Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the genus Homatula: [ 32 ] records

acuticephala, Paracobitis Zhou [W.] & He [J.-C.] 1993:5 [English p. 9], Fig. 1 [Zoological Research v. 14 (no. 1); ref. 21971] Haixihai near Niujie, Eryuan County, Yunnan Province, China. Holotype: MZYU 784141. Paratypes: (7); one at KIZ 784141. Type catalog: Du et al. 2013:350 [ref. 32939]. Species sometimes dated to 1992. •Valid as Paracobitis acuticephala Zhou & He 1993 -- (Min et al. 2010:202 [ref. 30726] as acuticephalus). •Valid as Homatula acuticephala (Zhou & He 1993) -- (Hu & Zhang 2010:52 [ref. 30780], Gu & Zhang 2012:xx [ref. 31632], Min et al. 2012:313 [ref. 32341], Min et al. 2012:81 [ref. 32392], Kottelat 2012:83 [ref. 32367], Kottelat 2013:199 [ref. 32989], Zhang et al. 2016:128 [ref. 34477], Endruweit et al. 2018:565 [ref. 35763], Li et al. 2019:346 [ref. 37175], Zhou et al. 2021:12 [ref. 38386], Nguyen et al. 2021:9 [ref. 38733], Min et al. 2022:112 [ref. 39113], Li et al. 2022:17 [ref. 39800]). Current status: Valid as Homatula acuticephala (Zhou & He 1993). Nemacheilidae. Distribution: East Asia: upper Mekong River basin, Yunnan (China). IUCN (2024): Vulnerable. Habitat: freshwater.

anguillioides, Paracobitis Zhu [S.-Q.] & Wang [S.-H.] 1985:210 [218], Figs. 1-4 [Acta Zootaxonomica Sinica v. 10 (no. 2); ref. 8120] Longtan spring, near Yousuo, 26°1.204'N, 100°3.206'E [not 26°N, 99°50'E], Eryuan County, Yunnan Province, China. Holotype: NIGLAS 820134 (now INB 820134). Paratypes: (19). English description on p. 218. New latitude and longitude is from Rui Min via R. Winterbottom, 7/25/2011. •Valid as Paracobitis anguillioides Zhu & Wang 1985 -- (Zhu 1989:34 [ref. 17744], Zhou & He 1993:5 [ref. 21971], Bănărescu & Nalbant 1995:444 [ref. 23187] with uncertain genus placement, Zhu 1995:109 [ref. 25213], Chen & Zhang in Chen 1998:50 [ref. 23556], Min et al. 2010:202 [ref. 30726]). •Valid as Homatula anguillioides (Zhu & Wang 1985) -- (Hu & Zhang 2010:52 [ref. 30780], Gu & Zhang 2012:597 [ref. 31632], Min et al. 2012:313 [ref. 32341], Min et al. 2012:81 [ref. 32392], Kottelat 2012:83 [ref. 32367], Kottelat 2013:199 [ref. 32989], Zhang et al. 2016:128 [ref. 34477], Endruweit et al. 2018:565 [ref. 35763], Li et al. 2019:346 [ref. 37175], Zhou et al. 2021:12 [ref. 38386], Nguyen et al. 2021:9 [ref. 38733], Min et al. 2022:112 [ref. 39113], Li et al. 2022:17 [ref. 39800]). Current status: Valid as Homatula anguillioides (Zhu & Wang 1985). Nemacheilidae. Distribution: East Asia: upper Mekong River drainage, Yunnan (China). IUCN (2024): Not Assessed. Habitat: freshwater.

anteridorsalis, Homatula Li [X.], Che [X.-J.] & Zhou [W.] 2019:337, Figs. 2A, 6-7A [Zootaxa 4711 (no. 2); ref. 37175] Bingmen Village, Lujiang Town, Longyang District, Baoshan City, Yunnan, China, 25°4'11.23"N, 98°49'57.77"E. Holotype: SWFC 1610006. Paratypes: SWFC. Misspelled in fig. 6 as anteriordorsalis. •Valid as Homatula anteridorsalis Li, Che & Zhou 2019 -- (Zhou et al. 2021:11 [ref. 38386], Li et al. 2022:17 [ref. 39800]). Current status: Valid as Homatula anteridorsalis Li, Che & Zhou 2019. Nemacheilidae. Distribution: East Asia: upper Salween River basin [Nujiang], Yunnan (China). IUCN (2024): Not Assessed. Habitat: freshwater.

