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Eschmeyer's Catalog of Fishes
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Species in the family or subfamily Cottidae: [ 533 ] records


abyssalis, Asprocottus herzensteini Taliev [D. N.] 1955:322, Figs. 128-129 [ ref. 4316] Lake Baikal, Russia, depth below 450 meters. Syntypes: ZIN (none found in 1996). Taliev specimens: ZIN 46867 (1), ? 46633 (1). Appeared without distinguishing features and not available in Taliev in Taliev & Koryakov 1947:1838 [ref. 6543]. Sideleva 2001:58 [ref. 26181] names a neotype [ZIN 46633], but if that is the first time a neotype was named it is unavailable under the Code. That neotype repeated in Sideleva 2003:181 [ref. 27815]. •Valid as Asprocottus abyssalis Taliev 1955 -- (Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702] with authorship and date as Taliev & Korjakov 1947, Sideleva 1998:155 [ref. 23595], Bogutskaya et al. 2001:47 [ref. 26178] with authorship and date as Taliev & Korjakov 1947, Sideleva 2001:61 [ref. 26181] with authorship as Taliev & Korjakov 1947, Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:17, 181 [ref. 27815] with authorship as Taliev & Korjakov 1947, Bogutskaya & Naseka 2004:194 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Asprocottus abyssalis Taliev 1955. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

acuticeps, Oligocottus Gilbert [C. H.] 1896:432 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Unalaska harbor, Unalaska, Aleutian Islands. Syntypes: (4) CAS-SU 3027 (2), USNM 48608 (1). Additional material: (3) Vancouver I. Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Clinocottus acuticeps (Gilbert 1896) -- (Bolin 1944:84 [ref. 6379], Lee et al. 1980:800 [ref. 22416], Eschmeyer & Herald 1983:165 [ref. 9277], Matarese et al. 1989:422 [ref. 26885], McAllister 1990:196 [ref. 14674], Mecklenburg et al. 2002:502 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:39 [ref. 34694], Kells et al. 2016:166 [ref. 35888]). Current status: Valid as Clinocottus acuticeps (Gilbert 1896). Cottidae. Distribution: Eastern North Pacific: Aleutian Islands to central California, U.S.A. Habitat: freshwater, brackish, marine.

aenaeus, Cottus Mitchill [S. L.] 1814:8 [Report, in part, of Samuel L. Mitchill; ref. 3030] New York, U.S.A. No types known. Appeared as Cottus aeneus in Mitchill 1815:380 [ref. 13292]. •Valid as Myoxocephalus aenaeus (Mitchill 1814) -- (Neyelov 1979:123 [ref. 3152], Smith 1986:449 [ref. 22186], Robins & Ray 1986:282 [ref. 23100] as aeneus, Scott & Scott 1988:500 [ref. 25518], McAllister 1990:200 [ref. 14674], Klein-MacPhee in Collette & Klein-MacPhee 2002:349 [ref. 26158], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Afonso et al. 2016:46, 225 [ref. 36199]). Current status: Valid as Myoxocephalus aenaeus (Mitchill 1814). Cottidae. Distribution: Northwestern Atlantic. Habitat: brackish, marine.

affinis, Centridermichthys Steindachner [F.] & Döderlein [L.] 1887:257 [1] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 53 (1. abth.); ref. 4249] Tango, Japan Sea and Kanagawa, Japan. Syntypes: (4) NMW. •Synonym of Pseudoblennius cottoides (Richardson 1848) -- (Watanabe 1960:152 [ref. 11959], Lindberg & Krasyukova 1987:252 [ref. 15964], Kim & Youn 1992:74 [ref. 22287]). Current status: Synonym of Pseudoblennius cottoides (Richardson 1848). Cottidae. Habitat: marine.

affinis, Cottus Heckel [J. J.] 1839:150 [Annalen des Wiener Museums der Naturgeschichte v. 2 (no. 1); ref. 2079] Mörkö Island, near Stockholm, Sweden. Syntypes: whereabouts unknown. •Synonym of Cottus gobio Linnaeus 1758 -- (Kottelat 1997:169 [ref. 22952], Freyhof et al. 2005:124, 132 [ref. 28259]). Current status: Synonym of Cottus gobio Linnaeus 1758. Cottidae. Habitat: freshwater, brackish.

albomaculatus, Crossias Schmidt [P. Yu.] 1916:619, Fig. 4 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 3935] Medny Island, Commander Islands, Bering Sea. Lectotype: ZIN 18966. Paralectotypes: ZIN 18964 (lost), 18965 (1), 19193 (2), ?21866 (?). Lectotype established by Neyelov 1979:136 [ref. 3152]. •Synonym of Porocottus bradfordi Rutter 1898, but a valid subspecies -- (Neyelov 1979:136 [ref. 3152], Muto et al. 2002:231 [ref. 26331]). •Synonym of Porocottus mednius (Bean 1898) -- (Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:486 [ref. 25968], Parin et al. 2002:S84 [ref. 28300], Parin et al. 2014:269 [ref. 33547]). Current status: Synonym of Porocottus mednius (Bean 1898). Cottidae. Habitat: marine.

alcicornis, Centridermichthys Herzenstein [S. M.] 1890:115 [25] [Mélanges Biologiques, tirés du Bulletin physico-mathématique de l'Académie Impériale des Sciences de St. Pétersbourg v. 13; ref. 2149] Tokyo, Japan. Holotype (unique): ZIN 8718. On p. 25 in Bull. Acad. Imp. Sci. St. Pétersb. (n. s.) v. 2. •Valid as Alcichthys alcicornis (Herzenstein 1890) -- (Watanabe 1960:131 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Kim & Youn 1992:70 [ref. 22287], Shinohara et al. 1996:175 [ref. 26605], Sokolovskaya et al. 1998:9 [ref. 24670], Youn 2002:262, 573 [ref. 26218]). •Synonym of Alcichthys elongatus (Steindachner 1881) -- (Parin et al. 2002:S68 [ref. 28300], Parin et al. 2014:233 [ref. 33547]). Current status: Synonym of Alcichthys elongatus (Steindachner 1881). Cottidae. Habitat: marine.

aleuticus, Cottus Gilbert [C. H.] 1896:418 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Streams at Iliuliuk, Unalaska, Bering Sea. Syntypes: ANSP 39139 (1); BMNH 1896.7.23.59 (1), 1900.7.11.2 (1); MCZ 28368 (1); ?NMW; CAS-SU 2610 (19); USNM 48716 (1); ZMUO J1417 (1). Type catalog: Böhlke 1953:128 [ref. 12291], Pethon 1969:7 [ref. 19268], Böhlke 1984:65 [ref. 13621]. •See Coffie 1998:126 [ref. 24964]. •Valid as Cottus aleuticus Gilbert 1896 -- (Lee et al. 1980:801 [ref. 22416], McAllister 1990:196 [ref. 14674], Page & Burr 1991:247 [ref. 18983], Swift et al. 1993:129 [ref. 22379], Chereshnev 1996:605 [ref. 24659], Scott & Crossman 1998:820 [ref. 24570], Mecklenburg et al. 2002:470 [ref. 25968], Moyle 2002:349 [ref. 26449], Wydoski & Whitney 2003:201 [ref. 27816], Nelson et al. 2004:119 [ref. 27807], McPhail 2007:450 [ref. 31199], Scharpf 2008:7 [ref. 30399], Page & Burr 2011:471 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[9] [ref. 33456], Pietsch & Orr 2015:40 [ref. 34694], Kells et al. 2016:168 [ref. 35888]). Current status: Valid as Cottus aleuticus Gilbert 1896. Cottidae. Distribution: Eastern North Pacific: Bristol Bay, Alaska Peninsula, and Aleutian Islands to California. Habitat: freshwater, brackish.

alexandrae, Cottocomephorus grewingki Taliev [D. N.] 1935:64, Pl. 1 (fig. 4) [Trudy Baikal'skoi limnologischeskoi stantsii = Travaux de la Station limnologique du Lac Baikal v. 6; ref. 15741] Lake Baikal near Marituy village, Russia, depth 500 meters. Syntypes: (7, ex Baikal Limnological Station) ZIN (none found in 1996 and not in The Baikal Museum). Original grewingki should have been grewingkii. Sideleva 2001:58 [ref. 26181] and 2003:165 [ref. 27815] names a lectotype [ZIN 36608] (in 2006 called neotype), but that specimen was not captured until 1949; it would probably make the best neotype if one is needed. •Synonym of Cottocomephorus grewingkii (Dybowski 1874), but a valid subspecies as described -- (Taliev 1955:287 [ref. 4316], Sideleva 1998:150 [ref. 23595]). •Valid as Cottocomephorus alexandrae Taliev 1935 -- (Bogutskaya et al. 2001:46 [ref. 26178], Sideleva 2001:58 [ref. 26181], Sideleva 2003:16, 165 [ref. 27815], Bogutskaya & Naseka 2004:192 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). •Synonym of Cottocomephorus grewingkii (Dybowski 1874) -- (Berg 1949:1173 [ref. 20662] as grewingki, Bogdanov 2019:231 [ref. 36814]). Current status: Synonym of Cottocomephorus grewingkii (Dybowski 1874). Cottidae. Habitat: freshwater.

allisi, Crossias Jordan [D. S.] & Starks [E. C.] 1904:296, Fig. 28 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate, Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7711. Paratypes: AMS I.6915 (6); BMNH 1905.2.4.385-391 (7); CAS-SU 7579 (34, now 32); USNM 50922 (19), 51240 (2). Type catalog: Böhlke 1953:128 [ref. 12291]. •Valid as Crossias allisi Jordan & Starks 1904 -- (Watanabe 1960:162 [ref. 11959]). •Valid as Porocottus allisi (Jordan & Starks 1904) -- (Neyelov 1979:136 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:223 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:646 [ref. 25086], Mecklenburg et al. 2002:404 [ref. 25968], Nakabo 2002:646 [ref. 26001], Youn 2002:263, 577 [ref. 26218], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S83 [ref. 28300], Parin et al. 2014:268 [ref. 33547]). Current status: Valid as Porocottus allisi (Jordan & Starks 1904). Cottidae. Distribution: Western North Pacific. Habitat: marine.

altaicus, Cottus poecilopus Kaschenko [N.] 1899:151 [Results of the Altai Zoological Expedition; ref. 19630] Katun River, Bol'shoi Uiman, Katun River basin, Altai, Siberia, Russia. Neotype: ZIN 53968. Syntypes: (15) whereabouts unknown. Syntypes (15) lost; neotype selected by Sideleva & Goto 2009:601 [ref. 30591]. •Synonym of Cottus poecilopus Heckel 1837 -- (Freyhof et al. 2005:167 [ref. 28259]). •Valid as Cottus altaicus Kaschenko 1899 -- (Bogutskaya & Naseka 2004:185 [ref. 28183], Kottelat 2006:90 [ref. 28831], Sideleva & Goto 2009:601 [ref. 30591], Goto et al. 2014:[9] [ref. 33456], Parin et al. 2014:241 [ref. 33547], Romanov et al. 2017:826 [ref. 35687]). Current status: Valid as Cottus altaicus Kaschenko 1899. Cottidae. Distribution: Irtysh River basin, Altai, western Siberia, Russia. Habitat: freshwater, brackish.

altaicus, Cottus sibiricus Li [S.-C.] & Ho [C.-W.] in Li, Tai, Chang, Ma, Ho & Kao 1966:49 (English p. 55), Fig. 2 [Acta Zoologica Sinica v. 18 (no. 1); ref. 9042] Altai, northern Sinkiang, China. Holotype: ASZIB [now NZMC] 56824 [ex NK 64-997]. Paratypes: ASZIB [now NZMC] 56825 [ex ZB 64-1085]; ZB 64-979 to 980 (2), 64-983 to 985 (3), 64-988 (1), 64-992 (1); Acad. Sinica, Sinkiang, Bureau of Water Conservancy and Dept. Biol. Sinkiang Univ. 64-981 to 982 (2), 64-986 to 987 (2), 64-989 to 991 (3). Type catalog: Ye et al. 2015:96 [ref. 33808]. Objectively invalid; preoccupied by Cottus poecilopus altaicus Kashchenko 1899. Replaced by Cottus dzungaricus Kottelat 2006:60 [ref. 28831]. •Incorrectly treated as valid -- (Zhu 1995:189 [ref. 25213]). •Synonym of Cottus dzungaricus Kottelat 2006 -- (Ye et al. 2015:96 [ref. 33808]). •Synonym of Cottus sibiricus, subspecies altaicus Li & Ho 1966 -- (Zhang et al. 2016:193 [ref. 34477] as altieus). Current status: Synonym of Cottus dzungaricus Kottelat 2006. Cottidae. Habitat: freshwater.

alvordii, Cottus Girard [C. F.] 1851:46, Pl. 1 (figs. 7-8) [Smithsonian Contributions to Knowledge v. 3 (art. 3) (for 1850); ref. 18124] Fort Gratiot, Lake Huron, Michigan, U.S.A. Holotype (unique): USNM. •Synonym of Cottus bairdii Girard 1850. Current status: Synonym of Cottus bairdii Girard 1850. Cottidae. Habitat: freshwater.

amblystomopsis, Cottus Schmidt [P. Yu.] 1904:89, Pl. 2 (figs. 1-3) [Fishes of the eastern seas of the Russian Empire; ref. 3946] Lyutoga River, Sakhalin Island, Russia. Holotype (unique): ZIN 12763. •Valid as Cottus amblystomopsis Schmidt 1904 -- (Berg 1949:1142 [ref. 20662], Yabe in Masuda et al. 1984:326 [ref. 6441], Lindberg & Krasyukova 1987:171 [ref. 15964], Andoh & Goto 1988:176 [ref. 22495], Kawanabe & Mizuno 1989:660 [ref. 25598], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:150 [ref. 23595], Nakabo 2000:638 [ref. 25086], Goto et al. 2001:70 [ref. 25596], Pietsch et al. 2001:144 [ref. 25688], Bogutskaya et al. 2001:46 [ref. 26178], Nakabo 2002:638 [ref. 26001], Parin et al. 2002:S72 [ref. 28300], Bogutskaya & Naseka 2004:186 [ref. 28183], Fujii et al. 2005:12 [ref. 28191], Goto et al. 2014:[9] [ref. 33456], Parin et al. 2014:241 [ref. 33547], Dyldin & Orlov 2017:62 [ref. 35138]). Current status: Valid as Cottus amblystomopsis Schmidt 1904. Cottidae. Distribution: Western North Pacific. Habitat: freshwater, brackish, marine.

anahaze, Pseudoblennius Bleeker [P.] 1879:11 [Verhandelingen der Koninklijke Akademie van Wetenschappen, Afdeeling Natuurkunde (Amsterdam) v. 18; ref. 460] Unexplained new name for Pseudoblennius pseudoclinus Temminck & Schlegel [see under Pseudoblennius] and Pseudoblennius percoides Günther 1861. •Synonym of Pseudoblennius percoides Günther 1861. Current status: Synonym of Pseudoblennius percoides Günther 1861. Cottidae. Habitat: marine.

analis, Oligocottus Girard [C. F.] 1858:201 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 9 (no. 15) (for 1857); ref. 1811] Monterey, California, U.S.A. Syntypes: FMNH 210 (1); USNM 486 (orig. 7, now 1). •Valid as Clinocottus analis (Girard 1858) -- (Bolin 1944:73 [ref. 6379], Eschmeyer & Herald 1983:166 [ref. 9277], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Kells et al. 2016:166 [ref. 35888]). Current status: Valid as Clinocottus analis (Girard 1858). Cottidae. Distribution: Eastern North Pacific: northern California to central Baja California. Habitat: marine.

anceps, Cottus (Acanthocottus) Sauvage [H.-E.] 1878:145, Pl. 1 (fig. 13) [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris (Série 2) v. 1; ref. 3880] U.S.A. Syntypes: MNHN 0000-7241 (1), 0000-7243 (1). Type catalog: Blanc & Hureau 1968:50 [ref. 20738]. •Synonym of Myoxocephalus aenaeus (Mitchill 1814) -- (Smith 1986:450 [ref. 22186]). Current status: Synonym of Myoxocephalus aenaeus (Mitchill 1814). Cottidae. Habitat: brackish, marine.

annae, Cottus Jordan [D. S.] & Starks [E. C.] in Jordan 1896:223, Pl. 27 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 2395] Eagle River at Gypsum, Colorado, U.S.A. Syntypes: CAS-SU 1305 and 1308-10 [but all in 1309] (4). Type catalog: Böhlke 1953:128 [ref. 12291]. Lectotype established in Jordan 1896:244 [ref. 2395] if specimen is identifiable. •Synonym of Cottus beldingii Eigenmann & Eigenmann 1891 -- (Bailey & Bond 1963:1 [ref. 3934], Lee et al. 1980:806 [ref. 22416]). Current status: Synonym of Cottus beldingii Eigenmann & Eigenmann 1891. Cottidae. Habitat: freshwater.

anoplus, Thyriscus Gilbert [C. H.] & Burke [C. V.] 1912:43, Fig. 4 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off Attu Island, Bering Sea, Albatross station 4782, depth 57 fathoms. Holotype (unique): USNM 74368. •Valid as Thyriscus anoplus Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:450 [ref. 25968], Parin et al. 2002:S87 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:274 [ref. 33547]). Current status: Valid as Thyriscus anoplus Gilbert & Burke 1912. Cottidae. Distribution: North Pacific: Aleutian Islands and Commander Islands to Kuril Islands. Habitat: marine.

ansatus, Centridermichthys Richardson [J.] 1845:74, Pl. 54 (figs. 6-10) [Ichthyology.--Part 2. The zoology of the voyage of H. M. S. Sulphur; ref. 20574] Harbor of Woosung, mouth of the Yangtze Kiang Kew, northern China, 31°N. Syntypes: BMNH 1865.5.13.- (2). •Placed in synonymy of Cottus uncinatus Temminck & Schlegel 1843 by Richardson 1845 (Oct.):150 [ref. 20575]. •Synonym of Trachidermus fasciatus Heckel 1839 -- (Okada 1961:709 [ref. 6363], Youn 2002:573 [ref. 26218]). •Not treated in Zhang et al. 2016:194 [ref. 34477]. Current status: Synonym of Trachidermus fasciatus Heckel 1839. Cottidae. Habitat: freshwater, brackish, marine.

antilope, Artediellus Schmidt [P. Yu.] 1937:572 [Comptes Rendus (Doklady) de l'Académie des Sciences de l'U.R.S.S. v. 15 (no. 9); ref. 15302] 55°31'N, 149°10'E, Okhotsk Sea, depth 335 meters. Lectotype: ZIN 24463 (female). Paralectotypes: ZIN 24463 (2 males). Lectotype selected by Neyelov 1979:160 [ref. 3152]. Appeared first as name only in Schmidt 1935. •Valid as Artediellina antilope (Schmidt 1937) -- (Neyelov 1979:160 [ref. 3152], Sheiko & Fedorov 2000:27 [ref. 25650], Parin et al. 2002:S69 [ref. 28300], Fedorov et al. 2003:65 [ref. 27579], Parin et al. 2014:235 [ref. 33547]). Current status: Valid as Artediellina antilope (Schmidt 1937). Cottidae. Distribution: Western North Pacific. Habitat: marine.

aporosus, Artediellus Soldatov [V. K.] 1922:321 [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23; ref. 15430] Tatar Strait, northern Sea of Japan; Cape Bellingshausen and Cape Terpeniya, eastern Sakhalin Island, Okhotsk Sea. Lectotype: ZIN 20613 (1 of 6). Paralectotypes: (15) ZIN 12907 (1), 12909 (3), 20613 (5 of 6). Possibly published in 1921. Lectotype selected by Neyelov 1979:154 [ref. 3152]. •Valid as Artediellus aporosus Soldatov 1922 -- (Neyelov 1979:154 [ref. 3152], Nelson 1986:43 [ref. 5808], Lindberg & Krasyukova 1987:232 [ref. 15964], Sheiko & Fedorov 2000:27 [ref. 25650], Parin et al. 2002:S69 [ref. 28300], Fedorov et al. 2003:65 [ref. 27579], Inagawa & Yabe 2006:192 [ref. 28854], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:235 [ref. 33547]). Current status: Valid as Artediellus aporosus Soldatov 1922. Cottidae. Distribution: Western North Pacific: Northern Sea of Japan, Okhotsk Sea; Russia and Japan. Habitat: marine.

arachlensis, Paracottus kessleri Tarkhova [Yu. N.] 1962:103 [Kratkiie soobscenjia Burjatskij Kompleksnogo Naucno-Issledovatel'skogo Instituta, Akademija Nauk SSR, Sibirskoe Otdelenie, Burjatskij Kompleksnyi Nauno-Issledovatel'skij Institut. No. 3; ref. 22756] Lake Arakhlei, Lake Baikal Basin, Russia. Neotype: ZIN 34220. Original not seen. Neotype designated by Sideleva 2003:154 [ref. 27815]. •Synonym of Cottus kesslerii Dybowski 1874, but a valid subspecies -- (Sideleva 1982 [ref. 14469], Sideleva 1998:152 [ref. 23595]). •Synonym of Leocottus kesslerii (Dybowski 1874), but a valid subspecies arahlensis (Tarkhova 1962) as described -- (Sideleva 2001:60 [ref. 26181], Sideleva 2003:154 [ref. 27815], Kottelat 2006:60 [ref. 28831]). Current status: Synonym of Leocottus kesslerii (Dybowski 1874). Cottidae. Habitat: freshwater.

argenteus, Centridermichthys Döderlein [L.] in Steindachner & Döderlein 1887:257 [1] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 53 (1. abth.); ref. 4249] Tokyo Bay, Japan. Holotype (unique): NMW 76418. •Valid as Pseudoblennius argenteus (Döderlein 1887) -- (Nakabo 2000:649 [ref. 25086], Nakabo 2002:649, 1526 [ref. 26001]). Current status: Valid as Pseudoblennius argenteus (Döderlein 1887). Cottidae. Distribution: Western North Pacific. Habitat: marine.

armata, Ochotskia Schmidt [P. Yu.] 1916:612, Fig. 1 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 3935] Near St. Jona Island, 56°18'N, 145°04'E, Okhotsk Sea, depth 175 fathoms. Holotype (unique): ZIN 18968. •Valid as Icelus armatus (Schmidt 1916) -- (Nelson 1984:22 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S76 [ref. 28300], Fedorov et al. 2003:71 [ref. 27579], Parin et al. 2014:253 [ref. 33547]). Current status: Valid as Icelus armatus (Schmidt 1916). Cottidae. Distribution: Western North Pacific. Habitat: marine.

armatus, Leptocottus Girard [C. F.] 1854:131 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] San Francisco, California, U.S.A. Syntypes: MCZ 36019 [ex USNM 310] (1), ?UMMZ 143015 (1, missing), USNM 310 (4). •Valid as Leptocottus armatus Girard 1854 -- (Bolin 1944:97 [ref. 6379], Lee et al. 1980:825 [ref. 22416], Eschmeyer & Herald 1983:175 [ref. 9277], Matarese et al. 1989:446 [ref. 26885], McAllister 1990:199 [ref. 14674], Swift et al. 1993:132 [ref. 22379], Castro-Aguirre et al. 1999:247 [ref. 24550], Mecklenburg et al. 2002:467 [ref. 25968], Moyle 2002:361 [ref. 26449], Wydoski & Whitney 2003:215 [ref. 27816], Nelson et al. 2004:120 [ref. 27807], Miller 2006:325 [ref. 28615], Page et al. 2013:124 [ref. 32708], Goto et al. 2014:[9] [ref. 33456], Pietsch & Orr 2015:43 [ref. 34694], Kells et al. 2016:180 [ref. 35888]). Current status: Valid as Leptocottus armatus Girard 1854. Cottidae. Distribution: Eastern North Pacific: southeastern Bering Sea to northern Baja Calfiornia. Habitat: brackish, marine.

asper, Cottus Richardson [J.] (ex Gairdner) 1836:295, 313, Pl. 95 (fig. 1) [Part 3; ref. 3731] Columbia River at Fort Vancouver, southern Washington, U.S.A. Holotype (unique): BMNH. Apparent holotype is at the BMNH (pers. comm. from C. Bond, 6/99). •Valid as Cottus asper Richardson 1836 -- (Lee et al. 1980:802 [ref. 22416], Eschmeyer & Herald 1983:167 [ref. 9277], Matarese et al. 1989:444 [ref. 26885], McAllister 1990:196 [ref. 14674], Page & Burr 1991:247 [ref. 18983], Swift et al. 1993:127 [ref. 22379], Scott & Crossman 1998:823 [ref. 24570], Fuller et al. 1999:339 [ref. 25838], Mecklenburg et al. 2002:469 [ref. 25968], Moyle 2002:346 [ref. 26449], Wydoski & Whitney 2003:203 [ref. 27816], Nelson et al. 2004:119 [ref. 27807], McPhail 2007:457 [ref. 31199], Scharpf 2008:7 [ref. 30399], Page & Burr 2011:472 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[9] [ref. 33456], Pietsch & Orr 2015:40 [ref. 34694], Kells et al. 2016:168 [ref. 35888]). Current status: Valid as Cottus asper Richardson 1836. Cottidae. Distribution: Eastern North Pacific/western North America: Seward, Alaska, to Ventura River, southern California, U.S.A. Habitat: freshwater, brackish, marine.

asperrima, Cottus Rutter [C. L.] 1908:144, Fig. 3 [Bulletin of the Bureau of Fisheries v. 27 (Doc. 637) (for 1907); ref. 12415] Fall River at Dana and Fall River Mills, California, U.S.A. Holotype: USNM 58500. Paratypes: (5) CAS-SU 9968 (2), USNM 126463 (1). Type catalog: Böhlke 1953:128 [ref. 12291]. •Valid as Cottus asperrimus Rutter 1908 -- (Lee et al. 1980:803 [ref. 22416], Page & Burr 1991:249 [ref. 18983], Moyle 2002:360 [ref. 26449], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:7 [ref. 30399], Page & Burr 2011:474 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[9] [ref. 33456]). Current status: Valid as Cottus asperrimus Rutter 1908. Cottidae. Distribution: California, U.S.A. Habitat: freshwater.

asperulus, Artedius Starks [E. C.] 1896:553 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 4195] Port Ludlow, Washington, U.S.A. Syntypes: CAS-SU 5046 (3). Type catalog: Böhlke 1953:127 [ref. 12291]. •Synonym of Artedius fenestralis Jordan & Gilbert 1883 -- (Bolin 1944:48 [ref. 6379]). Current status: Synonym of Artedius fenestralis Jordan & Gilbert 1883. Cottidae. Habitat: marine.

asprellus, Radulinus Gilbert [C. H.] 1890:88 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off northwestern U.S.A., depth 43-77 fathoms. Lectotype: USNM 43096. Paralectotypes: BMNH 1900.7.11.11 (1), CAS-SU 25 (4), ZMB 13940 (1), ZMUO J1479 (1). Type catalog: Böhlke 1953:132 [ref. 12291], Paepke & Fricke 1992:286 [ref. 19946]. Lectotype established in Jordan & Evermann 1898:1921 [ref. 2444]. •Valid as Radulinus asprellus Gilbert 1890 -- (Bolin 1944:40 [ref. 6379], Eschmeyer & Herald 1983:181 [ref. 9277], Matarese et al. 1989:406 [ref. 26885], McAllister 1990:201 [ref. 14674], Yabe & Maruyama 2001:59 [ref. 25245], Mecklenburg et al. 2002:446 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:44 [ref. 34694], Kells et al. 2016:188 [ref. 35888]). Current status: Valid as Radulinus asprellus Gilbert 1890. Cottidae. Distribution: Eastern North Pacific: Bering Sea to Aleutian Islands, Alaska, to northern Baja California. Habitat: marine.

asundensis, Cottus quadricornis Lönnberg [E.] 1932:17, Pl. (figs. 7 a-b) [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Åsunden, Sweden. Syntypes: NRM 20329-32 (1, 5, 3, 3). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

atlanticus, Artediellus Jordan [D. S.] & Evermann [B. W.] 1898:1906 [Bulletin of the United States National Museum No. 47; ref. 2444] Massachusetts Bay, U.S.A. Syntypes (?): CAS-SU 448 (3), USNM 21069 (?1). Type catalog: Böhlke 1953:126 [ref. 12291]. Based on literature and apparently one specimen; the described specimen would make the best lectotype if identifiable. •Valid as Artediellus atlanticus Jordan & Evermann 1898 -- (Neyelov 1973:594 [ref. 7219], Neyelov 1979:153 [ref. 3152], Fedorov in Whitehead et al. 1986:1244 [ref. 13677], Nelson 1986:43 [ref. 5808], Scott & Scott 1988:490 [ref. 25518], McAllister 1990:192 [ref. 14674], Yabe in Okamura et al. 1995:161 [ref. 22531], Klein-MacPhee in Collette & Klein-MacPhee 2002:346 [ref. 26158], Parin et al. 2002:S69 [ref. 28300], Moore et al. 2003:222 [ref. 27076], Nelson et al. 2004:118 [ref. 27807], Møller et al. 2010:51 [ref. 31169], Page et al. 2013:121 [ref. 32709], Mecklenburg et al. 2014:135 [ref. 33463], Parin et al. 2014:236 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Afonso et al. 2016:40, 99 [ref. 36199], Mecklenburg et al. 2016:49 [ref. 34440], Mecklenburg et al. 2018:186 [ref. 36370]). Current status: Valid as Artediellus atlanticus Jordan & Evermann 1898. Cottidae. Distribution: North Atlantic and Arctic Ocean, in North Pacific. Habitat: marine.

aturi, Cottus Freyhof [J.], Kottelat [M.] & Nolte [A.] 2005:132, Fig. 12 [Ichthyological Exploration of Freshwaters v. 16 (no. 2); ref. 28259] Adour drainage, Dept. Hautes-Pyrénées, River Adour in Tarbes (Séméac), 43°19'39"B, 0°05'27"E, France. Holotype: MNHN 2005-0750. Paratypes: CMK 17577 (5), 18327 (7); FSFJ 1653 (2); MNHN 2005-0751 (3), 2005-0752 (2). Name is a noun in apposition. •Valid as Cottus aturi Freyhof, Kottelat & Nolte 2005 -- (Goto et al. 2014:[10] [ref. 33456], Sideleva et al. 2018:9 [ref. 36393]). Current status: Valid as Cottus aturi Freyhof, Kottelat & Nolte 2005. Cottidae. Distribution: Adour and Nivelle drainages, France and Spain. Habitat: freshwater.

auriculatus, Artedielloides Soldatov [V. K.] 1922:353, Fig. (p. 353) [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23; ref. 4160] Peter the Great Bay [n. Japanese Sea], Russia, depth 29-30 meters. Holotype (unique): ZIN 20666. See Nakabo 2002:1526 [ref. 26001]. •Valid as Artedielloides auriculatus Soldatov 1922 -- (Neyelov 1979:159 [ref. 3152], Lindberg & Krasyukova 1987:236 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Parin et al. 2002:S69 [ref. 28300], Parin et al. 2014:235 [ref. 33547]). Current status: Valid as Artedielloides auriculatus Soldatov 1922. Cottidae. Distribution: Western North Pacific. Habitat: marine.

australis, Clinocottus analis Hubbs [C. L.] 1926:11 [Occasional Papers of the Museum of Zoology University of Michigan No. 171; ref. 2235] Point Loma, San Diego, California, U.S.A. Holotype: UMMZ 55002 [?ex FMNH 9256]. Paratypes: AMNH 5600 (1); ANSP; BMNH 1932.2.16.507-516 (40), FMNH 42048-51 [ex UMMZ] (57, 13, 44, 32); MCZ 36085 [ex USNM 66097] (5); CAS-SU (missing); UMMZ 64210 (now 0), 64417-20 (2, 3, 0, 1), 65495-500 (5, 50, 7, 106, 35, 0), 66001-08 (0, 37, 69, 63, 57, 107, 43, 130), 66097 (201); USNM 83501 (13), 117522 (7). Type catalog: Ibarra & Stewart 1987:27 [ref. 12367]. •Synonym of Clinocottus analis (Girard 1858) -- (Bolin 1944:73 [ref. 6379]). Current status: Synonym of Clinocottus analis (Girard 1858). Cottidae. Habitat: marine.

australis, Icelinus Eigenmann [C. H.] & Eigenmann [R. S.] 1889:131 [8] [West American Scientist v. 6 (no. 48); ref. 1251] Cortez Banks, off San Diego, California, U.S.A., depth 45 fathoms. Syntypes: BMNH 1891.5.19.59 (1, poor condition), USNM 41917 (1, poor condition). On p. 8 of separate. See Bolin 1936:159 [ref. 506]. •Unidentifiable, not an Icelinus -- (Rosenblatt & Smith 2004:560 [ref. 27912]). Cottidae. Habitat: marine.

australis, Leptocottus armatus Hubbs [C. L.] 1921:5 [Occasional Papers of the Museum of Zoology University of Michigan No. 94; ref. 19137] Lagoon at the mouth of Ventura River at Ventura, California, U.S.A. Holotype: UMMZ 55055. Paratypes: FMNH 1954 (4), 9109-10 (112, 3), 9140-41 (17, 1), 9143 (27), 9149 (3), 9163-64 (3, 1), 9166 (1); UMMZ ?80367 (7), 142922-25 (12, 2, 8, 20), 142927-28 (3, 1). Type catalog: Ibarra & Stewart 1987:52 [ref. 12367]. •Synonym of Leptocottus armatus Girard 1854 -- (Lee et al. 1980:825 [ref. 22416], Castro-Aguirre et al. 1999:247 [ref. 24550]). •Synonym of Leptocottus armatus Girard 1854, but a valid subspecies as described -- (Ruiz-Campos et al. 2000:71 [ref. 24940]). Current status: Synonym of Leptocottus armatus Girard 1854. Cottidae. Habitat: brackish, marine.

axillaris, Boreocottus Gill [T. N.] 1859:166 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 11; ref. 1760] Awatska Bay, Kamchatka, Russia, Bering Strait. Holotype (unique): ?USNM 24027 (poor condition). •Synonym of Myoxocephalus scorpioides (Fabricius 1780) -- (Neyelov 1973:599 [ref. 7219], Neyelov 1979:122 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:479 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:264 [ref. 33547], Mecklenburg et al. 2016:83 [ref. 34440]). Current status: Synonym of Myoxocephalus scorpioides (Fabricius 1780). Cottidae. Habitat: brackish, marine.

baicalensis, Cottus Dybowski [B. N.] 1874:386, Pl. 3 (fig. 2) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 24 (no. 3-4); ref. 1172] Lake Baikal, Russia. Syntypes: ZMB 7810 (4). Type catalog: Paepke & Fricke 1992:284 [ref. 19946]. •Valid as Batrachocottus baicalensis (Dybowski 1874) -- (Berg 1949:1165 [ref. 20662], Taliev 1955:257 [ref. 4316], Sideleva 1982:28 [ref. 14469], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:149 [ref. 23595], Sideleva 1999:149 [ref. 24864], Vasil'eva 1999:154 [ref. 24864], Bogutskaya et al. 2001:46 [ref. 26178], Sideleva 2001:58 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:15, 154 [ref. 27815], Bogutskaya & Naseka 2004:196 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Batrachocottus baicalensis (Dybowski 1874). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

baikalensis, Callionymus Pallas [P. S.] 1776:290, 707 [Reise durch verschiedene Provinzen des russischen Reiches; ref. 18509] Lake Baikal, Russia. Neotype: ZIN 16321. Originally as baïkalensis. Also described as new in Pallas 1787:258, Pl. 9 (figs. 2-3) [ref. 18506] as baicalensis, and Pallas 1787:349, Pl. 9 (figs. 2-3) [ref. 18507] as baicalensis. Sideleva 2001:58 [ref. 26181] names a neotype [ZIN 16321], but if that is the first time a neotype was named it is unavailable under the Code; same neotype mentioned in Sideleva 2003:171 [ref. 27815]. •Valid as Comephorus baicalensis [sic] (Pallas 1776) -- (Berg 1949:1175 [ref. 20662], Taliev 1955:367 [ref. 4316], Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:154 [ref. 23595], Yabe & Sideleva 2000:306 [ref. 25698], Bogutskaya et al. 2001:46 [ref. 26178], Fricke 2002:72 [ref. 26547] as baikalensis, Sideleva 2001:60 [ref. 26181], Kontula et al. 2003:149 [ref. 26791] as baicalensis, Sideleva 2003:16, 171 [ref. 27815] as baicalensis, Bogutskaya & Naseka 2004:193 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Comephorus baikalensis (Pallas 1776). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

baileyi, Cottus Robins [C. R.] 1961:311, Figs. 1b, c [Copeia 1961 (no. 3); ref. 9373] Middle fork of Holston River, 6 miles east-northeast of Marion, Smyth County, Virginia, U.S.A. Holotype: CUMV 25914. Paratypes: CUMV 22849 (58), 25915 (1, allotype); UMMZ 167171 (17). Additional material: CUMV 19179 (10), 21512 (4), 24418 (7); UMMZ 129084 (1), 129106 (1), 157471 (29); USNM 102433 (1). •Valid as Cottus baileyi Robins 1961 -- (Lee et al. 1980:804 [ref. 22416], Page & Burr 1991:250 [ref. 18983], Jenkins & Burkhead 1994:644 [ref. 21581], Kinziger et al. 2000:1007 [ref. 25019], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:7 [ref. 30399], Page & Burr 2011:476 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus baileyi Robins 1961. Cottidae. Distribution: Upper Tennessee River draiange, Virginia and Tennessee, U.S.A. Habitat: freshwater.

bairdii, Cottus Girard [C. F.] 1850:410 [Proceedings of AAAS (The American Association for the Advancement of Science) 2nd meeting, 1849; ref. 18123] Mahoning River, Portland, Ohio; tributaries to Ohio River in Pennsylvania, U.S.A. Syntypes: none known. See also Girard 1850:189 [ref. 1814]. •Valid as Cottus bairdii Girard 1850 [often incorrectly as bairdi; sometimes with subspecies] -- (Bailey & Bond 1963:1 [ref. 3934] as bairdi, Lee et al. 1980:805 [ref. 22416], Smith 1986:355 [ref. 22186], McAllister 1990:196 [ref. 14674] as bairdi, Page & Burr 1991:250 [ref. 18983], Boschung 1992:102 [ref. 23239], Etnier & Starnes 1993:384 [ref. 22809], Jenkins & Burkhead 1994:639 [ref. 21581], La Rivers 1994:585 [ref. 23389], Baxter & Stone 1995:220 [ref. 23374], Stauffer et al. 1995:250 [ref. 23376], Andersen & Deacon 1996:124 [ref. 24717], Chereshnev 1996:605 [ref. 24659], Mettee et al. 1996:491 [ref. 25836], Sigler & Sigler 1996:230 [ref. 25943], Scott & Crossman 1998:826 [ref. 24570] as bairdi, Fuller et al. 1999:340 [ref. 25838] as bairdi, Broughton 2000:653 [ref. 24593] as bairdi, Lyons et al. 2000:42 [ref. 26594] as bairdi, Markle & Hill 2000:202 [ref. 24886] as bairdi, Kinziger et al. 2000:1007 [ref. 25019], Gasper et al. 2001:264 [ref. 25578] as bairdi, Wydoski & Whitney 2003:205 [ref. 27816] as bairdi, Bailey et al. 2004:170 [ref. 27882], Nelson et al. 2004:119 [ref. 27807], Boschung & Mayden 2004:419 [ref. 27995], Robins 2005:4 [ref. 28230], McPhail 2007:463 [ref. 31199], Scharpf 2008:7, 9 [ref. 30399], Minckley & Marsh 2009:308 [ref. 31114] as bairdi, Kinziger & Wood 2010:59 [ref. 30701], Page & Burr 2011:475 [ref. 31215], Page et al. 2013:122 [ref. 32708], LeMone et al. 2014:249 [ref. 33134] as cf. bairdii, Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus bairdii Girard 1850. Cottidae. Distribution: Widespread in North America. Habitat: freshwater.

