markakulensis, Nemacheilus barbatulus Menshikov [M. I.] 1939:141, Fig. [Ucenye zapiski / Permskij Ordena Trudovogo Krasnogo Znameni Gosudarstvennyj Universitet Imeni A. M. Gor'kogo, Ministerstvo Vyssego i Srednego Special'nogo Obrazovanija RSFSR. v. 3 (no. 2); ref. 21170] Lake Marka-kul', Irtysh basin, Kazakhstan. Syntypes: ZIN 26864 (6). Author also seen as Men'shikov. •Synonym of Nemacheilus barbatulus (Linnaeus 1758) -- (Berg 1949:872 [ref. 20661] as Nemachilus). •Synonym of Noemacheilus barbatulus (Linnaeus 1758), but a valid subspecies markakulensis Manshikov 1939 -- (Mitrofanov & Dukravec 1989:60 [ref. 37748]). •Synonym of Barbatula toni (Dybowski 1869) -- (Kottelat 2006:53 [ref. 28831]). •Synonym of Orthrias barbatulus (Linnaeus 1758), subspecies toni (Dybowski 1869) -- (Prokofiev 2007:17 [ref. 30343]). •Synonym of Barbatula barbatula (Linnaeus 1758) -- (authors). •Valid as Barbatula markakulensis (Menshikov 1939) -- (Kottelat 2012:78 [ref. 32367] with question, Prokofiev 2015:461 [ref. 34065], Prokofiev 2016:396 [ref. 34693] as makakulensis, Prokofiev 2018:273 [ref. 35878]). Current status: Valid as Barbatula markakulensis (Menshikov 1939). Nemacheilidae. Distribution: North Asia: Markakul Lake (Kazakhstan). Habitat: freshwater.