berezowskii, Nemachilus Günther [A.] 1896:217 [19], Pl. 2 (fig. C) [Ezhegodnik Zoologicheskogo Muzeya Imperatorskoy Akademii Nauk 1896, v. 1 (no. 3); ref. 2019] Hui-hsien [now Hui County of Gansu Province], upper Chang Jiang [Yangtze River] drainage, China. Holotype (unique): whereabouts unknown. Original genus is correctly Nemacheilus. On p. 19 of separate. •Synonym of Paracobitis variegatus (Dabry de Thiersant 1874) -- (Zhu 1989:32 [ref. 17744], Wu & Wu 1992:140 [ref. 21205], Yang & Chen 1994:60 [ref. 28152], Chen & Zhang in Chen 1998:49 [ref. 23556]). •Synonym of Homatula variegata (Dabry de Thiersant 1874) -- (Bănărescu & Nalbant 1995:444 [ref. 23187]). •Valid as Homatula berezowskii (Günther 1896) -- (Hu & Zhang 2010:56, 57 [ref. 30780], Gu & Zhang 2012:597 [ref. 31632], Kottelat 2012:83 [ref. 32367], Li et al. 2019:346 [ref. 37175], Du et al. 2021:4 [ref. 38600], Zhou et al. 2021:12 [ref. 38386], Guo 2021:493 [ref. 38618], Nguyen et al. 2021:8 [ref. 38733], Che et al. 2023:284 [ref. 40280], Xiao et al. 2025:263 [ref. 41908], Cao et al. 2025:707 [ref. 42007]). Current status: Valid as Homatula berezowskii (Günther 1896). Nemacheilidae. Distribution: East Asia: upper Yangtze River basin, Gansu and Shaanxi Provinces, China. IUCN (2024): Not Assessed. Habitat: freshwater.

change, Homatula Endruweit [M.] 2015:66, Figs. 1, 3-4 [Ichthyological Exploration of Freshwaters v. 26 (no. 1); ref. 33868] Yunnan, Puer Prefecture, Jiangcheng County, hill stream close to Yiwanshui village, a tributary to Mengyejiang River, Black River, Red River basin, 22°35.759'N, 101°45.488'E, China, elevation 953 meters. Holotype: KIZ 2012004205. Paratypes: KIZ, ZFMK, ZRC. •Valid as Homatula change Endruweit 2015 -- (Endruweit et al. 2018:565 [ref. 35763], Li et al. 2019:346 [ref. 37175], Zhou et al. 2021:11 [ref. 38386], Nguyen et al. 2021:8 [ref. 38733], Min et al. 2022:112 [ref. 39113], Li et al. 2022:17 [ref. 39800], Che et al. 2023:284 [ref. 40280], Endruweit 2025:142 [ref. 42401] in subgenus Plerotula). Current status: Valid as Homatula change Endruweit 2015. Nemacheilidae. Distribution: Southeast Asia: upper Black River drainage, Red River basin, Yunnan Province (China). Habitat: freshwater.

coccinocola, Homatula Endruweit [M.], Min [R.] & Yang [J.-X.] 2018:558, Figs. 1-2 [Zootaxa 4375 (no. 4); ref. 35763] Tengtiaojiang River, Red River basin, Jiache town, Honghe City, Yunnan Province, China, 23°17'27.4"N, 102°11'19.1"E, elevation 1220 meters. Holotype: KIZ 2011002847. Paratypes: KIZ. •Valid as Homatula coccinocola Endruweit, Min & Yang 2018 -- (Li et al. 2019:346 [ref. 37175], Zhou et al. 2021:12 [ref. 38386], Nguyen et al. 2021:8 [ref. 38733], Min et al. 2022:112 [ref. 39113], Li et al. 2022:17 [ref. 39800], Che et al. 2023:284 [ref. 40280], Endruweit 2025:143 [ref. 42401] in subgenus Plerotula). Current status: Valid as Homatula coccinocola Endruweit, Min & Yang 2018. Nemacheilidae. Distribution: Southeast Asia: upper Black River, Red River basin, Yunnan Province (China). Habitat: freshwater.

cryptoclathrata, Homatula Li [X.], Che [X.-J.] & Zhou [W.] 2019:340, Figs. 2B, 4A, 5B, 7B, 9 [Zootaxa 4711 (no. 2); ref. 37175] Manging Village, Gengga Town, Changning County, Baoshan City, Yunnan, China, 24°19'42.65"N, 99°32'15.39"E. Holotype: SWFC 1002001. Paratypes: SWFC. •Valid as Homatula cryptoclathrata Li, Che & Zhou 2019 -- (Zhou et al. 2021:12 [ref. 38386], Min et al. 2022:112 [ref. 39113], Li et al. 2022:17 [ref. 39800]). Current status: Valid as Homatula cryptoclathrata Li, Che & Zhou 2019. Nemacheilidae. Distribution: East Asia: upper Salween River basin [Nujiang], Yunnan Province, China. Habitat: freshwater.

disparizona, Homatula Min [R.], Yang [J.-X.] & Chen [X.-Y.] 2013:351, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 23 (no. 4) (for 2012); ref. 32529] Panlong River (Red River drainage), Wenshan City, Xichou County, Yunnan Province, China. Holotype: KIZ 2012000623. Paratypes: KIZ 2012000622, 0624-0634 (12), 2012001859-1865 (7). •Valid as Homatula disparizona Min, Yang & Chen 2013 (Kottelat 2013:199 [ref. 32989], Endruweit 2015:70 [ref. 33868], Endruweit et al. 2018:565 [ref. 35763], Yang et al. 2017:1141 [ref. 35923], Li et al. 2019:346 [ref. 37175], Zhou et al. 2021:11 [ref. 38386], Nguyen et al. 2021:8 [ref. 38733], Nguyen et al. 2021:8 [ref. 38733], Min et al. 2022:112 [ref. 39113], Che et al. 2023:284 [ref. 40280], Cao et al. 2025:708 [ref. 42007], Endruweit 2025:141 [ref. 42401] in subgenus Litula). Current status: Valid as Homatula disparizona Min, Yang & Chen 2013. Nemacheilidae. Distribution: Southeast Asia: Panlong River drainage, upper Red River basin, Yunnan Province (China). Habitat: freshwater.