bathybius, Cottus Günther [A.] 1878:180 [13] [Annals and Magazine of Natural History (Series 5) v. 2 (nos 7/8/9)(art. 2/22/28); ref. 2010] Off Misaki, Japanese Seas, Challenger station 235, depth 565 fathoms. Holotype (unique): BMNH 1879.5.14.201. On p. 13 of separate. See also Günther 1887:62, Pl. 10 (fig. C) [ref. 2013]. •Valid as Zesticelus bathybius (Günther 1878) -- (Neyelov 1979:161 [ref. 3152], Yabe & Okamura 1993:369 [ref. 22281], Yabe 1995:19 [ref. 22666], Nakabo 2000:640 [ref. 25086], Shinohara et al. 2001:319 [ref. 25995], Nakabo 2002:640 [ref. 26001], Parin et al. 2002:S89 [ref. 28300], Parin et al. 2014:279 [ref. 33547]). Current status: Valid as Zesticelus bathybius (Günther 1878). Cottidae. Distribution: Western North Pacific: Bering Sea to northern Baja California and to southeastern Kamchatka. Habitat: marine.

batrachoides, Myoxocephalus Gilbert [C. H.] & Burke [C. V.] 1912:58, Fig. 11 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Codfish banks west of southern part of Kamchatka, Russia, Albatross station 4798, 51°37'N, 156°21'E, depth 25 fathoms. Holotype (unique): USNM 71809. •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Neyelov 1979:122 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Cottidae. Habitat: marine.

bauntovi, Cottus kessleri Taliev [D. N.] 1946:744, Fig. 2 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 52 (no. 8); ref. 13135] Baunt Lake, east of Lake Baikal, Vitim River basin, Buryatia, Russia. Syntypes: (3) not at ZIN. Evidently type locality is in error. •Synonym of Paracottus kessleri (Dybowski 1874), but a valid subspecies as described -- (Taliev 1955:249 [ref. 4316]). •Synonym of Cottus kesslerii Dybowski 1874 -- (Sideleva 1998:151 [ref. 23595]). •Synonym of Cottus kesslerii Dybowski 1874, but a valid subspecies as described -- (Berg 1949:1154 [ref. 20662]). •Synonym of Leocottus kesslerii (Dybowski 1874) -- (Sideleva 2001:59 [ref. 26181], Sideleva 2003:16, 151 [ref. 27815], Kottelat 2006:60 [ref. 28831]). Current status: Synonym of Leocottus kesslerii (Dybowski 1874). Cottidae. Habitat: freshwater.

beani, Triglops Gilbert [C. H.] 1896:426, Pl. 28 (upper) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Overall localities -- North and south of Aleutian Islands and in Bristol Bay, Alaska, U.S.A., depth 7.25-42 fathoms. Lectotype: ? CAS-SU 2422. Paralectotypes: BMNH 1900.7.11.10 (1); MCZ 28355, 28356 (1); CAS-SU 2418-19 (6, 5), 2421 (5), 2423-24 (2, 3). Type catalog: Böhlke 1953:132 [ref. 12291]. Lectotype established (as figured specimen from Albatross sta. 3220) in caption to Pl. 288, p. 3291 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable; Pietsch 1994:375, caption to fig. 17 regarded a male from SU 2422 as probably the figured specimen. •Valid as Triglops beani Gilbert 1896 -- (Watanabe 1960:16 [ref. 11959]). •Synonym of Triglops pingelii Reinhardt 1837 -- (Neyelov 1973:601 [ref. 7219], Lindberg & Krasyukova 1987:174 [ref. 15964], Pietsch 1994:374 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:428 [ref. 25968], Parin et al. 2002:S88 [ref. 28300], Parin et al. 2014:277 [ref. 33547], Mecklenburg et al. 2016:91 [ref. 34440]). Current status: Synonym of Triglops pingelii Reinhardt 1837. Cottidae. Habitat: marine.

beldingii, Cottus Eigenmann [C. H.] & Eigenmann [R. S.] 1891:1132 [American Naturalist v. 25 (no. 300); ref. 11063] Lake Tahoe and Donner Lake, California, U.S.A. Syntypes: BMNH 1890.11.15.94-97 (4), 1890.11.15.98-101 (4); not CAS 504, 505, and 702; UMMZ 92669 (2). CAS type specimens would have been lost in the fire of 1906. •Valid as Cottus beldingii Eigenmann & Eigenmann 1891 [often incorrectly as beldingi] -- (Bailey & Bond 1963:1 [ref. 3934] as beldingi, Lee et al. 1980:806 [ref. 22416], Page & Burr 1991:251 [ref. 18983], La Rivers 1994:588 [ref. 23389], Baxter & Stone 1995:222 [ref. 23374], Sigler & Sigler 1996:232 [ref. 25943] as beldingi, Moyle 2002:357 [ref. 26449] as beldingi, Wydoski & Whitney 2003:206 [ref. 27816] as beldingi, Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:8 [ref. 30399], Page & Burr 2011:479 [ref. 31215], Page et al. 2013:122 [ref. 32708], LeMone et al. 2014:249 [ref. 33134], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus beldingii Eigenmann & Eigenmann 1891. Cottidae. Distribution: Western U.S.A. Habitat: freshwater.

bendirei, Potamocottus Bean [T. H.] 1881:27 [Proceedings of the United States National Museum v. 4 (no. 190); ref. 16046] Rattlesnake Creek near Camp Harney, Oregon, U.S.A. Holotype (unique): USNM 24196. •Synonym of Cottus bairdii Girard 1850 -- (Bailey & Bond 1963:1 [ref. 3934]). •Valid as Cottus bendirei (Bean 1881) -- (Markle & Hill 2000:202 [ref. 24886], Nelson et al. 2004:119 [ref. 27807], McPhail 2007:476 [ref. 31199] with question, Scharpf 2008:8 [ref. 30399], Page & Burr 2011:479 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus bendirei (Bean 1881). Cottidae. Distribution: Northwestern U.S.A.: Oregon, Idaho, and questionably in s. Canada [on ref. 31199]; usually as Oregon only. Habitat: freshwater.

bergi, Limnocottus godlewskii var. Dybowski [B. N.] 1908:555 [20], Fig. 15 [Kosmos Lwów v. 33; ref. 17422] Lake Baikal, Russia. Syntypes: whereabouts unknown. •Synonym of Limnocottus godlewskii (Dybowski 1874) -- (Berg 1949:1160 [ref. 20662]). Current status: Synonym of Limnocottus godlewskii (Dybowski 1874). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

bergianus, Limnocottus Taliev [D. N.] 1935:61, Pl. 1 (fig. 1), Pl. 2 [Trudy Baikal'skoi limnologischeskoi stantsii = Travaux de la Station limnologique du Lac Baikal v. 6; ref. 15741] Off Listvenichnoe, depth about 700 meters, and Barjuzin Bay, depth 619 meters, southern Lake Baikal, Russia. Lectotype: ZIN 32562. Paralectotypes and/or Taliev specimens: ZIN 32562 (1). Lectotype apparently designated by Sideleva 2001:68 [ref. 26181] but not in conformance with the fourth edition of the Code. •Valid as Abyssocottus bergianus (Taliev 1935) -- (Taliev 1955:362 [ref. 4316]). •Valid as Limnocottus bergianus Taliev 1935 -- (Berg 1949:1161 [ref. 20662], Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:156 [ref. 23595], Sideleva 2001:68 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:18, 196 [ref. 27815], Bogutskaya & Naseka 2004:198 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Limnocottus bergianus Taliev 1935. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

bergii, Taurocottus Soldatov [V. K.] & Pavlenko [M. N.] 1915:149-151, Pl. 4 (figs. 1-2) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 4165] 47°55'00"N, 139°39'30"E, Primorye, Russia, Sea of Japan, depth 66 meters. Lectotype: ZIN 18637. Paralectotypes: (7) ZIN 18636-37 (2, 1). Lectotype selected by Neyelov 1979:143 [ref. 3152]. •Valid as Taurocottus bergii Soldatov & Pavlenko 1915 -- (Neyelov 1979:143 [ref. 3152], Yabe in Masuda et al. 1984:326 [ref. 6441] as bergi, Lindberg & Krasyukova 1987:227 [ref. 15964] as bergi, Kim & Youn 1992:70 [ref. 22287], Sokolovskaya et al. 1998:10 [ref. 24670] as bergi, Nakabo 2000:644 [ref. 25086] as bergi, Mecklenburg et al. 2002:474 [ref. 25968], Nakabo 2002:644 [ref. 26001] as bergi, Youn 2002:263, 578 [ref. 26218], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:273 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331]). Current status: Valid as Taurocottus bergii Soldatov & Pavlenko 1915. Cottidae. Distribution: Western North Pacific. Habitat: marine.

beringi, Crossias Soldatov [V. K.] 1916:218, Pl. 13 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 21; ref. 18735] Abrek Bay and Ayan Bay, Bol. Shantar Island, Okhotsk Sea; also Olga Bay, Sea of Japan. Syntypes: (many) ZIN 19066-67 (3, 4). Two new forms described on p. 221. •Synonym of Porocottus tentaculatus (Kner 1868) -- (Lindberg & Krasyukova 1987:224 [ref. 15964], Parin et al. 2014:270 [ref. 33547] in part). •Synonym of Porocottus minutus (Pallas 1814) or Porocottus tentaculatus (Kner 1868) -- (Neyelov 1979:137 [ref. 3152], Parin et al. 2002:S84 [ref. 28300], Parin et al. 2014:269 [ref. 33547]). Current status: Synonym of Porocottus minutus (Pallas 1814). Cottidae. Habitat: marine.

beringianum, Stelgistrum Gilbert [C. H.] & Burke [C. V.] 1912:52, Fig. 9 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Petrel Bank, Aleutian Islands, Albatross station 4777, depth 43-52 fathoms. Holotype: USNM 74373. Paratypes: CAS-SU 22245 (1). Type catalog: Böhlke 1953:132 [ref. 12291]. •Valid as Stelgistrum beringianum Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:449 [ref. 25968], Parin et al. 2002:S86 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:272 [ref. 33547]). Current status: Valid as Stelgistrum beringianum Gilbert & Burke 1912. Cottidae. Distribution: North Pacific: southern Bering Sea, Commander Islands, and Aleutian Islands. Habitat: marine.

beringianus, Artediellus scaber Schmidt [P. Yu.] 1927:8 [Proceedings of the United States National Museum v. 71 (no. 2685); ref. 15822] Tkatchen Bay, 64°25'N, 172°48'03"W, northern Bering Sea. Holotype: ZIN 21492 (1 of 2 adults). Paratypes: ZIN 21492 (1 of 2 adults). •Synonym of Artediellus scaber Knipowitsch 1907 -- (Neyelov 1979:154 [ref. 3152], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:491 [ref. 25968], Parin et al. 2002:S71 [ref. 28300], Parin et al. 2014:239 [ref. 33547], Mecklenburg et al. 2016:52 [ref. 34440]). Current status: Synonym of Artediellus scaber Knipowitsch 1907. Cottidae. Habitat: brackish, marine.

beringianus, Icelus bicornis Schmidt [P. Yu.] 1927:7 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 19815] Providence Bay, Bering Strait, 63°27' to 64°25'N, 168°48' to 176°27'E, depth 8-40 fathoms. Syntypes: ZIN 21914 (1). •Synonym of Icelus spatula Gilbert & Burke 1912 -- (Neyelov 1973:597 [ref. 7219], Nelson 1984:36 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:256 [ref. 33547]). Current status: Synonym of Icelus spatula Gilbert & Burke 1912. Cottidae. Habitat: marine.

bicolor, Procottus jeittelesi var. Dybowski [B. N.] 1908:557 [22], Fig. 18 [Kosmos Lwów v. 33; ref. 17422] Lake Baikal, Russia. Syntypes: whereabouts unknown. •Synonym of Procottus jeittelesii (Dybowski 1874) -- (Berg 1949:1168 [ref. 20662]). Current status: Synonym of Procottus jeittelesii (Dybowski 1874). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

bicornis, Cottus Reinhardt [J. C. H.] 1840:9 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) v. 7 (1839); ref. 18617] East Greenland. No types known. Type catalog: Nielsen 1974:81 [ref. 9588]. Published two more times in 1840-41. Cottus bicornis Anonymous (Volta) 1796:clxiv, Pl. 39 (fig. 4) [ref. 30407] from the 'monti del Veronese' (mountains of Province Verona, Italy) is an homonym in fossil fishes; Agassiz 1835:293 [ref. 30408] noted that the original material is lost and the illustration in Volta 1796 is unidentifiable; the name Cottus bicornis Anonymous (Volta) 1796 was not used as valid since 1899, therefore reversal of precedence according to ICZN Art. 23.2 and 23.9 in favor of Cottus bicornis Reinhardt 1840. •Valid as Icelus bicornis (Reinhardt 1840) -- (Neyelov 1973:596 [ref. 7219], Nelson 1984:35 [ref. 5391], Fedorov in Whitehead et al. 1986:1250 [ref. 13677], Robins & Ray 1986:282 [ref. 23100], Scott & Scott 1988:498 [ref. 25518], McAllister 1990:198 [ref. 14674], Yabe in Okamura et al. 1995:164 [ref. 22531], Mecklenburg et al. 2002:456 [ref. 25968], Parin et al. 2002:S77 [ref. 28300], Nelson et al. 2004:120 [ref. 27807], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Dolgov 2013:918 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:253 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:43, 173 [ref. 36199], Mecklenburg et al. 2016:67 [ref. 34440], Mecklenburg et al. 2018:205 [ref. 36370]). Current status: Valid as Icelus bicornis (Reinhardt 1840). Cottidae. Distribution: Cumpolar in Arctic Ocean, North Atlantic and North Pacific. Habitat: marine.

biseriatus, Archaulus Gilbert [C. H.] & Burke [C. V.] 1912:36, Fig. 1 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Petrel Bank, Bering Sea, Albatross station 4778, depth 43 fathoms. Holotype: USNM 74365. Paratypes: CAS-SU 22238 (4); USNM 70879 (1), 74503-04 (3, 3). Type catalog: Böhlke 1953:126 [ref. 12291]. •Valid as Archaulus biseriatus Gilbert & Burke 1912 -- (Parin et al. 2002:S68 [ref. 28300]). •Valid as Archistes biseriatus (Gilbert & Burke 1912) -- (Sheiko & Fedorov 2000:26 [ref. 25650], Mecklenburg et al. 2002:443 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Parin et al. 2014:233 [ref. 33547], Kells et al. 2016:162 [ref. 35888]). Current status: Valid as Archistes biseriatus (Gilbert & Burke 1912). Cottidae. Distribution: North Pacific: Kuril Islands; southern Bering Sea off Aleutian Islands. Habitat: marine.

bison, Aspicottus Girard [C. F.] 1854:130 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] San Francisco, California, U.S.A. No types known. •Valid as Aspicottus bison Girard 1854 -- (Neyelov 1979:148 [ref. 3152]). •Valid as Enophrys bison (Girard 1854) -- (Bolin 1944:91 [ref. 6379], Sandercock & Wilimovsky 1968:844 [ref. 26414], Eschmeyer & Herald 1983:168 [ref. 9277], Matarese et al. 1989:390 [ref. 26885], McAllister 1990:197 [ref. 14674], Mecklenburg et al. 2002:471 [ref. 25968], Nelson et al. 2004:119 [ref. 27807], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:40 [ref. 34694], Kells et al. 2016:170 [ref. 35888]). Current status: Valid as Enophrys bison (Girard 1854). Cottidae. Distribution: Eastern North Pacific: western Gulf of Alaska to central California. Habitat: marine.

bispinis, Icelus spatula Andriashev [A. P.] 1937:270, Pl. 6 (fig. 5) [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] Penzhina Bay to Chanjanga; northeastern Sachalin Island; and northwestern Kamchatka Peninsula, Okhotsk Sea, depth 55-150 meters and to 240 meters. Syntypes: ZIN 17787 (1), 17790-91 (1, 3), 18060 (1), 21920-23 (5, 6, 6, 2). •Synonym of Icelus bicornis (Reinhardt 1840) -- (Nelson 1984:42 [ref. 5391]). •Synonym of Icelus spatula Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:256 [ref. 33547]). Current status: Synonym of Icelus spatula Gilbert & Burke 1912. Cottidae. Habitat: marine.

boleoides, Cottus Girard [C. F.] 1850:411 [Proceedings of AAAS (The American Association for the Advancement of Science) 2nd meeting, 1849; ref. 18123] Windsor, Vermont, U.S.A. See also Girard 1850:189 [ref. 1814] and Girard 1851:56, Pl. 2 (figs. 7-8) [ref. 18124]. •Synonym of Cottus cognatus Richardson 1837. Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

boleoides, Radulinus Gilbert [C. H.] in Jordan & Evermann 1898:1919 [Bulletin of the United States National Museum No. 47; ref. 2444] Off Santa Catalina Island, California, U.S.A., Albatross station 3664, depth 59 fathoms. Holotype (unique): USNM 48795 [also = 48879]. Author is C. H. Gilbert. Also appeared as new in Gilbert 1899:26, Pl. 1 [ref. 21322]. •Valid as Radulinus boleoides Gilbert 1898 -- (Bolin 1944:38 [ref. 6379], Eschmeyer & Herald 1983:181 [ref. 9277], Matarese et al. 1989:408 [ref. 26885], McAllister 1990:201 [ref. 14674], Yabe & Maruyama 2001:59 [ref. 25245], Mecklenburg et al. 2002:445 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:44 [ref. 34694], Kells et al. 2016:188 [ref. 35888]). Current status: Valid as Radulinus boleoides Gilbert 1898. Cottidae. Distribution: Eastern North Pacific: Kodiak Island, Alaska to southern California, U.S.A. Habitat: marine.

borealis, Crossias beringi forma Soldatov [V. K.] 1916:220, Pl. 13 (upper) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 21; ref. 18735] Okhotsk Sea. Syntypes: ZIN 19066 (1). Treated as infrasubspecific in Eschmeyer et al. 1998; now regarded as subspecific. Muto et al. 2002:231 [ref. 26331] list one syntype. •Synonym of Porocottus minutus (Pallas 1814) -- (Neyelov 1979:137 [ref. 3152], Parin et al. 2002:S84 [ref. 28300]). Current status: Synonym of Porocottus minutus (Pallas 1814). Cottidae. Habitat: marine.

borealis, Icelinus Gilbert [C. H.] 1896:415, Pl. 25 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] North and south of Aleutians and in Bristol Bay, Alaska, U.S.A. Lectotype: not determined. Paralectotypes (incl. potential lectotype): MCZ 28373 (6); MNHN 1896-0491 [ex USNM] (4); CAS-SU 2291-92 (6, 34); USNM 53037 (6); ZMUO J1454 (3, poor condition) station 3265, ZMUO J6331 (3) station 3213. Type catalog: Böhlke 1953:129 [ref. 12291], Blanc & Hureau 1968:53 [ref. 20738], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen) in caption to Pl. 284, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. •Valid as Icelinus borealis Gilbert 1896 -- (Eschmeyer & Herald 1983:172 [ref. 9277], Peden 1984:79 [ref. 5193], McAllister 1990:198 [ref. 14674], Mecklenburg et al. 2002:441 [ref. 25968], Parin et al. 2002:S76 [ref. 28300], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:252 [ref. 33547], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:174 [ref. 35888]). Current status: Valid as Icelinus borealis Gilbert 1896. Cottidae. Distribution: North Pacific: Bristol Bay and Aleutian Islands to Puget Sound, Washington, U.S.A. Habitat: marine.

borealis, Oligocottus Jordan [D. S.] & Snyder [J. O.] in Jordan 1896:225 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 2395] Neah Bay, Puget Sound, Washington, U.S.A. Syntypes: ANSP 39141 (1, missing); BMNH 1897.1.12.17-18 (2); CAS-SU 2985 (4), 3396 (27). Type catalog: Böhlke 1953:131 [ref. 12291], Böhlke 1984:65 [ref. 13621]. •Synonym of Oligocottus maculosus Girard 1856 -- (Bolin 1944:65 [ref. 6379], Mecklenburg et al. 2002:500 [ref. 25968]). Current status: Synonym of Oligocottus maculosus Girard 1856. Cottidae. Habitat: marine.

borkensis, Cottus quadricornis Lönnberg [E.] 1939:4 [Arkiv för Zoologi v. 31 B (no. 2); ref. 17199] Lake Borken [Bork], parishes of Yxnerum and Björsäter, Osthrogothland Province, Sweden. Syntypes: NRM 20327 (2). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

boulengeri, Abyssocottus Berg [L. S.] 1906:908 [Zoologischer Anzeiger v. 30 (no. 26); ref. 265] Lake Baikal, Russia. Syntypes: whereabouts unknown. Sideleva 2001:68 [ref. 26181] lists ZIN 13736 as holotype. •Valid as Cottinella boulengeri (Berg 1906) -- (Berg 1949:1159 [ref. 20662], Taliev 1955:341 [ref. 4316], Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:156 [ref. 23595], Sideleva 2001:68 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:18, 189 [ref. 27815], Bogutskaya & Naseka 2004:197 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Cottinella boulengeri (Berg 1906). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

bradfordi, Porocottus Rutter [C. L.] in Jordan & Evermann 1898:2862 [Bulletin of the United States National Museum No. 47; ref. 2445] Rock pools at Karluk, Kodiak Island, Alaska, U.S.A. Syntypes: BMNH 1900.9.29.209-210 (2), CAS-SU 6063 (57), USNM 60328 (1), USFC. Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Porocottus bradfordi Rutter 1898 -- (Neyelov 1979:136 [ref. 3152], Eschmeyer & Herald 1983:179 [ref. 9277], Muto et al. 2002:231 [ref. 26331]). •Synonym of Porocottus mednius (Bean 1898) -- (Mecklenburg et al. 2002:404 [ref. 25968]). Current status: Synonym of Porocottus mednius (Bean 1898). Cottidae. Distribution: North Pacific: Bering Strait to Commander-Aleutian chain to southeastern Alaska and to Kuril Islands. Habitat: marine.

brandtii, Cottus Steindachner [F.] 1867:119 [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Classe v. 4 (no. 14) (for 16 May 1867); ref. 21724] Mouth of Amur River, Russia. Holotype (unique): NMW 11676. Species also appeared on p. 79 of English version [ref. 18764] as brandtii; later illustrated and described in more detail in Steindachner 1867:706, Pl. 3 (figs. 1-2) [p. 6 of separate] [ref. 6393] as brandti in text and brandtii on plate. •Valid as Myoxocephalus brandtii (Steindachner 1867) -- (Watanabe 1960:73 [ref. 11959], Neyelov 1979:121 [ref. 3152] as brandti, Yabe in Masuda et al. 1984:326 [ref. 6441] as brandti, Lindberg & Krasyukova 1987:210 [ref. 15964] as brandti, Sokolovsky & Sokolovskaya 1997:60 [ref. 24691] as brandti, Sokolovskaya et al. 1998:10 [ref. 24670] as brandti, Nakabo 2000:645 [ref. 25086] as brandti, Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:404 [ref. 25968], Nakabo 2002:645 [ref. 26001] as brandti, Parin et al. 2002:S80 [ref. 28300], Fedorov et al. 2003:75 [ref. 27579], Parin et al. 2014:260 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138], Prokofiev 2017:648 [ref. 35721]). Current status: Valid as Myoxocephalus brandtii (Steindachner 1867). Cottidae. Distribution: Western North Pacific. Habitat: brackish, marine.

brashnikovi, Trichocottus Soldatov [V. K.] & Pavlenko [M. N.] 1915:151, Pl. 4 (fig. 3) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 4165] 50°04'50"N, 140°47'E, Tatar Strait, Primorye, Russia, Sea of Japan. Lectotype: ZIN 18640. Paralectotypes: ZIN 18639 (1). Lectotype selected by Neyelov 1979:143 [ref. 3152]. •Valid as Trichocottus brashnikovi Soldatov & Pavlenko 1915 -- (Neyelov 1979:143 [ref. 3152] as brashnicovi, Lindberg & Krasyukova 1987:228 [ref. 15964] as brashnicovi, Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:474 [ref. 25968], Parin et al. 2002:S87 [ref. 28300], Fedorov et al. 2003:81 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:274 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Mecklenburg et al. 2016:86 [ref. 34440], Mecklenburg et al. 2018:223 [ref. 36370]). Current status: Valid as Trichocottus brashnikovi Soldatov & Pavlenko 1915. Cottidae. Distribution: Western North Pacific and Arctic: northern Bering Sea, Alaska, to Okhotsk Sea and Tatar Strait, Japan Sea. Habitat: marine.

brevis, Asprocottus herzensteini var. Taliev [D. N.] 1948:113 [Trudy Baikal'skoi Limnologicheskoi Stantsii v. 12; ref. 19628] Not available, manuscript name mentioned in table; described as a new subspecies, Asprocottus herzensteini platycephalus in Taliev 1955:324 [ref. 4316]. •In the synonymy of Asprocottus pulcher Taliev 1955 -- (Sideleva 2003:17 [ref. 27815]). Current status: Synonym of Asprocottus pulcher Taliev 1955. Cottidae.

bryosus, Blennicottus globiceps Jordan [D. S.] & Starks [E. C.] 1895:808 [Proceedings of the California Academy of Sciences (Series 2) v. 5; ref. 2522] Near Neah Bay, Washington, U.S.A. Syntypes: NMW 6549 (1), CAS-SU 3404 (6). Type catalog: Böhlke 1953:127 [ref. 12291]. •Synonym of Clinocottus globiceps (Girard 1858) -- (Bolin 1944:81 [ref. 6379]). Current status: Synonym of Clinocottus globiceps (Girard 1858). Cottidae. Habitat: marine.

bubalis, Cottus Euphrasen [B. A.] 1786:65, Pl. 3 (figs. 2-3) [Kongliga Vetenskaps-Academiens Handlingar, Stockholm v. 7 (for 1786); ref. 17465] Bohuslän, Sweden. No types known. Also described by Euphrasen 1787:64, pl. 3, fig. 2-3 [ref. 30157]. •Valid as Enophrys bubalis (Euphrasen 1786) -- (Sandercock & Wilimovsky 1968:841 [ref. 26414]). •Valid as Taurulus bubalis (Euphrasen 1786) -- (Neyelov 1973:600 [ref. 7219], Neyelov 1979:149 [ref. 3152], Fedorov in Whitehead et al. 1986:1256 [ref. 13677], Parin et al. 2002:S86 [ref. 28300], Fricke 2007:29 [ref. 30577], Parin et al. 2014:273 [ref. 33547]). Current status: Valid as Taurulus bubalis (Euphrasen 1786). Cottidae. Distribution: Baltic Sea, North Sea, northwestern Mediterranean Sea, eastern North Atlantic: Iceland and Norway south to Gibraltar. Habitat: brackish, marine.

burchami, Icelinus Evermann [B. W.] & Goldsborough [E. L.] 1907:297, Fig. 48 [Bulletin of the Bureau of Fisheries v. 26 (Doc. 624) (for 1906); ref. 6532] Off Loring, Alaska, U.S.A., Albatross station 4228, depth 41-134 fathoms. Holotype: USNM 57822. Paratypes: ANSP 33003 (1, disintegrated); FMNH 6117 (not found); CAS-SU 22614 [not 20010 as published] (1); USNM 57825 (2), 124970 [ex USBF 1637 = 5229]. Type catalog: Böhlke 1953:129 [ref. 12291], Böhlke 1984:115 [ref. 13621]. •Valid as Icelinus burchami Evermann & Goldsborough 1907 -- (Bolin 1944:30 [ref. 6379], Eschmeyer & Herald 1983:172 [ref. 9277], Peden 1984:79 [ref. 5193], McAllister 1990:198 [ref. 14674], Mecklenburg et al. 2002:440 [ref. 25968], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:176 [ref. 35888]). Current status: Valid as Icelinus burchami Evermann & Goldsborough 1907. Cottidae. Distribution: Eastern North Pacific: southeastern Alaska to southern California, U.S.A. Habitat: marine.

caeruleomentum, Cottus Kinziger [A. P.], Raesly [R. L.] & Neely [D. A.] 2000:1009, Figs. 1-2 [Copeia 2000 (no. 4); ref. 25019] Evitts Creek at I68 bridge, east of Cumberland, Potomac River drainage, Allegany County, Maryland, U.S.A., 39°40'21.322"N, 78°43'25.728"W. Holotype: USNM 350112. Paratypes: USNM 350113 (41). •Valid as Cottus caeruleomentum Kinziger, Raesly & Neely 2000 -- (Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:8 [ref. 30399], Page & Burr 2011:476 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus caeruleomentum Kinziger, Raesly & Neely 2000. Cottidae. Distribution: Eastern U.S.A. Habitat: freshwater.

camchaticus, Artediellus Gilbert [C. H.] & Burke [C. V.] 1912:46, Fig. 6 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off eastern coast of Kamchatka, Russia, 52°46'50"N, Albatross station 4795, depth 48-69 fathoms. Holotype: USNM 74370. Paratypes: CAS-SU 22384 (7); USNM 70927 (10), 74482-83 (27, 23). Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Artediellus camchaticus Gilbert & Burke 1912 -- (Neyelov 1979:154 [ref. 3152], Yabe in Masuda et al. 1984:328 [ref. 6441], Nelson 1986:43 [ref. 5808], Nakabo 2000:641 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:493 [ref. 25968], Nakabo 2002:641 [ref. 26001], Fedorov et al. 2003:65 [ref. 27579], Parin et al. 2002:S70 [ref. 28300], Mecklenburg et al. 2011:122 [ref. 31212], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:236 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033]). Current status: Valid as Artediellus camchaticus Gilbert & Burke 1912. Cottidae. Distribution: Western North Pacific. Habitat: marine.

camtschaticus, Crossias Schmidt [P. Yu.] 1916:617, Fig. 3 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 3935] Mouth of Osernaya River, western Kamchatka, Russia. Lectotype: ZIN 18963. Paralectotypes: ZIN 18962 (now 7). Also spelled kamtschaticus by authors. Lectotype selected by Neyelov 1979:137 [ref. 3152]. •Valid as Porocottus camtschaticus (Schmidt 1916) -- (Neyelov 1979:137 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:485 [ref. 25968], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S83 [ref. 28300], Fedorov et al. 2003:79 [ref. 27579], Parin et al. 2014:268 [ref. 33547]). Current status: Valid as Porocottus camtschaticus (Schmidt 1916). Cottidae. Distribution: Western North Pacific. Habitat: marine.

canaliculatus, Icelus Gilbert [C. H.] 1896:412, Pl. 24 (lower) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] North of Unalaska, Albatross station 3329, depth 399 fathoms. Lectotype: USNM or CAS-SU 2182. Paralectotypes (incl. potential lectotype): BMNH 1896.7.23.80 (1); MCZ 28346 (1); CAS-SU 2182 (1, poor condition), 18828 (3); USNM 48590 (1), 126722 (1); ZMUO J1459 (1). Type catalog: Böhlke 1953:130 [ref. 12291], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen from Albatross sta. 3329) in caption to Pl. 286, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable; SU 2182 designated lectotype by Nelson 1984:32 [ref. 5391]. •Valid as Icelus canaliculatus Gilbert 1896 -- (Yabe in Amaoka et al. 1983:281, 337 [ref. 8379], Nelson 1984:32 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:634 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:452 [ref. 25968], Nakabo 2002:634 [ref. 26001], Parin et al. 2002:S77 [ref. 28300], Fedorov et al. 2003:71 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:253 [ref. 33547], Kells et al. 2016:178 [ref. 35888]). Current status: Valid as Icelus canaliculatus Gilbert 1896. Cottidae. Distribution: North Pacific: northern Bering Sea to Bristol Bay and to Okhotsk Sea off Hokkaido. Habitat: marine.

carolinae, Potamocottus Gill [T. N.] 1861:41 [Proceedings of the Boston Society of Natural History v. 8 (1861-1862); ref. 1785] Maysville, Kentucky, U.S.A. Syntypes: MCZ 33438 (1), USNM 2859 (lost). •Valid as Cottus carolinae (Gill 1861) -- (Lee et al. 1980:807 [ref. 22416], Robison & Buchanan 1988:476 [ref. 23393], Page & Burr 1991:246 [ref. 18983], Boschung 1992:102 [ref. 23239], Etnier & Starnes 1993:386 [ref. 22809], Jenkins & Burkhead 1994:665 [ref. 21581], Stauffer et al. 1995:251 [ref. 23376], Cross & Collins 1995:183 [ref. 23394], Mettee et al. 1996:493 [ref. 25836], Ross et al. 2001:390 [ref. 25978], Nelson et al. 2004:119 [ref. 27807], Miller & Robison 2004:284 [ref. 27931], Robins 2005:4 [ref. 28230], Proudlove 2006:189 [ref. 28992], Neely et al. 2007:651 [ref. 29262] as "c. carolinae", Scharpf 2008:8 [ref. 30399] with subspecies and remarks, Kinziger & Wood 2010:59 [ref. 30701], Page & Burr 2011:470 [ref. 31215] with 3 subspecies, Adams et al. 2013:491 [ref. 32429], Page et al. 2013:122 [ref. 32708],. Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus carolinae (Gill 1861). Cottidae. Distribution: Eastern U.S.A. Habitat: freshwater.

cataphractus, Agonocottus Pavlenko [M. N.] 1910:23, Figs. 2-3 [Trudy Obshchestva Estestvoispytatelej Imperatorskom' Kazanskom' Universiteti, Kazan v. 42 (no. 2); ref. 3393] Near Cape Povorotnyi, Peter the Great Bay, Russia, Sea of Japan, depth 812-875 feet (= 116-125 Russian fathoms). Holotype (unique): not researched. •Synonym of Icelus spiniger Gilbert 1896, but a valid supspecies -- (Lindberg & Krasyukova 1987:190 [ref. 15964]). •Valid as Icelus cataphractus (Pavlenko 1910) -- (Nelson 1984:28 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Kim & Youn 1992:66 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:633 [ref. 25086], Nakabo 2002:633 [ref. 26001], Youn 2002:262, 575 [ref. 26218], Parin et al. 2002:S77 [ref. 28300], Shinohara et al. 2011:43 [ref. 31715], Song et al. 2012:305 [ref. 36295], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:254 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331]). Current status: Valid as Icelus cataphractus (Pavlenko 1910). Cottidae. Distribution: Northwestern Pacific. Habitat: marine.

caulias, Sigmistes Rutter [C. L.] in Jordan & Evermann 1898:2863 [Bulletin of the United States National Museum No. 47; ref. 2445] Rock pools at Karluk, Kodiak Island, Alaska, U.S.A. Holotype: CAS-SU 5884 (1 of 4). Paratypes: (5) CAS-SU 5884 (3 of 4), USNM 106612 [ex CAS-SU 5884] (1). Type catalog: Böhlke 1953:132 [ref. 12291]. •Valid as Sigmistes caulias Rutter 1898 -- (Yabe et al. 2001:70 [ref. 25246], Mecklenburg et al. 2002:489 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:272 [ref. 33547]). Current status: Valid as Sigmistes caulias Rutter 1898. Cottidae. Distribution: North Pacific: Pribilof Islands and Aleutian Islands, Alaska, to western Gulf of Alaska. Habitat: marine.

cavifrons, Icelinus Gilbert [C. H.] 1890:83 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., Albatross station 2907 and 2945, depth 30-44 fathoms. Syntypes: CAS-SU 69 (1), 148 (1), 28111 (1); USNM 44405 (2), 48247 (1). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelinus cavifrons Gilbert 1890 -- (Bolin 1944:28 [ref. 6379], Eschmeyer & Herald 1983:172 [ref. 9277], Peden 1984:79 [ref. 5193], Castro-Aguirre et al. 1993:92 [ref. 21807], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:176 [ref. 35888]). Current status: Valid as Icelinus cavifrons Gilbert 1890. Cottidae. Distribution: Eastern North Pacific: central California (U.S.A.) to southern Baja California. Habitat: marine.

cayuga, Cottus gracilis Meek [S. E.] 1889:315 [Annals of the New York Academy of Sciences v. 4 (art. 13) (for 1888); ref. 12390] Cayuga Lake, New York, U.S.A. •Synonym of Cottus cognatus Richardson 1837 -- (Smith 1986:358 [ref. 22186]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

centropleura, Uranidea semiscabra var. Eigenmann [C. H.] & Eigenmann [R. S.] 1889:149 [12] [West American Scientist v. 6 (no. 49) (for Nov. 1889); ref. 12402] Allen Springs, Lake County, California, U.S.A. Syntypes: BMNH 1891.5.19.53 [from Eigenmann] (1), USNM 41919 (1). On p. 12 of separate. •Synonym of Cottus gulosus (Girard 1854). Current status: Synonym of Cottus gulosus (Girard 1854). Cottidae. Habitat: freshwater.

centropomus, Podabrus Richardson [J.] 1848:11, Pl. 1 (figs. 7-11) [The zoology of the voyage of H. M. S. Samarang; ref. 3744] Quelpart, Korea Straits. Holotype (unique): BMNH 1982.6.10.1. •Valid as Vellitor centropomus (Richardson 1848) -- (Watanabe 1960:165 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Kim & Youn 1992:66 [ref. 22287], Nakabo 2000:647 [ref. 25086], Shinohara et al. 2000:177 [ref. 25987], Nakabo 2002:647 [ref. 26001], Youn 2002:263, 579 [ref. 26218]). Current status: Valid as Vellitor centropomus (Richardson 1848). Cottidae. Distribution: Western North Pacific. Habitat: marine.

cephaloides, Cottus Gray [J. E.] in Cuvier & Valenciennes 1829:194 [Histoire naturelle des poissons v. 4; ref. 998] Name mentioned in passing under Cottus ventralis, not available; from Kamchatka, Russia. Cottidae.

cervus, Synanceja Tilesius [W. G. von] 1811:278, Pl. 13 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 3; ref. 4408] Kamchatka, Russia. No types known. Syntypes: MNHN 0000-7249 (1). Type information: Svetovidov 1978:27 [ref. 6025]. •Synonym of Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414], Neyelov 1979:146 [ref. 3152], Parin et al. 2002:S73 [ref. 28300], Shinohara et al. 2014:249 [ref. 33330]). Current status: Synonym of Enophrys diceraus (Pallas 1787). Cottidae. Habitat: marine.

chaka, Cottus Hamilton [F.] in McClelland 1839:222 [Asiatic Researches v. 19 (pt 2); ref. 2923] Not available, name only. Cottidae.

chamberlaini, Cottus Evermann [B. W.] & Goldsborough [E. L.] 1907:308, Fig. 66 [Bulletin of the Bureau of Fisheries v. 26 (Doc. 624) (for 1906); ref. 6532] Probably from Loring, Alaska, U.S.A. Holotype: USNM 57823. Paratypes: CAS-SU 20011 (1); UMMZ 156705 [ex USNM 61052] (5); USNM 61052 (213), 124973 [ex USBF 1628 =5230] (1). Type catalog: Böhlke 1953:128 [ref. 12291]. •Synonym of Cottus cognatus Richardson 1837 -- (Lee et al. 1980:808 [ref. 22416], Mecklenburg et al. 2002:468 [ref. 25968]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

chattahoochee, Cottus Neely [D. A.], Williams [J. D.] & Mayden [R. L.] 2007:649, Fig. 1B [Copeia 2007 (no. 3); ref. 29262] 10 kilometers west-northwest of Hamilton,, Blue Springs Branch at Lower Blue Springs road, 32°42'49"N, 84°58'08"W, Harris County, Georgia, USA. Holotype: UAIC 12371.14. Paratypes: many at ANSP, AUM, GMNH, INHS, TU, UF, UMMZ and USNM. Plus many non-type specimens mentioned in Material Examined. •Valid as Cottus chattahoochee Neely, Williams & Mayden 2007 -- (Scharpf 2008:9 [ref. 30399], Page & Burr 2011:477 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus chattahoochee Neely, Williams & Mayden 2007. Cottidae. Distribution: Chattahoochee River drainage above the Fall Line in Georgia, U.S.A. Habitat: freshwater.

clarki, Blennicottus Evermann [B. W.] & Goldsborough [E. L.] 1907:323, Fig. 85 [Bulletin of the Bureau of Fisheries v. 26 (Doc. 624) (for 1906); ref. 6532] Bering Sea, Albatross station 3598. Holotype: USNM 57824. Paratypes: USNM 124987 (1). •Synonym of Artediellus pacificus Gilbert 1896 -- (Neyelov 1979:154 [ref. 3152], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:492 [ref. 25968], Parin et al. 2002:S71 [ref. 28300], Parin et al. 2014:238 [ref. 33547]). Current status: Synonym of Artediellus pacificus Gilbert 1896. Cottidae. Habitat: marine.

