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Eschmeyer's Catalog of Fishes
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         Comments: Richard van der Laan, Ron Fricke

    Please look in Eschmeyer's Catalog of Fishes for authorships of sections, Families of Fishes (all), and related information.

Species in the genus Systomus: [ 46 ] records


asoka, Puntius Kottelat [M.] & Pethiyagoda [R.] 1989:472, Figs. 1, 4 [Die Aquarien- und Terrarienzeitschrift (DATZ) v. 42 (no. 8); ref. 14110] Sitawaka River, 6°55'25"N, 80°20'02"E, Kegala District, Sabaragamuwa Province, Sri Lanka. Holotype: ZSM 27188 [now ZRC 38480]. Paratypes: ZMA 120334 (1); CMK 6402 (9), 6522 (1). Type catalog: Neumann 2011:255 [ref. 31652]. The holotype is currently at ZRC. •Valid as Puntius asoka Kottelat & Pethiyagoda 1989 -- (Pethiyagoda 1991:93 [ref. 20075], Silva et al. 2008:62 [ref. 29719]). •Valid as Systomus asoka Kottelat & Pethiyagoda 1989 -- (Pethiyagoda et al. 2012:77 [ref. 32071], Sudasinghe et al. 2020:[4] [ref. 37737]). Current status: Valid as Systomus asoka (Kottelat & Pethiyagoda 1989). Cyprinidae: Smiliogastrinae. Distribution: South Asia: southwestern Sri Lanka. Habitat: freshwater.

binduchitra, Barbus (Puntius) Hora [S. L.] 1937:327, Fig. 4 [Records of the Indian Museum (Calcutta) v. 39 (pt 4); ref. 20745] Road-side drains and small streams at Sandoway, Lower Myanmar. Holotype: ZSI F 12478/1. Paratypes: ZSI F12478/1 (9). Type catalog: Menon & Yazdani 1968:116 [ref. 20743]. See Das 2003:354 [ref. 27706]. •Synonym of Puntius sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Talwar & Jhingran 1991:282 [ref. 20764]). •Synonym of Barbodes sarana (Hamilton 1822) subspecies orphoides (Valenciennes 1842) -- (Menon 1999:70 [ref. 24904]). •Synonym of Puntius sarana (Hamilton 1822). •Synonym of Systomus sarana (Hamilton 1822) based on placement of sarana. •Valid as Systomus binduchitra (Hora 1937) -- (Kottelat 2013:165 [ref. 32989]). Current status: Valid as Systomus binduchitra (Hora 1937). Cyprinidae: Smiliogastrinae. Distribution: Southeast Asia: lower Myanmar. Habitat: freshwater, brackish.

caudimarginatus, Barbus Blyth [E.] 1860:157 [Journal of the Asiatic Society of Bengal v. 29 (no. 2); ref. 477] Tenasserim provinces, Myanmar. No types known. •Synonym of Puntius sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Talwar & Jhingran 1991:282 [ref. 20764], Menon 1999:69 [ref. 24904] but as genus Barbodes). •Synonym of Systomus orphoides (Valenciennes 1842) -- (Rainboth 1996:104 [ref. 22772]). •Synonym of Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus rubripinnis (Valenciennes 1842). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

chryseus, Systomus Plamoottil [M.] 2014:140, Fig. 12 [International Journal of Fauna and Biological Studies v. 1 (no. 6) Part C; ref. 33650] Keezhvaipur, Manimala River, Kerala, India. Holotype: ZSI/FF 4625. Paratypes: ZSI/WGRC/IR/V. •Valid as Systomus chryseus Plamoottil 2014 -- (Plamoottil 2016:95 [ref. 34814], Arunachalam et al. 2018:45 [ref. 36280], Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Valid as Systomus chryseus Plamoottil 2014. Cyprinidae: Smiliogastrinae. Distribution: South Asia: Manimala River drainage, Kerala, India. Habitat: freshwater.

chrysopoma, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:165, Pl. 466 [Histoire naturelle des poissons v. 16; ref. 1009] Coast of Malabar, India. Syntypes: MNHN 0000-5776 (2). Type catalog: Bertin & Estève 1948:24 [ref. 19574]. •Synonym of Barbodes sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Menon 1999:72 [ref. 24904]). •Synonym of Puntius sarana (Hamilton 1822). •Valid as Systomus chrysopoma (Valenciennes 1842) -- (Plamoottil 2014:145 [ref. 33650], Bleher 2018:788 [ref. 36647]). •Synonym of Systomus subnasutus (Valenciennes 1842) -- (Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989], Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

chrysosomus, Systomus McClelland [J.] 1839:284 [Asiatic Researches v. 19 (pt 2); ref. 2923] Bengal, India. No types known. •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904] as chrysopomus). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

