Institute for Biodiversity Science and Sustainability
Online Version, Updated 3 September 2019

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Eschmeyer's Catalog of Fishes
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ungulus, Creagrutus Vari [R. P.] & Harold [A. S.] 2001:209, Fig. 90 [Smithsonian Contributions to Zoology No. 613; ref. 25842] Quebrada Soga, tributary Of Río Madre de Dios, 1 kilometer upstream from Erika, opposite Salvacion, about 12°53'S, 71°12'W, Provincia Manu, Madre de Dios, Peru. Holotype: MUSM 8878. Paratypes: MNRJ 14481 (6); MUSM 8879-80 (7, 4); USNM 302788 (3), 303066 (10, 3 c&s). •Valid as Creagrutus ungulus Vari & Harold 2001 -- (Vari & Harold in Reis et al. 2003:126 [ref. 27061], Mirande et al. 2013:763 [ref. 33366], Thomaz et al. 2015:Add. 5 [ref. 33893]). Current status: Valid as Creagrutus ungulus Vari & Harold 2001. Characidae: Stevardiinae. Distribution: Madre de Dios River basin, Peru. Habitat: freshwater.