dotui, Homatula Nguyen [D. T.], Wu [H.], Cao [L.] & Zhang [E] 2021:6, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 31 (no. 1) IEF-1142; ref. 38733] Hang 21 Cave, circa 36 km from Tourist Center of Phong Nka Ke Bang National Park, Son River drainage, Gianh River basin, Quang Binh Province, central Vietnam, 106°12'46"E, 17°24'53"N. Holotype: IHB 201300440. Paratypes: IEBR 00439 (1). •Valid as Homatula dotui Nguyen, Wu, Cao & Zhang 2021 -- (Cao et al. 2025:706 [ref. 42007], probably in a different genus). Current status: Valid as Homatula dotui Nguyen, Wu, Cao & Zhang 2021. Nemacheilidae. Distribution: Southeast Asia: Son River drainage, Gianh River basin, Quang Binh Province, central Vietnam. Habitat: freshwater.

erhaiensis, Paracobitis Zhu [S.-Q.] & Cao [W.-X.] 1988:95 [100], Figs. 1-2 [Acta Zootaxonomica Sinica v. 13 (no. 1); ref. 17743] Wase, eastern bank of Erhai Lake, Yunnan Province, China. Holotype: IHASW 64-VI-0012. Paratypes: IHASW 64-VI-0001 to 11 (11), 64-VI-0013 to 15 (3), 64-VI-0775 to 77 (3), 64-VI-0779 (1). English description on p. 100. •Valid as Paracobitis erhaiensis Zhu & Cao 1988 -- (Zhu 1989:36 [ref. 17744], Zhou & He 1993:5 [ref. 21971], Bănărescu & Nalbant 1995:444 [ref. 23187] with uncertain genus placement, Zhu 1995:109 [ref. 25213], Chen & Zhang in Chen 1998:51 [ref. 23556], Min et al. 2010:202 [ref. 30726]). •Synonym of Homatula anguillioides (Zhu & Wang 1985) -- (Endruweit et al. 2018:564 [ref. 35763]). •Valid as Homatula erhaiensis (Zhu & Cao 1988) -- (Hu & Zhang 2010:52 [ref. 30780], Gu & Zhang 2012:xx [ref. 31632], Min et al. 2012:313 [ref. 32341], Kottelat 2012:83 [ref. 32367], Kottelat 2013:199 [ref. 32989], Zhang et al. 2016:128 [ref. 34477], Li et al. 2022:16 [ref. 39800]). Current status: Valid as Homatula erhaiensis (Zhu & Cao 1988). Nemacheilidae. Distribution: East Asia: upper Mekong River basin, Yunnan, China. Habitat: freshwater.

gelao, Homatula Xiao [G.-B.] in Xiao, Guo, Cao, Lin, Deng, Xiao, Zhou, Zhang & Fang 2025:259, Figs. 2, 3A, 4A, 5A, 6-7, 9 [Zoosystematics and Evolution v. 101 (no. 1); ref. 41908] Huaixi-He, a stream tributary to Furong-jiang of Wujiang basin, at Huaiping Village, Daozheng County, Zunyi City, Guizhou Province, China, 28°43'39.2124"N, 107°45'2.7072"E, elevation about 633 meters. Holotype: ZVT 20200601. Paratypes: ZVT 20200602-6029 (28). •Valid as Homatula gelao Xiao 2025. Current status: Valid as Homatula gelao Xiao 2025. Nemacheilidae. Distribution: East China: Wujiang River drainage, Yangtze River basin, Guizhou Province, China. Habitat: freshwater.

geminusclathratus, Homatula Li [X.], Yang [B.], Guo [Y.] & Zhou [W.] 2022:7, Figs. 1, 2A, 3A [PLoS ONE v. 17 (no. 11) (art. e0276847); ref. 39800] Xiaobahe on the eastern slope of Wuliangshan, Chuan-he drainage, Wenjing Town, Jingdong County, Pu'er City, Yunnan, China, 24°19'53.97"N, 100°48'37.17"E. Holotype: SWFC 0309250. Paratypes: SWFC 039251-72 (22). Combined with a feminine genus, the adjective should be geminusclathrata. •Valid as Homatula geminusclathrata Li, Yang, Guo & Zhou 2022 -- (Endruweit 2025:143 [ref. 42401] in subgenus Pleroula) Current status: Valid as Homatula geminusclathrata Li, Yang, Guo & Zhou 2022. Nemacheilidae. Distribution: Southeast Asia: upper Black River drainage, Red River basin, Yunnan (China). Habitat: freshwater.

