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Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information. Species in the genus Squalidus: [ 33 ] records argentatus, Gobio Sauvage [H.-E.] & Dabry de Thiersant [P.] 1874:9 [Annales des Sciences Naturelles, Paris (Zoologie et Paléontologie) (Sér. 6) v. 1 (art. 5); ref. 15665] Chang Jiang [Yangtze River], China. Holotype: MNHN 0000-5038. Paratypes: MNHN 0000-2064 (1), 0000-5039 (1). Type catalog: Bertin & Estève 1948:86-87 [ref. 19574]. •Valid as Gnathopogon argentatus (Sauvage & Dabry de Thiersant 1874) -- (Luo et al. 1977:487 [ref. 2850]). •Questionably the same as Squalidus mantschuricus (Mori 1927) -- (Bogutskaya et al. 2008:329 [ref. 30085]). •Valid as Squalidus argentatus (Sauvage & Dabry de Thiersant 1874) -- (Chen & Li in Chu & Chen 1989:106 [ref. 13584], Zheng et al. 1989:133 [ref. 21202], Jin in Pan et al. 1991:204 [ref. 23876], Ding 1994:268 [ref. 21531], Zhu 1995:79 [ref. 25213], Cheng & Zhou 1997:165 [ref. 26385], Yue in Chen et al. 1998:314 [ref. 23878], Tang et al. 2001:13 [ref. 25373], Kottelat 2001:42 [ref. 25482] as cf. argentatus, Shedko & Shedko 2003:323 [ref. 27966], Bogutskaya & Naseka 2004:73 [ref. 28183] as cf. argentatus, Chen & Chang 2007:70 [ref. 29029], Kottelat 2013:164 [ref. 32989], Zhang et al. 2016:80 [ref. 34477], Zhang et al. 2019:230 [ref. 36699], Dyldin et al. 2020:92, 94 [ref. 37569], Guo 2021:236 [ref. 38617], Endruweit 2025:92 [ref. 42401]). Current status: Valid as Squalidus argentatus (Sauvage & Dabry de Thiersant 1874). Gobionidae. Distribution: East Asia: northern Vietnam, southern and eastern China, Taiwan. IUCN (2012): Data Deficient. Habitat: freshwater, brackish. atromaculatus, Gnathopogon Nichols [J. T.] & Pope [C. H.] 1927:351 [Bulletin of the American Museum of Natural History v. 54 (art. 2); ref. 5747] Nodoa, Hainan Island, China. Holotype: AMNH 8442. Paratypes: AMNH 11007 (122), ?MCZ 32641 [ex AMNH] (2), ZMA 115792 [ex AMNH] (3). Type catalog: Nijssen et al. 1982:28 [ref. 19236]. •Valid as Gnathopogon atromaculatus Nichols & Pope 1927 -- (Luo et al. 1977:492 [ref. 2850]). •Synonym of Gnathopogon wolterstorffi (Regan 1908) -- (Yu in Kuang et al. 1986:98 [ref. 26639]). •Synonym of Squalidus wolterstorffi (Regan 1908) -- (Jin in Pan et al. 1991:203 [ref. 23876]). •Valid as Squalidus atromaculatus (Nichols & Pope 1927) -- (Zheng et al. 1989:134 [ref. 21202], Zhu 1995:80 [ref. 25213], Yue in Chen et al. 1998:318 [ref. 23878], Kottelat 2001:43 [ref. 25482], Kottelat 2001:81 [ref. 25780], Kottelat 2013:164 [ref. 32989], Zhang et al. 2016:80 [ref. 34477]). Current status: Valid as Squalidus atromaculatus (Nichols & Pope 1927). Gobionidae. Distribution: Southeast Asia: southern China, including Hainan (China). IUCN (2012): Least Concern. Habitat: freshwater. banarescui, Squalidus Chen [I-S.] & Chang [Y.