Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the genus Lycodes: [ 105 ] records

adolfi, Lycodes Nielsen [J. G.] & Fosså [S. A.] 1993:39, Figs. 1-2 [Cybium v. 17 (no. 1); ref. 20655] Greenland, 68°49'N, 19°44'W, depth 1371-1380 meters. Holotype: ZMUC P761186. Paratypes: ZIN 50515 [ex.ZUMC 761188] (1); ZMUC P761187 (1), P751189-90 (3), P761191 (1), P761192 (1). Type catalog: Balushkin et al. 2011:987 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. •Valid as Lycodes adolfi Nielsen & Fosså 1993 -- (Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:23 [ref. 27603], Møller et al. 2010:60 [ref. 31169], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin et al. 2011:987 [ref. 32449], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:140 [ref. 33463], Parin et al. 2014:406 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:63, 195 [ref. 36199], Mecklenburg et al. 2016:160 [ref. 34440], Dolgov et al. 2018:202 [ref. 41055], Mecklenburg et al. 2018:321 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:182 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes adolfi Nielsen & Fosså 1993. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

agnostus, Lycodes Jensen [A. S.] 1902:209 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Kara-Havet, Kara Sea, depth 46-100 fathoms. Syntypes: ZIN 40714 [ex. ZUMC P22] (1); ZMUC 3, 6, 11, 16, 20, 22, 24-27, 29. Type catalog: Nielsen 1974:76 [ref. 9588], Balushkin et al. 2011:1006 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. Specimens ZMUO J4490 (3) are labeled types but are not types according to Pethon 1969:11 [ref. 19268]. •Synonym of Lycodes polaris (Sabine 1824) -- (Andriashev 1973:545 [ref. 9214], Scott & Scott 1988:410 [ref. 25518], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:710 [ref. 25968], Anderson & Fedorov 2004:28 [ref. 27603], Balushkin et al. 2011:1006 [ref. 32449], Parin et al. 2014:418 [ref. 33547]). Current status: Synonym of Lycodes polaris (Sabine 1824). Zoarcidae: Lycodinae. Habitat: marine.

agulhensis, Lycodes Andriashev [A. P.] 1959:465, Figs. 1-3 [Zoologicheskii Zhurnal v. 38 (no. 3); ref. 11975] Agulhas Bank, 35°44.2-46'S, 22°33.5 -36.2'E, South Africa, southeastern Atlantic, depth 1400 meters. Holotype (unique): ZIN 35812. Type catalog: Balushkin et al. 2011:987 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. •Synonym of Lycodes terraenovae Collett 1896 -- (Møller 1997:48 [ref. 22879], Møller 2000:847 [ref. 24585], Carl 2002:65 [ref. 26925], Anderson & Fedorov 2004:30 [ref. 27603]). •Valid as Lycodes agulhensis Andriachev 1959 -- (Anderson 1986:342 [ref. 5633], Anderson & Almeida 1990:888 [ref. 19328], Anderson 1994:118 [ref. 21438], Balushkin et al. 2011:987 [ref. 32449]). Current status: Valid as Lycodes agulhensis Andriashev 1959. Zoarcidae: Lycodinae. Distribution: Agulhas Bank, South Africa. Habitat: marine.

akuugun, Lycodes Stevenson [D. E.] & Orr [J. W.] 2006:78, Figs. 1-2 [Copeia 2006 (no. 1); ref. 28543] South of Seguam Island, 52.210°N, 172.206°W, central Aleutian Islands, depth 347 meters. Holotype: UW 112519. Paratypes: CAS 222668 (1); LACM 56292-1 (1); UW 112520 (1), 112521 (2), 112522 (1), 112523 (1 c&s), 112524 (1), 112525 (1); USNM 384244 (1). •Valid as Lycodes akuugun Stevenson & Orr 2006 -- (Balushkin et al. 2011:1026 [ref. 32449], Page et al. 2013:162 [ref. 32708], Love et al. 2021:182 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes akuugun Stevenson & Orr 2006. Zoarcidae: Lycodinae. Distribution: North Pacific: central Aleutian Islands (U.S.A.). Habitat: marine.

albolineatus, Lycodes Andriashev [A. P.] 1955:397, Figs. 4-6 [Trudy Zoologicheskogo Instituta, Akademii Nauk SSSR v. 21; ref. 9472] Near Cape Lopatka, southeastern coast of Kamchatka, 50°59'N, 157°54"E, depth 241-270 meters. Holotype: ZIN 34582. Paratypes: BMNH 1979.8.4.5 [ex ZIN] (1), ZIN 34581 (4, now 3). Type catalog: Balushkin et al. 2011:987 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667].. Holotype established in Fig. 4, 605 mm. •Valid as Lycodes albolineatus Andriashev 1955 -- (Toyoshima 1985:232 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Møller 2001:973 [ref. 25812], Tokranov & Orlov 2002:605 [ref. 26464], Anderson & Fedorov 2004:23 [ref. 27603], Balushkin et al. 2011:987 [ref. 32449], Parin et al. 2014:406 [ref. 33547]). Current status: Valid as Lycodes albolineatus Andriashev 1955. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

andriashevi, Lycodes Fedorov [V. V.] 1966:160, Figs. 1-2 [Voprosy Ikhtiologii v. 6 (no. 1); ref. 20765] Bering Sea, 54°26'N, 116°13'W, depth 550 meters. Holotype (unique): ZIN 37515. Type catalog: Balushkin et al. 2011:992 [ref. 32449], Balushkin et al. 2012:156 [ref. 33667]. •Valid as Lycodes andriashevi Fedorov 1966 -- (Toyoshima 1985:235 [ref. 5722]). •Synonym of Lycodes concolor Gill & Townsend 1897 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:725 [ref. 25968], Møller 2001:975 [ref. 25812], Anderson & Fedorov 2004:24 [ref. 27603], Balushkin et al. 2011:992 [ref. 32449], Parin et al. 2014:410 [ref. 33547]). Current status: Synonym of Lycodes concolor Gill & Townsend 1897. Zoarcidae: Lycodinae. Habitat: marine.

arcticus, Lycodes palearis Taranetz [A. Ya.] & Andriashev [A. P.] in Andriashev 1937:333, Pl. 1 (figs. 9-10) [Issled. Morey SSSR, Leningrad v. 25; ref. 20242] Gulf of Anadyr, Bering Sea; Chukchi Sea, depth 84-110 meters. Syntypes: ZIN 26885 (1), 29985 (5). Additional material: ZIN 37927 (9). Type catalog: Balushkin et al. 2011:1002 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. Name mentioned in Taranetz & Andriashev 1935:253 (footnote 4) [ref. 15744]. •Synonym of Lycodes palearis Gilbert 1896 -- (Toyoshima 1985:212 [ref. 5722], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:719 [ref. 25968], Anderson & Fedorov 2004:27 [ref. 27603], Mecklenburg et al. 2007:182 [ref. 29448], Balushkin et al. 2011:1002 [ref. 32449], Parin et al. 2014:417 [ref. 33547]). Current status: Synonym of Lycodes palearis Gilbert 1896. Zoarcidae: Lycodinae. Habitat: marine.

atlanticus, Lycodes Jensen [A. S.] 1902:207 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Coast of North America, off Virginia, U.S.A., 39°34'45"N, 71°21'30"W, depth 516-1423 fathoms. Holotype (unique): ZMUC 536. Type catalog: Nielsen 1974:76 [ref. 9588]. •Valid as Lycodes atlanticus Jensen 1902 -- (Scott & Scott 1988:406 [ref. 25518], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Moore et al. 2003:229 [ref. 27076]). •Synonym of Lycodes terraenovae Collett 1896 -- (Møller 1997:47 [ref. 22879], Møller 2000:847 [ref. 24585], Anderson & Fedorov 2004:30 [ref. 27603], Balushkin et al. 2011:1015 [ref. 32449]). Current status: Synonym of Lycodes terraenovae Collett 1896. Zoarcidae: Lycodinae. Habitat: marine.

atlanticus, Lycodes turneri Vladykov [V. D.] & Tremblay [J.-L.] 1936:38, Pl. 7 (figs. 14-16) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Saint Lawrence River estuary, Quebec, Canada. Syntypes: (18) NMC 60-160 (1). Type catalog: McAllister 1965:9 [ref. 22019] as paratypes. Objectively invalid; preoccupied by Lycodes atlanticus Jensen 1902. •Synonym of Lycodes turneri Bean 1879, subspecies atlanticus Vladykov & Tremblay 1936 -- (Backus 1951:291 [ref. 41140]). •Synonym of Lycodes polaris (Sabine 1824) -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603], Parin et al. 2014:418 [ref. 33547]). 1°Homonym. Current status: Synonym of Lycodes polaris (Sabine 1824). Zoarcidae: Lycodinae. Habitat: marine.

atratus, Lycodes Vladykov [V. D.] & Tremblay [J.-L.] 1936:26, Pl. 2 (fig. 6) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Saint Lawrence Estuary, Canada. Holotype (unique): whereabouts unknown. •Synonym of Lycodes atlanticus Jensen 1902 -- (Anderson 1994:118 [ref. 21438]). •Synonym of Lycodes terraenovae Collett 1896 -- (Møller 1997:48 [ref. 22879], Møller 2000:847 [ref. 24585], Anderson & Fedorov 2004:30 [ref. 27603]). Current status: Synonym of Lycodes terraenovae Collett 1896. Zoarcidae: Lycodinae. Habitat: marine.

attenuatus, Lycodes Knipowitsch [N. M.] 1906:73, Figs. 13-14 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Laptev Sea, Russia, 75°-77°N, 114°-139°E [3 stations], depth 19-38 meters. Syntypes: ZIN 14249-51 (1, 1, 1). Type catalog: Balushkin et al. 2011:988 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667]. Also described as new in Knipowitsch 1907:36 [ref. 15152]. •Synonym of Lycodes pallidus Collett 1879 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:27 [ref. 27603]). •Valid as Lycodes attenuatus Knipowitsch 1906 -- (Balushkin et al. 2011:988 [ref. 32449], Parin et al. 2014:407 [ref. 33547], but treated as a synonym of Lycodes pallidus (Collett 1879) on p. 417, Smirnova et al. 2019:491 [ref. 36948]). Current status: Valid as Lycodes attenuatus Knipowitsch 1906. Zoarcidae: Lycodinae. Distribution: Arctic and North Pacific, Habitat: marine.

bathybius, Lycodes Schmidt [P. Yu.] 1950:105 [91], Pl. 9 (fig. 2) [Trudy Tichookeanskogo Komiteta, Akademija Nauk S.S.S.R. v. 6; ref. 12471] Near southern extremity of Kamchatka, Russia, Okhotsk Sea, 51°09'N, 154°17'E, depth 591 meters. Holotype (unique): ZIN 24829. Type catalog: Balushkin et al. 2011:988 [ref. 32449], Balushkin et al. 2012:157 [ref. 33667]. On p. 91 of English translation. Appeared as name only in Schmidt 1935. •Status uncertain -- (Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:48 [ref. 27603]). •Valid as Lycodes bathybius Schmidt 1950 -- (Toyoshima 1985:203 [ref. 5722], Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:112 [ref. 27579], Balushkin et al. 2011:988 [ref. 32449], Parin et al. 2014:407 [ref. 33547]). Current status: Valid as Lycodes bathybius Schmidt 1950. Zoarcidae: Lycodinae. Distribution: Western North Pacific. Habitat: marine.

beringi, Lycodes (Furcimanus) diapterus Andriashev [A. P.] 1935:115, Fig. 1 [Comptes Rendus (Doklady) de l'Académie des Sciences de l'URSS v. 4 [9] (nos 1-2); ref. 13870] Off Commander Island, 6 miles west of Cape Yuchina, depth 200-250 meters. Lectotype: ZIN 29850. Paralectotypes: ZIN 34838 (4), 54299 (now 7). Type catalog: Balushkin et al. 2011:988 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. Subgenus Furcimanus misspelled in original. USNM paralectotypes of Lycodes diapterus are this species. Lectotype selected by Stevenson & Sheiko 2009:132 [ref. 30070]. •Synonym of Lycodes diapterus Gilbert 1892 -- (Toyoshima 1985:230 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:724 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603]). •Valid as Lycodes beringi Andriashev 1935 -- (Stevenson & Sheiko 2009:130 [ref. 30070], Balushkin et al. 2011:988 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:407 [ref. 33547], Pietsch & Orr 2015:58 [ref. 34694], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes beringi Andriashev 1935. Zoarcidae: Lycodinae. Distribution: North Pacific: Kamchatka Peninsula (Russia) to Puget Sound (Canada). Habitat: marine.

brashnikovi, Lycodes Soldatov [V. K.] 1918:112, Fig. 1 [An. Mus. Zool. Acad. Sci. Petrograd v. 22; ref. 21830] 53°17'N, 154°47'E and Senyavine Cape, eastern coast of Sakhalin Island, Okhotsk Sea. Syntypes: ZIN 19167 (3), ZIN (1, not found in 1996). Non-types: ZIN 13010 (2) questionably included. Type catalog: Balushkin et al. 2011:989 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. Also in Soldatov 1922:113, Fig. 1 [ref. 15429]. •Synonym of Lycodes palearis Gilbert 1896, subspecies schmidtii Gratzianov 1907 -- (Lindberg & Krasyukova 1975:158 [ref. 7348]). •Synonym of Lycodes fasciatus (Schmidt 1904) -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Synonym of Lycodes schmidti Gratzianov 1907 -- (Toyoshima 1985:214 [ref. 5722], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:29 [ref. 27603]). •Valid as Lycodes brashnikovi Soldatov 1918 -- (Nazarkin 2010:603 [ref. 31019], Balushkin et al. 2011:989 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:408 [ref. 33547], Dyldin & Orlov 2021:837 [ref. 38925]). Current status: Valid as Lycodes brashnikovi Soldatov 1918. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk Sea. Habitat: marine.

brevicauda, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] 1935:248, Figs. 4-5 [Zoologischer Anzeiger v. 112 (nos 9-10); ref. 15744] Okhotsk Sea. Syntypes: (25) ZIN 24825 (1), 29993 (1), ?30015 (2), 30190 (6), 30196 (3), 30986 (3), 31292 (2), 31393 (1). Type catalog: Balushkin et al. 2011:990 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667].. •Synonym of Lycodes tanakai Jordan & Thompson 1914 -- (Anderson 1994:119 [ref. 21438] as tanakae). •Synonym of Lycodes tanakai Jordan & Thompson 1914 -- (Toyoshima 1985:193 [ref. 5722], Anderson & Fedorov 2004:30 [ref. 27603]). •Valid as Lycodes brevicaudus Taranetz & Andriashev 1935 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:113 [ref. 27579], Balushkin et al. 2011:990 [ref. 32449], Parin et al. 2014:408 [ref. 33547], but as a synonym of Lycodes tanakae on p. 423, Dyldin & Orlov 2021:837 [ref. 38925] with question). Current status: Valid as Lycodes brevicaudus Taranetz & Andriashev 1935. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk Sea. Habitat: marine.