guanheensis, Homatula Zhou [C.-J.], Ma [W.-W.], Wang [X.], Tang [Y.-T.], Meng [X.-L.] & Nie [G.-X.] 2021:5, Fig. 1 [Biodiversity Data Journal 9: e65130; ref. 38386] Guanhe River, tributary of Hanjiang River, Yangtze River basin, Xixia County, Henan Province, China, 33°52'79.3"N, 111°70'94"E. Holotype: HNU 010048. Paratypes: HNU 010049-56 (8). •Valid as Homatula guanheensis Zhou, Ma, Wang, Tang, Meng & Nie 2021 -- (Min et al. 2022:112 [ref. 39113], Xiao et al. 2025:269 [ref. 41908], Cao et al. 2025:708 [ref. 42007]). Current status: Valid as Homatula guanheensis Zhou, Ma, Wang, Tang, Meng & Nie 2021. Nemacheilidae. Distribution: East Asia: Hanjiang River drainage, Yangtze River basin, Henan and Shaanxi Provinces, China. Habitat: freshwater.

laxiclathra, Homatula Gu [J.-H.] & Zhang [E] 2012:592, Fig. 1a, 4a [Environmental Biology of Fishes v. 94 (no. 4); ref. 31632] Wei River, Yellow River drainage, Zhouzhi County, Shaanxi Province, China. Holotype: IHB 73V10738. Paratypes: 11 lots, 21 specimens at IHB. •Valid as Homatula laxiclathra Gu & Zhang 2012 -- (Min et al. 2012:313 [ref. 32341], Kottelat 2012:83 [ref. 32367], Zhang et al. 2016:128 [ref. 34477], Zhou et al. 2021:12 [ref. 38386], Nguyen et al. 2021:9 [ref. 38733], Min et al. 2022:112 [ref. 39113], Che et al. 2023:284 [ref. 40280], Xiao et al. 2025:269 [ref. 41908], Cao et al. 2025:708 [ref. 42007]). Current status: Valid as Homatula laxiclathra Gu & Zhang 2012. Nemacheilidae. Distribution: East Asia: Weihe River drainage, Yellow River basin, Shaanxi Province, China. Habitat: freshwater.

longibarbatus, Homatula Li [X.], Yang [B.], Guo [Y.] & Zhou [W.] 2022:11, Figs. 2B, 3B, 6 [PLoS ONE v. 17 (no. 11) (art. e0276847); ref. 39800] Pingpo Town, Yangbi County, Dali Prefecture, Yunnan, China, 25°35'53.36"N, 100°3'36.10"E. Holotype: SWFC 0309082. Paratypes: SWFC. Plus additional non-type material. Combined with a feminine genus, the adjective should be longibarbata. •Valid as Homatula longibarbata Li, Yang, Guo & Zhou 2022. Current status: Valid as Homatula longibarbata Li, Yang, Guo & Zhou 2022. Nemacheilidae. Distribution: East Asia: upper Mekong River basin, Yunnan, China. Habitat: freshwater.

longidorsalis, Paracobitis variegatus Yang [J.-X.], Chen [Y.-R.] & Kottelat [M.] 1994:63, Figs. 2B, 3 [Zoological Research v. 15, Supplement; ref. 28152] Jiuxiang, 25°06'N, 103°24'E, Yiliang County, Yunnan Province, China. Holotype: KIZ 874048. Paratypes: CMK 5871 (? 1 or 4 or 5); KIZ 874042-43 (2), 874045-47 (3), 874050 (1), 874198-216 (19). Plus non-type material. Type catalog: Du et al. 2013:351 [ref. 32939]. Cao et al. 2025:709 [ref. 42007] have holotype KIZ 1987003989. •Synonym of Paracobitis variegatus (Dabry de Thiersant 1874) but a valid subspecies as described -- (Chen et al. 1998:60 [ref. 23721], Min et al. 2010:202 [ref. 30726]). •Synonym of Homatula variegata (Dabry de Thiersat 1874) -- (Hu & Zhang 2010:59 [ref. 30780]). •Valid as Homatula longidorsalis (Yang, Chen & Kottelat 1994) -- (Gu & Zhang 2012:597 [ref. 31632], Min et al. 2012:80 [ref. 32392], Kottelat 2012:83 [ref. 32367], Endruweit et al. 2018:565 [ref. 35763], Li et al. 2019:347 [ref. 37175], Zhou et al. 2021:12 [ref. 38386], Nguyen et al. 2021:9 [ref. 38733], Min et al. 2022:111 [ref. 39113], Che et al. 2023:284 [ref. 40280], Xiao et al. 2025:270 [ref. 41908], Cao et al. 2025:709 [ref. 42007]). Current status: Valid as Homatula longidorsalis (Yang, Chen & Kottelat 1994). Nemacheilidae. Distribution: East Asia: upper Pearl River basin, Yunnan Province, China. Habitat: freshwater.

microcephala, Homatula Li [X.], Yang [B.], Guo [Y.] & Zhou [W.] 2022:14, Figs. 2C, 3C, 7 [PLoS ONE v. 17 (no. 11) (art. e0276847); ref. 39800] Shili-he, Jiancao Town, Yunlong County, Dali Prefecture, Yunnan, China, 26°3'13.58"N, 99°19'17.94"E. Holotype: SWFC 0612057. Paratypes: SWFC. Plus additional non-type material. The species epithet has to be treated as an adjective. •Valid as Homatula microcephala Li, Yang, Guo & Zhou 2022. Current status: Valid as Homatula microcephala Li, Yang, Guo & Zhou 2022. Nemacheilidae. Distribution: East Asia: upper Bi-jiang River drainage, upper Mekong River basin, Yunnan, China. Habitat: freshwater.