compressiformis, Barbus Cockerell [T. D. A.] 1913:133 [Bulletin of the Bureau of Fisheries v. 32 (Doc. 779) (for 1912); ref. 870] Fort Stedman, southern Shan States, Indochina, elevation 3000 feet. Syntypes: BMNH 1893.6.30.43-52 (10). Replacement name for Barbus compressus Boulenger 1893, preoccupied by Barbus (Barboides) compressus Day 1870. •Synonym of Puntius clavatus (McClelland 1845) -- (Talwar & Jhingran 1991:265 [ref. 20764]). •Valid as Puntius compressiformis (Cockerell 1913) -- (Doi 1997:11 [ref. 22832]). •Valid as Systomus compressiformis (Cockerell 1913) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Kottelat 2013:165 [ref. 32989], Fuke et al. 2021:[3] [ref. 38221]). Replacement Name. Current status: Valid as Systomus compressiformis (Cockerell 1913). Cyprinidae: Smiliogastrinae. Distribution: South Asia: Myanmar. Habitat: freshwater.

compressus, Barbus Boulenger [G. A.] 1893:202 [Annals and Magazine of Natural History (Series 6) v. 12 (no. 69); ref. 14522] Fort Stedman, southern Shan States, Indochina, elevation 3000 feet. Syntypes: BMNH 1893.6.30.43-52 (10). Objectively invalid; preoccupied by Barbus (Barbodes) compressus Day 1870, replaced by Barbus compressiformis Cockerell 1913. Also replaced by Barbus liui Fowler 1958. •Synonym of Puntius clavatus (McClelland 1845) -- (Talwar & Jhingran 1991:265 [ref. 20764]). •Synonym of Systomus compressiformis (Cockerell 1913) -- (Kottelat 2013:165 [ref. 32989]). 1°Homonym. Current status: Synonym of Systomus compressiformis (Cockerell 1913). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

diliciosus, Barbus McClelland [J.] 1839:272, 342, Pl. 39 (fig. 3) [Asiatic Researches v. 19 (pt 2); ref. 2923] Upper Assam, India. No types known. Spelled diliciosus on pp. 272, 343 and plate caption (p. 272); spelled deliciosus in index. •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904] as deliciosus). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). •Valid as Systomus diliciosus (McClelland 1839) -- (Ramalingam & Parasivan 2020:1748 [ref. 38780] as deliciosus). Current status: Valid as Systomus diliciosus (McClelland 1839). Cyprinidae: Smiliogastrinae. Distribution: South Asia: upper Assam (India). Habitat: freshwater, brackish.

duvaucelii, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:167 [Histoire naturelle des poissons v. 16; ref. 1009] Bengal, India. Holotype (unique): MNHN 0000-3817. Type catalog: Bertin & Estève 1948:23 [ref. 19574]. •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904] as duvancelli). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

gardonides, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:156, Pl. 465 [Histoire naturelle des poissons v. 16; ref. 1009] Calcutta and Mumbai, India; Java, Indonesia. Lectotype: MNHN 0000-3819. Syntypes: MNHN 0000-3820 to 0000-3821 (1, 2), B-1214 (1). Type catalog: Bertin & Estève 1948:24-25 [ref. 19574] with MNHN 3819 as holotype. Lectotype selected by Kottelat 2013:166 [ref. 32989]. •Java specimens are questionably a synonym of Puntius orphoides (Valenciennes 1842) -- (Roberts 1993:21 [ref. 21799]). •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904] as gardonoides). •Synonym of Puntius orphoides (Valenciennes 1842) -- (Zhang et al. 2016:97 [ref. 34477]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

gibbosus, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:155 [Histoire naturelle des poissons v. 16; ref. 1009] Aleppey, Kerala, India. Holotype (unique): MNHN 0000-3816. Type catalog: Bertin & Estève 1948:24 [ref. 19574]. •Synonym of Barbodes sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Menon 1999:72 [ref. 24904]). •Synonym of Systomus subnasutus (Valenciennes 1842) -- (Plamoottil & Maji 2020:1036 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

gracilus, Systomus Plamoottil [M.] & Maji [D.] 2020:1034, Figs. 1-2 [Journal of Experimental Zoology of India v. 23 (no. 2); ref. 37708] Ganges River near Naihati, West Bengal, India, 22.8895°N, 88.422°E. Holotype: ZSI/ANRC M-23593. Paratypes: ZSI PUNE P-5540 (3). •Valid as Systomus gracilus Plamoottil & Maji 2020. Current status: Valid as Systomus gracilus Plamoottil & Maji 2020. Cyprinidae: Smiliogastrinae. Distribution: South Asia: lower Ganges River basin, West Bengal, India. Habitat: freshwater.