claviger, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:195, Pl. 79 (fig. 2) [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka, Russia. Holotype (unique): BMNH 2011.5.17.1. Type catalog: Blanc & Hureau 1968:60 [ref. 20738]. •Synonym of Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414], Neyelov 1979:146 [ref. 3152], Lindberg & Krasyukova 1987:229 [ref. 15964], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:472 [ref. 25968], Parin et al. 2002:S74 [ref. 28300], Parin et al. 2014:245 [ref. 33547]). Current status: Synonym of Enophrys diceraus (Pallas 1787). Cottidae. Habitat: marine.

cognatus, Cottus Richardson [J.] 1836:40 [Part 3; ref. 3731] Great Bear Lake, Canada, 64.5°N. Syntypes: (many) MNHN 0000-6540 (1). Type catalog: Blanc & Hureau 1968:50-51 [ref. 20738]. •Valid as Cottus cognatus Richardson 1836, subspecies cognatus -- (Chereshnev et al. 2001:118 [ref. 26736]). •Valid as Cottus cognatus Richardson 1836 -- (Lee et al. 1980:808 [ref. 22416], Sideleva 1982:82 [ref. 14469], Smith 1986:356 [ref. 22186], McAllister 1990:196 [ref. 14674], Page & Burr 1991:249 [ref. 18983], Jenkins & Burkhead 1994:647 [ref. 21581], Stauffer et al. 1995:252 [ref. 23376], Chereshnev 1996:606 [ref. 24659], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:151 [ref. 23595], Scott & Crossman 1998:830 [ref. 24570], Kinziger et al. 2000:1007 [ref. 25019], Lyons et al. 2000:42 [ref. 26594], Sheiko & Fedorov 2000:27 [ref. 25650], Bogutskaya et al. 2001:46 [ref. 26178], Mecklenburg et al. 2002:468 [ref. 25968], Parin et al. 2002:S72 [ref. 28300], Wydoski & Whitney 2003:207 [ref. 27816], Bailey et al. 2004:171 [ref. 27882], Nelson et al. 2004:119 [ref. 27807], Bogutskaya & Naseka 2004:186 [ref. 28183], McPhail 2007:462 [ref. 31199], Scharpf 2008:9 [ref. 30399], Page & Burr 2011:474 [ref. 31215], Page et al. 2013:122 [ref. 32708], LeMone et al. 2014:249 [ref. 33134], Goto et al. 2014:[10] [ref. 33456], Parin et al. 2014:242 [ref. 33547]). Current status: Valid as Cottus cognatus Richardson 1836. Cottidae. Distribution: Canada and U.S.A. (but eastern North Asia and Western North America and Arctic on ref. 33547). Habitat: freshwater, brackish.

comephoroides, Cottus Berg [L. S.] 1900:338 [13], Pl. 8 (fig. 3 [not 1]) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 5; ref. 14714] Lake Baikal at Selenge River mouth, Russia, depth 255 meters. Syntypes: BMNH 1905.12.4.18 (1); ZIN 11531-32 (19, 9), 11533-35 (20, 3, 4). On p. 13 of separate. •Synonym of Cottocomephorus inermis (Yakovlev 1890) -- (Taliev 1955:292 [ref. 4316], Sideleva 1982:29 [ref. 14469], Sideleva 1998:150 [ref. 23595], Sideleva 2003:166 [ref. 27815]). •Valid as Cottocomephorus comephoroides (Berg 1900) -- (Berg 1949:1173 [ref. 20662], Bogdanov 2019:230 [ref. 36814] with date 1901). Current status: Valid as Cottocomephorus comephoroides (Berg 1900). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

concinnum, Stelgistrum Andriashev [A. P.] 1935:292, Fig. 1 [Zoologischer Anzeiger v. 111 (nos 11/12); ref. 13868] Cape Olutorsky, Bering Sea, depth 32 meters. Holotype: ZIN 24934. Paratypes: ZIN 21456 (2). •Valid as Stelgistrum concinnum Andriashev 1935 -- (Mecklenburg et al. 2002:448 [ref. 25968], Parin et al. 2002:S86 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:272 [ref. 33547]). Current status: Valid as Stelgistrum concinnum Andriashev 1935. Cottidae. Distribution: North Pacific: southern Bering Sea and Aleutian Islands. Habitat: marine.

confusus, Cottus Bailey [R. M.] & Bond [C. E.] 1963:12, Figs. 1c, 3a, 4c [Occasional Papers of the Museum of Zoology University of Michigan No. 634; ref. 3934] Salmon River, tributary to Snake River, 25 miles northwest of Ketchum, Blaine County, Idaho, U.S.A. Holotype: UMMZ 177377. Paratypes: UMMZ 161840 (39). Additional material at UBC, MSU, OS, UMMZ, and UW. •Valid as Cottus confusus Bailey & Bond 1963 -- (Lee et al. 1980:809 [ref. 22416], McAllister 1990:196 [ref. 14674], Page & Burr 1991:251 [ref. 18983], Scott & Crossman 1998:835 [ref. 24570], Wydoski & Whitney 2003:209 [ref. 27816], Nelson et al. 2004:119 [ref. 27807], McPhail 2007:469 [ref. 31199], Scharpf 2008:9 [ref. 30399], Page & Burr 2011:480 [ref. 31215], Page et al. 2013:122 [ref. 32708], LeMone et al. 2014:249 [ref. 33134], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus confusus Bailey & Bond 1963. Cottidae. Distribution: Columbia River drainage and Puget Sound drainage, southwestern Canada and northwestern U.S.A. Habitat: freshwater.

copei, Cottus Abbott [C. C.] 1860:326 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 12; ref. 15620] Brandywine Creek tributaries in Pennsylvania, U.S.A. Holotype: ANSP 22713 (poor condition). Paratypes: ANSP 22714 (1). Type catalog: Böhlke 1984:65 [ref. 13621]. •Synonym of Cottus cognatus Richardson 1837. Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater.

corallinus, Allartedius Hubbs [C. L.] 1926:8 [Occasional Papers of the Museum of Zoology University of Michigan No. 170; ref. 6069] California, U.S.A. Holotype: UMMZ 141868. •Valid as Artedius corallinus (Hubbs 1926) -- (Bolin 1944:53 [ref. 6379], Eschmeyer & Herald 1983:161 [ref. 9277], Nelson et al. 2004:118 [ref. 27807], Pietsch & Orr 2015:72 [ref. 34694], Kells et al. 2016:162 [ref. 35888]). Current status: Valid as Artedius corallinus (Hubbs 1926). Cottidae. Distribution: Eastern North Pacific: Washington, U.S.A. to Baja California. Habitat: marine.

corniger, Artediellus europeus Andriashev [A. P.] 1954:407, Fig. 228 [ ref. 6547] Northern parts of Barents [not Bering], Kara, and Laptev seas. Syntypes: ZIN 32202 (?). •Synonym of Artediellus atlanticus, but a valid subspecies -- (Parin et al. 2002:S70 [ref. 28300]). •Synonym of Artediellus atlanticus Jordan & Evermann 1898 -- (Neyelov 1973:594 [ref. 7219], Neyelov 1979:153 [ref. 3152], Parin et al. 2014:236 [ref. 33547]). Current status: Synonym of Artediellus atlanticus Jordan & Evermann 1898. Cottidae. Habitat: marine.

coronatus, Porocottus Yabe [M.] 1992:361, Figs. 1-3 [Japanese Journal of Ichthyology v. 38 (no. 4); ref. 19519] Near Konbumori, Pacific coast of Hokkaido, 42°56.5'N, 144°38.5'E, depth 0.5 meters. Holotype: HUMZ 111575. Paratypes: HUMZ 109999-110001 (3), 117471 (1); NSMT-P 35034 (1). •Valid as Porocottus coronatus Yabe 1992 -- (Nakabo 2000:646 [ref. 25086], Nakabo 2002:646 [ref. 26001], Muto et al. 2002:231 [ref. 26331]). Current status: Valid as Porocottus coronatus Yabe 1992. Cottidae. Distribution: Western North Pacific. Habitat: marine.

cottoides, Podabrus Richardson [J.] 1848:13, Pl. 1 (figs. 1-6) [The zoology of the voyage of H. M. S. Samarang; ref. 3744] China Sea. Syntypes: BMNH 1848.3.16.102 (1), 1855.9.19.399 (1). •Valid as Pseudoblennius cottoides (Richardson 1848) -- (Watanabe 1960:150 [ref. 11959], Nakabo in Masuda et al. 1984:330 [ref. 6441], Lindberg & Krasyukova 1987:252 [ref. 15964], Kim & Youn 1992:74 [ref. 22287], Mizuno et al. 1999:7 [ref. 25311], Nakabo 2000:650 [ref. 25086], Shimizu 2001:28 [ref. 25799], Nakabo 2002:650 [ref. 26001], Youn 2002:263, 577 [ref. 26218]). Current status: Valid as Pseudoblennius cottoides (Richardson 1848). Cottidae. Distribution: Western North Pacific. Habitat: marine.

crassus, Icelus uncinalis Andriashev [A. P.] 1937:265, Pl. 5 (fig. 2) [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] Penzhina Bay, northeastern Sakhalin Island, Okhotsk Sea. Syntypes: ZIN 21900 (1), 21905 (6), 21910 (2), 25281 (1). "Typus" given on caption to Pl. 5 but specimen not distinguished in text. •Synonym in part of Icelus bicornis (Reinhardt 1840) -- (Nelson 1984:42 [ref. 5391]). •Synonym in part of Icelus ochotensis Schmidt 1927 -- (Nelson 1984:50 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300]). •Synonym in part of Icelus spatula Gilbert & Burke 1912 -- (Sheiko & Fedorov 2000:28 [ref. 25650]). •Synonym of Icelus ochotensis Schmidt 1927 -- (Parin et al. 2014:255 [ref. 33547]). Current status: Synonym of Icelus ochotensis Schmidt 1927. Cottidae. Habitat: marine.

creaseri, Ruscariops Hubbs [C. L.] 1926:12 [Occasional Papers of the Museum of Zoology University of Michigan No. 170; ref. 6069] Rock pool at Bird Rock, San Diego, California, U.S.A. Holotype: UMMZ 141866. Paratypes: (9) UMMZ 141863-65 (1, 1, 6), ?141865 (6). •Valid as Artedius creaseri (Hubbs 1926) -- (Bolin 1944:43 [ref. 6379], Eschmeyer & Herald 1983:162 [ref. 9277]). •Valid as Ruscarius creaseri (Hubbs 1926) -- (Begle 1989:648 [ref. 12739], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Kells et al. 2016:190 [ref. 35888]). Current status: Valid as Ruscarius creaseri (Hubbs 1926). Cottidae. Distribution: Eastern North Pacific: central California (U.S.A.), to central Baja California. Habitat: marine.

criniger, Cottus Günther [A.] 1860:522 [Catalogue of the fishes in the British Museum v. 2; ref. 1963] Monterey, California, U.S.A. Syntypes: (3) BMNH. •Synonym of Clinocottus analis (Girard 1858) -- (Bolin 1944:73 [ref. 6379]). Current status: Synonym of Clinocottus analis (Girard 1858). Cottidae. Habitat: marine.

ctenopterygius, Phasmatocottus Bolin [R. L.] 1936:33, Fig. 7 [Proceedings of the United States National Museum v. 84 (no. 3000); ref. 505] Off Sendai Bay, Japan, 38°11'30"N, 142°08'00"E, Albatross station 5050, depth 266 fathoms. Holotype (unique): USNM 102107. •Valid as Phasmatocottus ctenopterygius Bolin 1936 -- (Yabe in Masuda et al. 1984:328 [ref. 6441], Nakabo 2000:639 [ref. 25086], Yabe & Fujii 2001:47 [ref. 25387], Nakabo 2002:639 [ref. 26001], Parin et al. 2014:267 [ref. 33547], Prokofiev 2017:648 [ref. 35721]). Current status: Valid as Phasmatocottus ctenopterygius Bolin 1936. Cottidae. Distribution: Western North Pacific. Habitat: marine.

czerskii, Cottus Berg [L. S.] 1913:17, Pl. [Zapiski obshchestva izucheniya Amurskogo Kraya [The Acta of the Society for the Study of the Amur Region] v. 13; ref. 14943] Sedanka River, Vladivostok, Russia. Holotype: ZIN 15853. Second specimen mentioned on p. 21 [ZISP 14983]. •Valid as Cottus czerskii Berg 1913 -- (Berg 1949:1140 [ref. 20662], Sideleva 1982:82 [ref. 14469], Lindberg & Krasyukova 1987:171 [ref. 15964], Kim & Youn 1992:60 [ref. 22287], Zhu 1995:189 [ref. 25213], Kim 1997:406 [ref. 24032], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:151 [ref. 23595], Sokolovskaya et al. 1998:9 [ref. 24670], Kim & Park 2002:326 [ref. 25913], Bogutskaya et al. 2001:46 [ref. 26178], Youn 2002:262, 572 [ref. 26218], Parin et al. 2002:S73 [ref. 28300], Bogutskaya & Naseka 2004:186 [ref. 28183], Fujii et al. 2005:12 [ref. 28191], Goto et al. 2014:[9] [ref. 33456], Parin et al. 2014:242 [ref. 33547], Zhang et al. 2016:193 [ref. 34477]). Current status: Valid as Cottus czerskii Berg 1913. Cottidae. Distribution: Continental coast of the Sea of Japan from Serebryanka River, Russia to the North Nandai River, Korea. Habitat: freshwater.

dabryi, Centridermichthys Sauvage [H.-E.] 1878:146, Pl. 1 (fig. 14) [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris (Série 2) v. 1; ref. 3880] South China. Holotype (unique): MNHN 0000-7359. Type catalog: Blanc & Hureau 1968:49 [ref. 20738]. Cottidae. Habitat: marine.

decastrensis, Cottus Kner [R.] 1865:2, Pl. 2 (fig. 1) [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 24; ref. 6174] Decastris Bay at Amur River mouth, northeastern Asia. Holotype (unique): NMW 80072. Appeared as Cottus decastrensis n. sp.? •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Berg 1949:1137 [ref. 20662], Neyelov 1979:121 [ref. 3152], Lindberg & Krasyukova 1987:207 [ref. 15964], Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:265 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138] with question). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Cottidae. Habitat: brackish, marine.

delacyi, Artedius Hubbs [C. L.] & Schultz [L. P.] 1941:4 [Occasional Papers of the Museum of Zoology University of Michigan No. 431; ref. 10913] Tidepools at Uyak Bay, Kodiak Island, Alaska, U.S.A. Holotype: UMMZ 126490. Paratypes: UMMZ 126491 (1), USNM 117494 (1). •Synonym of Artedius lateralis (Girard 1854) -- (Mecklenburg et al. 2002:459 [ref. 25968]). Current status: Synonym of Artedius lateralis (Girard 1854). Cottidae. Habitat: marine.

derjavini, Radulinopsis Soldatov [V. K.] & Lindberg [G. U.] 1930:185, Figs. 30-31 [Izvestiia Tikhookeanskogo nauchnogo instituta rybnogo khoziaistva [Bulletins of the Pacific Science Institute] v. 5; ref. 4164] Peter the Great Bay, Russia. Syntypes: (2) not at ZIN. •Valid as Radulinopsis derjavini Soldatov & Lindberg 1930 -- (Lindberg & Krasyukova 1987:187 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670] as derzhavini, Yabe & Maruyama 2001:53 [ref. 25245], Parin et al. 2002:S85 [ref. 28300], Parin et al. 2014:270 [ref. 33547]). Current status: Valid as Radulinopsis derjavini Soldatov & Lindberg 1930. Cottidae. Distribution: Western North Pacific. Habitat: marine.

derjugini, Icelus bicornis Soldatov [V. K.] in Knipowitsch 1926:124 [Trudy Nauchno-issledovatel'skogo instituta po izucheniya Severea. Moskva [= Transactions of the Institute for Scientific Exploration of the North.] No. 27; ref. 22166] Naiden, White Sea. Holotype: NIISM (whereabouts unknown). •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neyelov 1973:596 [ref. 7219], Nelson 1984:36 [ref. 5391], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:253 [ref. 33547]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Cottidae. Habitat: marine.

derjugini, Radulinopsis Soldatov [V. K.] in Soldatov & Lindberg 1930:189, Figs. 32-33 [Izvestiia Tikhookeanskogo nauchnogo instituta rybnogo khoziaistva [Bulletins of the Pacific Science Institute] v. 5; ref. 4164] Peter the Great Bay, Russia. Syntypes: (2) not at ZIN. •Valid as Radulinopsis derjugini Soldatov 1930 -- (Lindberg & Krasyukova 1987:188 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670]). •Synonym of Radulinopsis derjavini Soldatov & Lindberg 1930 -- (Yabe & Maruyama 2001:53 [ref. 25245]). Current status: Synonym of Radulinopsis derjavini Soldatov & Lindberg 1930. Cottidae. Habitat: marine.

detrisus, Gymnocanthus Gilbert [C. H.] & Burke [C. V.] 1912:61, Fig. 13 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off west coast of Kamchatka, Russia, 51°37'N, 156°21'E, Albatross station 4798, depth 25 fathoms. Holotype: USNM 74376. Paratypes: CAS-SU 22361 (2), USNM 74499 (2). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Gymnocanthus detrisus Gilbert & Burke 1912 -- (Watanabe 1960:61 [ref. 11959], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:200 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:629 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Mecklenburg et al. 2002:463 [ref. 25968], Nakabo 2002:629 [ref. 26001], Parin et al. 2002:S74 [ref. 28300] as Gymnacanthus, Fedorov et al. 2003:66 [ref. 27579] in Gymnacanthus, Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:246 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331]). Current status: Valid as Gymnocanthus detrisus Gilbert & Burke 1912. Cottidae. Distribution: Circumpolar: North Pacific, primarily western North Pacific: Bering Sea off Alaska to Pacific and Okhotsk coasts of Hokkaido. Habitat: marine.

diceraus, Cottus Pallas [P. S.] 1787:354, Pl. 10 (fig. 7) [Nova Acta Academiae Scientiarum Imperialis Petropolitanae v. 1 (Physica) (for 1783); ref. 18507] Kamchatka, Russia. Syntypes: ZIN 5380 (1); ZMB 5229 (not found), 6488 (1, dry) Type catalog: Paepke & Fricke 1992:282 [ref. 19946]. Date may be 1788. Type information: Svetovidov 1978:27 [ref. 6025]. •Valid as Ceratocottus diceraus (Pallas 1787) -- (Watanabe 1960:110 [ref. 11959]). •Valid as Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414], Neyelov 1979:146 [ref. 3152], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:229 [ref. 15964], Kim & Youn 1992:61 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:630 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:472 [ref. 25968], Nakabo 2002:630 [ref. 26001], Youn 2002:262, 574 [ref. 26218], Parin et al. 2002:S73 [ref. 28300], Fedorov et al. 2003:68 [ref. 27579] dated 1788, Nelson et al. 2004:119 [ref. 27807] dated 1788, Mecklenburg et al. 2007:174 [ref. 29448], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:245 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Tohkairin et al. 2015:7 [ref. 35331], Mecklenburg et al. 2016:53 [ref. 34440], Kells et al. 2016:170 [ref. 35888], Mecklenburg et al. 2018:192 [ref. 36370]). Current status: Valid as Enophrys diceraus (Pallas 1787). Cottidae. Distribution: North Pacific and adjacent Arctic. Habitat: marine.

distoechus, Stlengis Bolin [R. L.] 1936:328, Pl. 34 (fig. A); Fig. 26 [Proceedings of the United States National Museum v. 83 (no. 2987); ref. 504] Wakayama, Japan, 33°24'50"N, 135°38'40"E, Albatross station 4968, depth 253 fathoms. Holotype: USNM 94728. Paratypes: CAS-SU 28727 (1), USNM 94729 (3). Type catalog: Böhlke 1953:132 [ref. 12291]. •Valid as Stlengis distoechus Bolin 1936 -- (Watanabe 1960:37 [ref. 11959], Fricke & Brunken 1984:46 [ref. 5262], Nakabo 2000:630 [ref. 25086], Shinohara et al. 2001:319 [ref. 25995], Nakabo 2002:630 [ref. 26001]). Current status: Valid as Stlengis distoechus Bolin 1936. Cottidae. Distribution: Western North Pacific. Habitat: marine.

dorothy, Triglops Pietsch [T. W.] & Orr [J. W.] 2006:239, Fig. 1 [Fishery Bulletin v. 104 (no. 2); ref. 28666] Aniva Bay, 46°17'N, 143°09'E, Albatross station 5013, depth 77 meters. Holotype: USNM 74578. Paratypes: CAS-SU 22305 (3); HUMZ 41524 (1), 41525 (1); USNM 74575 (2), 74576 (4), 74577 (1), 74579 (8). •Valid as Triglops dorothy Pietsch & Orr 2006 -- (Parin et al. 2014:275 [ref. 33547]). Current status: Valid as Triglops dorothy Pietsch & Orr 2006. Cottidae. Distribution: Sea of Okhotsk, Japan and Kuril Islands. Habitat: marine.

dracunculus, Cottus Forsskål [P. S.] 1775:xviii [Descriptiones animalium (Forsskål); ref. 1351] Not available, name only; from Malta. Cottidae.

duranii, Cottus Freyhof [J.], Kottelat [M.] & Nolte [A.] 2005:136, Fig. 14 [Ichthyological Exploration of Freshwaters v. 16 (no. 2); ref. 28259] Stream Epie, tributary of Tryuètr. at Valette, Dept. Cantal, Dordogne drainage, France. Holotype: MNHN 2005-0754. Paratypes: CMK 18276 (3), 18270 (4); MNHN 2005-0755 (2), 2005-0756 (2). Plus non-type material. •Valid as Cottus duranii Freyhof, Kottelat & Nolte 2005 -- (Kottelat & Freyhof 2007:508 [ref. 29996], Sideleva 2009:211 [ref. 30173], Goto et al. 2014:[10] [ref. 33456], Sideleva et al. 2015:33 [ref. 35276], Sideleva et al. 2018:9 [ref. 36393]). Current status: Valid as Cottus duranii Freyhof, Kottelat & Nolte 2005. Cottidae. Distribution: France. Habitat: freshwater.

dybowskii, Comephorus Korotneff [A. de] 1904:17, Figs. 9-10 [Archives de zoologie expérimentale et générale (Paris) ser. 4, v. 2; ref. 15188] Lake Baikal, Russia. Holotype (?): ZIN 12474. Correct spelling may be dybowski; two publications may be involved; date may be 1905. •Valid as Comephorus dybowskii Korotneff 1904 -- (Berg 1949:1176 [ref. 20662], Taliev 1955:375 [ref. 4316], Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:759 [ref. 24702] as dybowski, Sideleva 1998:154 [ref. 23595], Yabe & Sideleva 2000:306 [ref. 25698], Kontula et al. 2003:149 [ref. 26791] as dybowski, Sideleva 2003:17, 172 [ref. 27815] as dybowski and date as 1905:12, Bogutskaya & Naseka 2004:193 [ref. 28183] as dybowski, Goto et al. 2014:[15] [ref. 33456] as dybowskii). Current status: Valid as Comephorus dybowskii Korotneff 1904. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

dybowskii, Cottus Hilgendorf [F. M.] 1879:106 [Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1879; ref. 21467] No locality [Japan]. Holotype (unique): ZMB 10628. Type catalog: Paepke & Fricke 1992:282 [ref. 19946]. •Synonym of Myoxocephalus brandtii (Steindachner 1867) -- (Prokofiev 2017:648 [ref. 35721]). Current status: Synonym of Myoxocephalus brandtii (Steindachner 1867). Cottidae.

dydymovi, Artediellus Soldatov [V. K.] 1915:157, Figs. 1 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 15068] Tatar Strait, 49°28'10"N, 140°39'30"E, Russia, northern Sea of Japan. Holotype: ZIN 18693 (male of 3). Paratypes: ZIN 18653 (6), 18655-63 (6, 6, 4, 3, 2, 1, 1, 1, 1), 18693 (2 females), 19435 (1). •Valid as Artediellus dydymovi Soldatov 1915, subspecies dydymovi Soldatov 1915 -- (Nakabo 2000:642 [ref. 25086], Nakabo 2002:642 [ref. 26001]). •Valid as Artediellus dydymovi Soldatov 1915 -- (Neyelov 1979:155 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Nelson 1986:43 [ref. 5808], Lindberg & Krasyukova 1987:235 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Sheiko & Fedorov 2000:27 [ref. 25650], Parin et al. 2002:S70 [ref. 28300], Fedorov et al. 2003:66 [ref. 27579], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:236 [ref. 33547], Dyldin & Orlov 2017:61 [ref. 35138]). Current status: Valid as Artediellus dydymovi Soldatov 1915. Cottidae. Distribution: Western North Pacific. Habitat: brackish, marine.

dzungaricus, Cottus Kottelat [M.] 2006:60, 90 [Fishes of Mongolia; ref. 28831] Altai, northern Sinkiang, China. Holotype: ASIZB NK 64-997. Paratypes: ASIZB 64-979 to 980 (2), 64-983 to 985 (3), 64-988 (1), 64-992 (1); Acad. Sinica, Sinkiang, Bureau of Water Conservancy and Dept. Biol. Sinkiang Univ. 64-981 to 982 (2), 64-986 to 987 (2), 64-989 to 991 (3). Relacement name for Cottus sibiricus altaicus Li & Ho 1966, preoccupied by Cottus poecilopus altaicus Kaschenko 1899. •Valid as Cottus dzungaricus Kottelat 2006 -- (Goto et al. 2014:[9] [ref. 33456]). •Possible synonym of Cottus sibiricus Warpachowski 1889 -- (Sideleva 2017:827 [ref. 35715]). Current status: Valid as Cottus dzungaricus Kottelat 2006. Cottidae. Distribution: Chinese Altai, China. Habitat: freshwater.

echinatus, Cottus Bailey [R. M.] & Bond [C. E.] 1963:2, Figs. 1a, 2a, 4a [Occasional Papers of the Museum of Zoology University of Michigan No. 634; ref. 3934] Utah Lake, mouth of Provo River, Utah County, Utah, U.S.A. Holotype: UMMZ 177376. Paratypes: UMMZ 156794 (2); USNM 27414 (2), 197681 (2). •Valid as Cottus echinatus Bailey & Bond 1963 -- (Lee et al. 1980:810 [ref. 22416], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:9 [ref. 30399], Page & Burr 2011:474 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus echinatus Bailey & Bond 1963. Cottidae. Distribution: Utah Lake, Utah, U.S.A [extinct]. Habitat: freshwater.

ecornis, Icelus Tsutsui [D.] & Yabe [M.] 1996:302, Fig. 2 [Ichthyological Research v. 43 (no. 3); ref. 22432] Kitami-Yamato Bank, southwestern Okhotsk Sea, off Abashiri, Hokkaido, Japan, 44°30.1'N, 144°14.5'E, depth 208 meters. Holotype: HUMZ 125990. Paratypes: CAS 82510 (2); HUMZ 124100-01 (1, 1), 124326 (1), 124832-34 (1, 1, 1), 124836 (1), 124838 (1), 124845 (1), 124848 (1), 124852 (1), 125966-67 (1, 1), 125975 (1), 125980 (1), 125985 (1), 125991 (1), 125999 (1), 126000 (1), 126002 (1), 126006 (1), 126018 (1), 126020-21 (1, 1); NSMT-P 47286-87 (1, 1); UMMZ 230706 (2). •Valid as Icelus ecornis Tsutsui & Yabe 1996 -- (Nakabo 2000:635 [ref. 25086], Nakabo 2002:635 [ref. 26001], Tsuruoka et al. 2006:51 [ref. 28501]). Current status: Valid as Icelus ecornis Tsutsui & Yabe 1996. Cottidae. Distribution: Western North Pacific. Habitat: marine.

edomius, Myoxocephalus Jordan [D. S.] & Starks [E. C.] 1904:274, Fig. 18 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Aino village of Edomo, near Mororan, Japan. Holotype: CAS-SU 7707. Paratypes: USNM 50919 (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Myoxocephalus edomius Jordan & Starks 1904 -- (Watanabe 1960:68 [ref. 11959]). •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Neyelov 1979:122 [ref. 3152], Lindberg & Krasyukova 1987:212 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:261 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Cottidae. Habitat: marine.

elegans, Antipodocottus Fricke [R.] & Brunken [H.] 1984:41, Figs. 1-2 [Journal of Natural History v. 18 (no. 1); ref. 5262] Off Wollongong, New South Wales, Australia, 34°21'S, 151°24'E, depth 220 fathoms. Holotype: AMS I.18795-007. •Valid as Antipodocottus elegans Fricke & Brunken 1984 -- (Paxton et al. 1989:477 [ref. 12442], Nelson 1990:841 [ref. 15720], Nelson & Carpenter 1999:2426 [ref. 24809], Paxton et al. 2006:952 [ref. 28994], Fricke et al. 2014:56 [ref. 33932], Stewart 2015:1118 [ref. 34254]). Current status: Valid as Antipodocottus elegans Fricke & Brunken 1984. Cottidae. Distribution: Western Pacific: Australia, Papua New Guinea and New Zealand region. Habitat: marine.

elegans, Centridermichthys Steindachner [F.] 1881:185 [7], Pl. 6 (figs. 1, 1a) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 83 (1. Abth.); ref. 4231] Strietok, near Vladivostock, Russia, Sea of Japan. Syntypes: NMW 6374 (3), 6578 (2), 78819 (1). On p. 7 of separate. •Valid as Bero elegans (Steindachner 1881) -- (Watanabe 1960:134 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:250 [ref. 15964], Kim & Youn 1992:72 [ref. 22287], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:648 [ref. 25086], Nakabo 2002:648 [ref. 26001], Youn 2002:262, 573 [ref. 26218], Parin et al. 2002:S72 [ref. 28300], Parin et al. 2014:240 [ref. 33547]). Current status: Valid as Bero elegans (Steindachner 1881). Cottidae. Distribution: Western North Pacific. Habitat: marine.

elegans, Cottus Gray [J. E.] in Cuvier & Valenciennes 1829:195 [Histoire naturelle des poissons v. 4; ref. 998] Name only mentioned under Cottus claviger. •In the synonymy of Enophrys diceraus (Pallas 1787). Current status: Synonym of Enophrys diceraus (Pallas 1787). Cottidae.

elegans, Limnocottus megalops Taliev [D. N.] 1948:113 [Trudy Baikal'skoi Limnologicheskoi Stantsii v. 12; ref. 19628] Not available, manuscript name mentioned in table; mentioned in synonymy in Taliev 1955:329 [ref. 4316]; from Lake Baikal. Current status: Synonym of Asprocottus. Cottidae.

elochini, Abyssocottus Taliev [D. N.] 1955:350, Fig. 147 [ ref. 4316] Northern Lake Baikal, Russia, depth 250-300 meters. Syntypes: (3) ZIN 46661 (1). Sideleva 2001:60 [ref. 26181] lists a holotype [ZIN 46661], but that is incorrect; that specimen is a "surviving" syntype and not the holotype. •Valid as Abyssocottus elochini Taliev 1955 -- (Taliev 1955:350 [ref. 4316], Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:155 [ref. 23595], Bogutskaya et al. 2001:47 [ref. 26178], Sideleva 2001:60 [ref. 26181], Sideleva 2003:17, 177 [ref. 27815], Bogutskaya & Naseka 2004:194 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Abyssocottus elochini Taliev 1955. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

elongatus, Centridermichthys Steindachner [F.] 1881:186 [8] [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 83 (1. Abth.); ref. 4231] Sea of Japan. On p. 8 of separate. Apparently figured as Centridermichthys glaber n. sp. in Pl. 6 (fig. 2). See account of Centridermichthys glaber Steindachner 1881. •Valid as Alcichthys elongatus (Steindachner 1881) -- (Lindberg & Krasyukova 1987:243 [ref. 15964] as Elophichthys, Nakabo 2000:648 [ref. 25086], Nakabo 2002:648, 1526 [ref. 26001], Parin et al. 2002:S68 [ref. 28300], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:232 [ref. 33547]). Current status: Valid as Alcichthys elongatus (Steindachner 1881). Cottidae. Distribution: Western North Pacific. Habitat: marine.

embryum, Oligocottus Jordan [D. S.] & Starks [E. C.] 1895:808, Pl. 82 [Proceedings of the California Academy of Sciences (Series 2) v. 5; ref. 2522] Several miles east of Neah Bay, Washington, U.S.A. Lectotype: CAS-SU 3128. Paralectotypes: ?BMNH 1897.1.12.16 (1), CAS-SU 3128 (1 of 2). Additional material: (1). Type catalog: Böhlke 1953:131 [ref. 12291]. Lectotype established (as figured specimen) in caption to Pl. 303, p. 3294 in Jordan & Evermann 1900 [ref. 2446]. •Valid as Clinocottus embryum (Jordan & Starks 1895) -- (Bolin 1944:78 [ref. 6379], Eschmeyer & Herald 1983:166 [ref. 9277], Matarese et al. 1989:424 [ref. 26885], McAllister 1990:196 [ref. 14674], Mecklenburg et al. 2002:503 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:39 [ref. 34694], Kells et al. 2016:168 [ref. 35888]). Current status: Valid as Clinocottus embryum (Jordan & Starks 1895). Cottidae. Distribution: Eastern North Pacific: Aleutian Islands to northern Baja California. Habitat: marine.

emeljanovi, Cottus Lindberg [G. U.] 1927:7, fig. [Trudy Gosudarstvennogo Dal'nevostocnogo Universiteta. (Ser. 8. Biologija) No. 2. No. 2; ref. 16497] Botchi River, Primorye, southeastern Russia. Lectotype: ZIN 21988. Paralectotypes: (1). Lectotype selected by Berg 1949:1147 [ref. 20662]. •Synonym of Cottus amblystomopsis Schmidt 1904 -- (Berg 1949:1142 [ref. 20662], Sideleva 1998:150 [ref. 23595], Parin et al. 2002:S72 [ref. 28300], Parin et al. 2014:241 [ref. 33547], Dyldin & Orlov 2017:62 [ref. 35138]). Current status: Synonym of Cottus amblystomopsis Schmidt 1904. Cottidae. Habitat: freshwater, brackish, marine.

ensiger, Ainocottus Jordan [D. S.] & Starks [E. C.] 1904:283, Fig. 23 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate,Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7709. Paratypes: USNM 50921 (not found in 1980). Additional material: CAS-SU 7596 (4). Type catalog: Böhlke 1953:126 [ref. 12291]. •Valid as Ainocottus ensiger Jordan & Starks 1904 -- (Watanabe 1960:87 [ref. 11959], Kim & Youn 1992:63 [ref. 22287]). •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Neyelov 1979:121 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Youn 2002:576 [ref. 26218], Nakabo 2002:1526 [ref. 26001], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547], Mecklenburg et al. 2016:79 [ref. 34440], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Cottidae. Habitat: marine.

europaeus, Artediellus Knipowitsch [N. M.] 1907:17 [Zapiski Imperatorskoj Akademii Nauk po Fiziko-Matematiceskomu Otdelniji = Mémoires de l'Académie Impériale des Sciences de St. Petersbourg, Classe Physico-Mathématique (8 Sér.) v. 18 (no. 5); ref. 15152] One of: west coast of Norway, Svalbard, and European Arctic Ocean. Lectotype: ZIN 28393. Paralectotypes: (19) ZIN 28393 (4, with lectotype), 28449 (6+). Lectotype selected by Neyelov 1979:154 [ref. 3152]. •Synonym of Artediellus atlanticus Jordan & Evermann 1898, but a valid subspecies -- (Neyelov 1973:594 [ref. 7219], Neyelov 1979:154 [ref. 3152], Fedorov in Whitehead et al. 1986:1245 [ref. 13677] as europeus, Yabe in Okamura et al. 1995:161 [ref. 22531], Parin et al. 2002:S70 [ref. 28300], Dolgov 2013:918 [ref. 33112] as europeus). •Synonym of Artediellus atlanticus Jordan & Evermann 1898 -- (Parin et al. 2014:236 [ref. 33547]). Current status: Synonym of Artediellus atlanticus Jordan & Evermann 1898. Cottidae. Habitat: marine.

euryops, Icelus Bean [T. H.] 1890:41 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Off Trinity Island, Alaska, U.S.A., Albatross station 2853, 56°N, 154°20'W, depth 159 fathoms. Lectotype: USNM 45367. Paralectotypes: USNM 227363 [ex USNM 45367] (2). Lectotype selected by Nelson 1984:30 [ref. 5391]. •Valid as Icelus euryops Bean 1890 -- (Nelson 1984:30 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:453 [ref. 25968], Parin et al. 2002:S77 [ref. 28300], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:254 [ref. 33547]). Current status: Valid as Icelus euryops Bean 1890. Cottidae. Distribution: North Pacific: northern Bering Sea to western Gulf of Alaska. Habitat: marine.

euryptera, Bolinia Yabe [M.] 1991:329, Figs. 1-3 [Copeia 1991 (no. 2); ref. 17338] Near Amukta Pass, Aleutian Islands, 52°51'N, 171°13'W, depth 205-225 meters. Holotype: HUMZ 68566. Paratypes: CAS 68097-98 (2); HUMZ 68293 (1), 68296-98 (3), 68552-53 (2), 68555-60 (6), 68567-68 (2), 88847-48 (2), 88869-72 (4), 89107-111 (5), 89113-123 (11), 89049-53 (5), 110007-10 (4), 110015-17 (3); NSMT-P 45082-83 (2), OSOU 10878-79 (5, 2); plus additional non-type material. •Valid as Bolinia euryptera Yabe 1991 -- (Mecklenburg et al. 2002:488 [ref. 25968]). Current status: Valid as Bolinia euryptera Yabe 1991. Cottidae. Distribution: North Pacific: Aleutian Islands. Habitat: marine.

eurystomus, Asprocottus megalops Taliev [D. N.] 1955:195, 332, Figs. 70, 138-139 [ ref. 4316] Lake Baikal, Russia. Neotype: ZIN 46624. Genus Cyphocottus was not available from 2001, two included species after 1930 with no designation of a type species (Art. 13.3); dates to Sideleva 2003:193 [ref. 27815]. Neotype designation also does not conform to code requirements. •Valid as Limnocottus eurystomus (Taliev 1955) -- (Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:157 [ref. 23595]). •Valid as Cyphocottus eurystomus (Taliev 1955) -- (Sideleva 2001:67 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:18, 193 [ref. 27815], Bogutskaya & Naseka 2004:198 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Cyphocottus eurystomus (Taliev 1955). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

evermanni, Cottus Gilbert [C. H.] 1898:11, Fig. [Bulletin of the U. S. Fish Commission v. 17 (art. 1) [1897]; ref. 12349] Lost River near Lostine, Oregon, U.S.A. Holotype (unique): USNM 48228. •Synonym of Cottus princeps Gilbert 1898 -- (Lee et al. 1980:820 [ref. 22416]). Current status: Synonym of Cottus princeps Gilbert 1898. Cottidae. Habitat: freshwater.

extensus, Cottus Bailey [R. M.] & Bond [C. E.] 1963:8, Figs. 1b, 2b, 4b [Occasional Papers of the Museum of Zoology University of Michigan No. 634; ref. 3934] Along eastern shore of Bear Lake, south of the South Eden Delta, Rich County, Utah, U.S.A. Holotype: UMMZ 141840. Paratypes: UMMZ 141841 (15). Additional material at UMMZ. •Valid as Cottus extensus Bailey & Bond 1963 -- (Lee et al. 1980:811 [ref. 22416], Page & Burr 1991:249 [ref. 18983], Sigler & Sigler 1996:235 [ref. 25943], Fuller et al. 1999:341 [ref. 25838], Broughton 2000:653 [ref. 24593], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:9 [ref. 30399], Page & Burr 2011:474 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus extensus Bailey & Bond 1963. Cottidae. Distribution: Bear Lake, Idaho/ Utah, introductions elsewhere, U.S.A. Habitat: freshwater.

