Institute for Biodiversity Science and Sustainability
Online Version, Updated 6 April 2020

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Eschmeyer's Catalog of Fishes
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umbratile, Cephaloscyllium Jordan [D. S.] & Fowler [H. W.] 1903:602, Fig. 1 [Proceedings of the United States National Museum v. 26 (no. 1324); ref. 2460] Nagasaki, Japan, 32°43'N, 129°50'E. Holotype (unique): CAS-SU 12693 (stuffed dry skin). Type catalog: Böhlke 1953:8-9 [ref. 12291]. •Questionably a synonym of Cephaloscyllium isabellum (Bonnaterre 1788) -- (Compagno 1984:298 [ref. 6846]). •Valid as Cephaloscyllium umbratile Jordan & Fowler 1903 -- (Nakaya in Masuda et al. 1984:4 [ref. 6441], Nakaya in Okamura & Kitajima 1984:39, 294 [ref. 8057], Chen et al. 1997:2 [ref. 26476], Cheng & Zhou 1997:26 [ref. 26385], Compagno 1999:479 [ref. 25589], Nakabo 2000:129 [ref. 25086], Compagno in Randall & Lim 2000:579 [ref. 25122], Shinohara et al. 2001:287 [ref. 25995], Nakabo 2002:129 [ref. 26001], Choi et al. 2003:51, 470 [ref. 26218], Shinohara et al. 2005:392 [ref. 28370], Inoue & Nakaya 2006:86 [ref. 28852], Schaaf-Da Silva & Ebert 2008:13 [ref. 29807], Ho & Shao 2011:17 [ref. 31432], Shinohara et al. 2011:38 [ref. 31715], Yamashita et al. 2012:121 [ref. 32408], Naylor et al. 2012:49 [ref. 32267] see remarks, Ebert et al. 2013:317 [ref. 33045], Akhilesh et al. 2014:4 [ref. 33217], Shinohara et al. 2014:232 [ref. 33330], Dyldin 2015:49 [ref. 34524], Weigmann 2016:37-38 [ref. 34211]). Current status: Valid as Cephaloscyllium umbratile Jordan & Fowler 1903. Scyliorhinidae. Distribution: Western North Pacific: Taiwan, Hong Kong, Gulf of Tongking, Vietnam, Malaysia. Habitat: marine.