nanpanjiangensis, Paracobitis Min [R.], Chen [X.-Y.] & Yang [J.-X.] 2010:200, Fig. 1 [Environmental Biology of Fishes v. 87 (no. 3); ref. 30726] Tributary of upper Nanpanjiang River drainage, Niujie River, 24°57'46.9"N, 104°13'09.2"E, at Nanjie village, Luoping County, Qujing City, Yunnan, China. Holotype: KIZ 1994000023. Paratypes: KIZ 194000018-22 (5), 1994000024-37 (14). Type catalog: Du et al. 2013:351 [ref. 32939]. •Valid as Homatula nanpanjiangensis (Min, Chen & Yang 2010) -- (Gu & Zhang 2012:597 [ref. 31632], Min et al. 2012:313 [ref. 32341], Kottelat 2012:83 [ref. 32367], Min et al. 2013:354 [ref. 32529], Zhang et al. 2016:128 [ref. 34477], Endruweit et al. 2018:565 [ref. 35763], Zhou et al. 2021:11 [ref. 38386], Nguyen et al. 2021:8 [ref. 38733], Min et al. 2022:111 [ref. 39113], Che et al. 2023:284 [ref. 40280], Cao et al. 2025:709 [ref. 42007]). Current status: Valid as Homatula nanpanjiangensis (Min, Chen & Yang 2010). Nemacheilidae. Distribution: East Asia: Nanpanjiang River drainage, upper Pearl River basin, Yunnan Province, China. Habitat: freshwater.

nigra, Homatula Li [X.], Che [X.-J.] & Zhou [W.] 2019:341, Figs. 2C, 7C, 10 [Zootaxa 4711 (no. 2); ref. 37175] Xiangshui Village, Gengga Town, Changning County, Baoshan City, Yunnan, China, 24°29'7.26"N, 99°29'39.94"E. Holotype: SWFC 1901005. Paratypes: SWFC. •Valid as Homatula nigra Li, Che & Zhou 2019 -- (Zhou et al. 2021:11 [ref. 38386], Li et al. 2022:16 [ref. 39800] as niger). Current status: Valid as Homatula nigra Li, Che & Zhou 2019. Nemacheilidae. Distribution: East Asia: upper Salween River basin [Nujiang], Yunnan, China. Habitat: freshwater.

oligolepis, Paracobitis variegatus Cao [W.-X.] & Zhu [S.-Q.] in Zheng et al. 1989:48 [Engl. p. 410], Fig. 27 [Fishes of the Zhujiang River; ref. 21202] Yangzonghai, Yunnan Province, China. Holotype: IHASW 63-V-553. Paratypes: IHASW 63-V-465 (1), 63-V-551 to 552 (1, 1), 63-V-559 (1), 63-V-1077 (1). •Valid species, may be in genus Homatula -- (Bănărescu & Nalbant 1995:444 [ref. 23187]). •Synonym of Paracobitis variegatus (Dabry de Thiersant 1874), but a valid subspecies as described -- (Zheng et al. 1989:48 [ref. 21202]). •Valid as Paracobitis oligolepis Cao & Zhu 1989 -- (Zhu 1989:35 [ref. 17744], Zhu 1995:109 [ref. 25213], Min et al. 2010:202 [ref. 30726]). •Valid as Homatula oligolepis (Cao & Zhu 1989) -- (Hu & Zhang 2010:52 [ref. 30780], Gu & Zhang 2012:597 [ref. 31632], Min et al. 2012:313 [ref. 32341], Kottelat 2012:84 [ref. 32367], Min et al. 2013:354 [ref. 32529], Zhang et al. 2016:128 [ref. 34477], Endruweit et al. 2018:565 [ref. 35763], Li et al. 2019:347 [ref. 37175], Zhou et al. 2021:11 [ref. 38386], Nguyen et al. 2021:8 [ref. 38733], Min et al. 2022:111 [ref. 39113], Che et al. 2023:284 [ref. 40280], Cao et al. 2025:709 [ref. 42007]). Current status: Valid as Homatula oligolepis (Cao & Zhu 1989). Nemacheilidae. Distribution: East Asia: Yangzong Lake drainage, upper Mekong River basin, Yunnan Province, China. Habitat: freshwater.

