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Species in the genus Pungitius: [ 39 ] records

alberti, Pungitius d'Annone [J. J.] (ex Jonston) 1760:302 [Acta Helvetica, Physico-Mathematico-Anatomico-Botanico-Medica (Basile) v. 4; ref. 38366] France, Germany, Italy. No types known. Based on 'De ... Pungitio Alberti' of Jonston 1649:158 [ref. 30579], which was based on the "Pisciculus aculeatus" of Rondelet 1555:206 [ref. 30378]. Composite species, also involving Gasterosteus aculeatus and Pungitius laevis. •Synonym of Pungitius pungitius (Linnaeus 1758). Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

aralensis, Gasterosteus platygaster var. Kessler [K. F.] 1877:4 [The Aralo-Caspian Expedition; ref. 2597] Aral Sea and mouth of Amu Darya River, Uzbekistan. Syntypes: ZIN 3712 (3). •Synonym of Pungitius platygaster (Kessler 1859) -- (Monod 1973:286 [ref. 7193], Keivany & Nelson 2000:117 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856]). •Synonym of Pungitius platygaster (Kessler 1859), but a valid subspecies aralensis Kessler 1877 -- (Berg 1949:970 [ref. 20662], Mitrofanov & Dukravec 1989:114 [ref. 37748], Reshetnikov et al. 1997:747 [ref. 24702], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Jouladeh-Roudbar et al. 2020:286 [ref. 37983], Çiçek et al. 2024:376 [ref. 40789]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish.

blanchardi, Gasterosteus (Gasterostea) Sauvage [H.-E.] 1874:32, Pl. 1 (fig. 16-16a) [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 10; ref. 3904] New York, U.S.A. Syntypes: MNHN 0000-7111 (2) New York, 0000-9383 to 9385 (20); USNM 21139 [ex MNHN] (1). Type catalog: Bertin & Estève 1950:40 [ref. 19576]. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies occidentalis (Cuvier 1829) [unavailable] -- (Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Monod 1973:284 [ref. 7193], Denys et al. 2022:400 [ref. 40085]). Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

brachypoda, Gasterosteus pungitius Bean [T. H.] 1879:129 [Bulletin of the United States National Museum v. 15; ref. 16032] Oosooadlin Mountain, Cumberland Gulf, Greenland, elevation 1500 feet. Syntypes: NMC 58-13 [ex USNM 21773 and Victoria Mem. Mus. 154] (1); USNM 21738 (1), 21766 (1), 21768 (1), 21771 (1), 21773 (1), 21773a (1), 21773c (1), 21773d (1). Type catalog: McAllister 1965:10 [ref. 22019]. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies pungitius -- (Okada 1961:616 [ref. 6363]). •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies occidentalis (Cuvier 1829) [unavailable] -- (Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Berg 1949:966 [ref. 20662], Monod 1973:284 [ref. 7193], Denys et al. 2022:400 [ref. 40085]). Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

breviceps, Gasterosteus Blanchard [C. E.] 1866:245, Fig. 43 [Les poissons des eaux douces de la France; ref. 310] Caen, Normandy, France. Holotype (unique): MNHN 0000-7100. Type catalog: Bertin & Estève 1950:39 [ref. 19576]. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies laevis (Cuvier 1829) -- (Monod 1973:285 [ref. 7193], Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Paepke in Bănărescu & Paepke 2002:277 [ref. 25856]). •Synonym of Pungitius laevis (Cuvier 1829) -- (Kottelat 1997:166 [ref. 22952], Denys et al. 2017:17 [ref. 35585]). Current status: Synonym of Pungitius laevis (Cuvier 1829). Gasterosteidae. Habitat: freshwater, brackish, marine.

brevispinosus, Pungitius Otaki [K.] 1908:87 [Transactions of the Sapporo Natural History Society v. 2 (pts 1-2); ref. 18503] Sapporo, Hokkaido, Japan. Syntypes: whereabouts unknown. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies pungitius -- (Okada 1961:615 [ref. 6363]). •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies sinensis (Guichenot 1869) -- (Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius sinensis (Guichenot 1869) -- (Dyldin & Orlov 2017:58 [ref. 35138]). Current status: Synonym of Pungitius sinensis (Guichenot 1869). Gasterosteidae. Habitat: freshwater, brackish, marine.

burgundianus, Gasterosteus Blanchard [C. E.] 1866:240, Fig. 40 [Les poissons des eaux douces de la France; ref. 310] Dijon, Côte d'Or, France. Syntypes: MNHN 0000-7103 [or 7109] (5). Type catalog: Bertin & Estève 1950:39 [ref. 19576]. •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Monod 1973:284 [ref. 7193], Paepke in Bănărescu & Paepke 2002:277 [ref. 25856], Denys et al. 2017:14 [ref. 35585], Denys et al. 2022:399 [ref. 40085]). •Synonym of Pungitius laevis (Cuvier 1829) -- (Kottelat 1997:166 [ref. 22952]). •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies laevis (Cuvier 1829) -- (Keivany & Nelson 2000:118 [ref. 24720]). Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

