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Eschmeyer's Catalog of Fishes
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Species in the genus Gila: [ 51 ] records


affinis, Gila Abbott [C. C.] 1860:474 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 12; ref. 15618] Bridger Pass, Wyoming, U.S.A. Lectotype: ANSP 4196. Paralectotype: USNM 3557 (1) Type catalog: Böhlke 1984:68 [ref. 13621] with "Kansas" locality in error, Gilbert 1998:35-36 [ref. 23395]. Lectotype established by Fowler 1925:398, fig. 6. •Synonym of Gila robusta Baird & Girard 1853 -- (Gilbert 1998:36 [ref. 23395], Copus et al. 2018:11 [ref. 36179]). Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

alutaceus, Acrocheilus Agassiz [L.] & Pickering [C.] in Agassiz 1855:99 [29] [American Journal of Science and Arts (Series 2) v. 19 (no. 55); ref. 71] Willamette Falls, Oregon City, Clackamas County, Oregon; Wallawalla River at Whitman, Walla Walla County, Washington, U.S.A. Syntypes: USNM 83357 (2, 1 a skin) Walla Walla R. Type catalog: Gilbert 1998:38 [ref. 23395]. On p. 29 of separate. •Valid as Acrocheilus alutaceus Agassiz & Pickering 1855 -- (Lee et al. 1980:140 [ref. 22416], McAllister 1990:58 [ref. 14674], Page & Burr 1991:70 [ref. 18983], La Rivers 1994:384 [ref. 23389], Simons & Mayden 1997:195 [ref. 24948], Gilbert 1998:38 [ref. 23395], Scott & Crossman 1998:386 [ref. 24570], Coffie 1998:154 [ref. 25003], Wydoski & Whitney 2003:118 [ref. 27816], Nelson et al. 2004:68 [ref. 27807], Scharpf 2005:10 [ref. 28940], McPhail 2007:56 [ref. 31199], Page & Burr 2011:163 [ref. 31215], Page et al. 2013:68 [ref. 32708]). •Valid as Gila alutacea (Agassiz & Pickering 1855) -- (Page et al. 2023:70 [ref. 40505]). Current status: Valid as Gila alutacea (Agassiz & Pickering 1855). Leuciscidae: Laviniinae. Distribution: Western North America: British Columbia (Canada) and western U.S.A. states. Habitat: freshwater.

atraria, Siboma Girard [C. F.] 1856:208 [44] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Fish Spring, Tooele County, Utah, U.S.A. Holotype (unique): USNM 236 (pharangeas areches as # 2789). Type catalog: Gilbert 1998:45 [ref. 23395]. On p. 44 of separate. Name placed on Official List in Opinion 1872. •Valid as Gila atraria (Girard 1856) -- (Lee et al. 1980:160 [ref. 22416], Page & Burr 1991:76 [ref. 18983], La Rivers 1994:395 [ref. 23389], Baxter & Stone 1995:68 [ref. 23374], Andersen & Deacon 1996:124 [ref. 24717], Sigler & Sigler 1996:74 [ref. 25943], Gilbert 1998:45 [ref. 23395], Fuller et al. 1999:72 [ref. 25838], Broughton 2000:651 [ref. 24593], Johnson 2002:948 [ref. 26052], Nelson et al. 2004:70 [ref. 27807], Scharpf 2005:18 [ref. 28940], Minckley & Marsh 2009:180 [ref. 31114], Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505]). Current status: Valid as Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Distribution: Western North America: western U.S.A. Habitat: freshwater.

bicolor, Tigoma Girard [C. F.] 1856:206 [42] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Klamath Lake, Oregon, U.S.A. Syntypes: USNM 234 (3, plus one pair pharyngeal arches as # 2788). Type catalog: Gilbert 1998:49 [ref. 23395]. Jordan & Evermann 1896:232 [ref. 2443] qualify as first revisers, selecting Tigoma bicolor over Cheonda coerulea (see Bailey & Uyeno 1964:238 [ref. 23390]). Bailey & Uyeno 1964:238 [ref. 23390] serve as first revisers selecting Algansea bicolor over Tigoma bicolor only when both are in Gila; therefore, Cheonda coerulea was reinstated. As this name was used as valid for more than 50 years, it is now retained even when in Klamathella for the sake of stability. •Synonym of Gila coerulea (Girard 1856) -- (Bailey & Uyeno 1964:239 [ref. 23390], Gilbert 1998:49 [ref. 23395], Nelson et al. 2004:70 [ref. 27807]). Current status: Synonym of Gila coerulea (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

bivittatus, Hybopsis Cope [E. D.] 1872:474 [U. S. Geological Survey Territitories Annual Report v. 5 (pt 4); ref. 923] Warm Springs, Utah, U.S.A. No types known. Type catalog: Böhlke 1984:70 [ref. 13621], Gilbert 1998:49-50 [ref. 23395]. •Synonym of Gila atraria (Girard 1856) -- (La Rivers 1994:395 [ref. 23389], Gilbert 1998:49-50 [ref. 23395]). Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

brevicauda, Gila Norris [S. M.], Fischer [J. M.] & Minckley [W. L.] 2003:20, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 14 (no. 1); ref. 26757] Río Basaseachic (= Candameña), 1-2 kilometers above Cascada de Basaseachic, Chihuahua, Mexico. Holotype: UMMZ 240260. Paratypes: ASU 8447 (4), 9936 (4); UMMZ 208179 (86), 21311 [ex UAZ 75-102] (39); UANL 15012 (8); UAZ 75-103 (32); UNAM 13105 (8). •Valid as Gila brevicauda Norris, Fischer & Minckley 2003 -- (Nelson et al. 2004:70 [ref. 27807], Scharpf 2005:18 [ref. 28940], Miller et al. 2006:120 [ref. 28615], Minckley & Marsh 2009:120 [ref. 31114], Page et al. 2013:71 [ref. 32708], Ballesteros-Córdova et al. 2020:216 [ref. 37371], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:172 [ref. 41622]). Current status: Valid as Gila brevicauda Norris, Fischer & Minckley 2003. Leuciscidae: Laviniinae. Distribution: Southern North America: Mayo River basin, Chihuahua (Mexico) endemic. Habitat: freshwater.

caerulea, Cheonda Girard [C. F.] 1857:30 [Report on fishes collected on the survey. Route near the 38th and 39th parallels; ref. 21360] Lost River, Oregon. See Cheonda coerulea, not a new description. Misspelling. Current status: Synonym of Gila coerulea (Girard 1856). Leuciscidae: Laviniinae.

