Institute for Biodiversity Science and Sustainability
Online Version, Updated 5 August 2019

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Eschmeyer's Catalog of Fishes
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johanna, Crenicichla Heckel [J. J.] 1840:425 [Annalen des Wiener Museums der Naturgeschichte v. 2; ref. 2064] Rio Guaporé, Mato Grosso, Brazil. Lectotype: NMW 16431. Paralectotypes: ?SMF 5409 (1, dry). Type catalog: Kullander 1986:101 [ref. 12439]. Spelled joanna by Cope 1868:697 [ref. 945]. Kullander lists holotype, but Ploeg 1991:68 [ref. 20196] claims that the original description indicated more than one specimen; he designated the only specimen found at NMW as lectotype. •Valid as Crenicichla johanna Heckel 1840 -- (Ortega & Vari 1986:20 [ref. 9708], Kullander 1986:98 [ref. 12439], Ploeg 1991:68 [ref. 20196], Kullander 1997:285 [ref. 22746], Keith et al. 2000:168 [ref. 24547], Lasso et al. 2001:101 [ref. 25928], Kullander in Reis et al. 2003:628 [ref. 27061], Barriga S. 2014:118 [ref. 35745], Sarmiento et al. 2014:189 [ref. 35004], Kullander & Varella 2015:512 [ref. 33921], DoNascimiento et al. 2017:97 [ref. 35633], Varella & Ito 2018:11 [ref. 35884]). Current status: Valid as Crenicichla johanna Heckel 1840. Cichlidae: Cichlinae. Distribution: Widespread in South America: Bolivia, Brazil, Ecuador, Colombia, French Guiana, Guyana, Peru and Venezuela. Habitat: freshwater.