brevipes, Lycodes Bean [T. H.] 1890:38 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Between Unga Island and Nagai Island, Aleutian Islands, Alaska, U.S.A., 55°10'N, 160°18'W, Albatross station 2848, depth 110 fathoms [201 meters]. Lectotype: USNM 45362. Paralectotypes: USNM 162712 (16). Type catalog: Springer & Anderson 1997:14-15 [ref. 22953], Balushkin et al. 2011:991 [ref. 32449]. Lectotype selected by Springer & Anderson 1997:14-15 [ref. 22953] (lectotype also figured by them). •Valid as Lycodes brevipes Bean 1890 -- (Eschmeyer & Herald 1983:105 [ref. 9277], Toyoshima 1985:233 [ref. 5722], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:716 [ref. 25968], Anderson & Fedorov 2004:23 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Love et al. 2005:142 [ref. 37547], Balushkin et al. 2011:991 [ref. 32449], Page et al. 2013:162 [ref. 32708], Shinohara et al. 2014:265 [ref. 33330], Parin et al. 2014:408 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Pietsch & Orr 2015:58 [ref. 34694], Kells et al. 2016:272 [ref. 35888], Dyldin & Orlov 2021:837 [ref. 38925], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes brevipes Bean 1890. Zoarcidae: Lycodinae. Distribution: North Pacific and adjacent Arctic Ocean: eastern Chukchi Sea and Bering sea south to central California (U.S.A.). Habitat: marine.

brunneofasciatus, Lycodes Suvorov [E. K.] 1935:437 [English p. 440], Fig. 3 [Bulletin de l'Academie des Sciences de l'U.R.S.S., Classe des Sciences Mathématiques et Naturelles 1935; ref. 4300] Western Kamchatka, Russia, 53°35'N, 154°22'E, depth 250 meters. Holotype (unique): ZIN 25074. Type catalog: Balushkin et al. 2011:991 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. •Valid as Lycodes brunneofasciatus Suvorov 1935 -- (Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:234 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1037 [ref. 25182], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:715 [ref. 25968], Nakabo 2002:1037 [ref. 26193], Tokranov & Orlov 2002:605 [ref. 26464], Fedorov et al. 2003:113 [ref. 27579], Anderson & Fedorov 2004:24 [ref. 27603], Love et al. 2005:142 [ref. 37547], Balushkin et al. 2011:991 [ref. 32449], Shinohara et al. 2014:265 [ref. 33330], Parin et al. 2014:409 [ref. 33547], Sonoyama et al. 2020:94 [ref. 37637], Dyldin & Orlov 2021:837 [ref. 38925], Love et al. 2021:183 [ref. 39279] as XLycodes brunneofasciatus, Balanov & Kurbanov 2024:531 [ref. 41624]). Current status: Valid as Lycodes brunneofasciatus Suvorov 1935. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Sea of Japan and Sea of Okhotsk north to Kamchatka (Russia); Commander Islands (Russia), straying to Unalaska Island (Alaska, U.S.A.). Habitat: marine.

brunneus, Lycodes Fowler [H. W.] 1944:73, Fig. [The Fish Culturist v. 23 (no. 10); ref. 17879] North of Bahamas, east of Florida, Albatross station 2660, 28°40'N, 78°46'W, depth 504 fathoms. Holotype: USNM 119446. Paratypes: USNM 119447 (1). Type catalog: Springer & Anderson 1997:15 [ref. 22953]. •Synonym of Lycodes atlanticus Jensen 1902 -- (Anderson 1994:118 [ref. 21438]). •Synonym of Lycodes terraenovae Collett 1896 -- (Møller 1997:48 [ref. 22879], Møller 2000:847 [ref. 24585], Anderson & Fedorov 2004:30 [ref. 27603]) Current status: Synonym of Lycodes terraenovae Collett 1896. Zoarcidae: Lycodinae. Habitat: marine.

caudimaculatus, Lycodes Matsubara [K.] 1936:115, Fig. 1 [Journal of the Imperial Fisheries Institute Tokyo v. 31 (no. 2); ref. 13718] Off Owase, Mie Prefecture, Japan, depth 80-160 fathoms. Holotype: not at FAKU. •Valid as Lycodes caudimaculatus Matsubara 1936 -- (Lindberg & Krasyukova 1975:144 [ref. 7348], Yamakawa in Okamura et al. 1982:243, 385 [ref. 8256], Toyoshima in Amaoka et al. 1983:139, 209 [ref. 8379], Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:220 [ref. 5722], Anderson 1994:118 [ref. 21438], Shinohara et al. 1996:182 [ref. 26605], Nakabo 2000:1031 [ref. 25182], Shinohara et al. 2001:328 [ref. 25995], Nakabo 2002:1031 [ref. 26193], Anderson & Fedorov 2004:24 [ref. 27603], Anderson & Fedorov 2004:24 [ref. 27603], Balushkin et al. 2011:992 [ref. 32449], Shinohara et al. 2014:265 [ref. 33330]). Current status: Valid as Lycodes caudimaculatus Matsubara 1936. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Japan. Habitat: marine.

celatus, Lycodes Jensen [A. S.] 1902:208 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Kara-Havet, depth 59 fathoms. Syntypes: ZMUC. Type catalog: Nielsen 1974:76 [ref. 9588]. •Synonym of Lycodes rossi Malmgren 1865 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603], Balushkin et al. 2011:1010 [ref. 32449], Parin et al. 2014:420 [ref. 33547]). Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae. Habitat: marine.

coccineus, Lycodes Bean [T. H.] 1881:144 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Big Diomede Island [Chukotsky District, Chukotka Autonomous Okrug, Russia], Bering Strait. Holotype (unique): USNM 27748. Type catalog: Springer & Anderson 1997:16 [ref. 22953]. •Questionably a synonym of Lycodes mucosus Richardson 1855 (Balushkin et al. 2011:1002 [ref. 32449]). •Synonym of Lycodes mucosus Richardson 1855 -- (Toyoshima 1985:200 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:708 [ref. 25968], Anderson & Fedorov 2004:26 [ref. 27603], Parin et al. 2014:417 [ref. 33547]). Current status: Synonym of Lycodes mucosus Richardson 1855. Zoarcidae: Lycodinae. Habitat: marine.

colletti, Lycodes Popov [A. M.] 1931:143, Pl. 2 (fig. 6) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] Okhotsk Sea, 59°11.5'N, 148°19.5'E, depth 141 meters, 57°40'N, 147°50.6'E, depth 144 meters. Syntypes: ZIN 23758 (1), 29996 (1). Type catalog: Balushkin et al. 2011:1015 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667], •Valid as Lycodes colletti Popov 1931 -- (Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:113 [ref. 27579], Anderson & Fedorov 2004:30 [ref. 27603]), •Synonym of Lycodes uschakovi Popov 1931 -- (Lindberg & Krasyukova 1975:147 [ref. 7348], Anderson 1994:119 [ref. 21438], Balushkin et al. 2011:1015 [ref. 32449], Parin et al. 2014:423 [ref. 33547]). Current status: Synonym of Lycodes uschakovi Popov 1931. Zoarcidae: Lycodinae. Habitat: marine.

concolor, Lycodes Gill [T. N.] & Townsend [C. H.] 1897:233 [Proceedings of the Biological Society of Washington v. 11; ref. 1749] Bering Sea, 55°19'N, 168°11'W, Albatross station 3608, depth 276 fathoms. Holotype (unique): USNM 48764. Type catalog: Springer & Anderson 1997:16 [ref. 22953]. •Valid as Lycodes concolor Gill & Townsend 1897 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:725 [ref. 25968], Møller 2001:975 [ref. 25812], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Love et al. 2005:142 [ref. 37547], Balushkin et al. 2011:991 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:409 [ref. 33547], Kells et al. 2016:272 [ref. 35888], Dyldin & Orlov 2021:837 [ref. 38925], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes concolor Gill & Townsend 1897. Zoarcidae: Lycodinae. Distribution: North Pacific: Sea of Okhotsk and northern Kuril Islands north to western Bering Sea, straying to eastern Chukchi Sea; eastern Bering Sea from Aleutian Islands to western Gulf of Alaska (U.S.A.). Habitat: marine.

corteziana, Aprodon Gilbert [C. H.] 1890:107 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Cortez Banks off San Diego, California, U.S.A., 33°53'30"N, 119°41'30"W, Albatross station 2948, depth 266 fathoms. Lectotype: USNM 46457. Paralectotypes: (5) CAS-SU 2780 (1); USNM 47056-57 (1, 1), 47180 (1), 47198 (1). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:16 [ref. 22953]. Lectotype established by Jordan & Evermann 1898:2461 [ref. 2445] and as figured specimen in caption to Pl. 349, p. 3304 in Jordan & Evermann 1900 [ref. 2446]. •Valid as Aprodon cortezianus Gilbert 1890 -- (Brock 1943:186 [ref. 41494], Toyoshima 1985:237 [ref. 5722], Toyoshima 1985:237 [ref. 5722], Balushkin et al. 2011:973 [ref. 32499]). •Valid as Lycodes cortezianus (Gilbert 1890) -- (Eschmeyer & Herald 1983:105 [ref. 9277], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:705 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Love et al. 2005:143 [ref. 37547], Maslenikov et al. 2013:17 [ref. 37947], Page et al. 2013:162 [ref. 32708], Pietsch & Orr 2015:58 [ref. 34694], Kells et al. 2016:272 [ref. 35888], Burton & Lea 2019:84 [ref. 37205], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes cortezianus (Gilbert 1890). Zoarcidae: Lycodinae. Distribution: Eastern Pacific: southeastern Alaska (U.S.A.) south to southern California (U.S.A.). Habitat: marine.

diapteroides, Lycodes brevipes Taranetz [A. Ya.] & Andriashev [A. P.] in Andriashev 1937:253 [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] Bering Sea and Bering Sea at 62°08'N, 178°32'W, depth 98 meters. Syntypes: ZIN 26886 (1), 30011 (3). Type catalog: Balushkin et al. 2011:991 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. •Valid as Lycodes diapteroides Taranetz & Andriashev 1937 -- (Toyoshima 1985:191 [ref. 5722]). •Synonym of Lycodes brevipes Bean 1890 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650] with question, Mecklenburg et al. 2002:716 [ref. 25968], Anderson & Fedorov 2004:23 [ref. 27603], Balushkin et al. 2011:991 [ref. 32449], Parin et al. 2014:409 [ref. 33547]). Current status: Synonym of Lycodes brevipes Bean 1890. Zoarcidae: Lycodinae. Habitat: marine.

diapterus, Lycodes Gilbert [C. H.] 1892:564 [Proceedings of the United States National Museum v. 14 (no. 880); ref. 1626] Off California, 33°55'30"N, 120°28'00"W, Albatross station 2896, depth about 688 meters. Lectotype: USNM 44385. Paralectotypes: BMNH 1900.9.29.245 (1) from Jordan; CAS-SU 176 (2); USNM 46716 (1), 125538 (1). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:17 [ref. 22953]. Lectotype established by Jordan & Evermann 1898:2473 [ref. 2445]. •Valid as Lycodes diapterus Gilbert 1892 -- (Lindberg & Krasyukova 1975:150 [ref. 7348], Eschmeyer & Herald 1983:105 [ref. 9277], Toyoshima 1985:230 [ref. 5722], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Mecklenburg et al. 2002:724 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:157 [ref. 27807], Love et al. 2005:143 [ref. 37547], Shinohara & Shirai 2005:172 [ref. 28232], Stevenson & Sheiko 2009:127 [ref. 30070], Balushkin et al. 2011:993 [ref. 32449], Pietsch & Orr 2015:72 [ref. 34694], Kells et al. 2016:272 [ref. 35888], Burton & Lea 2019:84 [ref. 37205], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes diapterus Gilbert 1892. Zoarcidae: Lycodinae. Distribution: North Pacific: Sea of Japan and Sea of Okhotsk north to northern Bering Sea, then Aleutian Islands (U.S.A.) south to southern California (U.S.A.). Habitat: marine.

digitatus, Lycodes Gill [T. N.] & Townsend [C. H.] 1897:232 [Proceedings of the Biological Society of Washington v. 11; ref. 1749] Bering Sea, 56°14'N, 164°09'W, Albatross station 3541, depth 49 fathoms. Holotype (unique): USNM 48765. Type catalog: Springer & Anderson 1997:17 [ref. 22953]. •Synonym of Lycodes palearis Gilbert 1896 -- (Toyoshima 1985:212 [ref. 5722], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:719 [ref. 25968], Anderson & Fedorov 2004:27 [ref. 27603], Parin et al. 2014:417 [ref. 33547]). Current status: Synonym of Lycodes palearis Gilbert 1896. Zoarcidae: Lycodinae. Habitat: marine.