coerulea, Cheonda Girard [C. F.] 1856:207 [43] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Lost River, Oregon, U.S.A. Holotype (unique): USNM 237 (plus pharyngeal arches as #2790). Type catalog: Gilbert 1998:61 [ref. 23395]. On p. 43 of separate. Clearly spelled "oe" in text and index [see "ae" spelling for other species in same work]. •Valid as Klamathella coerulea (Girard 1856) -- (Smith et al. 2002: 211 [ref. 36187], Tan & Armbruster 2018:18 [ref. 36121]). •Valid as Gila coerulea (Girard 1856) -- (Lee et al. 1980:162 [ref. 22416], Page & Burr 1991:76 [ref. 18983], Gilbert 1998:61-62 [ref. 23395], Fuller et al. 1999:74 [ref. 25838], Moyle 2002:128 [ref. 26449], Nelson et al. 2004:70 [ref. 27807], Scharpf 2005:18 [ref. 28940], Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505]). Current status: Valid as Gila coerulea (Girard 1856). Leuciscidae: Laviniinae. Distribution: Western North America: Oregon and California (U.S.A.). Habitat: freshwater.

conformis, Lavinia Baird [S. F.] & Girard [C. F.] in Girard 1854:137 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] Posa Creek, just north of Bakersfield, San Joaquin basin, California, U.S.A. Holotype (unique): USNM 231 (plus 1 pharyngeal arch as #2785). Type catalog: Gilbert 1998:64 [ref. 23395]. Author is S. F. Baird and Girard. •Synonym of Gila crassicauda (Baird & Girard 1854) -- (Gilbert 1998:64 [ref. 23395]). Current status: Synonym of Gila crassicauda (Baird & Girard 1854). Leuciscidae: Laviniinae. Habitat: freshwater.

conspersa, Gila Garman [S.] 1881:91 [Bulletin of the Museum of Comparative Zoology v. 8 (no. 3); ref. 1530] Río Nazas, probably state of Coahuila, Mexico. Syntypes: MCZ 24879-80 (9, 1); UMMZ 86635 [ex MCZ 24879] (3); USNM [ex MCZ 24879] (3). Type catalog: Gilbert 1998:64 [ref. 23395]. •Valid as Gila conspersa Garman 1881 -- (Espinosa Pérez et al. 1993:15 [ref. 22290], Gilbert 1998:64 [ref. 23395], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:70 [ref. 27807], Miller et al. 2006:120 [ref. 28615], Scharpf 2005:18 [ref. 28940], Page et al. 2013:71 [ref. 32708], Ballesteros-Córdova et al. 2020:216 [ref. 37371], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:172 [ref. 41622]). Current status: Valid as Gila conspersa Garman 1881. Leuciscidae: Laviniinae. Distribution: Southern North America: endorheic Nazas-Aguanaval basin, Coahuila, Durango and Zacatecas (Mexico) endemic. Habitat: freshwater.

coriacea, Moapa Hubbs [C. L.] & Miller [R. R.] 1948:8, Fig. 1B; Pl. 1 (fig. 1) [Occasional Papers of the Museum of Zoology University of Michigan No. 507; ref. 2258] Home Ranch, one of the sources of Moapa River [Muddy River], Clark County, Nevada, U.S.A. Holotype: UMMZ 143186. Paratypes: CAS-SU 18474 [ex UMMZ 124822] (7); UMMZ 124822 (267, now 257), 133545 (10, now 0); USNM 112067 [ex UMMZ 133545] (10). Type catalog: Gilbert 1998:66 [ref. 23395]. •Valid as Moapa coriacea Hubbs & Miller 1948 -- (Lee et al. 1980:207 [ref. 22416], Page & Burr 1991:79 [ref. 18983], La Rivers 1994:436 [ref. 23389], Gilbert 1998:66 [ref. 23395], Fuller et al. 1999:95 [ref. 25838], Nelson et al. 2004:73 [ref. 27807], Scharpf 2005:24 [ref. 28940], Minckley & Marsh 2009:156 [ref. 31114], Page & Burr 2011:175 [ref. 31215], Page et al. 2013:73 [ref. 32708]). •Valid as Gila coriacea (Hubbs & Miller 1948) -- (Smith et al. 2002: 211 [ref. 36187], Tan & Armbruster 2018:18 [ref. 36121], Page et al. 2023:70 [ref. 40505]). Current status: Valid as Gila coriacea (Hubbs & Miller 1948). Leuciscidae: Laviniinae. Distribution: North America: southeastern Nevada (U.S.A.). Habitat: freshwater.

crassa, Tigoma Girard [C. F.] 1856:207 [43] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Sacramento River, near Fort Reading [Redding], Shasta County, California, U.S.A. Holotype: USNM 216 (plus pharyngeal arches as #2777). Type catalog: Gilbert 1998:67 [ref. 23395]. On p. 43 of separate. •Synonym of Gila crassicauda (Baird & Girard 1854) -- (Gilbert 1998:67 [ref. 23395]). Current status: Synonym of Gila crassicauda (Baird & Girard 1854). Leuciscidae: Laviniinae. Habitat: freshwater.

crassicauda, Lavinia Baird [S. F.] & Girard [C. F.] in Girard 1854:137 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] San Francisco; San Joaquin River, California, U.S.A. Syntypes: ?ANSP 4197 (1) San Joaquin R., USNM 235 (2, plus pair pharyngeal arches as #2772) San Joaquin. Type catalog: Böhlke 1984:74 [ref. 13621], Gilbert 1998:67-68 [ref. 23395]. Author is S. F. Baird and Girard. •Valid as Gila crassicauda (Baird & Girard 1854) [but apparently extinct] -- (Lee et al. 1980:164 [ref. 22416], Gilbert 1998:67-68 [ref. 23395], Moyle 2002:127 [ref. 26449], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:18 [ref. 28940], Page & Burr 2011:165 [ref. 31215], Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505]). Current status: Valid as Gila crassicauda (Baird & Girard 1854). Leuciscidae: Laviniinae. Distribution: Western North America: California (U.S.A.). Habitat: freshwater.

