messinicus, Gasterosteus aculeatus var. Stephanidis [A.] 1971:202, Fig. 10 [Biologia Gallo-Hellenica v. 3 (no. 2) (for 1970); ref. 7540] Not available, infrasubspecific; from Pamissos R. near Messini, Greece. •In the synonymy of Gasterosteus gymnurus Cuvier 1829 -- (Kottelat 1997:165 [ref. 22952]). •Synonym of Gasterosteus aculeatus Linnaeus 1758 -- (Paepke in Bănărescu & Paepke 2002:209 [ref. 25856], Denys et al. 2022:398 [ref. 40085]). Infrasubspecific. Current status: Synonym of Gasterosteus aculeatus Linnaeus 1758. Gasterosteidae. Habitat: freshwater, brackish, marine.