bussei, Gasterosteus Warpachowski [N. A.] in Warpachowski & Herzenstein 1888:13, Pl. (fig. 1) [Trudy St.-Peterburgskogo Obscestva Estestvoispytatelej = Travaux de la Société des Naturalistes de St. Pétersbourg. v. 19 (no. 8) (for 1887); ref. 2152] Ilistaya [Lefu] River, tributary to Khanka Lake, Russia. Syntypes: USNM 57470 (1), ZIN 7100 (4, now 3). •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies sinensis (Guichenot 1869) -- (Berg 1949:967 [ref. 20662], Okada 1961:621 [ref. 6363], Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius sinensis (Guichenot 1869) -- (Bogutskaya & Naseka 1996:57 [ref. 22798], Reshetnikov & Savvaitove 1998:112 [ref. 23593]). •Valid as Pungitius bussei (Warpachowski 1888) -- (Bogutskaya et al. 2001:46 [ref. 26178], Bogutskaya & Naseka 2004:181 [ref. 28183] dated 1887, Takahashi et al. 2016:49 [ref. 34337], Dyldin et al. 2020:114 [ref. 37569], Matsumoto et al. 2021:14 [ref. 38508], Wang et al. 2021:[3] [ref. 38800]). Current status: Valid as Pungitius bussei (Warpachowski 1888). Gasterosteidae. Distribution: East Asia: Amur River basin (Russia). Habitat: freshwater.

carinata, Pygosteus pungitius forma Bertin [L.] 1925:122 [Annales de l'Institut Océanographique Monaco N. S. v. 2 (fasc. 1); ref. 23054] Not available, infrasubspecific; as a forme. •In the synonymy of Pungitius pungitius (Linnaeus 1758) -- (Kottelat 1997:167 [ref. 22952], Paepke in Bănărescu & Paepke 2002:277 [ref. 25856], Denys et al. 2022:400 [ref. 40085]). Infrasubspecific. Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae.

caucasicus, Gasterosteus platygaster var. Kessler [K. F.] 1877:3 [The Aralo-Caspian Expedition; ref. 2597] Transcaucasia. Syntypes: ZIN 2401 (12+). •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Kottelat 1997:166 [ref. 22952], Keivany & Nelson 2000:116 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Parin et al. 2014:205 [ref. 33547], Jouladeh-Roudbar et al. 2020:286 [ref. 37983], Çiçek et al. 2024:376 [ref. 40789]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

concinnus, Gasterosteus Richardson [J.] 1836:57 [Fauna Boreali-Americana Part 3; ref. 3731] Saskatchewan (53°N) to Great Bear Lake (65°N), Canada. Syntypes: (many) whereabouts unknown. •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Monod 1973:281 [ref. 7193]). •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies occidentalis (Cuvier 1829) [unavailable] -- (Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Monod 1973:284 [ref. 7193], Scott & Crossman 1998:674 [ref. 24570], Denys et al. 2022:399 [ref. 40085]). Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

danubica, Gasterosteus platygaster var. Steindachner [F.] 1899:542 [4] [Sitzungsberichte der Mathematisch-Naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften v. 108 (pt. 1) (no. 7) (for July 1899); ref. 10017] Danube swamps near Negotin, Srebia; Save [Sava] River at Belgrad, Serbia. Syntypes: NMW 6823 (6). On p. 4 of separate. •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Kottelat 1997:166 [ref. 22952], Keivany & Nelson 2000:117 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Jouladeh-Roudbar et al. 2020:286 [ref. 37983]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

dekayi, Gasterosteus Agassiz [L.] 1850:311 [Lake Superior; ref. 66] New York and Massachusetts, U.S.A. No types known. Based on literature sources. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies occidentalis (Cuvier 1829) [unavailable] -- (Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Monod 1973:284 [ref. 7193], Denys et al. 2022:399 [ref. 40085]). Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

globiceps, Gasterosteus Sauvage [H.-E.] 1874:35, Pl. 1 (fig. 17) [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 10; ref. 3904] Stated as North America but apparently erroneous. Syntypes: MNHN 0000-4081 (2). Type catalog: Bertin & Estève 1950:40 [ref. 19576]. •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Monod 1973:284 [ref. 7193], Paepke in Bănărescu & Paepke 2002:277 [ref. 25856] with question). •Tentatively a synonym of Pungitius laevis (Cuvier 1829) -- (Kottelat 1997:166 [ref. 22952]). •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies occidentalis (Cuvier 1829) [unavailable] -- (Keivany & Nelson 2000:118 [ref. 24720]). Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

hellenicus, Pungitius Stephanidis [A.] 1971:228, Figs. 7-8 [Biologia Gallo-Hellenica v. 3 (no. 2); ref. 7539] Sperchios River Valley, Greece. Syntypes: (40) MNHN 1975-0867 (8). •Synonym of Pungitius pungitius (Linnaeus 1758), but a valid subspecies -- (Lelek 1987:267 [ref. 22962]). •Valid as Pungitius hellenicus Stephanidis 1971 -- (Economidis 1995:206 [ref. 23025], Keivany et al. 1997:558 [ref. 22904], Kottelat 1997:166 [ref. 22952], Keivany & Nelson 1998:458 [ref. 24681], Keivany & Nelson 2000:116 [ref. 24720], McLennan & Mattern 2001:11 [ref. 25222], Paepke in Bănărescu & Paepke 2002:261 [ref. 25856], Kottelat & Freyhof 2007:496 [ref. 29996], Barbieri et al. 2015:97 [ref. 35029], Matsumoto et al. 2021:17 [ref. 38508], Wang et al. 2021:[3] [ref. 38800]). Current status: Valid as Pungitius hellenicus Stephanidis 1971. Gasterosteidae. Distribution: Europe: central Greece. Habitat: freshwater.