esmarkii, Lycodes Collett [R.] 1875:95 [Norges fiske; ref. 884] Varanger Fjord, northern Norway. Syntypes: BMNH 1890.11.1.1 (1) from Collett; MNHN 1890-0369 [ex ZMUO] (1); ZMUO J4496 (1), J4498 (1). Type catalog: Pethon 1969:11 [ref. 19268]. •Valid as Lycodes esmarkii Collett 1875 -- (Andriashev 1973:543 [ref. 9214] as esmarki, Andriashev in Whitehead et al. 1986:1138 [ref. 13677] as esmarki, Scott & Scott 1988:407 [ref. 25518] as esmarki, McAllister 1990:159 [ref. 14674] as esmarki, Anderson 1994:118 [ref. 21438], Saito & Okamura in Okamura et al. 1995:194 [ref. 22531], Møller 2000:847 [ref. 24585], Moore et al. 2003:228 [ref. 27076], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Møller et al. 2010:60 [ref. 31169], Balushkin et al. 2011:993 [ref. 32449], Page et al. 2013:162 [ref. 32708], Dolgov 2013:920 [ref. 33112], Parin et al. 2014:410 [ref. 33547], Afonso et al. 2016:63, 196 [ref. 36199], Dolgov et al. 2018:203 [ref. 41055], Mecklenburg et al. 2018:322 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Simian et al. 2022:376 [ref. 40776], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes esmarkii Collett 1875. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

eudipleurostictus, Lycodes Jensen [A. S.] 1902:206 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Faroe Islands to Spitsbergen, northern Atlantic, depth 350-471 fathoms. Syntypes: NRM 25527 (1); ZMUC 7, 138; ZMUO (4). Type catalog: Nielsen 1974:77 [ref. 9588]. •Valid as Lycodes eudipleurostictus Jensen 1902 -- (Andriashev 1973:543 [ref. 9214], McAllister et al. 1981:830 [ref. 5431], Andriashev in Whitehead et al. 1986:1138 [ref. 13677], Anderson 1994:118 [ref. 21438], Saito & Okamura in Okamura et al. 1995:195 [ref. 22531], Mecklenburg et al. 2002:723 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603], Love et al. 2005:143 [ref. 37547], Shinohara & Shirai 2005:171 [ref. 28232], Møller et al. 2010:60 [ref. 31169], Balushkin et al. 2011:993 [ref. 32449], Page et al. 2013:162 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:410 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:63, 197 [ref. 36199], Mecklenburg et al. 2016:162 [ref. 34440], Dolgov et al. 2018:205 [ref. 41055], Mecklenburg et al. 2018:324 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes eudipleurostictus Jensen 1902. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

fasciatus, Lycenchelys Schmidt [P. Yu.] 1904:203, Pl. 6 (fig. 2) [Fishes of the eastern seas of the Russian Empire; ref. 3946] Aniva Bay, southern Sakhalin Island, Russia, Okhotsk Sea, depth 126-161 feet (17-22 Russian fathoms). Syntypes: ZIN 12446 (1, lost), 13092-93 (1, 1). Type catalog: Balushkin et al. 2011:993 [ref. 32449], Balushkin et al. 2012:159 [ref. 33667]. Appeared first without distinguishing features in Schmidt 1903:520 (18 of separate) [ref. 3945]. •Synonym of Lycodes palearis Gilbert 1896, but a valid subspecies -- (Lindberg & Krasyukova 1975:157 [ref. 7348], Sokolovskaya et al. 1998:12 [ref. 24670], Shinohara et al. 2011:51 [ref. 31715], Shinohara et al. 2014:266 [ref. 33330]). •Valid as Lycodes fasciatus (Schmidt 1904) -- (Toyoshima 1985:214 [ref. 5722], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:720 [ref. 25968], Sheiko & Fedorov 2000:36 [ref. 25650], Møller 2001:111 [ref. 25378], Fedorov et al. 2003:113 [ref. 27579], Anderson & Fedorov 2004:24 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Balushkin et al. 2011:993 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:410 [ref. 33547], Mecklenburg et al. 2016:173 [ref. 34440], Dyldin & Orlov 2021:838 [ref. 38925], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes fasciatus (Schmidt 1904). Zoarcidae: Lycodinae. Distribution: Northwestern Pacific. Habitat: marine.

frigidus, Lycodes Collett [R.] 1879:45 [Forhandlinger i Videnskabs-selskabet i Christiania (for 1878) No. 14; ref. 885] Bear Island and Spitsbergen, Arctic Ocean, depth 475-2438 meters. Syntypes: BMNH 1885.12.2.1 (1) from Collett; MNHN 1890-0370 (1); NMW 77810 (1); USNM 22977 (1), ZMUB 1688 (1), 2223 (1); ZMUC 24 (1); ZMUO J4517-22 (6), plus 1 destroyed and some not traced. Type catalog: Pethon 1969:11 [ref. 19268], Nielsen 1974:77 [ref. 9588], Frøiland 1979:150 [ref. 21547], Springer & Anderson 1997:18 [ref. 22953]. •Valid as Lycodes frigidus Collett 1879 -- (Andriashev 1973:543 [ref. 9214], McAllister et al. 1981:829 [ref. 5431], Andriashev in Whitehead et al. 1986:1139 [ref. 13677], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Saito & Okamura in Okamura et al. 1995:196 [ref. 22531], Mecklenburg et al. 2002:726 [ref. 25968], Anderson & Fedorov 2004:25 [ref. 27603], Møller et al. 2010:60 [ref. 31169], Balushkin et al. 2011:995 [ref. 32449], Mecklenburg et al. 2014:144 ,145 [ref. 33463], Parin et al. 2014:411 [ref. 33547], Parin et al. 2014:411 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:63, 198 [ref. 36199], Mecklenburg et al. 2016:164 [ref. 34440], Dolgov et al. 2018:207 [ref. 41055], Mecklenburg et al. 2018:326 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:183 [ref. 39279]). Current status: Valid as Lycodes frigidus Collett 1879. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

fulvus, Lycodes Toyoshima [M.] 1985:216, Fig. 52 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Okhotsk Sea, 55°19'N, 142°34'E, depth 178 meters. Holotype: HUMZ 60246. Paratypes: HUMZ 58291-93 (3), 60244 (1), 60248-52 (5), 60254-58 (5), 60278 (1), 61434 (1), 61466 (1). •Valid as Lycodes fulvus Toyoshima 1985 -- (Anderson 1994:118 [ref. 21438], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:114 [ref. 27579], Anderson & Fedorov 2004:25 [ref. 27603], Balushkin et al. 2012:995 [ref. 32449], Parin et al. 2014:411 [ref. 33547], Dyldin & Orlov 2021:838 [ref. 38925]). •Synonym of Lycodes ygreknotatus Schmidt 1950 -- (Nazarkin et al. 2024:119 [ref. 41547]). Current status: Synonym of Lycodes ygreknotatus Schmidtb 1950. Zoarcidae: Lycodinae. Habitat: marine.

gracilis, Lycodes Sars [M.] 1867:40, 45, Pl. 1 (figs. 1-3) [Forhandlinger i Videnskabs-selskabet i Christiania (for 1866); ref. 18657] Drøbaksundet, Oslo Fjord, Norway, depth 94-113 meters. Holotype: ZMUO J4525. Type catalog: Pethon 1969:11 [ref. 19268]. •Synonym of Lycodes vahlii Reinhardt 1831, but a valid subspecies -- (Andriashev 1973:542 [ref. 9214] as vahli, Andriashev in Whitehead et al. 1986:1146 [ref. 13677] as vahli, Saito & Okamura in Okamura et al. 1995:201 [ref. 22531]). •Synonym of Lycodes vahlii Reinhardt 1831 -- (Anderson 1994:119 [ref. 21438]). •Valid as Lycodes gracilis Sars 1867 -- (Carl 2002:75 [ref. 26925], Anderson & Fedorov 2004:25 [ref. 27603], Møller et al. 2010:61 [ref. 31169], Mecklenburg 2011:124 [ref. 31212] as vahli, Balushkin et al. 2011:995 [ref. 32449], Page et al. 2013:162 [ref. 32708], Parin et al. 2014:411 [ref. 33547], Mecklenburg et al. 2018:328 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes gracilis Sars 1867. Zoarcidae: Lycodinae. Distribution: Arctic and North Atlantic. Habitat: marine.

hacheyi, Lycodes reticulatus Vladykov [V. D.] 1933:25 [Contributions to Canadian biology and fisheries: being studies from the biological stations of Canada (Ser. A, General) v. 8 (no. 2); ref. 23244] Hudson Bay, Canada. Syntypes: (2) whereabouts unknown. Also in Vladykov & Tremblay 1936:37, Pl. 6 (fig. 13) [ref. 11811]. •Synonym of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Habitat: marine.

heinemanni, Lycodes Soldatov [V. K.] 1916:215, Fig. (p. 215) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 21; ref. 15428] Northern Okhotsk Sea, 58°17'N, 143°15'E, depth 60 meters. Syntypes: ZIN 19169 (3). Type catalog: Balushkin et al. 2011:997 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. •Valid as Lycodes heinemanni Soldatov 1916 -- (Toyoshima 1985:202 [ref. 5722], Anderson 1994:118 [ref. 21438], Fedorov et al. 2003:114 [ref. 27579] dated 1917, Anderson & Fedorov 2004:25 [ref. 27603], Balushkin et al. 2011:997 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:412 [ref. 33547], Dyldin & Orlov 2021:838 [ref. 38925]). Current status: Valid as Lycodes heinemanni Soldatov 1916. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: northern Okhotsk Sea (Russia). Habitat: marine.

hubbsi, Lycodes Matsubara [K.] 1955:776 [Fish morphology and hierarchy Part 1; ref. 18463] Off Kinkazan and off Kushiro, Japan. No types known. •Status uncertain -- (Anderson 1994:118 [ref. 21438]). •Valid as Lycodes hubbsi Matsubara 1955 -- (Toyoshima in Amaoka et al. 1983:139, 208, 261, 330 [ref. 8379], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:226 [ref. 5722], Nakabo 2000:1029 [ref. 25182], Sheiko & Fedorov 2000:36 [ref. 25650], Nakabo 2002:1029 [ref. 26193], Anderson & Fedorov 2004:25 [ref. 27603], Shinohara & Shirai 2005:171 [ref. 28232], Balushkin et al. 2011:997 [ref. 32449], Parin et al. 2014:412 [ref. 33547], Dyldin & Orlov 2021:838 [ref. 38925]). Current status: Valid as Lycodes hubbsi Matsubara 1955. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific. Habitat: marine.

intermedia, Lycodes rossi forma Knipowitsch [N. M.] 1906:28 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Not available, infrasubspecific; publication in an available way as a species or subspecies not investigated [Knipowitsch used v. [variety] and f. [forma] -- in his work, the former is considered subspecific the latter infrasubspecific; see Article 45(g)]. From Barents Sea, Russia. •In the synonymy of Lycodes rossii Malmgren 1865 -- (Anderson 1994:119 [ref. 21438] as rossi, Anderson & Fedorov 2004:48 [ref. 27603], Parin et al. 2014:420 [ref. 33547]). Infrasubspecific. Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae.

japonicus, Lycodes Matsubara [K.] & Iwai [T.] 1951:368, Figs. 1-3 [Japanese Journal of Ichthyology v. 1 (no. 6); ref. 12785] Near the sea-coast of Uozu, southeast of Toyama Bay, Japan, depth 303 meters. Holotype: FAKU 13117 (not seen in 1997). Paratypes: (not seen in 1997) FAKU 12773-87 (15), 13115-16 (2), 13118-38 (21), 13150 (1). •Valid as Lycodes japonicus Matsubara & Iwai 1951 -- (Lindberg & Krasyukova 1975:143 [ref. 7348], Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:191 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1037 [ref. 25182], Mecklenburg et al. 2002:714 [ref. 25968], Nakabo 2002:1037 [ref. 26193], Anderson & Fedorov 2004:25 [ref. 27603], Love et al. 2005:143 [ref. 37547], Kim et al. 2006:274 [ref. 29191], Nazakin & Fedorov 2007:705 [ref. 29424], Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:998 [ref. 32449], Shinohara et al. 2014:265 [ref. 33330], Parin et al. 2014:412 [ref. 33547], Kim et al. 2020:155 [ref. 39118], Sonoyama et al. 2020:94 [ref. 37637], Dyldin & Orlov 2021:838 [ref. 38925], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes japonicus Matsubara & Iwai 1951. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia) north to Kuril Islands and Sakhalin (Russia); straying to eastern Bering Sea (Alaska, U.S.A.). Habitat: marine.

jenseni, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] 1935:243, Figs. 1-2 [Zoologischer Anzeiger v. 112 (nos 9-10); ref. 15744] Okhotsk Sea, east of Sakhalin Island, 50°03'N, 144°08'E. Holotype: ZIN 24836. Paratypes: ZIN 36937 (1). Type catalog: Balushkin et al. 2011:997 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. •Valid as Lycodes jenseni Taranetz & Andriashev 1935 -- (Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:211 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1035 [ref. 25182], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:36 [ref. 25650], Nakabo 2002:1035 [ref. 26193], Anderson & Fedorov 2004:25 [ref. 27603], Nazarkin 2010:9 [ref. 30733], Balushkin et al. 2011:998 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:413 [ref. 33547], Dyldin & Orlov 2021:838 [ref. 38925]). Current status: Valid as Lycodes jenseni Taranetz & Andriashev 1935. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific. Habitat: marine.

jugoricus, Lycodes Knipowitsch [N. M.] 1906:18, Figs. 4-5 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Proliv Yugorsky Shar, strait between Kara Sea and Barents Sea, Novaya Zemlya, Russia, depth 9 meters. Holotype (unique): ZIN 13506. Type catalog: Balushkin et al. 2011:997 [ref. 32449], Balushkin et al. 2012:160 [ref. 33667]. •Valid as Lycodes jugoricus Knipowitsch 1906 -- (Andriashev 1973:543 [ref. 9214], Andriashev in Whitehead et al. 1986:1139 [ref. 13677], McAllister 1990:159 [ref. 14674], Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:706 [ref. 25968], Anderson & Fedorov 2004:25 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Love et al. 2005:143 [ref. 37547], Balushkin et al. 2011:998 [ref. 32449], Page et al. 2013:162 [ref. 32708], Dolgov 2013:920 [ref. 33112], Parin et al. 2014:413 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:63, 199 [ref. 36199], Mecklenburg et al. 2016:166 [ref. 34440], Dolgov et al. 2018:209 [ref. 41055], Mecklenburg et al. 2018:330 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes jugoricus Knipowitsch 1906. Zoarcidae: Lycodinae. Distribution: Arctic Ocean. Habitat: marine.

knipowitschi, Lycodes Popov [A. M.] 1931:138, Pl. 2 (fig. 4); Fig. 1 [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] Okhotsk Sea, 59°31.2'N, 150°29.9'E, depth 40 meters. Holotype: ZIN 30009. Paratypes: ? ZIN 30009 (2 juv.). Type catalog: Balushkin et al. 2011:999 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. •Synonym of Lycodes mucosus Richardson 1855 -- (Anderson 1994:118 [ref. 21438], Mecklenburg et al. 2002:708 [ref. 25968]). •In part synonym of Lycodes raridens Taranetz & Andriashev 1937 -- (Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:711 [ref. 25968]). •Valid as Lycodes knipowitschi Popov 1931 -- (Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:195 [ref. 5722], Nakabo 2000:1038 [ref. 25182], Sheiko & Fedorov 2000:36 [ref. 25650], Nakabo 2002:1038 [ref. 26193], Fedorov et al. 2003:114 [ref. 27579], Anderson & Fedorov 2004:27 [ref. 27603], Balushkin et al. 2011:999 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:413 [ref. 33547]). Current status: Valid as Lycodes knipowitschi Popov 1931. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea. Habitat: marine.

laurentianus, Lycodes reticulatus Vladykov [V. D.] & Tremblay [J.-L.] 1936:34, Pls. 5 (fig. 11), 6 (fig. 12) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Saint Lawrence River estuary, Quebec, Canada. Syntypes: (8) NMC 60-159 (1). Type catalog: McAllister 1965:9 [ref. 22019] as paratypes. •Synonym of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Habitat: marine.