immaculatus, Systomus McClelland [J.] 1839:284, 380, Pl. 44 (fig. 5) [Asiatic Researches v. 19 (pt 2); ref. 2923] Assam, India Neotype: AMS B.7920. Neotype selected by Pethiyagoda et al. 2012:77 [ref. 32071], a Day specimen. •Synonym of Puntius sarana (Hamilton 1822) -- (Coad 1981:11 [ref. 22348]). •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904]). •Synonym of Systomus sarana (Hamilton 1822) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Kottelat 2013:166 [ref. 32989]). •Valid as Systomus immaculatus McClelland 1839 -- (Bleher 2018:792 [ref. 36647], Ramalingam & Parasivan 2020:1753 [ref. 38780], Çiçek et al. 2023:52 [ref. 40326]). Current status: Valid as Systomus immaculatus McClelland 1839. Cyprinidae: Smiliogastrinae. Distribution: South Asia: Assam (India). Habitat: freshwater.

jacobusboehlkei, Barbus Fowler [H. W.] 1958:11 [Notulae Naturae (Philadelphia) No. 310; ref. 1470] Huey Melin, a branch of the Menam Mao, northern Thailand. Holotype (unique): USNM 119452. Originally as jacobus-boehlkei. Replacement name for Puntius simus H. M. Smith 1945, secondarily preoccupied in Barbus by Barbus (Systomus) simus Sauvage & Dabry de Thiersant 1874. •Synonym of Systomus orphoides (Valenciennes 1842) -- (Rainboth 1996:104 [ref. 22772]). •Valid as Puntius jacobusboehlkei (Fowler 1958) -- (Doi 1997:11 [ref. 22832], Kottelat 1998:49 [ref. 23436], Freyhof et al. 2000:95 [ref. 25184], Kottelat 2001:74 [ref. 25780], Kottelat 2013:166 [ref. 32989]). •Valid as Barbodes jacobusboehlkei (Fowler 1958) -- (Kottelat 2015:40 [ref. 33971]). •Valid as Systomus jacobusboehlkei (Fowler 1958) -- (Taki et al. 2021:127 [ref. 39830]). Replacement Name. Current status: Valid as Systomus jacobusboehlkei (Fowler 1958). Cyprinidae: Smiliogastrinae. Distribution: Southeast Asia: Mekong and Chao Phraya River basins, Thailand, Laos and Cambodia. Habitat: freshwater.

kadoon, Barbus Bleeker [P.] 1853:60 [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 25 (art. 8); ref. 341] Coromandel, India. No types known. Appeared as "Barbus ? kadon Blkr., Kadoon Russ. no. 206" which is Russell 1803: Pl. 206 [ref. 13862]. •Synonymy of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [re. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

kadoon, Cyprinus Heckel [J. J.] 1843:1018 [28] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Vizagapatam, India. No types known. On p. 28 of separate. Based on Russell 1803, Pl. 206 [ref. 13862]. •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

kakus, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:153 [Histoire naturelle des poissons v. 16; ref. 1009] Tank near Tartoor, Vizagapatam, India. No types known. Bases on Russell, Coromondal 2 1803, 83, no. 205. •Synonym of Barbodes sarana (Hamilton 1822), subspecies sarana -- (Menon 1999:70 [ref. 24904]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

kunama, Cyprinus Cuvier [G.] in Russell 1829:273 (footnote 1) [Le Règne Animal (Edition 2) v. 2; ref. 995] Vizagapatham, India. As "C. kunama Russel, 204", in reference to Russel 1803:fig. 204. •Synonym of Systomus sarana (Hamilton 1822) with question -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

kunamo, Cyprinus McClelland [J.] in Russell 1839:272, 340 [Asiatic Researches v. 19 (pt 2); ref. 2923] Not available, name mentioned in synonymy. •In the synonymy of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Nomen Nudum. Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae.

laticeps, Systomus Plamoottil [M.] 2016:92, Figs. 1-4 [International Journal of Fauna and Biological Studies v. 3 (no. 5) Part B; ref. 34814] Freshwater stream at Thiruvalla, Pathanamthitta district, Kerala, India. Holotype (unique): FBRC/ZSI F2316. •Valid as Systomus laticeps Plamoottil 2016 -- (Arunachalam et al. 2018:45 [ref. 36280], Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Valid as Systomus laticeps Plamoottil 2016. Cyprinidae: Smiliogastrinae. Distribution: South Asia: Kerala, India. Habitat: freshwater.

liui, Barbus Fowler [H. W.] 1958:12 [Notulae Naturae (Philadelphia) No. 310; ref. 1470] Fort Stedman, southern Shan States, Myanmar, elevation 3000 feet. Syntypes: BMNH 1893.6.30.43-52. Unneeded replacement name for Barbus compressus Boulenger 1898, preoccupied by Barbus (Barbodes) compressus Day 1869, but replaced earlier by Barbus compressiformis Cockerell 1913. •Synonym of Puntius clavatus (McClelland 1845) -- (Talwar & Jhingran 1991:265 [ref. 20764] as lini). •Synonym of Systomus compressiformis (Cockerell 1913) -- (Kottelat 2013:165 [ref. 32989]). Replacement Name. Current status: Synonym of Systomus compressiformis (Cockerell 1913). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

