] Balneario La Concha, 9 kilometers southwest of Peñón Blanco, Durango, Mexico. Holotype: UANL 8302. Paratypes: AMNH 47834 [ex UANL 8303 & 11131] (4); BMNH 1990.6.23.3-6 [ex UANL 8310] (4); CAS [ex UANL 8303 & 11131] (4), 54426 (1); IPN P4436 [ex UANL 8310] (4); TNHC 17459 [ex UANL 8310] (4); UANL 8303 (276, now 256), 8310 (192, now 168), 8315-16 (12, 1), 8266 (1), 11131 (2, ? now 0); TU 157269 [ex UANL 8310] (4); UALA [ex UANL 8310] 9115.01 (4); UMMZ [ex UANL 8303 & 11131] 214693 (8); UNAM [ex UANL 8303 & 11131] 3239 (4); USNM 308111 [ex UANL 8310] (4). Type catalog: Gilbert 1998:38 [ref. 23395], Contreras-Balderas et al. 1998:2-3 [ref. 23752]. •Valid as Cyprinella alvarezdelvillari Contreras-Balderas & Lozano-Vilano 1994 -- (Gilbert 1998:38 [ref. 23395], Nelson et al. 2004:68 [ref. 27807], Miller 2006:109 [ref. 28615], Scharpf 2005:12 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page et al. 2013:69 [ref. 32708], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella alvarezdelvillari Contreras-Balderas & Lozano-Vilano 1994. Leuciscidae: Pogonichthyinae. Distribution: Southern North America: tributary of the Nazas River, endorheic Río Penon Blanco system, from Ojo la Concha spring to 8 km downstream (Durango, Mexico) endemic. Habitat: freshwater.

analostana, Cyprinella Girard [C. F.] 1859:59 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 11; ref. 1821] Rock Creek, tributary of Potomac River, opposite Analostan Island (Theodore Roosevelt Island), Washington, D.C., U.S.A. Syntypes: apparently lost. Type catalog: Gilbert 1978:26 [ref. 7042], Gilbert 1998:40 [ref. 23395]. •Valid as Notropis analostanus (Girard 1859) -- (Gibbs 1963:520 [ref. 21327], Gilbert 1978:24 [ref. 7042], Lee et al. 1980:226 [ref. 22416], Smith 1986:176 [ref. 22186]). •Valid as Cyprinella analostana Girard 1859 -- (Mayden 1989:73 [ref. 12555], Page & Burr 1991:120 [ref. 18983], Jenkins & Burkhead 1994:361 [ref. 21581], Stauffer et al. 1995:68 [ref. 23376], Gilbert 1998:40 [ref. 23395], Fuller et al. 1999:62 [ref. 25838], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:12 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:237 [ref. 31215], Page et al. 2013:69 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella analostana Girard 1859. Leuciscidae: Pogonichthyinae. Distribution: North America: northeastern U.S.A. and southeastern Canada. Habitat: freshwater.

aurata, Moniana Girard [C. F.] 1856:200 [36] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Pinto Creek, 15-20 miles southeast of Del Rio, Kinney County, Texas, U.S.A. Syntypes: ANSP 2833-34 [ex USNM 118] (2); MCZ 1689 [ex USNM 118] (2); MNHN 0000-0378 [ex USNM 118] (2); USNM 118 (43 + 1 left arch), 125084 [ex USBF 100] (4). Type catalog: Böhlke 1984:69 [ref. 13621], Gilbert 1998:46 [ref. 23395]. On p. 36 of separate. •Synonym of Notropis proserpinus (Girard 1856) -- (Gilbert 1978:28 [ref. 7042]). •Synonym of Cyprinella proserpina (Girard 1856) -- (Mayden 1989:63 [ref. 12555], Gilbert 1998:46 [ref. 23395]). Current status: Synonym of Cyprinella proserpina (Girard 1856). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

beckwithi, Cyprinella Girard [C. F.] 1856:197 [33] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Sluices of Arkansas River near Fort Makee, Kansas, U.S.A. Syntypes: (9) ANSP 3053 (1, missing), MCZ 1801 (1), MNHN 0000-0374 (1, lost), USNM 135 (orig. 7, now 0, 3 pharyngeal arches as # 2714). Type catalog: Gilbert 1978:29 [ref. 7042], Böhlke 1984:70 [ref. 13621], Gilbert 1998:48 [ref. 23395]. On p. 33 of separate. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:29 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:49 [ref. 12555], Gilbert 1998:48 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

billingsiana, Cyprinella Cope [E. D.] 1871:439 [Preliminary report of the United States Geological Survey of Wyoming and portions of contiguous territories. Part 4 (art. 8); ref. 19655] Missouri River at St. Joseph, Buchanan County, Missouri, U.S.A. Lectotype: ANSP 2952. Paralectotypes: ANSP 2953-91 (39). Type catalog: Gilbert 1978:29 [ref. 7042], Böhlke 1984:70 [ref. 13621] as holotype and paratypes, Gilbert 1998:49 [ref. 23395]. Lectotype established by Fowler 1910:280 [ref. 19840]. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:29 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:50 [ref. 12555], Gilbert 1998:49 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

blairi, Notropis lutrensis Hubbs [C. L.] 1940:6 [Transactions of the Texas Academy of Sciencs v. 23 (1938-1939); ref. 10707] Monument Spring, tributary to Peña Colorado Creek, 12-13 miles south-southwest of Marathon, Texas, U.S.A., elevation 3700 feet. Holotype: UMMZ 104170. Paratypes: BU uncat. [ex UMMZ 127335] (3); UMMZ 104171 (15), 120330 (10), 127335 (22) Type catalog: Gilbert 1978:29-30 [ref. 7042], Gilbert 1998:50 [ref. 23395]. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:30 [ref. 7042]). •Now extinct, sometimes recognized as a valid subspecies. •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Matthews 1987:616 [ref. 6774], Mayden 1989:50 [ref. 12555], Gilbert 1998:50 [ref. 23395]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) but a valid extinct subspecies -- (Scharpf 2005:13 [ref. 28940]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

bocagrande, Notropis Chernoff [B.] & Miller [R. R.] 1982:514, Figs. 2-3 [Copeia 1982 (no. 3); ref. 8493] Ojo Solo, about 39 kilometers west of Villa Ahumada, Chihuahua, Mexico. Holotype: UMMZ 208249. Paratypes: UMMZ 208250 (28, now 27, 2 c&s), 208265 (5); USNM 226900 (4). Type catalog: Gilbert 1998:50 [ref. 23395]. •Valid as Cyprinella bocagrande (Chernoff & Miller 1982) -- (Mayden 1989:57 [ref. 12555], Mayden & Hillis 1990:1005 [ref. 19122], Espinosa Pérez et al. 1993:11 [ref. 22290], Gilbert 1998:50 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:12 [ref. 28940], Miller 2006:110 [ref. 28615], Schönhuth & Mayden 2009:78 [ref. 38584], Page et al. 2013:69 [ref. 32708], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella bocagrande (Chernoff & Miller 1982). Leuciscidae: Pogonichthyinae. Distribution: Southern North America: Ojo Solo, about 39 kilometers west of Villa Ahumada (Chihuahua, Mexico) endemic. Habitat: freshwater.

bubalinus, Leuciscus Baird [S. F.] & Girard [C. F.] 1853:391 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 6; ref. 165] Otter Creek, Arkansas [= tributary to North Fork of Red River, either Kiowa or Tillman County, southwestern Oklahoma], U.S.A. Holotype: USNM 125 (lost, 1 pair pharyngeal arches as #2798). Type catalog: Gilbert 1978:32 [ref. 7042], Gilbert 1998:52 [ref. 23395]. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:32 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:49 [ref. 12555], Gilbert 1998:52 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

caeruleus, Photogenis Jordan [D. S.] 1877:338 [Annals of the Lyceum of Natural History of New York v. 11 (nos 11-12) (art. 29); ref. 2373] Etowah River, Rome, Floyd County, Georgia, U.S.A. Lectotype: USNM 17883. Paralectotypes: ANSP 19839-41 (3, missing), BMNH 1880.1.21.10 (1), MCZ 24396 (1), MNHN A-1306 (1), USNM 20114 (2). Type catalog: Bertin & Estève 1948:66 [ref. 19574], Gilbert 1978:33 [ref. 7042], Böhlke 1984:71 [ref. 13621], Gilbert 1998:54 [ref. 23395]. Lectotype selected by Jordan & Evermann 1896:277 [ref. 2443] and by Gibbs in Gilbert 1978:33 [ref. 7042]. •Valid as Notropis caeruleus (Jordan 1877) -- (Gilbert 1978:33 [ref. 7042], Lee et al. 1980:244 [ref. 22416]). •Valid as Cyprinella caerulea (Jordan 1877) -- (Mayden 1989:85 [ref. 12555], Page & Burr 1991:126 [ref. 18983], Boschung 1992:45 [ref. 23239], Etnier & Starnes 1993:147 [ref. 22809], Mettee et al. 1996:157 [ref. 25836], Gilbert 1998:54 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Boschung & Mayden 2004:162 [ref. 27995], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:244 [ref. 31215], Page et al. 2013:69 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Bagley et al. 2023:313 [ref. 41241], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella caerulea (Jordan 1877). Leuciscidae: Pogonichthyinae. Distribution: North America: southeastern Tennessee, Georgia and Alabama (southeastern U.S.A.). Habitat: freshwater.

callisema, Episema Jordan [D. S.] 1877:363 [Annals of the Lyceum of Natural History of New York v. 11 (nos 11-12) (art. 29); ref. 2373] South Fork of Ocmulgee River, Flat Rock, DeKalb County, Georgia, U.S.A. Lectotype: USNM 17864. Paralectotypes: ?BMNH 1880.1.21.20 (1), 1883.12.14.210; MCZ 24384 (1); USNM 20126 (1), 163954 [ex USNM 17864] (3). Type catalog: Gilbert 1978:33-34 [ref. 7042], Gilbert 1998:55 [ref. 23395], Gilbert 2009:161 [ref. 30728]. Lectotype selected by Gibbs in Gilbert 1978:33 [ref. 7042]. •Valid as Notropis callisema (Jordan 1877) -- (Gilbert 1978:34 [ref. 7042], Lee et al. 1980:246 [ref. 22416]). •Valid as Cyprinella callisema (Jordan 1877) -- (Mayden 1989:95 [ref. 12555], Page & Burr 1991:127 [ref. 18983], Gilbert 1998:55 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:246 [ref. 31215], Page et al. 2013:69 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella callisema (Jordan 1877). Leuciscidae: Pogonichthyinae. Distribution: North America: Georgia (eastern U.S.A.). Habitat: freshwater.

