Institute for Biodiversity Science and Sustainability
Online Version, Updated 4 December 2019

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Eschmeyer's Catalog of Fishes
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quadricornis, Aspidophorus Valenciennes [A.] in Cuvier & Valenciennes 1829:221, Pl. 80 [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka, Russia. Holotype (unique): BMNH 2007.12.7.1 (very poor condition). Type catalog: Blanc & Hureau 1968:61 [ref. 20738]. Author is correctly Valenciennes; see Sheiko & Mecklenburg 2004:3 [ref. 27591] for comments on authorship. •Valid as Hypsagonus quadricornis (Valenciennes 1829) [as Cuvier 1829]-- (Eschmeyer & Herald 1983:188 [ref. 9277], Kanayama in Masuda et al. 1984:331 [ref. 6441], Lindberg & Krasyukova 1987:295 [ref. 15964], McAllister 1990:205 [ref. 14674], Kanayama 1991:17 [ref. 19261], Sheiko 1993:74 [ref. 21227], Busby 1998:13 [ref. 25508], Nakabo 2000:660 [ref. 25086], Sheiko & Fedorov 2000:30 [ref. 25650], Mecklenburg et al. 2002:531 [ref. 25968], Nakabo 2002:660 [ref. 26001], Youn 2002:272, 581 [ref. 26218], Parin et al. 2002:S98 [ref. 28300], Sheiko & Mecklenburg 2004:3 [ref. 27591], Nelson et al. 2004:122 [ref. 27807], Mecklenburg et al. 2011:125 [ref. 31212], Shinohara et al. 2011:45 [ref. 31715], Page et al. 2013:126 [ref. 32708], Shinohara et al. 2014:252 [ref. 33330], Parin et al. 2014:289 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Pietsch & Orr 2015:49 [ref. 34694], Mecklenburg et al. 2016:108 [ref. 34440], Kells et al. 2016:202 [ref. 35888], Mecklenburg et al. 2018:250 [reg. 36370]). Current status: Valid as Hypsagonus quadricornis (Valenciennes 1829). Agonidae: Hypsagoninae. Distribution: North Pacific and Arctic areas: southern Chukchi Sea to northern Okhotsk Sea, northern Kuril Islands, and Puget Sound, Washington. Habitat: marine.