martenstyni, Puntius Kottelat [M.] & Pethiyagoda [R.] 1991:303, Figs. 6-8 [Freshwater fishes of Sri Lanka; ref. 16752] Kalu River about 2 kilometers north of Pallegama, 7°33'N, 80°50'E, Mahaweli River basin, Marale District, Sri Lanka. Holotype: ZSM 27966 [now ZRC 38482]. Paratypes: CMK 6539 (2), 7078 (2), 7168 (11). Type catalog: Neumann 2011:255 [ref. 31652]. The holotype is at ZRC. •Valid as Systomus martenstyni (Kottelat & Pethiyagoda 1991) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Ren et al. 2020:[5] [ref. 37805], Sudasinghe et al. 2020:[4] [ref. 37737]). Current status: Valid as Systomus martenstyni (Kottelat & Pethiyagoda 1991). Cyprinidae: Smiliogastrinae. Distribution: South Asia: central highlands of Sri Lanka. Habitat: freshwater.

mclellandi, Cyprinus Valenciennes [A.] in Cuvier & Valenciennes 1842:390 [Histoire naturelle des poissons v. 16; ref. 1009] Assam streams and probably occasionally in the large rivers, India. No type known. Original as M'clellandi. Possibly changed to macclellandi as named for McClelland. Unnecessary replacement name for Systomus immaculatus McClelland 1839. •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). •Synonym of Systomus immaculatus McClelland 1839. Replacement Name. Current status: Synonym of Systomus immaculatus McClelland 1839. Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

morehensis, Puntius Arunkumar [L.] & Tombi Singh [H.] 1998:254, Fig.3 [Journal of Nature Conservation: an International journal devoted to natural resource conservation and environment. v. 10 (no. 2); ref. 24931] Lairok Maru stream of Lokchao River near Moreh, 2 kilometers from Moreh Bazar, Manipur, India. Holotype: MUMF 240/1A. Paratypes: MUMF 240/3A (19). •Valid as Puntius morehensis Arunkumar & Tombi Singh 1998 -- (Arunkumar & Tombe Singh 2003:481 [ref. 26926]). •Synonym of Systomus sewelli (Prashad & Mukerji 1929) -- (Kotteat 2013:167 [ref. 32989] with question). Current status: Synonym of Systomus sewelli (Prashad & Mukerji 1929). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

orphoides, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:193 [Histoire naturelle des poissons v. 16; ref. 1009] Java, Indonesia. No types known. Type information: Roberts 1993:21-22 [ref. 21799]. Simultaneous subjective synonym of Barbus rubripinnis Valenciennes 1842; Bleeker 1860:338 [ref. 380] gave preference to rubripinnis, see Kottelat 2013:166 [ref. 32989]. •Synonym of Puntius sarana (Hamilton 1822), but a valid subspecies -- (Talwar & Jhingran 1991:282 [ref. 20764], Arunkumar & Tombi Singh 1998:256 [ref. 24931], Arunachalam & Sankaranarayanan 1999:336 [ref. 24216], Karmakar 2000:29 [ref. 25662]). •Synonym of Barbodes sarana (Hamilton 1822), but a valid subspecies -- (Menon 1999:69 [ref. 24904]). •Valid as Puntius orphoides (Valenciennes 1842) -- (Roberts 1993:21 [ref. 21799], Kottelat et al. 1993:43 [ref. 23448], Kottelat & Lim 1996:233 [ref. 23210], Doi 1997:11 [ref. 22832], Doi 2000:396 [ref. 25097], Kottelat 2001:74 [ref. 25780], Vishwanath & Laisram 2004:136 [ref. 27863], Shantakumar & Vishwanath 2006:2279 [ref. 28676], Zhang et al. 2016:97 [ref. 34477]). •Valid as Systomus orphoides (Valenciennes 1842) -- (Rainboth 1996:104 [ref. 22772], Parenti & Lim 2005:186 [ref. 28285], Pethiyagoda et al. 2012:77 [ref. 32071], Plamoottil 2014:145 [ref. 33650], Plamoottil 2016:96 [ref. 34814], Bleher 2018:794 [ref. 36647], Ren et al. 2020:[5] [ref. 37805]). •Synonym of Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus rubripinnis (Valenciennes 1842). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

pinnauratus, Cyclocheilichthys Day [F.] 1865:300 [Proceedings of the Zoological Society of London 1865 (pt 1); ref. 5295] Small pond, Cochin, India. Holotype (unique): ?ZSI 2728. Day specimens: BMNH 1889.2.1.4332-33 (2), FMNH 2309 (2), NMW 54468-70 (3), RMNH 2582 (1), ZIN 8216 (2). Type catalog: Whitehead & Talwar 1976:156 [ref. 23762]. Also appeared in Day 1865:209, Pl. 15, fig. 2 [ref. 1074]. •Mention under Puntius sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Talwar & Jhingran 1991:285 [ref. 20764], Indra 1993:184 [ref. 24009], Indra 1994:416 [ref. 24015]). •Synonym of Barbodes sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Menon 1999:72 [ref. 24904]). •Valid as Systomus pinnauratus (Day 1865) -- (Plamoottil 2014:142 [ref. 33650], Plamoottil 2016:92 [ref. 34814]). •Synonym of Systomus subnasutus (Valenciennes 1842) -- (Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989], Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