cruoreus, Squalius Jordan [D. S.] & Gilbert [C. H.] 1881:460 [Proceedings of the United States National Museum v. 3 (no. 175); ref. 10606] Provo River, tributary to Utah Lake, Provo, Utah County, Utah, U.S.A. Lectotype: USNM 30806. Paralectotypes: (numerous) BMNH 1883.12.14.203 [ex USNM 30805] (1), USNM 27408 (orig. 6, now 4). Type catalog: Gilbert 1998:68 [ref. 23395]. Lectotype selected by Gilbert 1998:68 [ref. 23395]. •Synonym of Gila atraria (Girard 1856) -- (La Rivers 1994:395 [ref. 23389], Gilbert 1998:68 [ref. 23395]). Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

cypha, Gila Miller [R. R.] 1946:410, Fig. 1 [Journal of the Washington Academy of Sciences v. 36 (no. 12); ref. 13129] Near Phantom Ranch, western end of Grand Canyon, Arizona, U.S.A. (presumably from Colorado River, at mouth of Bright Angel Creek). Holotype: USNM 131839. Other specimens: USNM 15349 (1, head only), 20083 (1). Type catalog: Gilbert 1998:69-70 [ref. 23395]. •Valid as Gila cypha Miller 1946 -- (Lee et al. 1980:165 [ref. 22416], Suttkus & Clemmer 1977 [ref. 21045], Page & Burr 1991:74 [ref. 18983], McElroy & Douglas 1995:636 [ref. 22405], Simons & Mayden 1997:195 [ref. 24948], Sigler & Sigler 1996:79 [ref. 25943], Tyus 1998:190 [ref. 24138], Gilbert 1998:69 [ref. 23395], Douglas et al. 2001:389 [ref. 25233], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:18 [ref. 28940], Minckley & Marsh 2009:121 [ref. 31114], Page & Burr 2011:168 [ref. 31215], Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505]). Current status: Valid as Gila cypha Miller 1946. Leuciscidae: Laviniinae. Distribution: Western North America: western U.S.A. Habitat: freshwater.

ditaenia, Gila Miller [R. R.] 1945:106, Pl. 1 [Copeia 1945 (no. 2); ref. 3016] Río Magdalena, 500 yards west of La Casita, Sonora, Mexico. Holotype: USNM 129954. Paratypes: (476) UMMZ 141945 (241, 11 c&s); USNM 45431 (orig. 40, now 53), 129953 (172), 129955 (9), 130668 (14). Non-type material: USNM 130669 (8). Type catalog: Gilbert 1998:72 [ref. 23395]. •Valid as Gila ditaenia Miller 1945 -- (Branson et al. 1960:219 [ref. 41482]Lee et al. 1980:166 [ref. 22416], Page & Burr 1991:75 [ref. 18983], DeMarais & Minckley 1992 [ref. 19868], Espinosa Pérez et al. 1993:15 [ref. 22290], Gilbert 1998:72 [ref. 23395], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:18 [ref. 28940], Miller et al. 2006:121 [ref. 28615], Minckley & Marsh 2009:124 [ref. 31114], Page & Burr 2011:171 [ref. 31215], Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:172 [ref. 41622]). Current status: Valid as Gila ditaenia Miller 1945. Leuciscidae: Laviniinae. Distribution: Southern North America: southern Arizona (U.S.A.) and Sonora (northwestern Mexico) endemic. Habitat: freshwater.

domninus, Protoporus Cope [E. D.] 1872:473 [U. S. Geological Survey Territitories Annual Report v. 5 (pt 4); ref. 923] Snake River (or tributary thereof), Fort Hall, north of Plcatello, Bingham County, Idaho, U.S.A. Syntypes: (many) whereabouts unknown. Type catalog: Böhlke 1984:75 [ref. 13621] as none at ANSP, spelled dominus, Gilbert 1998:73 [ref. 23395]. •Synonym of Gila atraria (Girard 1856) -- (La Rivers 1994:395 [ref. 23389], Gilbert 1998:73 [ref. 23395]). Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

elegans, Gila Baird [S. F.] & Girard [C. F.] 1853:369 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 6; ref. 166] Zuni River, New Mexico, U.S.A. [actual: Little Colorado River, below Grand Falls, Coconino County, Arizona, U.S.A.]. Holotype (unique): USNM 20079 [orig. USNM 251] plus 1 pharyngeal arch. Type catalog: Gilbert 1998:74-75 [ref. 23395]. •Synonym of Gila robusta Baird & Girard 1853, but a valid subspecies -- (La Rivers 1994:391 [ref. 23389]). •Valid as Gila elegans Baird & Girard 1853 -- (Lee et al. 1980:167 [ref. 22416], Kaeding et al. 1986:1021 [ref. 6781], Page & Burr 1991:74 [ref. 18983], Espinosa Pérez et al. 1993:15 [ref. 22290], Swift et al. 1993:115 [ref. 22379], Baxter & Stone 1995:115 [ref. 23374], Sigler & Sigler 1996:84 [ref. 25943], Tyus 1998:190 [ref. 24138], Gilbert 1998:74-75 [ref. 23395], Minckley & DeMarais 2000:251 [ref. 24298], Moyle 2002:132 [ref. 26449], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:18 [ref. 28940], Miller et al. 2006:122 [ref. 28615], Minckley & Marsh 2009:126 [ref. 31114], Page & Burr 2011:168 [ref. 31215], Page et al. 2013:71 [ref. 32708], Copus et al. 2018:6 [ref. 36179], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:172 [ref. 41622]). Current status: Valid as Gila elegans Baird & Girard 1853. Leuciscidae: Laviniinae. Distribution: Western North America: Río Colorado system, southwestern U.S.A. and Baja California and S(southwestern U.S. and northern Mexico) (BCN, SON) Habitat: freshwater.