lavalaei, Lycodes Vladykov [V. D.] & Tremblay [J.-L.] 1936:31, Pls. 4 (fig. 9), 5 (figs. 10-11) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Gulf of Saint Lawrence and Newfoundland, Canada. Syntypes: (11) NMC 60-157 (1), 60-158 (1). Type catalog: McAllister 1965:9 [ref. 22019] as paratypes. Misspelled lavalei in earlier editions of the Catalog of Fishes. •Valid as Lycodes lavalaei Vladykov & Tremblay 1936 -- (Andriashev 1973:544 [ref. 9214], Andriashev in Whitehead et al. 1986:1140 [ref. 13677], Robins & Ray 1986:237 [ref. 23100], Scott & Scott 1988:408 [ref. 25518], McAllister 1990:161 [ref. 14674], Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:26 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Mecklenburg et al. 2011:128 [ref. 31212], Balushkin et al. 2011:999 [ref. 32449], Page et al. 2013:162 [ref. 32708], Afonso et al. 2016:64, 200 [ref. 36199], Mecklenburg et al. 2018:331 [ref. 36370], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes lavalaei Vladykov & Tremblay 1936. Zoarcidae: Lycodinae. Distribution: Northwestern Atlantic and adjacent Arctic Region. Habitat: marine.

lindbergi, Lycodes Popov [A. M.] 1931:142, Pl. 2 (fig. 5) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] 58°11.5'N, 148°19.5'N, Okhotsk Sea, depth 141 meters. Holotype (unique): whereabouts unknown. •Synonym of Lycodes semenovi Popov 1931 -- (Balushkin et al. 2011:1011 [ref. 32449]). •Synonym of Lycodes uschakovi Popov 1931 -- (Lindberg & Krasyukova 1975:147 [ref. 7348], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:30 [ref. 27603], Parin et al. 2014:423 [ref. 33547]). Current status: Synonym of Lycodes uschakovi Popov 1931. Zoarcidae: Lycodinae. Habitat: marine.

luetkenii, Lycodes Collett [R.] 1880:103, Pl. 3 (fig. 25) [The Norwegian North-Atlantic Expedition; ref. 887] 115 kilometers west of Norskøerne, Spitsbergen, North Atlantic, depth 459 fathoms. Holotype (unique): ZMUO J4536. Type catalog: Pethon 1969:12 [ref. 19268]. Original as lütkenii. •Valid as Lycodes luetkenii Collett 1880 -- (Andriashev 1973:544 [ref. 9214] as leutkeni, Andriashev in Whitehead et al. 1986:1140 [ref. 13677] as leutkeni, Anderson 1994:118 [ref. 21438], Saito & Okamura in Okamura et al. 1995:197 [ref. 22531] as luetkeni, Møller & Petersen 1997:290 [ref. 23332] as luetkeni, Anderson & Fedorov 2004:26 [ref. 27603], Møller et al. 2010:61 [ref. 31169], Balushkin et al. 2011:999 [ref. 32449], Page et al. 2013:162 [ref. 32708], Dolgov 2013:920 [ref. 33112] as leutkeni, Parin et al. 2014:413 [ref. 33547], Afonso et al. 2016:64, 201 [ref. 36199], Dolgov et al. 2018:211 [ref. 41055], Mecklenburg et al. 2018:333 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes luetkenii Collett 1880. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic. Habitat: marine.

lugubris, Lycodes Lütken [C. F.] 1880:315 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1879-1880; ref. 19479] Iceland. Syntypes: ZMUC 10 (1), 11 (skeleton), 13 (1), 14 (1). Type catalog: Nielsen 1974:77 [ref. 9588]. •Synonym of Lycodes vahlii Reinhardt 1831 -- (Andriashev 1973:542 [ref. 9214], Anderson 1994:119 [ref. 21438]). •Synonym of Lycodes gracilis Sars 1867 -- (Carl 2002:75 [ref. 26925], Anderson & Fedorov 2004:25 [ref. 27603]). Current status: Synonym of Lycodes gracilis Sars 1867. Zoarcidae: Lycodinae. Habitat: marine.

macrocephalus, Lycodes reticulatus var. Jensen [A. S.] 1904:258, Pl. 13 (figs. 2 a, b) [Meddelelser om Grønland v. 29; ref. 20203] Eastern Greenland. Syntypes: (11) whereabouts unknown. •Synonym of Lycodes reticulatus Reinhardt 1835, but a valid subspecies as described -- (Andriashev 1973:545 [ref. 9214], Andriashev in Whitehead et al. 1986:1143 [ref. 13677]). •Synonym of Lycodes reticulatus Reinhardt 1835 -- (Anderson & Fedorov 2004:28 [ref. 27603], Balushkin et al. 2011:1009 [ref. 32449]), •In the synonymy of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Parin et al. 2014:419 [ref. 33547]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Habitat: marine.

macrochir, Lycodes Schmidt [P. Yu.] in Taranetz 1937:162, 164 [Izvestiia Tikhookeanskogo Nauchno-Issledovatel'skogo Instituta Rybnogo Khoziaistva i Okeanografii [Proceedings, Pacific Research Institute of Fisheries and Oceanography, TINRO, Moskva] v. 11; ref. 13384] Okhotsk Sea. Syntypes: ZIN 25273 (1), 29092 (1), 29142 (1). Type catalog: Balushkin et al. 2011:999 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667].. •Valid as Lycodes macrochir Schmidt 1937 -- (Toyoshima 1985:205 [ref. 5722], Anderson 1994:118 [ref. 21438], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:36 [ref. 25650], Fedorov et al. 2003:114 [ref. 27579], Anderson & Fedorov 2004:26 [ref. 27603], Shinohara & Shirai 2005:171 [ref. 28232], Balushkin et al. 2011:999 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Parin et al. 2014:414 [ref. 33547], Dyldin & Orlov 2021:838 [ref. 38925]). Current status: Valid as Lycodes macrochir Schmidt 1937. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk Sea (Russia). Habitat: marine.

macrolepis, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] 1935:251, Figs. 6-7 [Zoologischer Anzeiger v. 112 (nos 9-10); ref. 15744] Okhotsk Sea, northwest of St. Jonas Island, 57°02'N, 141°40'E, depth 147 meters. Holotype: ZIN 24837. Paratypes: ZIN 24837 (2 of 3), 29998 (1). Type catalog: Balushkin et al. 2011:1000 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. •Valid as Lycodes macrolepis Taranetz & Andriashev 1935 -- (Lindberg & Krasyukova 1975:153 [ref. 7348], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:204 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Nakabo 2002:1039 [ref. 26193], Fedorov et al. 2003:115 [ref. 27579], Anderson & Fedorov 2004:26 [ref. 27603], Nazarkin 2010:596 [ref. 31019], Balushkin et al. 2011:1000 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Shinohara et al. 2014:265 [ref. 33330], Dyldin & Orlov 2021:838 [ref. 38925]). Current status: Valid as Lycodes macrolepis Taranetz & Andriashev 1935. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk and Japan seas. Habitat: marine.

maculatus, Lycodes vahlii Vladykov [V. D.] & Tremblay [J.-L.] 1936:27, Pl. 3 (figs. 7-8) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Saint Lawrence River estuary, Quebec, Canada. Syntypes: (219) NMC 60-161 (1), 60-162 (orig. 23, now 21); ROM (1); UBC (1). Type catalog: McAllister 1965:10 [ref. 22019] as paratypes. •Synonym of Lycodes vahlii Reinhardt 1831 -- (Anderson 1994:119 [ref. 21438], Carl 2002:69 [ref. 26925] as vahli, Anderson & Fedorov 2004:31 [ref. 27603], Balushkin et al. 2011:1016 [ref. 32449]). Current status: Synonym of Lycodes vahlii Reinhardt 1831. Zoarcidae: Lycodinae. Habitat: marine.

marginalis, Lycodes brevipes Taranetz [A. Ya.] & Andriashev [A. P.] in Taranetz 1937:163 [Izvestiia Tikhookeanskogo Nauchno-Issledovatel'skogo Instituta Rybnogo Khoziaistva i Okeanografii [Proceedings, Pacific Research Institute of Fisheries and Oceanography, TINRO, Moskva] v. 11; ref. 13384] Not available, no distinguishing features. Apparently misprint for Lycodes breviceps diapteroides. •Nomen nudum, in the synonymy of Lycodes brevipes Bean 1890 -- (Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Anderson & Fedorov 2004:48 [ref. 27603], Parin et al. 2014:408 [ref. 33547]). Nomen Nudum. Current status: Synonym of Lycodes brevipes Bean 1890. Zoarcidae: Lycodinae.

marisalbi, Lycodes Knipowitsch [N. M.] 1906:48, Figs. 9-12 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] White Sea, Russia, 65°13'N, 39°07'E, depth 91 meters. Lectotype: ZIN 40511. Paralectotypes: ZIN 13500-502 (4, 3, 3). Type catalog: Balushkin et al. 2011:1000 [ref. 32449], Balushkin et al. 2012:161 [ref. 33667]. Originally as maris-albi. Lectotype selected by Andriashev 1973:545 [ref. 7214]. •Synonym of Lycodes pallidus Collett 1879, but a valid subspecies -- (Andriashev 1973:545 [ref. 9214], Andriashev in Whitehead et al. 1986:1142 [ref. 13677]). •Synonym of Lycodes pallidus Collett 1879 -- (Anderson 1994:119 [ref. 21438]). •Valid as Lycodes marisalbi Knipowitsch 1906 -- (Møller 2000:1406 [ref. 40921], Møller 2001:111 [ref. 25378], Anderson & Fedorov 2004:26 [ref. 27603], Balushkin et al. 2011:1000 [ref. 32449], Page et al. 2013:163 [ref. 32708], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:414 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:64, 202 [ref. 36199], Mecklenburg et al. 2016:168 [ref. 34440] see remarks, Mecklenburg et al. 2018:335 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes marisalbi Knipowitsch 1906. Zoarcidae: Lycodinae. Distribution: Arctic Ocean: White Sea (Russia, Norway); Canada, Alaska (U.S.A.). Habitat: marine.

matsubarai, Lycodes Toyoshima [M.] 1985:218, Fig. 53 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Kitami-yamato Bank, 44°47'N, 144°01.5'E, southern Okhotsk Sea. depth 200 meters. Holotype: HUMZ 33970. Paratypes: HUMZ 33974 (1), 33948 (1), 33971-72 (2), 33974-79 (6), 33981 (1), 33983 (1), 33950 (2), 33960 (1), 33968-69 (2), 40945-46 (2). Lycodes Matsubarai and L. yamatoi were simutaneouly described in ref. 5722; Nazarkin & Shinohara 2012:34 [ref. 32106] act as first revisers selecting matsubarai over yamatoi. •Synonym of Lycodes microlepidotis Schmidt 1950 -- (Balushkin et al. 2011:1000 [ref. 24994], Nazarkin et al. 2014:192 [ref. 33213]). •Valid as Lycodes matsubarai Toyoshima 1985 -- (Anderson 1994:118 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:36 [ref. 25650], Nakabo 2002:1039 [ref. 26193], Anderson & Fedorov 2004:26 [ref. 27603], Nazarkin 2010:9 [ref. 30733], Nazarkin & Shinohara 2012:34 [ref. 32106], Shinohara et al. 2014:265 [ref. 33330], Parin et al. 2014:414 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331], Sonoyama et al. 2020:94 [ref. 37637], Dyldin & Orlov 2021:839 [ref. 38925]). Current status: Valid as Lycodes matsubarai Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk Sea and Sea of Japan to Peter the Great Bay (Russia); Yamato Bank (Japan and Russia). Habitat: marine.

mcallisteri, Lycodes Møller [P. R.] 2001:112, Figs. 2-4 [Ichthyological Research v. 48 (no. 2); ref. 25378] Western Baffin Bay, 70°02'N, 65°32'W, eastern Arctic Canada, depth 579 meters. Holotype: ZMUC P764746. Paratypes: CMNFI 2000-0055(1); HUMZ 172115 (1); NMC 90-0223 (1); USNM 362128 (1); ZIN 52125 (1); ZMUC P765161 (1), P765163 (1), P765168 (1). •Valid as Lycodes mcallisteri Møller 2001 -- (Anderson & Fedorov 2004:26 [ref. 27603], Møller et al. 2010:61 [ref. 31169], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin et al. 2011:1000 [ref. 32449], Afonso et al. 2016:65, 203 [ref. 36199], Mecklenburg et al. 2018:337 [ref. 36370], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes mcallisteri Møller 2001. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and Labrador (Canada). Habitat: marine.

megalocephala, Lycodes rossi forma Knipowitsch [N. M.] 1906:27 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Syntypes: ZIM 13406 (1), 13407 (5), 13411 (3), 13412 (2), 13420 (1), 13421 (3), 13422 (3), 13423 (1), 13434 (2). Type cataqlog: Balushkin et al. 2011:1010 [ref. 32449], Balushkin et al. 2012:164 [ref. 33667]. Not available, infrasubspecific; publication in an available way as a species or subspecies not investigated [Knipowitsch used v. [variety] and f. [forma] -- in his work, the former is considered subspecific the latter infrasubspecific; see Article 45(g)]. From Barents Sea, Russia. •Infrasubspecific, in the synonymy of Lycodes rossii Malmgren 1865 -- (Anderson 1994:119 [ref. 21438] as rossi, Anderson & Fedorov 2004:48 [ref. 27603], Balushkin et al. 2011:1010 [ref. 32449], Parin et al. 2014:420 [ref. 33547]). Infrasubspecific. Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae.

microlepidotus, Lycodes Schmidt [P. Yu.] 1950:91 [78], Pl. 6 (fig. 1) [Trudy Tichookeanskogo Komiteta, Akademija Nauk S.S.S.R. v. 6; ref. 12471] East coast of Sakhalin and west coast of Kamchatka, Russia, Okhotsk Sea. Syntypes: ZIN 24827 (4), 24844 (1), 25268 (1), Type catalog: Balushkin et al. 2011:1001 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. On p. 78 of English translation. •Valid as Lycodes microlepidotus Schmidt 1950 -- (Toyoshima 1985:210 [ref. 5722], Anderson 1994:118 [ref. 21438], Møller 2001:111 [ref. 25378], Sheiko & Fedorov 2000:37 [ref. 25650], Anderson & Fedorov 2004:26 [ref. 27603], Shinohara & Shirai 2005:171 [ref. 28232], Nazarkin 2010:4 [ref. 30733], Balushkin et al. 2011:1001 [ref. 32449], Parin et al. 2014:415 [ref. 33547], Dyldin & Orlov 2021:839 [ref. 38925]). •Synonym of Lycodes ygreknotatus Schmidt 1950 -- (Nazarkin et al. 2024:119 [ref. 41547]). Current status: Synonym of Lycodes ygreknotatus Schmidt 1950. Zoarcidae: Lycodinae. Habitat: marine.