callistius, Photogenis Jordan [D. S.] 1877:337 [Annals of the Lyceum of Natural History of New York v. 11 (nos 11-12) (art. 29); ref. 2373] Tributaries of Etowah and Oostanaula rivers, near Rome, Georgia, U.S.A. Lectotype: MCZ 24363. Paralectotypes: ANSP 19658 (1), 19837-38 (2); BMNH 1880.1.21.67 (1); MNHN A-1289 (1); USNM 17882 (1), 31148 (2), 163965-66 (1, 1). Type catalog: Bertin & Estève 1948:66 [ref. 19574], Gilbert 1978:34 [ref. 7042], Böhlke 1984:71 [ref. 13621], Gilbert 1998:55 [ref. 23395], ilbert 2009:158 [ref. 30728]. Lectotype selected by Gibbs in Gilbert 1978:34 [ref. 7042]. •Valid as Notropis callistius (Jordan 1877) -- (Gilbert 1978:34 [ref. 7042], Lee et al. 1980:246 [ref. 22416]). •Valid as Cyprinella callistia (Jordan 1877) -- (Mayden 1989:90 [ref. 12555], Page & Burr 1991:124 [ref. 18983], Boschung 1992:46 [ref. 23239], Etnier & Starnes 1993:148 [ref. 22809], Mettee et al. 1996:159 [ref. 25836], Gilbert 1998:55 [ref. 23395], Ross et al. 2001:136 [ref. 25978], Nelson et al. 2004:69 [ref. 27807], Boschung & Mayden 2004:164 [ref. 27995], Scharpf 2005:13 [ref. 28940], Page & Burr 2011:242 [ref. 31215], Page et al. 2013:69 [ref. 32708], Stout et al. 2022:7 [ref. 39738] as 'Cyprinella' callistia, Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella callistia (Jordan 1877). Leuciscidae: Pogonichthyinae. Distribution: Eastern North America: southeastern U.S.A. Habitat: freshwater.

callitaenia, Notropis Bailey [R. M.] & Gibbs [R. H., Jr.] 1956:2, Pl. 1 [Occasional Papers of the Museum of Zoology University of Michigan No. 576; ref. 12177] Flint River, about 1 mile south of the outlet of Radium Springs, 5.5 miles south of Albany, Dougherty County, Georgia, U.S.A. Holotype: UMMZ 168938 [ex UMMZ 163922]. Paratypes: ANSP 73864 (20); UMMZ 163922 (76, now 70); USNM 171351 (6). Additional non-type material at CUMV, UF, UMMZ, USNM. Type catalog: Gilbert 1978:34 [ref. 7042], Böhlke 1984:71 [ref. 13621], Gilbert 1998:55 [ref. 23395]. •Valid as Notropis callitaenia Bailey & Gibbs 1956 -- (Gilbert 1978:34 [ref. 7042], Lee et al. 1980:247 [ref. 22416]). •Valid as Cyprinella callitaenia (Bailey & Gibbs 1956) -- (Mayden 1989:97 [ref. 12555], Page & Burr 1991:127 [ref. 18983], Boschung 1992:46 [ref. 23239], Mettee et al. 1996:161 [ref. 25836], Gilbert 1998:55 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Boschung & Mayden 2004:165 [ref. 27995], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:246 [ref. 31215], Page et al. 2013:69 [ref. 32708], Robins et al. 2018:80 [ref. 35886], Bagley et al. 2023:313 [ref. 41241], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella callitaenia (Bailey & Gibbs 1956). Leuciscidae: Pogonichthyinae. Distribution: Eastern North America: Georgia, Alabama and Florida (southeastern U.S.A.). Habitat: freshwater.

calliura, Cyprinella Jordan [D. S.] 1877:61 [Bulletin of the United States National Museum [v. 1] No. 10; ref. 2374] Black Warrior River, Alabama, U.S.A. Lectotype: USNM 6865. Paralectotypes: USNM 163951 [ex USNM 6865] (7), 214661 (1). Type catalog: Gilbert 1978:34 [ref. 7042], Gilbert 1998:55 [ref. 23395] with note on locality. Appeared as name only in Jordan 1877:339 [ref. 2373]. Lectotype selected by Gibbs 1957:192 [ref. 21244]. •Synonym of Notropis venustus (Girard 1856) as intergrades between two subspecies -- (Gibbs 1957:195 [ref. 21244], Gilbert 1978:34 [ref. 7042]). •Synonym of Cyprinella venusta Girard 1856 -- (Mayden 1989:78 [ref. 12555]). •Synonym of Cyprinella stigmatura (Jordan 1877) -- (Gilbert 1998:55 [ref. 23395]). Current status: Synonym of Cyprinella stigmatura (Jordan 1877). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

camura, Cliola Jordan [D. S.] & Meek [S. E.] 1884:474 [Proceedings of the United States National Museum v. 7 (no. 450); ref. 10636] Arkansas River at Ft. Lyon, Colorado, U.S.A. [in error, probably Lyons County, Kansas, U.S.A.]. Lectotype: USNM 15256. Paralectotypes: USNM 163962 [ex USNM 15256] (1). Type catalog: Gilbert 1978:34 [ref. 7042], Gilbert 1998:55-56 [ref. 23395] with locality information. Lectotype selected by Gibbs 1961:345 [ref. 21240]. •Valid as Notropis camurus (Jordan & Meek 1884) -- (Gibbs 1961:344 [ref. 21240], Gilbert 1978:35 [ref. 7042], Lee et al. 1980:248 [ref. 22416], Dimmick 1987:316 [ref. 11406], Robison & Buchanan 1988:205 [ref. 23393]). •Valid as Cyprinella camura (Jordan & Meek 1884) -- (Mayden 1989:68 [ref. 12555], Page & Burr 1991:122 [ref. 18983], Etnier & Starnes 1993:149 [ref. 22809], Warren et al. 1994:132 [ref. 25557], Cross & Collins 1995:55 [ref. 23394], Gilbert 1998:55-56 [ref. 23395], Ross et al. 2001:138 [ref. 25978], Nelson et al. 2004:69 [ref. 27807], Miller & Robison 2004:90 [ref. 27931], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:239 [ref. 31215], Page et al. 2013:69 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella camura (Jordan & Meek 1884). Leuciscidae: Pogonichthyinae. Distribution: North America: Kentucky to Louisiana, Missouri, Kansas, Arkansas and Oklahoma (central U.S.A.). Habitat: freshwater.

cercostigma, Cyprinella Cope [E. D.] 1868:157 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 19; ref. 7316] Pearl River at Monticello, Lawrence County, Mississippi, U.S.A. Syntypes: USNM (4, not found). Type catalog: Böhlke 1984:72 [ref. 13621], Gilbert 1998:58 [ref. 23395]. •Synonym of Notropis venustus (Girard 1856), but a valid subspecies -- (Gibbs 1957:189 [ref. 21244], Gilbert 1978:35 [ref. 7042], Lee et al. 1980:321 [ref. 22416], Robison & Buchanan 1988:239 [ref. 23393]). •Synonym of Cyprinella venusta Girard 1856 -- (Mayden 1989:77 [ref. 12555]). •Synonym of Cyprinella venusta Girard 1856, but a valid subspecies -- (Gilbert 1998:58 [ref. 23395], Ross et al. 2001:150 [ref. 25978], Scharpf 2005:14 [ref. 28940], Page & Burr 2011:239 [ref. 31215], Robins et al. 2018:84 [ref. 35886] with 3 subspecies). Current status: Synonym of Cyprinella venusta Girard 1856. Leuciscidae: Pogonichthyinae. Distribution: Gulf drainages east of the Mississippi River [subspecies cercostigma]. Habitat: freshwater.

chickasavensis, Luxilus Hay [O. P.] 1881:506 [Proceedings of the United States National Museum v. 3 (no. 179); ref. 2053] Chickasawha River at Enterprise, Clark County, Mississippi, U.S.A. Lectotype: USNM 27419. Paralectotypes: USNM 163953 (8). Type catalog: Gilbert 1978:36 [ref. 7042], Gilbert 1998:58-59 [ref. 23395]. Lectotype selected by Gibbs 1957:190 [ref. 21244]. •Synonym of Notropis venustus (Girard 1856), subspecies cercostigma (Cope 1868) -- (Gibbs 1957:189 [ref. 21244], Gilbert 1978:36 [ref. 7042]). •Synonym of Cyprinella venusta Girard 1856 -- (Mayden 1989:78 [ref. 12555], Ross et al. 2001:150 [ref. 25978]). •Synonym of Cyprinella venusta Girard 1856, but a valid subspecies -- (Gilbert 1998:58-59 [ref. 23395]). Current status: Synonym of Cyprinella venusta Girard 1856. Leuciscidae: Pogonichthyinae. Habitat: freshwater.

chloristia, Codoma Jordan [D. S.] & Brayton [A. W.] 1878:21 [Bulletin of the United States National Museum No. 12; ref. 2436] Tributary of Saluda River, Farr's Mills, west of Greenville, Greenville County, North Carolina, U.S.A. Lectotype: MCZ 24380. Paralectotypes: BMNH 1880.1.21.43 (1), INHS (1, lost). Type catalog: Sabaj et al. 1997:286 [ref. 23225], Gilbert 1998:59 [ref. 23395]. Lectotype selected by Gibbs 1963:522 [ref. 21327]. •Synonym of Notropis analostanus (Girard 1859), but a valid subspecies -- (Gibbs 1963:522 [ref. 21327]). •Valid as Notropis chloristius (Jordan & Brayton 1878) -- (Gilbert 1978:36 [ref. 7042], Lee et al. 1980:253 [ref. 22416]). •Valid as Cyprinella chloristia (Jordan & Brayton 1878) -- (Mayden 1989:76 [ref. 12555], Page & Burr 1991:121 [ref. 18983], Gilbert 1998:59 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:238 [ref. 31215], Page et al. 2013:69 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella chloristia (Jordan & Brayton 1878). Leuciscidae: Pogonichthyinae. Distribution: Eastern North America: North Carolina and South Carolina (eastern U.S.A.). Habitat: freshwater.