fabricii, Cottus Girard [C. F.] 1850:411 [Proceedings of AAAS (The American Association for the Advancement of Science) 2nd meeting, 1849; ref. 18123] Greenland. Based on Cottus gobio of Fabricius 1780. See also Girard 1850:189 [ref. 1814] and Girard 1851:59 [ref. 18124]. •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Neyelov 1973:595 [ref. 7219] as Gymnacanthus). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Cottidae. Habitat: marine.

fasciatus, Ainocottus Pavlenko [M. N.] 1910:30 [Trudy Obshchestva Estestvoispytatelej Imperatorskom' Kazanskom' Universiteti, Kazan v. 42 (no. 2); ref. 3393] Peter the Great Bay, Russia, Sea of Japan. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Neyelov 1979:122 [ref. 3152], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Cottidae. Habitat: marine.

fasciatus, Trachidermus Heckel [J. J.] 1839:160, Pl. 9 (fig. 2) [Annalen des Wiener Museums der Naturgeschichte v. 2 (no. 1); ref. 2079] Philippines [=? China]. Holotype (unique): NMW 92826. Spelled efasciatus in main heading, otherwise as fasciatus. •Valid as Trachidermus fasciatus Heckel 1839 -- (Watanabe 1960:114 [ref. 11959], Okada 1961:709 [ref. 6363], Sideleva 1982:82 [ref. 14469], Yabe in Masuda et al. 1984:326 [ref. 6441], Kawanabe & Mizuno 1989:654 [ref. 25598], Kim & Youn 1992:62 [ref. 22287], Takita & Chikamoto 1994:123 [ref. 22342], Zhu 1995:188 [ref. 25213], Cheng & Zhou 1997:437 [ref. 26385] dated 1840, Kim 1997:408 [ref. 24032], Yue & Chen 1998:240 [ref. 26415], Onikura et al. 1999:31 [ref. 23924], Nakabo 2000:637 [ref. 25086], Wang et al. 2001:300 [ref. 26566] dated 1840, Kim & Park 2002:328 [ref. 25913], Nakabo 2002:637 [ref. 26001], Youn 2002:263, 573 [ref. 26218], Goto et al. 2014:[6] [ref. 33456], Zhang et al. 2016:194 [ref. 34477]). Current status: Valid as Trachidermus fasciatus Heckel 1839. Cottidae. Distribution: Western North Pacific, including China, Korea and Japan. Habitat: freshwater, brackish, marine.

fenestralis, Artedius Jordan [D. S.] & Gilbert [C. H.] 1883:577 [Proceedings of the United States National Museum v. 5 (no. 326); ref. 10614] Commencement Bay near New Tacoma, Washington, U.S.A. Syntypes: USNM 27146 (7), 27206 (2). •Valid as Artedius fenestralis Jordan & Gilbert 1883 -- (Bolin 1944:48 [ref. 6379], Eschmeyer & Herald 1983:162 [ref. 9277], Begle 1989:646 [ref. 12739], Matarese et al. 1989:414 [ref. 26885], McAllister 1990:194 [ref. 14674], Mecklenburg et al. 2002:460 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:164 [ref. 35888]). Current status: Valid as Artedius fenestralis Jordan & Gilbert 1883. Cottidae. Distribution: Eastern North Pacific: eastern Aleutian Islands to southern California. Habitat: marine.

ferrugineus, Cottus Heckel [J. J.] & Kner [R.] 1857:34, Figs. 14-15 [Die Süsswasserfische der Österreichischen Monarchie; ref. 2078] Garda Lake, Mailand, Treviso, Italy and Xegar, Dalmatia [Croatia]. Lectotype: NMW 6419. Paralectotypes: NMW 6036 (4), 6419 (? now 4), 6562-63 (2, 3), 6673 (6), 78799 (4). Appeared first as a nomen nudum in Bonaparte 1846:62. •See Lelek 1987:274 [ref. 22962]. •Synonym of Cottus gobio Linnaeus 1758 -- (Berg 1949:1145 [ref. 20662], Neyelov 1973:593 [ref. 7219], Freyhof et al. 2005:132 [ref. 28259], Bianco & Delmastro 2011:6 [ref. 31648] Bianco 2013:10 [ref. 32897]). •Valid as Cottus ferrugineus Heckel & Kner 1857 -- (Kottelat 1997:169 [ref. 22952], Sideleva et al. 2015:33 [ref. 35276]). Current status: Valid as Cottus ferrugineus Heckel & Kner 1857. Cottidae. Distribution: Southern Europe. Habitat: freshwater.

filamentosus, Cottus Sauvage [H.-E.] in Vaillant & Sauvage 1875:279 [Revue et Magasin de Zoologie (Sér. 3) v. 3; ref. 17853] "Hawaiian Islands" [error]. Syntypes: MNHN 0000-9090 (2). Type catalog: Blanc & Hureau 1968:51 [ref. 20738]. •Apparently a synonym of Gymnocanthus intermedius (Temminck & Schlegel 1843) -- (see Mundy 2005:336 [ref. 28379]). Current status: Synonym of Gymnocanthus intermedius (Temminck & Schlegel 1843). Cottidae. Habitat: marine.

filamentosus, Icelinus Gilbert [C. H.] 1890:85 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., Albatross station 2893 and 2959, depths 145 and 55 fathoms. Syntypes: CAS-SU 118 (1), 28109 (1); USNM 44407 (1), 48245 (2), 125011 [ex USBF 449] (2). Type catalog: Böhlke 1953:130 [ref. 12291]. Lectotype established (as figured specimen) in Jordan 1896:244 [ref. 2395] if specimen is identifiable. •Valid as Icelinus filamentosus Gilbert 1890 -- (Bolin 1944:24 [ref. 6379], Eschmeyer & Herald 1983:173 [ref. 9277], Peden 1984:79 [ref. 5193], McAllister 1990:198 [ref. 14674], Mecklenburg et al. 2002:439 [ref. 25968], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:176 [ref. 35888]). Current status: Valid as Icelinus filamentosus Gilbert 1890. Cottidae. Distribution: Eastern North Pacific: western Gulf of Alaska to southern California. Habitat: marine.

fimbriatus, Icelinus Gilbert [C. H.] 1890:87 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., Albatross station 2893 and 2975, depths 45 and 36 fathoms. Syntypes: USNM 43087 (1), 48246 (1). •Valid as Icelinus fimbriatus Gilbert 1890 -- (Bolin 1944:32 [ref. 6379], Eschmeyer & Herald 1983:173 [ref. 9277], Peden 1984:67 [ref. 5193], McAllister 1990:198 [ref. 14674], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:176 [ref. 35888]). Current status: Valid as Icelinus fimbriatus Gilbert 1890. Cottidae. Distribution: Eastern North Pacific: southern British Columbia to southern Baja California. Habitat: marine.

forficata, Elanura Gilbert [C. H.] 1896:430, Pl. 30 (top) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] South of Sannak Island, Aleutian Islands, Albatross station 3214, depth 70 meters. Lectotype: MCZ 28266. Paralectotypes: BMNH 1896.7.23.149 (1); NMW 11468 (2); CAS-SU 2420 (3); USNM 48673 (1), 59404 (1), 119654 (1); ZMUO J1442 (1) station 3214. Type catalog: Böhlke 1953:129 [ref. 12291], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen from Albatross sta. 3214) in caption to Pl. 289, p. 3291 in Jordan & Evermann 1900 [ref. 2446]; this is the specimen figured with the original description and is apparently MCZ 28266. This specimen was also designated lectotype by Pietsch 1994:354 [ref. 21189]. •Valid as Triglops forficatus (Gilbert 1896) -- (Pietsch 1994:354 [ref. 21189] as forficata, Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:424 [ref. 25968], Parin et al. 2002:S87 [ref. 28300], Fedorov et al. 2003:81 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:275 [ref. 33547], Kells et al. 2016:192 [ref. 35888]). Current status: Valid as Triglops forficatus (Gilbert 1896). Cottidae. Distribution: North Pacific: Bering Sea to northwestern Gulf of Alaska and to Kuril Islands; one record from southeastern Alaska. Habitat: marine.

formosus, Cottus Girard [C. F.] 1851:58 [Smithsonian Contributions to Knowledge v. 3 (art. 3) (for 1850); ref. 18124] Lake Ontario off Oswego, New York, U.S.A. Holotype (unique): USNM. •Synonym of Cottus cognatus Richardson 1837 -- (Smith 1986:358 [ref. 22186], Scott & Crossman 1998:835 [ref. 24570]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

franklini, Cottus Agassiz [L.] 1850:303 [Lake Superior; ref. 66] East and south shores of Lake Superior, North America. Syntypes: ?MCZ 35977 [ex USNM] (2). •Synonym of Cottus cognatus Richardson 1837 -- (Scott & Crossman 1998:835 [ref. 24570]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

frykenensis, Cottus quadricornis Lönnberg [E.] 1932:14, Pl. (fig. 5) [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Fryken, Sweden. Syntypes: NRM 30315 (1), Mus. of Karlstad's school (1), Uppsala Univ. Zool. Mus. (1). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

fujiyamae, Rheopresbe Jordan [D. S.] & Starks [E. C.] 1904:270, Fig. 16 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Odowara, Sagami Bay, Japan (from English label [Japanese label states "locality unknown"]). Holotype (unique): CAS-SU 7706. Type catalog: Böhlke 1953:132 [ref. 12291]. •Synonym of Cottus kazika Jordan & Starks 1904 -- (Watanabe 1960:117 [ref. 11959], Okada 1961:696 [ref. 6363], Goto et al. 2014:[9] [ref. 33456]). Current status: Synonym of Cottus kazika Jordan & Starks 1904. Cottidae. Habitat: freshwater, brackish, marine.

furciger, Icelus Malm [A. W.] 1865:410 [Förhandlingar vid de Skandinaviske Naturforskarnes. v. 9; ref. 17596] Löken, Gullmarfjärden, Sweden. No types known. •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neyelov 1973:596 [ref. 7219], Nelson 1984:36 [ref. 5391]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Cottidae. Habitat: marine.

fuscescens, Icelinus Gilbert [C. H.] 1915:340, Pl. 17 (fig. 9) [Proceedings of the United States National Museum v. 48 (no. 2075); ref. 1632] Off Santa Barbara Island, California, U.S.A., Albatross station 4418, depth 260-310 fathoms. Holotype: USNM 75812. Paratypes: CAS-SU 22981 (1), 22985 (1), 22991 (1), 23001 (6). Type catalog: Böhlke 1953:130 [ref. 12291]. •Synonym of Icelinus burchami Evermann & Goldsborough 1907 -- (Bolin 1944:30 [ref. 6379], Mecklenburg et al. 2002:440 [ref. 25968]). •Synonym of Icelinus burchami Evermann & Goldsborough 1907, but a valid subspecies -- (Peden 1984:79 [ref. 5193]). Current status: Synonym of Icelinus burchami Evermann & Goldsborough 1907. Cottidae. Habitat: marine.

fuscimentus, Artediellus (Artediellops) Nelson [D. W.] 1986:41, Fig. 10 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 138 (no. 1); ref. 5808] Southern Sea of Japan, 36°34'N, 132°50'20"E, Albatross station 4844, depth 202 meters. Holotype: USNM 74464. Paratypes: CAS-SU 22230 (24, 1 c&s); USNM 74491 (5), 74493-95 (1, 11, 3), 74497-98 (1, 1). •Valid as Artediellus fuscimentus Nelson 1986 -- (Nakabo 2000:642 [ref. 25086], Nakabo 2002:642, 1526 [ref. 26001], Shinohara et al. 2011:43 [ref. 31715], Shinohara et al. 2014:248 [ref. 33330], Shin et al. 2016:209 [ref. 34833]). Current status: Valid as Artediellus fuscimentus Nelson 1986. Cottidae. Distribution: Western North Pacific. Habitat: marine.

galatheae, Antipodocottus Bolin [R. L.] 1952:432, Fig. 1 [Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, Kjøbenhavn. v. 114; ref. 511] Tasman Sea off New Zealand, 42°10'S, 170°10'E, Galathea station 626, depth 594 [or 610] meters. Holotype: ZMUC P811234. Paratypes: CAS-SU 17360 (1), ZMUC P811235-36 (2). Type catalog: Böhlke 1953:126 [ref. 12291], Nielsen 1974:81 [ref. 9588]. •Valid as Antipodocottus galatheae Bolin 1952 -- (Nelson 1975:80 [ref. 30370], Fricke & Brunken 1984:46 [ref. 5262], Paxton et al. 1989:477 [ref. 12442], Paulin et al. 1989:172 [ref. 24556], Paxton et al. 2006:952 [ref. 28994], Stewart 2015:1119 [ref. 34254], Girard & Smith 2016:556 [ref. 34786]). Current status: Valid as Antipodocottus galatheae Bolin 1952. Cottidae. Distribution: Southwestern Pacific: Australia and New Zealand. Habitat: marine.

galeatus, Gymnacanthus Bean [T. H.] 1881:153 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Iliuliuk, Unalaska Island, Aleutian Islands, Alaska, U.S.A. Lectotype: USNM 28097. Paralectotypes: USNM 28097 (2 of 3, orig. 5). Original genus should have been spelled Gymnocanthus. Lectotype established (as figured specimen) in caption to Pl. 301, p. 3293 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. •Valid as Gymnocanthus galeatus Bean 1881 -- (Watanabe 1960:59 [ref. 11959], Eschmeyer & Herald 1983:169 [ref. 9277], Yabe in Masuda et al. 1984:328 [ref. 6441], McAllister 1990:197 [ref. 14674], Nakabo 2000:629 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Mecklenburg et al. 2002:466 [ref. 25968], Nakabo 2002:629 [ref. 26001], Fedorov et al. 2003:69 [ref. 27579] as Gymnacanthus, Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:247 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Mecklenburg et al. 2016:57 [ref. 34440], Kells et al. 2016:172 [ref. 35888], Balanov & Matveev 2018:367 [431] [ref. 36463], Mecklenburg et al. 2018:196 [ref. 36370]). Current status: Valid as Gymnocanthus galeatus Bean 1881. Cottidae. Distribution: North Pacific: Bering Sea south to British Columbia (Canada), Sea of Okhotsk and to northern Japan Sea off Hokkaido; also adjacent Arctic Region. Habitat: marine.

gibbosus, Abyssocottus Berg [L. S.] 1906:908 [Zoologischer Anzeiger v. 30 (no. 26); ref. 265] Lake Baikal, opposite Baikalskoye village, Russia. Holotype (unique): ZIN 13737. •Valid as Asprocottus gibbosus (Berg 1906) -- (Taliev 1955:336 [ref. 4316]). •Valid as Abyssocottus gibbosus Berg 1906 -- (Berg 1949:1157 [ref. 20662], Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:155 [ref. 23595], Bogutskaya et al. 2001:47 [ref. 26178], Sideleva 2001:60 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:17, 178 [ref. 27815], Bogutskaya & Naseka 2004:194 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Abyssocottus gibbosus Berg 1906. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

gibbsii, Hemilepidotus Gill [T. N.] 1862:13 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 14; ref. 1654] San Francisco, California, U.S.A., and other areas. •Synonym of Hemilepidotus hemilepidotus (Tilesius 1811) -- (Bolin 1944:17 [ref. 6379]). Current status: Synonym of Hemilepidotus hemilepidotus (Tilesius 1811). Cottidae. Habitat: marine.

gigas, Cottus Kner [R.] 1864:186 [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Classe v. 1 (no. 24); ref. 2637] Cottidae.

gilberti, Hemilepidotus Jordan [D. S.] & Starks [E. C.] 1904:254, Fig. 10 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate,Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7446. Paratypes: USNM 50916 (1). Additional material or paratypes: AMS I.6950 (1), ?BMNH 1923.2.26.484 (1), CAS-SU 7573 (2). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Hemilepidotus gilberti Jordan & Starks 1904 -- (Watanabe 1960:48 [ref. 11959], Peden 1979:36 [ref. 24080], Yabe in Masuda et al. 1984:323 [ref. 6441], Matarese et al. 1989:372 [ref. 26885], Shinohara et al. 1996:176 [ref. 26605] as jordani but corrected to gilberti with errata insert, Lindberg & Krasyukova 1987:181 [ref. 15964], Kim & Youn 1992:68 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:631 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:433 [ref. 25968], Nakabo 2002:631 [ref. 26001], Youn 2002:262, 575 [ref. 26218], Parin et al. 2002:S75 [ref. 28300], Tokranov et al. 2003:299 [ref. 26972], Fedorov et al. 2003:70 [ref. 27579], Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:250 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331]). Current status: Valid as Hemilepidotus gilberti Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

gilberti, Icelus Taranetz [A. Ya.] 1936:149, Fig. [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (pt 13) (No. 3, art. 107); ref. 6509] Sea of Japan (Vladmir and Olga Bays and Yuzhnyy Cape) and Okhotsk Sea (Aniva Cape), depth 56-180 meters. Syntypes: (many) ZIN 25514 (4), 25517 (1), 25328 (3). •Valid as Icelus gilberti Taranetz 1936 -- (Nelson 1984:21 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:192 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:636 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Nakabo 2002:636 [ref. 26001], Parin et al. 2002:S77 [ref. 28300], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:254 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331]). Current status: Valid as Icelus gilberti Taranetz 1936. Cottidae. Distribution: Western North Pacific. Habitat: marine.

gilberti, Oncocottus hexacornis Fowler [H. W.] 1905:366, Figs. 2-5 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 57; ref. 15353] Point Barrow, northern Alaska, U.S.A. Holotype: ANSP not found [ex WIAP 7203]. Paratypes: WIAP (1). Type catalog: Böhlke 1984:65 [ref. 13621] as types not found, specimens perhaps never deposited at ANSP. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies hexacornis (Richardson 1823) -- (Berg 1949:1129 [ref. 20662]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

gilli, Synchirus Bean [T. H.] 1890:642 [Proceedings of the United States National Museum v. 12 (no. 787); ref. 228] Barkley Sound [Barclay Sound], west coast of Vancouver Island, British Columbia, Canada. Syntypes: USNM 41820 (now 4). •Valid as Synchirus gilli Bean 1890 -- (Eschmeyer & Herald 1983:183 [ref. 9277], Matarese et al. 1989:410 [ref. 26885], McAllister 1990:201 [ref. 14674], Mecklenburg et al. 2002:418 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:125 [ref. 32708], Pietsch & Orr 2015:45 [ref. 34694], Kells et al. 2016:192 [ref. 35888]). Current status: Valid as Synchirus gilli Bean 1890. Cottidae. Distribution: Eastern North Pacific: Unalaska Island and Gulf of Alaska to southern California (U.S.A.). Habitat: marine.

girardi, Cottus Robins [C. R.] 1961:306, Figs. 1a, d [Copeia 1961 (no. 3); ref. 9373] Conochocheague Creek, 9.6 miles northeast of Chambersburg on rt. 11, Franklin County, Pennsylvania, U.S.A. Holotype: CUMV 25607. Paratypes: CUMV 1680 (13), 7363 (10), 25605 (1); UMMZ 167174 (4). Additional material: CUMV (15 lots, 16). •Valid as Cottus girardi Robins 1961 -- (Lee et al. 1980:812 [ref. 22416], Page & Burr 1991:246 [ref. 18983], Jenkins & Burkhead 1994:670 [ref. 21581], Stauffer et al. 1995:255 [ref. 23376], Nelson et al. 2004:119 [ref. 27807], Robins 2005:4 [ref. 28230], Scharpf 2008:9 [ref. 30399], Page & Burr 2011:471 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus girardi Robins 1961. Cottidae. Distribution: Eastern U.S.A. Habitat: freshwater.

glaber, Centridermichthys Steindachner [F.] 1881:219 [41], Pl. 6 (fig. 2) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 83 (1. Abth.); ref. 4231] Appeared as "Fig. 2. Centridermichthys glaber n. sp." in figure captions on p. 41; this corresponds to Centridermichthys elongatus on p. 186 [8]. Regarded as two names for the same taxon; first reviser not researched. •Synonym of Alcichthys elongatus (Steindachner 1881) -- (Parin et al. 2002:S68 [ref. 28300], Parin et al. 2014:233 [ref. 33547]). Current status: Synonym of Alcichthys elongatus (Steindachner 1881). Cottidae. Habitat: marine.

glacialis, Cottus Richardson [J.] 1855:349, Pl. 24 [The last of the Arctic voyages in search of Sir J. Franklin; ref. 18631] Northumberland Sound, 76°53'N, western Greenland, depth 9 fathoms. No types known. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Cottidae. Habitat: brackish, marine.

globiceps, Oligocottus Girard [C. F.] 1858:58 [General report upon zoology of the several Pacific railroad routes, 1857; ref. 4911] Southern Farallones Island, off San Francisco, California, U.S.A. Syntypes: USNM 300 (orig. 4, now 2) Also appeared in Girard 1858:202 [ref. 1811]. •Valid as Clinocottus globiceps (Girard 1858) -- (Bolin 1944:81 [ref. 6379], Eschmeyer & Herald 1983:166 [ref. 9277], Matarese et al. 1989:426 [ref. 26885], McAllister 1990:196 [ref. 14674], Mecklenburg et al. 2002:504 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:39 [ref. 34694], Kells et al. 2016:168 [ref. 35888]). Current status: Valid as Clinocottus globiceps (Girard 1858). Cottidae. Distribution: Eastern North Pacific: western Gulf of Alaska to southern California (U.S.A). Habitat: marine.

gobio, Cottus Linnaeus [C.] 1758:265 [Systema Naturae, Ed. X v. 1; ref. 2787] Stream Freudenbach at Lockstädt, 53°12'N, 12°01'E, Lower Elbe River drainage, Germany. Neotype: NRM 51803. Neotype designated by Fricke 1999:38 [ref. 24101], but withdrawn in Fricke 2000:639 [ref. 24537]. For a discussion of the types and neotype designation see Freyhof et al. 2005:129 [ref. 28259]. •Valid as Cottus gobio Linnaeus 1758 -- (Berg 1949:1145 [ref. 20662], Neyelov 1973:593 [ref. 7219], Sideleva 1982:81 [ref. 14469], Fedorov in Whitehead et al. 1986:1247 [ref. 13677], Lelek 1987:274 [ref. 22962], Mrakovcic et al. 1995:184 [ref. 23079], Chereshnev 1996:606 [ref. 24659], Kottelat 1997:169 [ref. 22952], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:151 [ref. 23595], Fricke 1999:38 [ref. 24101], Bogutskaya et al. 2001:46 [ref. 26178], Parin et al. 2002:S73 [ref. 28300], Hanel 2003:61 [ref. 27894], Bogutskaya & Naseka 2004:186 [ref. 28183], Freyhof et al. 2005:124 [ref. 28259], Hanel & Lusk 2005:337 [ref. 28804], Fricke 2007:26 [ref. 30577], Shedko & Miroshnichenko 2007:24 [ref. 29369], Kottelat & Freyhof 2007:508 [ref. 29996], Sideleva 2009:210 [ref. 30173], Bianco 2013:10 [ref. 32897], Goto et al. 2014:[10] [ref. 33456], Parin et al. 2014:243 [ref. 33547], Sideleva et al. 2015:430 [ref. 33770], Sideleva et al. 2018:8 [ref. 36393], Ćaleta et al. 2019:186 [ref. 37026]). Current status: Valid as Cottus gobio Linnaeus 1758. Cottidae. Distribution: Europe. Habitat: freshwater, brackish.

gobioides, Cottus Girard [C. F.] 1850:411 [Proceedings of AAAS (The American Association for the Advancement of Science) 2nd meeting, 1849; ref. 18123] Burlington, Vermont, U.S.A. See also Girard 1850:189 [ref. 1814] and Girard 1851:55, Pl. 2 (figs. 3-4) [ref. 18124]. •Synonym of Cottus cognatus Richardson 1837. Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

godlewskii, Cottus Dybowski [B. N.] 1874:385, Pl. 4 (fig. 2) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 24 (no. 3-4); ref. 1172] Lake Baikal, Russia, depth 100-300 meters. Syntypes: not found at ZIN. •Valid as Abyssocottus godlewskii (Dybowski 1874) -- (Taliev 1955:351 [ref. 4316]). •Valid as Limnocottus godlewskii (Dybowski 1874) -- (Berg 1949:1160 [ref. 20662], Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:157 [ref. 23595], Sideleva 2001:68 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:18, 194 [ref. 27815], Bogutskaya & Naseka 2004:198 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Limnocottus godlewskii (Dybowski 1874). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

gomojunovi, Artediellus Taranetz [A. Ya.] 1933:71 [Bulletin of the Far Eastern Branch of the Academy of Sciences U.S.S.R. 1933 (nos 1-2-3); ref. 15743] Gulf of Anadyr and Olytorski, northwestern Bering Sea. Lectotype: ZIN 25337 (male). Paralectotypes: (4+) ZIN 25337 (female). Lectotype selected by Neyelov 1979:155 [ref. 3152]. •Valid as Artediellus gomojunovi Taranetz 1933 -- (Neyelov 1979:155 [ref. 3152], Nelson 1986:43 [ref. 5808], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:490 [ref. 25968], Parin et al. 2002:S70 [ref. 28300], Nelson et al. 2004:118 [ref. 27807], Mecklenburg et al. 2011:122 [ref. 31212] with comments, Page et al. 2013:121 [ref. 32708], Parin et al. 2014:237 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033]). Current status: Valid as Artediellus gomojunovi Taranetz 1933. Cottidae. Distribution: North Pacific: Bering Sea off Alaska to Kuril Islands. Habitat: marine.

gonez, Cottiusculus Jordan [D. S.] & Starks [E. C.] (ex Schmidt) 1904:298, Fig. 29 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Aniva Bay, Sakhalin Island, Russia. Lectotype: CAS-SU 7713. Paralectotypes: CAS-SU 69887 [ex CAS-SU 7713 [ex ZIN]] (4). Type catalog: Böhlke 1953:127 [ref. 12291]. Appeared without distinguishing features in Schmidt 1903:518 [p. 16 of separate] [ref. 3945]. Apparently appeared first in an available way as above, then in Schmidt 1904:108, Pl. 11 [ref. 3946]. Types are specimens used by Jordan and not by Schmidt; Schmidt's specimens are ZISP 12736-37 (20, 6), 12739-43 (1, 2, 8, 2, 4). Lectotype selected by Kai & Nakabo 2009:223 [ref. 30299]. •Valid as Cottiusculus gonez Jordan & Starks 1904 [sometimes with author as Schmidt] -- (Watanabe 1960:170 [ref. 11959], Neyelov 1979:162 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:238 [ref. 15964], Kim & Youn 1992:71 [ref. 22287], Cheng & Zhou 1997:436 [ref. 26385] with author as Schmidt, Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:643 [ref. 25086], Nakabo 2002:643 [ref. 26001], Youn 2002:262, 573 [ref. 26218], Parin et al. 2002:S72 [ref. 28300], Fedorov et al. 2003:67 [ref. 27579] as Schmidt in Jordan & Starks 1904, Kai & Nakabo 2009:222 [ref. 30299], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:240 [ref. 33547]). Current status: Valid as Cottiusculus gonez Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

gotoi, Procottus Sideleva [V. G.] 2001:69, Fig. 2 [Proceedings of the Zoological Institute, Academy of Sciences, St.-Petersburg No. 287; ref. 26181] Lake Baikal, near Bolshiye Koty Village, depth 10 meters. Holotype: ZIN 52803. Paratypes: ZIN 52084-85 (1, 2). •Valid as Procottus gotoi Sideleva 2001 -- (Bogutskaya et al. 2001:47 [ref. 26178], Sideleva 2003:19, 204 [ref. 27815], Bogutskaya & Naseka 2004:200 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Procottus gotoi Sideleva 2001. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

gracilis, Cottus Heckel [J. J.] 1839:148 [Annalen des Wiener Museums der Naturgeschichte v. 2 (no. 1); ref. 2079] New York, U.S.A. Holotype (unique): NMW 6422. •Synonym of Cottus cognatus Richardson 1837 -- (Smith 1986:358 [ref. 22186], Scott & Crossman 1998:835 [ref. 24570], Scharpf 2008:9 [ref. 30399]). •Synonym of Cottus cognatus Richardson 1837, but a valid subspecies -- (Chereshnev 1996:606 [ref. 24659], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:151 [ref. 23595]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

gratzianowi, Cottus Sideleva [V. G.], Naseka [A. M.] & Zhidkov [Z. V.] 2015:420, Figs. 2-3, 5a-6a [Zootaxa 3949 (no. 3); ref. 33770] Ukhtomitsa River village of Korotetskaya (Vologda Province), 60°18′19″N, 38°40′30″E, Onega River drainage, White Sea basin, Arctic basin. Holotype: ZIN 55596. Paratypes: ZIN 55597. •Valid as Cottus gratzianowi Sideleva, Naseka & Zhidkov 2015. Current status: Valid as Cottus gratzianowi Sideleva, Naseka & Zhidkov 2015. Cottidae. Distribution: Onega River drainage, White Sea basin, Arctic basin. Habitat: freshwater.

greenei, Uranidea Gilbert [C. H.] & Culver [G. B.] in Jordan & Evermann 1898:1965 [Bulletin of the United States National Museum No. 47; ref. 2444] Thousand Springs, Snake River, near mouth of Salmon Fall River, Gooding County, Idaho, U.S.A. Holotype: CAS-SU uncat. Paratypes: CAS-SU uncat. (incl. the type, marked drawn). Type series found by D. Neely 1/2008 at CAS. •Valid as Cottus greenei (Gilbert & Culver 1898) -- (Lee et al. 1980:813 [ref. 22416], Page & Burr 1991:248 [ref. 18983], Fuller et al. 1999:341 [ref. 25838], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:9 [ref. 30399], Page & Burr 2011:473 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus greenei (Gilbert & Culver 1898). Cottidae. Distribution: Snake River basin, Idaho,U.S.A. Habitat: freshwater.

grewingkii, Cottus Dybowski [B. N.] 1874:384, Pl. 1 (fig. 1) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 24 (no. 3-4); ref. 1172] Lake Baikal, Russia. Lectotype (?): BMNH 1897.7.5.4 [ex ZIN]. Paralectotypes: ZMB 7808 (8). Type catalog: Paepke & Fricke 1992:285 [ref. 19946]. Lectotype selected by Dybowski 1908:559 [ref. 17422]. •Valid as Cottocomephorus grewingkii [sometimes as gerwingki or grewingki] (Dybowski 1874) -- (Berg 1949:1171 [ref. 20662], Taliev 1955:275 [ref. 4316], Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:150 [ref. 23595], Bogutskaya et al. 2001:46 [ref. 26178], Sideleva 2001:58 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:15, 163 [ref. 27815], Bogutskaya & Naseka 2004:192 [ref. 28183], Goto et al. 2014:[15] [ref. 33456], Bogdanov 2019:231 [ref. 36814]). Current status: Valid as Cottocomephorus grewingkii (Dybowski 1874). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

griseus, Abyssocottus godlewskii Taliev [D. N.] 1955:355, Figs. 20, 150-151 [ ref. 4316] Lake Baikal, Russia, depth 16-180 to 700 meters. Syntypes: none found at ZIN. Appeared without distinguishing features and not available in Taliev 1948:113 [ref. 19628]. Neotype [ZIN 50804] apparently designated by Sideleva but not in conformance with the fourth edition of the Code. •Valid as Limnocottus griseus (Taliev 1955) -- (Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702] dating to 1948, Sideleva 1998:157 [ref. 23595], Bogutskaya et al. 2001:47 [ref. 26178] dating to 1948, Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:18, 197 [ref. 27815] dated to 1948, Bogutskaya & Naseka 2004:199 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Limnocottus griseus (Taliev 1955). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

groenlandicus, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:185 [Histoire naturelle des poissons v. 4; ref. 998] Greenland. No types known. Type catalog: Blanc & Hureau 1968:60 [ref. 20738]. Based on Cottus scorpius of Fabricius. •Synonym of Myoxocephalus scorpius (Linnaeus 1758), but a valid subspecies -- (Neyelov 1973:599 [ref. 7219], Neyelov 1979:122 [ref. 3152], Fedorov in Whitehead et al. 1986:1255 [ref. 13677], Mecklenburg et al. 2007:176 [ref. 29448]). •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Scott & Scott 1988:509 [ref. 25518], Yabe in Okamura et al. 1995:165 [ref. 22531], Mecklenburg et al. 2002:478 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Mecklenburg et al. 2011:123 [ref. 31212], Parin et al. 2014:265 [ref. 33547], Mecklenburg et al. 2016:85 [ref. 34440]). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Cottidae. Habitat: brackish, marine.

groenlandicus, Gymnacanthus tricuspis Schmidt [P. Yu.] 1927:28 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 15821] Arctic. Syntypes: ZMUC (2), ZIN 1930-31 (1, 1). •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Neyelov 1973:595 [ref. 7219] as Gymnacanthus, Mecklenburg et al. 2016:62 [ref. 34440]). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Cottidae. Habitat: marine.

gruvintii, Centridermichthys Sauvage [H.-E.] (ex Dybowski) 1878:146 [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris (Série 2) v. 1; ref. 3880] Lake Baikal, Russia. Apparently a misspelling of Cottus grewingkii Dybowsky 1874. Current status: Synonym of Cottocomephorus grewingkii (Dybowski 1874). Cottidae. Distribution: Lake Baikal, Russia.

gulosus, Cottopsis Girard [C. F.] 1854:129 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] San Joaquin River, California, U.S.A. Syntypes: USNM 291 (2). •Valid as Cottus gulosus (Girard 1854) -- (Lee et al. 1980:814 [ref. 22416], Page & Burr 1991:252 [ref. 18983], Mecklenburg et al. 2002:404 [ref. 25968], Moyle 2002:350 [ref. 26449], Wydoski & Whitney 2003:210 [ref. 27816], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:9 [ref. 30399], Page & Burr 2011:481 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[9] [ref. 33456]). Current status: Valid as Cottus gulosus (Girard 1854). Cottidae. Distribution: Western U.S.A.: Washington, Oregon and California. Habitat: freshwater.

gurwicii, Metacottus Taliev [D. N.] 1946:90, Figs. 1-5 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 54 (no. 1); ref. 4315] Lake Baikal at Maritui station, Russia, depth 93 meters. Holotype (unique): ZIN 46660. Sideleva 2001:69 [ref. 26181] and 2003:205 [ref. 27815] regards the holotype as the neotype which does not conform to the code. •Valid as Metacottus gurwicii Taliev 1946 -- (Berg 1949:1170 [ref. 20662], Taliev 1955:300 [ref. 4316]). •Valid as Procottus gurwicii (Taliev 1946) -- (Sideleva 1982:31 [ref. 14469] as gurwici, Reshetnikov et al. 1997:761 [ref. 24702], Sideleva 1998:158 [ref. 23595] as gurwici, Sideleva 2001:72 [ref. 26181] as gurwici, Bogutskaya et al. 2001:47 [ref. 26178] as gurwici, Sideleva 2003:19, 206 [ref. 27815] as gurwici, Bogutskaya & Naseka 2004:200 [ref. 28183], Goto et al. 2014:[15] [ref. 33456] as gurwicii). Current status: Valid as Procottus gurwicii (Taliev 1946). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

gussinensis, Paracottus kessleri Tarkhova [Yu. N.] 1962:108 [Kratkiie soobscenjia Burjatskij Kompleksnogo Naucno-Issledovatel'skogo Instituta, Akademija Nauk SSR, Sibirskoe Otdelenie, Burjatskij Kompleksnyi Nauno-Issledovatel'skij Institut. No. 3; ref. 22756] Gusinor Lake, Lake Baikal Basin, Russia. Lectotype: ZIN 52232. Paralectotypes: ZIN 52233 (1). Lectotype selected by Sideleva 2003:152 [ref. 27815]. •Synonym of Cottus kesslerii Dybowski 1874, but a valid subspecies -- (Sideleva 1982 [ref. 14469], Sideleva 1998:152 [ref. 23595]). •Synonym of Leocottus kesslerii (Dybowski 1874), but a valid subspecies gussinensis (Tarkhova 1962) as described -- (Sideleva 2001:60 [ref. 26181], Sideleva 2003:152 [ref. 27815] with author as Tarkhova). •Synonym of Leiocottus kesslerii (Dybowski 1874) -- (Kottelat 2006:61 [ref. 28831]). Current status: Synonym of Leocottus kesslerii (Dybowski 1874). Cottidae. Habitat: freshwater.

haemusi, Cottus gobio Marinov [B. T.] & Dikov [C. I.] 1986:19 [Acta Zoologica Bulgarica v. 31; ref. 8968] Upper Beli Vit River, above village Ribarica, Danube River basin, Bulgaria. Holotype: DHISUB 6039-1. Paratypes: DHISUB 6016-1 to 6038-1 (23), 6040-1 to 6045-1 (6). •Tentatively a synonym of Cottus gobio Linnaeus 1758 -- (Kottelat 1997:169 [ref. 22952]). •Synonym of Cottus gobio Linnaeus 1758, but a valid subspecies as described -- (Vassilev & Pehlivanov 2005:184 [ref. 28369]). •Valid as Cottus haemusi Marinov & Dikov 1986 -- (Freyhof et al. 2005:160 [ref. 28259], Kottelat & Freyhof 2007:510 [ref. 29996] Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus haemusi Marinov & Dikov 1986. Cottidae. Distribution: Bulgaria. Habitat: freshwater, brackish.

haitej, Cottus Dybowski [B. N.] 1869:949, Pl. 14 (fig. 2) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 19; ref. 1169] Onon and Ingoda rivers and their tributaries, Russia. Syntypes: (2) ?BMNH 1871.7.19.9 (1), ZMB 7106 (1). Type catalog: Paepke & Fricke 1992:282 [ref. 19946]. Original as haïtej, correction to haitej required. •Valid as Mesocottus haitej (Dybowski 1869) -- (Berg 1949:1138 [ref. 20662], Sideleva 1982:82 [ref. 14469], Zhu 1995:188 [ref. 25213], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:153 [ref. 23595], Pietsch et al. 2001:145 [ref. 25688], Bogutskaya et al. 2001:46 [ref. 26178], Parin et al. 2002:S80 [ref. 28300], Bogutskaya & Naseka 2004:190 [ref. 28183], Kottelat 2006:61 [ref. 28831], Ocock et al. 2006:58 [ref. 28933], Bogutskaya et al. 2008:352 [ref. 30085], Goto et al. 2014:[6] [ref. 33456], Parin et al. 2014:259 [ref. 33547], Zhang et al. 2016:194 [ref. 34477], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Valid as Mesocottus haitej (Dybowski 1869). Cottidae. Distribution: China, Mongolia and Russia. Habitat: freshwater, brackish.

hamatus, Icelus Krøyer [H. N.] 1845:253 [Naturhistorisk Tidsskrift (Kjøbenhavn) (n. s.) v. 1; ref. 2689] Belsund, Spitzbergen. Lectotype: ZMUC P811869. Paralectotypes: ?ZMUC 10-11 (1 of 2). Type catalog: Nielsen 1974:81 [ref. 9588]. Lectotype selected by Nelson 1984:35 [ref. 5391]. •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neyelov 1973:596 [ref. 7219], Nelson 1984:35 [ref. 5391], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:253 [ref. 33547]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Cottidae. Habitat: marine.

hangiongensis, Cottus Mori [T.] 1930:48 [10], Fig. [Journal of the Chosen Natural History Society No. 11; ref. 16505] Kai-nei, Korea. Holotype: whereabouts unknown. Additional specimens: (3). On p. 10 of separate. •Valid as Cottus hangiongensis Mori 1930 -- (Watanabe 1960:126 [ref. 11959], Okada 1961:704 [ref. 6363], Yabe in Masuda et al. 1984:326 [ref. 6441], Kawanabe & Mizuno 1989:658 [ref. 25598], Kim & Youn 1992:60 [ref. 22287], Zhu 1995:189 [ref. 25213], Kim 1997:404 [ref. 24032], Nakabo 2000:638 [ref. 25086], Matsuura et al. 2000:202 [ref. 26000], Pietsch et al. 2001:144 [ref. 25688], Bogutskaya et al. 2001:46 [ref. 26178], Kim & Park 2002:325 [ref. 25913], Nakabo 2002:638 [ref. 26001], Youn 2002:262, 572 [ref. 26218], Hagiwara 2003:37 [ref. 27188], Bogutskaya & Naseka 2004:187 [ref. 28183], Fujii et al. 2005:12 [ref. 28191], Sideleva & Goto 2009:600 [ref. 30591], Goto et al. 2014:[9] [ref. 33456], Parin et al. 2014:243 [ref. 33547], Zhang et al. 2016:193 [ref. 34477], Dyldin & Orlov 2017:63 [ref. 35138]). Current status: Valid as Cottus hangiongensis Mori 1930. Cottidae. Distribution: Western North Pacific. Habitat: freshwater, brackish, marine.