pleurotaenia, Puntius (Barbodes) Bleeker [P.] 1863:242 [Verslagen en Mededeelingen der Koninklijke Akademie van Wetenschappen. Afdeling Natuurkunde. v. 15; ref. 17011] Sri Lanka. Holotype (unique): whereabouts unknown. Also appeared in Bleeker 1864:13, Pl. 3 (fig. 2) [ref. 13925]. •Valid as Puntius pleurotaenia Bleeker 1863 -- (Pethiyagoda 1991:111 [ref. 20075], Talwar & Jhingran 1991:279 [ref. 20764], Silva et al. 2008:62 [ref. 29719]). •Valid as Systomus pleurotaenia (Bleeker 1863) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Ren et al. 2020:[5] [ref. 37805], Sudasinghe et al. 2020:[4] [ref. 37737]). Current status: Valid as Systomus pleurotaenia (Bleeker 1863). Cyprinidae: Smiliogastrinae. Distribution: South Asia: southwestern Sri Lanka. Habitat: freshwater.

polydori, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:170 [Histoire naturelle des poissons v. 16; ref. 1009] Mumbai, India. Holotype (unique): MNHN 0000-3827. Type catalog: Bertin & Estève 1948:25 [ref. 19574]. •Synonym of Barbodes sarana (Hamilton 1822) -- (Menon 1999:70 [ref. 24904]). •Uncertain in genus Puntius -- (Plamoottil & Johnson 2020:99 [ref. 37958]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

roseipinnis, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:169 [Histoire naturelle des poissons v. 16; ref. 1009] Puducherry [Pondicherry], India. Syntypes: MNHN 1405 (2). •Valid as Puntius roseipinnis (Valenciennes 1842) -- (Talwar & Jhingran 1991:281 [ref. 20764]). •Synonym of Barbodes sarana (Hamilton 1822), subspecies subnasutus (Valenciennes 1842) -- (Menon 1999:72 [ref. 24904]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 3289]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

rubripinna, Barbus van Hasselt [J. C.] 1823:132 [Algemeene Konst- en Letter-bode voor het Jaar II Deel (no. 35); ref. 5963] Not available, name only; from Batavia, Java (see Alfred 1961:85 [ref. 20553], Kottelat 1987:370 [ref. 5962]). Also in van Hasselt 1824:375 [ref. 5964]. •In the synonymy of Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989]). Nomen Nudum. Current status: Synonym of Systomus rubripinnis (Valenciennes 1842). Cyprinidae: Smiliogastrinae.

rubripinnis, Barbus Valenciennes [A.] (ex Kuhl & van Hasselt) in Cuvier & Valenciennes 1842:194 [Histoire naturelle des poissons v. 16; ref. 1009] Java, Indonesia. Syntypes: whereabouts unknown. Simultaneous subjective synonym of Barbus orphoides Valenciennes 1842. Bleeker 1860:338 [ref. 380] gave precedence to Barbus rubripinnis. •Synonym of Barbodes sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Menon 1999:69 [ref. 24904]). •Synonym of Puntius sarana (Hamilton 1822). •Valid as Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989], Ng et al. 2019:525 [ref. 36606], Taki et al. 2021:127 [ref. 39830], Hasan et al. 2022:6035 [ref. 39888], Ahmad et al. 2023:76 [ref. 41243], Chungthanawong et al. 2025:65 [ref. 41920]). Current status: Valid as Systomus rubripinnis (Valenciennes 1842). Cyprinidae: Smiliogastrinae. Distribution: Southeast Asia: Sundaland, Malaysia, Myanmar, Chao Phraya and Mekong River basin, Thailand, Laos, Cambodia and Vietnam; introduced elsewhere. IUCN (2019): Data Deficient. Habitat: freshwater.

rufus, Systomus Plamoottil [M.] 2014:142, Fig. 17 [International Journal of Fauna and Biological Studies v. 1 (no. 6) Part C; ref. 33650] Venpala, Manimala River, Kerala, India. Holotype: ZSI/WGRC/IR/2367. Paratypes: ZSI/WGRC/IR/2368 (1). •Valid as Systomus rufus Plamoottil 2014 -- (Plamoottil 2016:95 [ref. 34814], Arunachalam et al. 2018:45 [ref. 36280], Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Sudasinghe et al. 2020:[14] [ref. 37737]). Current status: Valid as Systomus rufus Plamoottil 2014. Cyprinidae: Smiliogastrinae. Distribution: South Asia: Manimala River drainage, Kerala, India. Habitat: freshwater.

