Institute for Biodiversity Science and Sustainability
Online Version, Updated 6 July 2020

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Eschmeyer's Catalog of Fishes
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billi, Helcogramma Hansen [P. E. H.] 1986:329, Fig. 10 [Bulletin of Marine Science v. 38 (no. 2); ref. 5810] Rocky area opposite Fort Frederick, Trincomalee (outside harbor), Sri Lanka. Holotype: USNM 222377. Paratypes: USNM 222367 (6, 1 c&s), 222368 (1), 222374 (3), 222380 (17, not 27). Type catalog: Springer & Orrell 1996:25 [ref. 22274]. •Valid as Helcogramma billi Hansen 1986 -- (Fricke 1997:577 [ref. 23339], Manilo & Bogorodsky 2003:S116 [ref. 27377], Holleman 2007:58 [ref. 29195], Fricke 2009:48 [ref. 30394], Patzner et al. 2009:446 [ref. 30396]). Current status: Valid as Helcogramma billi Hansen 1986. Tripterygiidae: Tripterygiinae. Distribution: Northern Indian Ocean: Sri Lanka to southern India. Habitat: marine.