hankinsoni, Parartedius Hubbs [C. L.] 1926:4 [Occasional Papers of the Museum of Zoology University of Michigan No. 170; ref. 6069] Tide-pool, near the end of Point Loma, California, U.S.A. Holotype (unique): UMMZ 55001. •Valid as Artedius hankinsoni (Hubbs 1926) -- (Bolin 1944:57 [ref. 6379]). •Synonym of Artedius lateralis (Girard 1854) -- (Begle 1989:648 [ref. 12739]). Current status: Synonym of Artedius lateralis (Girard 1854). Cottidae. Habitat: marine.

harringtoni, Axyrias Starks [E. C.] 1896:554, Pl. 74 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 4195] Vicinity of Port Ludlow, Washington, U.S.A. Holotype (unique): CAS-SU 5047. Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Artedius harringtoni (Starks 1896) -- (Bolin 1944:45 [ref. 6379], Eschmeyer & Herald 1983:162 [ref. 9277], Begle 1989:646 [ref. 12739], Matarese et al. 1989:416 [ref. 26885], McAllister 1990:194 [ref. 14674], Mecklenburg et al. 2002:461 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:164 [ref. 35888]). Current status: Valid as Artedius harringtoni (Starks 1896). Cottidae. Distribution: Eastern North Pacific: eastern Aleutian Islands to southern California (U.S.A.). Habitat: marine.

hemilepidotus, Cottus Tilesius [W. G. von] 1811:262 [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 3; ref. 4408] Sea of Okhotsk and Bering Sea, Alaska, U.S.A. Syntypes: MNHN 0000-7155 (1) Kamchatka. •Valid as Hemilepidotus hemilepidotus (Tilesius 1811) -- (Bolin 1944:17 [ref. 6379], Peden 1979:24 [ref. 24080], Eschmeyer & Herald 1983:170 [ref. 9277], Matarese et al. 1989:374 [ref. 26885], McAllister 1990:197 [ref. 14674], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:435 [ref. 25968], Parin et al. 2002:S75 [ref. 28300], Tokranov et al. 2003:300 [ref. 26972], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:250 [ref. 33547], Pietsch & Orr 2015:40 [ref. 34694], Kells et al. 2016:172 [ref. 35888]). Current status: Valid as Hemilepidotus hemilepidotus (Tilesius 1811). Cottidae. Distribution: North Pacific: southern Bering Sea and Aleutian and Commander Islands to central California (U.S.A.) and to southeastern Kamchatka. Habitat: marine.

herzensteini, Asprocottus Berg [L. S.] 1906:907 [Zoologischer Anzeiger v. 30 (no. 26); ref. 265] Lake Baikal, Russia, depth 45-1800 meters. Holotype: ZIN 13760. Sideleva 2001:61 [ref. 26181] recognizes a holotype. •Valid as Asprocottus herzensteini Berg 1906 -- (Berg 1949:1167 [ref. 20662], Taliev 1955:316 [ref. 4316], Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:155 [ref. 23595], Sideleva 2001:61 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:17, 180 [ref. 27815], Bogutskaya & Naseka 2004:195 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Asprocottus herzensteini Berg 1906. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

herzensteini, Gymnocanthus Jordan [D. S.] & Starks [E. C.] 1904:294, Fig. 27 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate,Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7710. Paratypes: CAS-SU 7572 (1). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Gymnocanthus herzensteini Jordan & Starks 1904 -- (Watanabe 1960:51 [ref. 11959], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:197 [ref. 15964], Kim & Youn 1992:62 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:629 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Nakabo 2002:629 [ref. 26001], Youn 2002:262, 574 [ref. 26218], Parin et al. 2002:S74 [ref. 28300] as Gymnacanthus, Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:247 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331]). Current status: Valid as Gymnocanthus herzensteini Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

hexacornis, Cottus Richardson [J.] 1823:726 [22] [Narrative of a journey to the shores of the Polar Sea; ref. 18624] Mouth of Tree River near Coppermine River, Arctic Sea about Herschel Island. On p. 22 of separate. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1129 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Mecklenburg et al. 2002:477 [ref. 25968], Parin et al. 2014:262 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

hilgendorfii, Cottus Döderlein [L.] in Steindachner & Döderlein 1884:208 [40] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 49 (1. Abth.); ref. 4248] Tokyo, Japan. Holotype (unique): NMW. See also Watanabe 1960:121 [ref. 11959]. Both Watanabe and Okada described a new species Cottus japonicus but with hilgendorfii in synonymy [as hilgendorfi]. •Synonym of Cottus japonicus Okada 1961 -- (Okada 1961:699 [ref. 6363]). •Synonym of Cottus pollux Günther 1873 -- (Okada 1961:699 [ref. 6363], Zhang et al. 2016:193 [ref. 34477] with authorship as Steindachner & Doderlein). Current status: Synonym of Cottus. Cottidae. Habitat: freshwater.

hirundo, Leiocottus Girard [C. F.] 1856:133 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1809] San Miguel Island, near Santa Barbara, California, U.S.A. Syntypes: USNM 313 (1). •Valid as Leiocottus hirundo Girard 1856 -- (Bolin 1944:86 [ref. 6379], Eschmeyer & Herald 1983:175 [ref. 9277], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:124 [ref. 32708], Kells et al. 2016:180 [ref. 35888]). Current status: Valid as Leiocottus hirundo Girard 1856. Cottidae. Distribution: Eastern North Pacific: southern California to northern Baja California. Habitat: marine.

hispaniolensis, Cottus gobio Bǎcescu [M. C.] & Bǎcescu-Meşter [L.] 1964:444 [Vie et Milieu Suppl. No. 17; ref. 1238] Garona River at Viella, eastern Pyreneans, Spain. Syntypes: ?MGAB. •Synonym of Cottus gobio Linnaeus 1758 -- (Kottelat 1997:169 [ref. 22952]). •Valid as Cottus hispaniolensis Bǎcescu & Bǎcescu-Meşter 1964 -- (Freyhof et al. 2005:134 [ref. 28259], Kottelat & Freyhof 2007:511 [ref. 29996], Sideleva 2009:211 [ref. 30173], Goto et al. 2014:[11] [ref. 33456], Sideleva et al. 2018:9 [ref. 36393]). Current status: Valid as Cottus hispaniolensis Bǎcescu & Bǎcescu-Meşter 1964. Cottidae. Distribution: Garonne drainage, Spain and France. Habitat: freshwater.

hopliticus, Paricelinus Eigenmann [C. H.] & Eigenmann [R. S.] 1889:131 [9] [West American Scientist v. 6 (no. 48); ref. 1251] Cortez Bank, off San Diego, California, U.S.A., depth 45 fathoms. Holotype (unique): USNM 41881. On p. 9 of separate. •Valid as Paricelinus hopliticus Eigenmann & Eigenmann 1889 -- (Bolin 1944:11 [ref. 6379], Eschmeyer & Herald 1983:179 [ref. 9277], Matarese et al. 1989:404 [ref. 26885], McAllister 1990:201 [ref. 14674], Mecklenburg et al. 2002:429 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:44 [ref. 34694], Kells et al. 2016:188 [ref. 35888]). Current status: Valid as Paricelinus hopliticus Eigenmann & Eigenmann 1889. Cottidae. Distribution: Eastern North Pacific: Gulf of Alaska to southern California, U.S.A. Habitat: marine.

hoyi, Uranidea Putnam [F. W.] in Nelson 1876:41 [Bulletin of the Illinois Museum of Natural History v. 1 (no. 1); ref. 3154] Lake Michigan, 12 miles off Racine, Wisconsin, U.S.A., depth 42 fathoms. Holotype (unique): USNM 30903. •Synonym of Cottus cognatus Richardson 1837. Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

hubbsi, Cottus Bailey [R. M.] & Dimick [M. F.] 1949:2, Pl. 1 [Occasional Papers of the Museum of Zoology University of Michigan No. 513; ref. 12998] Entiat River, Chelan County, Washington, U.S.A. Holotype: UMMZ 145161. Paratypes: UMMZ 98675 (2), 98802 (2), 129736 (3), 145160 (2), 145162 (2); USNM 142967-70 (2, 1, 2, 2). •Synonym of Cottus bairdii Girard 1850 -- (Bailey & Bond 1963:1 [ref. 3934], Scott & Crossman 1998:830 [ref. 24570] as bairdi). •Valid as Cottus hubbsi Bailey & Dimick 1949 -- (Markle & Hill 2000:202 [ref. 24886], Nelson et al. 2004:119 [ref. 27807], McPhail 2007:475 [ref. 31199], Scharpf 2008:10 [ref. 30399], Page & Burr 2011:478 [ref. 31215], Page et al. 2013:122 [ref. 32708], LeMone et al. 2014:249 [ref. 33134], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus hubbsi Bailey & Dimick 1949. Cottidae. Distribution: Southwestern Canada and northwestern U.S.A. Habitat: freshwater.

hudsonius, Gymnocanthus tricuspis Vladykov [V. D.] 1933:16 [Contributions to Canadian biology and fisheries: being studies from the biological stations of Canada (Ser. A, General) v. 8 (no. 2); ref. 23244] Nottingham Island and Port Burwell, Hudson Bay region, Canada. Syntypes: (4). •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Scott & Scott 1988:496 [ref. 25518], Mecklenburg et al. 2016:62 [ref. 34440]). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Cottidae. Habitat: marine.

humilis, Cottus Bean [T. H.] 1881:149 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Chamisso Island, Spafarief Bay at mouth of Eschscholtz Bay, Kotzebue Sound, Alaska, U.S.A., Chukchi Sea, Arctic Ocean. Holotype (unique): USNM 27972. Non-types: MNHN 1890-0465 and 0466 [ex USNM] (2) station 3242. Type catalog: Blanc & Hureau 1968:51 [ref. 20738]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Mecklenburg et al. 2016:75 [ref. 34440]). •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Neyelov 1979:122 [ref. 3152], Mecklenburg et al. 2002:481 [ref. 25968], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Cottidae. Habitat: marine.

hypselurus, Cottus Robins [C. R.] & Robison [H. W.] 1985:361, Figs. 1-2 [American Midland Naturalist v. 114 (no. 2); ref. 17268] Bennett Springs, 11 miles northwest of Lebanon, Missouri, U.S.A. Holotype: UMMZ 150419. Paratypes: CUMV 23478 (26), 37382 (25). Additional material: ANSP 67275-77 (3); CUMV (5 lots, 143); KU (4 lots, 99); NLU (8 lots, 242; now 6 lots); SAU uncat (5 lots, 99); UMMZ (19 lots, 268); USNM 93272 (7). •Valid as Cottus hypselurus Robins & Robison 1985 -- (Robison & Buchanan 1988:477 [ref. 23393], Page & Burr 1991:250 [ref. 18983], Kinziger et al. 2000:1007 [ref. 25019], Gasper et al. 2001:264 [ref. 25578], Kinziger & Wood 2003:624 [ref. 27132] and also recognized a Cottus sp. cf. cypselurus, Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:10 [ref. 30399], Kinziger & Wood 2010:58 [ref. 30701], Page & Burr 2011:478 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus hypselurus Robins & Robison 1985. Cottidae. Distribution: Missouri and Arkansas, U.S.A. Habitat: freshwater.

ictalops, Pegedictis Rafinesque [C. S.] 1820:85 [Ichthyologia Ohiensis; ref. 3592] Near Lexington, Kentucky, U.S.A. No types known. •Synonym of Cottus bairdii Girard 1850 -- (Smith 1986:356 [ref. 22186] as bairdi). Current status: Synonym of Cottus bairdii Girard 1850. Cottidae. Habitat: freshwater.

ijimai, Porocottus Tanaka [S.] 1908:39 [244], Pl. 9 (figs. 3-4) [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 20; ref. 14360] Mouth of Susuya River, Sakhalin Island, Russia. Holotype: ZUMT 21700 [ex ZUMT 1388]. Paratypes: (3) FMNH 71846 [ex ZUMT 1389] (1), ZUMT (2 or 3, lost). Appeared first as above in Japanese. Then in English version in Tanaka 1908:244, Pl. 9 (figs. 3-4) [see ref. 14360]. •Synonym of Myoxocephalus platycephalus (Pallas 1814), subspecies taeniopterus (Kner 1868) -- (Berg 1949:1128 [ref. 20662]). •Valid as Porocottus ijimai Tanaka 1908 -- (Watanabe 1960:95 [ref. 11959]). •Synonym of Megalocottus platycephalus (Pallas 1814), subspecies taeniopterus (Kner 1868) -- (Neyelov 1979:130 [ref. 3152]). •Synonym of Megalocottus platycephalus (Pallas 1814) -- (Parin et al. 2002:S79 [ref. 28300]). •Synonym of Megalocottus taeniopterus (Kner 1868) -- (Parin et al. 2014:258 [ref. 33547], Dyldin & Orlov 2017:64 [ref. 35138] with question). Current status: Synonym of Megalocottus taeniopterus (Kner 1868). Cottidae. Habitat: brackish, marine.

immaculatus, Cottus Kinziger [A. P.] & Wood [R. M.] 2010:51, Figs. 1, 2B, 3 [Zootaxa No. 2340; ref. 30701] Current River at Baptist Camp Access, 2.7 air kilometers east of Montauk State Park, 37°26.111'N, 91°39.447'W, Dent County, Missouri, U.S.A. Holotype: USNM 369996. Paratypes: INHS 102783 (10), USNM 396997 (10), UW 118923 (5). Plus non-type specimens. •Valid as Cottus immaculatus Kinziger & Wood 2010 -- (Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus immaculatus Kinziger & Wood 2010. Cottidae. Distribution: Arkansas and Missouri, U.S.A. Habitat: freshwater.

incitus, Myoxocephalus Watanabe [M.] 1958:294, Figs. 121-122, Pl. 21 [Studies of Japanese cottid fishes; ref. 21357] Same, Aomori Prefecture, Japan, depth 10 fathoms. Holotype (unique): not researched. Also described as new in Watanabe 1960:85, Pl. 27 (fig. 1) [ref. 11959]. •Valid as Myoxocephalus incitus Watanabe 1958 -- (Yabe in Masuda et al. 1984:327 [ref. 6441]). •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Cottidae. Distribution: Western North Pacific. Habitat: brackish, marine.

inermis, Acanthocottus Ayres [W. O.] 1854:170 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] San Francisco Bay, California, U.S.A. No types known. Originally as Acanthocottus (?) inermis. Appeared first as above on 1 Sept. •Synonym of Leptocottus armatus Girard 1854. Current status: Synonym of Leptocottus armatus Girard 1854. Cottidae. Habitat: brackish, marine.

inermis, Cottus Yakovlev [V. E.] 1890:52 [4] [Izvestiia Vostochno-Sibirskogo otdiela Imperatorskago geograficheskago obshchestva. v. 21 (no. 3); ref. 19625] Irkut River, tributary of Angara River, Lake Baikal, Russia (Angara River near Irkutsk). Neotype: ZIN 6350. On p. 4 of separate. Sideleva 2001:58 [ref. 26181] names a neotype [ZIN 6350], but if that is the first time a neotype was named it is unavailable under the Code. Neotype designated by Sideleva 2003:166 [ref. 27815]. •Synonym of Cottocomephorus grewingkii (Dybowski 1874) -- (Berg 1949:1171 [ref. 20662]). •Valid as Cottocomephorus inermis (Yakovlev 1890) -- (Taliev 1955:292 [ref. 4316], Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:150 [ref. 23595], Bogutskaya et al. 2001:46 [ref. 26178], Sideleva 2001:58 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:16, 166 [ref. 27815], Bogutskaya & Naseka 2004:192 [ref. 28183], Goto et al. 2014:[15] [ref. 33456], Bogdanov 2019:230 [ref. 36814]). Current status: Valid as Cottocomephorus inermis (Yakovlev 1890). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

infernatis, Cottus carolinae Williams [J. D.] & Robins [C. R.] 1970:371, Fig. 1 [American Midland Naturalist v. 83 (no. 2); ref. 7791] Chewacla Creek, Chewacla State Park, about 3 miles south of Auburn, Lower Alabama river system, Lee County, Alabama, U.S.A. Holotype: USNM 203762. Paratypes: CUMV 53321 (4); TU 32497 (5); UAIC 1237 (4), 2118 (3); UMMZ 111208 (5); USNM 20078 (4), 203336-39 (1, 1, 1, 1). Type catalog: Bart & Taylor 1993:66-67 [ref. 20906]. •Synonym of Cottus carolinae (Gill 1861) -- (Lee et al. 1980:807 [ref. 22416], Robison & Buchanan 1988:476 [ref. 23393]). •Synonym of Cottus carolinae (Gill 1861), but a valid subspecies as described -- (Boschung & Mayden 2004:423 [ref. 27995] as infernatus, Neely et al. 2007:651 [ref. 29262] as inferenatus, Scharpf 2008:8 [ref. 30399], Page & Burr 2011:470 [ref. 31215]). Current status: Synonym of Cottus carolinae (Gill 1861). Cottidae. Distribution: Alabama, U.S.A. [subspecies infernatis]. Habitat: freshwater.

ingens, Artediellus Nelson [D. W.] 1986:35, Fig. 7 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 138 (no. 1); ref. 5808] Cape Rollin, Simushir Island, Kurils. [Kuril], Albatross station 4804, 46°42'N, 151°47'E, depth 429 meters. Holotype: USNM 74460. Paratypes: CAS-SU 22237 (4); USNM 74484-85 (13, 20). •Valid as Artediellus ingens Nelson 1986 -- (Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:404 [ref. 25968], Parin et al. 2002:S70 [ref. 28300], Parin et al. 2014:237 [ref. 33547]). Current status: Valid as Artediellus ingens Nelson 1986. Cottidae. Distribution: Western North Pacific. Habitat: marine.

insularis, Paracottus Taliev [D. N.] 1955:42 et seq., 241, Figs. 2 et seq., Fig. 92 [ ref. 4316] Lake Baikal, Russia. Syntypes: none found at ZIN. Appeared as Cottus insularis in Taliev 1948:113 [ref. 19628] without distinguishing features. As Paracottus (Paracottus) insularis on p. 235 and 241. •Synonym of Paracottus knerii (Dybowski 1874) -- (Sideleva 1982:28 [ref. 14469], Sideleva 1998:153 [ref. 23595], Sideleva 2001:59 [ref. 26181], Sideleva 2003:16, 168 [ref. 27815]). Current status: Synonym of Paracottus knerii (Dybowski 1874). Cottidae. Habitat: freshwater.

intermedius, Asprocottus herzensteini Taliev [D. N.] 1955:319, Figs. 132-133 [ ref. 4316] Lake Baikal, Russia. Syntypes: no types found at ZIN. Appeared without distinguishing features and not available in Taliev 1948:113 [ref. 19628] as Asprocottus intermedius. •Valid as Asprocottus intermedius Taliev 1955 -- (Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702] dating to 1948, Sideleva 1998:156 [ref. 23595], Bogutskaya et al. 2001:47 [ref. 26178], Bogutskaya & Naseka 2004:195 [ref. 28183]). Current status: Valid as Asprocottus intermedius Taliev 1955. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

intermedius, Cottus Temminck [C. J.] & Schlegel [H.] 1843:38 [Fauna Japonica Parts 2-4; ref. 4371] Coast of Yezo, Japan. Holotype: RMNH D652 (dry skin, poor condition). Type information: Boeseman 1947:48 [ref. 12876]. •Valid as Gymnocanthus intermedius (Temminck & Schlegel 1843) -- (Watanabe 1960:54 [ref. 11959], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:198 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:629 [ref. 25086], Nakabo 2002:629 [ref. 26001], Youn 2002:262, 575 [ref. 26218], Parin et al. 2002:S74 [ref. 28300] as Gymnacanthus, Mundy 2005:336 [ref. 28379], Shinohara et al. 2014:249 [ref. 33330]). Current status: Valid as Gymnocanthus intermedius (Temminck & Schlegel 1843). Cottidae. Distribution: Western North Pacific. Habitat: marine.

intermedius, Icelus spiniger Lindberg [G. U.] & Andriashev [A. P.] 1938:517, Figs. 1-2 [Trudy Gidrobiologicheskoi Ekspeditsii Zoologicheskogo instituta AN SSSR No. 1; ref. 19731] Okhotsk Sea, southeastern Kamchatka, Russia, 52°33.3'N, 154°54'E, depth 185-195 meters. Holotype: ZIN 24470 (185 mm). Paratypes: ZIN 24470 (1, with holotype), 27962 (2), 27969 (3). Appeared first as a nomen nudum in Andriashev 1937:301[not researched]. •Synonym of Icelus spiniger Gilbert 1896 -- (Nelson 1984:24 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:257 [ref. 33547]). Current status: Synonym of Icelus spiniger Gilbert 1896. Cottidae. Habitat: marine.

ishikawae, Furcina Jordan [D. S.] & Starks [E. C.] 1904:303, Fig. 32 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Rikuchu, Myiako, Japan. Holotype: CAS-SU 7715. Paratypes: AMS I.6953 (6); BMNH 1905.2.4.365-372 (8?); CAS 31363 [ex IU 10719] (3, c&s), 58402 [ex IU 10719] (6); CAS-SU 7580 (18), 7582 (3, dry), 7587 (4); USNM 50924 (32), 74788 (1). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Furcina ishikawae Jordan & Starks 1904 -- (Watanabe 1960:137 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:244 [ref. 15964], Kim & Youn 1992:72 [ref. 22287], Nakabo 2000:648 [ref. 25086], Shimizu 2001:27 [ref. 25799], Nakabo 2002:648 [ref. 26001], Youn 2002:262, 574 [ref. 26218]). Current status: Valid as Furcina ishikawae Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

islandicus, Triglops pingelii Jensen [A. S.] 1944:16, 29 [Spolia Zoologica Musei Hauniensis v. 4; ref. 13060] Iceland. Syntypes: ZMUC 56 (1), 158 (1), P811667-735 (69). Type catalog: Nielsen 1974:82 [ref. 9588]. •Synonym of Triglops murrayi Günther 1888 -- (Pietsch 1994:369 [ref. 21189], Parin et al. 2014:276 [ref. 33547]). Current status: Synonym of Triglops murrayi Günther 1888. Cottidae. Habitat: marine.

jaok, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:172 [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka coast, Russia. Holotype (unique): ZMB 31656. Type catalog: Blanc & Hureau 1968:60 [ref. 20738], Paepke & Fricke 1992:284 [ref. 19946]. •Valid as Myoxocephalus jaok (Cuvier 1829) -- (Watanabe 1960:75 [ref. 11959], Neyelov 1979:122 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:212 [ref. 15964], Kim & Youn 1992:64 [ref. 22287], Sokolovsky & Sokolovskaya 1997:55 [ref. 24691], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:645 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:481 [ref. 25968], Nakabo 2002:645 [ref. 26001], Youn 2002:262, 576 [ref. 26218], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:75 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Bogutskaya & Naseka 2004:190 [ref. 28183], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:261 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Tohkairin et al. 2015:7 [ref. 35331], Mecklenburg et al. 2016:76 [ref. 34440], Kells et al. 2016:182 [ref. 35888], Dyldin & Orlov 2017:65 [ref. 35138], Mecklenburg et al. 2018:213 [ref. 36370]). Current status: Valid as Myoxocephalus jaok (Cuvier 1829). Cottidae. Distribution: North Pacific and adjacent Arctic. Habitat: brackish, marine.

japonicus, Centridermichthys Steindachner [F.] 1870:625 [3], Pl. 1 (fig. 3) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 61 (1. Abth.); ref. 4218] Nagasaki, Japan. Holotype (unique): NMW. On p. 3 of separate. Also described and figured as a new species in Steindachner 1881:187 [9 of separate] [ref. 4231]; 1881 description is based on a different specimen. •Synonym of Pseudoblennius cottoides (Richardson 1848) -- (Watanabe 1960:152 [ref. 11959], Lindberg & Krasyukova 1987:252 [ref. 15964], Kim & Youn 1992:74 [ref. 22287]). Current status: Synonym of Pseudoblennius cottoides (Richardson 1848). Cottidae. Habitat: marine.

japonicus, Centridermichthys Steindachner [F.] 1881:187 [9], Pl. 7 (figs. 1, 1a) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 83 (1. Abth.); ref. 4231] Kanagava, Japan. On p. 9 of separate. Apparently described independently of Centridermichthys japonicus Steindachner 1870. Objectively invalid; preoccupied by Centridermichthys japonicus Steindachner 1870. Cottidae. Habitat: freshwater.

japonicus, Cottus Watanabe [M.] 1960:121, Pl. 7 (fig. 1) [Fauna Japonica: Cottidae (Pisces); ref. 11959] Hei River, Miyako, & Koshi River, Kamaishi-gun, Iwate Prefecture; and Haguro River, Kitamurayama-gun, Yamagata Prefecture, Japan. Syntypes: (51). See also Cottus japonicus Okada 1961. •Objectively invalid; preoccupied by Cottus japonicus Pallas 1769. •Status uncertain. Current status: Synonym of Cottus. Cottidae. Habitat: freshwater.

japonicus, Cottus Okada [Y.] 1961:699, Figs. 121 a-e [Studies on the freshwater fishes of Japan.; ref. 6363] Japan. Syntypes: whereabouts unknown. Objectively invalid; preoccupied by Cottus japonicus Pal/las 1769 and Cottus japonicus Watanabe 1960. Clearly described as a new species, but perhaps intended to be the same as Cottus japonicus Watanabe 1960 with same common name "Kajika" and based on same sources. Okada's work was apparently to predate Watanabe's work, but Okada dates to 1961 and Watanabe to 1960; perhaps last minute changes resulted in the description of two new species. •Objectively invalid; preoccupied by Cottus japonicus Pallas 1769. •Status uncertain. Current status: Synonym of Cottus. Cottidae. Habitat: freshwater.

japonicus, Icelinus Yabe [M.], Tsumura [K.] & Katayama [M.] 1980:106, Figs. 1-2 [Japanese Journal of Ichthyology v. 27 (no. 2); ref. 8640] Mano Bay, off Sado Island, Niigata Prefecture, Japan, 37°57'N, 138°19'E. Holotype: HUMZ 79035. Paratypes: HUMZ 75466 (1), 77562 (1), 77563 (1, dissected); NSMT-P 18575 (1). Other material: HUMZ 77561 (1). •Valid as Icelinus japonicus Yabe, Tsumura & Katayama 1980 -- (Yabe in Masuda et al. 1984:325 [ref. 6441], Peden 1984:79 [ref. 5193], Suzuki & Uno 1992:4 [ref. 24139], Nakabo 2000:636 [ref. 25086], Nakabo 2002:636 [ref. 26001], Rosenblatt & Smith 2004:560 [ref. 27912]). Current status: Valid as Icelinus japonicus Yabe, Tsumura & Katayama 1980. Cottidae. Distribution: Western North Pacific. Habitat: marine.

japonicus, Neohemilepidotus Sakamoto [K.] 1932:4, Fig. 2 [Journal of the Imperial Fisheries Institute v. 27 (no. 1); ref. 3860] Northern Japan. Holotype: whereabouts unknown. Paratypes: (1) whereabouts unknown. •Valid as Neohemilepidotus japonicus Sakamoto 1932 -- (Watanabe 1958 [ref. 21357] and 1960:45 [ref. 11959] and Lindberg & Krasyukova 1987:182 [ref. 15964] but as pacificus by mistake and not japonicus). •Synonym of Melletes papilio Bean 1880 -- (Parin et al. 2002:S79 [ref. 28300]). •Synonym of Hemilepidotus papilio (Bean 1880) -- (Peden 1979:41 [ref. 24080], Parin et al. 2014:252 [ref. 33547], Dyldin & Orlov 2017:64 [ref. 35138]). Current status: Synonym of Hemilepidotus papilio (Bean 1880). Cottidae. Habitat: marine.

japonicus, Porocottus Schmidt [P. J.] in Taranetz 1935:177 [Vestnik Dal'nevostocnogo Filiala Akademii Nauk SSSR = Bulletin of the Far Eastern Branch of the Academy of Sciences of the USSR No. 14; ref. 21496] DeCastri Bay and western coast of Sakhalin. Lectotype: ZIN 26314 or 6314. Paralectotypes: ZIN 18962. Lectotype designation not researched, not designated here. •Valid as Porocottus japonicus Schmidt 1935 -- (Neyelov 1979:137 [ref. 3152], Lindberg & Krasyukova 1987:221 [ref. 15964], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S84 [ref. 28300], Parin et al. 2014:268 [ref. 33547], Dyldin & Orlov 2017:67 [ref. 35138]). Current status: Valid as Porocottus japonicus Schmidt 1935. Cottidae. Distribution: Western North Pacific. Habitat: marine.

japonicus, Zesticelus Oshima [M.] 1957:2, Fig. 1 [Japanese Journal of Ichthyology v. 6 (nos 1-2); ref. 12161] Offshore of Niigata, Japan. Holotype (unique): Japan Sea Reg. Fish. Lab. Niigata No. 1335 (lost). •Valid as Zesticelus japonicus Oshima 1957 -- (Neyelov 1979:161 [ref. 3152], Yabe 1995:19 [ref. 22666]). •Mention as Zeticelus japonicus Oshima 1957 -- (Yabe & Okamura 1993:370 [ref. 22281]). •Status uncertain -- (Nakabo 2002:1526 [ref. 26001]). Current status: Uncertain as Zesticelus japonicus Oshima 1957. Cottidae. Distribution: Western North Pacific. Habitat: marine.

jaxartensis, Cottus gobio Berg [L. S.] 1916:437 [Les Poissons des eaux douces de la Russie; ref. 277] Syrdarja, Ugam River, tributary of Chirchik River, Syr-Darya basin, Uzbekistan. Holotype (unique): not researched. •Synonym of Cottus gobio Linnaeus 1758, but a valid subspecies as described -- (Berg 1949:1148 [ref. 20662]). •Tentatively a synonym of Cottus gobio Linnaeus 1758 -- (Kottelat 1997:169 [ref. 22952]). Current status: Synonym of Cottus gobio Linnaeus 1758. Cottidae. Habitat: freshwater, brackish.

jeittelesii, Cottus Dybowski [B. N.] 1874:386, Pl. 3 (fig. 1) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 24 (no. 3-4); ref. 1172] Lake Baikal, Russia. Lectotype: ZIN 3240. Paralectotypes: ? ZMB 7946 (1). Type catalog: Paepke & Fricke 1992:285 [ref. 19946]. Lectotype selected by Sideleva 2003:203 [ref. 27815]. •Valid as Procottus jeittelesii (Dybowski 1874) -- (Berg 1949:1168 [ref. 20662], Taliev 1955:302 [ref. 4316], Sideleva 1982:31 [ref. 14469] as jeittelesi, Reshetnikov et al. 1997:761 [ref. 24702], Sideleva 1998:158 [ref. 23595], Sideleva 2001:69 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:19, 203 [ref. 27815], Bogutskaya & Naseka 2004:200 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Procottus jeittelesii (Dybowski 1874). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

jordani, Elanura Schmidt [P. Yu.] 1903:518 [16] [Izvestiya Imperatorskago Russkago Geograficheskago Obshchestva v. 38 (pt 5) (for 1902); ref. 3945] On p. 16 of separate. Not available, name and localities but no description. See Prionistius jordani Schmidt in Jordan & Starks 1904. Invalid lectotype designation by Pietsch 1994:351 [ref. 21189] as SU 7704. •Treated as valid as Elanura jordani Schmidt 1903 -- (Kim & Youn 1992:66 [ref. 22287]). •Treated as valid as Triglops jordani (Schmidt 1904) -- (Yabe in Masuda et al. 1984:325 [ref. 6441], Pietsch 1994:351 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Nakabo 2000:628 [ref. 25086], Mecklenburg et al. 2002:422 [ref. 25968], Nakabo 2002:628 [ref. 26001], Youn 2002:263, 578 [ref. 26218], Shinohara et al. 2011:44 [ref. 31715]). Current status: Synonym of Triglops jordani (Schmidt 1904). Cottidae.

jordani, Hemilepidotus Bean [T. H.] 1881:153 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Iliuliuk [Ilyulyuk], Unalaska Island, Aleutian Islands, Alaska, U.S.A. Lectotype: USNM 27598. Paralectotype: USNM 201606 [ex 27598] (1). Non-type: MNHN 1896-0477 [ex USNM] (1) station 3333. Lectotype established (as figured specimen) in caption to Pl. 290, p. 3291 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. •Valid as Hemilepidotus jordani Bean 1881 -- (Peden 1979:32 [ref. 24080], Eschmeyer & Herald 1983:171 [ref. 9277], Yabe in Masuda et al. 1984:324 [ref. 6441], Matarese et al. 1989:376 [ref. 26885], Shinohara et al. 1996:176 [ref. 26605] but corrected to gilberti with insert, Nakabo 2000:631 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:436 [ref. 25968], Nakabo 2002:631 [ref. 26001], Parin et al. 2002:S76 [ref. 28300], Tokranov et al. 2003:300 [ref. 26972], Fedorov et al. 2003:70 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:251 [ref. 33547], Mecklenburg & Steinke 2015:114 [ref. 34033], Mecklenburg et al. 2016:63 [ref. 34440], Kells et al. 2016:174 [ref. 35888], Mecklenburg et al. 2018:201 [ref. 36370]). Current status: Valid as Hemilepidotus jordani Bean 1881. Cottidae. Distribution: North Pacific and margin of adjacent Arctic Region. Habitat: marine.

jordani, Prionistius Schmidt [P. J.] in Jordan & Starks 1904:252, Fig. 9 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Peter the Great Bay, Vladivostok, Russia. Holotype: CAS-SU 7704 [from Schmidt]. Type catalog: Böhlke 1953:131 [ref. 12291]. Appeared first without distinguishing features in Schmidt 1903:518 (p. 16 of separate) [ref. 3945]. Appeared in 1904 in Jordan & Starks as above and as Prionistius jordani, with the description based solely on one "cotype" received from Schmidt; this cotype is the holotype and authorship is Jordan & Starks as above. Pietsch 1994:351 [ref. 21189] regarded Schmidt 1903 as the original description and named SU 7704 as the lectotype. •Valid as Triglops jordani (Jordan & Starks 1904) -- (Lindberg & Krasyukova 1987:177 [ref. 15964], Pietsch 1994:351 [ref. 21189], Sokolovskaya et al. 1998:10 [ref. 24670], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:422 [ref. 25968], Parin et al. 2002:S87 [ref. 28300], Fedorov et al. 2003:81 [ref. 27579], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:275 [ref. 33547], Dyldin & Orlov 2017:68 [ref. 35138] as Schmidt in Jordan & Starks 1904). Current status: Valid as Triglops jordani (Schmidt 1904). Cottidae. Distribution: Western North Pacific: southern Kamchatka to Okhotsk Sea and Japan Sea off Korea. Habitat: brackish, marine.

kaganowskii, Cottus Berg [L. S.] 1932:17, Figs. 1-2 [Copeia 1932 (no. 1); ref. 16356] Belaya River to the mouth, Anadyr River system, northeastern Siberia, Russia. Syntypes: ZIN 23140-41 (1, 1), 23805-09 (23, 20, 2, 1, 1). •Valid as Cottus kaganowskii Berg 1932 -- (Berg 1949:1152 [ref. 20662]). •Synonym of Cottus cognatus Richardson 1837 -- (Lee et al. 1980:808 [ref. 22416], Sideleva 1998:151 [ref. 23595], Sheiko & Fedorov 2000:27 [ref. 25650], Parin et al. 2002:S73 [ref. 28300]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

kallavesensis, Cottus quadricornis race Lönnberg [E.] 1933:7, Pl. 1 (fig. 1) [Arkiv för Zoologi v. 24 A (pt 3) (no. 12) [1932]; ref. 22149] Lake Kallavesi; Lake Niinivesi in Rautalampi; Lake Virmavesi in Karttula, Lake Päijänne basin, Finland. Syntypes: MZH (9). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies onegensis Berg & Popov 1932 -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

kanawhae, Cottus Robins [C. R.] 2005:1, Fig. 1 [Zootaxa No. 987; ref. 28230] Creek in Tony's Cave, tributary of Sinking Creek, Giles County, Virginia, U.S.A. Holotype: CUMV 25655. Paratypes: CUMV 12453 (19), 21946 (5), 22085 (2), 22087 (2), 22832 (13), 22833 (31), 22835 (1), 22840 (11), 24397 (1); USNM 162191 (6). Plus non-type material. •Valid as Cottus kanawhae Robins 2005 -- (Scharpf 2008:10 [ref. 30399], Page & Burr 2011:470 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus kanawhae Robins 2005. Cottidae. Distribution: New River system, Virginia and West Virginia, U.S.A. Habitat: freshwater.

karaensis, Icelus Soldatov [V. K.] 1923:31 [English p. 74], Fig. 34 [Russkij gidrobiologiceskij zurnal: organ Obscestva Issledovatelej Vody i ee Zizni = Russian hydrobiologische Zeitschrift izd. pri Volzkoj Biologicaskoj Stancii - Saratow v. 3; ref. 15569] Kara Sea, 70°08'N, 56°26'E, Russia. Syntypes: ZIN 22027 (3). •Synonym of Icelus spatula Gilbert & Burke 1912 -- (Neyelov 1973:597 [ref. 7219], Nelson 1984:42 [ref. 5391], Parin et al. 2002:S78 [ref. 28300], Parin et al. 2014:256 [ref. 33547]). Current status: Synonym of Icelus spatula Gilbert & Burke 1912. Cottidae. Habitat: marine.

kazika, Cottus Jordan [D. S.] & Starks [E. C.] 1904:265, Fig. 15 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Niigata, Japan. Holotype: CAS-SU 7705. Paratypes: USNM 50918 (1), 57795 (2). Non-types: ?BMNH 1905.2.4.397 (1), CAS-SU 7578 (5). Type catalog: Böhlke 1953:128 [ref. 12291]. The species name was spelled kazika in three places, but in the derivation it is stated, "(kajika, the Japanese name of river sculpins.)". •Valid as Cottus kazika Jordan & Starks 1904 -- (Watanabe 1960:117 [ref. 11959] as kajika, Okada 1961:696 [ref. 6363], Yabe in Masuda et al. 1984:326 [ref. 6441], Kawanabe & Mizuno 1989:655 [ref. 25598], Takahashi et al. 1997:45 [ref. 24971], Sato et al. 1998:39 [ref. 24972], Nakabo 2000:637 [ref. 25086], Matsuura et al. 2000:203 [ref. 26000], Nakabo 2002:637 [ref. 26001], Fujii et al. 2005:12 [ref. 28191], Goto et al. 2014:[9] [ref. 33456]). Current status: Valid as Cottus kazika Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: freshwater, brackish, marine.

kesslerii, Cottus Dybowski [B. N.] 1874:384, Pl. 2 (fig. 1) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 24 (no. 3-4); ref. 1172] Lake Baikal; Angara, Irkut and Selenga rivers, Siberia, Russia. Syntypes: BMNH 1897.7.5.5 (1), ZMB 7807 (4). Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. •Valid as Paracottus kessleri (Dybowski 1874) -- (Taliev 1955:243 [ref. 4316]). •Valid as Cottus kesslerii Dybowski 1874 -- (Berg 1949:1153 [ref. 20662] as kessleri, Sideleva 1982:27 [ref. 14469] as kessleri, Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:151 [ref. 23595]). •Valid as Leocottus kesslerii (Dybowski 1874) -- (Bogutskaya et al. 2001:46 [ref. 26178], Sideleva 2001:59 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:16, 151 [ref. 27815], Bogutskaya & Naseka 2004:189 [ref. 28183], Kottelat 2006:60 [ref. 28831], Ocock et al. 2006:58 [ref. 28933], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Leocottus kesslerii (Dybowski 1874). Cottidae. Distribution: Lake Baikal and tributaries, Russia and Mongolia. Habitat: freshwater.