russellii, Barbus Günther [A.] 1868:121 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Indus River, Sabzilkot, India. Syntypes: BMNH 1860.3.19.732 (1), 1860.3.19.856 (1). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

saberi, Puntius Datta [A. K.] & Karmakar [A. K.] 1981:179, Fig. 1 [Bulletin of the Zoological Survey of India v. 3 (no. 3); ref. 8616] Cheroguru Talao, Pujaripal village, 23 kilometers from Sukma, Bastar District, Madhya Pradesh, India. Holotype: ZSI FF1398. Paratypes: ZSI FF1399 (12). See Das 2003:360 [ref. 27706]. •Synonym of Puntius sarana (Hamilton 1822) -- (Talwar & Jhingran 1991:283 [ref. 20764]). •Synonym of Barbodes sarana (Hamilton 1822), subspecies sarana -- (Menon 1999:72 [ref. 24904]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

sarana, Cyprinus Hamilton [F.] 1822:307, 388 [An account of the fishes found in the river Ganges; ref. 2031] Ponds and rivers of Bengal. No types known. Hamilton's original, unpublished illustration reproduced by Britz 2019:Pl. 180 [ref. 37532]. •Valid as Puntius sarana (Hamilton 1822), subspecies sarana -- (Karmakar 2000:29 [ref. 25662], Shrestha 2008:80 [ref. 29923], Gurung et al. 2013:4882 [ref. 32969], Arunkumar & Manimekalan 2018:1739 [ref. 36172]). •Valid as Barbodes sarana (Hamilton 1822) -- (Rema Devi & Raghunathan 1999:150 [ref. 24759], Rema Devi & Raghunathan 1999:167 [ref. 24760], Rema Devi et al. 1999:155 [ref. 24758], Menon 1999:70 [ref. 24904], Rafique 2000:323 [ref. 25220], Yadav 2003:13 [ref. 27711], Ataur Rahman 2003:145 [ref. 31338]). •Valid as Puntius sarana (Hamilton 1822) -- (Shrestha 1978:36 [ref. 15970], Ataur Rahman 1989:126 [ref. 24860], Coad 1981:11 [ref. 22348], Pethiyagoda 1991:113 [ref. 20075], Talwar & Jhingran 1991:283 [ref. 20764], Indra 1991:239 [ref. 24005], Sen 1995:515 [ref. 23778], Doi 1997:11 [ref. 22832], Vu & Nguyen 1997:26 [ref. 26438], Arunkumar & Tombi Singh 1998:256 [ref. 24931], Manimekalan 1998:436 [ref. 24989], Arunachalam & Sankaranarayanan 1999:336 [ref. 24216], Bijukumar & Sushama 2000:185 [ref. 25703], Arunkumar & Tombe Singh 2003:481 [ref. 26926], Rafique et al. 2003:97 [ref. 27173], Vishwanath & Laisram 2004:136 [ref. 27863], Shantakumar & Vishwanath 2006:2279 [ref. 28676]). •Valid as Systomus sarana (Hamilton 1822) -- (Mirza 2003:9 [ref. 27265], Pethiyagoda et al. 2012:77 [ref. 32071], Kottelat 2013:166 [ref. 32989], Plamoottil 2014:145 [ref. 33650], Plamoottil 2016:95 [ref. 34814], Arunachalam et al. 2018:45 [ref. 36280], Suresh et al. 2018:113 [ref. 36495], Bleher 2018:798 [ref. 36647], Plamoottil & Maji 2020:1035 [ref. 37708], Ren et al. 2020:[5] [ref. 37805], Sudasinghe et al. 2020:[4] [ref. 37737], Kosygin et al. 2022:657 [ref. 40736], Tudu et al. 2022:43 [ref. 40484], Çiçek et al. 2023:52 [ref. 40326], Das & Sivaperuman 2024:154 [ref. 41718]). Current status: Valid as Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Distribution: South Asia: Nepal, Bhutan, India, Sri Lanka, Myanmar, Pakistan and Bangladesh. Introduced in Andaman Islands (India). Habitat: freshwater, brackish.