emoryi, Gila Baird [S. F.] & Girard [C. F.] 1853:388 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 6; ref. 15636] Near mouth of Gila River, Arizona, U.S.A. Holotype (unique): USNM 247. Type catalog: Gilbert 1998:75-76 [ref. 23395]. •Synonym of Gila robusta Baird & Girard 1853, subspecies elegans Baird & Girard 1853 -- (La Rivers 1994:391 [ref. 23389]). •Synonym of Gila elegans Baird & Girard 1853 -- (Gilbert 1998:75-76 [ref. 23395], Minckley & Marsh 2009:130 [ref. 31114]). Current status: Synonym of Gila elegans Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

eremica, Gila DeMarais [B. D.] 1991:179, Figs. 1-2 [Copeia 1991 (no. 1); ref. 19508] Río Matape, San Jose de Pimas, Sonora, Mexico, elevation 350 meters. Holotype: UMMZ 216819. Paratypes: ANSP 165627 (10); ASU 9036 (108), 9924 (146); UANL 9282 (25); UMMZ 161468 (9), 161587 (1), 216820 (50). Type catalog: Gilbert 1998:76 [ref. 23395], Contreras-Balderas et al. 1998:3 [ref. 23752]. •Valid as Gila eremica DeMarais 1991 -- (Espinosa Pérez et al. 1993:15 [ref. 22290], Gilbert 1998:76 [ref. 23395], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:18 [ref. 28940], Miller et al. 2006:122 [ref. 28615], Minckley & Marsh 2009:130 [ref. 31114], Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:173 [ref. 41622]). Current status: Valid as Gila eremica DeMarais 1991. Leuciscidae: Laviniinae. Distribution: Southern North America: Sonora (Pacific slope of northwestern Mexico) endemic. Hybridizes with Gila ditaenia Miller 1945 in Sonora (Mexico). Habitat: freshwater.

gibbosa, Gila Baird [S. F.] & Girard [C. F.] 1854:28 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 168] Rio Santa Cruz, tributary of Gila River, Arizona, U.S.A. Syntypes: [all originally USNM 223] ANSP 4198 (1); MCZ 1806 (orig. 2, now 1), 1984 (1), 153935 (pharyngeal arch); MNHN 0000-0391 (1); UMMZ 87006 [ex MCZ 1806] (1); USNM 223 (15, plus 2 pharyngeal arches as #2776), 344867 (1). Type catalog: Böhlke 1984:77 [ref. 13621], Gilbert 1998:84-85 [ref. 23395]. •Synonym of Gila intermedia (Girard 1856) -- (Gilbert 1998:84-85 [ref. 23395]). •Synonym of Gila robusta Baird & Girard 1853 -- (Copus et al. 2018:11 [ref. 36179], Fricke et al. 2024:174 [ref. 41622]). Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

gibbosus, Leuciscus Ayres [W. O.] 1854:1, col. 1 [Daily Placer Times and Transcript [newspaper] v. 5 (no. 1284); ref. 6368] San Francisco, California, U.S.A. Syntypes: ANSP 14733 (1). Type catalog: Böhlke 1984:77 [ref. 13621], Gilbert 1998:85 [ref. 23395]. Appeared above on 30 May 1854. Also appeared as Lavinia gibbosa in Ayres 1854:14 [ref. 157] in Dec. 1854. Objectively invalid; preoccupied by Leuciscus gibbosus Storer 1848. •Synonym of Gila crassicauda (Baird & Girard 1854) -- (Gilbert 1998:85 [ref. 23395]). 1°Homonym. Current status: Synonym of Gila crassicauda (Baird & Girard 1854). Leuciscidae: Laviniinae. Habitat: freshwater.

gracilis, Gila Baird [S. F.] & Girard [C. F.] 1853:369 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 6; ref. 166] "Zuni River, New Mexico" [possibly Little Colorado River, at Grand Falls, Coconino County, Arizona, U.S.A.]. Syntypes: MCZ 35926 [ex USNM 245] (1), USNM 245 (orig. 9, now 8, plus 1 pharyngeal arch as #2797). Type catalog: Gilbert 1998:86 [ref. 23395]. •Synonym of Gila robusta Baird & Girard 1853, subspecies robusta -- (Gilbert 1998:86 [ref. 23395]). •Synonym of Gila robusta Baird & Girard 1853 -- (Minckley & Marsh 2009:142 [ref. 31114], Copus et al. 2018:11 [ref. 36179]). Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

gracilis, Tigoma Girard [C. F.] 1856:206 [42] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] No locality [believed to be Utah Creek, vicinity of Fort Garland, Costilla County, Colorado, U.S.A.]. Syntypes: MNHN 0000-0395 [ex USNM 230] (1), USNM 230 (orig. 2, now 0, 1 pharyngeal arch as #2784). Type catalog: Gilbert 1998:87-88 [ref. 23395]. On p. 42 of separate. Subjectively invalid; secondarily preoccupied in Gila by Gila gracilis Baird & Girard 1853. •Synonym of Gila pandora (Cope 1872) -- (Gilbert 1998:88 [ref. 23395]). Current status: Synonym of Gila pandora (Cope 1872). Leuciscidae: Laviniinae. Habitat: freshwater.

grahamii, Gila Baird [S. F.] & Girard [C. F.] 1853:389 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 6; ref. 15636] Rio San Pedro, Gila River basin, Arizona, U.S.A. Syntypes: ANSP 4185-86 [ex USNM 252] (2); MCZ 153937 (1 pharyngeal arch); MNHN 0000-0384 [ex USNM 252] (1), USNM 252 (orig. 6, now 1, plus pharyngeal arch as #2801), 253 (1, plus pharyngeal arch as #2802). Type catalog: Böhlke 1984:78 [ref. 13621] as grahami, Gilbert 1998:88 [ref. 23395]. See Minckley & DeMarais 2000:254 [ref. 24298]. •Synonym of Gila robusta Baird & Girard 1853, but a valid subspecies -- (Lee et al. 1980:172 [ref. 22416], Gilbert 1998:88 [ref. 23395]). •Synonym of Gila robusta Baird & Girard 1853 -- (La Rivers 1994:388 [ref. 23389], Copus et al. 2018:11 [ref. 36179]). Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

gula, Gila Cope [E. D.] in Cope & Yarrow 1875:661, Pl. 30 (figs. 2, 2a-b) [Engineer Department, United States Army, Report upon the Geographical and Geological Explorations and Surveys west of the one hundredth Meridian v. 5 (Zoology) Chapter 6; ref. 968] Rio de Acama and near Fort Wingate, New Mexico, U.S.A. Syntypes: ANSP 18700 (1), MCZ 35915 [ex USNM 16979] (1), USNM 16979 (orig. 6, now 4). Type catalog: Böhlke 1984:78 [ref. 13621], Gilbert 1998:89 [ref. 23395]. •Synonym of Gila pandora (Cope 1872) -- (Gilbert 1998:89 [ref. 23395]). Current status: Synonym of Gila pandora (Cope 1872). Leuciscidae: Laviniinae. Habitat: freshwater.