microporus, Lycodes Toyoshima [M.] in Amaoka et al. 1983:263, Pl. 152: Figs. 23-24 [Fishes from the north-eastern Sea of Japan and the Okhotsk Sea off Hokkaido; ref. 8379] Abashiri, northeastern Hokkaidoo, Okhotsk Sea, 44°11'N, 144°47'E, depth 650-700 meters. Holotype: HUMZ 77795. Paratypes: HUMZ 77783-87 (5), 77807 (1), 78782 (1), 79662-63 (2). •Valid as Lycodes microporus Toyoshima 1983 -- (Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:221 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1034 [ref. 26193], Fedorov et al. 2003:115 [ref. 27579], Anderson & Fedorov 2004:26 [ref. 27603], Ikeda et al. 2008:357 [ref. 29896], Balushkin et al. 2011:1001 [ref. 32449], Parin et al. 2014:415 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331], Dyldin & Orlov 2021:839 [ref. 38925]). Current status: Valid as Lycodes microporus Toyoshima 1983. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk Sea (northern Japan, Russia). Habitat: marine.

mucosus, Lycodes Richardson [J.] 1855:362, Pl. 26 [The last of the Arctic voyages in search of Sir J. Franklin; ref. 18631] Northumberland Sound, Perry Island, Arctic Canada. Syntypes: (2) BMNH 1855.9.19.760. •Valid as Lycodes mucosus Richardson 1855 -- (Toyoshima 1985:200 [ref. 5722], McAllister 1990:161 [ref. 14674], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:708 [ref. 25968], Anderson & Fedorov 2004:26-27 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Love et al. 2005:143 [ref. 37547], Mecklenburg et al. 2007:181 [ref. 29448], Møller et al. 2010:61 [ref. 31169], Balushkin et al. 2011:1002 [ref. 32449], Page et al. 2013:163 [ref. 32708], Parin et al. 2014:416 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:65, 204 [ref. 36199], Kells et al. 2016:272 [ref. 35888], Mecklenburg et al. 2016:170 [ref. 34440] see remarks, Mecklenburg et al. 2018:338 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:183 [ref. 39279], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes mucosus Richardson 1855. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and adjacent North Pacific and North Atlantic. Habitat: marine.

multifasciatus, Lycodes palearis Schmidt [P. Yu.] 1950:95 [82], Pl. 7 (fig. 2) [Trudy Tichookeanskogo Komiteta, Akademija Nauk S.S.S.R. v. 6; ref. 12471] West coast of Kamchatka, Russia, opposite the mouths of the Ozernaya and Bol'shaya rivers, depth 97-250 meters. Syntypes: ZIN 29102 (1), 29102a (3). Type catalog: Balushkin et al. 2011:989 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. On p. 82 of English translation. •Valid as Lycodes multifasciatus Schmidt 1950 -- (Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:214 [ref. 5722], Anderson 1994:118 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Møller 2001:111 [ref. 25378], Nakabo 2002:1039 [ref. 26193]). •Questionably a synonym of Lycodes fasciatus (Schmidt 1904) -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Synonym of Lycodes fasciatus (Schmidt 1904) -- (Mecklenburg et al. 2002:720 [ref. 25968], Anderson & Fedorov 2004:24 [ref. 27603]). •Synonym of Lycodes brashnikovi Soldatov 1918 -- (Nazarkin 2010:603 [ref. 31019], Balushkin et al. 2011:989 [ref. 32449], Parin et al. 2014:408 [ref. 33547], Mecklenburg et al. 2016:173 [ref. 34440]). Current status: Synonym of Lycodes brashnikovi Soldatov 1918. Zoarcidae: Lycodinae. Habitat: marine.

nakamurae, Furcimanus Tanaka [S.] 1914:303, Pl. 82 (fig. 276) [Figures and descriptions of the fishes of Japan v. 18; ref. 14935] Off Niigata, Japan, Sea of Japan. Holotype (unique): ZUMT (lost). The spelling nakamurae is acceptable; early workers almost certainly Latinized the name first, and then added an 'e' (Art. 31.1.1) [WNE, 12 Mar. 2012] -- we suggest using nakamurae. Plate is in vol. 17 [ref. 14934]. •Synonym of Lycodes diapterus Gilbert 1892, but a valid subspecies nakamurai -- (Lindberg & Krasyukova 1975:149 [ref. 7348], Sheiko 1983:18 [ref. 37041]). •Valid as Furcimanus nakamurae Tanaka 1914 -- (Anderson 1994:119 [ref. 21438]). •Valid as Lycodes nakamurai (Tanaka 1914) -- (Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:224 [ref. 5722], Sokolovskaya et al. 1998:12 [ref. 24670], Nakabo 2000:1029 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:724 [ref. 25968], Nakabo 2002:1029 [ref. 26193], Choi et al. 2003:375, 640 [ref. 26218], Anderson & Fedorov 2004:27 [ref. 27603], Shinohara & Shirai 2005:172 [ref. 28232], Kim et al. 2006:276 [ref. 29191], Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:1002 [ref. 32449], Nazarkin et al. 2014:177 [ref. 33213], Shinohara et al. 2014:265 [ref. 33330], Saveliev et al. 2014:125 [ref. 33340], Parin et al. 2014:416 [ref. 33547], Kim et al. 2020:155 [ref. 39118], Sonoyama et al. 2020:94 [ref. 37637], Dyldin & Orlov 2021:839 [ref. 38925]). Current status: Valid as Lycodes nakamurae (Tanaka 1914). Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia). Habitat: marine.

nigricans, Lycodes Jensen [A. S.] 1952:21, Pl. 1 (fig. 1) [Meddelelser om Grønland v. 142 (no. 7); ref. 12776] Amerdlok fjord near Holsteinsborg, Greenland, 340-473 meters. Holotype: ZMUC P76803. Paratypes: ZMUC P76799-802 (4), P76804-15 (12). Type catalog: Nielsen 1974:77 [ref. 9588]. •Synonym of Lycodes seminudus Reinhardt 1837 -- (Andriashev 1973:546 [ref. 9214], Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:29 [ref. 27603], Balushkin et al. 2011:1011 [ref. 32449], Mecklenburg et al. 2014:142 [ref. 33463], Parin et al. 2014:421 [ref. 33547]). Current status: Synonym of Lycodes seminudus Reinhardt 1837. Zoarcidae: Lycodinae. Habitat: marine.

nishimurai, Lycodes Shinohara [G.] & Shirai [S. M.] 2005:172, Figs. 1-3 [Ichthyological Research v. 52 (no. 2); ref. 28232] Yamato Bank, 39°28.36'N, 135°0.31'E to 39°27.24'N, 134°59.02'E, Sea of Japan, depth 642-669 meters. Holotype: NSMT-P 63795. Paratypes: NSMT-P 63797-99 (3). •Valid as Lycodes nishimurai Shinohara & Shirai 2005 -- (Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:1002 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Saveliev et al. 2014:152 [ref. 33340], Parin et al. 2014:416 [ref. 33547]). Current status: Valid as Lycodes nishimurai Shinohara & Shirai 2005. Zoarcidae: Lycodinae. Distribution: Sea of Japan. Habitat: marine.

obscurus, Lycodes Toyoshima [M.] 1985:235, Figs. 65-66 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Southern Okhotsk Sea, 44°59.7'N, 144°21.2'E, depth 340 meters. Holotype: HUMZ 76006. Paratypes: HUMZ 92885-87 (3). •Valid as Lycodes obscurus Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1034 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1034 [ref. 26193], Anderson & Fedorov 2004:27 [ref. 27603], Ikeda et al. 2008:360 [ref. 29896]). •Synonym of Lycodes microporus Toyoshima 1983 -- (Balushkin et al. 2011:1001 [ref. 32449], Parin et al. 2014:416 [ref. 33547]). Current status: Synonym of Lycodes microporus Toyoshima 1983. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk Sea (if valid). Habitat: marine.

ocellatus, Lycodes Toyoshima [M.] 1985:223, Fig. 57 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Off Kushiro, eastern coast of Hokkaido, Japan, depth 600 meters. Holotype: HUMZ 36817. Paratypes: HUMZ 36811 (1), 36815 (1), 36823 (1) 36824 (1, missing in 1997), 36826 (1), 36829 (1), 36830 (1, missing in 1997), 36844 (1). •Valid as Lycodes ocellatus Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1037 [ref. 25182], Nakabo 2002:1037 [ref. 26193], Anderson & Fedorov 2004:27 [ref. 27603], Ikeda et al. 2008:361 [ref. 29896], Balushkin et al. 2011:1002 [ref. 32449]). Current status: Valid as Lycodes ocellatus Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Hokkaido, Japan. Habitat: marine.

ochotensis, Lycodes brevipes Schmidt [P. Yu.] 1950:89 [76] [Trudy Tichookeanskogo Komiteta, Akademija Nauk S.S.S.R. v. 6; ref. 12471] East coast of Sakhalin, 53°17'N, 144°10'E, Okhotsk Sea, depth 165 meters. Holotype (unique): ZIN 25271. Type catalog: Balushkin et al. 2011:1001 [ref. 32449], Balushkin et al. 2012:158 [ref. 33667]. On p. 76 of English translation. Appeared first without description as Schmidt in Taranetz 1937:163 [ref. 13384]. •Synonym of Lycodes brevipes Bean 1890, but a valid subspecies -- (Lindberg & Krasyukova 1975:155 [ref. 7348]). •Valid as Lycodes ochotensis Schmidt 1950 -- (Toyoshima 1985:211 [ref. 5722], Sheiko & Fedorov 2000:37 [ref. 25650], Møller 2001:111 [ref. 25378], Fedorov et al. 2003:115 [ref. 27579]). •Synonym of Lycodes brevipes Bean 1890 -- (Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:23 [ref. 27603], Nazarkin 2010:4 [ref. 30733] as Lycodes breviceps ochotensis on p. 2). •Synonym of Lycodes microlepidotus Schmidt 1950 -- (Balushkin et al. 2011:1001 [ref. 32449], Parin et al. 2014:415 [ref. 33547]). •Synonym of Lycodes ygreknotatus Schmidt 1950 -- (Nazarkin et al. 2024:119 [ref. 41547]). Current status: Synonym of Lycodes ygreknotatus Schmidt 1950. Zoarcidae: Lycodinae. Habitat: marine.

paamiuti, Lycodes Møller [P. R.] 2001:977, Figs. 4-5 [Copeia 2001 (no. 4); ref. 25812] Davis Strait, 66°25'N, 57°04'W, Atlantic, depth 678 meters. Holotype: ZMUC P762724. Paratypes: BMNH 1999.7.9.1-2 (2); BSKU 47297 (1), 47551 (1), 80158 (1), 80527 (1), 80530 (1); CNMFI 1999-0020 (4); ISH 804-1964 (2), 807-1964 (12), 147-1973 (3), 149-1973 (3), 6-1993 (10), 7-1994 (5); ROM 72564 (2); USNM 357279 (2); ZIN 51961 (2); ZMH 8421 (7); ZMUC P761327 (1), P761364 (1), P761646 (1), P761664-68 (5), P762215 (1), P762697-98 (2, the larger c&s), P762699-705 (7), P762707-17 (11), P762718-20 (3), P762723 (1), P762725-28 (4), P762730-32 (3), P763053-56 (4), 763057 (1), P763182 (1), P763184 (1), P763185-86 (2), P764086-91 (6, 1c&s). Additional paratypes at ISH, NHMT, ZMUB, ZMUC and ZMUO. Type catalog: Balushkin et al. 2011:1002 [ref. 32449], Balushkin et al. 2012:162 [ref. 33667]. •Valid as Lycodes paamiuti Møller 2001 -- (Anderson & Fedorov 2004:27 [ref. 27603], Møller et al. 2010:62 [ref. 31169], Mecklenburg et al. 2011:129 [ref. 31212], Balushkin et al. 2011:1002 [ref. 32449], Afonso et al. 2016:65, 205 [ref. 36199], Mecklenburg et al. 2018:340 [ref. 36370], Page et al. 2023:212 [ref. 40505]). Current status: Valid as Lycodes paamiuti Møller 2001. Zoarcidae: Lycodinae. Distribution: North Atlantic and Arctic Ocean: Davis Strait to Greenland and Norwegian seas. Habitat: marine.

pacificus, Lycodes Collett [R.] 1879:381, Fig. [Proceedings of the Zoological Society of London 1879 (pt 2) (art. 4); ref. 886] Said to be from Japan (probably from Alaska). Holotype (unique): ZMB 8280. Described in Lycodes, placed in Lycodopsis on p. 382. •Valid as Lycodopsis pacificus (Collett 1879) -- (Toyoshima 1985:238 [ref. 5722], Balushkin et al. 2011:1018 [ref. 32449] as pacifica, Page et al. 2013:163 [ref. 32708]). •Valid as Lycodes pacificus Collett 1879 -- (Eschmeyer & Herald 1983:105 [ref. 9277], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:704 [ref. 25968], Anderson & Fedorov 2004:27 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Love et al. 2005:143 [ref. 37547], Pietsch & Orr 2015:58 [ref. 34694], Kells et al. 2016:274 [ref. 35888], Burton & Lea 2019:84 [ref. 37205], Love et al. 2021:184 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes pacificus Collett 1879. Zoarcidae: Lycodinae. Distribution: Northeastern Pacific: Aleutian Islands (U.S.A.) and Gulf of Alaska (U.S.A.) south to Pacific coast of northern Baja California (Mexico). Habitat: marine.

palearis, Lycodes Gilbert [C. H.] 1896:454 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bristol Bay, Alaska, U.S.A., Albatross station 3253, 3254, depth 36-46 fathoms. Syntypes: CAS-SU 3165 (1), 3486 (1); USNM 48592 (1). Type catalog: Böhlke 1953:101 [ref. 12291], Springer & Anderson 1997:20 [ref. 22953]. •Valid as Lycodes palearis Gilbert 1896 -- (Lindberg & Krasyukova 1975:157 [ref. 7348], Eschmeyer & Herald 1983:106 [ref. 9277], Toyoshima 1985:212 [ref. 5722], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:719 [ref. 25968], Møller 2001:111 [ref. 25378], Fedorov et al. 2003:115 [ref. 27579], Anderson & Fedorov 2004:27 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Love et al. 2005:143 [ref. 37547], Mecklenburg et al. 2007:182 [ref. 29448], Nazarkin 2010:603 [ref. 31019], Balushkin et al. 2011:1002 [ref. 32449], Page et al. 2013:163 [ref. 32708], Parin et al. 2014:417 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Pietsch & Orr 2015:58 [ref. 34694], Mecklenburg et al. 2016:172 [ref. 34440], Kells et al. 2016:274 [ref. 35888], Mecklenburg et al. 2018:342 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:184 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes palearis Gilbert 1896. Zoarcidae: Lycodinae. Distribution: North Pacific and adjacent Arctic. Habitat: marine.

