Institute for Biodiversity Science and Sustainability
Online Version, Updated 4 December 2019

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Eschmeyer's Catalog of Fishes
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paulus, Cottus Williams [J. D.] 2000:302 [Copeia 2000 (no. 1); ref. 24258] Coldwater Spring, 5.7 miles west of Oxford about 500 yards north of U.S. highway 78, tributary of Coosa River, Calhoun County, Alabama, U.S.A. Holotype: UMMZ 187462. Paratypes: ALA 2031 (12), 1138 (39), 1665 (68); CUMV 52064 [ex ALA 1063] (26); TU 34396 (47); UMMZ 187463 (25); USNM 201404 [ex ALA 1063] (26). Replacement name for Cottus pygmaeus Williams 1968, preoccupied by Cottus quadricornis pygmaeus Lönnberg 1932. •Valid as Cottus paulus Williams 2000 -- (Kinziger et al. 2000:1007 [ref. 25019], Nelson et al. 2004:119 [ref. 27807], Boschung & Mayden 2004:424 [ref. 27995], Neely et al. 2007:651 [ref. 29262], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:476 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus paulus Williams 2000. Cottidae. Distribution: Coldwater Spring, Coosa River system, Alabama, U.S.A. Habitat: freshwater.