margaritaceus, Aspius Siebold [C. T. E. von] 1863:168 [Die Süsswasserfische von Mitteleuropa; ref. 4021] Not available, proposed for a hybrid. •Hybrid between Scardinius erythrophthalmus x Alburnus alburnus -- (Kottelat 1997:183 [ref. 22952]). Hybrid. Leuciscidae: Leuciscinae.