hologymna, Pygosteus pungitius forma Bertin [L.] 1925:122 [Annales de l'Institut Océanographique Monaco N. S. v. 2 (fasc. 1); ref. 23054] Not available, infrasubspecific; as a forma. •In the synonymy of Pungitius laevis (Cuvier 1829) -- (Kottelat 1997:166 [ref. 22952]). •In the synonymy of Pungitius pungitius (Linnaeus 1758) -- (Paepke in Bănărescu & Paepke 2002:277 [ref. 25856]). Infrasubspecific. Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae.

japonicus, Gasterosteus Steindachner [F.] 1881:264 [27], Pl. 3 (fig. 2) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 82 (1. Abth., no. 2); ref. 4230] Gulf of Strielok near Vladivostok, Sea of Japan. Syntypes: (several) MSNG 14075 (1, not found in 1995). Type catalog: Tortonese 1963:312 [ref. 10351]. On p. 27 of separate. Objectively invalid; preoccupied by Gasterosteus japonicus Houttuyn 1782, replaced by Gasterosteus steindachneri Jordan & Snyder 1900. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies sinensis (Guichenot 1869) -- (Berg 1949:967 [ref. 20662], Okada 1961:621 [ref. 6363], Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitis sinensis (Guichenot 1869) -- (Bogutskaya et al. 2008:350 [ref. 30085], Parin et al. 2014:206 [ref. 33547], Dyldin & Orlov 2017:58 [ref. 35138]). 1°Homonym. Current status: Synonym of Pungitius sinensis (Guichenot 1869). Gasterosteidae. Habitat: freshwater, brackish, marine.

kaibarae, Pygosteus Tanaka [S.] 1915:565 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 27 (no. 325); ref. 4324] Kichisho-in, southwest of Kyoto, Japan. Lectotype: ZUMT 8197. Paralectotypes: ZUMT 59848-54 (7). Lectotype established by Tanaka 1918:491 [ref. 14911], as ZUMT 8197 (identity of specimen not researched). •Synonym of Pungitius sinensis (Guichenot 1869), but a valid subspecies -- (Ida in Masuda et al. 1984:84 [ref. 6441]). •See Kim 1997:392 [ref. 24032]. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies sinensis (Guichenot 1869) -- (Okada 1961:622 [ref. 6363], Keivany & Nelson 2000:118 [ref. 24720]). •Valid as Pungitius kaibarae (Tanaka 1915) -- (Nakabo 2000:514 [ref. 25086], Kim & Park 2002:314 [ref. 25913], Nakabo 2002:514 [ref. 26001], Choi et al. 2003:231, 553 [ref. 26218], Jang et al. 2003:120 [ref. 27681], Bogutskaya et al. 2008:349 [ref. 30085], Parin et al. 2014:204 [ref. 33547], Takahashi et al. 2016:49 [ref. 34337], Matsumoto et al. 2021:16 [ref. 38508], Wang et al. 2021:[3] [ref. 38800]). Current status: Valid as Pungitius kaibarae (Tanaka 1915). Gasterosteidae. Distribution: Northwestern Pacific: Sea of Japan (Hyogo, Kyoto and Osaka prefectures, Japan and eastern side of Korean Peninsulae). IUCN (2021): Not assessed. Habitat: freshwater, brackish.

kessleri, Gasterosteus pungitius var. Yakovlev [V. E.] 1870:110 [10] [On the new and little-known species of fish found in the mouths of the Volga; ref. 19610] Lakes near Astrakhan, Volga River Delta, Russia. Syntypes: whereabouts unknown. On p. 10 of separate. •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Coad 1995:27 [ref. 23608], Kottelat 1997:166 [ref. 22952], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Keivany & Nelson 2000:116 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Parin et al. 2014:205 [ref. 33547], Jouladeh-Roudbar et al. 2020:286 [ref. 37983]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