-C.] 2007:71, Fig. 3 [Raffles Bulletin of Zoology Suppl. No. 14; ref. 29029] Wu (Ta-du) River, Taichung County, Taiwan. Holotype: NTOU P-2005-07-017. Paratypes: NTOU P-2005-07-018 (11). Type catalog: Ho & Shao 2011:31 [ref. 31432]. •Valid as Squalidus banarescui Chen & Chang 2007 -- (Ho & Shao 2011:31 [ref. 31432], Zhang et al. 2016:80 [ref. 34477] [not China]). Current status: Valid as Squalidus banarescui Chen & Chang 2007. Gobionidae. Distribution: East Asia: Taiwan. IUCN (2024): Least Concern. Habitat: freshwater. biwae, Gobio Jordan [D. S.] & Snyder [J. O.] 1900:340, Pl. 9 (fig. 1) [Proceedings of the United States National Museum v. 23 (no. 1213); ref. 2502] Lake Biwa, near Matsubara, Japan. Holotype: USNM 49399. Paratypes: CAS-SU 6273 (2, missing in 1995). Type catalog: Böhlke 1953:31 [ref. 12291]. •Valid as Gnathopogon biwae (Jordan & Snyder 1900) -- (Kuroda 1953:275 [ref. 39545], Okada 1961:433 [ref. 6363]). •Valid as Squalidus biwae (Jordan & Snyder 1900) -- (Sawada in Masuda et al. 1984:56 [ref. 6441], Okamura & Machida 1987:104 [ref. 22279], Bănărescu 1992:316 [ref. 40251]). •Synonym of Squalidus chankaensis Dybowski 1872, but a valid subspecies -- (Kawanabe & Mizuno 1989:318 [ref. 25598], Sato 1995:51 [ref. 25531], Watanabe 1998:263 [ref. 23491], Sato et al. 1998:36 [ref. 24972], Nakabo 2000:271 [ref. 25086], Matsuura et al. 2000:125 [ref. 26000], Nakabo 2002:271 [ref. 26001]). Current status: Synonym of Squalidus chankaensis Dybowski 1872. Gobionidae. Habitat: freshwater. brevicirris, Gobio ussuriensis morpha Berg [L. S.] 1914:476, Fig. 70 [Faune de la Russie et des pays limitrophes. Poissons, Marsipobranchii et Pisces.; ref. 274] Not available, infrasubspecific, from Ussuri River drainage, Viti River, Russia. •In the synonymy of Squalidus chankaensis Dybowski 1872 -- (Kottelat 2006:50 [ref. 28831], Bogutskaya et al. 2008:329 [ref. 30085]). Infrasubspecific. Current status: Synonym of Squalidus chankaensis Dybowski 1872. Gobionidae. chankaensis, Squalidus Dybowski [B. N.] 1872:215 [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 22; ref. 1170] Lake Khanka [Chanka], Amur River drainage, southeastern Russia. Syntypes: ZMB 7940 [from Dybowski] (2). Original as chankaënsis. •Valid as Gobio chankaensis (Dybowski 1872) -- (Berg 1949:658 [ref. 20661]). •Valid as Gnathopogon chankaensis (Dybowski 1872) -- (Luo et al. 1977:480 [ref. 2850]). •Valid as Squalidus chankaensis Dybowski 1872 -- (Mai 1978:190 [ref. 13511], Hosoya & Jeon 1984:41 [ref. 17431] with subspecies, Zhu 1995:79 [ref. 25213], Bogutskaya & Naseka 1996:43 [ref. 22798], Naseka 1996:160 [ref. 22943], Kim 1997:214 [ref. 24032], Reshetnikov et al. 1997:741 [ref. 24702], Watanabe 1998:263 [ref. 23491] with subspecies, Naseka 1998:87 [ref. 23590], Yue in Chen et al. 1998:313 [ref. 23878], Bogutskaya et al. 2001:44 [ref. 26178], Shedko & Shedko 2003:323 [ref. 27966], Bogutskaya & Naseka 2004:73 [ref. 28183], Ocock et al. 2006:46 [ref. 28933], Bogutskaya et al. 2008:329 [ref. 30085], Zhang et al. 2016:80 [ref. 34477], Dyldin et al. 2023:[24] [ref. 40328] with question). Current status: Valid as Squalidus chankaensis Dybowski 1872. Gobionidae. Distribution: Eastern Asia: China, Russia and Mongolia. Habitat: freshwater. coreanus, Leucogobio Berg [L. S.] 1906:394 [Annals and Magazine of Natural History (Series 7) v. 18 (no. 107) (art. 57); ref. 15643] Sambau River, Kyong-sang-do