oxygnathus, Nemachilus Regan [C. T.] 1908:357 [Annals and Magazine of Natural History (Series 8) v. 2 (no. 10); ref. 14350] Yunnan Fu [Kunming City], China. Lectotype: BMNH 1908.2.27.23. Paralectotype: BMNH 1908.2.27.24 (1). Original genus is correctly Nemacheilus. Yang et al. 1994:62 [ref. 28152] mention a lectotype for this species, but evidently no lectotype was designated; they mention syntypes on p. 60. Lectotype selected as BMNH 1908.2.27-24 by Bǎnǎrescu & Nalbant 1974, which is 1908.2.27.23. •Synonym of Paracobitis variegatus (Dabry de Thiersant 1874) -- (Zhu 1989:32 [ref. 17744], Wu & Wu 1992:140 [ref. 21205], Yang & Chen 1994:59 [ref. 28152], Chen & Zhang in Chen 1998:49 [ref. 23556]). •Synonym of Homatula variegata (Dabry de Thiersant 1874) -- (Bănărescu & Nalbant 1995:444 [ref. 23187], Kottelat 2012:84 [ref. 32367], Zhang et al. 2019:364 [ref. 36699]). •Valid as Homatula oxygnathus (Regan 1908) -- (Liu et al. 2022:[9] [ref. 39342] as oxygnathra, Xiao et al. 2025:263 [ref. 41908] as oxygnathra, Cao et al. 2025:709 [ref. 42007] as oxygnathra). Current status: Valid as Homatula oxygnathus (Regan 1908). Nemacheilidae. Distribution: East Asia: upper Yangtze River basin, Sichuan and Yunnan Provinces, China. Habitat: freshwater.

potanini, Nemachilus Günther [A.] 1896:218 [20] [Ezhegodnik Zoologicheskogo Muzeya Imperatorskoy Akademii Nauk 1896, v. 1 (no. 3); ref. 2019] Ya River, China. Syntypes: ZISP 10005 (3). Original genus is correctly Nemacheilus. On p. 20 of separate. •Valid as Paracobitis potanini (Günther 1896) -- (Zhu 1989:37 [ref. 17744], Wu & Wu 1992:142 [ref. 21205], Ding 1994:54 [ref. 21531], Zhu 1995:109 [ref. 25213], Chen & Zhang in Chen 1998:51 [ref. 23556], Tang et al. 2001:14 [ref. 25373], Min et al. 2010:203 [ref. 30726]). •Valid as Homatula potanini (Günther 1896) -- (Bănărescu & Nalbant 1995:444 [ref. 23187], Hu & Zhang 2010:52 [ref. 30780], Gu & Zhang 2012:597 [ref. 31632], Min et al. 2012:313 [ref. 32341], Min et al. 2012:82 [ref. 32392], Kottelat 2012:84 [ref. 32367], Zhang et al. 2016:129 [ref. 34477], Endruweit et al. 2018:565 [ref. 35763], Li et al. 2019:347 [ref. 37175], Zhang et al. 2019:366 [ref. 36699], Du et al. 2021:4 [ref. 38600], Zhou et al. 2021:10 [ref. 38386], Guo 2021:496 [ref. 38618], Nguyen et al. 2021:9 [ref. 38733], Min et al. 2022:112 [ref. 39113], Che et al. 2023:284 [ref. 40280], Xiao et al. 2025:269 [ref. 41908], Cao et al. 2025:709 [ref. 42007]). Current status: Valid as Homatula potanini (Günther 1896). Nemacheilidae. Distribution: East Asia: upper Yangtze River basin (Sichuan Province, China). Habitat: freshwater.

pycnolepis, Homatula Hu [Y.-T.] & Zhang [E] 2010:52, Figs. 1, 2a, 3a, 5a, 6a, 7a [Ichthyological Exploration of Freshwaters v. 21 (no. 1); ref. 30780] Yangbi River, tributary of upper Mekong at Shaxi Town, about 26°19'N, 99°51'E, Jianchuan County, Yunnan Province, China. Holotype: IHASW 814045. Paratypes: IHASW 814042-43 (2), 814046 (1), 814047 (1, stained), 814048-51 (4). •Valid as Homatula pycnolepis Hu & Zhang 2010 -- (Gu & Zhang 2012:597 [ref. 31632], Min et al. 2012:313 [ref. 32341], Min et al. 2012:81 [ref. 32392], Kottelat 2012:84 [ref. 32367], Kottelat 2013:199 [ref. 32989], Endruweit 2015:70 [ref. 33868], Zhang et al. 2016:129 [ref. 34477], Endruweit et al. 2018:565 [ref. 35763], Li et al. 2019:343 [ref. 37175], Zhou et al. 2021:12 [ref. 38386], Nguyen et al. 2021:9 [ref. 38733], Min et al. 2022:112 [ref. 39113], Li et al. 2022:17 [ref. 39800], Xiao et al. 2025:270 [ref. 41908], Zeng et al. 2025:18 [ref. 42440]). Current status: Valid as Homatula pycnolepis Hu & Zhang 2010. Nemacheilidae. Distribution: Southeast Asia: upper Mekong River basin and Irrawaddy River basin, Yunnan (China). Habitat: freshwater.

robusta, Homatula Min [R.], Zhao [Y.-H.], Shi [J.-S.] & Yang [J.-X.] 2022:113, Figs. 1-3A, 4A [ZooKeys No. 1089; ref. 39113] Nanpanjiang River at Changdi village, upper Pearl River, Luoping County, Qujing City, Yunnan Province, China, 25°02'N, 104°30'E, elevation circa 1,210 meters. Holotype: KIZ 2009000125. Paratypes: KIZ 2009000122 (1), 2009000144 (1) 2009000146 (1). •Valid as Homatula robusta Min, Zhao, Shi & Yang 2022. Current status: Valid as Homatula robusta Min, Zhao, Shi & Yang 2022. Nemacheilidae. Distribution: East Asia: Nanpanjiang River drainage, upper Pearl River basin, Yunnan Province, China. Habitat: freshwater.