laevis, Gasterosteus Cuvier [G.] 1829:170 [Le Règne Animal (Edition 2) v. 2; ref. 995] Seine basin at Bobigny near Paris, France. Lectotype: MNHN 2016-0604 [ex 0000-7103]. Paralectotypes: MNHN 0000-7102 (5), 0000-7103 (2). Lectotype selected by Denys et al. 2017:17 [ref. 35585]. •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Paepke in Bănărescu & Paepke 2002:277 [ref. 25856]). •Synonym of Pungitius pungitius (Linnaeus 1758), but a valid subspecies -- (Monod 1973:284 [ref. 7193], Reshetnikov et al. 1997:747 [ref. 24702], Keivany & Nelson 2000:118 [ref. 24720]). •Valid as Pungitius laevis (Cuvier 1829) -- (Berg 1949:969 [ref. 20662], Kottelat 1997:166 [ref. 22952], Kottelat & Freyhof 2007:497 [ref. 29996], Denys et al. 2017:17 [ref. 35585], Matsumoto et al. 2021:9 [ref. 38508], Wang et al. 2021:[3] [ref. 38800]). Current status: Valid as Pungitius laevis (Cuvier 1829). Gasterosteidae. Distribution: Western Europe: coastal Atlantic basins (Seine, Scheldt, Loire) of France. Habitat: freshwater.

lotharingus, Gasterosteus Blanchard [C. E.] 1866:244, Fig. 42 [Les poissons des eaux douces de la France; ref. 310] Meuse River and adjacent brooks, near Saint-Mihiel, France. Syntypes: MNHN 0000-7090 (2). Type catalog: Bertin & Estève 1950:39 [ref. 19576]. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies laevis (Cuvier 1829) -- (Monod 1973:284 [ref. 7193], Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Paepke in Bănărescu & Paepke 2002:277 [ref. 25856]). •Synonym of Pungitius laevis (Cuvier 1829) -- (Berg 1949:969 [ref. 20662], Kottelat 1997:166 [ref. 22952], Denys et al. 2017:17 [ref. 35585]). Current status: Synonym of Pungitius laevis (Cuvier 1829). Gasterosteidae. Habitat: freshwater, brackish, marine.

mainensis, Gasterosteus Storer [D. H.] 1837:465 [Boston Journal of Natural History v. 1 (no. 4) (art. 23); ref. 18821] Kennebeck County, Maine, U.S.A. No types known. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies occidentalis (Cuvier 1829) [unavailable] -- (Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Monod 1973:284 [ref. 7193], Denys et al. 2022:399 [ref. 40085]). Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

modestus, Pungitius Matsumoto [T.], Matsuura [K.] & Hanzawa [N.] 2021:3, Figs. 2-4 [Zootaxa 5005 (no. 1); ref. 38508] Tributary of the Mogami-gawa River, Tendo, Yamagata Prefecture, Japan. Holotype: NSMT-P 133674. Paratypes: (17) NSMT. Plus additional non-type material. •Valid as Pungitius modestus Matsumoto, Matsuura & Hanzawa 2021. Current status: Valid as Pungitius modestus Matsumoto, Matsuura & Hanzawa 2021. Gasterosteidae. Distribution: Yamagata Prefecture, northern Honshu, Japan. Habitat: freshwater.

nebulosus, Gasterosteus Agassiz [L.] 1850:310, Pl. 4 (fig. 2) [not fig. 4] [Lake Superior; ref. 66] Mouth of Pic River, Lake Superior. Syntypes: MCZ 6696 (10, now 8), ?MCZ 58765 (2), USNM 120414 [ex MCZ 6696] (2). •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies occidentalis (Cuvier 1829) [unavailable] -- (Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Monod 1973:284 [ref. 7193], Denys et al. 2022:399 [ref. 40085]). Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

niger, Gasterosteus pungitius var. Yakovlev [V. E.] 1870:110 [10] [On the new and little-known species of fish found in the mouths of the Volga; ref. 19610] Lakes near Astrakhan, Volga River Delta, Russia. Syntypes: whereabouts unknown. On p. 10 of separate. Objectively invalid; preoccupied by Gasterosteus niger Cuvier 1829. •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Coad 1995:27 [ref. 23608], Kottelat 1997:166 [ref. 22952], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Keivany & Nelson 2000:116 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Parin et al. 2014:205 [ref. 33547], Jouladeh-Roudbar et al. 2020:286 [ref. 37983]). 1°Homonym. Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

nuda, Pygosteus platygaster Berg [L. S.] (ex Nikolskii) 1905:218 [Izviestii Turkestanskago otdiela Russkago geograficheskago obschchestva v. 4; ref. 15338] Lake Charkal, Ural River basin, Russia. Syntypes: ZIN 10613. •Synonym of Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Kottelat 1997:167 [ref. 22952], Keivany & Nelson 2000:117 [ref. 24720], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Parin et al. 2014:205 [ref. 33547], Jouladeh-Roudbar et al. 2020:286 [ref. 37983]). Current status: Synonym of Pungitius platygaster (Kessler 1859). Gasterosteidae. Habitat: freshwater, brackish, marine.

occidentalis, Gasterosteus Cuvier [G.] in Cuvier & Valenciennes 1829:509 [Histoire naturelle des poissons v. 4; ref. 998] Newfoundland. Holotype (unique): MNHN 0000-7112. Type catalog: Bertin & Estève 1950:39 [ref. 19576]. See Mecklenburg et al. 2002:334 [ref. 25968]. •This name is preoccupied by Gasterosteus occidentalis Linnaeus 1758 and should not be used. •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Monod 1973:284 [ref. 7193], Smith 1986:278 [ref. 22186]). •Synonym of Pungitius pungitius (Linnaeus 1758), but a valid subspecies -- (Keivany & Nelson 2000:118 [ref. 24720], Krueger in Collette & Klein-MacPhee 2002:319 [ref. 26158], Scharpf 2007:35 [ref. 30398], Denys et al. 2022:399 [ref. 40085]). 1°Homonym. Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish.

