Institute for Biodiversity Science and Sustainability
Online Version, Updated 5 August 2019

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Eschmeyer's Catalog of Fishes
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pungitius, Gasterosteus Linnaeus [C.] 1758:296 [Systema Naturae, Ed. X v. 1; ref. 2787] Europe. Syntypes: LS 34-35 (2, left half-skins). Type catalog: Wheeler 1985:44 [ref. 20705]. This taxon may involve a number of valid species and subspecies. •Valid as Pungitius pungitius (Linnaeus 1758), subspecies pungitius -- (Okada 1961:615 [ref. 6363], Kawanabe & Mizuno 1989:445 [ref. 25598], Sheiko & Fedorov 2000:25 [ref. 25650], Pietsch et al. 2001:144 [ref. 25688], Dolgov 2013:917 [ref. 33112]). •Valid as Pungitius pungitius (Linnaeus 1758) -- (Berg 1949:965 [ref. 20662], Monod 1973:284 [ref. 7193], Lee et al. 1980:566 [ref. 22416], Ida in Masuda et al. 1984:83 [ref. 6441], Smith 1986:277 [ref. 22186], Banister in Whitehead et al. 1986:642 [ref. 13676], Robins & Ray 1986:122 [ref. 23100], Lelek 1987:271 [ref. 22962], Scott & Scott 1988:343 [ref. 25518], McAllister 1990:219 [ref. 14674], Page & Burr 1991:242 [ref. 18983], Haglund et al. 1992 [ref. 20881], Chereshnev 1996:605 [ref. 24659], Keivany et al. 1997:558 [ref. 22904], Kottelat 1997:167 [ref. 22952], Kim 1997:391 [ref. 24032], Reshetnikov et al. 1997:747 [ref. 24702], Reshetnikov & Savvaitove 1998:112 [ref. 23593], Scott & Crossman 1998:671 [ref. 24570], Keivany & Nelson 1998:458 [ref. 24681] with subspecies, Fuller et al. 1999:336 [ref. 25838], Keivany & Nelson 2000:117 [ref. 24720], Lyons et al. 2000:42 [ref. 26594], Nakabo 2000:514 [ref. 25086], Matsuura et al. 2000:197 [ref. 26000], Zemnukhov et al. 2001:420 [ref. 25412], Bogutskaya et al. 2001:46 [ref. 26178], Chereshnev et al. 2001:113 [ref. 26736], Mecklenburg et al. 2002:334 [ref. 25968], Paepke in Bănărescu & Paepke 2002:277 [ref. 25856], Kim & Park 2002:313 [ref. 25913], Nakabo 2002:514 [ref. 26001], Youn 2002:231, 554 [ref. 26218], Kolpakov 2002:708 [ref. 26469], Kolpakov 2003:847 [ref. 27537], Fedorov et al. 2003:57 [ref. 27579], Pichugin et al. 2004:15 [ref. 27623], Alekseyev et al. 2004:54 [ref. 27625], Bailey et al. 2004:169 [ref. 27882], Nelson et al. 2004:111 [ref. 27807], Bogutskaya & Naseka 2004:181 [ref. 28183], Fricke 2007:28 [ref. 30577], Kottelat & Freyhof 2007:498 [ref. 29996], Scharpf 2007:34 [ref. 30398], McPhail 2007:440 [ref. 31199], Bogutskaya et al. 2008:350 [ref. 30085], Mecklenburg et al. 2011:121 [ref. 31212], Page & Burr 2011:462 [ref. 31215], Page et al. 2013:113 [ref. 32708], Parin et al. 2014:205 [ref. 33547], Afonso et al. 2016:39, 259 [ref. 36199], Kells et al. 2016:122 [ref. 35888], Takahashi et al. 2016:49 [ref. 34337], Dyldin & Orlov 2017:57 [ref. 35138], Denys et al. 2017:14 [ref. 35585], Romanov et al. 2017:825 [ref. 35687], Artüz & Fricke 2019:555 [ref. 36612]). Current status: Valid as Pungitius pungitius (Linnaeus 1758). Gasterosteidae. Distribution: Nearly circumboreal. Habitat: freshwater, brackish.