complanata, Moniana Girard [C. F.] 1856:200 [36] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Rio Grande (or tributary), Brownsville, Cameron County, Texas, U.S.A. Syntypes: (3) USNM 94 (orig 3, now 0, 1 pharyngeal arch as # 2694). Type catalog: Gilbert 1978:38 [ref. 7042], Gilbert 1998:63 [ref. 23395]. On p. 36 of separate. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:38 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:50 [ref. 12555], Gilbert 1998:63 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

cooglei, Notropis Hildebrand [S. F.] & Towers [I. L.] 1928:118, Fig. 3 [Bulletin of the Bureau of Fisheries v. 43 (pt 2, 1927); ref. 15897] Pelucia Creek, south of Greenwood, Leflore County, Mississippi, U.S.A. Holotype: USNM 88379. Paratypes: (214) UMMZ 167162-66 [ex USNM] (8, 10, 8, 4, 4); USNM 127019-24 (29, 1, 46, 26, 63 or 62, 22). Type catalog: Gilbert 1978:38 [ref. 7042], Gilbert 1998:64-65 [ref. 23395]. •Synonym of Notropis venustus (Girard 1856) -- (Gibbs 1957:180 [ref. 21244], Gilbert 1978:38 [ref. 7042]). •Synonym of Cyprinella venusta Girard 1856 -- (Mayden 1989:78 [ref. 12555], Gilbert 1998:64-65 [ref. 23395]). Current status: Synonym of Cyprinella venusta Girard 1856. Leuciscidae: Pogonichthyinae. Habitat: freshwater.

couchi, Moniana Girard [C. F.] 1856:201 [37] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Río San Juan, near China, Nuevo León State, Mexico. Syntypes: MCZ 1795 (2); USNM 103 ? = 20227 (34 spec., 2 pharyngeal bones). Type catalog: Gilbert 1978:39 [ref. 7042], Gilbert 1998:66 [ref. 23395]. On p. 37 of separate. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:39 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:50 [ref. 12555], Gilbert 1998:66 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

eurystomus, Photogenis Jordan [D. S.] 1877:356 [Annals of the Lyceum of Natural History of New York v. 11 (nos 11-12) (art. 29); ref. 2373] Nancys Creek, tributary of Chattahoochee River, Atlanta, Fulton County Georgia, U.S.A. Lectotype: MCZ 24388. Paralectotypes: apparently lost. Type catalog: Gilbert 1978:42-43 [ref. 7042], Gilbert 1998:77-78 [ref. 23395], Gilbert 2009:161 [ref. 30728]. Lectotype dsignated by Gibbs 1957:190 [ref. 21244]. •Synonym of Notropis venustus (Girard 1856), subspecies cercostigma (Cope 1868) -- (Gibbs 1957:189 [ref. 21244], Gilbert 1978:43 [ref. 7042]). •Synonym of Cyprinella venusta Girard 1856 -- (Mayden 1989:78 [ref. 12555]). •Valid as Cyprinella eurystoma (Jordan 1877) -- (Gilbert 1998:78 [ref. 23395]). Current status: Valid as Cyprinella eurystoma (Jordan 1877). Leuciscidae: Pogonichthyinae. Distribution: Southeastern U.S.A. Habitat: freshwater.

forbesi, Cyprinella Jordan [D. S.] 1878:57 [Bulletin of the Illinois State Laboratory of Natural History v. 1 (no. 2); ref. 10444] Illinois River, Union County, southern Illinois, U.S.A. Lectotype: USNM 29869. Paralectotypes: (many) INHS 86401 (6), 86451 (21); CAS-SU 1290 (2). Type catalog: Böhlke 1953:31 [ref. 12291], Gilbert 1978:43 [ref. 7042], Sabaj et al. 1997:262-263 [ref. 23225], Gilbert 1998:80-81 [ref. 23395]. Lectotype designation by Jordan & Evermann 1896:272 [ref. 2443] but mistakenly reported as USNM 29867; if that is not accepted then lectotype established by Sabaj et al. 1997:262 [ref. 23225]. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:43 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Matthews 1987:616 [ref. 6774], Mayden 1989:50 [ref. 12555], Gilbert 1998:81 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

forlonensis, Nototropis Meek [S. E.] 1904:70, Fig. 20 [Field Columbian Museum, Zoölogical Series v. 5; ref. 2958] Río Forlón, tributary of River Pánuco, Forlón, Tamaulipas, Mexico. Holotype: FMNH 4478. Paratypes: CAS 33825 [ex IU 10914] (4); FMNH 4479 (25), 4495 (3); CAS-SU 22341 (3). Type catalog: Gilbert 1978:43 [ref. 7042], Ibarra & Stewart 1987:60 [ref. 12367], Gilbert 1998:81 [ref. 23395]. Misspelled forionensis in Zoological Record. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:43 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:50 [ref. 12555], Miller 2006:112 [ref. 28615]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853), but a valid subspecies -- (Gilbert 1998:81 [ref. 23395], Scharpf 2005:13 [ref. 28940]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

formosa, Moniana Girard [C. F.] 1856:201 [37] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Río Mimbres, Chihuahua, Mexico [= Mimbres River, north of Deming, Luna County, New Mexico, U.S.A.]. Syntypes: ANSP 3056 [ex USNM 114] (1); MCZ 1686 [ex USNM 114] (2); MNHN 0000-0438 [ex USNM 114] (2); USNM 113 (orig. 4, now 0), 114 (orig. 20, now 0, 1 pair pharyngeal arches as #2700). Type catalog: Gilbert 1978:44 [ref. 7042], Böhlke 1984:76 [ref. 13621], Gilbert 1998:81 [ref. 23395]. On p. 37 of separate. •Valid as Cyprinella formosa (Girard 1856) -- (Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:13 [ref. 28940] with subspecies, Miller 2006:110 [ref. 28615], Minckley & Marsh 2009:118, 177 [ref. 31114], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:240 [ref. 31215], Page et al. 2013:69 [ref. 32708], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella formosa (Girard 1856). Leuciscidae: Pogonichthyinae. Distribution: Southern North America: southwestern U.S.A. (New Mexico and Arizona) and Mexico. Habitat: freshwater.

fretensis, Hybopsis Cope [E. D.] 1867:382 [Transactions of the American Philosophical Society (New Series) v. 13 (pt 3) (art. 13); ref. 910] Near Detroit, Wayne County, Michigan, U.S.A. Holotype (unique): ANSP 2840. Type catalog: Gilbert 1978:44 [ref. 7042], Böhlke 1984:76 [ref. 13621], Gilbert 1998:82 [ref. 23395]; see also Fowler 1918:22 [ref. 11665]. •Synonym of Notropis spilopterus (Cope 1867) -- (Gibbs 1957:192 [ref. 12065], Gilbert 1978:44 [ref. 7042], Scott & Crossman 1998:468 [ref. 24570]). •Synonym of Cyprinella spiloptera (Cope 1867) -- (Mayden 1989:66 [ref. 12555], Gilbert 1998:82 [ref. 23395]). Current status: Synonym of Cyprinella spiloptera (Cope 1867). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

frigida, Moniana Girard [C. F.] 1856:200 [36] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Salado Creek, tributary to Rio San Antonio, just east of San Antonio, Bexar County, Texas, U.S.A. Lectotype: USNM 123. Paralectotypes: ANSP 5062-63 [ex USNM 123] (2); UMMZ 56287 [ex USNM 123] (1); USNM 97 (orig. 2, now 0), 98 (9), 99 (1, not found), 100 (orig. 3, now 0, 4 pharyngeal archs as #2696), 101 (orig. 10, now 0, plus 2 pharyngeal arches as #2697), 124 (orig. 60, now 47, plus 2 pharyngeal arches as #2707), 343586 [ex USNM 123] (orig. 100, now 92), 343588-89 [ex USNM 124] (1, 4). Type catalog: Gilbert 1978:44-45 [ref. 7042], Böhlke 1984:76 [ref. 13621], Gilbert 1998:82-83 [ref. 23395]. On p. 36 of separate. Lectotype selected by Gilbert 1998:82 [ref. 23395]. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:45 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:49 [ref. 12555]). •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:45 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853), subspecies lutrensis -- (Gilbert 1998:82-83 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

galacturus, Hypsilepis Cope [E. D.] 1868:160 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 19; ref. 7316] Tributary of the Holston River, Virginia, U.S.A. Lectotype: USNM 14981. Paralectotypes: ANSP 3381-3444 (65), MCZ 32255 (1), MNHN 0000-4853 (1). Type catalog: Bertin & Estève 1948:65 [ref. 19574], Gilbert 1978:45 [ref. 7042], Böhlke 1984:77 [ref. 13621], Gilbert 1998:83 [ref. 23395]. Original genus should have been Hypsolepis. Lectotype established by Jordan & Evermann 1896:279 [ref. 2443]; also designated by Gibbs 1961:339 [ref. 21240]; see also Gilbert 1978:45 [ref. 7042]. •Valid as Notropis galacturus (Cope 1868) -- (Gibbs 1961:338 [ref. 21240], Gilbert 1978:43 [ref. 7042], Lee et al. 1980:266 [ref. 22416], Robison & Buchanan 1988:211 [ref. 23393]). •Valid as Cyprinella galactura (Cope 1868) -- (Mayden 1989:80 [ref. 12555], Page & Burr 1991:124 [ref. 18983], Boschung 1992:47 [ref. 23239], Etnier & Starnes 1993:150 [ref. 22809], Jenkins & Burkhead 1994:356 [ref. 21581], Stauffer et al. 1995:70 [ref. 23376], Mettee et al. 1996:163 [ref. 25836], Gilbert 1998:83 [ref. 23395], Fuller et al. 1999:62 [ref. 25838], Ross et al. 2001:140 [ref. 25978], Nelson et al. 2004:69 [ref. 27807], Boschung & Mayden 2004:166 [ref. 27995], Boschung & Mayden 2004:166 [ref. 27995], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:242 [ref. 31215], Page et al. 2013:69 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Bagley et al. 2023:313 [ref. 41241], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella galactura (Cope 1868). Leuciscidae: Pogonichthyinae. Distribution: Eastern North America: eastern U.S.A. Habitat: freshwater.