shexiang, Homatula Cao [L.], Liu [Y.], Zeng [Z.] & Zhang [E] in Cao, Liu, Zeng, Yi & Zhang 2025:702, Figs. 4-5, 8B [Zoosystematics and Evolution v. 101 (no. 2); ref. 42007] Maodong-He, stream tributary to Maotiao-He, under Wugong Bridge, Xiuwen District, Guiyang City, Guizhou Province, China, 26°54'11"N,106°29'22"E. Holotype: IHB 202006048998. Paratypes: (16) IHB. •Valid as Homatula shexiang Cao, Liu, Zeng & Zhang 2025. Current status: Valid as Homatula shexiang Cao, Liu, Zeng & Zhang 2025. Nemacheilidae. Distribution: East Asia: Wujiang River drainage, upper Yangtze River basin, Guizhou Province, China. Habitat: freshwater.

submarginata, Homatula (Plerotula) Endruweit [M.] 2025:143, Fig. 141 [Fishes of the Red River - Cypriniformes, Siluriformes; ref. 42401] Baohua, ca. 24°54'N 100°30'E, Nanjian County, Dali prefecture, Yunnan (China). Holotype: KIZ 2015005112. Paratypes: KIZ 2016001924-9 (6), 2016001916-23 (8). •Valid as Homatula submarginata Endruweit 2025. Current status: Valid as Homatula submarginata Endruweit 2025. Nemacheilidae. Distribution: Southeast Asia: Black River system (China). Habitat: freshwater.

tigris, Homatula Che [X.-J.], Dao [W.], Chen [Y.-C.], Pan [X.-F.], Hua [Z.-X.], Liang [X.] & Wang [X.-A.] 2023:280, Figs. 1-2 [Ichthyological Exploration of Freshwaters v. 31 (no. 3): IEF-1187; ref. 40280] Muyang-he River, Panlong-jiang, Dianyuan Town, Panlong District, Yunnan Province, China, 25°14'02"N, 102°49'52"E, elevation 2,003 meters. Holotype: KIZ 202100004152. Paratypes: KIZ 202100004144-4151 (8). •Valid as Homatula tigris Che, Dao, Chen, Pan, Hua, Liang & Wang 2023. Current status: Valid as Homatula tigris Che, Dao, Chen, Pan, Hua, Liang & Wang 2023. Nemacheilidae. Distribution: East Asia: Panlong-jiang drainage, upper Yangtze River basin, Yunnan, China. Habitat: freshwater.

variegatus, Nemachilus Dabry de Thiersant [P.] in Sauvage & Dabry de Thiersant 1874:14 [Annales des Sciences Naturelles, Paris (Zoologie et Paléontologie) (Sér. 6) v. 1 (art. 5); ref. 15665] Tchen-si, Wei River, China. Lectotype: MNHN 0000-7854. Paralectotype: MNHN B-2641 [ex MNHN 0000-7854] (1). Type catalog: Bertin & Estève 1948:98 [ref. 19574]. Original genus is correctly Nemacheilus. Lectotype selected by Bănărescu & Nalbant 1974:96 [ref. 28153]. •Valid as Paracobitis variegatus (Dabry de Thiersant 1874) -- (Zheng et al. 1989:48 [ref. 21202], Zhu 1989:32 [ref. 17744], Wu & Wu 1992:140 [ref. 21205], Ding 1994:51 [ref. 21531], Yang & Chen 1994:60 [ref. 28152] with subspecies, Zhu 1995:108 [ref. 25213], Chen & Zhang in Chen 1998:49 [ref. 23556], Tang et al. 2001:14 [ref. 25373], Min et al. 2010:202 [ref. 30726] with subspecies). •Valid as Homatula variegata (Dabry de Thiersant 1874) -- (Bănărescu & Nalbant 1995:444 [ref. 23187], Hu & Zhang 2010:52 [ref. 30780], Gu & Zhang 2012:597 [ref. 31632], Min et al. 2012:313 [ref. 32341], Min et al. 2012:80 [ref. 32392] as variegatus, Kottelat 2012:84 [ref. 32367], Zhang et al. 2016:129 [ref. 34477] as variegatus, Endruweit et al. 2018:565 [ref. 35763], Li et al. 2019:347 [ref. 37175], Zhang et al. 2019:364 [ref. 36699], Zhou et al. 2021:12 [ref. 38386], Guo 2021:490 [ref. 38618] as variegatus, Nguyen et al. 2021:8 [ref. 38733], Liu et al. 2022:[3] [ref. 39342], Min et al. 2022:112 [ref. 39113], Che et al. 2023:284 [ref. 40280], Xiao et al. 2025:263 [ref. 41908], Cao et al. 2025:709 [ref. 42007]). Current status: Valid as Homatula variegata (Dabry de Thiersant 1874). Nemacheilidae. Distribution: East Asia: Weihe River drainage, middle Yellow River basin and Yangtze River basin (Shaanxi Province, China). Habitat: freshwater.

