perlo, Squalus Bonnaterre [J. P.] (ex Broussonet) 1788:10 [Tableau encyclopédique et méthodique des trois règnes de la nature... Ichthyologie; ref. 4940] [La Méditerranée] France, northwestern Mediterranean Sea. No types known. Based on 'Le Perlon' of Broussonet 1784:668 [ref. 20282]. •Valid as Heptranchias perlo (Bonnaterre 1788) -- (Garrick & Paul 1971 [ref. 21295], Boeseman 1973:9 [ref. 7161], Nakaya in Okamura et al. 1982:41, 308 [ref. 8256], Uyeno & Sasaki in Uyeno et al. 1983:46 [ref. 14275], Boeseman in Whitehead et al. 1984:72 [ref. 13675], Compagno 1984:17 [ref. 6474], Nakaya in Okamura & Kitajima 1984:37, 293 [ref. 8057], Nakaya & Shirai in Masuda et al. 1984:3 [ref. 6441], Bass et al. 1986:45 [ref. 5638], Capapé 1987:22 [ref. 39326], Castro-Aguirre & Garcia-Dominguez 1988:101 [ref. 19199], Gubanov & Timoxin 1989:220 [ref. 36138], Paxton et al. 1989:26 [ref. 12442], Paulin et al. 1989:11 [ref. 24556], Pequeño 1989:11 [ref. 14125], Nakaya in Amaoka et al. 1990:55 [ref. 16024], Springer 1990:4 [ref. 19315], Compagno et al. 1991:51 [ref. 21083], Bianchi & Carpenter in Bianchi et al. 1993:72 [ref. 25472] with author as Rafinesque, Gomon et al. 1994:87 [ref. 22532], Last & Stevens 1994:39 [ref. 23873], Herman et al. 1994:149 [ref. 27820], Santos et al. 1997:11 [ref. 23531], Arruda 1997:15 [ref. 24952], Adam et al. 1998:5 [ref. 23610], Compagno 1998:1210 [ref. 23785], McEachran & Fechhelm 1998:99 [ref. 23897], Chirichigno F. & Vélez D. 1998:31 [ref. 24555], Uiblein et al. 1999:59 [ref. 24045], Cervigón & Alcalá 1999:24 [ref. 24490], Compagno 1999:471 [ref. 25589], Nakabo 2000:142 [ref. 25086], Compagno in Randall & Lim 2000:577 [ref. 25122], Henderson & Williams 2001:707 [ref. 25602], Hutchins 2001:13 [ref. 25847], Shinohara et al. 2001:288 [ref. 25995], Soto 2001:65, 90 [ref. 26637], Bilecenoğlu et al. 2002:8 [ref. 26753], Nakabo 2002:142 [ref. 26001], Choi et al. 2003:59, 477 [ref. 26218], Compagno 2003:375 [ref. 26984], Gadig & Gomes in Menezes et al. 2003:25 [ref. 27192], Moore et al. 2003:168 [ref. 27076], Manilo & Bogorodsky 2003:S91 [ref. 27377], Espinosa Pérez et al. 2004:87 [ref. 27705], Nelson et al. 2004:49 [ref. 27807], McLaughlin & Morrissey 2004:482 [ref. 28052], Heemstra & Heemstra 2004:52 [ref. 28072], Shinohara et al. 2005:394 [ref. 28370], Compagno et al. 2005:47 [ref. 29145], Golani 2005:8 [ref. 37112], Hoese & Gates 2006:135 [ref. 28997], Fricke et al. 2007:15 [ref. 29533], White 2008:76 [ref. 30617], Fricke et al. 2009:11 [ref. 30213], Lipej & Dulčić 2010:5 [ref. 36649], Fricke et al. 2011:346 [ref. 31242], Castro 2011:28 [ref. 31457], Shinohara et al. 2011:38 [ref. 31715], Yamashita et al. 2012:126 [ref. 32408], Page et al. 2013:52 [ref. 32708], Ebert et al. 2013:281 [ref. 33045], Shinohara et al. 2014:233 [ref. 33330], Dyldin 2015:42 [ref. 34524], Duffy & Ahyong 2015:105 [ref. 36167], Stewart 2015:116 [ref. 34217], Del Moral-Flores et al. 2016:55 [ref. 34398], Weigmann 2016:51 [ref. 34211], Compagno 2016:1148 [ref. 34544], Vieira et al. 2016:[4] [ref. 34869], Nión et al. 2016:21 [ref. 35565], Biscoito et al. 2018:462 [ref. 37562], Ehemann et al. 2018:23 [ref. 36194], Fricke et al. 2018:18 [ref. 35805], Kumar et al. 2018:290 [ref. 36178], Almeida & Biscoito 2019:91 [ref. 37302], Carneiro et al. 2019:15 [ref. 37250], Ehemann et al. 2019:[3] [ref. 36861], Miyazaki et al. 2019:122 [ref. 37957], Reiner 2019:17 [ref. 38139], Bariche & Fricke 2020:14 [ref. 37515], Kabasakal 2020:39 [ref. 37952], Kim et al. 2020:41 [ref. 39118], Kovačić et al. 2020:4 [ref. 37519], Psomadakis et al. 2020:160 [ref. 37272], Serena et al. 2020:502 [ref. 37963], Sonoyama et al. 2020:10 [ref. 37637], Ebert et al. 2021:18 [ref. 38234], Giovos et al. 2021:5 [ref. 38895], Golani 2021:6 [ref. 38303], González-Acosta et al. 2021:6 [ref. 38361], Kovačić et al. 2021:8 [ref. 39014], Bañón & Mano 2022:3 [ref. 40121], Barone et al. 2022:7 [ref. 39527], Dragičević & Isajlović in Montesanto et al. 2022:982 [ref. 39977], Ebert 2022:412 [ref. 39657], Fermon et al. 2022:90 [ref. 39448], Gloerfelt-Tarp & Kailola 2022:1 [ref. 41223], Lipej & Mavrič in Kousteni et al. 2022:430 [ref. 39260], Balàka et al. 2023:7 [ref. 40410], Carvalho-Filho 2023:292 [ref. 40480], Castillo Domínguez et al. 2023:4 [ref. 40663], Page et al. 2023:36 [ref. 40505], Shakman et al. 2023:463 [ref. 40810], Deval in Digenis et al. 2024:102 [ref. 40958], Fricke et al. 2024:58 [ref. 41622], Gajić 2024:3 [ref. 41088], Leeney 2024:11 [ref. 40985]). Current status: Valid as Heptranchias perlo (Bonnaterre 1788). Hexanchidae. Distribution: Circumglobal in tropical through warm temperate sea (including Gulf of Mexico, Caribbean Sea, Mediterranean Sea, Sea of Japan). IUCN (2020): Near Threatened. Habitat: marine.