klamathensis, Cottus Gilbert [C. H.] 1898:10, Fig. [Bulletin of the U. S. Fish Commission v. 17 (art. 1) [1897]; ref. 12349] Upper Klamath Lake near Klamath Falls, Klamath County, Oregon, U.S.A. Holotype: USNM 48226. Paratypes: (20) ANSP 39138 (1), 39143 (1); BMNH 1900.9.29.195-196 (2); ?CAS-SU 6290 (63); ?UMMZ 130637 (19). Type catalog: Böhlke 1953:128 [ref. 12291], Böhlke 1984:65 [ref. 13621]. •Valid as Cottus klamathensis subspecies klamathensis Gilbert 1898 -- (Miller & Taylor 1984:938 [ref. 6809], Moyle 2002:356 [ref. 26449]). •Valid as Cottus klamathensis Gilbert 1898 -- (Lee et al. 1980:815 [ref. 22416], Daniels & Moyle 1984:954 [ref. 8977], Page & Burr 1991:253 [ref. 18983], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:10 [ref. 30399] with subspecies, Page & Burr 2011:482 [ref. 31215] with 3 subspecies, Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[9] [ref. 33456]). Current status: Valid as Cottus klamathensis Gilbert 1898. Cottidae. Distribution: Western U.S.A.: Oregon and California. Klamath River above Kalamath Falls [subspecies klamathensis]. Habitat: freshwater.

knerii, Cottus Dybowski [B. N.] 1874:385, Pl. 2 (fig. 2) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 24 (no. 3-4); ref. 1172] Lake Baikal; Angara, Irkut, Selenga rivers, Siberia, Russia. Syntypes: ZMB 7809 (1), 7947 (2). Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. •Valid as Cottus knerii Dybowski 1874 -- (Berg 1949:1151 [ref. 20662] as kneri). •Valid as Paracottus knerii (Dybowski 1874) -- (Taliev 1955:236 [ref. 4316], Sideleva 1982:28 [ref. 14469], Chereshnev 1996:606 [ref. 24659] as kneri, Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:153 [ref. 23595], Bogutskaya et al. 2001:46 [ref. 26178], Sideleva 2001:58 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:16, 168 [ref. 27815], Bogutskaya & Naseka 2004:193 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Paracottus knerii (Dybowski 1874). Cottidae. Distribution: Lake Baikal and tributaries, Russia. Habitat: freshwater.

kolymensis, Cottus Sideleva [V. G.] & Goto [A.] 2012 [Voprosy Ikhtiologii v. 52; ref. 32050] Holotype. Full account not yet seen. •Valid as Cottus kolymensis Sideleva & Goto 2012 -- (Parin et al. 2014:241 [ref. 33547], Parin et al. 2014:243 [ref. 33547]). Current status: Valid as Cottus kolymensis Sideleva & Goto 2012. Cottidae. Distribution: Kolyma, Dukcha nnd Glukhaya rivers, Russia. Habitat: freshwater.

koreanus, Cottus Fujii [R.], Choi [Y.] & Yabe [M.] 2005:8, Figs. 1-3 [Species Diversity v. 10 (no. 1); ref. 28191] Stream at Mt. Chiak, Namhan River system, Hackkok-ri, Socho-myon, Wonju-gun, Kangwon-do, Korea. Holotype: HUMZ 155725. Paratypes: CNUC 25009-10 (2); HUMZ 155722 (1), 155726-39 (14), 155764 (1); NSMT-P 60690-91 (2). Plus non-type material. •Valid as Cottus koreanus Fujii, Choi & Yabe 2005 -- (Sideleva & Goto 2009:607 [ref. 30591], Goto et al. 2014:[9] [ref. 33456], Parin et al. 2014:241 [ref. 33547]). Current status: Valid as Cottus koreanus Fujii, Choi & Yabe 2005. Cottidae. Distribution: Korea. Habitat: freshwater.

korjakovi, Asprocottus Sideleva [V. G.] 2001:61, Fig. 1 [Proceedings of the Zoological Institute, Academy of Sciences, St.-Petersburg No. 287; ref. 26181] Lake Baikal, Maloye Morye Strait, Zunduk Cape, Russia, depth 115-125 meters. Holotype: ZIN 50871. Paratypes: ZIN 52086 (1). •Valid as Asprocottus korjakovi Sideleva 2001, with subspecies -- (Sideleva 2001:61 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:17, 183 [ref. 27815], Bogutskaya & Naseka 2004:195 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Asprocottus korjakovi Sideleva 2001. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

korotneffi, Abyssocottus Berg [L. S.] 1906:908 [Zoologischer Anzeiger v. 30 (no. 26); ref. 265] Lake Baikal, Russia, depth 700 feet. Lectotype: ZIN 13733. Paralectotypes: ZIN 13733A (1). Sideleva 2001:61 [ref. 26181] lists a holotype [ZIN 50871], but that is invalid under the code. She designated a lectotype in Sideleva 2003:174 [ref. 27815]. •Valid as Abyssocottus korotneffi Berg 1906 -- (Berg 1949:1157 [ref. 20662], Taliev 1955:348 [ref. 4316], Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:155 [ref. 23595], Bogutskaya et al. 2001:47 [ref. 26178], Sideleva 2001:61 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:17 [ref. 27815], Bogutskaya & Naseka 2004:194 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Abyssocottus korotneffi Berg 1906. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

koshewnikowi, Cottus Gratzianov [V. I.] 1907:658 [Zoologischer Anzeiger v. 31 (no. 21/22); ref. 1872] Goredva River, Ligachevo village, Moscow Oblast, Russia. Lectotype: ZMMU P-2688. Paralectotypes: ZMMU P-2001 (4), P-2658 (20), P-2688 (2, now 1), P-3118 (1), P-3121 (3), P-3912 (1). Type catalog: Svetovidova 1978:259 [ref. 9011], Vasil'eva et al. 2001:79-80 [ref. 25714]. Also in Gratsianov 1907:315 [ref. 1871], earliest not researched. Name also seen as Gratsianow. Lectotype designated by Sideleva et al. 2015:33 [ref. 35276]. •Synonym of Cottus gobio Linnaeus 1758, but a valid subspecies -- (Berg 1949:1147 [ref. 20662], Lelek 1987:274 [ref. 22962], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:151 [ref. 23595]). •Synonym of Cottus gobio Linnaeus 1758 -- (Kottelat 1997:169 [ref. 22952], Tsvetkov et al. 2001:121 [ref. 26836]). •Valid as Cottus koshewnikowi Gratzianov 1907 -- (Bogutskaya & Naseka 2004:187 [ref. 28183], Freyhof et al. 2005:163 [ref. 28259], Fricke 2007:29 [ref. 30577], Kottelat & Freyhof 2007:511 [ref. 29996], Goto et al. 2014:[11] [ref. 33456], Sideleva et al. 2015:429 [ref. 33770], Sideleva et al. 2015:34 [ref. 35276]). Current status: Valid as Cottus koshewnikowi Gratzianov 1907. Cottidae. Distribution: Northern Europe. Habitat: freshwater, brackish.

kozovi, Limnocottus Taliev [D. N.] 1946:737, Fig. 1 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 52 (no. 8); ref. 13135] Baunt Lake, east of Lake Baikal, Vitim River basin, Buryatia, Russia [in error for Lake Baikal]. Holotype (unique): ZIN 49713. Sideleva (pers. comm.) indicates that this species does not occur in Baunt Lake. •Valid as Asprocottus kozovi (Taliev 1946) -- (Taliev 1955:339 [ref. 4316]). •Valid as Limnocottus kozovi Taliev 1946 -- (Berg 1949:1164 [ref. 20662]). •Valid as Asprocottus kozovi (Taliev 1946) -- (Taliev 1955:75 [ref. 4316]). •Synonym of Limnocottus megalops (Gratzianov 1902) -- (Sideleva 1998:157 [ref. 23595]). •Synonym of Cyphocottus megalops (Gratzianov 1902) -- (Sideleva 2003:191 [ref. 27815]). Current status: Synonym of Cyphocottus megalops (Gratzianov 1902). Cottidae. Habitat: freshwater.

krawchukii, Myoxocephalus quadricornis Mikhalev [Y. V.] 1962:137 Reference not traced, date 1964? •Probably a synonym of Myoxocephalus quadricornis (Linnaeus 1758). Current status: Uncertain as Myoxocephalus quadricornis (Linnaeus 1758). Cottidae.

kumlieni, Uranidea Hoy [P. R.] in Nelson 1876:41 [Bulletin of the Illinois Museum of Natural History v. 1 (no. 1); ref. 3154] Deep water in Lake Michigan, U.S.A. Holotype (unique): USNM 30904. Authorship uncertain from text, perhaps Nelson (ex Hoy), with description from one of Hoy's types. •Synonym of Cottus bairdii Girard 1850 -- (Smith 1986:356 [ref. 22186], Scott & Crossman 1998:826 [ref. 24570] as bairdi). •Synonym of Cottus bairdii Girard 1850, but a valid subspecies -- (Kinziger et al. 2000:1007 [ref. 25019] as bairdi, Scharpf 2008:7 [ref. 30399]). Current status: Synonym of Cottus bairdii Girard 1850. Cottidae. Distribution: Canada and northern U.S.A. [this subspecies]. Habitat: freshwater.

kuznetzovi, Cottus Berg [L. S.] 1903:110 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 8; ref. 263] Chuya River, Lena River basin [not Lake Baikal basin], Russia. Neotype: ZIN 53969. Unique holotype [ZIN 12476] lost; neotype selected by Sideleva & Goto 2009:607 [ref. 30591]. •Synonym of Cottus poecilopus Heckel 1837 -- (Berg 1949:1143 [ref. 20662], Fedorov in Whitehead et al. 1986:1248 [ref. 13677], Sideleva 1998:152 [ref. 23595], Freyhof et al. 2005:167 [ref. 28259]). •Valid as Cottus kuznetzovi berg 1903 -- (Sideleva & Goto 2009:607 [ref. 30591], Goto et al. 2014:[9] [ref. 33456], Parin et al. 2014:241 [ref. 33547], Parin et al. 2014:244 [ref. 33547]). Current status: Valid as Cottus kuznetzovi Berg 1903. Cottidae. Distribution: Lena River and its tributaries, Russia. Habitat: freshwater, brackish.

labradoricus, Acanthocottus Girard [C. F.] in Storer 1850:247, Pl. 7 (fig. 3) [Boston Journal of Natural History v. 6 (no. 2) (art. 15); ref. 18109] Lost. No types known. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1130 [ref. 20662]). •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Neyelov 1973:599 [ref. 7219], Neyelov 1979:122 [ref. 3152]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

lateralis, Scorpaenichthys Girard [C. F.] 1854:145 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 5769] Monterey and San Luis Obispo, California, U.S.A. Syntypes: USNM 328 (1). •Valid as Artedius lateralis (Girard 1854) -- (Bolin 1944:55 [ref. 6379], Eschmeyer & Herald 1983:162 [ref. 9277], Begle 1989:648 [ref. 12739], Matarese et al. 1989:418 [ref. 26885], McAllister 1990:194 [ref. 14674], Mecklenburg et al. 2002:459 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:164 [ref. 35888]). Current status: Valid as Artedius lateralis (Girard 1854). Cottidae. Distribution: Eastern North Pacific: western Gulf of Alaska to northern Baja California. Habitat: marine.

laticeps, Acanthocottus Gilbert [C. H.] 1896:422, Pls. 26-27 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Nushagak River [lectotype] and Herendeen Bay, Alaska, U.S.A. Lectotype. Paralectotypes (incl. potential lectotype): BMNH 1896.4.23.48 (1), MCZ 28218 (1), CAS-SU 3016 (4), USNM 48651 (1), ZMUO J1399 (1) station 3232. Type catalog: Böhlke 1953:126 [ref. 12291], Pethon 1969:7 [ref. 19268]. Lectotype established (as figured specimen) in caption to Pl. 299, p. 3293 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable •Synonym of Megalocottus platycephalus (Pallas 1814) -- (Neyelov 1979:130 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:475 [ref. 25968], Parin et al. 2002:S79 [ref. 28300], Parin et al. 2014:258 [ref. 33547], Mecklenburg et al. 2016:73 [ref. 34440]). Current status: Synonym of Megalocottus platycephalus (Pallas 1814). Cottidae. Habitat: brackish, marine.

latifrons, Cottus Gratzianov [V. I.] 1904:195, Pl. 11 [Zapiski Imperatorskago russkago geograficheskago obshchestva po obshchei geografii v. 41 (no. 1); ref. 21411] Western coast Kanin Peninsula, northern Russia, Barents Sea. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies labradoricus (Girard 1850) -- (Berg 1949:1130 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952], Parin et al. 2002:S89 [ref. 28300]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Neyelov 1973:597 [ref. 7219], Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

latifrons, Stelgidonotus Gilbert [C. H.] & Thompson [J. C.] 1905:977, Fig. 1 [Proceedings of the United States National Museum v. 28 (no. 1414); ref. 1640] Friday Harbor, Washington, U.S.A. Holotype (unique): USNM 53027. •Synonym of Oligocottus rimensis (Greeley 1899) -- (Bolin 1944:63 [ref. 6379], Mecklenburg et al. 2002:499 [ref. 25968]). Current status: Synonym of Oligocottus rimensis (Greeley 1899). Cottidae. Habitat: marine.

latissimus, Cottus Meuschen [F. C.] 1778:42 [Museum Gronovianum; ref. 17682] Original not seen. Not available, published in a rejected work (Opinion 260). Cottidae.

leiopomus, Cottus Gilbert [C. H.] & Evermann [B. W.] 1894:203, Pl. 20 (fig. 2) [Bulletin of the U. S. Fish Commission v. 14 (art. 16) (for 1894); ref. 12377] Big [not Little] Wood River, Shoshone, Idaho, U.S.A. Holotype: USNM 45389. Paratypes: BMNH 1897.1.12.15 (1), CAS-SU 1178 [not 1151 as published] (1). Type catalog: Böhlke 1953:128 [ref. 12291]. •Valid as Cottus leiopomus Gilbert & Evermann 1894 -- (Lee et al. 1980:816 [ref. 22416], Page & Burr 1991:252 [ref. 18983], Nelson et al. 2004:119 [ref. 27807], Meyer et al. 2008:648 [ref. 29820], Page & Burr 2011:483 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus leiopomus Gilbert & Evermann 1894. Cottidae. Distribution: Snake River system, Idaho, U.S.A. Habitat: freshwater.

leprops, Astrocottus Bolin [R. L.] 1936:331, Pl. 34 (fig. B); Fig. 27 [Proceedings of the United States National Museum v. 83 (no. 2987); ref. 504] Tsugaru Strait, Japan, 41°35'50"N, 140°36'45"E, Albatross station 4808, depth 47 fathoms. Holotype (unique): USNM 94730. •Valid as Astrocottus leprops Bolin 1936 -- (Yabe in Masuda et al. 1984:325 [ref. 6441], Shinohara et al. 1996:176 [ref. 26605], Nakabo 2000:632 [ref. 25086], Yabe & Maruyama 2001:61 [ref. 25245], Nakabo 2002:632 [ref. 26001], Tsuruoka et al. 2008:27 [ref. 29568]). Current status: Valid as Astrocottus leprops Bolin 1936. Cottidae. Distribution: Western North Pacific. Habitat: marine.

leptosomus, Porocottus Muto [F.], Choi [Y.] & Yabe [M.] 2002:229, Figs. 1-2 [Ichthyological Research v. 49 (no. 3); ref. 26331] Mohang, Taean, Chungnam, western coast of Korea, 36°47'N, 126°10'E, depth 0.2 meters. Holotype: HUMZ 172394. Paratypes: HUMZ 153700-04 (1, 1, 1, 1, 1), 172395-96 (1, 1), 172399 (1); KNUM 1051-52 (1, 1); NSMT-P 61563 (1). •Valid as Porocottus leptosomus Muto, Choi & Yabe 2002. Current status: Valid as Porocottus leptosomus Muto, Choi & Yabe 2002. Cottidae. Distribution: Yellow Sea, western coast of Korea. Habitat: marine.

lilljeborgii, Cottus Collett [R.] 1875:25, Pl. 1 (figs. 4-5) [Norges fiske; ref. 884] Western Norway. Syntypes: ZMUO J4163 (1) Kristiansund, J4160 (1) Hardangerfjord. Type catalog: Pethon 1969:8 [ref. 19268]. •Valid as Enophrys lilljeborgi [sic] (Collett 1875) -- (Sandercock & Wilimovsky 1968:839 [ref. 26414]). •Valid as Tautulus lilljeborgi [sic] (Collett 1875) -- (Neyelov 1973:600 [ref. 7219]). •Valid as Micrenophrys lilljeborgii [sometimes as lilljiborgi] (Collett 1875) -- (Neyelov 1979:150 [ref. 3152], Fedorov in Whitehead et al. 1986:1252 [ref. 13677], Parin et al. 2002:S80 [ref. 28300], Fricke 2007:29 [ref. 30577], Parin et al. 2014:259 [ref. 33547]). Current status: Valid as Micrenophrys lilljeborgii (Collett 1875). Cottidae. Distribution: Western Baltic Sea, North Sea, eastern North Atlantic, Arctic Ocean. Habitat: brackish, marine.

limbaughi, Icelinus Rosenblatt [R. H.] & Smith [W. L.] 2004:556, Fig. 1 [Copeia 2004 (no. 3); ref. 27912] La Jolla Submarine Canyon, California, U.S.A., 32°53.0'N, 117°15.0'W, depth 40 meters. Holotype: SIO 62-628. Paratypes: AMNH 233608 [ex SIO 58-510] (2, 1 c&s); CAS 218345 [ex SIO 62-634] (1); SIO 51-247 (1), 51-252 (1), 51-253 (2), 54-112 (18), 58-508 (1), 58-509 (2), 62-632 (1), 62-635 (1), 62-673 (1), 63-239-55A (1), 63-1024-55B (1), 74-164 (1); USNM 374474 [ex SIO 62-636] (1). Plus non-type material. •Valid as Icelinus limbaughi Rosenblatt & Smith 2004 -- (Page et al. 2013:123 [ref. 32708], Girard & Smith 2016:556 [ref. 34786]). Current status: Valid as Icelinus limbaughi Rosenblatt & Smith 2004. Cottidae. Distribution: Southern California, U.S.A. Habitat: marine.

lonnbergi, Cottus quadricornis Lönnberg [E.] 1919:167, 171 [Fauna och Flora (Uppsala) v. 14 (no. 5); ref. 23061] Lake Mälaren, Sweden. Syntypes: ?NRM. Name should not be changed since it is Swedish (see Koerber 2009:63 [ref. 30671] for spelling of Lönnberg and origin of his name.). •Validates the infrasubspecific Myoxocephalus quadricornis morpha lonnbergi Berg 1916. •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

lonnbergi, Myoxocephalus quadricornis morpha Berg [L. S.] 1916:431, Fig. 318 [Les Poissons des eaux douces de la Russie; ref. 277] Lake Ladoga, Russia; lake Mälaren, Sweden. No types known. Original as lönnbergi; should not be changed to loennbergi since the name is not German, but Swedish (see Koerber 2009:63 [ref. 30671] for spelling of Lönnberg and origin of his name.). Not available, as a morpha, infrasubspecific; dates to Lönnberg 1919 or to a later publication by Berg. •In the synonymy of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies as described -- (Berg 1949:1133 [ref. 20662]). •In the synonymy of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

lubricus, Paracottus (Leocottus) kessleri Taliev [D. N.] 1955:250, Figs. 97-98 [ ref. 4316] Lake Baikal, Russia. Syntypes: none at ZIN. •Synonym of Cottus kesslerii Dybowski 1874 -- (Sideleva 1982:15 [ref. 14469], Sideleva 1998:151 [ref. 23595]). •Synonym of Leocottus kesslerii (Dybowski 1874) -- (Sideleva 2001:59 [ref. 26181], Sideleva 2003:16, 151 [ref. 27815], Kottelat 2006:60 [ref. 28831]). Current status: Synonym of Leocottus kesslerii (Dybowski 1874). Cottidae. Habitat: freshwater.

lucasi, Ceratocottus Jordan [D. S.] & Gilbert [C. H.] in Jordan & Evermann 1898:1940 [Bulletin of the United States National Museum No. 47; ref. 2444] Near St. Paul Island, Bering Sea (stomach content). Holotype: USNM 48234. Paratypes: CAS-SU 5661 (1), USNM 48235 (1). Type catalog: Böhlke 1953:127 [ref. 12291]. Also appeared in Jordan & Gilbert 1899:459, Pl. 57 [ref. 2478]. •Synonym of Enophrys diceraus (Pallas 1788) -- (Neyelov 1979:146 [ref. 3152], Sheiko & Fedorov 2000:27 [ref. 25650], Parin et al. 2002:S74 [ref. 28300]). •Valid as Enophrys lucasi (Jordan & Gilbert 1898) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414], McAllister 1990:197 [ref. 14674], Mecklenburg et al. 2002:473 [ref. 25968], Nelson et al. 2004:119 [ref. 27807], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:246 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Mecklenburg et al. 2016:55 [ref. 34440], Kells et al. 2016:170 [ref. 35888], Mecklenburg et al. 2018:194 [ref. 36370]). Current status: Valid as Enophrys lucasi (Jordan & Gilbert 1898). Cottidae. Distribution: North Pacific and adjacent Arctic. Habitat: marine.

macellus, Prionistius Bean [T. H.] 1884:355 [Proceedings of the United States National Museum v. 6 (no. 387); ref. 225] Carter Bay, British Columbia, Canada. Holotype: USNM 31958. Paratypes: USNM 33793 (1). •Valid as Triglops macellus (Bean 1884) -- (Eschmeyer & Herald 1983:184 [ref. 9277], McAllister 1990:202 [ref. 14674], Pietsch 1994:347 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:423 [ref. 25968], Parin et al. 2002:S88 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:276 [ref. 33547], Pietsch & Orr 2015:45 [ref. 34694], Kells et al. 2016:192 [ref. 35888]). Current status: Valid as Triglops macellus (Bean 1884). Cottidae. Distribution: North Pacific: northeastern Bering Sea, Alaska, to Aleutian Islands and to Washington (U.S.A.). Habitat: marine.

macouni, Pterygiocottus Bean [B. A.] & Weed [A. C.] 1920:73, Pl. 3 [Transactions of the Royal Society of Canada (Series 3) v. 13 (sect. 5) (1919); ref. 221] Ucluelet, west coast of Vancouver Island, British Columbia, Canada. Holotype (unique): USNM 82169. •Synonym of Artedius harringtoni (Starks 1896) -- (Bolin 1944:45 [ref. 6379]). Current status: Synonym of Artedius harringtoni (Starks 1896). Cottidae. Habitat: marine.

macrops, Cottus Rutter [C. L.] 1908:146, Fig. 4 [Bulletin of the Bureau of Fisheries v. 27 (Doc. 637) (for 1907); ref. 12415] Fall River at Dana, California, U.S.A. Lectotype: USNM 58499. Paralectotypes: (18) CAS-SU 24186 (4); USNM 88464 (2), 126424 (4), 265225 (13). Type catalog: Böhlke 1953:128 [ref. 12291]. Lectotype selected by Daniels & Moyle 1984:951 [ref. 8977]. •Status uncertain -- (Lee et al. 1980:815 [ref. 22416]). •Synonym of Cottus klamathensis Gilbert 1898, but a valid subspecies macrops Rutter 1908 -- (Miller & Taylor 1984:957 [ref. 6809], Moyle 2002:356 [ref. 26449], Scharpf 2008:10 [ref. 30399], Page & Burr 2011:483 [ref. 31215]). Current status: Synonym of Cottus klamathensis Gilbert 1898. Cottidae. Distribution: Pit River system, California, U.S.A. [subspecies macrops]. Habitat: freshwater.

macrostomus, Cottus gobio var. Jeitteles [L. H.] 1863:14 [Jahres-Berichte des Olumtzer K. K. Gymnasium für das Schuljahr 1863; ref. 23055] Original not seen. Treated as infrasubspecific by Kottelat 1997:169 [ref. 22952]. Not available; if regarded as available the name is preoccupied by Cottus poecilopus var. macrostomus Jeitteles 1862. From Czech Republic. •In the synonymy of Cottus gobio Linnaeus 1758 -- (Kottelat 1997:169 [ref. 22952], Freyhof et al. 2005:125 [ref. 28259]). Current status: Synonym of Cottus gobio Linnaeus 1758. Cottidae.

macrostomus, Cottus poecilopus var. Jeitteles [L. H.] 1862:229, Pl. 2 (fig. 2) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 12; ref. 21181] Slovakia. Syntypes: whereabouts unknown. Original not seen. See Kottelat 1997:170 [ref. 22952] for more locality information. •Synonym of Cottus poecilopus Heckel 1839 -- (Kottelat 1997:170 [ref. 22952], Freyhof et al. 2005:167 [ref. 28259]). Current status: Synonym of Cottus poecilopus Heckel 1839. Cottidae. Habitat: freshwater, brackish.

maculatus, Cottus Fischer [J. G.] 1885:78, Pl. 2 (fig. 8) [Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten v. 2; ref. 18084] Reported as Barbados, but that locality in error. Holotype (unique): ZMH H138 [ex ZMH 3523]. Non-types: ZMH H190 [ex 3523] (1). Type catalog: Ladiges et al. 1958:167 [ref. 19181], lkens & Dohse 1993:408 [ref. 21161] with 138 as holotype. Lectotype selected by Ladiges et al. 1958:167 [ref. 19181] but original mentions only one specimen. •Synonym of Enophrys bubalis (Euphrasen 1786) -- (Sandercock & Wilimovsky 1968:842 [ref. 26414]). •Synonym of Taurulus bubalis (Euphrasen 1786) -- (Neyelov 1973:600 [ref. 7219], Neyelov 1979:149 [ref. 3152]). Current status: Synonym of Taurulus bubalis (Euphrasen 1786). Cottidae. Habitat: brackish, marine.

maculosus, Oligocottus Girard [C. F.] 1856:133 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1809] Tomales Bay; San Francisco, California, U.S.A. Syntypes: MCZ 13641 (2); USNM 297 (4, not found in 1980), 298 (6). Oligocottus alaskanus Miles 1918:93 was a lapsus calami for Oligocottus maculosus (Bolin 1944:65 [ref. 6379]). •Valid as Oligocottus maculosus Girard 1856 -- (Bolin 1944:65 [ref. 6379], Eschmeyer & Herald 1983:178 [ref. 9277], Yabe in Masuda et al. 1984:328 [ref. 6441], Matarese et al. 1989:430 [ref. 26885], McAllister 1990:201 [ref. 14674], Nakabo 2000:646 [ref. 25086], Mecklenburg et al. 2002:500 [ref. 25968], Nakabo 2002:646, 1526 [ref. 26001], Parin et al. 2002:S83 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:267 [ref. 33547], Pietsch & Orr 2015:43 [ref. 34694], Kells et al. 2016:186 [ref. 35888]). Current status: Valid as Oligocottus maculosus Girard 1856. Cottidae. Distribution: Eastern North Pacific: southeastern Bering Sea to southern California, U.S.A. Habitat: marine.

major, Procottus jeittelesi Taliev [D. N.] in Berg 1949:1169 [Freshwater fishes of the U.S.S.R. and adjacent countries No. 30; ref. 20662] Lake Baikal, Russia. No types at ZIN. Appeared without distinguishing features and not available in Taliev 1944:34 and Taliev 1948:113 [ref. 19628]. Also appeared in Taliev 1955:309 [ref. 4316] but dates to Taliev in Berg as above. Neotype [ZIN 52082] if designated first by Sideleva 2001:73 [ref. 26181] and 2003:208 [ref. 27815] is invalid. •Synonym of Procottus jeittelesii (Dybowski 1874), but a valid subspecies as described -- (Berg 1949:1169 [ref. 20662], Taliev 1955:309 [ref. 4316]). •Valid as Procottus major Taliev 1949 -- (Sideleva 1982:31 [ref. 14469], Reshetnikov et al. 1997:761 [ref. 24702], Sideleva 1998:158 [ref. 23595], Sideleva 2001:72 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:19, 208 [ref. 27815] dating to Taliev 1944, Bogutskaya & Naseka 2004:200 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Procottus major Taliev 1949. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

mandibularis, Icelus Yabe [M.] in Amaoka et al. 1983:283, Pl. 66; Figs. 28-30 [Fishes from the north-eastern Sea of Japan and the Okhotsk Sea off Hokkaido; ref. 8379] Okhotsk Sea, 44°47.0'N, 144°26.5'E, depth 905 meters. Holotype: HMMZ 78676. Paratypes: HUMZ 78675 (1), 78677-78 (2). •Valid as Icelus mandibularis Yabe 1983 -- (Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:634 [ref. 25086], Nakabo 2002:634 [ref. 26001], Tsuruoka et al. 2006:51 [ref. 28501]). Current status: Valid as Icelus mandibularis Yabe 1983. Cottidae. Distribution: Western North Pacific. Habitat: marine.

marginata, Uranidea Bean [T. H.] 1881:26 [Proceedings of the United States National Museum v. 4 (no. 190); ref. 16046] Tributaries of Walla Walla River, Washington, U.S.A. Syntypes: CAS-SU 1953 (1), USNM 24197 (orig. 9, now 8). Type catalog: Böhlke 1953:133 [ref. 12291]. •Valid as Cottus marginatus (Bean 1881) -- (Lee et al. 1980:817 [ref. 22416], Page & Burr 1991:252 [ref. 18983], Wydoski & Whitney 2003:213 [ref. 27816], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:10 [ref. 30399], Page & Burr 2011:480 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[9] [ref. 33456]). Current status: Valid as Cottus marginatus (Bean 1881). Cottidae. Distribution: Washington and Oregon, U.S.A. Habitat: freshwater.

marmoratus, Centridermichthys Döderlein [L.] in Steindachner & Döderlein 1884:210 [42] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 49 (1. Abth.); ref. 4248] Tokyo, Japan. Holotype (unique): NMW 92827. •Valid as Pseudoblennius marmoratus (Döderlein 1884) -- (Watanabe 1960:150 [ref. 11959] as marumoratus, Nakabo in Masuda et al. 1984:330 [ref. 6441], Kimura et al. 1988:19 [ref. 6692], Kim & Youn 1992:73 [ref. 22287], Nakabo 2000:650 [ref. 25086], Nakabo 2002:650 [ref. 26001], Youn 2002:263, 577 [ref. 26218], Yoshigou 2019:200 [ref. 36714]). Current status: Valid as Pseudoblennius marmoratus (Döderlein 1884). Cottidae. Distribution: Northwestern Pacific: Korea and Japan. Habitat: marine.

marmoratus, Cottus Valenciennes [A.] in Cuvier & Valenciennes 1832:497 [Histoire naturelle des poissons v. 8; ref. 1000] Kamchatka, Russia. Type catalog: Blanc & Hureau 1968:61 [ref. 20738]. •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:265 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Cottidae. Habitat: brackish, marine.

marmoratus, Hemitripterus Ayres [W. O.] 1854:174 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] California, U.S.A. No types known. Appeared first as above on 8 Sept.; then in Ayres 1855:4 [ref. 18923]. Original spelling of the genus by Ayres was Hemitripteras. Authorship of this species was established as Ayres 1854 rather than Girard 1854 in Opinion 1583 in 1990. •Synonym of and junior homonym of Scorpaenichthys marmoratus Girard 1854 -- (Bolin 1944:6 [ref. 6379]). •Valid as Scorpaenichthys marmoratus (Ayres 1854) -- (McAllister 1990:201 [ref. 14674], Mecklenburg 2002:430 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:44 [ref. 34694], Kells et al. 2016:192 [ref. 35888]). Current status: Valid as Scorpaenichthys marmoratus (Ayres 1854). Cottidae. Distribution: Eastern North Pacific: southeastern Alaska to central Baja California. Habitat: marine.

marmoratus, Scorpaenichthys Girard [C. F.] 1854:131 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 1817] San Francisco, California, U.S.A. Syntypes: USNM 314 (2). Scorpaenichthys marmoratus Girard 1854 and Hemitripteras [sic] marmoratus Ayres 1854 were described independently at about the same date. Authorship of this species was fixed as Ayres 1854 and not Girard 1854 in Opinion 1583 in 1990. •Valid as Scorpaenichthys marmoratus Girard 1854 -- (Bolin 1944:6 [ref. 6379], Eschmeyer & Herald 1983:182 [ref. 9277] implied, Matarese et al. 1989:382 [ref. 26885]). •Synonym of Scorpaenichthys marmoratus (Ayres 1854). Current status: Synonym of Scorpaenichthys marmoratus (Ayres 1854). Cottidae. Habitat: marine.

maschalis, Ocynectes Jordan [D. S.] & Starks [E. C.] 1904:307, Fig. 34 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Wakanoura, Wakayama Prefecture, Japan, Inland Sea. Holotype: CAS-SU 7717. Paratypes: USNM 50926 (1). Additional material: CAS-SU 7583 (6), CAS-SU 69836 [ex CAS-SU 7717] (1); USNM 50926. Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Ocynectes maschalis Jordan & Starks 1904 -- (Watanabe 1960:142 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:246 [ref. 15964], Kim & Youn 1992:72 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:647 [ref. 25086], Shinohara et al. 2000:177 [ref. 25987], Nakabo 2002:647 [ref. 26001], Youn 2002:262, 577 [ref. 26218], Parin et al. 2014:266 [ref. 33547]). Current status: Valid as Ocynectes maschalis Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

matsubarae, Astrocottus Katayama [M.] 1942:334, Fig. 1 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 54 (no. 8); ref. 22718] Kakogawa, Hyogo Prefecture, Japan, Inland Sea of Japan. Holotype: NSMT-P 18253. Paratypes: (1, missing). The spelling matsubarae is acceptable; early workers almost certainly Latinized the name first, and then added an 'e' (Art. 31.1.1) [WNE, 12 Mar. 2012] -- we suggest using matsubarae when the species is treated as valid. •Valid as Astrocottus matsubarae Katayama 1942 -- (Watanabe 1960:24 [ref. 11959], Yabe in Masuda et al. 1984:325 [ref. 6441], Nakabo 2000:632 [ref. 25086], Yabe & Maruyama 2001:61 [ref. 25245], Nakabo 2002:632 [ref. 26001], Tsuruoka et al. 2008:29 [ref. 29568]). Current status: Valid as Astrocottus matsubarae Katayama 1942. Cottidae. Distribution: Western North Pacific: Japan. Habitat: brackish, marine.

matsubarai, Myoxocephalus Watanabe [M.] 1958:289, Figs. 117-118, Pl. 20 [Studies of Japanese cottid fishes; ref. 21357] Funka Bay, Hokkaidô, Japan, depth 35 fathoms. Holotype (unique): not researched. Spelled matsubari on plate caption. Also described as new in Watanabe 1960:80, Pl. 26 (fig. 2) [ref. 11959]. •Valid as Myoxocephalus matsubarai Watanabe 1958 -- (Yabe in Masuda et al. 1984:327 [ref. 6441]). •Synonym of Myxocephalus jaok (Cuvier 1829) -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Cottidae. Distribution: Western North Pacific. Habitat: marine.

matuaensis, Microcottus Yabe [M.] & Pietsch [T. W.] 2003:276, Figs. 1-4 [Ichthyological Research v. 50 (no. 3); ref. 27215] Eastern Matua Island, central Kuril Islands, 48°04.2'N, 153°16.1'E, depth 0.5 meters. Holotype: HUMZ 172380. Paratypes: HUMZ 150599-600 (2), 172381-91 (11); UW 47239 (1), NSMT-P 66123 (1). •Valid as Microcottus matuaensis Yabe & Pietsch 2003 -- (Parin et al. 2014:259 [ref. 33547]). Current status: Valid as Microcottus matuaensis Yabe & Pietsch 2003. Cottidae. Distribution: Boreal North Pacific, Kuril Archipelago. Habitat: marine.

meanyi, Ruscarius Jordan [D. S.] & Starks [E. C.] 1895:805, Pl. 80 [Proceedings of the California Academy of Sciences (Series 2) v. 5; ref. 2522] Puget Sound, Washington, U.S.A. Lectotype: CAS-SU 3127 (with syntypes). Paralectotypes: CAS-SU 3127 (2). Type catalog: Böhlke 1953:132 [ref. 12291]. Lectotype established (as figured specimen) in caption to Pl. 285, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. •Valid as Artedius meanyi (Jordan & Starks 1895) -- (Eschmeyer & Herald 1983:163 [ref. 9277], Matarese et al. 1989:384 [ref. 26885], McAllister 1990:194 [ref. 14674]). •Valid as Ruscarius meanyi Jordan & Starks 1895 -- (Begle 1989:648 [ref. 12739], Robins et al. 1991:43 [ref. 14237], Mecklenburg et al. 2002:462 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:44 [ref. 34694], Kells et al. 2016:190 [ref. 35888]). Current status: Valid as Ruscarius meanyi Jordan & Starks 1895. Cottidae. Distribution: Eastern North Pacific: southeastern Alaska to northern California, U.S.A. Habitat: marine.

mednius, Myoxocephalus Bean [T. H.] in Jordan & Evermann 1898:1983 [Bulletin of the United States National Museum No. 47; ref. 2444] Medny Island, Commander Islands, Bering Sea. Holotype (unique): USNM 33863. Author is T. H. Bean. Also appeared as Bean in Jordan & Gilbert 1899:465, Pl. 64b [ref. 2478]. •Questionably a synonym of Porocottus bradfordi Rutter 1898, subspecies albomaculatus (Schmidt 1916) -- (Neyelov 1979:136 [ref. 3152]). •Valid as Porocottus mednius (Bean 1898) -- (Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:486 [ref. 25968], Parin et al. 2002:S84 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:269 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033]). Current status: Valid as Porocottus mednius (Bean 1898). Cottidae. Distribution: North Pacific: Bering Strait to Commander-Aleutian chain to southeastern Alaska and to Kuril Islands. Habitat: marine.

megacephalus, Andriashevicottus Fedorov [V. V.] 1990:746, Fig. [Voprosy Ikhtiologii v. 30 (no. 5); ref. 20136] Simushir Island, Kuril Islands, western Pacific, 46°46'N, 151°51'E, depth 100 meters. Holotype (unique): ZIN 49087. On p. 124 of English translation. •Valid as Andriashevicottus megacephalus Fedorov 1990 -- (Sheiko & Fedorov 2000:26 [ref. 25650], Parin et al. 2002:S68 [ref. 28300], Parin et al. 2014:233 [ref. 33547]). Current status: Valid as Andriashevicottus megacephalus Fedorov 1990. Cottidae. Distribution: Western North Pacific. Habitat: marine.

megacephalus, Chitonotus Lockington [W. N.] 1879:70 [Mining and Scientific Press. An illustrated journal of mining, popular sciences and general news v. 39 (no. 5) 2 Aug.; ref. 20591] San Francisco Bay, California, U.S.A. Syntypes: CAS (3, lost in 1906). Also appeared in Lockington 1881:142 [ref. 2819]. •Synonym of Chitonotus pugetensis (Steindachner 1876). Current status: Synonym of Chitonotus pugetensis (Steindachner 1876). Cottidae. Habitat: marine.

megalops, Antipodocottus DeWitt [H. H.] 1969:30, Fig. 1 [Copeia 1969 (no. 1); ref. 7811] Near Antipodes Islands, New Zealand Region, 49°40'S, 178°53'E, depth 439-540 meters. Holotype (unique): LACM 10991-1. •Valid as Antipodocottus megalops DeWitt 1969 -- (Fricke & Brunken 1984:46 [ref. 5262], Paulin et al. 1989:172 [ref. 24556], Stewart 2015:1120 [ref. 34254]). Current status: Valid as Antipodocottus megalops DeWitt 1969. Cottidae. Distribution: New Zealand region. Habitat: marine.

megalops, Cottocomephorus Pellegrin [J.] 1900:354 [Bulletin du Muséum National d'Histoire Naturelle (Série 1) v. 6 (no. 7); ref. 3398] Angara River, upstream of Irkutsk, Lake Baikal basin, Russia. Holotype (unique): MNHN 1897-0590. Type catalog: Blanc & Hureau 1968:49 [ref. 20738]. •Synonym of Cottocomephorus comephoroides (Berg 1900) -- (Berg 1949:1174 [ref. 20662]). •Synonym of Cottocomephorus inermis (Yakovlev 1890) -- (Taliev 1955:292 [ref. 4316], Bogdanov 2019:230 [ref. 36814]). Current status: Synonym of Cottocomephorus inermis (Yakovlev 1890). Cottidae. Habitat: freshwater.