wenshanensis, Homatula Li [C.-Q.], Yang [H.-F.], Li [W.-X.] & Liu [T.] in Yang, Li, Liu & Li 2017:1140, Figs. 1a-d, 2 [Journal of Yunnan Agricultural University (Natural Sciences) v. 32 (no. 6); ref. 35923] River in Xigu Township, Wenshan City, Yunnan, China, 23°23'N, 104°05'E. Holotype: HRA 2016072508. Paratypes: HRA [?]. •Valid as Homatula wenshanensis Li, Yang, Li & Liu 2017 -- (Li et al. 2019:347 [ref. 37175], Zhou et al. 2021:11 [ref. 38386], Min et al. 2022:112 [ref. 39113], Endruweit 2025:142 [ref. 42401] in subgenus Litula). •Not in Homatula -- (Nguyen et al. 2021:8 [ref. 38733]). Current status: Valid as Homatula wenshanensis Li, Yang, Li & Liu 2017. Nemacheilidae. Distribution: Southeast Asia: upper Red River basin, Wenshan Prefecture, Yunnan Provincem (China). Habitat: freshwater.

wujiangensis, Paracobitis Ding [R.-H.] & Deng [Q.-X.] 1990:287 (English p. 290), Figs. 1-3 [Zoological Research v. 11 (no. 4); ref. 21065] Wujiang River, Sichuan, China. Holotype: SPNRI 64274. Paratypes: SPNRI 64276-78 (1, 1, 1), 84200 (1), 87277-78 (1, 1). •Valid as Paracobitis wujiangensis Ding & Deng 1990 -- (Ding 1994:55 [ref. 21531], Bănărescu & Nalbant 1995:444 [ref. 23187] with uncertain genus placement, Zhu 1995:109 [ref. 25213], Tang et al. 2001:14 [ref. 25373], Min et al. 2010:202 [ref. 30726]). •Valid as Homatula wujiangensis (Ding & Deng 1990) -- (Hu & Zhang 2010:52 [ref. 30780], Gu & Zhang 2012:597 [ref. 31632], Min et al. 2012:313 [ref. 32341], Kottelat 2012:85 [ref. 32367], Kottelat 2012:85 [ref. 32367], Min et al. 2013:354 [ref. 32529], Zhang et al. 2016:129 [ref. 34477], Zhou et al. 2021:10 [ref. 38386], Guo 2021:499 [ref. 38618], Nguyen et al. 2021:8 [ref. 38733], Che et al. 2023:284 [ref. 40280], Xiao et al. 2025:269 [ref. 41908], Cao et al. 2025:709 [ref. 42007]). Current status: Valid as Homatula wujiangensis (Ding & Deng 1990). Nemacheilidae. Distribution: East Asia: Wujiang River drainage, upper Yangtze River basin, Sichuan and Chongqing Provinces, China. Habitat: freshwater.

wuliangensis, Homatula Min [R.], Yang [J.-X.] & Chen [X.-Y.] 2012:315, Fig. 1 [Zootaxa No. 3586; ref. 32341] Wuliang Mountain (24°17′26.6″N, 100°39′03.6″E-24°20′37.5″N, 100°41′18.2″E), Lancang River, at Jingdong County, Pu-Er City, China, altitude 1274-1781 meters. Holotype: KIZ 2008008158. Paratypes: many at KIZ (33). Type catalog: Du et al. 2013:351 [ref. 32939]. •Valid as Homatula wuliangensis Min, Yang & Chen 2012 -- (Kottelat 2013:200 [ref. 32989], Endruweit 2015:70 [ref. 33868], Zhang et al. 2016:129 [ref. 34477], Endruweit et al. 2018:565 [ref. 35763], Li et al. 2019:347 [ref. 37175], Zhou et al. 2021:12 [ref. 38386], Min et al. 2022:112 [ref. 39113], Li et al. 2022:17 [ref. 39800]). Current status: Valid as Homatula wuliangensis Min, Yang & Chen 2012. Nemacheilidae. Distribution: East Asia: upper Mekong River basin, Yunnan Province, China. Habitat: freshwater.

xiangzhi, Homatula Cao [L.], Liu [Y.], Zeng [Z.] & Zhang [E] in Cao, Liu, Zeng, Yi & Zhang 2025:698, Figs. 1-2, 8A [Zoosystematics and Evolution v. 101 (no. 2); ref. 42007] Baishui-He, tributary of Qingshui-He discharging into Wu-Jiang, at Shanglongjiao Village of Xiangzhi-Gou, Wudang District, Guiyang City, Guizhou Province, China, 26°46'21"N, 106°55'02"E. Holotype: IHB 202006048865. Paratypes: (7) IHB. •Valid as Homatula xiangzhi Cao, Liu, Zeng & Zhang 2025. Current status: Valid as Homatula xiangzhi Cao, Liu, Zeng & Zhang 2025. Nemacheilidae. Distribution: East Asia: Wu Jiang River drainage, upper Yangtze River basin, and upper Pearl River basin, Guizhou Province, China. Habitat: freshwater.