garmani, Notropis Jordan [D. S.] 1885:813 [25] [United States Commission of Fish and Fisheries, Report of the Commissioner v. 13 (1885); ref. 2385] Lago del Muerto, near Parras, 25°25'N, 102°11'W, Coahuila, Mexico. Syntypes: MCZ 25891 (2), 24892 (orig. 26, now 20); SMNS 3034 (2); UMMZ 86663 [ex MCZ 24892] (6); USNM 120257 [ex MCZ 14892] (6). Type catalog: Gilbert 1978:45 [ref. 7042], Gilbert 1998:84 [ref. 23395], Fricke 1995:11 [ref. 28100], Fricke 2005:31 [ref. 29864]. On p. 25 of separate. Replacement name for Cyprinella rubripinna Garman 1881, secondarily preoccupied by Argyreus rubripinnis Heckel 1843, only when classified in genus Notropis. •Valid as Notropis garmani Jordan 1885 -- (Chernoff & Miller 1986:182 [ref. 6782]). •Synonym of Notropis lutrensis (Baird & Girard 1853), but a valid subspecies -- (Gilbert 1978:46 [ref. 7042]). •Valid as Cyprinella garmani (Jordan 1885) -- (Mayden 1989:53 [ref. 12555], Espinosa Pérez et al. 1993:11 [ref. 22290], Gilbert 1998:84 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:13 [ref. 28940], Miller 2006:111 [ref. 28615], Schönhuth & Mayden 2009:78 [ref. 38584], Page et al. 2013:69 [ref. 32708], Page et al. 2023:68 [ref. 40505]). •Synonym of Cyprinella rubripinna Garman 1881 – (Fricke 2005: 32 [ref. 29864]). Replacement Name. Current status: Synonym of Cyprinella rubripinna Garman 1881. Leuciscidae: Pogonichthyinae. Distribution: Southern North America: Coahuila (Mexico) [if valid]. Habitat: freshwater.

gibbosa, Moniana Girard [C. F.] 1856:201 [37] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Rio Grande (or tributary) at Brownsville, Cameron County, Texas, U.S.A. Syntypes: USNM 105 [= USNM 31159] (orig. 3, now 1). Type catalog: Gilbert 1978:46 [ref. 7042], Gilbert 1998:85 [ref. 23395]. On p. 37 of separate. Subjectively invalid; secondarily preoccupied in Notropis by Leuciscus gibbosus Storer 1848. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:46 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:50 [ref. 12555]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) subspecies lutrensis -- (Gilbert 1998:85 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

gibbsi, Notropis Howell [W. M.] & Williams [J. D.] 1971:55, Figs. 1-3 [Copeia 1971 (no. 1); ref. 18313] Enitachope Creek, tributary to Hillabee Creek, 3.2 miles south-southwest of Ashland, Clay County, Alabama, U.S.A. Holotype: USNM 204933. Paratypes: CUMV 64311 (45), TU 61102 (28), UIAC 3020 (30), UF 15933 (28), 66881 [ex FSU 16881] (28), UMMZ 189199 (45), USNM 204934 (40). Non-types: UMMZ (2 lots), USNM (1 lot), ALA (35 lots). Type catalog: Gilbert 1974:112 [ref. 21035], Gilbert 1978:46 [ref. 7042], Bart & Taylor 1993:42 [ref. 20906], Gilbert 1998:85 [ref. 23395]. •Valid as Notropis gibbsi Howell & Williams 1971 -- (Gilbert 1978:46 [ref. 7042], Lee et al. 1980:267 [ref. 22416]). •Valid as Cyprinella gibbsi (Howell & Williams 1971) -- (Mayden 1989:87 [ref. 12555], Page & Burr 1991:126 [ref. 18983], Boschung 1992:46 [ref. 23239], Mettee et al. 1996:165 [ref. 25836], Gilbert 1998:85 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Boschung & Mayden 2004:167 [ref. 27995], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:243 [ref. 31215], Page et al. 2013:69 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Bagley et al. 2023:313 [ref. 41241], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella gibbsi (Howell & Williams 1971). Leuciscidae: Pogonichthyinae. Distribution: North America: Alabama and Georgia (southeastern U.S.A.). Habitat: freshwater.

gracilis, Moniana Girard [C. F.] 1856:201 [37] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Acapulco, near Monterrey, Nuevo León, Mexico. Syntypes: ANSP 2992 [ex USNM 116] (1), MCZ 1796 [ex USNM 116] (1), MNHN 0000-0432 [ex USNM 116] (1), USNM 116 (orig. 7, now 4). Type catalog: Gilbert 1978:47 [ref. 7042], Böhlke 1984:77 [ref. 13621], Gilbert 1998:87 [ref. 23395]. On p. 37 of separate. Subjectively invalid; secondarily preoccupied by Leuciscus gracilis Agassiz 1850 when both are in Notropis. •Synonym of Notropis lutrensus (Baird & Girard 1853) -- (Gilbert 1978:47 [ref. 7042]). •Synonym of Cyprinella rutila (Girard 1856) -- (Mayden 1989:60 [ref. 12555], Gilbert 1998:87 [ref. 23395]). 2°Homonym. Current status: Synonym of Cyprinella rutila (Girard 1856). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

gunnisoni, Cyprinella Girard [C. F.] 1856:197 [33] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Cottonwood River, about 5 miles northwest of Durham, Marion County, Kansas. Syntypes: (12) USNM 139 (24, plus 1 set of pharyngeal arches as #2718). Type catalog: Gilbert 1978:48 [ref. 7042] with locality correction. On p. 33 of separate. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:48 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:49 [ref. 12555]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853), subspecies lutrensis -- (Gilbert 1998:89 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

hypsisomatus, Notropis spilopterus Gibbs [R. H., Jr.] 1957:195, Fig. 4 (upper & lower middle) [Lloydia (Cincinnati) v. 20 (no. 3); ref. 12065] Bay at north end of Wonder Lake, McHenry County, Illinois, U.S.A. Holotype: CUMV 28541. Paratypes: CUMV 17931 (48), 17946 (15), 17960 (31), 17984 (6), 22305 (2), 22326 (12), 22330 (33). Type catalog: Gilbert 1978:50 [ref. 7042], Gilbert 1998:94 [ref. 23395]. •Synonym of Notropis spilopterus (Cope 1867) -- (Gilbert 1978:50 [ref. 7042], Lee et al. 1980:312 [ref. 22416], Robison & Buchanan 1988:233 [ref. 23393]). •Synonym of Cyprinella spiloptera (Cope 1867) -- (Mayden 1989:66 [ref. 12555], Gilbert 1998:94 [ref. 23395]). Current status: Synonym of Cyprinella spiloptera (Cope 1867). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

iris, Hypsilepis Cope [E. D.] in Cope & Yarrow 1875:653, Pl. 31 (figs. 4-4a) [Engineer Department, United States Army, Report upon the Geographical and Geological Explorations and Surveys west of the one hundredth Meridian v. 5 (Zoology) Chapter 6; ref. 968] Rio Grande at San Ildefonso, about 10 miles east of Los Alamos, Santa Fe County, New Mexico, U.S.A. Lectotype: USNM 16976. Paralectotypes: MCZ 35927 [ex USNM 16976] (5), 35953 (1); USNM 16977 (orig. 268, now 187), 16980 (98), 343590 [ex USNM 16976] (now 9), 343591 [ex USNM 16976] (now 98), 343592 (19), 343593 (8), 343594 (2), 343595 (50), 343596 (1) 343597 (6), 343598 (2), 343599 (27), 343601 (28). Type catalog: Gilbert 1978:50-51 [ref. 7042], Gilbert 1998:96-97 [ref. 23395]. Lectotype selected by Gilbert 1998:96 [ref. 23395]. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:51 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:50 [ref. 12555]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853), subspecies lutrensis (Gilbert 1998:97 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

jugalis, Moniana Cope [E. D.] 1871:439 [Preliminary report of the United States Geological Survey of Wyoming and portions of contiguous territories. Part 4 (art. 8); ref. 19655] Missouri River at St. Joseph, Missouri, U.S.A. Lectotype: ANSP 3144. Paralectotypes: ANSP 3145-50 (16). Type catalog: Gilbert 1978:52 [ref. 7042], Böhlke 1984:80 [ref. 13621] as holotype and paratypes, Gilbert 1998:98 [ref. 23395]. Lectotype established by Fowler 1910:279, Pl. 17 (fig. 20). •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:52 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:50 [ref. 12555]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) subspecies lutrensis -- (Gilbert 1998:98 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

labrosus, Ceratichthys Cope [E. D.] 1870:458 [Proceedings of the American Philosophical Society v. 11 (no. 84); ref. 913] Upper waters of Catawba River (Santee River system), McDowell and Burke counties, North Carolina, U.S.A. Lectotype: ANSP 2045. Paralectotypes: ANSP 2041-43 (3), 2044 (1), 2046 (1), 2047-2054 (8). Type catalog: Böhlke 1984:80 [ref. 13621], Gilbert 1998:100 [ref. 23395]. Lectotype established by Jenkins & Lachner in Gilbert 1978:91 [ref. 7042]. •Valid as Hybopsis labrosa (Cope 1870) -- (Gilbert 1978:91 [ref. 7042], Lee et al. 1980:189 [ref. 22416], Page & Burr 1991:162 [ref. 18983]). •Valid as Cyprinella labrosa (Cope 1870) -- (Jenkins & Burkhead 1994:350 [ref. 21581], Gilbert 1998:100 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:248 [ref. 31215], Page et al. 2013:69 [ref. 32708], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella labrosa (Cope 1870). Leuciscidae: Pogonichthyinae. Distribution: North America: Virginia, North Carolina and South Carolina (eastern U.S.A.). Habitat: freshwater.