sarananella, Barbus Bleeker [P.] 1849:16 [Verhandelingen van het Bataviaasch Genootschap van Kunsten en Wetenschappen. v. 23 (no. 12); ref. 16883] Serayu River, Banjumas; Gombong; Bogowonto River, Purworedjo, Java, Indonesia. •Synonym of Barbodes sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Menon 1999:69 [ref. 24904] as saranella). •Synonym of Puntius sarana (Hamilton 1822). •Synonym of Systomus sarana (Hamilton 1822) based on placement of sarana. •Synonym of Systomus rubripinnis (Valenciennes 1842) -- (Kottelat 2013:166 [ref. 32989]). •Synonym of Systomus orphoides (Valenciennes 1842). Current status: Synonym of Systomus rubripinnis (Valenciennes 1842). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

sewelli, Barbus Prashad [B.] & Mukerji [D. D.] 1929:197, Pl. 9 (figs. 1, 1a-b) [Records of the Indian Museum (Calcutta) v. 31 (pt 3); ref. 3558] Indawgyi Lake, Myitkyina District, Upper Myanmar. Holotype (unique): ZSI F10910/1. Type catalog: Menon & Yazdani 1968:106 [ref. 20743]. See Das 2003:354 [ref. 27706] with date as 1930. •Synonym of Puntius sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Talwar & Jhingran 1991:282 [ref. 20764]). •Synonym of Barbodes sarana (Hamilton 1822), subspecies orphoides (Valenciennes 1842) -- (Menon 1999:70 [ref. 24904]). •Synonym of Puntius sarana (Hamilton 1822). •Valid as Systomus sewelli (Prashad & Mujkerji 1929) -- (Kottelat 2013:167 [ref. 32989], Chen et al. 2024:78 [ref. 41792]). Current status: Valid as Systomus sewelli (Prashad & Mukerji 1929). Cyprinidae: Smiliogastrinae. Distribution: Southeast Asia: Irrawaddy, Sittaung and Tanintharyi River basins, Myanmar. Habitat: freshwater, brackish.

simus, Puntius Smith [H. M.] 1945:185, Fig. 32 [Bulletin of the United States National Museum No. 188; ref. 4056] Huey Melin [Maelin], a tributary of the Mao River, northern Thailand. Holotype (unique): USNM 119452. Type catalog: Monkolprasit 1969:13 [ref. 25088], Vilasri et al. 2018:53 [ref. 36451]. Secondarily preoccupied in Barbus by Barbus (Systomus) simus Sauvage & Dabry de Thiersant 1874. •Synonym of Systomus orphoides (Valenciennes 1842) -- (Rainboth 1996:104 [ref. 22772]). •Synonym of Barbodes jacobusboehlkei (Fowler 1958) -- (Vilasri et al. 2018:53 [ref. 36451]). •Synonym of Systomus jacobusboehlkei (Fowler 1958) -- (Kottelat 2013:166 [ref. 32989]). 2°Homonym. Current status: Synonym of Systomus jacobusboehlkei (Fowler 1958). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

spilurus, Barbus Günther [A.] 1868:114 [Catalogue of the fishes in the British Museum v. 7; ref. 1990] Sri Lanka. Syntypes: BMNH 1852.2.19.124 (1), 1858.10.19.86-91 (6), 1859.5.31.47 (1). •Synonym of Puntius sarana (Hamilton 1822), but a valid subspecies -- (Talwar & Jhingran 1991:284 [ref. 20764], Arunkumar & Manimekalan 2018:1739 [ref. 36172] as spirulus). •Valid as Systomus spilurus (Günther 1868) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Plamoottil 2014:145 [ref. 33650], Plamoottil 2016:96 [ref. 34814], Arunachalam et al. 2018:46 [ref. 36280], Ren et al. 2020:[5] [ref. 37805], Bhakta et al. 2022:[2] 194 [ref. 39348]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989], Sudasinghe et al. 2020:[16] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

stedmanensis, Barbus Annandale [N.] (ex Boulenger) 1918:47 [Records of the Indian Museum (Calcutta) v. 14; ref. 127] Unneeded replacement name for Barbus compressus Boulenger 1893, preoccupied by Barbus compressus Day 1870; replaced earlier by Barbus compressiformis Cockerell 1913. •Synonym of Puntius clavatus (McClelland 1845) -- (Talwar & Jhingran 1991:265 [ref. 20764]). •Synonym of Systomus compresssiformis (Cockerell 1913) -- (Kottelat 2013:165 [ref. 32989], Fuke et al. 2021:[3] [ref. 38221]). Replacement Name. Current status: Synonym of Systomus compressiformis (Cockerell 1913). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

subnasutus, Barbus Valenciennes [A.] in Cuvier & Valenciennes 1842:154 [Histoire naturelle des poissons v. 16; ref. 1009] Puducherry [Ponducherry], India. Holotype (unique): MNHN 0000-3823. Type catalog: Bertin & Estève 1948:24 [ref. 19574]. •Synonym of Puntius sarana (Hamilton 1822), but a valid subspecies -- (Talwar & Jhingran 1991:285 [ref. 20764], Indra 1993:184 [ref. 24009], Rema Devi 1993:198 [ref. 24010], Indra 1994:416 [ref. 24015], Manimekalan 1998:436 [ref. 24989], Arunachalam et al. 2001:128 [ref. 25692], Bijukumar & Sushama 2000:185 [ref. 25703]). •Synonym of Barbodes sarana (Hamilton 1822), but a valid subspecies -- (Rema Devi et al. 1996:132 [ref. 24021], Rema Devi & Raghunathan 1999:150 [ref. 24759], Rema Devi & Raghunathan 1999:167 [ref. 24760], Rema Devi et al. 1999:155 [ref. 24758], Menon 1999:70 [ref. 24904], Raju Thomas et al. 2000:444 [ref. 25319], Raju Thomas et al. 2002:49 [ref. 26359], Yadav 2003:13 [ref. 27711]). •Valid as Systomus subnasutus (Valenciennes 1842) -- (Plamoottil 2014:145 [ref. 33650], Plamoottil 2016:95 [ref. 34814], Arunachalam et al. 2018:45 [ref. 36280], Bleher 2018:800 [ref. 36647], Plamoottil & Maji 2020:1035 [ref. 37708]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:166 [ref. 32989], Sudasinghe et al. 2020:[12] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