intermedia, Tigoma Girard [C. F.] 1856:206 [42] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Rio San Pedro, Gila River basin, Arizona, U.S.A. Syntypes: [all originally USNM 232] ANSP 4187 (1), MCZ 1963 [ex USNM 232] (1), MNHN 0000-0396 (1), NMW 48718 (1), USNM 232 (orig. 20, now 12, plus 1 pair pharyngeal arches as #2786). Type catalog: Böhlke 1984:79 [ref. 13621], Gilbert 1998:96 [ref. 23395]. On p. 42 of separate. •Valid as Gila intermedia (Girard 1856) -- (Espinosa Pérez et al. 1993:15 [ref. 22290], Gilbert 1998:96 [ref. 23395], Minckley & DeMarais 2000:251 [ref. 24298], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:18 [ref. 28940], Miller et al. 2006:123 [ref. 28615], Minckley & Marsh 2009:131 [ref. 31114], Page & Burr 2011:165 [ref. 31215], Page & Burr 2011:167 [ref. 31215], Page et al. 2013:71 [ref. 32708], Moran et al. 2017:1 [ref. 35521], Page et al. 2023:70 [ref. 40505]). •Synonym of Gila robusta Baird & Girard 1853 -- (Page et al. 2017:459 [ref. 35584], Carter et al. 2018:289 [ref. 35930], Copus et al. 2018:11 [ref. 36179], Suchocki et al. 2023:1 [ref. 40513], Fricke et al. 2024:174 [ref. 41622]). Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

jordani, Gila Tanner [V. M.] 1950:32, Fig. 1 [Great Basin Naturalist, Provo Utah v. 10 (nos 1-4); ref. 12978] Crystal Spring, Pahranagat Valley, Lincoln County, Nevada, U.S.A. Holotype: BYU 9959. Paratypes: BYU 8980-81 (1, 1), 9084 (1), 9958 (1), 9960 (1). Type catalog: Tanner 1970:220 [ref. 23524]. •Synonym of Gila robusta Baird & Girard 1853, but a valid subspecies -- (Lee et al. 1980:172 [ref. 22416], La Rivers 1994:393 [ref. 23389], Gilbert 1998:98 [ref. 23395], Scharpf 2005:19 [ref. 28940], Page & Burr 2011:165 [ref. 31215]). •Valid as Gila jordani Tanner 1950 -- (Minckley & Marsh 2009:133 [ref. 31114], Page et al. 2013:71 [ref. 32708], Copus et al. 2018:6 [ref. 36179], Page et al. 2023:70 [ref. 40505]). Current status: Valid as Gila jordani Tanner 1950. Leuciscidae: Laviniinae. Distribution: North America: Nevada (U.S.A.). Habitat: freshwater.

lemmoni, Squalius Smith [R.] 1884:3-4 [Bulletin of the California Academy of Sciences No. 1; ref. 12300] Rillitto Creek, small stream in Santa Catalina Mts., 7 miles north of Tucson, Pimz County, Arizona, U.S.A. Syntypes: CAS-SU 24128 (orig. 4, now 1 [3 lost in 1906 earthquake and fire]). Type catalog: Böhlke 1953:39 [ref. 12291], Gilbert 1998:101-102 [ref. 23395]. •Synonym of Gila intermedia (Girard 1856) -- (Gilbert 1998:102 [ref. 23395], Minckley & Marsh 2009:133 [ref. 31114]). •Synonym of Gila robusta Baird & Girard 1853 -- (Copus et al. 2018:12 [ref. 36179]). Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

lineata, Tigoma Girard [C. F.] 1856:206 [42] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Near 38°N [not Humboldt River, Nevada, U.S.A.]. Syntypes: [all originally USNM 229] ANSP 4188 (1), MCZ 1807 (1), MNHN 0000-0394 (1), USNM 229 (orig. 6, now 0, 1 pharyngeal arch as #2783). Type catalog: Böhlke 1984:81 [ref. 13621], Gilbert 1998:104 [ref. 23395]. On p. 42 of separate. Not available, placed on Official Index in Opinion 1872 (see Appendix B). •In the synonymy of Gila atraria (Girard 1856) -- (La Rivers 1994:395 [ref. 23389] with question, Gilbert 1998:104 [ref. 23395]). Suppressed. Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae.

longiceps, Siboma atraria var. Cope [E. D.] in Cope & Yarrow 1875:668, Pl. 29 (fig. 4) [Engineer Department, United States Army, Report upon the Geographical and Geological Explorations and Surveys west of the one hundredth Meridian v. 5 (Zoology) Chapter 6; ref. 968] Snake Creek Valley, near Utah line, near Sevier Basin, Lake Bonneville drainage, Nevada, U.S.A. Syntypes: USNM 15767 (4), 15770 (1). Non-types: MCZ 18609 [ex USNM 12910] (4), 36000 [ex USNM 12910] (2); USNM 12910 (orig. 15, now 9), 15759 (1), 15782 (4). Type catalog: Gilbert 1998:105 [ref. 23395]. •Synonym of Gila atraria (Girard 1856) -- (Gilbert 1998:105 [ref. 23395]). Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

minacae, Gila Meek [S. E.] 1902:80, Pl. 17 lower [Field Columbian Museum, Zoölogical Series v. 3 (no. 6); ref. 2957] Río Paphigochic at Miñaca, Río Yaqui basin, Chihuahua, Mexico. Holotype (unique): FMNH 3573. Type catalog: Ibarra & Stewart 1987:39 [ref. 12367], Gilbert 1998:115 [ref. 23395]. •Synonym of Gila robusta Baird & Girard 1853, but a valid subspecies -- (Gilbert 1998:115 [ref. 23395]). •Valid as Gila minacae Meek 1902 -- (Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:18 [ref. 28940], Miller et al. 2006:123 [ref. 28615], Minckley & Marsh 2009:135 [ref. 31114], Page et al. 2013:71 [ref. 32708], Ballesteros-Córdova et al. 2020:217 [ref. 37371], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:173 [ref. 41622]). Current status: Valid as Gila minacae Meek 1902. Leuciscidae: Laviniinae. Distribution: Southern North America: Sierra Madre Occidental, Sinaloa, Sonora (Mexico) endemic; ?Arizona (U.S.A.). Habitat: freshwater.

