Institute for Biodiversity Science and Sustainability
Online Version, Updated 3 June 2019

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Eschmeyer's Catalog of Fishes
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asper, Cottus Richardson [J.] (ex Gairdner) 1837:295, 313, Pl. 95 (fig. 1) [Part 3; ref. 3731] Columbia River at Fort Vancouver, southern Washington, U.S.A. Holotype (unique): BMNH. Apparent holotype is at the BMNH (pers. comm. from C. Bond, 6/99). •Valid as Cottus asper Richardson 1837 -- (Lee et al. 1980:802 [ref. 22416], Eschmeyer & Herald 1983:167 [ref. 9277], Matarese et al. 1989:444 [ref. 26885], McAllister 1990:196 [ref. 14674], Page & Burr 1991:247 [ref. 18983], Swift et al. 1993:127 [ref. 22379], Scott & Crossman 1998:823 [ref. 24570], Fuller et al. 1999:339 [ref. 25838], Mecklenburg et al. 2002:469 [ref. 25968], Moyle 2002:346 [ref. 26449], Wydoski & Whitney 2003:203 [ref. 27816], Nelson et al. 2004:119 [ref. 27807], McPhail 2007:457 [ref. 31199], Scharpf 2008:7 [ref. 30399], Page & Burr 2011:472 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[9] [ref. 33456], Pietsch & Orr 2015:40 [ref. 34694], Kells et al. 2016:168 [ref. 35888]). Current status: Valid as Cottus asper Richardson 1837. Cottidae. Distribution: Eastern North Pacific/western North America: Seward, Alaska, to Ventura River, southern California, U.S.A. Habitat: freshwater, brackish, marine.