Institute for Biodiversity Science and Sustainability
Online Version, Updated 4 November 2019

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Eschmeyer's Catalog of Fishes
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testudineus, Anthias Bloch [M. E.] 1792:121, Pl. 322 [Naturgeschichte der ausl√§ndischen Fische v. 6; ref. 470] "Japan" [actual is southeastern Asia]. Lectotype: ZMB 1370. Paralectotypes: ZMB 6578 (1, lost). Type catalog: Paepke 1994:313 [ref. 21516], Paepke 1999:44 [ref. 24282]. On p. 104 of Bloch's Ichthyologie, v. 9 [ref. 21381]. Spelled testudinaceus in Rendahl 1928:176 [ref. 3702]. Lectotype selected by Paepke 1994:313 [ref. 21516]. Sometimes dated to 1795. •Valid as Anabas testudineus (Bloch 1792) -- (Shrestha 1978:41 [ref. 15970], Feng in Kuang et al. 1986:322 [ref. 26639] dated 1795, Vierke 1986:60 [ref. 36452], Vierke 1988:89 [ref. 36453], Roberts 1989:171 [ref. 6439], Ataur Rahman 1989:284 [ref. 24860], Pethiyagoda 1991:258 [ref. 20075], Talwar & Jhingran 1991:996 [ref. 20764], Zhong in Pan et al. 1991:507 [ref. 23876], Kottelat et al. 1993:160 [ref. 23448], Indra 1994:425 [ref. 24015], Kottelat & Lim 1995:248 [ref. 23210], Sen 1995:590 [ref. 23778], Zhu 1995:185 [ref. 25213], Rainboth 1996:214 [ref. 22772], Rema Devi et al. 1996:143 [ref. 24021], Banik & Choudhury 1996:580 [ref. 24736], Wakiyama et al. 1997:100 [ref. 25519], Vu & Nguyen 1997:27 [ref. 26438], Kottelat 1998:115 [ref. 23436], Ng & Tan 1999:363 [ref. 24159], Rema Devi et al. 1999:163 [ref. 24758], Menon 1999:296 [ref. 24904], Fuller et al. 1999:468 [ref. 25838], Freyhof et al. 2000:97 [ref. 25184], Karmakar 2000:35 [ref. 25662], Kottelat 2001:62 [ref. 25482], Britz & Cambray 2001:280 [ref. 25647], Doi et al. 2001:17 [ref. 25691], Allen et al. 2000:160 [ref. 27467], Kottelat 2001:158 [ref. 25780], Raju Thomas et al. 2002:51 [ref. 26359], Yadav 2003:24 [ref. 27711], Ataur Rahman 2003:309 [ref. 31338], Tan & Lim 2004:111 [ref. 27716], Larson & Pidgeon 2004:197 [ref. 28107], Tan & Ng 2005:116 [ref. 28283], Parenti & Lim 2005:194 [ref. 28285], Shrestha 2008:226 [ref. 29923], Allen et al. 2008:197 [ref. 30265], Valis'eva & Vasil'ev 2012:664 [ref. 32477], Kottelat 2013:447 [ref. 32989], Kottelat 2015:48 [ref. 33971], Miesen et al. 2016:95 [ref. 34492], Zhang et al. 2016:208 [ref. 34477], Labatos 2017:5 [ref. 36239], Suresh et al. 2018:308 [ref. 36495], Sulaiman et al. 2018:37 [ref. 35802], Ng et al. 2019:534 [ref. 36606]). Current status: Valid as Anabas testudineus (Bloch 1792). Anabantidae. Distribution: South, Southeast and East Asia; India and Sri Lanka to Indonesia, Philippines and China; introduced widely elsewhere. Habitat: freshwater, brackish.