pallida, Lycodes vahlii forma Smitt [F. A.] 1901:24 [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 27 (no. 4); ref. 18732] Not intended as new; this is a subsequent treatment of Lycodes pallidus Collett 1879 as a subspecific name. •In the synonymy of Lycodes vahlii Reinhardt 1831 -- (Anderson 1994:119 [ref. 21438]). •In the synonymy of Lycodes pallidus Collett 1879 -- (Møller 2001:983 [ref. 25812], Carl 2002:65 [ref. 26925]). •In the synonymy of Lycodes vahlii or Lycodes pallidus Collett 1879 -- (Anderson & Fedorov 2004:48 [ref. 27603]). Current status: Synonym of Lycodes. Zoarcidae: Lycodinae.

pallidus, Lycodes Collett [R.] 1879:70 [Forhandlinger i Videnskabs-selskabet i Christiania (for 1878) No. 14; ref. 885] West of Norskøyane, Spitsbergen, depth 475 and 839 meters. Syntypes: ZMUO J4540 (1), J4541 (1). Type catalog: Pethon 1969:12 [ref. 19268]. Also appeared as new in Collett 1880:110, Pl. 3 (figs. 26-27) [ref. 887]. •Valid as Lycodes pallidus Collett 1879 -- (Andriashev 1973:544 [ref. 9214], Andriashev in Whitehead et al. 1986:1141 [ref. 13677], Robins & Ray 1986:237 [ref. 23100], Scott & Scott 1988:409 [ref. 25518], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Saito & Okamura in Okamura et al. 1995:199 [ref. 22531], Mecklenburg et al. 2002:721 [ref. 25968], Møller 2001:982 [ref. 25812], Carl 2002:65 [ref. 26925], Anderson & Fedorov 2004:27 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Møller et al. 2010:62 [ref. 31169], Balushkin et al. 2011:1003 [ref. 32449], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Parin et al. 2014:417 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:65, 206 [ref. 36199], Mecklenburg et al. 2016:174 [ref. 34440] see remarks, Dolgov et al. 2018:213 [ref. 41055],, Mecklenburg et al. 2018:344 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:184 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes pallidus Collett 1879. Zoarcidae: Lycodinae. Distribution: Arctic Ocean: circumpolar, and North Atlantic. Habitat: marine.

panthera, Lycodes knipowitschi Schmidt [P. Yu.] 1950:103 [89], Pl. 8 (fig. 2) [Trudy Tichookeanskogo Komiteta, Akademija Nauk S.S.S.R. v. 6; ref. 12471] Bank northeast of Iona Island, Okhotsk Sea, depth 35-75 meters. Syntypes: ?ZIN 24839 (2), 24824 (2), 24842 (1), 30000 (1). Type catalog: Balushkin et al. 2011:999 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667] with original description dated 1937. On p. 89 of English translation. •Synonym of Lycodes knipowitschi Popov 1931 -- (Toyoshima 1985:195 [ref. 5722], Sheiko & Fedorov 2000:36 [ref. 25650], Balushkin et al. 2011:999 [ref. 32449]). •Synonym of Lycodes mucosus Richardson 1855 -- (Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:27 [ref. 27603], Parin et al. 2014:416 [ref. 33547]). Current status: Synonym of Lycodes mucosus Richardson 1855. Zoarcidae: Lycodinae. Habitat: marine.

paucidens, Leurynnis Lockington [W. N.] 1879:102 [Mining and Scientific Press. An illustrated journal of mining, popular sciences and general news v. 39 (no. 7) 16 Aug.; ref. 20592] Off San Francisco, California, U.S.A. Lectotype: USNM 23502. Paralectotypes: USNM 336667 [ex USNM 23502] (1). Type catalog: Springer & Anderson 1997:20 [ref. 22953]. Original genus as Leurhynnis as above, Leurynnis in ref. 2815. Also appeared in Lockington 1880:326 [ref. 2815]. Lectotype selected by Springer & Anderson 1997:20 [ref. 22953]. •Valid as Lacodopsis paucidens Lockington 1879 -- (Herrera 1896:67 [ref. 39336] with author as Gill). •Synonym of Lycodes pacificus Collett 1879 -- (Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:704 [ref. 25968], Anderson & Fedorov 2004:27 [ref. 27603]). Current status: Synonym of Lycodes pacificus Collett 1879. Zoarcidae: Lycodinae. Habitat: marine.

paucidens, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] in Taranetz 1937:163 [Izvestiia Tikhookeanskogo Nauchno-Issledovatel'skogo Instituta Rybnogo Khoziaistva i Okeanografii [Proceedings, Pacific Research Institute of Fisheries and Oceanography, TINRO, Moskva] v. 11; ref. 13384] Not available, no distinguishing features. •In the synonymy of Lycodes raridens Taranetz & Andriashev 1937 -- (Anderson 1994:119 [ref. 21438]). •In the synonymy of Lycodes brevicaudus Taranetz & Andriashev 1935 -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Nomen nudum in the family Zoarcidae, genus Lycodes -- (Anderson & Fedorov 2004:48 [ref. 27603], Parin et al. 2014:419 [ref. 33547]). Nomen Nudum. Current status: Synonym of Lycodes raridens Taranetz & Andriashev 1937. Zoarcidae: Lycodinae. Habitat: marine.

paucilepidotus, Lycodes Toyoshima [M.] 1985:196, Fig. 43 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Northern Okhotsk Sea, 58°33'N, 147°25'E, depth 138 meters. Holotype: HUMZ 58117. Paratypes: HUMZ 55142 (1), 55144 (1), 55159 (1), 57658-59 (2), 57792 (1), 57862 (1), 57865 (1), 57867-69 (3), 58058 (1), 58117 (1), 58222 (1), 58224 (1), 60756 (1), 60814-20 (7), 60822-23 (2), 60844 (1), 60846 (1), 61071 (1), 61458 (1), 61480 (1), 61497 (1), 61506 (1), 61509 (1), 61512 (1), 68063 (1). •Valid as Lycodes paucilepidotus Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1038 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1038 [ref. 26193], Fedorov et al. 2003:116 [ref. 27579], Anderson & Fedorov 2004:28 [ref. 27603], Balushkin et al. 2011:1005 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:418 [ref. 33547], Dyldin & Orlov 2021:840 [ref. 38925]). Current status: Valid as Lycodes paucilepidotus Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk Sea and Sea of Japan. Habitat: marine.

pectoralis, Lycodes Toyoshima [M.] 1985:228, Fig. 60 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Southern Okhotsk Sea, 45°37'N, 143°53'E, depth 290-480 meters. Holotype: HUMZ 49083. Paratypes: HUMZ 33951 (1), 33956 (1), 33964 (1), 49071 (1), 49075-77 (3), 49082 (1), 49084-91 (8). •Valid as Lycodes pectoralis Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1029 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:724 [ref. 25968], Nakabo 2002:1029 [ref. 26193], Fedorov et al. 2003:116 [ref. 27579], Anderson & Fedorov 2004:28 [ref. 27603], Shinohara & Shirai 2005:172 [ref. 28232], Kim et al. 2006:275 [ref. 29191], Balushkin et al. 2011:1005 [ref. 32449], Nazarkin at al. 2014:177 [ref. 33213], Saveliev et al. 2014:152 [ref. 33340], Parin et al. 2014:418 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331], Kim et al. 2020:155 [ref. 39118], Dyldin & Orlov 2021:840 [ref. 38925]). Current status: Valid as Lycodes pectoralis Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk Sea and Kuril Islands (presence in Korea is a misidentification [ref. 33340]). Habitat: marine.

perspicillum, Lycodes Krøyer [H. N.] 1845:140 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1844 (no. 8); ref. 19477] Newfoundland, Canada. Syntypes: ZMUC 23 (1) Greenland, ZMUC (1 lost). Type catalog: Nielsen 1974:77 [ref. 9588]. •In part a synonym of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603], Parin et al. 2014:419 [ref. 33547]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Habitat: marine.

polaris, Blennius Sabine [E.] 1824:212 [ccxii] [A supplement to the appendix of Captain Parry's voyage; ref. 18649] North Georgia, Boothia Peninsula, Northwest Territories, Canada. No types known. •Valid as Lycodes polaris (Sabine 1824) -- (Andriashev 1973:545 [ref. 9214], Andriashev in Whitehead et al. 1986:1142 [ref. 13677], Scott & Scott 1988:409 [ref. 25518], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:710 [ref. 25968], Anderson & Fedorov 2004:28 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Love et al. 2005:143 [ref. 37547], Møller et al. 2010:62 [ref. 31169], Balushkin et al. 2011:1006 [ref. 32449], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:144, 145 [ref. 33463], Parin et al. 2014:418 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:66, 207 [ref. 36199], Kells et al. 2016:274 [ref. 35888], Mecklenburg et al. 2016:176 [ref. 34440] see remarks, Dolgov et al. 2018:214 [ref. 41055], Mecklenburg et al. 2018:346 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:184 [ref. 39279], Chernova et al. 2022:8 [ref. 39177], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes polaris (Sabine 1824). Zoarcidae: Lycodinae. Distribution: Arctic Ocean, North Atlantic and North Pacific: nearly circumpolar. Habitat: marine.

raridens, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] in Andriashev 1937:335, Figs. 13, 14 [Issled. Morey SSSR, Leningrad v. 25; ref. 20242] Bering Sea, Gulf of Anadry, 61°52'N, 173°11'W, depth 66 meters. Holotype: ZIN 25070. Paratypes: ZIN 13009 (1), 23757 (1), 24845 (6), 25071 (1), 29999 (2), 30007 (2), 30008 (1), 30010 (2). Type catalog: Balushkin et al. 2011:1008 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. •Questionably a synonym of Lycodes brevicaudus Taranetz & Andriashev 1935 -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Valid as Lycodes raridens Taranetz & Andriashev 1937 -- (Lindberg & Krasyukova 1975:161 [ref. 7348], Toyoshima 1985:198 [ref. 5722], Scott & Scott 1988:408 [ref. 25518], Anderson 1994:119 [ref. 21438], Chikilev & Kharitonov 2000:100 [ref. 24916], Mecklenburg et al. 2002:711 [ref. 25968], Fedorov et al. 2003:116 [ref. 27579], Anderson & Fedorov 2004:28 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Love et al. 2005:144 [ref. 37547], Mecklenburg et al. 2007:182 [ref. 29448], Mecklenburg et al. 2011:128 [ref. 31212], Balushkin et al. 2011:1008 [ref. 32449], Page et al. 2013:163 [ref. 32708], Parin et al. 2014:419 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Mecklenburg et al. 2016:178 [ref. 34440] see remarks, Kells et al. 2016:274 [ref. 35888], Mecklenburg et al. 2018:348 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Dyldin & Orlov 2021:840 [ref. 38925], Love et al. 2021:184 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes raridens Taranetz & Andriashev 1937. Zoarcidae: Lycodinae. Distribution: Arctic and adjacent North Pacific. Habitat: marine.

reticulatus, Lycodes Reinhardt [J. C. H.] 1835:7 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1834-35; ref. 3695] Greenland. Syntypes: ZMUC 17 (1) Umanak, ZMUC 21 (1) Fiskenaesset. Type catalog: Nielsen 1974:78 [ref. 9588]. Also appeared in Reinhardt 1837:115, 195, Pl. 6 [33, 113, Pl. 6 of separate] [ref. 3691]. •Valid as Lycodes reticulatus Reinhardt 1835 -- (Andriashev 1973:545 [ref. 9214], Andriashev in Whitehead et al. 1986:1143 [ref. 13677], Robins & Ray 1986:237 [ref. 23100], Scott & Scott 1988:410 [ref. 25518], McAllister 1990:161 [ref. 14674], Saito & Okamura in Okamura et al. 1995:198 [ref. 22531], Mecklenburg et al. 2002:712 [ref. 25968], Anderson & Fedorov 2004:28 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Love et al. 2005:144 [ref. 37547], Møller et al. 2010:62 [ref. 31169], Mecklenburg et al. 2011:128 [ref. 31212] with comments, Balushkin et al. 2011:1009 [ref. 32449], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:419 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:66, 208 [ref. 36199], Mecklenburg et al. 2016:180 [ref. 34440] see remarks, Dolgov et al. 2018:216 [ref. 41055], Mecklenburg et al. 2018:350 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:184 [ref. 39279], Simian et al. 2022:376 [ref. 40776], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Distribution: North Atlantic and Arctic Ocean. Habitat: marine.

rossi, Lycodes Malmgren [A. J.] 1865:516 [Öfversigt af Kongliga Vetenskaps-Akademiens förhandlingar, Kungliga Svenska Vetenskapsakademien. 1864 (no. 10) [v. 21]; ref. 17598] Treurenberg Bay, Spitsbergen, 5 fathoms and other localities. Syntypes: NRM 9398 (1, juvenile), NRM not found (1). Based in part Blennius polaris of J.C. Ross, Append. to Parry's Attempt to reach the North-pole, 1827, p. 200. Sometimes as rossii but rossi is correct. •Synonym of Lycodes reticulatus Reinhardt 19835 with question -- (Meckenburg et al. 2011:127 [ref. 31212]). •Valid as Lycodes rossi Malmgren 1865 -- (Andriashev 1973:546 [ref. 9214], McAllister et al. 1981:825 [ref. 5431], Andriashev in Whitehead et al. 1986:1143 [ref. 13677], McAllister 1990:161 [ref. 14674], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:713 [ref. 25968], Anderson & Fedorov 2004:28 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Love et al. 2005:144 [ref. 37547], Balushkin et al. 2011:1010 [ref. 32449], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:420 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Mecklenburg et al. 2016:182 [ref. 34440] see remarks, Dolgov et al. 2018:218 [ref. 41055], Mecklenburg et al. 2018:352 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948] as rossii, Love et al. 2021:184 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae. Distribution: Arctic and North Atlantic. Habitat: marine.

sagittarius, Lycodes McAllister [D. E.] 1976:9, Figs. 1-3 [Publications in Biological Oceanography, National Museum Natural Science Canada No. 9; ref. 7446] Beaufort Sea, 50 miles north-northeast of Brownlow Point, 70°51'30"N, 145°17'W, depth 357 meters. Holotype: NMC 74-282. Paratypes: BMNH 1974.10.5.1 (1); NMC 74-275 (1), 74-282A (1); ROM 30520 (1); USNM 212282 (1); ZIN 32050 (spec. 1 of 2); ZMUC P761071 (1). Type catalog: Springer & Anderson 1997:22 [ref. 22953], Balushkin et al. 2011:1010 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667]. •Valid as Lycodes sagittarius McAllister 1976 -- (Andriashev in Whitehead et al. 1986:1144 [ref. 13677], McAllister et al. 1981:828 [ref. 5431], McAllister 1990:163 [ref. 14674], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:717 [ref. 25968], Møller 2001:111 [ref. 25378], Anderson & Fedorov 2004:29 [ref. 27603], Love et al. 2005:144 [ref. 37547], Balushkin et al. 2011:1011 [ref. 32449], Dolgov 2013:920 [ref. 33112], Mecklenburg 2014:145 [ref. 33463], Parin et al. 2014:420 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:67, 209 [ref. 36199], Mecklenburg et al. 2016:184 [ref. 34440] see remarks, Dolgov et al. 2018:220 [ref. 41055], Mecklenburg et al. 2018:354 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:184 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes sagittarius McAllister 1976. Zoarcidae: Lycodinae. Distribution: Arctic Ocean. Habitat: marine.

