Institute for Biodiversity Science and Sustainability
Online Version, Updated 3 September 2019

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Eschmeyer's Catalog of Fishes
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tutul, Himantura Borsa [P.], Durand [J.-D.], Shen [K.-N.], Alyza [I. A.], Solihin [D. D.] & Berrebi [P.] 2013:89, Fig. 5 [Comptes Rendus Biologies v. 336; ref. 32736] Mkoani fish landing, Pemba Island, Tanzania. Holotype: MNHN ICTI-5184 (2 sections of tail). Paratypes: CSIRO H 5284.05 (2). Plus non-type specimens. See also Borsa 2016:[1] 1170 [ref. 35022] and Weigmann 2016:4 [ref. 35023]. This species has been confused with Himantura leoparda Manjaji-Matsumoto & Last 2008 (Borsa 2016:[2]1171 [ref. 35022]). •Valid as Himantura tutul Borsa, Durand, Shen, Alyza, Solihin & Berrebi 2013 -- (Borsa 2016:[4] 1173 [ref. 35022], Miesen et al. 2016:80 [ref. 34492], Fernando et al. 2019:208 [ref. 36642]). •Synonym of Himantura uarnak (Gmelin 1789) -- (Last et al. 2016:563 [ref. 35010], Weigmann 2016:[4] 1179 [ref. 34211] see remarks, Weigmann 2016:1177 [ref. 35023]). Current status: Synonym of Himantura uarnak (Gmelin 1789). Dasyatidae: Urogymninae. Distribution: Indian Ocean: East Africa east to Indonesia (if valid). Habitat: brackish, marine.