Province, South Korea. Syntypes: ZIN 13801 (3). Spelled coreianus by Koller 1927:44 [ref. 5622]. •Valid as Squalidus coreanus (Berg 1906) -- (Naseka 1996:160 [ref. 22943]). •Synonym of Squalidus japonicus Sauvage 1883, but a valid subspecies -- (Hosoya & Jeon 1984:41 [ref. 17431], Kim 1997:213 [ref. 24032], Kim & Lee 2001:46 [ref. 25334] as koreanus, Kim & Park 2002:118 [ref. 25913], Choi et al. 2003:158, 507 [ref. 26218]). Current status: Synonym of Squalidus japonicus (Sauvage 1883). Gobionidae. Habitat: freshwater. gracilis, Capoeta Temminck [C. J.] & Schlegel [H.] 1846:201, Pl. 100 (figs. 2, 2a-b) [Fauna Japonica Parts 10-14; ref. 4374] Streams near Nagasaki, Japan. Holotype (unique): RMNH 2499. Type catalog: Boeseman 1947:157 [ref. 12876]. Subjectively invalid; secondarily preoccupied by Barbus gracilis Jerdon 1849 when Günther 1868 placed gracilis Temminck & Schlegel in Barbus. Replaced by and objective synonym of Barbus homozonus Günther 1868; the name homozonus should be used for this species unless gracilis is treated as a nomen protectum. •Valid as Gnathopogon gracilis (Temminck & Schlegel 1846) -- (Okada 1961:436 [ref. 6363]). •Valid as Squalidus gracilis (Temminck & Schlegel 1846), subspecies gracilis -- (Kawanabe & Mizuno 1989:321 [ref. 25598], Sato 1995:51 [ref. 25531], Nakabo 2000:271 [ref. 25086], Matsuura et al. 2000:126 [ref. 26000], Nakabo 2002:271 [ref. 26001], Miuchi & Kunimatsu 2021:18 [ref. 38479], Saiki et al. 2023:69 [ref. 40766]). •Valid as Capoeta gracilis Temminck & Schlegel 1846 -- (Faal et al. 2025:184 [ref. 42154]). •Valid as Squalidus gracilis (Temminck & Schlegel 1846) -- (Sawada in Masuda et al. 1984:56 [ref. 6441], Hosoya & Jeon 1984:41 [ref. 17431], Okamura & Machida 1987:104 [ref. 22279], Kim 1997:211 [ref. 24032], Watanabe 1998:263 [ref. 23491], Sato et al. 1998:36 [ref. 24972]). 2°Homonym. Current status: Valid as Squalidus gracilis (Temminck & Schlegel 1846). Gobionidae. Distribution: East Asia: Japan. IUCN (2023): Least Concern. Habitat: freshwater. guentheri, Abbottina Bănărescu [P. M.] & Nalbant [T. T.] 1973:240, Figs. 127, 127a [Das Tierreich v. 93; ref. 173] Probably fish market, Shanghai, lower Chang Jiang [Yangtze River] estuary, China. Holotype (unique): BMNH 1873.7.30.104. •Synonym of Squalidus nitens (Günther 1873) -- (Yue in Chen et al. 1998:320 [ref. 23878]). •Not in Zhang et al. 2016:81 [ref. 34477]. Current status: Synonym of Squalidus nitens (Günther 1873). Gobionidae. Habitat: freshwater. homozonus, Barbus Günther [A.] 1868:137 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Japan. Replacement name for Capoeta gracilis Temminck & Schlegel 1846, preoccupied by Barbus gracilis Jerdon 1849 when Günther placed Capoeta gracilis in Barbus. Perhaps qualifies as a nomen protectum. •Treated as a synonym of Gnathopogon gracilis (Temminck & Schlegel 1846) -- (Okada 1961:436 [ref. 6363]). •Treated as a synonym of Squalidus gracilis (Temminck & Schlegel 1846) -- (Okamura & Machida 1987:104 [ref. 22279]). Current status: Synonym of Squalidus gracilis (Temminck & Schlegel 1846). Gobionidae. Distribution: Japan. Habitat: freshwater. hsui, Gobio (Leucogobio) Wu [H.