platygaster, Gasterosteus Kessler [K. F.] 1859:202 [Bulletin de la Société Impériale des Naturalistes de Moscou v. 32 (no. 3); ref. 18391] Odessa and side arm of Dnieper River in Aleschki, Ukraine. Syntypes: BMNH 1897.7.5.2 [ex ZIN] (2), ZIN 2350-51 (6, 6+). Additional material: (14) ZIN 2350-51 (6, 6+) Dnieper River near Aleschki, Ukraine. Also appeared in Kessler 1860:17, Pl. 1 (fig. 1) [ref. 18393]. •May be a subspecies of Pungitius pungitius (Linnaeus 1758) -- (Coad 1995:27 [ref. 23608]). •Valid as Pungitius platygaster (Kessler 1859), subspecies platygaster Kessler 1859 -- (Mitrofanov & Dukravec 1989:112 [ref. 37748]). •Valid as Pungitius platygaster (Kessler 1859) -- (Berg 1949:969 [ref. 20662], Monod 1973:285 [ref. 7193], Coad 1981:16 [ref. 22348], Banister in Whitehead et al. 1986:642 [ref. 13676], Coad 1995:26 [ref. 23608], Economidis 1995:208 [ref. 23025], Keivany et al. 1997:558 [ref. 22904], Kottelat 1997:166 [ref. 22952], Reshetnikov et al. 1997:747 [ref. 24702], Reshetnikov & Savvaitove 1998:111 [ref. 23593], Coad 1998:103 [ref. 23869], Keivany & Nelson 1998:458 [ref. 24681], Keivany & Nelson 2000:116 [ref. 24720], McLennan & Mattern 2001:11 [ref. 25222], Bogutskaya et al. 2001:46 [ref. 26178], Bilecenoğlu et al. 2002:64 [ref. 26753], Paepke in Bănărescu & Paepke 2002:267 [ref. 25856], Bogutskaya & Naseka 2004:181 [ref. 28183], Vassilev & Pehlivanov 2005:175 [ref. 28369], Fricke et al. 2007:73 [ref. 29533], Kottelat & Freyhof 2007:498 [ref. 29996], Vasil'eva 2007:71 [ref. 30517], Esmaeili et al. 2010:375 [ref. 33391], Parin et al. 2014:205 [ref. 33547] as subspecies platycephalus, Jouladeh-Roudbar et al. 2015:895 [ref. 34062], Barbieri et al. 2015:97 [ref. 35029], Çiçek et al. 2015:151 [ref. 35151], Takahashi et al. 2016:49 [ref. 34337], Esmaeili et al. 2017:95 [ref. 35254], Esmaeili et al. 2018:77 [ref. 36089], Artüz & Fricke 2019:555 [ref. 36612], Jouladeh-Roudbar et al. 2020:286 [ref. 37983], Kuljanishvili et al. 2020:[5] [ref. 37535], Matsumoto et al. 2021:9 [ref. 38508], Wang et al. 2021:[3] [ref. 38800], Çiçek et al. 2023:55 [ref. 40326], Çiçek et al. 2024:376 [ref. 40789]). Current status: Valid as Pungitius platygaster (Kessler 1859). Gasterosteidae. Distribution: Eurasia: Black, Caspian and Aral Sea basins. Habitat: freshwater, brackish.

polyakovi, Pungitius Shedko [S. V.], Shedko [M. B.] & Pietsch [T. W.] 2005:223, Figs. 1-2 [Flora and Fauna of Sakhalin Island; ref. 29561] 46°47.035'N, 143°47.337'E, southeastern Sakhalin Island, Russia. Holotype: ZIN 53441. Paratypes: ZIN 53442 (25). •Possibly the same as Pungitius tymensis (Nikolskii 1889) -- (Dyldin & Orlov 2017:57 [ref. 35138]). •Valid as Pungitius polyakovi Shedko, Shedko & Pietsch 2005 -- (Takahashi et al. 2016:49 [ref. 34337], Dyldin & Orlov 2017:57 [ref. 35138], Dyldin et al. 2020:108, 114 [ref. 37569], Dyldin et al. 2021:58 [ref. 38804], Matsumoto et al. 2021:18 [ref. 38508], Wang et al. 2021:[3] [ref. 38800], Dyldin & Orlov 2022:35 [ref. 38972]). Current status: Valid as Pungitius polyakovi Shedko, Shedko & Pietsch 2005. Gasterosteidae. Distribution: East Asia: southeastern Sakhalin Island (Russia). Habitat: freshwater, brackish.