laetabilis, Moniana Girard [C. F.] 1856:200 [36] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Hurah Creek (a tributary to Pecos River), about 10 miles northeast of Santa Rosa, Guadalupe County, New Mexico, U.S.A. Syntypes: ANSP 3029 (1), MCZ 1691 (1), USNM 120 (10+, now 0, 1 set of pharyngeal arches as # 2705). Type catalog: Gilbert 1978:53 [ref. 7042], Böhlke 1984:80 [ref. 13621], Gilbert 1998:100 [ref. 23395]. On p. 36 of separate. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:53 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:50 [ref. 12555]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) subspecies lutrensis -- (Gilbert 1998:100 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

leedsi, Notropis Fowler [H. W.] 1942:2, Figs. 3-4 [Notulae Naturae (Philadelphia) No. 107; ref. 13199] Ohoopee River, about 2.5 miles north of Oak Park, Emanuel County, Georgia, U.S.A. Holotype (unique): ANSP 69969. Type catalog: Gilbert 1978:53 [ref. 7042], Böhlke 1984:81 [ref. 13621], Gilbert 1998:101 [ref. 23395]. •Valid as Notropis leedsi Fowler 1942 -- (Gilbert 1978:53 [ref. 7042], Lee et al. 1980:280 [ref. 22416]). •Valid as Cyprinella leedsi (Fowler 1942) -- (Mayden 1989:93 [ref. 12555], Page & Burr 1991:128 [ref. 18983], Gilbert 1998:101 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:247 [ref. 31215], Page et al. 2013:69 [ref. 32708], Robins et al. 2018:82 [ref. 35886], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella leedsi (Fowler 1942). Leuciscidae: Pogonichthyinae. Distribution: North America: South Carolina, Georgia and Florida (southeastern U.S.A.). Habitat: freshwater.

leonina, Moniana Girard [C. F.] 1856:199 [35] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Leon River [Leon Creek], tributary to San Antonio River, Bexar County, Texas, U.S.A. Syntypes: (18) MCZ 1687 [ex USNM 115] (1), MNHN 0000-0377 [ex USNM 115] (1), USNM 115 (orig. 10, now 0, plus 1 set of pharyngeal bones as # 2701). Type catalog: Gilbert 1978:53-54 [ref. 7042]. On p. 35 of separate •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:54 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:49 [ref. 12555]). •Synonym of Cyprinella lutrensis, subspecies lutrensis (Baird & Girard 1853) -- (Gilbert 1998:102 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

lepida, Cyprinella Girard [C. F.] 1856:197 [33] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Rio Frio, a tributary to Rio Nueces, Real or Uvalde County, Texas, U.S.A. Syntypes: (7) MCZ 1692 [ex USNM 126] (1), MNHN 0000-0369 [ex USNM 126] (1), USNM 126 (orig. 7, now 0, 1 set of pharyngeal arches as # 2709). Type catalog: Gilbert 1978:54 [ref. 7042], Gilbert 1998:102 [ref. 23395]. On p. 33 of separate. •Valid as Notropis lepidus (Girard 1856) -- (Gilbert 1978:54 [ref. 7042]). •Valid as Cyprinella lepida Girard 1856 -- (Matthews 1987:616 [ref. 6774], Mayden 1989:58 [ref. 12555], Page & Burr 1991:124 [ref. 18983], Contreras-Balderas & Lozano-Vilano 1994:904 [ref. 22061], Richardson & Gold 1995:28 [ref. 22412], Gilbert 1998:102 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:13 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:241 [ref. 31215], Page et al. 2013:70 [ref. 32708], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella lepida Girard 1856. Leuciscidae: Pogonichthyinae. Distribution: North America: Texas (southeastern U.S.A.). Habitat: freshwater.

leucopus, Photogenis Jordan [D. S.] & Brayton [A. W.] 1878:41 [Bulletin of the United States National Museum No. 12; ref. 2436] Chattahoochee River at Shallow Ford, northwest of Gainesville, Hall County, Georgia, U.S.A. Lectotype: USNM 31124. Paralectotypes: MCZ 35935 (1); USNM 163963-64 (1, 2). Type catalog: Gilbert 1978:54 [ref. 7042], Gilbert 1998:103 [ref. 23395]. Lectotype selected by Bailey & Gibbs 1956:2 [ref. 12177]; they regard USNM 163964 as a different species. •Synonym of Notropis venustus (Girard 1856) subspecies cercostigma (Cope 1868) -- (Gibbs 1957:189 [ref. 21244], Gilbert 1978:54 [ref. 7042]). •Synonym of Cyprinella venusta (Girard 1856) -- (Mayden 1989:78 [ref. 12555]). •Synonym of Cyprinella eurystoma (Jordan 1877) -- (Gilbert 1998:103 [ref. 23395]). Current status: Synonym of Cyprinella eurystoma (Jordan 1877). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

lutrensis, Leuciscus Baird [S. F.] & Girard [C. F.] 1853:391 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 6; ref. 165] Otter Creek, Arkansas [= tributary to North Fork of Red River, either Kiowa or Tillman County, southwestern Oklahoma], U.S.A. Syntypes: (5) USNM 104 (orig. 5, now 1). Subsequent material: MCZ 1685 [ex USNM 107] (1, discarded in 1938). Type catalog: Gilbert 1978:57 [ref. 7042], Gilbert 1998:107 [ref. 23395]. •Valid as Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:57 [ref. 7042], Lee et al. 1980:285 [ref. 22416], Chernoff & Miller 1986:182 [ref. 6782], Robison & Buchanan 1988:216 [ref. 23393]). •Valid as Cyprinella lutrensis (Baird & Girard 1853) -- (Matthews 1987:616 [ref. 6774], Mayden 1989:49 [ref. 12555], Page & Burr 1991:123 [ref. 18983], Boschung 1992:46 [ref. 23239], Espinosa Pérez et al. 1993:11 [ref. 22290], Etnier & Starnes 1993:152 [ref. 22809], Warren et al. 1994:132 [ref. 25557], Richardson & Gold 1995:28 [ref. 22412], Baxter & Stone 1995:64 [ref. 23374], Cross & Collins 1995:56 [ref. 23394], Schmidt & Gold 1995:200 [ref. 22413], Mettee et al. 1996:161 [ref. 25836], Sigler & Sigler 1996:68 [ref. 25943], Fuller et al. 1999:63 [ref. 25838], Gilbert 1998:107 [ref. 23395], Lyons et al. 2000:64 [ref. 26594], Ross et al. 2001:142 [ref. 25978], Moyle 2002:168 [ref. 26449], Grose & Wiley 2002:1092 [ref. 26513], Nelson et al. 2004:69 [ref. 27807], Miller & Robison 2004:92 [ref. 27931], Boschung & Mayden 2004:168 [ref. 27995], Scharpf 2005:13 [ref. 28940] with subspecies, Miller 2006:112 [ref. 28615], Minckley & Marsh 2009:178 [ref. 31114], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:240 [ref. 31215], Page et al. 2013:70 [ref. 32708], Conway & Kim 2016:334 [ref. 34355], Stout et al. 2022:7 [ref. 39738], Bagley et al. 2023:313 [ref. 41241], Page et al. 2023:68 [ref. 40505]). Current status: Valid as Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Distribution: North America: central U.S.A. to northern Mexico; widely introduced. Habitat: freshwater.

luxiloides, Cyprinella Girard [C. F.] 1856:198 [34] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Rio San Antonio, Bexar County, Texas, U.S.A. Neotype: TCWC 14075.01. Syntypes: USNM 131 (orig. 2, now 0). Type catalog: Gilbert 1978:57-58 [ref. 7042], Gilbert 1998:107-108 [ref. 23395]. Neotype designated by Conway & Kim 2016:332 [ref. 34355]. On p. 34 of separate. Syntypes lost, see Conway & Kim 2016:329 [ref. 34355]. •Probably a synonym of Notropis amabilis (Girard 1856) -- (Gilbert 1978:58 [ref. 7042], Gilbert 1998:107-108 [ref. 23395]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Conway & Kim 2016:332 [ref. 34355]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

mearnsi, Notropis Snyder [J. O.] 1915:582, Fig. [Proceedings of the United States National Museum v. 49 (no. 2125); ref. 10255] Río San Bernardino (tributary to Río Yaqui), near monument 77 of the international boundary, Sonora, Mexico. Holotype: USNM 76163. Paratypes: USNM 76164 (4). Type catalog: Gilbert 1978:60 [ref. 7042], Gilbert 1998:111 [ref. 23395]. •Synonym of Notropis formosus (Girard 1856) -- (Gilbert 1978:60 [ref. 7042], Chernoff & Miller 1982:519 [ref. 8493]). •Synonym of Cyprinella formosa (Girard 1856) -- (Mayden 1989:55 [ref. 12555], Gilbert 1998:111 [ref. 23395], Minckley & Marsh 2009:120 [ref. 31114]). •Synonym of Cyprinella formosa (Girard 1856), but a valid subspecies -- (Scharpf 2005:13 [ref. 28940]). Current status: Synonym of Cyprinella formosa (Girard 1856). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

montiregis, Cliola Cope [E. D.] 1885:168 [Proceedings of the American Philosophical Society v. 22 (for 19 Dec. 1884); ref. 18038] Monterrey, Nuevo León, Mexico. Holotype (unique): ANSP 19344. Type catalog: Böhlke 1984:84 [ref. 13621], Gilbert 1998:117 [ref. 23395]. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:62 [ref. 7042]). •Synonym of Cyprinella rutila (Girard 1856) -- (Mayden 1989:60 [ref. 12555], Gilbert 1998:1176 [ref. 23395]). Current status: Synonym of Cyprinella rutila (Girard 1856). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

niveus, Hybopsis Cope [E. D.] 1870:460 [Proceedings of the American Philosophical Society v. 11 (no. 84); ref. 913] Upper waters of Catawba River, North Carolina, U.S.A. Lectotype: ANSP 2930. Paralectotypes: ANSP 2931-47 (17), 2948 (1, missing). Type catalog: Gilbert 1978:64 [ref. 7042], Böhlke 1984:85 [ref. 13621]. Lectotype established by Fowler 1910: Pl. 18 (fig. 27) [ref. 19840]. •Valid as Notropis niveus (Cope 1870) -- (Gilbert 1978:64 [ref. 7042], Lee et al. 1980:288 [ref. 22416], Cloutman & Harrell 1987:1037 [ref. 22448]). •Valid as Cyprinella nivea (Cope 1870) -- (Mayden 1989:92 [ref. 12555], Page & Burr 1991:127 [ref. 18983], Gilbert 1998:121 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:14 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:247 [ref. 31215], Page et al. 2013:70 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella nivea (Cope 1870). Leuciscidae: Pogonichthyinae. Distribution: North America: North Carolina and Georgia (southeastern U.S.A.). Habitat: freshwater.