-W.] & Wang [K. F.] 1931:227, Fig. 3 [Contributions from the Biological Laboratory of the Science Society of China. (Zoological Series) v. 7 (no. 6); ref. 16351] Katin, Sichuan, China. Holotype (unique): not researched. Originally as hsüi; not of German origin so apparently remains as hsui. Kottelat's (2013) correction of Eschmeyer came after it had been corrected online many months earlier. •Synonym of Gnathopogon argentatus (Sauvage & Dabry de Thiersant 1874) -- (Luo et al. 1977:487 [ref. 2850], Yue in Chen et al. 1998:315 [ref. 23878]). •Synonym of Squalidus argentatus (Sauvage & Dabry de Thiersant 1874) -- (Chen & Chang 2007:70 [ref. 29029], Kottelat 2013:164 [ref. 32989], Zhang et al. 2016:80 [ref. 34477], Zhang et al. 2019:230 [ref. 36699] as hsüi, Endruweit 2025:92 [ref. 42401]). Current status: Synonym of Squalidus argentatus (Sauvage & Dabry de Thiersant 1874). Gobionidae. Habitat: freshwater, brackish. huapingensis, Gnathopogon wolterstorffi Wu [Y.-F.] & Wu [C.-Z.] 1989:68, Fig. 1 [Acta Biologica Plateau Sinica No. 9; ref. 23994] Jinsha River, Chang Jiang [Yangtze River] basin, Huaping, Yunnan Province, China. Holotype: NPIB 19804131. Paratypes: NPIB 19804128 (1), 19804131-33 (2), 19804139 (1), 19804141-43 (3). Species epithet was misspelled in original as woltestorffi. •Synonym of Ganthopogon wolterstorffi (Regan 1908), but a valid subspecies as described -- (Wu & Wu 1992:278 [ref. 21205] as waltestorffi). •Synonym of Squalidus wolterstorffi (Regan 1908) -- (Kottelat 2013:164 [ref. 32989], Zhang et al. 2016:81 [ref. 34477], Zhang et al. 2019:232 [ref. 36699]). Current status: Synonym of Squalidus wolterstorffi (Regan 1908). Gobionidae. Habitat: freshwater, brackish. iijimae, Gnathopogon Oshima [M.] 1919:219, Pl. 51 (fig. 2) [Annals of the Carnegie Museum v. 12 (nos 2-4); ref. 3312] Tozen River, Shin-chu County, Taiwan. Holotype (unique): FMNH 59096 [ex CM 8234]. Type catalog: Henn 1928:71 [ref. 12305], Ibarra & Stewart 1987:40 [ref. 12367], Ho & Shao 2011:27 [ref. 31432]. •Valid as Gnathopogon iijimae Oshima 1919 -- (Luo et al. 1977:490 [ref. 2850]). •Valid as Squalidus iijimae (Oshima 1919) -- (Hosoya & Jeon 1984:41 [ref. 17431], Zhu 1995:80 [ref. 25213], Yue in Chen et al. 1998:322 [ref. 23878], Chen & Chang 2007:71 [ref. 29029], Ho & Shao 2011:27 [ref. 31432], Zhang et al. 2016:80 [ref. 34477] [not China]). Current status: Valid as Squalidus iijimae (Oshima 1919). Gobionidae. Distribution: Taiwan. Habitat: freshwater. intermedius, Gnathopogon Nichols [J. T.] 1929:3, Fig. 2 [American Museum Novitates No. 377; ref. 15912] Tsinan, Shantung Province, China. Holotype: AMNH 9652. Paratypes: (10) AMNH 11021 (3), 11024 (6, 1 c&s), 11026 (1). •Valid as Gnathopogon intermedius Nichols 1929 -- (Luo et al. 1977:489 [ref. 2850]). •Valid as Squalidus intermedius (Nichols 1929) -- (Hosoya & Jeon 1984:41 [ref. 17431], Zhu 1995:79 [ref. 25213], Cheng & Zhou 1997:166 [ref. 26385], Yue in Chen et al. 1998:321 [ref. 23878], Chen & Chang 2007:72 [ref. 29029], Zhang et al. 2016:80 [ref. 34477]). Current status: Valid as Squalidus intermedius (Nichols 1929). Gobionidae. Distribution: China. Habitat: freshwater. ishikawae, Gnathopogon Jordan [D. S.] & Thompson [W. F.] 1914:215, Pl. 24 (fig. 4) [Memoirs of the Carnegie Museum v. 6 (no. 4); ref. 2543] Chikugo River at Kurume, Japan. Holotype: CAS-SU 22607. Paratypes: FMNH 57077 (3), CAS-SU 7355 (10). Type catalog: Henn 1928:72 [ref. 12305], Böhlke 1953:32 [ref. 12291], Ibarra & Stewart 1987:40 [ref. 12367]. Böhlke 1953:32 [ref. 12291] has SU 22607 as paratype; see comment by Ibarra & Stewart 1987:40 [ref. 12367]. The spelling ishikawae is acceptable; early workers almost certainly Latinized the name first, and then added an 'e' (Art. 31.1.1) [WNE, 12 Mar. 2012]. •Synonym of Gnathopogon gracilis (Temminck & Schlegel 1846) -- (Okada 1961:436 [ref. 6363]). •Synonym of Squalidus gracilis (Temminck & Schlegel 1846) -- (Okamura & Machida 1987:104 [ref. 22279]). Current status: Synonym of Squalidus gracilis (Temminck & Schlegel 1846). Gobionidae. Habitat: freshwater. japonicus, Squalius Sauvage [H.-E.] 1883:147 [Bulletin de la Société philomathique de Paris (7th Série) v. 7; ref. 3896] Lake Biwa, Japan. Syntypes: MNHN A-6496 (4). Type catalog: Bertin & Estève 1948:88 [ref. 19574]. •Valid as Gnathopogon japonicus (Sauvage 1883) -- (Kuroda 1953:275 [ref. 39545], Okada 1961:430 [ref. 6363]). •Valid as Squalidus japonicus (Sauvage 1883) -- (Sawada in Masuda et al. 1984:56 [ref. 6441], Hosoya & Jeon 1984:41 [ref. 17431], Okamura & Machida 1987:104 [ref. 22279], Naseka 1996:160 [ref. 22943], Kim 1997:213 [ref. 24032], Watanabe 1998:263 [ref. 23491], Nakabo 2000:271 [ref. 25086]). •Valid as Squalidus japonicus (Sauvage 1883), subspecies japonicus -- (Kawanabe & Mizuno 1989:300 [ref. 25598], Matsuura et al. 2000:128 [ref. 26000], Nakabo 2002:271 [ref. 26001]). Current status: Valid as Squalidus japonicus (Sauvage 1883). Gobionidae. Distribution: East Asia: Japan. Habitat: freshwater. longicirris, Gobio ussuriensis morpha Berg [L. S.] 1914:476, Fig. 71 [Faune de la Russie et des pays limitrophes. Poissons, Marsipobranchii et Pisces.; ref. 274] Not available, infrasubspecific; from Ussuri River, Russia. •In the synonymy of Squalidus mantschuricus (Mori 1927) -- (Bogutskaya et al. 2008:329 [ref. 30085]). Infrasubspecific. Current status: Synonym of Squalidus mantschuricus (Mori 1927). Gobionidae. longifilis, Gnathopogon Jordan [D. S.] & Hubbs [C. L.] 1925:169 [Memoirs of the Carnegie Museum v. 10 (no. 2); ref. 2486] Ping-yang River, northern Korea. Holotype (unique): FMNH 58711 [CM 7817, not 7819]. Type catalog: Henn 1928:72 [ref. 12305], Ibarra & Stewart 1987:40 [ref. 12367]. •Synonym of Squalidus gracilis (Temminck & Schlegel 1846) -- (Hosoya & Jeon 1984:41 [ref. 17431]). Current status: Synonym of Squalidus gracilis (Temminck & Schlegel 1846). Gobionidae. Habitat: freshwater. maii, Parasqualidus Doi [A.] 2000:234, Figs. 1-10 [Raffles Bulletin of Zoology v. 48 (no. 2); ref. 25282] Vugia River, near Ha Tan, Da Nang, central Vietnam. Holotype: NSMT-P 58547. Paratypes: NSMT-P 58548 (1), 58549 (1, c&s). •Valid as Squalidus maii (Doi 2000) -- (Kottelat 2013:164 [ref. 32989]). Current