megalops, Cottus Gratzianov [V. I.] 1902:38 [Dnevnik' Zoologicheskogo Otdelenya Obshchestva Lyubiteley Estestvoznaniya, Antropologii i Etnografii, Moskva v. 3 (no. 3); ref. 1868] Kicherskaya Bay, Lake Baikal, Russia. Holotype (unique): ZIN 9971. Genus Cyphocottus was not available from 2001, two included species after 1930 with no designation of a type species (Art. 13.3); dates to Sideleva 2003:193 [ref. 27815]. •Valid as Asprocottus megalops (Gratzianov 1902) -- (Taliev 1955:329 [ref. 4316]). •Valid as Limnocottus megalops (Gratzianov 1902) -- (Berg 1949:1162 [ref. 20662], Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:157 [ref. 23595]). •Valid as Cyphocottus megalops (Gratzianov 1902) -- (Sideleva 2001:67 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Sideleva 2003:18, 191 [ref. 27815] as Gratzianow, Bogutskaya & Naseka 2004:198 [ref. 28183] as Gratzianov, Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Cyphocottus megalops (Gratzianov 1902). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

meridionalis, Cottus Girard [C. F.] 1850:410 [Proceedings of AAAS (The American Association for the Advancement of Science) 2nd meeting, 1849; ref. 18123] James River, Virginia, U.S.A. No types known. See also Girard 1850:189 [ref. 1814] and Girard 1851:47, Pl. 1 (figs. 9-10) [ref. 18124]. •Synonym of Cottus cognatus Richardson 1837 -- (Smith 1986:358 [ref. 22186]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

meridionalis, Crossias beringi forma Soldatov [V. K.] 1916:220, Pl. 13 (lower) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 21; ref. 18735] Sea of Japan. Lectotype: ZIN 19067. Paralectotypes: ZIN 19067 (3). Treated as infrasubspecific in Eschmeyer et al. 1998; now regarded as subspecific. Muto et al. 2002:231 [ref. 26331] list a lectotype and 3 paralectotypes as ZISP 19067. •Synonym of Porocottus tentaculatus (Kner 1868) -- (Lindberg & Krasyukova 1987:224 [ref. 15964], Parin et al. 2002:S84 [ref. 28300], Parin et al. 2014:270 [ref. 33547], Dyldin & Orlov 2017:67 [ref. 35138]). Current status: Synonym of Porocottus tentaculatus (Kner 1868). Cottidae. Habitat: brackish, marine.

mertensii, Cottus Valenciennes [A.] in Cuvier & Valenciennes 1832:496 [Histoire naturelle des poissons v. 8; ref. 1000] Petropaulski, Kamchatka. Type catalog: Blanc & Hureau 1968:61 [ref. 20738]. •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Neyelov 1979:122 [ref. 3152], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:261 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Cottidae. Habitat: marine.

mesembrinus, Stlengis Fricke [R.] & Brunken [H.] 1983:10, Figs. 1-2 [Japanese Journal of Ichthyology v. 30 (no. 1); ref. 8347] Kai Islands, Indonesia, 5°46'00"S, 132°49'35"E, depth 352 meters. Holotype (unique): ZMUC P81868. •Valid as Antipodocottus mesembrinus (Fricke & Brunken 1983) -- (Fricke & Brunken 1984:46 [ref. 5262], Nelson 1990:845 [ref. 15720], Nelson & Carpenter 1999:2426 [ref. 24809]). Current status: Valid as Antipodocottus mesembrinus (Fricke & Brunken 1983). Cottidae. Distribution: Indonesia. Habitat: marine.

metae, Cottus Freyhof [J.], Kottelat [M.] & Nolte [A.] 2005:156, Figs. 25-26 [Ichthyological Exploration of Freshwaters v. 16 (no. 2); ref. 28259] River Iscica at Ig, 45°57.444'N, 14°32.088'E, Slovenia. Holotype: ZMB 33592. Paratypes: CMK 18551 (10), FSJF 1062 (20). •Valid as Cottus metae Freyhof, Kottelat & Nolte 2005 -- (Kottelat & Freyhof 2007:512 [ref. 29996], Goto et al. 2014:[11] [ref. 33456], Sideleva et al. 2015:430 [ref. 33770], Sideleva et al. 2018:9 [ref. 36393]). Current status: Valid as Cottus metae Freyhof, Kottelat & Nolte 2005. Cottidae. Distribution: Upper Save system, Danube drainage, Slovenia. Habitat: freshwater.

metopias, Triglops Gilbert [C. H.] & Burke [C. V.] 1912:50, Fig. 8 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Petrel Bank, Bering Sea, Albatross station 4777, depth 52 fathoms. Holotype: USNM 74372. Paratypes: CAS-SU 22231 (4); USNM 70851 (2), 70870 (1). Type catalog: Böhlke 1953:133 [ref. 12291]. •Valid as Triglops metopias Gilbert & Burke 1912 -- (Pietsch 1994:365 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:427 [ref. 25968], Parin et al. 2002:S88 [ref. 28300], Nelson et al. 2004:122 [ref. 27807], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:276 [ref. 33547]). Current status: Valid as Triglops metopias Gilbert & Burke 1912. Cottidae. Distribution: North Pacific: Bering Sea, Aleutian Islands to southeastern Alaska. Habitat: marine.

miacanthus, Artediellus Gilbert [C. H.] & Burke [C. V.] 1912:47, Fig. 7 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off east coast of Kamchatka, Russia, 52°46'50"N,148°44'E, Albatross station 4795, depth 48-69 fathoms. Holotype: USNM 74371. Paratypes: CAS-SU 22229 (4), 22243 (4); USNM 74480 (1). Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Artediellus miacanthus Gilbert & Burke 1912 -- (Neyelov 1979:154 [ref. 3152], Nelson 1986:43 [ref. 5808], Lindberg & Krasyukova 1987:231 [ref. 15964], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:495 [ref. 25968], Parin et al. 2002:S70 [ref. 28300], Inagawa & Yabe 2006:196 [ref. 28854], Parin et al. 2014:237 [ref. 33547]). Current status: Valid as Artediellus miacanthus Gilbert & Burke 1912. Cottidae. Distribution: North Pacific: Bering Sea off Alaska to northern Kuril Islands. Habitat: marine.

microcephalus, Cottus gobio var. Kessler [K. F.] 1868:44 [Trudy i Serda Russkich Estestvoispytatelej Prilozenie Suppl.; ref. 23057] Onega District, Russia. No types known. Original not seen. This name is suppressed in favor of koshewnikowi by Freyhof et al. 2005:165 [ref. 28259]; koshewnikowi is in prevailing usage. •Synonym of Cottus poecilopus Heckel 1839 -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Cottus koshewnikowi Gratzianov 1907 -- (Freyhof et al. 2005:163 [ref. 28259], Sideleva et al. 2015:34 [ref. 35276]). Current status: Synonym of Cottus koshewnikowi Gratzianov 1907. Cottidae. Habitat: freshwater, brackish.

microps, Icelinus Starks [E. C.] 1896:552 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 4195] Name only, not available; mentioned in passing under Icelinus strabo Starks 1896. •In the synonymy of Icelinus borealis Gilbert 1896 -- (Bolin 1936:159 [ref. 506]). Current status: Synonym of Icelinus borealis Gilbert 1896. Cottidae.

microstoma, Uranidea Lockington [W. N.] 1880:58 [Proceedings of the United States National Museum v. 3 (no. 121); ref. 17549] St. Paul, Kodiak Island, Alaska, U.S.A. Lectotype: ?USNM 26992. Paralectotypes: (2). Additional material: specimens from Aleutian Is. mentioned in passing. Lectotype established (as figured specimen) in caption to Pl. 293, p. 3292 under Cottus aleuticus in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. Subjectively invalid; secondarily preoccupied in Cottus by Cottus microstomus Heckel 1837; replaced by Cottus philoniips Eigenmann & Eigenmann Lectotype established (as figured specimen) in caption to Pl. 293, p. 3292 under Cottus aleuticus in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. Secondarily preoccupied in Cottus by Cottus microstomus Heckel 1837; replaced by Cottus philoniips Eigenmann & Eigenmann 1892. •Synonym of Cottus aleuticus Gilbert 1896 -- (Scott & Crossman 1998:822 [ref. 24570], Mecklenburg et al. 2002:470 [ref. 25968]). Current status: Synonym of Cottus aleuticus Gilbert 1896. Cottidae. Habitat: freshwater, brackish.

microstomus, Cottus Heckel [J. J.] 1839:147, Pl. 8 (figs. 3-4) [Annalen des Wiener Museums der Naturgeschichte v. 2 (no. 1); ref. 2079] Vicinity of Krakow, Poland. Holotype (unique): NMW 98212. •Synonym of Cottus gobio Linnaeus 1758 -- (Berg 1949:1145 [ref. 20662], Neyelov 1973:593 [ref. 7219], Kottelat 1997:169 [ref. 22952], Sideleva 1998:151 [ref. 23595]). •Valid as Cottus microstomus Heckel 1839 -- (Bogutskaya & Naseka 2004:187 [ref. 28183], Freyhof et al. 2005:160 [ref. 28259], Sideleva 2009:211 [ref. 30173], Goto et al. 2014:[11] [ref. 33456], Sideleva et al. 2018:3 [ref. 36393]). Current status: Valid as Cottus microstomus Heckel 1839. Cottidae. Distribution: Black Sea basin (Dniestr River system) and southern Baltic basin (Vistula River system): Poland and Lithuania. Habitat: freshwater.

microstomus, Cottus poecilopus var. Jeitteles [L. H.] 1862:292, Pl. 2 (fig. 3) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 12; ref. 21181] Creek in Zips, Slovakia. Syntypes: whereabouts unknown. Original not seen. Objectively invalid; preoccupied by Cottus microstomus Heckel 1837. See Kottelat 1997:170 [ref. 22952] for more locality information. •Synonym of Cottus poecilopus Heckel 1839 -- (Kottelat 1997:170 [ref. 22952], Freyhof et al. 2005:167 [ref. 28259]). Current status: Synonym of Cottus poecilopus Heckel 1839. Cottidae. Habitat: freshwater, brackish.

milvensis, Cottus gobio Soldatov [V. K.] 1924:57 [Trudy Severnoi naucho-promyslovoi ekspeditsii / Vysshii sovet narodnogo khoziaistv; ref. 23056] Pechora River, near confluence with Mylva River, Russia. No types known. The species was spelled milvensis and milviensis by Soldatov; Freyhof et al. 2005:166 [ref. 28259] serve as first revisers selecting milvensis. •Synonym of Cottus gobio Linnaeus 1758 -- (Kottelat 1997:169 [ref. 22952] as molvensis). •Synonym of Cottus koshewnikowi Gratzianov 1907 -- (Freyhof et al. 2005:163 [ref. 28259] as milvensis, Sideleva et al. 2015:34 [ref. 35276]). Current status: Synonym of Cottus koshewnikowi Gratzianov 1907. Cottidae. Habitat: freshwater.

minor, Asprocottus korjakovi Sideleva [V. G.] 2001:64 [Proceedings of the Zoological Institute, Academy of Sciences, St.-Petersburg No. 287; ref. 26181] Lake Baikal, Selenga River delta, Russia, depth 250 meters. Holotype: ZIN 49704. Paratypes: ZIN (5). •Synonym of Asprocottus korjakovi Sideleva 2001, but as a valid subspecies as described -- (Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:17, 184 [ref. 27815], Goto et al. 2014:[15] [ref. 33456]). •Valid as Asprocottus minor Sideleva 2001 -- (Bogutskaya & Naseka 2004:195 [ref. 28183]). Current status: Valid as Asprocottus minor Sideleva 2001. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

minor, Cottiusculus Watanabe [M.] 1958:410, Figs. 198-199, Pl. 43 [Studies of Japanese cottid fishes; ref. 21357] Off Funakawa, Akita Prefecture, Japan, depth 70 fathoms. Syntypes: (4). Also described as new in Watanabe 1960:172, Pl. 37 (fig. 2) [ref. 11959]. •Valid as Cottiusculus minor Watanabe 1958 -- (Watanabe 1960:172 [ref. 11959]). •Valid as Artediellus minor (Watanabe 1958) -- (Nakabo 2000:642 [ref. 25086], Nakabo 2002:642, 1526 [ref. 26001], Parin et al. 2014:238 [ref. 33547]). Current status: Valid as Artediellus minor (Watanabe 1958). Cottidae. Distribution: Western North Pacific. Habitat: marine.

minor, Procottus jeittelesi Taliev [D. N.] 1946:91 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 54 (no. 1); ref. 4315] Lake Baikal, Russia, depth 70-200 meters. No types at ZIN. Questionably available, virtually no distinguishing features. •Synonym of Procottus jeittelesii (Dybowski 1874), but a valid subspecies as described -- (Berg 1949:1169 [ref. 20662], Taliev 1955:307 [ref. 4316]). •Synonym of Procottus gurwicii [often as gurwici] (Taliev 1946) -- (Sideleva 1982:27 [ref. 14469], Sideleva 1998:158 [ref. 23595], Sideleva 2001:72 [ref. 26181], Bogutskaya & Naseka 2004:200 [ref. 28183]). Current status: Synonym of Procottus gurwicii (Taliev 1946). Cottidae. Habitat: freshwater.

minutus, Cottus Pallas [P. S.] 1814:145 [Zoographia Rosso-Asiatica v. 3; ref. 3351] Talak Island, Okhotsk Sea. No types known. •Valid as Porocottus minutus (Pallas 1814) -- (Neyelov 1979:137 [ref. 3152], Sheiko & Fedorov 2000:29 [ref. 25650], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S84 [ref. 28300], Fedorov et al. 2003:79 [ref. 27579], Parin et al. 2014:269 [ref. 33547]). Current status: Valid as Porocottus minutus (Pallas 1814). Cottidae. Distribution: Western North Pacific. Habitat: marine.

minutus, Vellitor Iwata [A.] 1983:2, Figs. 1, 2 (A-B), 3 (A, C) [Japanese Journal of Ichthyology v. 30 (no. 1); ref. 5794] Kominato, Chiba Prefecture, Japan, 35°07'N, 140°11'E. Holotype: NSMT-P 19799. Paratypes: (39) BMNH 1981.9.5.13-18 (6); HUMZ 92517-21 (5); MTUF 24661-62 (2), 24663 (1, c&s), 24664-66 (3); NSMT-P 19793-98 (6, 1 ea.); USNM 228073-77 (5), YCM P.2372 (1), P.10010-10015 (6); ZUMT 54267-71 (5). Type catalog: Shibata et al. 2011:13 [ref. 31469]. •Valid as Vellitor minutus Iwata 1983 -- (Yabe in Masuda et al. 1984:329 [ref. 6441], Nakabo 2000:647 [ref. 25086], Nakabo 2002:647 [ref. 26001], Shibata et al. 2011:13 [ref. 31469]). Current status: Valid as Vellitor minutus Iwata 1983. Cottidae. Distribution: Western North Pacific. Habitat: marine.

misakia, Schmidtia Jordan [D. S.] & Starks [E. C.] 1904:237, Fig. 2 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Manazuru Point, Sagami Bay, Japan, Albatross station 3698, depth 153 fathoms. Holotype: USNM 50913. Paratypes: CAS-SU 7506 (1). Type catalog: Böhlke 1953:132 [ref. 12291]. Types: Shinohara & Williams 2006:557 [ref. 31656]. •Valid as Stlengis misakia (Jordan & Starks 1904) -- (Watanabe 1960:35 [ref. 11959], Yamakawa in Okamura et al. 1982:291, 403 [ref. 8256], Yabe in Masuda et al. 1984:325 [ref. 6441], Nakabo 2000:630 [ref. 25086], Shinohara et al. 2001:319 [ref. 25995], Nakabo 2002:630 [ref. 26001]). Current status: Valid as Stlengis misakia (Jordan & Starks 1904). Cottidae. Distribution: Western North Pacific. Habitat: marine.

mitchilli, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:188 [Histoire naturelle des poissons v. 4; ref. 998] New York, U.S.A. •Synonym of Myoxocephalus aenaeus (Mitchill 1814) -- (Scott & Scott 1988:502 [ref. 25518]). Current status: Synonym of Myoxocephalus aenaeus (Mitchill 1814). Cottidae. Habitat: brackish, marine.

modestus, Ocynectes Snyder [J. O.] 1911:539 [Proceedings of the United States National Museum v. 40 (no. 1836); ref. 4152] Same, Mutsu, Japan. Holotype: USNM 68240. Paratypes: CAS-SU 21426 (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Ocynectes modestus Snyder 1911 -- (Watanabe 1960:144 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Shiogaki 1987:222 [ref. 6708], Lindberg & Krasyukova 1987:248 [ref. 15964], Nakabo 2000:647 [ref. 25086], Nakabo 2002:647 [ref. 26001]). Current status: Valid as Ocynectes modestus Snyder 1911. Cottidae. Distribution: Western North Pacific. Habitat: marine.

mororane, Stelgistrum Jordan [D. S.] & Seale [A.] 1906:146, Fig. 4 [Proceedings of the United States National Museum v. 30 (no. 1445); ref. 2496] Mororan harbor, Hokkaido Island, Japan. Holotype (unique): CAS-SU 9260. Type catalog: Böhlke 1953:132 [ref. 12291]. •Synonym of Stelgistrum stejnegeri Jordan & Gilbert 1898 -- (Watanabe 1960:32 [ref. 11959] as morarane, Lindberg & Krasyukova 1987:195 [ref. 15964] as mororanae). •Valid as Icelus mororanis (Jordan & Seale 1906) -- (Tsuruoka et al. 2009:97 [ref. 32107], Parin et al. 2014:255 [ref. 33547]). Current status: Valid as Icelus mororanis (Jordan & Seale 1906). Cottidae. Distribution: Japan and ? Kuril Islands. Habitat: marine.

mucosus, Acanthocottus Ayres [W. O.] 1854:194 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] New England coast, U.S.A. No types known. Appeared first as above on 13 Oct.; then in Ayres 1854:12 [ref. 18923]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Cottidae. Habitat: brackish, marine.

multiradiatus, Batrachocottus nikolskii var. Berg [L. S.] 1907:53, Pl. 2 (fig. 2) [Die Cataphracti des Baikal-Sees; ref. 266] Lake Baikal, Russia, depth 525 meters. Lectotype: ZIN 13750. Paralectotypes: ZIN 13750-52 (1A, 1, 1). Sideleva 2001:57 [ref. 26181] apparently wrongly lists a holotype as ZIN 13750. Lectotype selected by Sideleva 2003:156 [ref. 27815]. •Valid as Batrachocottus multiradiatus Berg 1907 -- (Berg 1949:1166 [ref. 20662], Taliev 1955:270 [ref. 4316], Sideleva 1982:28 [ref. 14469], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:149 [ref. 23595], Sideleva 1999:152 [ref. 24864], Vasil'eva 1999:154 [ref. 24864], Bogutskaya et al. 2001:46 [ref. 26178], Sideleva 2001:57 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:15, 156 [ref. 27815], Bogutskaya & Naseka 2004:197 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Batrachocottus multiradiatus Berg 1907. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

murrayi, Triglops Günther [A.] 1888:209, Pl. 4 (fig. A) [Proceedings of the Royal Society of Edinburgh v. 15; ref. 14482] Northeastern coast of Scotland, depth 64 fathoms. Lectotype: BMNH 1889.6.28.5. Paralectotypes: BMNH 1889.6.28.2-4 (3). Appeared on Pl. 4 as Inglops murrayi. Lectotype selected by Pietsch 1994:368 [ref. 21189] -- the original figured specimen. •Valid as Triglops murrayi Günther 1888 -- (Neyelov 1973:602 [ref. 7219], Fedorov in Whitehead et al. 1986:1257 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:509 [ref. 25518], McAllister 1990:202 [ref. 14674], Pietsch 1994:368 [ref. 21189], Yabe in Okamura et al. 1995:166 [ref. 22531], Klein-MacPhee in Collette & Klein-MacPhee 2002:356 [ref. 26158], Parin et al. 2002:S88 [ref. 28300], Moore et al. 2003:222 [ref. 27076], Nelson et al. 2004:122 [ref. 27807], Fricke 2007:29 [ref. 30577], Møller et al. 2010:53 [ref. 31169], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:276 [ref. 33547], Afonso et al. 2016:51, 289 [ref. 36199], Mecklenburg et al. 2018:225 [ref. 36370]). Current status: Valid as Triglops murrayi Günther 1888. Cottidae. Distribution: North Atlantic anf adjacent Arctic. Habitat: marine.

namiyei, Ceratocottus Jordan [D. S.] & Starks [E. C.] 1904:259, Fig. 13 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Nemuro, Hokkaido, Japan. Holotype: CAS-SU 8108. Paratypes: USNM 50917 (1, dry). Type catalog: Böhlke 1953:127 [ref. 12291]. •Synonym of Ceratocottus diceraus (Pallas 1787) -- (Watanabe 1960:110 [ref. 11959]). •Synonym of Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:836 [ref. 26414], Neyelov 1979:146 [ref. 3152], Kim & Youn 1992:62 [ref. 22287], Parin et al. 2002:S74 [ref. 28300], Parin et al. 2014:245 [ref. 33547]). Current status: Synonym of Enophrys diceraus (Pallas 1787). Cottidae. Habitat: marine.

nasalis, Cottus Berg [L. S.] 1933:701, Figs. 671-673 [Les poissons des eaux douces de l'U.R.S.S. Part 2; ref. 279] Upper Syr-Darya River, Siberia, Russia. Original not seen. •Valid as Cottus nasalis Berg 1933 -- (Berg 1949:1150 [ref. 20662], Goto et al. 2014:[11] [ref. 33456]). Current status: Valid as Cottus nasalis Berg 1933. Cottidae. Distribution: Russia. Habitat: freshwater.

nebulosus, Hemilepidotus Ayres [W. O.] 1854:166 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] Originally as H. ? nebulosus. Appeared first as above on 25 Aug. and is available from that publication; not published in the Proceedings (Ayres 1854 [ref. 18923]; see Ayres 1855:76 [ref. 159]). •Synonym of Artedius notospilotus Girard 1856. Current status: Synonym of Artedius notospilotus Girard 1856. Cottidae. Habitat: marine.

neyelovi, Artediellus Muto [F.], Yabe [M.] & Amaoka [K.] 1994:275, Figs. 1-3 [Japanese Journal of Ichthyology v. 41 (no. 3); ref. 21556] Eastern end of Tsugaru Strait, 41°57'N, 141°08'E, Japan, depth 137 meters. Holotype: HUMZ 103874. Paratypes: HUMZ 93974 (1), 96598-601 (4), 96628 (1), 99540 (1), 100981-88 (8), 100990 (1), 101261-67 (7), 103849 (1), 103876-79 (4), 103881 (1), 103948 (1), 103976 (1), 103990 (1); MNHN 1994-0507 to 0508 (1, 1); NSMT-P 45944-45 (1, 1); USNM 330305-06 (2). •Valid as Artediellus neyelovi Muto, Yabe & Amaoka 1994 -- (Nakabo 2000:641 [ref. 25086], Nakabo 2002:641 [ref. 26001]). Current status: Valid as Artediellus neyelovi Muto, Yabe & Amaoka 1994. Cottidae. Distribution: Western North Pacific. Habitat: marine.

niger, Cottus Bean [T. H.] 1881:151 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Saint Paul Island, Pribilof Group, Bering Sea, Alaska, U.S.A. Lectotype: USNM 27971. Paralectotypes: MCZ 36021 [ex USNM 27952] (1); USNM 23929 (1), 27952 (orig. 12, now 1). Lectotype established (as figured specimen) in caption to Pl. 298, p. 3293 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. Objectively invalid; preoccupied by Cottus niger Lacepède 1801 but we recommend suppressing that name that has not been used in 100 years and its status is uncertain. •Valid as Myoxocephalus niger (Bean 1881) -- (Watanabe 1960:78 [ref. 11959], Neyelov 1979:123 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:213 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:476 [ref. 25968], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:76 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:124 [ref. 32708], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:262 [ref. 33547], Kells et al. 2016:182 [ref. 35888]). Current status: Valid as Myoxocephalus niger (Bean 1881). Cottidae. Distribution: North Pacific: southern Bering Sea to western Gulf of Alaska and to Okhotsk Sea and northern Japan Sea. Habitat: marine.

nigrescens, Porocottus Tanaka [S.] 1908:37 [241], Pl. 9 (figs. 1-2) [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 20; ref. 14360] Southern part of Sakhalin Island, western Okhotsk Sea. Holotype (unique): ZUMT 1387 [= ZUMT 21699]. Appeared first as above in Japanese. Then in English version in Tanaka 1908:241, Pl. 9 (figs. 1-2) [see ref. 14360]. •Valid as Porocottus nigrescens Tanaka 1908 -- (Watanabe 1960:93 [ref. 11959]). •Synonym of Myoxocephalus platycephalus (Pallas 1814), subspecies taeniopterus (Kner 1868) -- (Berg 1949:1128 [ref. 20662]). •Synonym of Megalocottus platycephalus (Pallas 1814), subspecies taeniopterus (Kner 1868) -- (Neyelov 1979:130 [ref. 3152]). •Synonym of Megalocottus platycephalus (Pallas 1814) -- (Parin et al. 2002:S79 [ref. 28300]). •Synonym of Megalocottus taeniopterus (Kner 1868) -- (Parin et al. 2014:258 [ref. 33547], Dyldin & Orlov 2017:64 [ref. 35138] with question). Current status: Synonym of Megalocottus taeniopterus (Kner 1868). Cottidae. Habitat: brackish, marine.

nigricans, Cottus Lesueur [C. A.] in Vaillant 1896:26 [12], Pl. 19 (unedited) [Bulletin de la Société philomathique de Paris (8th Série) v. 8 (no. 1); ref. 13482] No locality. On p. 12 of separate. •Synonym of Myoxocephalus aenaeus (Mitchill 1814). Current status: Synonym of Myoxocephalus aenaeus (Mitchill 1814). Cottidae. Habitat: brackish, marine.

nigricans, Cottus Commerson [P.] in Lacepède 1801:250 [Histoire naturelle des poissons (Lacepède) v. 3; ref. 2710] Appeared as "Cottus nigricans, squamosus, scaber, aculeo obscuro ..." in account of Cottus niger; regarded as non-binominal and not available. See ICZN Opinion 89. Cottidae.

nigripinnis, Artediellus Schmidt [P. Yu.] 1937:571 [Comptes Rendus (Doklady) de l'Académie des Sciences de l'U.R.S.S. v. 15 (no. 9); ref. 15302] Between St. Jona Island (55°40'N, 143°04'E) and western coast of Kamchatka (51°09'N, 154°17'E), Okhotsk Sea, depth 500-592 meters. Lectotype: ZIN 24464 (1 of 3). Paralectotypes: MCZ 34085 [ex ZIN] (1), ZIN (2 of 3). Lectotype selected by Neyelov 1979:160 [ref. 3152]. Appeared first as name only in Schmidt 1935. •Valid as Artediellichthys nigripinnis (Schmidt 1937) -- (Sheiko & Fedorov 2000:26 [ref. 25650], Mecklenburg et al. 2002:489 [ref. 25968], Parin et al. 2002:S69 [ref. 28300], Fedorov et al. 2003:64 [ref. 27579], Parin et al. 2014:234 [ref. 33547]). Current status: Valid as Artediellichthys nigripinnis (Schmidt 1937). Cottidae. Distribution: North Pacific: Bering Sea and Aleutian Islands to eastern Gulf of Alaska and to Okhotsk Sea and Kuril Islands. Habitat: marine.

nihonkaiensis, Cottiusculus Kai [Y.] & Nakabo [T.] 2009:215, Figs. 1-3 [Ichthyological Research v. 56 (no. 3); ref. 30299] Off Tango Peninsula, Kyoto Prefecture, Japan. Holotype: FAKU 130664. Paratypes: (31) at FAKU, NSMT. •Valid as Cottiusculus nihonkaiensis Kai & Nakabo 2009 -- (Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:240 [ref. 33547]). Current status: Valid as Cottiusculus nihonkaiensis Kai & Nakabo 2009. Cottidae. Distribution: Sea of Japan and northern Hokkaido and Korean Peninsula. Habitat: marine.

nikolskii, Cottus Berg [L. S.] 1900:346 [21], Pl. 8 (figs. 1-2) [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 5; ref. 14714] Lake Baikal near Selenga River mouth, Russia, depth 840 feet (120 Russian fathoms). Holotype (unique): ZIN 11676. On p. 21 of separate. •Valid as Batrachocottus nikolskii (Berg 1900) -- (Berg 1949:1165 [ref. 20662], Taliev 1955:266 [ref. 4316], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:150 [ref. 23595], Sideleva 1999:149 [ref. 24864], Vasil'eva 1999:154 [ref. 24864], Bogutskaya et al. 2001:46 [ref. 26178], Sideleva 2001:57 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:15, 158 [ref. 27815], Bogutskaya & Naseka 2004:197 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Batrachocottus nikolskii (Berg 1900). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

nipponicus, Hemitripterus Ishikawa [C.] 1904:14, Pl. 7 (fig. 2) [Proceedings of the Department of Natural History, Tokyo Imperial Museum v. 1; ref. 9847] Hakodate, Oshima Subprefecture, Hokkaido, Japan. Holotype (unique): whereabouts unknown. Cottidae.

nivosus, Cottus Herzenstein [S. M.] 1890:113 [23] [Mélanges Biologiques, tirés du Bulletin physico-mathématique de l'Académie Impériale des Sciences de St. Pétersbourg v. 13; ref. 2149] St. Olga Bay, Russia, Sea of Japan. Holotype (unique): ZIN 8719. On p. 23 of Bull. Acad. Imp. Sci. St. Petersbourg (n. s.) v. 2. •Synonym of Myoxocephalus brandtii (Steindachner 1867) -- (Neyelov 1979:121 [ref. 3152], Lindberg & Krasyukova 1987:210 [ref. 15964] as brandti, Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S80 [ref. 28300], Parin et al. 2014:260 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus brandtii (Steindachner 1867). Cottidae. Habitat: marine.

notospilotus, Artedius Girard [C. F.] 1856:134 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 8; ref. 1809] Tomales Bay, California, U.S.A. Syntypes: USNM 329 (1), 7867 (not found in 1980). •Valid as Artedius notospilotus Girard 1856 -- (Bolin 1944:50 [ref. 6379], Eschmeyer & Herald 1983:163 [ref. 9277], Begle 1989:648 [ref. 12739], Mecklenburg et al. 2002:404 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:164 [ref. 35888]). Current status: Valid as Artedius notospilotus Girard 1856. Cottidae. Distribution: Eastern North Pacific: Puget Sound, Washington, U.S.A. to central Baja California. Habitat: marine.

nozawae, Cottus Snyder [J. O.] 1911:537 [Proceedings of the United States National Museum v. 40 (no. 1836); ref. 4152] Ishikari River at Sapporo, Hokkaido, Japan. Holotype: USNM 68237. Paratypes: CAS-SU 21425 (2); USNM 74550 (1), 74590 (2). Type catalog: Böhlke 1953:128 [ref. 12291]. The name nozawae is the correct original spelling and should not be emended (not excluded by Art. 32); the spelling nozawai has not been established as prevailing usage, so the spelling nozawae is continued. •Synonym of Cottus amblystomopsis Schmidt 1904 -- (Berg 1949:1142 [ref. 20662]). •Synonym of Cottus pollux Günther 1873 -- (Watanabe 1960:119 [ref. 11959], Okada 1961:693 [ref. 6363], Zhang et al. 2016:193 [ref. 34477]). •Valid as Cottus nozawae Snyder 1911 -- (Yabe in Masuda et al. 1984:326 [ref. 6441], Andoh & Goto 1988:176 [ref. 22495], Kawanabe & Mizuno 1989:662 [ref. 25598], Okumura et al. 1996 [ref. 22114], Nakabo 2000:638 [ref. 25086], Goto et al. 2001:69 [ref. 25596], Pietsch et al. 2001:144 [ref. 25688], Bogutskaya et al. 2001:46 [ref. 26178], Yokoyama & Goto 2002:147 [ref. 26160], Nakabo 2002:638 [ref. 26001], Goto et al. 2002:322 [ref. 26495], Bogutskaya & Naseka 2004:187 [ref. 28183], Fujii et al. 2005:12 [ref. 28191], Goto et al. 2014:[9] [ref. 33456], Dyldin & Orlov 2017:63 [ref. 35138]). Current status: Valid as Cottus nozawae Snyder 1911. Cottidae. Distribution: Western North Pacific: Japan and Korea. Habitat: freshwater.

nudithorax, Ricuzenius Bolin [R. L.] 1936:27, Fig. 5 [Proceedings of the United States National Museum v. 84 (no. 3000); ref. 505] Yeso Strait, Japan, 44°04'N, 145°32'E, Albatross station 5031, depth 86 fathoms. Holotype (unique): USNM 102104. •Valid as Ricuzenius nudithorax Bolin 1936 -- (Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:186 [ref. 15964], Nakabo 2000:636 [ref. 25086], Nakabo 2002:636 [ref. 26001], Parin et al. 2002:S85 [ref. 28300], Parin et al. 2014:271 [ref. 33547]). Current status: Valid as Ricuzenius nudithorax Bolin 1936. Cottidae. Distribution: Western North Pacific. Habitat: marine.

nudus, Centridermichthys Döderlein [L.] in Steindachner & Döderlein 1887:258 [2] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 53 (1. abth.); ref. 4249] Tokyo Bay, Japan. Holotype (unique): NMW. •Synonym of Vellitor centropomus (Richardson 1848) -- (Watanabe 1960:165 [ref. 11959]). Current status: Synonym of Vellitor centropomus (Richardson 1848). Cottidae. Habitat: marine.

nudus, Cottus kessleri var. Dybowski [B. N.] 1908:545 [10] [Kosmos Lwów v. 33; ref. 17422] Lake Baikal, Russia. Syntypes: whereabouts unknown. •Synonym of Leocottus kesslerii (Dybowski 1874) -- (Kottelat 2006:60 [ref. 28831]). Current status: Synonym of Leocottus kesslerii (Dybowski 1874). Cottidae. Habitat: freshwater.

nybelini, Triglops Jensen [A. S.] 1944:24, Pl. 3 (figs. a-b) [Spolia Zoologica Musei Hauniensis v. 4; ref. 13060] Northwestern Greenland, depth 490 meters. Lectotype: ZMUC P81232. Paralectotypes: ZMUC P81182-231 (52), P81233-95 (60). Type catalog: Nielsen 1974:81 [ref. 9588]. Lectotype selected by Pietsch 1994:360 [ref. 21189] -- the figured specimen in Jensen 1944, Pl. 3a. •Valid as Triglops nybelini Jensen 1944 -- (Neyelov 1973:602 [ref. 7219], Fedorov in Whitehead et al. 1986:1258 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:510 [ref. 25518], McAllister 1990:202 [ref. 14674], Pietsch 1994:361 [ref. 21189], Yabe in Okamura et al. 1995:167 [ref. 22531], Mecklenburg et al. 2002:426 [ref. 25968], Parin et al. 2002:S88 [ref. 28300], Nelson et al. 2004:122 [ref. 27807], Møller et al. 2010:53 [ref. 31169], Mecklenburg et al. 2011:124 [ref. 31212], Page et al. 2013:125 [ref. 32708], Dolgov 2013:918 [ref. 33112], Mecklenburg et al. 2014:136 [ref. 33463], Parin et al. 2014:277 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:51, 290 [ref. 36199], Mecklenburg et al. 2016:88 [ref. 34440], Mecklenburg et al. 2018:227 [ref. 36370]). Current status: Valid as Triglops nybelini Jensen 1944. Cottidae. Distribution: Arctic Ocean and North Atlantic; nearly circumpolar. Habitat: marine.

obtusus, Phallocottus Schultz [L. P.] 1938:188, Fig. 70 [Proceedings of the United States National Museum v. 85 (no. 3038); ref. 3951] Igitkin Island, Aleutian Islands, Alaska, U.S.A. Holotype: USNM 105280. Paratypes: USNM 105281 (5). •Valid as Phallocottus obtusus Schultz 1938 -- (Sheiko & Fedorov 2000:28 [ref. 25650], Yabe et al. 2001:70 [ref. 25246], Mecklenburg et al. 2002:496 [ref. 25968], Parin et al. 2002:S83 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:267 [ref. 33547]). Current status: Valid as Phallocottus obtusus Schultz 1938. Cottidae. Distribution: Eastern North Pacific: Aleutian Islands, Alaska and Kuril Islands. Habitat: marine.

occidentalis, Gymnacanthus tricuspis Schmidt [P. Yu.] 1927:29 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 15821] Barents Sea and White Sea. Syntypes: ZIN 6370 (2), 7083 (1), 10446 (1), 11262 (1), 20380 (1). •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Parin et al. 2002:S75 [ref. 28300] as Gymnacanthus, Parin et al. 2014:249 [ref. 33547], Mecklenburg et al. 2016:62 [ref. 34440]). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Cottidae. Habitat: marine.

ocellatus, Acanthocottus Storer [H. R.] 1850:253 [Boston Journal of Natural History v. 6 (no. 2) (art. 15); ref. 18109] Entrance to the Gut of Canso, Burial Island in Wilmot Harbor, Nova Scotia, Canada. Holotype (unique): whereabouts unknown. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Cottidae. Habitat: brackish, marine.

ochotensis, Artediellus Gilbert [C. H.] & Burke [C. V.] 1912:44, Fig. 5 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off western coast of Kamchatka, Russia, 51°37'N, Albatross station 4798, depth 25 fathoms. Holotype: USNM 22271 (?74369). Paratypes: CAS-SU 68870 (3), USNM 70910 (3). •Valid as Artediellus ochotensis Gilbert & Burke 1912 -- (Neyelov 1979:154 [ref. 3152], Yabe in Masuda et al. 1984:328 [ref. 6441], Nelson 1986:43 [ref. 5808], Lindberg & Krasyukova 1987:233 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:641 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:494 [ref. 25968], Nakabo 2002:641 [ref. 26001], Parin et al. 2002:S71 [ref. 28300], Fedorov et al. 2003:66 [ref. 27579], Nelson et al. 2004:118 [ref. 27807], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:121 [ref. 32709], Shinohara et al. 2014:248 [ref. 33330], Parin et al. 2014:238 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Dyldin & Orlov 2017:62 [ref. 35138]). Current status: Valid as Artediellus ochotensis Gilbert & Burke 1912. Cottidae. Distribution: North Pacific and adjacent Arctic: Chukchi Sea off Alaska to Okhotsk Sea and Japan Sea to Peter the Great Bay. Habitat: brackish, marine.

ochotensis, Icelus Schmidt [P. Yu.] 1927:4, Figs. 1-2 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 19815] Northern Okhotsk Sea, between 58°48.5'N to 53°56'N, 151°18.5'-137°32'E, depth 60-105 meters. Syntypes: (45) ZIN 21898-99 (1, 2), 21901-04 (3, 6, 6, 6). •Valid as Icelus ochotensis Schmidt 1927 -- (Nelson 1984:50 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Kim & Youn 1992:67 [ref. 22287], Suzuki & Uno 1992:4 [ref. 24139], Murai et al. 1994:343 [ref. 21908], Nakabo 2000:635 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Nakabo 2002:635 [ref. 26001], Youn 2002:262, 576 [ref. 26218], Parin et al. 2002:S77 [ref. 28300], Fedorov et al. 2003:71 [ref. 27579], Parin et al. 2014:255 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331]). Current status: Valid as Icelus ochotensis Schmidt 1927. Cottidae. Distribution: Western North Pacific. Habitat: marine.

ochotensis, Myoxocephalus niger var. Taranetz [A. Ya.] 1937:115 [Izvestiia Tikhookeanskogo nauchnogo instituta rybnogo khoziaistva [Bulletins of the Pacific Science Institute] v. 11; ref. 13384] Holotype: ZIN 25468. Objectively invalid; preoccupied by Myoxocephalus verrucosus subspecies ochotensis Schmidt 1929. •Synonym of Myoxocephalus aeneus (Mitchill 1814) -- (Neyelov 1979:123 [ref. 3152]). •Synonym of Myoxocephalus niger (Bean 1881) -- (Lindberg & Krasyukova 1987:213 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Parin et al. 2014:262 [ref. 33547]). Current status: Synonym of Myoxocephalus niger (Bean 1881). Cottidae. Habitat: marine.