notata, Cyprinella Girard [C. F.] 1856:198 [34] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Rio Seco (= Seco Creek) at D'Hanis, a tributary to Rio Nueces, Medina County, Texas, U.S.A. Lectotype: USNM 136 (plus pharyngeal arch as # 2715). Paralectotypes: ?USNM 162720 (1, not this species), 163955 [ex USNM 136] (1). Type catalog: Gilbert 1978:65 [ref. 7042], Gilbert 1998:122 [ref. 23395]. On p. 34 of separate. Lectotype selected by Gibbs 1957:180 [ref. 21244]. •Synonym of Notropis venustus (Girard 1856) -- (Gibbs 1957:179 [ref. 21244], Gilbert 1978:65 [ref. 7042]). •Synonym of Cyprinella venusta Girard 1856 -- (Mayden 1989:77 [ref. 12555]). •Synonym of Cyprinella venusta Girard 1856, subspecies venusta -- (Gilbert 1998:122 [ref. 23395]). Current status: Synonym of Cyprinella venusta Girard 1856. Leuciscidae: Pogonichthyinae. Habitat: freshwater.

panarcys, Notropis Hubbs [C. L.] & Miller [R. R.] 1978:582, Figs. 1-3 [Copeia 1978 (no. 4); ref. 8924] Río San Pedro at Meoqui, along highway 45, Chihuahua, Mexico, elevation 1115 meters. Holotype: UMMZ 198785. Paratypes: FMNH 83854 (10); TNHC 4903 (2); CAS-SU 47604 (11 or 22); UMMZ 161173 (4), 164695-96 (11), 198786 (100), USNM 218221 (10). Type catalog: Ibarra & Stewart 1987:61 [ref. 12367], Gilbert 1998:129 [ref. 23395]. •Valid as Cyprinella panarcys (Hubbs & Miller 1978) -- (Mayden 1989:65 [ref. 12555], Espinosa Pérez et al. 1993:12 [ref. 22290], Contreras-Balderas & Lozano-Vilano 1994:904 [ref. 22061], Gilbert 1998:129 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:14 [ref. 28940], Miller 2006:112 [ref. 28615], Schönhuth & Mayden 2009:78 [ref. 38584], Page et al. 2013:70 [ref. 32708], Pinion et al. 2018:1124 [ref. 36371], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella panarcys (Hubbs & Miller 1978). Leuciscidae: Pogonichthyinae. Distribution: Southern North America: northern Mexico, north to Rio Grande (Texas, U.S.A.). Habitat: freshwater.

proserpina, Moniana Girard [C. F.] 1856:200 [36] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Devils River, just about mouth into Rio Grande, Val Verde County, Texas, U.S.A. Syntypes: ANSP 3026 [ex USNM 117] (1); BMNH 1883.12.14.216 [ex USNM 117] (1); MCZ 1688 [ex USNM 117] (2); MNHN 0000-0433 [ex USNM 117] (2); USNM 117 (now 9, plus 1 phyrngeal arch as #2702), 125084 [ex USBF 100] (4). Type catalog: Gilbert 1978:72 [ref. 7042], Böhlke 1984:88 [ref. 13621], Gilbert 1998:135-136 [ref. 23395]. On p. 36 of separate. •Valid as Notropis proserpinus (Girard 1856) -- (Gilbert 1978:73 [ref. 7042], Lee et al. 1980:299 [ref. 22416]). •Valid as Cyprinella proserpina (Girard 1856) -- (Mayden 1989:63 [ref. 12555], Espinosa Pérez et al. 1993:12 [ref. 22290], Gilbert 1998:136 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:14 [ref. 28940], Miller 2006:113 [ref. 28615], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:241 [ref. 31215], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella proserpina (Girard 1856). Leuciscidae: Pogonichthyinae. Distribution: Southern North America: Texas (U.S.A.) and Río Grande system (Mexico). Habitat: freshwater.

pulchella, Moniana Girard [C. F.] 1856:200 [36] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Sugar Loaf Creek, tributary to Poteau River, about 20-25 miles south of Fort Smith, near Poteay, Oklahoma-Arkansas line, U.S.A. Girard specimens: ANSP 4085 [ex USNM 121] (1); MCZ 1797 [ex USNM 121] (2); MNHN 0000-0380 [ex USNM 121]; USNM 121 (now 0, 3 pharyngeal archesas # 2706), 122 (orig. 1, now 0). Type catalog: Gilbert 1978:73 [ref. 7042], Böhlke 1984:88 [ref. 13621], Gilbert 1998:136 [ref. 23395]. On p. 36 of separate. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:73 [ref. 7042]). •Synonym of Cyprinella lutrensis, subspecies lutrensis (Baird & Girard 1853) -- (Mayden 1989:50 [ref. 12555], Gilbert 1998:136 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

pyrrhomelas, Photogenis Cope [E. D.] 1870:463 [Proceedings of the American Philosophical Society v. 11 (no. 84); ref. 913] Tributary of upper Catawba River, North Carolina, U.S.A. Lectotype: ANSP 2631. Paralectotypes: ANSP 2632-2725 (94). Type catalog: Gilbert 1978:74 [ref. 7042], Böhlke 1984:88 [ref. 13621], Gilbert 1998:139 [ref. 23395]. Lectotype established by Fowler 1910:Pl. 18 (fig. 29) [ref. 19840]. •Valid as Notropis pyrrhomelas (Cope 1870) -- (Gilbert 1978:74 [ref. 7042], Lee et al. 1980:300 [ref. 22416]). •Valid as Cyprinella pyrrhomelas (Cope 1870) -- (Mayden 1989:82 [ref. 12555], Page & Burr 1991:126 [ref. 18983], Gilbert 1998:139 [ref. 23395], Fuller et al. 1999:65 [ref. 25838], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:14 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:245 [ref. 31215], Stout et al. 2022:7 [ref. 39738], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella pyrrhomelas (Cope 1870). Leuciscidae: Pogonichthyinae. Distribution: North America: North Carolina and South Carolina (U.S.A.). Introduced in Georgia (U.S.A.). Habitat: freshwater.

rubripinna, Cyprinella Garman [S.] 1881:91 [Bulletin of the Museum of Comparative Zoology v. 8 (no. 3); ref. 1530] Lago del Muerto, near Parras, 25°25'N, 102°11'W, Coahuila, Mexico. Syntypes: MCZ 25891 (2), 24892 (orig. 26, now 20); SMNS 3034 (2); UMMZ 86663 [ex MCZ 24892] (6); USNM 120257 [ex MCZ 14892] (6). Type catalog: Fricke 1995:11 [ref. 28100], Gilbert 1998:144 [ref. 23395], Fricke 2005:31 [ref. 29864]. Subjectively invalid; secondarily preoccupied in Notropis by Argyreus rubripinnis Heckel 1843; replaced by Notropis garmani Jordan 1885. Fricke 2005:32 [ref. 29864] suggests the name is available as the homonymy no longer exists. As the species is currently classified in genus Cyprinella, the original name is here reinstated. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:76 [ref. 7042]). •Synonym of Cyprinella garmani (Jordan 1885) -- (Mayden 1989:53 [ref. 12555], Gilbert 1998:144 [ref. 23395]). •Valid as Cyprinella rubripinna Garman 1881 – (Fricke 2005: 32 [ref. 29864]). 2°Homonym. Current status: Valid as Cyprinella rubripinna Garman 1881. Leuciscidae: Pogonichthyinae. Distribution: Southern North America: Coahuila (Mexico). Habitat: freshwater.

rutila, Moniana Girard [C. F.] 1856:201 [37] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Río Monterrey (tributary to Río San Juan), Cadereita [Cadereyta], Nuevo León, Mexico. Syntypes: (10) [all originally USNM 95] ANSP 4086 (1, in several pieces), MCZ 1794 (1), MNHN 0000-0429 (2), USNM 95 (orig. 10+, now 0, 2 sets of pharyngeal arches as #2698). Type catalog: Gilbert 1978:77 [ref. 7042], Böhlke 1984:90 [ref. 13621]. On p. 37 of separate. •Valid as Notropis rutilus (Girard 1856) -- (Gilbert 1978:77 [ref. 7042]). •Valid as Cyprinella rutila (Girard 1856) -- (Mayden 1989:60 [ref. 12555], Espinosa Pérez et al. 1993:12 [ref. 22290], Contreras-Balderas & Lozano-Vilano 1994:904 [ref. 22061], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:14 [ref. 28940], Miller 2006:113 [ref. 28615], Schönhuth & Mayden 2009:78 [ref. 38584], Page et al. 2013:70 [ref. 32708], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella rutila (Girard 1856). Leuciscidae: Pogonichthyinae. Distribution: Southern North America: Atlantic Slope of Mexico. Habitat: freshwater.