status: Valid as Squalidus maii (Doi 2000). Gobionidae. Distribution: Vietnam. Habitat: freshwater. majimae, Gnathopogon Jordan [D. S.] & Hubbs [C. L.] 1925:167, Pl. 9 (fig. 2) [Memoirs of the Carnegie Museum v. 10 (no. 2); ref. 2486] Ping-yang River, northwestern Korea. Holotype (unique): FMNH 58710 [ex CM 7816]. Type catalog: Henn 1928:72 [ref. 12305], Ibarra & Stewart 1987:40 [ref. 12367]. •Synonym of Squalidus gracilis (Temminck & Schlegel 1846), but a valid subspecies majimae (Jordan & Hubbs 1925) -- (Hosoya & Jeon 1984:41 [ref. 17431], Kim 1997:211 [ref. 24032], Kim & Lee 2001:46 [ref. 25334] as majinae, Kim & Park 2002:116 [ref. 25913], Choi et al. 2003:158, 507 [ref. 26218], Jang et al. 2003:119 [ref. 27681]). Current status: Synonym of Squalidus gracilis (Temminck & Schlegel 1846). Gobionidae. Distribution: Subspecies is endemic to Korea. Habitat: freshwater. mantschuricus, Leucogobio Mori [T.] 1927:30 [1] [Journal of the Chosen Natural History Society No. 5; ref. 16382] Hun River, tributary of Liao River, at Fushun, Manchuria, China. Holotype (unique): whereabouts unknown. On p. 1 of separate. •Probably a subspecies of Squalidus argentatus (Sauvage & Dabry de Thiersant 1874) -- (Naseka 1996:160 [ref. 22943]). •Valid as Squalidus mantschuricus (Mori 1927) -- (Bogutskaya et al. 2008:329 [ref. 30085], Dyldin et al. 2023:[24] [ref. 40328] with question). Current status: Valid as Squalidus mantschuricus (Mori 1927). Gobionidae. Distribution: East Asia: Amur and Liao He rivers (China and Russia). Habitat: freshwater. mayedae, Gobio Jordan [D. S.] & Snyder [J. O.] 1900:342, Pl. 9 (fig. 2) [Proceedings of the United States National Museum v. 23 (no. 1213); ref. 2502] Lake Biwa near Karasaki, Japan. Holotype: USNM 49400. Paratypes: CAS-SU 6272 (orig. 15, now 11). Type catalog: Böhlke 1953:32 [ref. 12291]. •Synonym of Gnathopogon japonicus (Sauvage 1883) -- (Okada 1961:430 [ref. 6363]). •Synonym of Squalidus japonicus (Sauvage 1883) -- (Okamura & Machida 1987:104 [ref. 22279], Naseka 1996:160 [ref. 22943]). Current status: Synonym of Squalidus japonicus (Sauvage 1883). Gobionidae. Habitat: freshwater. minkiangensis, Squalidus gracilis Bănărescu [P. M.] & Nalbant [T. T.] 1964:460, Figs. 3-4 [Bulletin du Muséum National d'Histoire Naturelle (Série 2) v. 36 (no. 4); ref. 9196] Mouth of Min-Kiang River, Foukien Province, China. Holotype (unique): MNHN 1939-0050. •Synonym of Squalidus gracilis (Temminck & Schlegel 1846), but a valid subspecies as described -- (Hosoya & Jeon 1984:41 [ref. 17431] as mikiangensis). Current status: Synonym of Squalidus gracilis (Temminck & Schlegel 1846). Gobionidae. Habitat: freshwater. minor, Leucogobio Harada [I.] 1943:42, Pl. 14 (figs. 52-53) [Freshwater fishes of Hainan Island; ref. 22794] Outskirts of Jinjiang, Qiong-san, Hainan Island, China. No types known. •See also Doi 2000:233 [ref. 25282]. •Valid as Squalidus minor (Harada 1943) -- (Jin in Pan et al. 1991:202 [ref. 23876], Zhu 1995:79 [ref. 25213], Yue in Chen et al. 1998:324 [ref. 23878], Yue & Chen 1998:140 [ref. 26415], Kottelat 2013:164 [ref. 32989], Zhang et al. 2016:80 [ref. 34477]). Current status: Valid as Squalidus minor (Harada 1943). Gobionidae. Distribution: China. Habitat: freshwater, brackish. multimaculatus, Squalidus Hosoya [K.] & Jeon [S.