schmidti, Lycodes Soldatov [V. K.] 1918:115, Fig. 2 [An. Mus. Zool. Acad. Sci. Petrograd v. 22; ref. 21830] Near Rymnik, eastern Sakhalin Island, Okhotsk Sea; western Sakhalin Island, 50°31'N, 141°56'E, Sea of Japan. Type catalog: Balushkin et al. 2011:1020 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667] Also appeared in Soldatov 1922:115, Fig. 2 [ref. 15429]. Objectively invalid; preoccupied by Lycodes schmidti Gratzianov 1907, replaced by Lycodes sigmatoides Lindberg & Krasyukova 1975. •Questionably a synonym of Lycodes brevicaudus Taranetz & Andriashev 1937 -- (Sheiko & Fedorov 2000:36 [ref. 25650]). •Objective synonym of Lycodes sigmatoides Lindberg & Krasyukova 1975 -- (Toyoshima 1985:204 [ref. 5722], Sheiko & Fedorov 2000:37 [ref. 25650], Anderson & Fedorov 2004:29 [ref. 27603]). •Synonym of Lycodes tanakae Jordan & Thompson 1914 -- (Balushkin et al. 2011:1014 [ref. 32449], Parin et al. 2014:422 [ref. 33547]). 1°Homonym. Current status: Synonym of Lycodes tanakae Jordan & Thompson 1914. Zoarcidae: Lycodinae. Habitat: marine.

semenovi, Lycodes Popov [A. M.] 1931:144, Pl. 1 (fig. 2) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] 56°09'N, 144°30'E, Okhotsk Sea, off St. Iona Island, depth 307 meters. Holotype (unique): ZIN 29995 [not 2995]. Type catalog: Balushkin et al. 2011:1011 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667]. •Valid as Lycodes semenovi Popov 1931 -- (Toyoshima 1985:205 [ref. 5722], Anderson 1994:119 [ref. 21438], Fedorov et al. 2003:116 [ref. 27579], Anderson & Fedorov 2004:29 [ref. 27603], Nazakin & Fedorov 2007:701 [ref. 29424], Ikeda et al. 2008:357 [ref. 29896], Balushkin et al. 2011:1011 [ref. 32449], Parin et al. 2014:420 [ref. 33547], Dyldin & Orlov 2021:840 [ref. 38925]). Current status: Valid as Lycodes semenovi Popov 1931. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk Sea (Russia). Habitat: marine.

seminuda, Lycodes reticulatus forma Smitt [F. A.] 1901:31 [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 27 (no. 4); ref. 18732] Not intended as new; subsequent treatment of Lycodes seminudus Reinhardt 1837 as a subspecific name. •Not intended as new, in the family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). •In the synonymy of Lycodes reticulatus Reinhardt 1835 -- (Anderson 1994:119 [ref. 21438], Parin et al. 2014:419 [ref. 33547]). Current status: Synonym of Lycodes reticulatus Reinhardt 1835. Zoarcidae: Lycodinae. Habitat: marine.

seminudus, Lycodes Reinhardt [J. C. H.] 1837:223 [117] [Det Kongelige Danske videnskabernes selskabs naturvidenskabelige og mathematiske afhandlinger v. 7; ref. 13398] Omenak, Greenland, 71°N. Holotype: ZMUC 15 [or 9]. Type catalog: Nielsen 1974:78 [ref. 9588]. On p. 117 of separate. •Valid as Lycodes seminudus Reinhardt 1837 -- (Andriashev 1973:546 [ref. 9214], McAllister et al. 1981:833 [ref. 5431], Andriashev in Whitehead et al. 1986:1145 [ref. 13677], McAllister 1990:163 [ref. 14674], Anderson 1994:119 [ref. 21438], Saito & Okamura in Okamura et al. 1995:200 [ref. 22531], Mecklenburg et al. 2002:707 [ref. 25968], Anderson & Fedorov 2004:29 [ref. 27603], Love et al. 2005:144 [ref. 37547], Møller et al. 2010:62 [ref. 31169], Balushkin et al. 2011:1011 [ref. 32449], Maslenikov et al. 2013:17 [ref. 37947], Page et al. 2013:163 [ref. 32708], Dolgov 2013:920 [ref. 33112], Mecklenburg et al. 2014:141 [ref. 33463], Parin et al. 2014:421 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:67, 210 [ref. 36199], Mecklenburg et al. 2016:186 [ref. 34440] see remarks, Dolgov et al. 2018:222 [ref. 41055], Mecklenburg et al. 2018:356 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:185 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes seminudus Reinhardt 1837. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and North Atlantic and North Pacific. Habitat: marine.

septentrionalis, Lycodes vahli var. Knipowitsch [N. M.] 1906:95 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Barents Sea, Russia. Syntypes: ZIN 13437-71 (1 lost, 1 lost, 1, 3, 3, 1, 2, 1 lost, 3, 1, 1, 1, 1 lost, 2, 3, 1, 1, 1 lost, 1, 1 lost, 2 lost, 3, 2, 1 lost, 1, 1 lost, 1 lost, 1, 1, 1, 2, 1, 1, 1, 4), 14248 (1). Type catalog: Balushkin et al. 2011:995 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. Spelled sententrionalis by Knipowitsch 1926:111 [ref. 22166]. •Synonym of Lycodes vahlii Reinhardt 1831 -- (Andriashev 1973:543 [ref. 9214], Anderson 1994:119 [ref. 21438]). •Synonym of Lycodes gracilis Sars 1867 -- (Carl 2002:75 [ref. 26925], Anderson & Fedorov 2004:25 [ref. 27603], Balushkin et al. 2011:995 [ref. 32449], Parin et al. 2014:411 [ref. 33547]). Current status: Synonym of Lycodes gracilis Sars 1867. Zoarcidae: Lycodinae. Habitat: marine.

sigmatoides, Lycodes Lindberg [G. U.] & Krasyukova [Z. V.] 1975:161, Fig. 127 [ ref. 7348] Near Rymnik, eastern Sakhalin Island, Okhotsk Sea; western Sakhalin Island, 50°31'N, 141°56'E, Sea of Japan. Syntypes: (5) ZIN 13009 (3). Non-types: ZIN 19165-66 (3, 1). Type catalog: Balushkin et al. 2011:1014 [ref. 32449], Balushkin et al. 2012:165 [ref. 33667] with ZIN 19165 as syntypes. Replacement name for Lycodes schmidti Soldatov 1918, preoccupied by Lycodes schmidti Gratzianov 1907. Appeared in error as sigmatus in the index. •Valid as Lycodes sigmatoides Lindberg & Krasyukova 1975 -- (Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:204 [ref. 5722], Anderson 1994:119 [ref. 21438], Sokolovskaya et al. 1998:12 [ref. 24670] as sygmatoides, Nakabo 2000:1038 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1038 [ref. 26193], Fedorov et al. 2003:117 [ref. 27579], Anderson & Fedorov 2004:29 [ref. 27603], Kim et al. 2006:276 [ref. 29191], Shinohara et al. 2011:51 [ref. 31715], Balushkin et al. 2011:1014 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:421 [ref. 33547], Kim et al. 2020:155 [ref. 39118]). Replacement Name. Current status: Valid as Lycodes sigmatoides Lindberg & Krasyukova 1975. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia) and Kuril Islands north to Sea of Okhotsk and Kamchatka (Russia). Habitat: marine.

similis, Lycodes Jensen [A. S.] 1902:205 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Jan Mayen, northeastern Atlantic, depth 371 fathoms. Syntypes: ZMUC 220-234 (25) (but see Møller 2001:985 [ref. 25812], who gives different catalog numbers). Type catalog: Nielsen 1974:78 [ref. 9588]. Appeared in Jensen 1904:39, 46-48, Pl. 5 (figs.2a-d), Pl. 6 (figs. 3a-d) [ref. 15186] as Lycodes pallidus var. similis according to Møller 2001:983 [ref. 25812]. •Synonym of Lycodes pallidus Collett 1879 -- (Andriashev 1973:544 [ref. 9214], Anderson 1994:119 [ref. 21438], Møller 2001:983 [ref. 25812], Anderson & Fedorov 2004:27 [ref. 27603], Parin et al. 2014:417 [ref. 33547]). Current status: Synonym of Lycodes pallidus Collett 1879. Zoarcidae: Lycodinae. Habitat: marine.

soldatovi, Lycodes Taranetz [A. Ya.] & Andriashev [A. P.] 1935:246, Fig. 3 [Zoologischer Anzeiger v. 112 (nos 9-10); ref. 15744] Okhotsk Sea, east of Cape Terpenija, 49°09.5'N, 145°17.5'E, depth 440 meters. Syntypes: ZIN 24846 (1), 25190 (2). Type catalog: Balushkin et al. 2011:1012 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. •Valid as Lycodes soldatovi Taranetz & Andriashev 1935 -- (Lindberg & Krasyukova 1975:153 [ref. 7348], Toyoshima in Amaoka et al. 1983:267, 331 [ref. 8379], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:207 [ref. 5722], Anderson 1994:119 [ref. 21438], Nakabo 2000:1035 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:718 [ref. 25968], Love et al. 2005:144 [ref. 37547], Møller 2001:111 [ref. 25378], Nakabo 2002:1035 [ref. 26193], Fedorov et al. 2003:117 [ref. 27579], Anderson & Fedorov 2004:29 [ref. 27603], Balushkin et al. 2011:1012 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:422 [ref. 33547], Dyldin & Orlov 2021:840 [ref. 38925], Love et al. 2021:185 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes soldatovi Taranetz & Andriashev 1935. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk Sea, Sea of Japan, and Bering seas (the latter doubtful on ref. 33547). Habitat: marine.

spitzbergensis, Lycodes celatus var. Jensen [A. S.] 1902:209 [Videnskabelige Meddelelser fra den Naturhistoriske Forening i Kjøbenhavn, Aaret 1901; ref. 14758] Spitsbergen, Arctic regions, depth 39-75 fathoms. No types known. •Synonym of Lycodes rossi Malmgren 1865 -- (Anderson 1994:119 [ref. 21438], Anderson & Fedorov 2004:28 [ref. 27603], Parin et al. 2014:420 [ref. 33547]). Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae. Habitat: marine.

squamiventer, Lycodes pallidus var. Jensen [A. S.] 1904:39, 46, Pl. 4 (fig. 2) [Danish Ingolf-Expedition v. 2 (no. 4); ref. 15186] Iceland. Lectotype: ZMUC 235. Paralectotypes: ZMUC 7-8 (2), 237239 (3); NMC 74-0395 (1), ZIN 40718 (1), ZMUB 1811 (2), ZMUO J4546 (2). Type catalog: Nielsen 1974:77 [ref. 9588]; Balushkin et al. 2011:1013 [ref. 32449], Balushkin et al. 2012:163 [ref. 33667]. •Valid as Lycodes squamiventer Jensen 1904 -- (Andriashev 1973:546 [ref. 9214], Andriashev in Whitehead et al. 1986:1145 [ref. 13677], McAllister et al. 1981:832 [ref. 5431], Anderson 1994:119 [ref. 21438], Mecklenburg et al. 2002:722 [ref. 25968], Møller 2001:985 [ref. 25812], Anderson & Fedorov 2004:29 [ref. 27603], Love et al. 2005:144 [ref. 37547], Møller et al. 2010:63 [ref. 31169], Balushkin et al. 2011:1013 [ref. 32449], Dolgov 2013:920 [ref. 33112], Mecklenburg 2014:145 [ref. 33463], Parin et al. 2014:422 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Afonso et al. 2016:67, 211 [ref. 36199], Mecklenburg et al. 2016:188 [ref. 34440], Mecklenburg et al. 2018:358 [ref. 36370] see remarks, Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:185 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes squamiventer Jensen 1904. Zoarcidae: Lycodinae. Distribution: North Atlantic and Arctic Ocean. Habitat: marine.

squamosa, Lycodes turneri atlanticus morpha Vladykov [V. D.] & Tremblay [J.-L.] 1936:39 [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] Described as a morpha, regarded as infrasubspecific and unavailable. •Infrasubspecific and unavailable, in family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Infrasubspecific. Zoarcidae: Lycodinae.

subarcticus, Lycodes rossi forma Knipowitsch [N. M.] 1906:27 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Syntypes: ZIN 13408 (1), 13409 (1), 13424 (2), 13425 (1), 13426 (2), 13427 (2). Type catalog: Balushkin et al. 2011:1010 [ref. 32449], Balushkin et al. 2012:164 [ref. 33667]. Not available, infrasubspecific; publication in an available way as a species or subspecies not investigated [Knipowitsch used v. [variety] and f. [forma] -- in his work, the former is considered subspecific the latter infrasubspecific; see Article 45(g).]. From Barents Sea, Russia. •In the synonymy of Lycodes rossii Malmgren 1865 -- (Anderson 1994:119 [ref. 21438] as rossi, Anderson & Fedorov 2004:48 [ref. 27603], Balushkin et al. 2011:1010 [ref. 32449], Parin et al. 2014:420 [ref. 33547]). Infrasubspecific. Current status: Synonym of Lycodes rossi Malmgren 1865. Zoarcidae: Lycodinae. Habitat: marine.

