Institute for Biodiversity Science and Sustainability
Online Version, Updated 4 December 2019

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Eschmeyer's Catalog of Fishes
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ustus, Callyodon Valenciennes [A.] in Cuvier & Valenciennes 1840:286, Pl. 405 [Histoire naturelle des poissons v. 14; ref. 4882] Brazil. Syntypes: MNHN 0000-2194 to 2195 (2, 1), A-8230 (1, dry), A-9290 (1, dry). Type catalog: Bauchot & GuibĂ© 1960:290-291 [ref. 20736]. Eastern Atlantic references are Nicholsina collettei, a separate species. •Synonym of Nicholsina collettei (Valenciennes 1840) -- (Westneat in Carpenter & De Angelis 2016:2736 [ref. 34618] from eastern Atlantic). •Valid as Nicholsina usta (Valenciennes 1840) -- (Robins & Ray 1986:208 [ref. 23100], Boschung 1992:157 [ref. 23239], Cervigón 1992:391 [ref. 23827], Bellwood 1994:63 [ref. 19116], Cervigón 1994:26 [ref. 24488], Aguilera 1998:52 [ref. 24221], Castro-Aguirre et al. 1999:399 [ref. 24550], Smith-Vaniz et al. 1999:298 [ref. 25013] as valid species and subspecies, Gushchin & Girardin 2000:103 [ref. 24893], Parenti & Randall 2000:54 [ref. 24943], Schmitter-Soto et al. 2000:164 [ref. 27754], Moura et al. 2001:519 [ref. 25371], Rocha & Rosa 2001:991 [ref. 25909], Westneat 2003:1728 [ref. 27105], Moura & Menezes in Menezes et al. 2003:93 [ref. 27192], Nelson et al. 2004:156 [ref. 27807], McEachran & Fechhelm 2005:532 [ref. 28815], Robertson et al. 2006:797 [ref. 31512], Wirtz et al. 2007:15 [ref. 30263], Parenti & Randall 2011:37 [ref. 31299], Page et al. 2013:160 [ref. 32708], Smith-Vaniz & Jelks 2014:63 [ref. 33341]). Current status: Valid as Nicholsina usta (Valenciennes 1840). Scaridae: Sparisomatinae. Distribution: Western Atlantic. Habitat: marine.