Institute for Biodiversity Science and Sustainability
Online Version, Updated 4 December 2019

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Eschmeyer's Catalog of Fishes
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aquila, Helcogramma Williams [J. T.] & McCormick [C. J.] 1990:1021, Fig. 3 [Copeia 1990 (no. 4); ref. 18992] White Beach, past Mahatae, Batan Island, Batanes, Philippines, 20°24'45"N, 121°55'02"E, depth 0-6 meters. Holotype: USNM 298405. Paratypes: AMS I.29427-001 (1); ANSP 164964 (1); BPBM 32855 (1); USNM 293945 (1), 297370 (4, 1 c&s), 298400 (4). Type catalog: Springer & Orrell 1996:24 [ref. 22274]. Species name should not be spelled aquilum -- see account of Helcogramma. •Valid as Helcogramma aquilum [sometimes as aquila] Williams & McCormick 1990 -- (Fricke 1997:409, 577 [ref. 23339], Fricke & Williams in Randall & Lim 2000:632 [ref. 25122], Williams & Fricke 2001:3534 [ref. 26288], Williams & Howe 2003:158 [ref. 27249], Myers & Donaldson 2003:638 [ref. 27495], Holleman 2006:102 [ref. 28864], Fricke 2009:48 [ref. 30394], Patzner et al. 2009:446 [ref. 30396] as aquilum, Allen & Erdmann 2012:772 [ref. 31980], Tashiro & Motomura 2014:103 [ref. 35481], Fricke & Erdmann 2017:41 [ref. 35190]). Current status: Valid as Helcogramma aquila Williams & McCormick 1990. Tripterygiidae: Tripterygiinae. Distribution: Western Pacific: Ryukyu Islands, Philippines and Guam. Habitat: marine.