pungitius, Gasterosteus Linnaeus [C.] 1758:296 [Systema Naturae, Ed. X v. 1; ref. 2787] Europe. Syntypes: LS 34-35 (2, left half-skins). Type catalog: Wheeler 1985:44 [ref. 20705]. •Valid as Pungitius pungitius (Linnaeus 1758), subspecies pungitius -- (Okada 1961:615 [ref. 6363], Kawanabe & Mizuno 1989:445 [ref. 25598], Sheiko & Fedorov 2000:25 [ref. 25650], Pietsch et al. 2001:144 [ref. 25688], Dolgov 2013:917 [ref. 33112]). •Valid as Pungitius pungitius (Linnaeus 1758) -- (Berg 1949:965 [ref. 20662], Monod 1973:284 [ref. 7193], Lee et al. 1980:566 [ref. 22416], Ida in Masuda et al. 1984:83 [ref. 6441], Smith 1986:277 [ref. 22186], Banister in Whitehead et al. 1986:642 [ref. 13676], Robins & Ray 1986:122 [ref. 23100], Lelek 1987:271 [ref. 22962], Scott & Scott 1988:343 [ref. 25518], Mitrofanov & Dukravec 1989:111 [ref. 37748], McAllister 1990:219 [ref. 14674], Page & Burr 1991:242 [ref. 18983], Haglund et al. 1992 [ref. 20881], Chereshnev 1996:605 [ref. 24659], Keivany et al. 1997:558 [ref. 22904], Kottelat 1997:167 [ref. 22952], Kim 1997:391 [ref. 24032], Reshetnikov et al. 1997:747 [ref. 24702], Reshetnikov & Savvaitove 1998:112 [ref. 23593], Scott & Crossman 1998:671 [ref. 24570], Keivany & Nelson 1998:458 [ref. 24681] with subspecies, Fuller et al. 1999:336 [ref. 25838], Keivany & Nelson 2000:117 [ref. 24720], Lyons et al. 2000:42 [ref. 26594], Matsuura et al. 2000:197 [ref. 26000], Nakabo 2000:514 [ref. 25086], Novikov et al. 2000:161 [ref. 40929], Zemnukhov et al. 2001:420 [ref. 25412], Bogutskaya et al. 2001:46 [ref. 26178], Chereshnev et al. 2001:113 [ref. 26736], Mecklenburg et al. 2002:334 [ref. 25968], Paepke in Bănărescu & Paepke 2002:277 [ref. 25856], Kim & Park 2002:313 [ref. 25913], Nakabo 2002:514 [ref. 26001], Choi et al. 2003:231, 554 [ref. 26218], Kolpakov 2002:708 [ref. 26469], Kolpakov 2003:847 [ref. 27537], Fedorov et al. 2003:57 [ref. 27579], Pichugin et al. 2004:15 [ref. 27623], Alekseyev et al. 2004:54 [ref. 27625], Bailey et al. 2004:169 [ref. 27882], Nelson et al. 2004:111 [ref. 27807], Bogutskaya & Naseka 2004:181 [ref. 28183], Love et al. 2005:71 [ref. 37547], Fricke 2007:28 [ref. 30577], Kottelat & Freyhof 2007:498 [ref. 29996], Scharpf 2007:34 [ref. 30398], McPhail 2007:440 [ref. 31199], Bogutskaya et al. 2008:350 [ref. 30085], Mecklenburg et al. 2011:121 [ref. 31212], Page & Burr 2011:462 [ref. 31215], Page et al. 2013:113 [ref. 32708], Parin et al. 2014:205 [ref. 33547], Afonso et al. 2016:39, 259 [ref. 36199], Kells et al. 2016:122 [ref. 35888], Takahashi et al. 2016:49 [ref. 34337], Dyldin & Orlov 2017:57 [ref. 35138], Denys et al. 2017:14 [ref. 35585], Romanov et al. 2017:825 [ref. 35687], Dolgov et al. 2018:109 [ref. 41055], Artüz & Fricke 2019:555 [ref. 36612], Dyldin et al. 2020:85 [ref. 37569], Love et al. 2021:102 [ref. 39279], Matsumoto et al. 2021:8 [ref. 38508], Wang et al. 2021:[3] [ref. 38800], Denys et al. 2022:399 [ref. 40085], Dyldin & Orlov 2022:35 [ref. 38972], Simian et al. 2022:375 [ref. 40776], Page et al. 2023:216 [ref. 40505]). Current status: Valid as Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Distribution: Central Europe. Habitat: freshwater, brackish.

semiarmata, Pygosteus pungitius forma Bertin [L.] 1925:122 [Annales de l'Institut Océanographique Monaco N. S. v. 2 (fasc. 1); ref. 23054] Not available, infrasubspecific; as a forme. •In the synonymy of Pungitius pungitius (Linnaeus 1758) -- (Kottelat 1997:167 [ref. 22952], Paepke in Bănărescu & Paepke 2002:277 [ref. 25856], Denys et al. 2022:400 [ref. 40085]). Infrasubspecific. Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish, marine.

