Institute for Biodiversity Science and Sustainability
Online Version, Updated 3 January 2020

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Eschmeyer's Catalog of Fishes
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lyolepis, Stolephorus Evermann [B. W.] & Marsh [M. C.] 1900:89, Fig. 13 [Bulletin of the U. S. Fish Commission v. 20 (pt 1) [for 1900]; ref. 14876] Culebra Island, Puerto Rico. Holotype: USNM 49528. Paratypes: (36) ?FMNH (5, not found). Type catalog: Ibarra & Stewart 1987:82 [ref. 12367]. •Valid as Anchoa lyolepis (Evermann & Marsh 1900) -- (Nelson 1986:895 [ref. 5735], Robins & Ray 1986:74 [ref. 23100], Whitehead et al. 1988:360 [ref. 5725], Cervigón 1991:110 [ref. 24472], Boschung 1992:43 [ref. 23239], Cervigón 1992:311 [ref. 23827], McEachran & Fechhelm 1998:353 [ref. 23897], Aguilera 1998:46 [ref. 24221], Castro-Aguirre et al. 1999:138 [ref. 24550], Smith-Vaniz et al. 1999:144 [ref. 25013], Schmitter-Soto et al. 2000:150 [ref. 27754], Camargo & Isaac 2001:140 [ref. 27639], Menezes & Figueiredo in Menezes et al. 2003:38 [ref. 27192] dated 1902, Nizinski & Munroe 2003:777 [ref. 26995], Nelson et al. 2004:66 [ref. 27807], Page et al. 2013:66 [ref. 32708], Smith-Vaniz & Jelks 2014:26 [ref. 33341], Raz-Guzmán et al. 2018:344 [ref. 36407], Spier et al. 2018:6 [ref. 36103], Ribeiro et al. 2019:5 [ref. 36717]). Current status: Valid as Anchoa lyolepis (Evermann & Marsh 1900). Engraulidae. Distribution: Western Atlantic. Habitat: marine.