tanakae, Lycodes Jordan [D. S.] & Thompson [W. F.] 1914:299, Pl. 37 (fig. 2) [Memoirs of the Carnegie Museum v. 6 (no. 4); ref. 2543] Noto, Hondo, Japan. Holotype (unique): FMNH 57068 [ex CM 6004]. Type catalog: Henn 1928:98 [ref. 12305], Ibarra & Stewart 1987:54 [ref. 12367]. The spelling tanakae is acceptable; early workers almost certainly Latinized the name first, and then added an 'e' (Art. 31.1.1) [WNE, 12 Mar. 2012] -- we now suggest using tanakae.. •Valid as Lycodes tanakae Jordan & Thompson 1914 -- (Lindberg & Krasyukova 1975:160 [ref. 7348] as tanakae, Sheiko 1983:18 [ref. 37041] as tanakai, Toyoshima in Amaoka et al. 1983:261, 330 [ref. 8379] as tanakae, Toyoshima in Masuda et al. 1984:307 [ref. 6441], Toyoshima 1985:193 [ref. 5722], Anderson 1994:119 [ref. 21438] as tanakae, Sokolovskaya et al. 1998:12 [ref. 24670], Nakabo 2000:1038 [ref. 25182] as tanakai, Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1038 [ref. 26193] as tanakai, Choi et al. 2003:375, 640 [ref. 26218] as tanakai, Anderson & Fedorov 2004:30 [ref. 27603], Kim et al. 2006:278 [ref. 29191], Nazarkin 2010:603 [ref. 31019], Shinohara et al. 2011:52 [ref. 31715], Balushkin et al. 2011:1013 [ref. 32449], Nazarkin et al. 2014:177 [ref. 33213], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:422 [ref. 33547], Tohkairin et al. 2015:12 [ref. 35331], Kim et al. 2020:155 [ref. 39118], Sonoyama et al. 2020:94 [ref. 37637], Dyldin & Orlov 2021:841 [ref. 38925]). Current status: Valid as Lycodes tanakae Jordan & Thompson 1914. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Sea of Japan (Korea, Japan, Russia) and Sea of Okhotsk. Habitat: marine.

taranetzi, Lycodes (Furcimanus) Andriashev [A. P.] in Lindberg & Krasyukova 1975:152, Fig. 119-A [ ref. 7348] Pacific side of Iturup Island, depth 415 meters. Holotype: ZIN 42247. Paratypes: ZIN 42248 (7), 42262 (1). Type catalog: Balushkin et al. 2011:977 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. Author is A. N. Andriashev. On p. 185 of English translation. Appeared first as name only in Lindberg 1959. •Synonym of Lycodes hubbsi Matsubara 1955 -- (Toyoshima 1985:226 [ref. 5722], Anderson 1994:118 [ref. 21438], Sheiko & Fedorov 2000:36 [ref. 25650], Anderson & Fedorov 2004:25 [ref. 27603], Balushkin et al. 2011:997 [ref. 32449], Parin et al. 2014:412 [ref. 33547]). Current status: Synonym of Lycodes hubbsi Matsubara 1955. Zoarcidae: Lycodinae. Habitat: marine.

terraenovae, Lycodes Collett [R.] 1896:54 [Poissons provenant des campagnes du Yacht "L'Hirondelle" (1885-1888) Fasc. 10; ref. 890] Newfoundland Banks, northwestern Atlantic, 46°04'40"N, 49°02'30"W, probably station 161 [not 162], depth 1267 meters. Lectotype: MOM-POI 295 [ex syntype b]. Paralectotypes: (3) MOM-POI 295 (1, former syntype a), 4651 (1, former syntype c, in 3 pieces), 3005 (former syntype d, stuffed, poor condition). Type catalog: Belloc 1949:14 [ref. 21544]. Originally as Terrae Novae; as a name "donnes ici provisoirement..." Lectotype selected by Møller 2000:847 [ref. 24585]. See Møller 2000:849 [ref. 24585] for information on the type locality. •Status uncertain -- (Anderson 1994:119 [ref. 21438]). •Valid as Lycodes terraenovae Collett 1896 -- (McAllister 1990:163 [ref. 14674], Møller 1997:47 [ref. 22879], Møller 2000:846 [ref. 24585], Carl 2002:65 [ref. 26925], Moore et al. 2003:228 [ref. 27076], Anderson 2003:1741 [ref. 27106], Anderson & Fedorov 2004:30 [ref. 27603], Møller et al. 2010:63 [ref. 31169], Balushkin et al. 2011:1015 [ref. 32449], Afonso et al. 2016:67, 212 [ref. 36199], Anderson in Carpenter & De Angelis 2016:2760 [ref. 34618], Bañón et al. 2021:7 [ref. 40417], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes terraenovae Collett 1896. Zoarcidae: Lycodinae. Distribution: Northwestern Atlantic, northeastern Atlantic, South Atlantic. Habitat: marine.

turneri, Lycodes Bean [T. H.] 1879:463 [Proceedings of the United States National Museum v. 1 (no. 59); ref. 16020] Saint Michaels Island, Alaska, U.S.A. Holotype (unique): USNM 21529. Type catalog: Springer & Anderson 1997:22 [ref. 22953]. •Valid as Lycodes turneri Bean 1879 -- (Toyoshima 1985:202 [ref. 5722], Robins & Ray 1986:237 [ref. 23100], Anderson 1994:119 [ref. 21438], Sheiko & Fedorov 2000:37 [ref. 25650], Mecklenburg et al. 2002:709 [ref. 25968], Anderson & Fedorov 2004:30 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Love et al. 2005:144 [ref. 37547], Mecklenburg et al. 2007:181 [ref. 29448], Balushkin et al. 2011:1015 [ref. 32449], Page et al. 2013:163 [ref. 32708], Parin et al. 2014:423 [ref. 33547], Mecklenburg & Steinke 2015:165 [ref. 34033], Mecklenburg et al. 2016:190 [ref. 34440], Kells et al. 2016:351 [ref. 35888], Mecklenburg et al. 2018:360 [ref. 36370], Smirnova et al. 2019:491 [ref. 36948], Love et al. 2021:185 [ref. 39279], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes turneri Bean 1879. Zoarcidae: Lycodinae. Distribution: Arctic and adjacent North Pacific. Habitat: marine.

typica, Lycodes rossi forma Knipowitsch [N. M.] 1906:27 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 19 (no. 1); ref. 15661] Not available, infrasubspecific; publication in an available way as a species or subspecies not investigated [Knipowitsch used v. [variety] and f. [forma] -- in his work, the former is considered subspecific, the latter infrasubspecific; see Article 45(g)]. Not treated by Anderson 1994 [ref. 21438]. Also, typica probably means nominal subspecies, "rossi rossi". •Infrasubspecific, in subfamily Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Infrasubspecific. Zoarcidae: Lycodinae.

typica, Lycodes vahlii forma Smitt [F. A.] 1901:26 [Bihang till Kongl. Svenska vetenskaps-akademiens handlingar. Stockholm v. 27 (no. 4); ref. 18732] Not intended as new; treatment of Lycodes vahlii Reinhardt 1831 as a subspecific name vahlii (i.e., typical) -- see Anderson & Fedorov 2004:48 [ref. 27603]. •In the synonymy of Lycodes pallidus Collett 1879 -- (Møller 2001:983 [ref. 25812]). •In the synonymy of Lycodes vahlii Reinhardt 1831 -- (Carl 2002:69 [ref. 26925] as vahli). •Not available, in family Zoarcidae -- (Anderson & Fedorov 2004:48 [ref. 27603]). Current status: Synonym of Lycodes vahlii Reinhardt 1831. Zoarcidae: Lycodinae.

uschakovi, Lycodes Popov [A. M.] 1931:141, Pl. 2 (fig. 7) [Explorations des mers d'U.R.S.S., Leningrad No. 14; ref. 5702] Okhotsk Sea, 58°11.5'N, 148°19.5'N, depth 141 meters. Holotype (unique): ZIN 29986. Type catalog: Balushkin et al. 2011:1015 [ref. 32449], Balushkin et al. 2012:166 [ref. 33667]. •Valid as Petroschmidtia uschakovi (Popov 1931) -- (Dyldin & Orlov 2021:842 [ref. 38925]). •Valid as Lycodes uschakovi Popov 1931 -- (Lindberg & Krasyukova 1975:147 [ref. 7348], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:205 [ref. 5722], Anderson 1994:119 [ref. 21438], Sokolovskaya et al. 1998:12 [ref. 24670], Nakabo 2000:1037 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Nakabo 2002:1037 [ref. 26193], Fedorov et al. 2003:117 [ref. 27579], Anderson & Fedorov 2004:30 [ref. 27603], Balushkin et al. 2011:1015 [ref. 32449], Nazarkin et al. 2014:192 [ref. 33213], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:423 [ref. 33547]). Current status: Valid as Lycodes uschakovi Popov 1931. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Okhotsk and Japan seas. Habitat: marine.

vachonii, Lycodes Vladykov [V. D.] & Tremblay [J.-L.] 1936:23, Pls. 1 (fig. 3, 4), 2 (fig. 5) [Fauna et flora laurentianae, Station Biologique du Satin-Laurent. Québec. No. 1; ref. 11811] St. Lawrence River estuary, Canada. Syntypes: (5) whereabouts unknown. •Synonym of Lycodes esmarki [sic] Collett 1875, but probably a valid subspecies -- (Andriashev in Whitehead et al. 1986:1138 [ref. 13677]). •Synonym of Lycodes esmarkii Collett 1875 -- (Scott & Scott 1988:407 [ref. 25518] as esmarki, Anderson 1994:118 [ref. 21438], Anderson & Fedorov 2004:24 [ref. 27603], Parin et al. 2014:410 [ref. 33547]). Current status: Synonym of Lycodes esmarkii Collett 1875. Zoarcidae: Lycodinae. Habitat: marine.

vahlii, Lycodes Reinhardt [J. C. H.] 1831:18 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1830-31; ref. 6533] Julianehåb, Greenland. Holotype: ZMUC 9 (skeleton). Type catalog: Nielsen 1974:78 [ref. 9588]. Also appeared in Reinhardt 1837:114, 123, 195, Pl. 5 [32, 41, 113, Pl. 5 of separate] [ref. 3691]. •Valid as Lycodes vahlii Reinhardt 1831 -- (Andriashev 1973:542 [ref. 9214] as vahli, Andriashev in Whitehead et al. 1986:1142 [ref. 13677] as vahli, Scott & Scott 1988:411 [ref. 25518], McAllister 1990:163 [ref. 14674] as vahli, Anderson 1994:119 [ref. 21438], Miki in Okamura et al. 1995:210 [ref. 22531], Møller 2000:847 [ref. 24585], Carl 2002:69 [ref. 26925] as vahli, Anderson & Fedorov 2004:31 [ref. 27603], Nelson et al. 2004:158 [ref. 27807], Møller et al. 2010:63 [ref. 31169] as vahli on p. 61, Balushkin et al. 2011:1016 [ref. 32449], Page et al. 2013:163 [ref. 32708], Afonso et al. 2016:67, 213 [ref. 36199], Mecklenburg et al. 2018:362 [ref. 36370], Simian et al. 2022:376 [ref. 40776], Page et al. 2023:213 [ref. 40505]). Current status: Valid as Lycodes vahlii Reinhardt 1831. Zoarcidae: Lycodinae. Distribution: Arctic Ocean and northwestern Atlantic. Habitat: marine.

yamatoi, Lycodes Toyoshima [M.] 1985:208, Fig. 49 [Memoirs of the Faculty of Fisheries Hokkaido University v. 32 (no. 2); ref. 5722] Sea of Japan off Kasumi, Hyogo Prefecture, Japan. Holotype: HUMZ 41094. Paratypes: HUMZ 42480 (1), 53191 (1), 53597-98 (2), 53610-11 (2), 53618 (1), 53620 (1), 53630 (1), 53634 (1), 53658 (1), 53668 (1), 53672 (1), 53682 (1), 53684 (1), 53726 (1), 53734-35 (2), 53737-38 (2), 53740-42 (3), 53798 (1), 53805 (1), 53817 (1), 53856 (1), 53866 (1), 53911 (1), 53917-18 (2), 53921 (1), 53923 (1). The species matsubarai and yamatoi were described in the same journal article; the first name was kept as the valid one in Parin et al. 2014:415 [ref. 33547], but see matsubarai. •Valid as Lycodes yamatoi Toyoshima 1985 -- (Anderson 1994:119 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Møller 2001:111 [ref. 25378], Nakabo 2002:1039 [ref. 26193], Anderson & Fedorov 2004:31 [ref. 27603], Nazarkin 2010:603 [ref. 31019], Shinohara et al. 2011:52 [ref. 31715], Balushkin et al. 2011:1016 [ref. 32449]). •Synonym of Lycodes matsubarai Toyoshima 1985 -- (Nazarkin & Shinohara 2012:33 [ref. 32106], Parin et al. 2014:415 [ref. 33547]). Current status: Synonym of Lycodes matsubarai Toyoshima 1985. Zoarcidae: Lycodinae. Distribution: Western North Pacific: Okhotsk and Japan seas (if valid). Habitat: marine.

ygreknotatus, Lycodes Schmidt [P. Yu.] 1950:92 [79] [Trudy Tichookeanskogo Komiteta, Akademija Nauk S.S.S.R. v. 6; ref. 12471] Shantar Islands, north of Cape Aleksandir, northwestern part of Okhotsk Sea, Russia, depth 135 meters. Holotype (unique): ZIN 24843. Type catalog: Balushkin et al. 2011:1017 [ref. 32449], Balushkin et al. 2012:167 [ref. 33667]. On p. 79 of English translation. Originally ygrek-notatus. Appeared first without description as Schmidt in Taranetz 1937:163 [ref. 13384]. •Valid as Lycodes ygreknotatus Schmidt 1950 -- (Lindberg & Krasyukova 1975:155 [ref. 7348], Toyoshima in Masuda et al. 1984:308 [ref. 6441], Toyoshima 1985:210 [ref. 5722], Anderson 1994:119 [ref. 21438], Nakabo 2000:1039 [ref. 25182], Sheiko & Fedorov 2000:37 [ref. 25650], Møller 2001:111 [ref. 25378], Nakabo 2002:1039 [ref. 26193], Fedorov et al. 2003:117 [ref. 27579] dated 1937, Anderson & Fedorov 2004:31 [ref. 27603], Nazarkin 2010:9 [ref. 30733], Balushkin et al. 2011:1017 [ref. 32449], Shinohara et al. 2014:266 [ref. 33330], Parin et al. 2014:424 [ref. 33547], Nazarkin et al. 2024:119 [ref. 41547]). Current status: Valid as Lycodes ygreknotatus Schmidt 1950. Zoarcidae: Lycodinae. Distribution: Northwestern Pacific: Sea of Okhotsk and Sea of Japan (Russia, Japan). Habitat: marine.

zoarchus, Lycodes Goode [G. B.] & Bean [T. H.] 1896:308, Pls. 79, 81 (figs. 283) [Special Bulletin U. S. National Museum No. 2; ref. 1848] Off Nova Scotia, Canada, 44°46'30"N, 59°55'45"W, Albatross [station 2499], depth 130 fathoms. Holotype: USNM 39298 (366 mm). Paratypes: USNM 336785 [ex USNM 39298] (3, poor condition), 39299 (1, poor condition). Type catalog: Springer & Anderson 1997:23-24 [ref. 22953]. •Synonym of Lycodes vahlii Reinhardt 1831 -- (Anderson 1994:119 [ref. 21438], Carl 2002:69 [ref. 26925] as vahli, Anderson & Fedorov 2004:31 [ref. 27603]). Current status: Synonym of Lycodes vahlii Reinhardt 1831. Zoarcidae: Lycodinae. Habitat: marine.