sinensis, Gasterosteus Guichenot [A.] 1869:204, Pl. 12 (fig. 4) [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 5 (fasc. 3); ref. 1952] Chang Jiang [Yangtze River] (?), China. Syntypes: MNHN 0000-5228 (6). Type catalog: Bertin & Estève 1950:39 [ref. 19576]. •Synonym of Pungitius pungitius (Linnaeus 1758), but a valid subspecies -- (Berg 1949:967 [ref. 20662], Okada 1961:621 [ref. 6363], Monod 1973:285 [ref. 7193], Kawanabe & Mizuno 1989:444 [ref. 25598], Keivany & Nelson 2000:118 [ref. 24720], Matsuura et al. 2000:198 [ref. 26000], Pietsch et al. 2001:144 [ref. 25688]). •Valid as Pungitius sinensis (Guichenot 1869) -- (Ida in Masuda et al. 1984:84 [ref. 6441], Wang 1984:88 [ref. 22951], Zhu 1995:165 [ref. 25213], Bogutskaya & Naseka 1996:56 [ref. 22798], Chereshnev 1996:605 [ref. 24659], Cheng & Zhou 1997:234 [ref. 26385], Kim 1997:388 [ref. 24032], Nikiforov et al. 1997:330 [ref. 24694], Reshetnikov et al. 1997:747 [ref. 24702], Reshetnikov & Savvaitove 1998:112 [ref. 23593], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:514 [ref. 25086], Bogutskaya et al. 2001:46 [ref. 26178], Chereshnev et al. 2001:115 [ref. 26736], Wang et al. 2001:196 [ref. 26566], Zemnukhov et al. 2001:420 [ref. 25412], Kim & Park 2002:310 [ref. 25913], Chereshnev 2002:133 [ref. 25962], Choi et al. 2003:231, 553 [ref. 26218], Fedorov et al. 2003:58 [ref. 27579], Pichugin et al. 2004:15 [ref. 27623], Alekseyev et al. 2004:54 [ref. 27625], Bogutskaya & Naseka 2004:182 [ref. 28183], Bogutskaya et al. 2008:350 [ref. 30085], Parin et al. 2014:206 [ref. 33547], Takahashi et al. 2016:49 [ref. 34337], Zhang et al. 2016:192 [ref. 34477], Dyldin & Orlov 2017:58 [ref. 35138], Dyldin et al. 2020:85 [ref. 37569], Dyldin et al. 2021:58 [ref. 38804], Matsumoto et al. 2021:9 [ref. 38508], Wang et al. 2021:[3] [ref. 38800], Dyldin & Orlov 2022:36 [ref. 38972]). Current status: Valid as Pungitius sinensis (Guichenot 1869). Gasterosteidae. Distribution: Northwestern Pacific: Kamchatka (Russia) south to Amur River and Tumannaya River (Tumen, China), including Kuril Islands. Habitat: freshwater, brackish, marine.

steindachneri, Pygosteus Jordan [D. S.] & Snyder [J. O.] 1901:747 [Proceedings of the United States National Museum v. 23 (no. 1235); ref. 2504] Replacement name for Gasterosteus japonicus Steindachner 1881, preoccupied by Gasterosteus japonicus Houttuyn 1782. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies sinensis (Guichenot 1869) -- (Okada 1961:621 [ref. 6363], Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius sinensis (Guichenot 1869) -- (Bogutskaya et al. 2008:350 [ref. 30085]). Replacement Name. Current status: Synonym of Pungitius sinensis (Guichenot 1869). Gasterosteidae. Habitat: freshwater, brackish, marine.

stenurus, Gasterosteus Kessler [K. F.] 1876:6, Pl. 3 (fig. 6) [v. 2 (pt 4); ref. 2595] Hu-lun [Dalai-Nor] Lake, southeastern Mongolia, China, elevation 4200 feet. Syntypes: (8) BMNH 1897.7.5.3 [ex ZIN] (1), ZIN 2471 (6). •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies sinensis (Guichenot 1869) -- (Berg 1949:967 [ref. 20662], Keivany & Nelson 2000:118 [ref. 24720]). •May be a distinct species -- (Bogutskaya et al. 2008:351 [ref. 30085], Matsumoto et al. 2021:10 [ref. 38508]) Current status: Valid as Pungitius stenurus (Kessler 1876). Gasterosteidae. Distribution: Northwest China. Habitat: freshwater.

trachura, Pygosteus pungitius forma Bertin [L.] 1925:122 [Annales de l'Institut Océanographique Monaco N. S. v. 2 (fasc. 1); ref. 23054] Not available, infrasubspecific; as a forme. •In the synonymy of Pungitius pungitius (Linnaeus 1758) -- (Kottelat 1997:167 [ref. 22952], Denys et al. 2022:400 [ref. 40085]). Infrasubspecific. Current status: Synonym of Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Habitat: freshwater, brackish, marine.

