Institute for Biodiversity Science and Sustainability
Online Version, Updated 1 March 2021

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Eschmeyer's Catalog of Fishes
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variabilis, Discognathus Heckel [J. J.] 1843:1069 [79] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Aleppo, Syria; Mosul, Iraq. Syntypes: NMW 53238-40 (3, 8, ca. 10), 53260-69 (1, 2, 2, 2, 3, 2, 2, 2, 2, 2), 53272 (4); SMF 403 [ex NMW in 1844] (4); ZMB 3301 [ex NMW] (3). Illustrated by Heckel 1843, Pl. 8 (fig. 1) [ref. 2067]. On p. 79 of separate [ref. 2067]. •Valid as Garra variabilis (Heckel 1843) -- (Coad 1991:17 [ref. 15702], Coad 1995:17 [ref. 23608], Coad 1998:102 [ref. 23869], Fricke et al. 2007:39 [ref. 29533], Coad 2010:180 [ref. 36620], Esmaeili et al. 2010:369 [ref. 33391], Hamidan et al. 2014:235 [ref. 33626], Sayyadzadeh et al. 2015:78 [ref. 34042], Jouladeh-Roudbar et al. 2015:871 [ref. 34062], Çiçek et al. 2015:144 [ref. 35151], Mousavi-Sabet & Eagderi 2016:45 [ref. 34341], Lyon et al. 2016:81 [ref. 34670], Freyhof 2016:500 [ref. 34805], Esmaeili et al. 2017:49 [ref. 35254], Esmaeili et al. 2018:28 [ref. 36089], Yoğurtçuoğlu et al. 2018:287 [ref. 36205], Bayçelebi 2020:753 [ref. 37964], Çiçek et al. 2020:248 [ref. 37644]). Current status: Valid as Garra variabilis (Heckel 1843). Cyprinidae: Labeoninae. Distribution: Asia Minor and Middle East: Tigris River basin (Turkey, Syria, Iraq). Habitat: freshwater.