modesta, Cheonda Garman [S.] 1881:92 [Bulletin of the Museum of Comparative Zoology v. 8 (no. 3); ref. 1530] Spring near Saltillo, Río Salinas drainage, Coahuila, Mexico. Syntypes: MCZ 24893 (2). •Valid as Gila modesta (Garman 1881) -- (Espinosa Pérez et al. 1993:15 [ref. 22290], Gilbert 1998:115 [ref. 23395], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:18 [ref. 28940], Miller et al. 2006:124 [ref. 28615], Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:173 [ref. 41622]). Current status: Valid as Gila modesta (Garman 1881). Leuciscidae: Laviniinae. Distribution: Southern North America: Chorro Stream in Río San Juan basin (Coahuila), possibly also in Iturbide (Nuevo León), Atlantic Slope (Mexico) endemic. Habitat: freshwater.

nacrea, Gila Cope [E. D.] 1871:441 [Preliminary report of the United States Geological Survey of Wyoming and portions of contiguous territories. Part 4 (art. 8); ref. 19655] Fork of Green River near Fort Bridger, Wyoming, U.S.A. No types known. Type catalog: Böhlke 1984:84 [ref. 13621], Gilbert 1998:117 [ref. 23395]. •Synonym of Gila robusta, subspecies robusta Baird & Girard 1853 -- (Gilbert 1998:117 [ref. 23395]). •Synonym of Gila robusta Baird & Girard 1853 -- (Minckley & Marsh 2009:142 [ref. 31114], Copus et al. 2018:12 [ref. 36179]). Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

nigra, Gila Cope [E. D.] in Cope & Yarrow 1875:663, Pl. 30 (figs. 3, 3a-b) [Engineer Department, United States Army, Report upon the Geographical and Geological Explorations and Surveys west of the one hundredth Meridian v. 5 (Zoology) Chapter 6; ref. 968] Ash Creek, Pinal County and San Carlos River, Gila County, Arizona, U.S.A. Syntypes: MCZ 35963 [ex USNM 16972] (1), USNM 16972 (orig. 8, now 5), 16987 (2). Type catalog: Böhlke 1984:84 [ref. 13621], Gilbert 1998:119-120 [ref. 23395]. •Synonym of Gila intermedia (Girard 1856) -- (Gilbert 1998:120 [ref. 23395]). •Valid as Gila nigra Cope 1875 -- (Minckley & DeMarais 2000:255 [ref. 24298], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:18 [ref. 28940], Minckley & Marsh 2009:136 [ref. 31114], Page & Burr 2011:165 [ref. 31215], Page et al. 2013:71 [ref. 32708], Moran et al. 2017:1 [ref. 35521], Page et al. 2023:70 [ref. 40505]). •Synonym of Gila robusta Baird & Girard 1853 -- (Page et al. 2017:459 [ref. 35584], Carter et al. 2018:289 [ref. 35930], Copus et al. 2018:12 [ref. 36179], Suchocki et al. 2023:1 [ref. 40513], Fricke et al. 2024:174 [ref. 41622]). Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

nigrescens, Tigoma Girard [C. F.] 1856:207 [43] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Boca Grande and Río Janos, state of Chihuahua, Mexico. Syntypes: ANSP 4183 [ex USNM 219] (1); MCZ 1804 [ex USNM 218 or 220] (1), 1805 [ex USNM 220] (1); MNHN 0000-0388 [ex USNM 220], NMW 48719 [ex USNM 220] (1); USNM 219 (2), 220 (orig. 10+, now 3, plus 1 pair of pharyngeal arches as #2774). Type catalog: Böhlke 1984:85 [ref. 13621]. On p. 43 of separate. •Valid as Gila nigrescens (Girard 1856) -- (Lee et al. 1980:168 [ref. 22416], Page & Burr 1991:74 [ref. 18983], Espinosa Pérez et al. 1993:15 [ref. 22290], Stefferud & Propst 1996:446 [ref. 25520], Gilbert 1998:120 [ref. 23395], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:19 [ref. 28940], Miller et al. 2006:124 [ref. 28615], Page & Burr 2011:170 [ref. 31215], Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:173 [ref. 41622]). Current status: Valid as Gila nigrescens (Girard 1856). Leuciscidae: Laviniinae. Distribution: Southern North America: New Mexico (U.S.A.) and Mexico. Habitat: freshwater.

orcuttii, Phoxinus (Tigoma) Eigenmann [C. H.] & Eigenmann [R. S.] 1890:2 [Proceedings of the California Academy of Sciences (Series 2) v. 3; ref. 1256] Temecula River and tributaries, Riverside, Riverside County, California, U.S.A. Syntypes: (over 100) BMNH 1890.11.15.340-342 (3); CAS 58621 [IU 4844] (4); FMNH 54985 (10); MCZ 27390 (4); CAS-SU 858 (15); UMMZ 87065 [ex MCZ 27390]; USNM 41886 (21), 41925 (17); CAS 1011 (old registry) destroyed in earthquake of 1906. Type catalog: Böhlke 1953:36 [ref. 12291], Ibarra & Stewart 1987:65 [ref. 12367], Gilbert 1998:126 [ref. 23395]. Jordan & Evermann 1896:242 [ref. 2443] list USNM 41925 as type, but that lot contained many specimens. •Valid as Gila orcuttii (Eigenmann & Eigenmann 1890) -- (Lee et al. 1980:169 [ref. 22416], Page & Burr 1991:75 [ref. 18983], Swift et al. 1993:115 [ref. 22379] as orcutti, Gilbert 1998:126 [ref. 23395], Fuller et al. 1999:76 [ref. 25838], Moyle 2002:130 [ref. 26449] as orcutti, Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:19 [ref. 28940], Page & Burr 2011:171 [ref. 31215], Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:173 [ref. 41622]). Current status: Valid as Gila orcuttii (Eigenmann & Eigenmann 1890). Leuciscidae: Laviniinae. Distribution: Western North America: California (U.S.A.). Introduced in Mexico. Habitat: freshwater.