tymensis, Gasterosteus Nikolskii [A. M.] 1889:293 [Sakhalin Island and its fauna of vertebarate animals; ref. 14493] Tym River, Sakhalin Island, Russia. Syntypes: (many) BMNH 1892.4.28.1 [ex ZIN]; ZIN 6593-95 (6+, 6+, 6+). Authorship sometimes as Nikolsky. •Synonym of Pungitius pungitius (Linnaeus 1758), but a valid subspecies -- (Okada 1961:619 [ref. 6363], Monod 1973:285 [ref. 7193], Ida in Masuda et al. 1984:83 [ref. 6441], Nikiforov et al. 1997:330 [ref. 24694], Keivany & Nelson 2000:119 [ref. 24720], Chereshnev 2002:704 [ref. 26468], Kolpakov 2003:847 [ref. 27537], Pichugin et al. 2004:15 [ref. 27623]). •Valid as Pungitius tymensis (Nikolskii 1889) -- (Berg 1949:968 [ref. 20662], Kawanabe & Mizuno 1989:442 [ref. 25598], Kim 1997:390 [ref. 24032], Reshetnikov et al. 1997:748 [ref. 24702], Reshetnikov & Savvaitove 1998:112 [ref. 23593], Nakabo 2000:514 [ref. 25086], Matsuura et al. 2000:199 [ref. 26000], McLennan & Mattern 2001:11 [ref. 25222], Pietsch et al. 2001:144 [ref. 25688], Kim & Park 2002:312 [ref. 25913], Bogutskaya et al. 2001:46 [ref. 26178], Nakabo 2002:514 [ref. 26001], Choi et al. 2003:231, 554 [ref. 26218], Bogutskaya & Naseka 2004:182 [ref. 28183], Takahashi et al. 2016:49 [ref. 34337], Dyldin & Orlov 2017:58 [ref. 35138], Dyldin et al. 2021:59 [ref. 38804], Matsumoto et al. 2021:15 [ref. 38508], Wang et al. 2021:[3] [ref. 38800], Dyldin & Orlov 2022:36 [ref. 38972]). Current status: Valid as Pungitius tymensis (Nikolskii 1889). Gasterosteidae. Distribution: East Asia: Russia and Japan. Habitat: freshwater.

undecimalis, Pygosteus Jordan [D. S.] & Starks [E. C.] 1902:62, Fig. 1 [Proceedings of the United States National Museum v. 26 (no. 1308); ref. 2525] Chitose, Hokkaido, Japan. Holotype: CAS-SU 7119. Paratypes: CAS-SU 7153 (5). Type catalog: Böhlke 1953:59 [ref. 12291]. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies tymensis (Nikolskii 1889) -- (Okada 1961:619 [ref. 6363], Monod 1973:285 [ref. 7193], Keivany & Nelson 2000:119 [ref. 24720]). •Synonym of Pungitius tymensis (Nikolskii 1889) -- (Berg 1949:968 [ref. 20662], Dyldin & Orlov 2017:58 [ref. 35138]). Current status: Synonym of Pungitius tymensis (Nikolskii 1889). Gasterosteidae. Habitat: freshwater.

vulgaris, Gasterosteus Mauduyt [F.] 1849:19 [Bulletin Société Académique d'Agriculture, Belles-Lettres, Sciences et Arts de Poitiers 1848, v. 3, fasc. 9-12; ref. 17615] Belle at Puy-Rabier, France. Neotype: UCBLZ 2012.9.1010. Neotype designated by Denys et al. 2017:19 [ref. 35585]. •Synonym of Pungitius laevis (Cuvier 1829) -- (Kottelat 1997:166 [ref. 22952]). •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies laevis (Cuvier 1829) -- (Keivany & Nelson 2000:118 [ref. 24720]). •Synonym of Pungitius pungitius (Linnaeus 1758) -- (Paepke in Bănărescu & Paepke 2002:277 [ref. 25856]). •Valid as Pungitius vulgaris (Mauduyt 1849) -- (Denys et al. 2017:18 [ref. 35585], Matsumoto et al. 2021:9 [ref. 38508]). Current status: Valid as Pungitius vulgaris (Mauduyt 1849). Gasterosteidae. Distribution: South-western France: Loire, Gironde and Dordogne basins. Habitat: freshwater, brackish.

wosnesenjenskyi, Gasterosteus Kessler [K. F.] 1876:9 [v. 2 (pt 4); ref. 2595] West coast of Kamchatka, Russia. Syntypes: ZIN 2377-78 (6, 6+) Spellings in literature differ some. •Synonym of Pungitius pungitius (Linnaeus 1758), subspecies sinensis (Guichenot 1869) -- (Berg 1949:967 [ref. 20662] as wosnessenskyi, Okada 1961:622 [ref. 6363] as wossnessenskyi, Keivany & Nelson 2000:118 [ref. 24720]). •Questionably a synonym of Pungitius pungitius subspecies pungitius (Linnaeus 1758) -- (Sheiko & Fedorov 2000:25 [ref. 25650] as wosnessenskyi). •Synonym of Pungitius sinensis (Guichenot 1869) -- (Bogutskaya et al. 2008:350 [ref. 30085], Dyldin & Orlov 2017:58 [ref. 35138]). Current status: Synonym of Pungitius sinensis (Guichenot 1869). Gasterosteidae. Habitat: freshwater, brackish, marine.