-R.] 1984:42, Figs. 1-3 [Korean Journal of Limnology v. 17 (nos 1-2); ref. 17431] Yongdok-Oship River, Namsan-ri, Yongdok-myon, Yondok-gun, Kyongsangbuk-do, South Korea. Holotype: EWNHM 80204 [ex SMU 301]. Paratypes: EWHNM 80205-06 [ex SMU 302-303] (2); FAKU 52310 (1, c&s), 52311 (1), 52312 (1, c&s), 52313-14 (2), 52315 (1, c&s), 52316 (1); MNHN 1983-0630 to 0631 (2); NSMT-P 23003-04 (2); USNM 263126 (2). Type catalog: Jeon 2001:89 [ref. 25360]. •Valid as Squalidus multimaculatus Hosoya & Jeon 1984 -- (Kim 1997:216 [ref. 24032], Kim & Park 2002:122 [ref. 25913], Choi et al. 2003:158, 507 [ref. 26218]). Current status: Valid as Squalidus multimaculatus Hosoya & Jeon 1984. Gobionidae. Distribution: Korea. Habitat: freshwater. nitens, Gobio Günther [A.] 1873:246 [Annals and Magazine of Natural History (Series 4) v. 12 (no. 69) (art. 31); ref. 2004] Shanghai, China. Syntypes: BMNH 1873.7.30.102 (1), 1873.7.30.103-104 (2). •Valid as Squalidus nitens (Günther 1873) -- (Yue in Chen et al. 1998:319 [ref. 23878], Chen & Chang 2007:72 [ref. 29029], Zhang et al. 2016:81 [ref. 34477], Guo 2021:241 [ref. 38617]). Current status: Valid as Squalidus nitens (Günther 1873). Gobionidae. Distribution: Yangtze River basin, China. Habitat: freshwater. punctatus, Gnathopogon Nichols [J. T.] 1925:7 [American Museum Novitates No. 181; ref. 15553] Near Nanping, Fukien Province, China. Holotype: AMNH 8423. •Synonym of Gnathopogon wolterstorffi (Regan 1908) -- (Luo et al. 1977:491 [ref. 2850]). •Synonym of Squalidus wolterstorffi (Regan 1908) -- (Bănărescu 1992:316 [ref. 40251], Yue in Chen et al. 1998:316 [ref. 23878], Kottelat 2013:164 [ref. 32989]). •Not in Zhang et al. 2016:81 [ref. 34477]. Current status: Synonym of Squalidus wolterstorffi (Regan 1908). Gobionidae. Habitat: freshwater, brackish. sihuensis, Gobio Chu [Y.-T.] 1932:22, Figs. 11, 23 [Fishes of the West Lake; ref. 16479] West Lake, Hangchow, China. No types known. •Valid as Gnathopogon sihuensis (Chu 1932) -- (Luo et al. 1977:489 [ref. 2850]). •Valid as Squalidus sihuensis (Chu 1932) -- (Ding 1994:270 [ref. 21531], Zhu 1995:80 [ref. 25213], Cheng & Zhou 1997:167 [ref. 26385]). •Synonym of Squalidus nitens (Günther 1873) -- (Yue in Chen et al. 1998:319 [ref. 23878], Zhang et al. 2016:81 [ref. 34477], Guo 2021:241 [ref. 38617]). Current status: Synonym of Squalidus nitens (Günther 1873). Gobionidae. Habitat: freshwater. similis, Gnathopogon Nichols [J. T.] 1929:4, Fig. 3 [American Museum Novitates No. 377; ref. 15912] Tsinan, Shantung Province, China. Holotype: AMNH 9653. •Synonym of Squalidus intermedius (Nichols 1929) -- (Yue in Chen et al. 1998:321 [ref. 23878]). Current status: Synonym of Squalidus intermedius (Nichols 1929). Gobionidae. Habitat: freshwater. tsuchigae, Gnathopogon Jordan [D. S.] & Hubbs [C. L.] 1925:170 [Memoirs of the Carnegie Museum v. 10 (no. 2); ref. 2486] Ping-yang River, northern Korea. Holotype (unique): FMNH 58712 [ex CM 7818]. Type