pandora, Clinostomus Cope [E. D.] 1872:475 [U. S. Geological Survey Territitories Annual Report v. 5 (pt 4); ref. 923] Sangre de Christo Pass, tributary to Rio Grande River, Costilla County, Colorado, U.S.A. Syntypes: ANSP 4195 (1). Type catalog: Böhlke 1984:86 [ref. 13621], Gilbert 1998:129 [ref. 23395]. •Valid as Gila pandora (Cope 1872) -- (Lee et al. 1980:170 [ref. 22416], Page & Burr 1991:75 [ref. 18983], Gilbert 1998:129 [ref. 23395], Fuller et al. 1999:76 [ref. 25838], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:19 [ref. 28940], Page & Burr 2011:170 [ref. 31215], Page et al. 2013:71 [ref. 32708], Copus et al. 2018:6 [ref. 36179], Page et al. 2023:70 [ref. 40505]). Current status: Valid as Gila pandora (Cope 1872). Leuciscidae: Laviniinae. Distribution: North America: Texas (southwestern U.S.A.). Introduced in Colorado and New Mexico (U.S.A.). Habitat: freshwater.

parovanus, Myloleucus Cope [E. D.] 1874:136 [8] [Proceedings of the American Philosophical Society v. 14; ref. 932] Beaver River, Beaver County, southwestern Utah, U.S.A. Syntypes: MCZ 87011 (orig. 4, now 3), USNM 15775 (11), UMMZ 87011 [ex MCZ 87011] (1). Type catalog: Böhlke 1984:86 [ref. 13621], Gilbert 1998:129-130 [ref. 23395]. On p. 8 of separate. •Synonym of Gila atraria (Girard 1856) -- (Gilbert 1998:129-130 [ref. 23395]). •Synonym of Siphateles bicolor (Girard 1856) -- (La Rivers 1994:410 [ref. 23389]). Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

pulchella, Gila Baird [S. F.] & Girard [C. F.] 1854:29 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 168] Rio Mimbres, tributary of Gila River, north of Deming, Luna County, New Mexico, U.S.A. Syntypes: USNM 233 (2, plus 1 pair of pharyngeal arches as #2787). Type catalog: Gilbert 1998:136 [ref. 23395]. Subjectively invalid; secondarily preoccupied in Leuciscus by Leuciscus pulchellus Storer 1839. •Synonym of Gila nigrescens (Girard 1856) -- (Gilbert 1998:136 [ref. 23395], Fricke et al. 2024:173 [ref. 41622]). Current status: Synonym of Gila nigrescens (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

pulchra, Tigoma Girard [C. F.] 1856:207 [43] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] Río Chihuahua and tributaries, Chihuahua, Mexico. Syntypes: MNHN 0000-0393 [ex USNM 227] (1); USNM 227 (orig. 3, now 2, plus 2 pairs of pharyngeal arches as #2781), 228 (5, plus 1 pair ofpharyngeal arches as #2782). Type catalog: Gilbert 1998:137 [ref. 23395]. On p. 43 of separate. •Valid as Gila pulchra (Girard 1856) -- (Espinosa Pérez et al. 1993:15 [ref. 22290], Gilbert 1998:137 [ref. 23395], Norris et al. 2003:22 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:19 [ref. 28940], Miller et al. 2006:125 [ref. 28615], Minckley & Marsh 2009:145 [ref. 31114] as cf. pulchra, Page et al. 2013:71 [ref. 32708], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:173 [ref. 41622]). Current status: Valid as Gila pulchra (Girard 1856). Leuciscidae: Laviniinae. Distribution: Southern North America: Río Conchos basin, Chihuahua and Durango (Mexico) endemic. Habitat: freshwater.

pulverulentus, Myloleucus Cope [E. D.] 1872:475 [U. S. Geological Survey Territitories Annual Report v. 5 (pt 4); ref. 923] Warm Springs, Utah, U.S.A. Syntypes: ANSP 18788 (1, missing). Type catalog: Böhlke 1984:88 [ref. 13621], Gilbert 1998:137 [ref. 23395]. •Synonym of Gila atraria (Girard 1856) -- (Gilbert 1998:137 [ref. 23395]). Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

purpurea, Tigoma Girard [C. F.] 1856:206 [42] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] San Bernardino Creek, upper Rio Huagui [Rio Yaqui], west of the Sierra Madre, Cochise County, southern Arizona, U.S.A. Lectotype: USNM 323519. Paralectotypes: USNM 224 (20, plus 1 pharyngeal arch as #2778), 301925 [ex USNM 224] (2). Type catalog: Gilbert 1998:138 [ref. 23395]. On p. 42 of separate. Lectotype not yet published; see Gilbert 1998:138 [ref. 23395], but specimen isolated in USNM 323519 [not here designated as such]. •Valid as Gila purpurea (Girard 1856) -- (Branson et al. 1960:219 [ref. 41482], Lee et al. 1980:171 [ref. 22416], Page & Burr 1991:76 [ref. 18983], Espinosa Pérez et al. 1993:15 [ref. 22290], Gilbert 1998:138 [ref. 23395], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:19 [ref. 28940], Miller et al. 2006:125 [ref. 28615], Minckley & Marsh 2009:138 [ref. 31114], Page & Burr 2011:171 [ref. 31215], Page et al. 2013:71 [ref. 32708], Ballesteros-Córdova et al. 2020:216 [ref. 37371], Page et al. 2023:70 [ref. 40505], Fricke et al. 2024:173 [ref. 41622]). Current status: Valid as Gila purpurea (Girard 1856). Leuciscidae: Laviniinae. Distribution: Southern North America: Arizona (U.S.A.) and upper Yaqui River basin (northern Mexico). Habitat: freshwater.

rhomaleus, Squalius Jordan [D. S.] & Gilbert [C. H.] 1881:461 [Proceedings of the United States National Museum v. 3 (no. 175); ref. 10606] Utah Lake at Provo, Utah County, Utah, U.S.A. Syntypes: (2) USNM 27364 (orig. 2, now 1). Type catalog: Gilbert 1998:140 [ref. 23395]. •Synonym of Gila atraria (Girard 1856) -- (La Rivers 1994:395 [ref. 23389], Gilbert 1998:140 [ref. 23395]). Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