santamariae, Notropis Evermann [B. W.] & Goldsborough [E. L.] 1902:147, Fig. 2 [Bulletin of the U. S. Fish Commission v. 21 [1901]; ref. 14875] Small pool near Lake Santa María, Chihuahua, Mexico. Holotype: USNM 50002. Paratypes: USNM 126485 [ex USBF 964] (1). Type catalog: Gilbert 1978:77-78 [ref. 7042], Gilbert 1998:145 [ref. 23395]. •Synonym of Notropis lutrensis (Baird & Girard 1853), but a valid subspecies -- (Gilbert 1978:78 [ref. 7042]). •Synonym of Notropis formosus (Girard 1856) -- (Chernoff & Miller 1982:519 [ref. 8493]). •Synonym of Cyprinella formosa (Girard 1856) -- (Mayden 1989:55 [ref. 12555], Gilbert 1998:145 [ref. 23395], Minckley & Marsh 2009:120 [ref. 31114]). •Synonym of Cyprinella lutrensis ssp. -- (Espinosa Pérez et al. 1993:12 [ref. 22290]). •Synonym of Cyprinella formosa (Girard 1856), but a valid subspecies -- (Scharpf 2005:13 [ref. 28940]). Current status: Synonym of Cyprinella formosa (Girard 1856). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

spilopterus, Photogenis Cope [E. D.] 1867:378 [Transactions of the American Philosophical Society (New Series) v. 13 (pt 3) (art. 13); ref. 910] St. Josephs River, southwestern Michigan, U.S.A. Holotype or lectotype: ANSP 22901 (1). Type catalog: Gilbert 1978:81 [ref. 7042], Böhlke 1984:91 [ref. 13621], Gilbert 1998:149-150 [ref. 23395]. Also appeared as Leuciscus spilopterus Cope in Günther 1868:254 [ref. 1990]. Lectotype not designated by Gibbs 1957:193 [ref. 12065]; see Gilbert 1998:149-150 [ref. 23395]; apparently 1 specimen originally. •Valid as Notropis spilopterus (Cope 1867) -- (Gibbs 1957:191 [ref. 12065], Gilbert 1978:81 [ref. 7042], Lee et al. 1980:312 [ref. 22416], Smith 1986:180 [ref. 22186], Schaefer & Cavender 1986:122 [ref. 22457], Robison & Buchanan 1988:232 [ref. 23393], McAllister 1990:63 [ref. 14674], Scott & Crossman 1998:466 [ref. 24570]). •Valid as Cyprinella spiloptera (Cope 1867) -- (Mayden 1989:66 [ref. 12555], Page & Burr 1991:120 [ref. 18983], Boschung 1992:47 [ref. 23239], Etnier & Starnes 1993:155 [ref. 22809], Jenkins & Burkhead 1994:365 [ref. 21581], Stauffer et al. 1995:71 [ref. 23376], Cross & Collins 1995:58 [ref. 23394], Mettee et al. 1996:171 [ref. 25836], Murdy et al. 1997:87 [ref. 23144], Gilbert 1998:150 [ref. 23395], Briolay et al. 1998:101 [ref. 25586], Fuller et al. 1999:65 [ref. 25838], Lyons et al. 2000:23 [ref. 26594], Ross et al. 2001:144 [ref. 25978], Nelson et al. 2004:69 [ref. 27807], Bailey et al. 2004:75 [ref. 27882], Miller & Robison 2004:94 [ref. 27931], Boschung & Mayden 2004:171 [ref. 27995], Scharpf 2005:14 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:236 [ref. 31215], Page et al. 2013:70 [ref. 32708], Bagley et al. 2023:313 [ref. 41241], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella spiloptera (Cope 1867). Leuciscidae: Pogonichthyinae. Distribution: North America: northeastern U.S.A. and southeastern Canada. Habitat: freshwater.

stigmaturus, Photogenis Jordan [D. S.] 1877:337 [Annals of the Lyceum of Natural History of New York v. 11 (nos 11-12) (art. 29); ref. 2373] Tributary of Etowah River (probably Silver Creek), near Rome, Floyd County, Georgia, U.S.A. Lectotype: USNM 17891. Paralectotypes: ANSP 19820-27 (8); MCZ 24371 (1); MNHN A-1283 (2); UF 173465 [ex Bulter Univ. 62] (55), 173474 [ex Butler Univ. 62] (17); USNM 20125 (1), 163938 (10), 163981 (1). Type catalog: Bertin & Estève 1948:66 [ref. 19574], Gilbert 1978:81 [ref. 7042], Böhlke 1984:92 [ref. 13621], Gilbert 1998:151 [ref. 23395], Gilbert 2009:158 [ref. 30728]. Lectotype selected by Gibbs 1957:192 [ref. 21244]. Gilbert 2009:159 [ref. 30728] reidentified UF 173474. •Synonym of Notropis venustus (Girard 1856), but a valid subspecies -- (Gibbs 1957:192 [ref. 21244], Gilbert 1978:81 [ref. 7042], Lee et al. 1980:321 [ref. 22416], Robison & Buchanan 1988:239 [ref. 23393]). •Synonym of Cyprinella venusta (Girard 1856) -- (Mayden 1989:78 [ref. 12555]). •Valid as Cyprinella stigmatura (Jordan 1877) -- (Gilbert 1998:151 [ref. 23395]). •Synonym of Cyprinella venusta (Girard 1856), but a valid subspecies -- (Ross et al. 2001:150 [ref. 25978], Scharpf 2005:14 [ref. 28940], Page & Burr 2011:239 [ref. 31215]). Current status: Valid as Cyprinella stigmatura (Jordan 1877). Leuciscidae: Pogonichthyinae. Distribution: Eastern U.S.A.: eastern part of range [subspecies stigmaturus]. Habitat: freshwater.

suavis, Cyprinella Girard [C. F.] 1856:197 [33] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Near San Antonio, Bexar County, Texas, U.S.A. Syntypes: ANSP 2769-70 [ex USNM 138] (2), MCZ 1694 [ex USNM 138] (2), MNHN 0000-0375 [ex USNM 138] (2), USNM 138 (orig. 20, now 0, 1 pair of pharyngeal arches as #2717). Type catalog: Gilbert 1978:79 [ref. 7042], Böhlke 1984:92 [ref. 13621]. On p. 33 of separate. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:82 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853), but a valid subspecies -- (Matthews 1987:616 [ref. 6774]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853) -- (Mayden 1989:49 [ref. 12555]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853), subspecies lutrensis -- (Gilbert 1998:153 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

trichroistia, Codoma Jordan [D. S.] & Gilbert [C. H.] in Jordan & Brayton 1878:50 [Bulletin of the United States National Museum No. 12; ref. 2436] Tributary of Etowah River, Georgia, U.S.A. (probably near Rome, Floyd County). Lectotype: USNM 31131. Paralectotypes: MCZ 24393 (1), MNHN A-1280 (1), USNM 163956 [ex USNM 31131] (6). Type catalog: Bertin & Estève 1948:67 [ref. 19574], Gilbert 1978:84 [ref. 7042], Gilbert 1998:157-158 [ref. 23395], Gilbert 2009:158 [ref. 30728]. Lectotype selected by Howell & Williams 1971:63 [ref. 18313]. •Valid as Notropis trichroistia (Jordan & Gilbert 1878) -- (Gilbert 1978:84 [ref. 7042], Lee et al. 1980:318 [ref. 22416]). •Valid as Cyprinella trichroistia (Jordan & Gilbert 1878) -- (Mayden 1989:89 [ref. 12555], Page & Burr 1991:125 [ref. 18983], Boschung 1992:47 [ref. 23239], Etnier & Starnes 1993:157 [ref. 22809], Mettee et al. 1996:173 [ref. 25836], Gilbert 1998:158 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Boschung & Mayden 2004:172 [ref. 27995], Scharpf 2005:14 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:243 [ref. 31215], Page et al. 2013:70 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Bagley et al. 2023:313 [ref. 41241], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella trichroistia (Jordan & Gilbert 1878). Leuciscidae: Pogonichthyinae. Distribution: North America: Tennessee, Georgia and Alabama (southeastern U.S.A.). Habitat: freshwater.

umbrosa, Cyprinella Girard [C. F.] 1856:197 [33] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Coal Creek, tributary to south fork of Canadian River, Arkansas, and also 20 miles west of Choctaw Agency, U.S.A. Syntypes: ANSP 2830 [ex USNM 134] (1); MCZ 1800 [ex USNM 134] (1); MNHN 0000-0373 [ex USNM 134] (1), USNM 133 (orig. 8, now 6, plus 1 pair pharyngeal arches as #2712) west of Choctaw Agency, 134 (orig. 8, now 0, plus 1 pair pharyngeal arches as #2713) Coal Creek. Type catalog: Gilbert 1978:86 [ref. 7042], Böhlke 1984:94 [ref. 13621], Gilbert 1998:160 [ref. 23395]. On p. 33 of separate. •Synonym of Notropis lutrensis (Baird & Girard 1853) -- (Gilbert 1978:86 [ref. 7042]). •Synonym of Cyprinella lutrensis (Baird & Girard 1853), subspecies lutrensis -- (Mayden 1989:49 [ref. 12555], Gilbert 1998:160 [ref. 23395]). Current status: Synonym of Cyprinella lutrensis (Baird & Girard 1853). Leuciscidae: Pogonichthyinae. Habitat: freshwater.

urostigma, Cliola Jordan [D. S.] & Meek [S. E.] 1884:475 [Proceedings of the United States National Museum v. 7 (no. 450); ref. 10636] San Saba River (tributary to Colorado River) at Fort McKavit [= McKavett], Menard County, Texas, U.S.A. Lectotype: USNM 20466. Paralectotypes: MCZ 35903 [ex USNM 20446] (1, disintegrated), 35948 [ex USNM 17812] (1); USNM 163952 [ex 20446] (orig. 20, now 8), 17812 (now 7). Type catalog: Gilbert 1978:86-87 [ref. 7042], Gilbert 1998:160 [ref. 23395]. •Synonym of Notropis venustus (Girard 1856) -- (Gibbs 1957:179 [ref. 21244], Gilbert 1978:87 [ref. 7042]). •Synonym of Cyprinella venusta Girard 1856 -- (Mayden 1989:78 [ref. 12555], Gilbert 1998:160 [ref. 23395]). Current status: Synonym of Cyprinella venusta Girard 1856. Leuciscidae: Pogonichthyinae. Habitat: freshwater.