catalog: Henn 1928:72 [ref. 12305], Ibarra & Stewart 1987:41 [ref. 12367]. •Synonym of Squalidus gracilis (Temminck & Schlegel 1846), but a valid subspecies -- (Hosoya & Jeon 1984:41 [ref. 17431], Kim 1997:214 [ref. 24032], Kim & Lee 2001:46 [ref. 25334], Kim & Park 2002:120 [ref. 25913]). •Synonym of Squalidus chankaensis Dybowski 1872, but a valid subspecies -- (Choi et al. 2003:158, 506 [ref. 26218]). Current status: Synonym of Squalidus gracilis (Temminck & Schlegel 1846). Gobionidae. Habitat: freshwater. ussuriensis, Gobio Berg [L. S.] 1914:473, Figs. 70-71 [Faune de la Russie et des pays limitrophes. Poissons, Marsipobranchii et Pisces.; ref. 274] Ussuri, lower Amur River, Russia. Syntypes: ZIN (many). •Valid as Gobio ussuriensis Berg 1914 -- (Berg 1949:657 [ref. 20661]). •Synonym of Gnathopogon chankaensis (Dybowski 1872) -- (Luo et al. 1977:486 [ref. 2850], Bogutskaya & Naseka 1996:43 [ref. 22798]). •Synonym of Squalidus chankaensis Dybowski 1872 -- (Reshetnikov et al. 1997:742 [ref. 24702], Naseka 1998:87 [ref. 23590], Yue in Chen et al. 1998:313 [ref. 23878], Kottelat 2006:50 [ref. 28831]). Current status: Synonym of Squalidus chankaensis Dybowski 1872. Gobionidae. Habitat: freshwater. vietnamensis, Squalidus chankaensis Bănărescu [P. M.] & Nalbant [T. T.] 1964:457, Fig. 1 [Bulletin du Muséum National d'Histoire Naturelle (Série 2) v. 36 (no. 4); ref. 9196] Nam-Lung River, tributary of Noire River at Phong-Thô, Vietnam. Holotype: MNHN 1935-0344. Paratypes: MNHN 1935-0345 (1), 1935-0346 (1). •Synonym of Squalidus chankaensis Dybowski 1872, but a valid subspecies as described -- (Mai 1978:191 [ref. 13511]). •Synonym of Squalidus atromaculatus (Nichols & Pope 1927) -- (Kottelat 2001:43 [ref. 25482], Kottelat 2001:81 [ref. 25780], Kottelat 2013:164 [ref. 32989]). •Valid as Squalidus vietnamensis Bănărescu & Nalbant 1964 -- (Endruweit 2025:92 [ref. 42401]). Current status: Valid as Squalidus vietnamensis Bănărescu & Nalbant 1964. Gobionidae. Distribution: Southeast Asia: Red River system (China and Vietnam). Habitat: freshwater. wolterstorffi, Gobio Regan [C. T.] 1908:110, Pl. 4 (fig. 2) [Annals and Magazine of Natural History (Series 8) v. 1 (no. 1) (art. 15); ref. 14335] Nankancho, near Tinghsiang, Shanxi, China. Holotype (unique): BMNH 1907.11.26.3. •Valid as Gnathopogon wolterstorffi (Regan 1908) -- (Luo et al. 1977:491 [ref. 2850], Wang 1984:26 [ref. 22951], Yu in Kuang et al. 1986:98 [ref. 26639], Wu & Wu 1992:278 [ref. 21205], Wang et al. 2001:131 [ref. 26566]). •Valid as Squalidus wolterstorffi (Regan 1908) -- (Zheng et al. 1989:135 [ref. 21202], Jin in Pan et al. 1991:202 [ref. 23876], Ding 1994:272 [ref. 21531], Zhu 1995:80 [ref. 25213], Cheng & Zhou 1997:168 [ref. 26385], Yue in Chen et al. 1998:316 [ref. 23878], Tang et al. 2001:13 [ref. 25373], Chen & Chang 2007:72 [ref. 29029], Kottelat 2013:164 [ref. 32989], Zhang et al. 2016:81 [ref. 34477], Zhang et al. 2019:232 [ref. 36699], Guo 2021:239 [ref. 38617]). Current status: Valid as Squalidus wolterstorffi (Regan 1908). Gobionidae. Distribution: Yangtze River basin, China. Habitat: freshwater, brackish. |