takhoaensis, Puntius Nguyen [V. H.] & Doan [L. H.] 1969:12, Fig. 10 in ref. 29795 [Some data on composition, origin, and distribution of cyprinid species in northern Vietnam; ref. 25483] Suoi Nan Ta Khoa stream, Son La Province, Vietnam. Lectotype: RIAH 140. Paralectotypes: (9) RIAH 140 (8). See Kottelat 2001:113, 119 for availability issues. Spelled in more than one way originally, Kottelat 2001:119 [ref. 25482] as first reviser selected takhoaensis. See Nguyen & Doan 2008 [ref. 29793] for an English translation of ref. 25483. Lectotype selected by Roberts & Catania 2008:91 [ref. 29794]. •Valid as Puntius takhoaensis Nguyen & Doan 1969 -- (Kottelat 2001:37 [ref. 25482]). •Should be referred to Systomus -- (Roberts & Catania 2008:96 [ref. 29794]). •Synonym of Systomus jacobusboehlkei (Fowler 1958) -- (Kottelat 2013:166 [ref. 32989]). •Valid as Puntius takhoaensis Mai 1978 -- (Endruweit 2024:15 [ref. 41040]). •Synonym of Barbodes jacobusboehlkei (Fowler 1958). Current status: Synonym of Systomus jacobusboehlkei (Fowler 1958). Cyprinidae: Smiliogastrinae. Habitat: freshwater.

takhoaensis, Puntius Mai [D. Y.] (ex Hao & Hoa) 1978:110 [Identification of the fresh-water fishes of North Viet Nam v. 1; ref. 13511] Northern Vietnam. Syntypes: DVZUT. Apparently unavailable from Hao & Hoa 1969 according to Kottelat 1989:1102 [ref. 14361], but Kottelat 2001 [ref. 25482] reversed this interpretation and dated to Nguyen & Doan [= Hao & Hoa] 1969. •See the account of Puntius takhoaensis Nguyen & Doan 1969. •In the synonymy of Puntius takhoaensis Nguyen & Doan 1969 -- (Kottelat 2001:37 [ref. 25482]). •Should be referred to the genus Systomus -- (Roberts & Catania 2008:94 [ref. 29794]). •Valid as Puntius takhoaensis Mai 1978 -- (Endruweit 2024:15 [ref. 41040]). •In the synonymy of Systomus jacobusboehlkei (Fowler 1958) 1°Homonym. Current status: Synonym of Systomus jacobusboehlkei (Fowler 1958). Cyprinidae: Smiliogastrinae.

timbiri, Puntius Deraniyagala [P. E. P.] 1963:63, Pl. 1 (figs. 2-3) [Spolia Zeylanica v. 30 (pt 1); ref. 4605] Valavé River at Timbirigas-mankada, Sri Lanka, depth about 5 feet. Holotype (unique): NMSL FF801. Type information: Pethiyagoda 1991:336 [ref. 20075]. •Synonym of Puntius sarana (Hamilton 1822), subspecies spilurus (Günther 1868) -- (Talwar & Jhingran 1991:285 [ref. 20764]). •Valid as Systomus timbiri (Deraniyagala 1963) -- (Pethiyagoda et al. 2012:77 [ref. 32071], Ren et al. 2020:[5] [ref. 37805]). •Synonym of Systomus sarana (Hamilton 1822) -- (Kottelat 2013:167 [ref. 32989] with question, Sudasinghe et al. 2020:[16] [ref. 37737]). Current status: Synonym of Systomus sarana (Hamilton 1822). Cyprinidae: Smiliogastrinae. Habitat: freshwater, brackish.

tripunctatus, Systomus Jerdon [T. C.] 1849:316 [Madras Journal of Literature and Science v. 15 (pt 2); ref. 4902] Near the coast in Canara [Karnataka], southern India. No types known. •Status uncertain -- (Kawate et al. 2014:242 [ref. 33433]). Current status: Uncertain as Systomus tripuinctatus Jerdon 1849. Cyprinidae: Smiliogastrinae. Habitat: freshwater.


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