robusta, Gila Baird [S. F.] & Girard [C. F.] 1853:369 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 6; ref. 166] Zuni River, New Mexico, U.S.A. [possibly Little Colorado River, below Grand Falls, Coconino County, Arizona, U.S.A.]. Syntypes: USNM 246 [= 47983] (3, plus 1 pharyngeal arch as #2798). Type catalog: Gilbert 1998:141 [ref. 23395]. •Valid as Gila robusta Baird & Girard 1853 -- (Lee et al. 1980:172 [ref. 22416], Page & Burr 1991:72 [ref. 18983], Espinosa Pérez et al. 1993:16 [ref. 22290], Swift et al. 1993:117 [ref. 22379], La Rivers 1994:388 [ref. 23389], McElroy & Douglas 1995:636 [ref. 22405], Baxter & Stone 1995:72 [ref. 23374], Sigler & Sigler 1996:86 [ref. 25943], Tyus 1998:190 [ref. 24138], Gilbert 1998:141 [ref. 23395], Fuller et al. 1999:77 [ref. 25838], Minckley & DeMarais 2000:251 [ref. 24298], Douglas et al. 2001:389 [ref. 25233], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:19 [ref. 28940] with subspecies, Miller et al. 2006:126 [ref. 28615], Minckley & Marsh 2009:140 [ref. 31114], Page & Burr 2011:165 [ref. 31215], Page & Burr 2011:168 [ref. 31215], Page et al. 2013:71 [ref. 32708], Moran et al. 2017:1 [ref. 35521], Page et al. 2017:459 [ref. 35584], Carter et al. 2018:289 [ref. 35930], Copus et al. 2018:11 [ref. 36179], Page et al. 2023:70 [ref. 40505], Suchocki et al. 2023:1 [ref. 40513], Fricke et al. 2024:174 [ref. 41622]). Current status: Valid as Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Distribution: North America: Colorado River basin: Arizona, New Mexico, Colorado, Wyoming, Utah and Nevada (western U.S.A.) and Chihuahua and Sonora (northwestern Mexico). Habitat: freshwater.

seminuda, Gila Cope [E. D.] & Yarrow [H. C.] 1875:666, Pl. 31 (figs. 1-1a) [Engineer Department, United States Army, Report upon the Geographical and Geological Explorations and Surveys west of the one hundredth Meridian v. 5 (Zoology) Chapter 6; ref. 968] Virgen [Virgin] River, an affluent of Colorado River, a little south of Washington, Washington County, southern Utah, U.S.A. Syntypes: USNM 16975 (orig. 6, now 5) Type catalog: Gilbert 1998:146 [ref. 23395]. •Synonym of Gila robusta Baird & Girard 1853, but a valid subspecies -- (Lee et al. 1980:172 [ref. 22416]). •Valid as Gila seminuda Cope & Yarrow 1875 -- (Gilbert 1998:146 [ref. 23395], Norris et al. 2003:24 [ref. 26757], Nelson et al. 2004:71 [ref. 27807], Scharpf 2005:19 [ref. 28940], Minckley & Marsh 2009:143 [ref. 31114], Page & Burr 2011:167 [ref. 31215], Page et al. 2013:71 [ref. 32708], Copus et al. 2018:6 [ref. 36179], Page et al. 2023:70 [ref. 40505]). Current status: Valid as Gila seminuda Cope & Yarrow 1875. Leuciscidae: Laviniinae. Distribution: Western North America: western U.S.A. Habitat: freshwater.

squamata, Tigoma Gill [T. N.] 1861:42 [Proceedings of the Boston Society of Natural History v. 8 (1861-1862); ref. 12586] Salt Lake, Utah, U.S.A. Syntypes: MNHN 0000-0390 [ex USNM 2607] (1), USNM 2607 (2). Type catalog: Gilbert 1998:150 [ref. 23395]. •Synonym of Gila atraria (Girard 1856) -- (La Rivers 1994:395 [ref. 23389], Gilbert 1998:150 [ref. 23395]). Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

timpanogensis, Hybopsis Cope [E. D.] 1874:134 [6] [Proceedings of the American Philosophical Society v. 14; ref. 932] Provo River, Provo, Utah County; pool off Sevier River, Gunnison, Sanpete County, Utah, U.S.A. Syntypes: (numerous) ANSP 23001-06 (6), MCZ 18586 [ex USNM 15769] (orig. 8, now 6), UMMZ 86443 [ex MCZ 18586] (2), USNM 15769 (now 33). Type catalog: Böhlke 1984:93 [ref. 13621], Gilbert 1998:156 [ref. 23395]. On p. 6 of separate. •Synonym of Gila atraria (Girard 1856) -- (La Rivers 1994:395 [ref. 23389], Gilbert 1998:156 [ref. 23395]). Current status: Synonym of Gila atraria (Girard 1856). Leuciscidae: Laviniinae. Habitat: freshwater.

utensis, Gila Jordan [D. S.] 1878:424 [Bulletin of the United States Geological and Geographical Survey of the Territories. v. 4 (no. 2, art. 18); ref. 2379] Type catalog: Gilbert 1998:160 [ref. 23395]. Appeared as "285. Gila utensis Jor. Utah. (Tigoma gracilis Grd.) (d.s.)." Not available; no distinguishing features, apparently not a replacement name. See Gilbert 1998:160 [ref. 23395]. Nomen Nudum. Leuciscidae: Laviniinae.

vorax, Ptychocheilus Girard [C. F.] 1856:209 [45] [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1810] [Precise locality not known] Colorado River drainage between Cochetopapa Creek and Green River, Colorado/Utah, U.S.A. Syntypes: USNM 202 (2, plus 1 pharyngeal arch as #2756) near 38°N. Type catalog: Gilbert 1998:163-164 [ref. 23395]. On p. 45 of separate. •Synonym of Gila robusta Baird & Girard 1853, but a valid subspecies vorax (Baird & Girard 1853) -- (La Rivers 1994:388 [ref. 23389], Gilbert 1998:164 [ref. 23395]). •Synonym of Gila robusta Baird & Girard 1853 -- (Minckley & Marsh 2009:142 [ref. 31114], Copus et al. 2018:11 [ref. 36179]). Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.

zunnensis, Leuciscus Günther [A.] 1868:241 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Zuni River, New Mexico [possibly Little Colorado River, at Grand Falls, Coconino County, Arizona, U.S.A.]. Syntypes: MCZ 35926 [ex USNM 245] (1), USNM 245 (orig. 9, now 8, plus 1 pharyngeal arch as #2727). Type catalog: Gilbert 1998:167 [ref. 23395]. Apparently a replacement name for Gila gracilis Baird & Girard 1853, preoccupied in Leuciscus by Cyprinus (Leuciscus) gracilis Richardson (on Günther p. 240). •Synonym of Gila robusta Baird & Girard 1853 -- (Gilbert 1998:166 [ref. 23395], Copus et al. 2018:11 [ref. 36179]). Replacement Name. Current status: Synonym of Gila robusta Baird & Girard 1853. Leuciscidae: Laviniinae. Habitat: freshwater.


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