venusta, Cyprinella Girard [C. F.] 1856:198 [34] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Rio Sabinal at Sabinal, Uvalde County, Texas, U.S.A. Lectotype: ANSP 3035 [ex USNM 140]. Paralectotypes: ANSP 3036 [ex USNM 140] (1), MCZ 1695 [ex USNM 140] (2), MNHN 0000-0376 [ex USNM 140] (2), USNM 140 (orig. 15+, now 0, 1 pair pharyngeal arches as #2719). Type catalog: Gilbert 1978:87 [ref. 7042], Böhlke 1984:94 [ref. 13621], Gilbert 1998:161 [ref. 23395]. On p. 34 of separate. Lectotype selected by Gibbs 1957:180 [ref. 21244]. •Valid as Notropis venustus (Girard 1856), subspecies venustus -- (Gibbs 1957:179 [ref. 21244], Gilbert 1978:87 [ref. 7042], Lee et al. 1980:321 [ref. 22416], Dimmick 1987:316 [ref. 11406], Robison & Buchanan 1988:238 [ref. 23393]). •Valid as Cyprinella venusta Girard 1856, subspecies venusta [most authors as subspecies] -- (Mayden 1989:77 [ref. 12555], Page & Burr 1991:122 [ref. 18983], Boschung 1992:47 [ref. 23239], Etnier & Starnes 1993:158 [ref. 22809], Taylor et al. 1994:72 [ref. 21896], Warren et al. 1994:132 [ref. 25557], Knight & Hastings 1994:149 [ref. 25560], Richardson & Gold 1995:32 [ref. 22412], Schmidt & Gold 1995:200 [ref. 22413], Kristmundsdóttir & Gold 1996:773 [ref. 22599], Mettee et al. 1996:175 [ref. 25836], Simons & Mayden 1997:195 [ref. 24948], Gilbert 1998:161 [ref. 23395], Fuller et al. 1999:66 [ref. 25838], Ross et al. 2001:147 [ref. 25978]). •Valid as Cyprinella venusta Girard 1856 -- (Nelson et al. 2004:69 [ref. 27807], Miller & Robison 2004:96 [ref. 27931], Boschung & Mayden 2004:173 [ref. 27995], Scharpf 2005:14 [ref. 28940] with subspecies, Miller 2006:114 [ref. 28615], Minckley & Marsh 2009:179 [ref. 31114], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:238 [ref. 31215] with 3 subspecies, Page et al. 2013:70 [ref. 32708], Robins et al. 2018:84 [ref. 35886] with 3 subspecies, Stout et al. 2022:7 [ref. 39738], Bagley et al. 2023:313 [ref. 41241], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella venusta Girard 1856. Leuciscidae: Pogonichthyinae. Distribution: North America: Florida to western Texas (southeastern U.S.A.). Mississippi River and drainages to the west [subspecies venusta]. Gulf drainages (except Mobile Bay) east of the Mississippi River [subspecies cercostigma]. Habitat: freshwater.

whipplii, Cyprinella Girard [C. F.] 1856:198 [34] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Sugarloaf Creek, tributary of Poteau River, about 20-25 miles south of Fort Smith, Arkansas, vicinity of Poteau, Oklahoma, U.S.A. Lectotype: USNM 137 (plus 1 pharyngeal arch as #2716). Paralectotypes: lost or none collected. Type catalog: Gilbert 1998:164-165 [ref. 23395]. On p. 34 of separate. Originally as whipplii, presumably named for Lt. A. W. Whipple by authors; but the spelling whipplii is correctly formed, but the spelling whipplei probably can be retained on the basis of Art. 33.2.3.1 [or Art. 33.3.1] of the Code. Lectotype selected by Gibbs 1963:513 [ref. 21327] but other syntypes unknown. •Valid as Notropis whipplei (Girard 1856) -- (Gibbs 1963:511 [ref. 21327], Gilbert 1978:88 [ref. 7042], Lee et al. 1980:324 [ref. 22416], Robison & Buchanan 1988:241 [ref. 23393]). •Valid as Cyprinella whipplei Girard 1856 -- (Mayden 1989:71 [ref. 12555], Burkhead & Jenkins 1991:348 [ref. 21804], Page & Burr 1991:120 [ref. 18983], Boschung 1992:47 [ref. 23239], Etnier & Starnes 1993:159 [ref. 22809], Jenkins & Burkhead 1994:359 [ref. 21581], Stauffer et al. 1995:73 [ref. 23376], Mettee et al. 1996:177 [ref. 25836], Gilbert 1998:165 [ref. 23395], Fuller et al. 1999:67 [ref. 25838], Ross et al. 2001:150 [ref. 25978], Nelson et al. 2004:69 [ref. 27807], Miller & Robison 2004:98 [ref. 27931], Boschung & Mayden 2004:175 [ref. 27995], Scharpf 2005:14 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:236 [ref. 31215], Page et al. 2013:70 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Bagley et al. 2023:313 [ref. 41241], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella whipplei Girard 1856. Leuciscidae: Pogonichthyinae. Distribution: Eastern North America: eastern-central U.S.A. Habitat: freshwater.

xaenurus, Minnilus Jordan [D. S.] 1877:79 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 29 (pt 1); ref. 2372] South Fork of Ocmulgee River at Flat Shoals, DeKalb County, Georgia, U.S.A. Lectotype: USNM 17862. Paralectotypes: BMNH 1880.1.21.82 (1), CAS-SU 1996 (1); MCZ 24365 (1), MNHN A-1313 (2), CAS-SU 1996 (1), UF 165650 [ex Butler Univ. 63] (49), 173579 [ex Butler Univ. 63] (225); USNM 163957 [ex USNM 17862] (4), 329284 (7). Type catalog: Bertin & Estève 1948:67 [ref. 19574], Böhlke 1953:34 [ref. 12291], Gilbert 1978:89 [ref. 7042], Gilbert 1998:165-166 [ref. 23395], Gilbert 2009:161 [ref. 30728]. Lectotype established in Gilbert 1978:89-90 [ref. 7042]. Gilbert 2009:161 [ref. 30728] reidentified UF 165650. •Valid as Notropis xaenurus (Jordan 1877) -- (Gilbert 1978:90 [ref. 7042], Lee et al. 1980:326 [ref. 22416]). •Valid as Cyprinella xaenura (Jordan 1877) -- (Mayden 1989:84 [ref. 12555], Page & Burr 1991:126 [ref. 18983], Gilbert 1998:166 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:14 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:245 [ref. 31215], Page et al. 2013:70 [ref. 32708], Stout et al. 2022:7 [ref. 39738], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella xaenura (Jordan 1877). Leuciscidae: Pogonichthyinae. Distribution: North America: Georgia (eastern U.S.A.). Habitat: freshwater.

xanthicara, Notropis Minckley [W. L.] & Lytle [G. L.] 1969:491, Fig. 1 (upper 2) [Proceedings of the Biological Society of Washington v. 82; ref. 21391] Río Puente Colorado, 8.5 kilometers south and 0.7 kilometers west of Cuatro Ciénegas, Coahuila, Mexico. Holotype: UMMZ 188782. Paratypes: ASU 969 (8), 2316 (185), 3728 (68); KU 7404 (72); UMMZ 179202 (46), 179834 (11), 179860 (53); UNAL 703 (6), 709 (105). Additional material: ASU (5 lots); KU (6 lots); UMMZ (4 lots); UNL (1 lot). Type catalog: Gilbert 1978:90 [ref. 7042], Contreras-Balderas et al. 1998:3 [ref. 23752]. •Valid as Notropis xanthicara Minckley & Lytle 1969 -- (Gilbert 1978:90 [ref. 7042], Gilbert 1998:166 [ref. 23395]). •Valid as Cyprinella xanthicara (Minckley & Lytle 1969) -- (Mayden 1989:61 [ref. 12555], Espinosa Pérez et al. 1993:12 [ref. 22290], Contreras-Balderas & Lozano-Vilano 1994:904 [ref. 22061], Gilbert 1998:166 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:15 [ref. 28940], Miller 2006:114 [ref. 28615], Schönhuth & Mayden 2009:78 [ref. 38584], Page et al. 2013:70 [ref. 32708], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella xanthicara (Minckley & Lytle 1969). Leuciscidae: Pogonichthyinae. Distribution: Southern North America: interior and Atlantic Slope of Mexico. Habitat: freshwater.

zanemus, Ceratichthys Jordan [D. S.] & Brayton [A. W.] 1878:24 [Bulletin of the United States National Museum No. 12; ref. 2436] Saluda River, at Farr's Mills, west of Greenville, Greenville County, South, U.S.A. Lectotype: USNM 31126 (poor condition). Paralectotypes: none found. Type catalog: Gilbert 1978:92 [ref. 7042], Gilbert 1998:166-167 [ref. 23395]. Lectotype established in Gilbert 1978:92 [ref. 7042]. •Valid as Hybopsis zanema (Jordan & Brayton 1878) -- (Gilbert 1978:92 [ref. 7042], Lee et al. 1980:197 [ref. 22416], Page & Burr 1991:162 [ref. 18983], Grose & Wiley 2002:1092 [ref. 26513]). •Valid as Cyprinella zanema (Jordan & Brayton 1878) -- (Gilbert 1998:166-167 [ref. 23395], Nelson et al. 2004:69 [ref. 27807], Scharpf 2005:15 [ref. 28940], Schönhuth & Mayden 2009:78 [ref. 38584], Page & Burr 2011:248 [ref. 31215], Page et al. 2013:70 [ref. 32708], Page et al. 2023:69 [ref. 40505]). Current status: Valid as Cyprinella zanema (Jordan & Brayton 1878). Leuciscidae: Pogonichthyinae. Distribution: North America: North Carolina and South Carolina (eastern U.S.A.). Habitat: freshwater.


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