ochotensis, Myoxocephalus verrucosus var. Schmidt [P. Yu.] 1929:417, Fig. 3 [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 30 (no. 3); ref. 16059] Sea of Okhotsk. •Valid as Myoxocephalus ochotensis Schmidt 1929 -- (Neyelov 1979:122 [ref. 3152], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:76 [ref. 27579], Parin et al. 2014:262 [ref. 33547], Mecklenburg et al. 2016:85 [ref. 34440]). Current status: Valid as Myoxocephalus ochotensis Schmidt 1929. Cottidae. Distribution: Western North Pacific. Habitat: marine.

ochotensis, Porocottus sellaris Schmidt [P. Yu.] 1929:411, Figs. 1-2 [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 30 (no. 3); ref. 16059] Penshinskaya Guba [Penzhinskii Bay], Shantar Islands, northern Okhotsk Sea. Syntypes: ZIN 21934-39 (3, 6, 4, 1, 1, 2). •Synonym of Microcottus sellaris (Gilbert 1896) -- (Neyelov 1979:135 [ref. 3152], Lindberg & Krasyukova 1987:219 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S80 [ref. 28300], Parin et al. 2014:260 [ref. 33547]). Current status: Synonym of Microcottus sellaris (Gilbert 1896). Cottidae. Habitat: brackish, marine.

ochotensis, Zesticelus Yabe [M.] 1995:17, Figs. 1-3 [Japanese Journal of Ichthyology v. 42 (no. 1); ref. 22666] Southwestern Okhotsk Sea off Cape Shiretoko, Japan, 44°27.2'N, 145°06.9'E, depth 1450 meters. Holotype: HUMZ 126133. Paratypes: HUMZ 121626 (1); NSMT-P 46437 (1). •Valid as Zesticelus ochotensis Yabe 1995 -- (Shinohara et al. 1996:176 [ref. 26605], Nakabo 2000:640 [ref. 25086], Sheiko & Fedorov 2000:29 [ref. 25650], Nakabo 2002:640 [ref. 26001], Parin et al. 2002:S90 [ref. 28300], Parin et al. 2014:279 [ref. 33547]). Current status: Valid as Zesticelus ochotensis Yabe 1995. Cottidae. Distribution: Western North Pacific. Habitat: marine.

octodecemspinosus, Cottus Mitchill [S. L.] 1814:7 [Report, in part, of Samuel L. Mitchill; ref. 3030] New York, U.S.A. No types known. Also in Mitchill 1815:380 [ref. 13292]. Originally as 18-spinosus; correction mandatory; as octodecem-spinosus in Mitchill 1815:380 [ref. 13292]. •Valid as Myoxocephalus octodecemspinosus (Mitchill 1814) -- (Neyelov 1979:123 [ref. 3152], Smith 1986:450 [ref. 22186], Robins & Ray 1986:282 [ref. 23100] as octodecimspinosus, Scott & Scott 1988:502 [ref. 25518], McAllister 1990:200 [ref. 14674], Murdy et al. 1997:147 [ref. 23144], Klein-MacPhee in Collette & Klein-MacPhee 2002:351 [ref. 26158], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Afonso et al. 2016:46, 226 [ref. 36199]). Current status: Valid as Myoxocephalus octodecemspinosus (Mitchill 1814). Cottidae. Distribution: Northwestern Atlantic. Habitat: brackish, marine.

oculatus, Icelinus Gilbert [C. H.] 1890:87 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., 32°44'30"N, 117°23'W, Albatross station 2935, depth 124 fathoms. Holotype (unique): CAS-SU 80. •Valid as Icelinus oculatus Gilbert 1890 -- (Bolin 1944:19 [ref. 6379], Peden 1984:67 [ref. 5193], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Girard & Smith 2016:556 [ref. 34786]). Current status: Valid as Icelinus oculatus Gilbert 1890. Cottidae. Distribution: Eastern North Pacific: southern California. Habitat: marine.

oernensis, Cottus quadricornis Lönnberg [E.] 1932:16, Figs. 6 a-b [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Örn, Sweden. Syntypes: (9) ?BMNH 1932.5.17.1 (1), NRM 20323-25 (1, 5, 2). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

ohmiensis, Cottus Watanabe [M.] 1960:123, Fig. 45, Pl. 30 (fig. 2) [Fauna Japonica: Cottidae (Pisces); ref. 11959] Chinai River, Takashima-gun, and Ado River, Takashima-gun, Shiga Prefecture; and Tama River, Nishitama-gun, Tôkyô-to, Japan. Syntypes: (46). Cottidae. Habitat: freshwater.

okadai, Porocottus Watanabe [M.] 1958:301, Figs. 126, 127, Pl. 22 [Studies of Japanese cottid fishes; ref. 21357] Off Date, Hokkaido, Japan, depth 30 fathoms. Holotype (unique): not researched. Also described as new in Watanabe 1960:90, Pl. 27 (fig. 2) [ref. 11959]. •Valid as Porocottus okadai Watanabe 1958 -- (Watanabe 1960:90 [ref. 11959]). •Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus polyacanthocephalus (Pallas 1814). Cottidae. Habitat: marine.

okiensis, Alcichthys Mori [T.] 1956:15, Fig. 1 [Japanese Journal of Ichthyology v. 5 (nos 1-2); ref. 12213] Off Oki Island coast, Japan Sea. Holotype (unique): Hyogo Univ. Agric. •Valid as Alcichthys okiensis Mori 1956 -- (Lindberg & Krasyukova 1987:243 [ref. 15964]). •Synonym of Alcichthys elongatus (Steindachner 1881) -- (Nakabo 2002:1526 [ref. 26001], Parin et al. 2002:S68 [ref. 28300], Parin et al. 2014:233 [ref. 33547]). Current status: Synonym of Alcichthys elongatus (Steindachner 1881). Cottidae. Habitat: marine.

ommatistius, Triglops Gilbert [C. H.] 1913:465, Pl. 64 [Proceedings of the United States National Museum v. 44 (no. 1963); ref. 14957] Gloucester, Massachusetts, U.S.A. Holotype: USNM 73921. Paratypes: CAS-SU 23064 (1); USNM 43067-68 (2, 1), 43954 (1), 45547 (6). Type catalog: Böhlke 1953:133 [ref. 12291]. •Synonym of Triglops murrayi Günther 1888, but a valid subspecies -- (Fedorov in Whitehead et al. 1986:1258 [ref. 13677]). •Synonym of Triglops murrayi Günther 1888 -- (Scott & Scott 1988:510 [ref. 25518], Pietsch 1994:368 [ref. 21189], Parin et al. 2014:276 [ref. 33547]). Current status: Synonym of Triglops murrayi Günther 1888. Cottidae. Habitat: marine.

omodakae, Icelus Tanaka [S.] 1915:616 [Dobutsugaku Zasshi = Zoological Magazine Tokyo v. 27; ref. 15023] 50 miles off Hinomisaki, Japan, depth 120 fathoms. Syntypes: (3) ZUMT 6420 (1, lost). •Synonym of Icelus cataphractus (Pavlenko 1910) -- (Nelson 1984:28 [ref. 5391]). Current status: Synonym of Icelus cataphractus (Pavlenko 1910). Cottidae. Habitat: marine.

onegensis, Myoxocephalus quadricornis Berg [L. S.] & Popov [A. A.] 1932:158 [Doklady Akademii Nauk SSSR 1932; ref. 16359] Onega Lake and Petrozavodsk, Onega Lake, Russia. Syntypes: ZIN 3842 (1), 9026a (1). Based on Berg 1916:432, fig. 319 and on specimens. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

ontariensis, Triglopsis Jordan [D. S.] & Thompson [W. F.] 1910:75, Fig. 1 [Proceedings of the United States National Museum v. 38 (no. 1728); ref. 10651] Lake Ontario (deep water), off Toronto, Canada. Holotype (unique): USNM 64599. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Scott & Crossman 1998:846 [ref. 24570]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

onychus, Cottus Eigenmann [C. H.] & Eigenmann [R. S.] 1892:963 [American Naturalist v. 26 (no. 311); ref. 12339] Bow River at Calgary, tributary of South Saskatchewan, Alberta, Canada. Holotype (unique): BMNH 1892.12.30.234. •Synonym of Cottus ricei (Nelson 1876) -- (Scott & Crossman 1998:842 [ref. 24570]). Current status: Synonym of Cottus ricei (Nelson 1876). Cottidae. Habitat: freshwater.

orientalis, Gymnacanthus tricuspis Schmidt [P. Yu.] 1927:29 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 15821] Kara Sea to Bering Sea. Syntypes: ZIN 14234-35 (1, 2), 20383 (3), 20689-90 (1, 2), 21761-74 (3, 1, 1, 6, 1, 1, 1, 1, 3, 5, 3, 1, 2, 2). •Synonym of Gymnocanthus tricuspis (Reinhardt 1930) -- (Sheiko & Fedorov 2000:27 [ref. 25650] as Gymnacanthus, Parin et al. 2002:S75 [ref. 28300] as Gymnacanthus, Parin et al. 2014:249 [ref. 33547], Mecklenburg et al. 2016:62 [ref. 34440]). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Cottidae. Habitat: marine.

osensis, Stlengis Jordan [D. S.] & Starks [E. C.] 1904:236, Fig. 1 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Ose Point, Suruga Bay, Japan, Albatross station 3788, depth 167 fathoms. Holotype (unique): USNM 50912. •Valid as Stlengis osensis Jordan & Starks 1904 -- (Watanabe 1960:40 [ref. 11959], Fricke & Brunken 1984:46 [ref. 5262], Yabe in Masuda et al. 1984:325 [ref. 6441], Nakabo 2000:630 [ref. 25086], Shinohara et al. 2001:319 [ref. 25995], Nakabo 2002:630 [ref. 26001]). Current status: Valid as Stlengis osensis Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

osimae, Furcina Jordan [D. S.] & Starks [E. C.] 1904:305, Fig. 33 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Hakodate, Oshima Subprefecture, Hokkaido, Japan. Holotype: CAS-SU 7716. Paratypes: USNM 50925 (1). Additional material: BMNH 1905.2.4.364 (1), CAS 58440 [ex IU 10724] (1), CAS-SU 7584 (1). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Furcina osimae Jordan & Starks 1904 -- (Watanabe 1960:139 [ref. 11959], Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:246 [ref. 15964], Kim & Youn 1992:72 [ref. 22287] as osihmae, Nakabo 2000:648 [ref. 25086], Shinohara et al. 2000:177 [ref. 25987], Nakabo 2002:648 [ref. 26001], Youn 2002:262, 574 [ref. 26218], Girard & Smith 2016:556 [ref. 34786]). Current status: Valid as Furcina osimae Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

oyamai, Astrocottus Watanabe [M.] 1958:221, Figs. 73-74, Pl. 4 [Studies of Japanese cottid fishes; ref. 21357] Misaki, Kanagawa Perf., Japan, depth 70 fathoms. Holotype (unique): apparently lost. Also described as new in Watanabe 1960:27, pl. 19 (fig. 1) [ref. 11959]. See Tsuruoka et al. 21008:36 [ref. 29568] for comments on discrepancies in the original description. •Valid as Astrocottus oyamai Watanabe 1958 -- (Yabe in Masuda et al. 1984:325 [ref. 6441], Nakabo 2000:632 [ref. 25086], Nakabo 2002:632 [ref. 26001]). •Nomen dubium, not in Astrocottus -- (Tsuruoka et al. 2008:36 [ref. 29568]). Cottidae. Distribution: Western North Pacific: Japan [if valid]. Habitat: marine.

pachypus, Cottus Günther [A.] 1860:161 [Catalogue of the fishes in the British Museum v. 2; ref. 1963] Port Leopold. Syntypes: (2) BMNH 1855.9.19.48 (1). Cottidae.

pachytus, Batrachocottus baicalensis Taliev [D. N.] 1955:262, Figs. 22B, 104 [ ref. 4316] Near settlement of B. Koty, Lake Baikal, Russia, depth 80-120 meters. Holotype: ZIN (lost). Appeared without distinguishing features in Taliev 1948:107, 113 [ref. 19628]. •Synonym of Batrachocottus baicalensis (Dybowski 1874) -- (Sideleva 1982:28 [ref. 14469], Sideleva 1998:149 [ref. 23595], Sideleva 1999:149 [ref. 24864], Sideleva 2001:57 [ref. 26181], Sideleva 2003:15 [ref. 27815]). Current status: Synonym of Batrachocottus baicalensis (Dybowski 1874). Cottidae. Habitat: freshwater.

pacificus, Artediellus Gilbert [C. H.] 1896:416 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bristol Bay; south of Sannak Island; north of Unalaska Island, Alaska. Syntypes: BMNH 1896.7.23.55-58 (4?), 1900.9.11.3 (1); MCZ 28328 (12); NMW 6832 (3); CAS-SU 2602 (5), 2604-05 (29, 41); USNM 48605 (1), 48657-58 (6, 3), 48664 (2), 48757 (1), 59410 (5, now 1); ZMUO J1403 (12), J1404 (10). Type catalog: Böhlke 1953:127 [ref. 12291], Pethon 1969:7 [ref. 19268]. •Valid as Artediellus pacificus Gilbert 1896 -- (Neyelov 1979:154 [ref. 3152], Eschmeyer & Herald 1983:161 [ref. 9277], Nelson 1986:43 [ref. 5808], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:492 [ref. 25968], Parin et al. 2002:S71 [ref. 28300], Nelson et al. 2004:118 [ref. 27807], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:121 [ref. 32708], Parin et al. 2014:238 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033]). Current status: Valid as Artediellus pacificus Gilbert 1896. Cottidae. Distribution: North Pacific and adjacent Arctic: northern Bering Sea to southeastern Alaska and to Okhotsk Sea and northern Japan Sea. Habitat: marine.

pacificus, Neohemilepidotus Watanabe [M.] 1958:15, Fig. 9 [Studies of Japanese cottid fishes; ref. 21357] A mistake for Hemilepidotus japonicus (Sakamoto 1932) and not a new species description (Peden 1977:41 [ref. 24080]); see Hemilepidotus japonicus Sakamoto 1932. •In the synonymy of Hemilepidotus papilio (Bean 1880) -- (Peden 1979:41 [ref. 24080], Nakabo 2002:1525 [ref. 26001] with wrong author). Current status: Synonym of Hemilepidotus papilio (Bean 1880). Cottidae.

pacificus, Triglops pingelii Schmidt [P. Yu.] 1930:515, Fig. 1a [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 30 (no. 4) (for 1929); ref. 16144] Okhotsk and Bering seas. Syntypes: (11) ZIN 21462-63 (4, 4), 21468 (1) •Synonym of Triglops pingelii Reinhardt 1837 -- (Pietsch 1994:374 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Parin et al. 2002:S88 [ref. 28300], Parin et al. 2014:277 [ref. 33547]). Current status: Synonym of Triglops pingelii Reinhardt 1837. Cottidae. Habitat: marine.

pallidus, Limnocottus Taliev [D. N.] 1948:113, 128 table [Trudy Baikal'skoi Limnologicheskoi Stantsii v. 12; ref. 19628] Lake Baikal, Russia. No types at ZIN. Can date to above reference based on morphometrics presented in the table. Also appeared as new in Taliev 1955:86, 121, 148, 200, 357 [ref. 4316] as Abyssocottus pallidus. Neotype [ZIN 13744] apparently designated by Sideleva but not in conformance with the fourth edition of the Code and is unavailable. •Valid as Limnocottus pallidus Taliev 1948 -- (Sideleva 1982:30 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:157 [ref. 23595], Sideleva 2001:68 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:18, 198 [ref. 27815], Bogutskaya & Naseka 2004:199 [ref. 28183] dated 1955, Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Limnocottus pallidus Taliev 1948. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

paltschevskii, Cottus (Pegedictis) Schmidt [P. Yu.] 1916:614, Fig. 2 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 3935] Sakhabe River, tributary to Ternay Bay, Tatar Strait, Russia. Syntypes: ZIN 18959 (3). Original spelling should be studied. •Synonym of Cottus czerskii Berg 1913 -- (Berg 1949:1140 [ref. 20662], Sideleva 1998:151 [ref. 23595], Parin et al. 2002:S73 [ref. 28300] as palischevskii, Parin et al. 2014:242 [ref. 33547]). Current status: Synonym of Cottus czerskii Berg 1913. Cottidae. Habitat: freshwater.

papilio, Melletes Bean [T. H.] 1880:354 [Proceedings of the United States National Museum v. 2 (no. 100); ref. 222] Saint Paul Island, Pribilof Group, Bering Sea, Alaska, U.S.A. Holotype (unique): USNM 23751. •Valid as Melletes papilio Bean 1880 -- (Lindberg & Krasyukova 1987:183 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S79 [ref. 28300], Tokranov et al. 2003:300 [ref. 26972], Fedorov et al. 2003:74 [ref. 27579]). •Valid as Hemilepidotus (Melletes) papilio (Bean 1880) -- (Mecklenburg et al. 2007:176 [ref. 29448]). •Valid as Hemilepidotus papilio (Bean 1880) -- (Peden 1979:41 [ref. 24080], Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:631 [ref. 25086], Mecklenburg et al. 2002:431 [ref. 25968], Nakabo 2002:631 [ref. 26001], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:251 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Tohkairin et al. 2015:7 [ref. 35331], Mecklenburg et al. 2016:65 [ref. 34440], Kells et al. 2016:174 [ref. 35888], Dyldin & Orlov 2017:64 [ref. 35138], Mecklenburg et al. 2018:203 [ref. 36370]). Current status: Valid as Hemilepidotus papilio (Bean 1880). Cottidae. Distribution: North Pacific and adjacent Arctic Region. Habitat: brackish, marine.

parmiferus, Asprocottus herzensteini Taliev [D. N.] 1955:194, 319, Figs. 43, 69, 127 [ ref. 4316] Lake Baikal, Russia. Taliev specimens: ZIN 33012 (1), 33017 (1), 34301 (5). Sideleva 2001:67 [ref. 26181] lists a neotype [ZIN 49702] but that is invalid under the code. This is again treated as neotype in Sideleva 2003:185 [ref. 27815]. •Valid as Asprocottus parmiferus Taliev 1955 -- (Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:156 [ref. 23595], Sideleva 2001:67 [ref. 26181], Bogutskaya et al. 2001:47 [ref. 26178], Sideleva 2003:17, 185 [ref. 27815], Bogutskaya & Naseka 2004:195 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Asprocottus parmiferus Taliev 1955. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

parvulus, Myoxocephalus Gilbert [C. H.] & Burke [C. V.] 1912:59, Fig. 12 [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Tidepool at Preobrazhenskoi Bay, Medny Island, Commander Islands, Bering Sea. Holotype: USNM 74375. Paratypes: CAS-SU 22369 (1), USNM 70829 (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Synonym of Microcottus sellaris (Gilbert 1896) -- (Neyelov 1979:135 [ref. 3152], Lindberg & Krasyukova 1987:219 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:483 [ref. 25968], Parin et al. 2002:S80 [ref. 28300], Parin et al. 2014:260 [ref. 33547], Mecklenburg et al. 2016:75 [ref. 34440]). Current status: Synonym of Microcottus sellaris (Gilbert 1896). Cottidae. Habitat: brackish, marine.

parvus, Cottopsis Girard [C. F.] 1854:144 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 7; ref. 5769] Presidio, San Francisco, California, U.S.A. Syntypes: USNM 294 (2, not found in 1980). •Synonym of Cottus asper Richardson 1837 -- (Scott & Crossman 1998:825 [ref. 24570]). Current status: Synonym of Cottus asper Richardson 1837. Cottidae. Habitat: freshwater, brackish, marine.

patris, Acanthocottus Storer [H. R.] 1850:250, Pl. 7 (fig. 2) [Boston Journal of Natural History v. 6 (no. 2) (art. 15); ref. 18109] Great Mecatina Island Harbor and Red Bay, Straits of Belle Isle, Labrador, Canada, depth 10-20 fathoms. Syntypes: (2). •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Neyelov 1973:595 [ref. 7219] as Gymnacanthus). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Cottidae. Habitat: marine.

paulus, Cottus Williams [J. D.] 2000:302 [Copeia 2000 (no. 1); ref. 24258] Coldwater Spring, 5.7 miles west of Oxford about 500 yards north of U.S. highway 78, tributary of Coosa River, Calhoun County, Alabama, U.S.A. Holotype: UMMZ 187462. Paratypes: ALA 2031 (12), 1138 (39), 1665 (68); CUMV 52064 [ex ALA 1063] (26); TU 34396 (47); UMMZ 187463 (25); USNM 201404 [ex ALA 1063] (26). Replacement name for Cottus pygmaeus Williams 1968, preoccupied by Cottus quadricornis pygmaeus Lönnberg 1932. •Valid as Cottus paulus Williams 2000 -- (Kinziger et al. 2000:1007 [ref. 25019], Nelson et al. 2004:119 [ref. 27807], Boschung & Mayden 2004:424 [ref. 27995], Neely et al. 2007:651 [ref. 29262], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:476 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus paulus Williams 2000. Cottidae. Distribution: Coldwater Spring, Coosa River system, Alabama, U.S.A. Habitat: freshwater.

pelagicus, Paracottus Taliev [D. N.] 1955:48 et seq., 252, Figs. 99-100 [ ref. 4316] Near town of Slyudyanka, Lake Baikal, Russia, depth 4-40 meters. Syntypes: ZIN (not found in 1996). As Paracottus (Leocottus) pelagicus on p. 252. •Valid as Paracottus pelagicus Taliev 1955 -- (Taliev 1955:114 [ref. 4316]). •Synonym of Paracottus knerii (Dybowski 1874) -- (Sideleva 1982:16 [ref. 14469]). •Synonym of Cottus kesslerii Dybowski 1874 -- (Sideleva 1998:152 [ref. 23595]). •Synonym of Leocottus kesslerii (Dybowski 1874) -- (Sideleva 2001:59 [ref. 26181], Sideleva 2003:16, 151 [ref. 27815]). Current status: Synonym of Leocottus kesslerii (Dybowski 1874). Cottidae. Habitat: freshwater.

pellegrini, Cottus gobio natio Vladykov [V. D.] 1931:353, Fig. 28 [Mémoires de la Société Zoologique de France v. 29 (no. 4); ref. 16343] Type catalog: Blanc & Hureau 1968:51 [ref. 20738] with MNHN 1930-220 to 223 (4) as syntypes; Sanda et al. 2010:30 [ref. 31086] with paralectotypes and a lectotype designated by Freyhof et al. 2005 [MNHN 1930-222].. Not available, infrasubspecific; from Tisza basin, Ukraine. Validly published by Bacescu & Nacescu-Mester 1964. •In the synonymy of Cottus gobio Linnaeus 1758 -- (Berg 1949:1331 [ref. 20662]). Current status: Synonym of Cottus gobio Linnaeus 1758. Cottidae.

pellegrini, Cottus gobio Bǎcescu [M. C.] & Bǎcescu-Meşter [L.] 1964:442 [Vie et Milieu Suppl. No. 17; ref. 1238] Carpathes range in Ukraine, Danube basin, Teresva, Ukraine. Lectotype: MNHN. Syntypes: CUP; MNHN 1930-0220 to 0223 (4, niow 3); many at NMP. Type catalog: Mlíkovský et al. 2011:154 [ref. 33498]. Lectotype apparently designated by Freyhof et al. 2005. Available as a subspecies by indication to infrasubspecific treatment by Vladykov 1931. •Synonym of Cottus gobio Linnaeus 1758 -- (Kottelat 1997:169 [ref. 22952], Freyhof et al. 2005:125 [ref. 28259]). Current status: Synonym of Cottus gobio Linnaeus 1758. Cottidae. Habitat: freshwater, brackish.

percoides, Pseudoblennius Günther [A.] 1861:297 [Catalogue of the fishes in the British Museum v. 3; ref. 1964] Omura, near Nagasaki, Japan. No types known. Based on Temminck & Schlegel 1844: Pl. 79A (figs. 2-3) [ref. 4372]. See Nakabo 2002:1526 [ref. 26001]. •Valid as Pseudoblennius percoides Günther 1861 -- (Watanabe 1960:147 [ref. 11959], Nakabo in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:251 [ref. 15964], Kimura et al. 1988:19 [ref. 6692], Kim & Youn 1992:74 [ref. 22287], Mizuno et al. 1999:8 [ref. 25311], Nakabo 2000:650 [ref. 25086], Shinohara et al. 2000:177 [ref. 25987], Nakabo 2002:650 [ref. 26001], Youn 2002:263, 577 [ref. 26218]). Current status: Valid as Pseudoblennius percoides Günther 1861. Cottidae. Distribution: Western North Pacific. Habitat: marine.

perifretum, Cottus Freyhof [J.], Kottelat [M.] & Nolte [A.] 2005:144, Figs. 18-20 [Ichthyological Exploration of Freshwaters v. 16 (no. 2); ref. 28259] Stream Zwaanebeek, 51°14'44N, 4°34'58'E, Province Flandern, Belgium. Holotype: ZMB 33588. Paratypes: FSJF 1383 (15). Plus non-type material. Previously misspelled perifrenatum. •Valid as Cottus perifretum Freyhof, Kottelat & Nolte 2005 -- (Kottelat & Freyhof 2007:513 [ref. 29996], Sideleva 2009:210 [ref. 30173], Goto et al. 2014:[11] [ref. 33456], Sideleva et al. 2015:33 [ref. 35276], Sideleva et al. 2018:9 [ref. 36393]). Current status: Valid as Cottus perifretum Freyhof, Kottelat & Nolte 2005. Cottidae. Distribution: Northern Europe. Habitat: freshwater.

perminovi, Icelus Taranetz [A. Ya.] 1936:151 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (pt 13) (No. 3, art. 107); ref. 6509] Okhotsk Sea, 56°12'N, 143°49'E. Holotype (unique): No types at ZIN. •Valid as Icelus perminovi Taranetz 1936 -- (Nelson 1984:34 [ref. 5391], Yabe in Masuda et al. 1984:324 [ref. 6441], Nakabo 2000:634 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Nakabo 2002:634 [ref. 26001], Parin et al. 2002:S78 [ref. 28300], Fedorov et al. 2003:72 [ref. 27579], Parin et al. 2014:255 [ref. 33547]). Current status: Valid as Icelus perminovi Taranetz 1936. Cottidae. Distribution: Western North Pacific. Habitat: marine.

perplexus, Cottus Gilbert [C. H.] & Evermann [B. W.] 1894:202, Pl. 20 (fig. 1) [Bulletin of the U. S. Fish Commission v. 14 (art. 16) (for 1894); ref. 12377] Shookumchuck River, near Chehalis, Washington, U.S.A. Holotype: USNM 45387. Paratypes: (12) BMNH 1897.1.12.13 (1), 1900.9.29.197 (1); CAS-SU 1324 (17), ?1343 (missing, wrong no.); NMW 11458 (1); USNM 45388 (24). Type catalog: Böhlke 1953:128 [ref. 12291]. •Valid as Cottus perplexus Gilbert & Evermann 1894 -- (Lee et al. 1980:818 [ref. 22416], Page & Burr 1991:253 [ref. 18983], Moyle 2002:354 [ref. 26449], Wydoski & Whitney 2003:212 [ref. 27816], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:481 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[9] [ref. 33456]). Current status: Valid as Cottus perplexus Gilbert & Evermann 1894. Cottidae. Distribution: Northwestern U.S.A.: Washington, Oregon and California. Habitat: freshwater.

petiti, Cottus Bǎcescu [M. C.] & Bǎcescu-Meşter [L.] 1964:432, Figs. 1-4; Pls. 1-2 [Vie et Milieu Suppl. No. 17; ref. 1238] Sources of river Lez, France. Holotype: MGAB 50. Paratypes: ?BMNH 1964.12.23.10 [from Bacescu] (1); MGAB 51 (1) + uncat. (15). See Lelek 1987:274 [ref. 22962]. •Valid as Cottus petiti Bǎcescu & Bǎcescu-Meşter -- (Kottelat 1997:170 [ref. 22952], Freyhof et al. 2005:142 [ref. 28259], Kottelat & Freyhof 2007:515 [ref. 29996], Sideleva 2009:211 [ref. 30173], Goto et al. 2014:[11] [ref. 33456], Sideleva et al. 2015:33 [ref. 35276], Sideleva et al. 2018:9 [ref. 36393]). Current status: Valid as Cottus petiti Bǎcescu & Bǎcescu-Meşter 1964. Cottidae. Distribution: France. Habitat: freshwater.

philonips, Cottus Eigenmann [C. H.] & Eigenmann [R. S.] 1892:963 [American Naturalist v. 26 (no. 311); ref. 12339] Kicking Horse River, Field, British Columbia, Canada. Syntypes: BMNH 1892.12.30.235-244 (10), 1892.12.20.245 (1); CAS-SU 527 (1); USNM 125483 (1). Type catalog: Böhlke 1953:128 [ref. 12291]. Text somewhat unclear; tentatively proposed for specimens that "are probably to be referred to the description quoted above"; quoted above are Cottus minutus Pallas and Uranidea microstomus Lockington 1880 (the latter noted as preoccupied in Cottus by Cottus microstomus Heckel 1837). We regard as an independent proposal of a new name (not a replacement name); however, Hubbs & Schultz 1932 state, "..we rule as first revisers that Cottus philonips was proposed as a substitute name for Cottus minutus Pallas and not for Uranidea microstoma Lockington." Under the circumstances, action of a first reviser is probably inappropriate; minutus was not in fact preoccupied. •Synonym of Cottus cognatus Richardson 1837 -- (Lee et al. 1980:808 [ref. 22416], Scott & Crossman 1998:835 [ref. 24570]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

pietschi, Icelinus Yabe [M.], Soma [A.] & Amaoka [K.] 2001:65, Figs. 1-3 [Ichthyological Research v. 48 (no. 1); ref. 25246] Tide pool on east side of Chirip Peninsula, Okhotsk coast of Iturup Island, 45°19.8'N, 148°00.3'E, depth 0.5-1 meters. Holotype: HUMZ 151942. Paratypes: HUMZ 151943-44 (1, 1), NSMT-P 58459 (1), UW 43755 (1). •Valid as Icelinus pietschi Yabe, Soma & Amaoka 2001 -- (Rosenblatt & Smith 2004:559 [ref. 27912]). Current status: Valid as Icelinus pietschi Yabe, Soma & Amaoka 2001. Cottidae. Distribution: Western North Pacific. Habitat: marine.

pietschmanni, Triglops pingelii Jensen [A. S.] 1944:15, 29, Pls. 1 (figs. a-b), 2 (fig. a) [Spolia Zoologica Musei Hauniensis v. 4; ref. 13060] Southwestern Greenland. Syntypes: (19) ZMUC P81371 (1), P81374-385 (12), P81482 (1), P81477 (1). Type catalog: Nielsen 1974:82 [ref. 9588]. •Synonym of Triglops murrayi Günther 1888 -- (Neyelov 1973:602 [ref. 7219], Pietsch 1994:369 [ref. 21189], Parin et al. 2014:276 [ref. 33547]). Current status: Synonym of Triglops murrayi Günther 1888. Cottidae. Habitat: marine.

pinetorum, Ricuzenius Jordan [D. S.] & Starks [E. C.] 1904:243, Fig. 5 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Kinkwazan Island, in Matsushima Bay, Japan, Albatross station 3773. Holotype (unique): USNM 50915. •Valid as Ricuzenius pinetorum Jordan & Starks 1904 -- (Watanabe 1960:19 [ref. 11959], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:185 [ref. 15964], Kim & Youn 1992:69 [ref. 22287], Shinohara et al. 1996:176 [ref. 26605], Nakabo 2000:632 [ref. 25086], Nakabo 2002:632 [ref. 26001], Youn 2002:263, 578 [ref. 26218], Shinohara et al. 2014:251 [ref. 33330]). Current status: Valid as Ricuzenius pinetorum Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

pingelii, Triglops Reinhardt [J. C. H.] 1837:114, 118 [32, 36] [Det Kongelige Danske videnskabernes selskabs naturvidenskabelige og mathematiske afhandlinger v. 7; ref. 3691] Quanneoen, south of Frederikshaab, western Greenland. Lectotype: ZMUC 1. Paralectotypes: ZMB 672 [ex ZMUC] (1), ZMUC 2-3 (2). Type catalog: Nielsen 1974:82 [ref. 9588], Paepke & Fricke 1992:284 [ref. 19946]. On pp. 32 & 36 of separate. Described but not named in 1830:17 [ref. 19480]. Lectotype established by Pietsch 1994:374 [ref. 21189]. •Valid as Triglops pingeli Reinhardt 1837, subspecies pingeli Reinhardt 1837 -- (Nakabo 2000:628 [ref. 25086], Nakabo 2002:628 [ref. 26001]). •Valid as Triglops pingelii Reinhardt 1837 [often as pingeli] -- (Neyelov 1973:601 [ref. 7219], Eschmeyer & Herald 1983:184 [ref. 9277], Yabe in Masuda et al. 1984:325 [ref. 6441], Fedorov in Whitehead et al. 1986:1259 [ref. 13677], Robins & Ray 1986:284 [ref. 23100], Lindberg & Krasyukova 1987:174 [ref. 15964], Scott & Scott 1988:511 [ref. 25518], McAllister 1990:202 [ref. 14674], Kim & Youn 1992:65 [ref. 22287], Pietsch 1994:374 [ref. 21189], Yabe in Okamura et al. 1995:168 [ref. 22531], Sokolovskaya et al. 1998:10 [ref. 24670], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:428 [ref. 25968], Youn 2002:263, 578 [ref. 26218] as pingeli, Parin et al. 2002:S88 [ref. 28300], Fedorov et al. 2003:82 [ref. 27579], Nelson et al. 2004:122 [ref. 27807], Pietsch & Orr 2006:244 [ref. 28666], Mecklenburg et al. 2007:177 [ref. 29448], Møller et al. 2010:53 [ref. 31169], Shinohara et al. 2011:44 [ref. 31715], Page et al. 2013:125 [ref. 32708], Dolgov 2013:918 [ref. 33112], Shinohara et al. 2014:251 [ref. 33330], Mecklenburg et al. 2014:137, 145 [ref. 33463], Parin et al. 2014:277 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Pietsch & Orr 2015:45 [ref. 34694], Tohkairin et al. 2015:7 [ref. 35331], Afonso et al. 2016:52, 291 [ref. 36199], Kells et al. 2016:194 [ref. 35888], Mecklenburg et al. 2016:90 [ref. 34440], Mecklenburg et al. 2018:229 [ref. 36370]). Current status: Valid as Triglops pingelii Reinhardt 1837. Cottidae. Distribution: Circumpolar in Arctic Ocean, adjacent North Pacific and North Atlantic. Habitat: marine.

pistilliger, Cottus Pallas [P. S.] 1814:143, Pl. 20 (figs. 3-4) [Zoographia Rosso-Asiatica v. 3; ref. 3351] Unalaska Island and Port Avatsch. Syntypes: ZMB 8213 (1). Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. Type information: Svetovidov 1978:28 [ref. 6025]. •Valid as Gymnocanthus pistilliger (Pallas 1814) -- (Watanabe 1960:56 [ref. 11959], Yabe in Masuda et al. 1984:328 [ref. 6441], Lindberg & Krasyukova 1987:199 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Sokolovskaya et al. 1998:9 [ref. 24670], Nakabo 2000:629 [ref. 25086], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Mecklenburg et al. 2002:465 [ref. 25968], Nakabo 2002:629 [ref. 26001], Youn 2002:262, 575 [ref. 26218], Parin et al. 2002:S75 [ref. 28300] as Gymnacanthus, Fedorov et al. 2003:69 [ref. 27579] as Gymnacanthus, Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Shinohara et al. 2014:249 [ref. 33330], Parin et al. 2014:248 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Mecklenburg et al. 2016:59 [ref. 34440], Kells et al. 2016:172 [ref. 35888], Dyldin & Orlov 2017:63 [ref. 35138], Balanov & Matveev 2018:366 [430] [ref. 36463], Mecklenburg et al. 2018:197 [ref. 36370]). Current status: Valid as Gymnocanthus pistilliger (Pallas 1814). Cottidae. Distribution: North Pacific and adjacent Arctic. Habitat: brackish, marine.

pitensis, Cottus Bailey [R. M.] & Bond [C. E.] 1963:20, Figs. 1d, 3b, 4d [Occasional Papers of the Museum of Zoology University of Michigan No. 634; ref. 3934] North Fork of Pit River, east of Alturas, Modoc County, California, U.S.A. Holotype: UMMZ 130558. Paratypes: UMMZ 130559 (6). Additional material at CAS 25963 (37), OSU, and UMMZ [ex CAS 25963] (4). •Valid as Cottus pitensis Bailey & Bond 1963 -- (Lee et al. 1980:819 [ref. 22416], Page & Burr 1991:253 [ref. 18983], Moyle 2002:353 [ref. 26449], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:482 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus pitensis Bailey & Bond 1963. Cottidae. Distribution: Western U.S.A.: Oregon and California. Habitat: freshwater.

platycephalus, Asprocottus herzensteini Taliev [D. N.] 1955:324, Figs. 130-131 [ ref. 4316] Lake Baikal, Russia. Neotype: ZIN 46863. Taliev specimens: ZIN 33038 (2). Appeared as Asprocottus herzensteini brevis, name only, in Taliev 1948:113 [ref. 4316]. •Valid as Asprocottus platycephalus Taliev 1955 [usually dated to 1948] -- (Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702] dating to 1948, Sideleva 1998:156 [ref. 23595], Bogutskaya et al. 2001:47 [ref. 26178], Sideleva 2001:67 [ref. 26181], Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:17, 187 [ref. 27815], Bogutskaya & Naseka 2004:196 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Asprocottus platycephalus Taliev 1955. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

platycephalus, Cottus Pallas [P. S.] 1814:135 [Zoographia Rosso-Asiatica v. 3; ref. 3351] Kamchatka, Russia; America; Okhotsk Sea. Lectotype: ZMB 4756. Syntypes: ZMB 4756 (1, dry). Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. Lectotype designated by Svetovidov 1978:27 [ref. 6025]. •Valid as Myoxocephalus platycephalus (Pallas 1814) -- (Berg 1949:1127 [ref. 20662]). •Valid as Megalocottus platycephalus (Pallas 1814) -- (Neyelov 1979:130 [ref. 3152], Pietsch et al. 2001:145 [ref. 25688], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:152 [ref. 23595], Sheiko & Fedorov 2000:28 [ref. 25650], Zemnukhov et al. 2001:420 [ref. 25412], Mecklenburg et al. 2002:475 [ref. 25968], Parin et al. 2002:S79 [ref. 28300], Fedorov et al. 2003:73 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Bogutskaya & Naseka 2004:190 [ref. 28183], Mecklenburg et al. 2007:183 [ref. 29448], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:257 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Mecklenburg et al. 2016:72 [ref. 34440], Kells et al. 2016:182 [ref. 35888], Mecklenburg et al. 2018:209 [ref. 36370]). Current status: Valid as Megalocottus platycephalus (Pallas 1814). Cottidae. Distribution: North Pacific and Arctic region. Habitat: brackish, marine.

pleurostictus, Triglops Cope [E. D.] 1865:81 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 17; ref. 907] Godhaven, western Greenland. Holotype (unique): ANSP (not found). Type catalog: Böhlke 1984:164 [ref. 13621]. •Synonym of Triglops pingelii Reinhardt 1837 -- (Neyelov 1973:601 [ref. 7219], Pietsch 1994:374 [ref. 21189]). Current status: Synonym of Triglops pingelii Reinhardt 1837. Cottidae. Habitat: marine.

plumarius, Archistes Jordan [D. S.] & Gilbert [C. H.] in Jordan & Evermann 1898:1900 [Bulletin of the United States National Museum No. 47; ref. 2444] Ushishir Island, Kurils [Kuril] Islands. Holotype (unique): USNM 48233. Also in Jordan & Gilbert 1899:454, Pl. 53 [ref. 2478]. •Valid as Archistes plumarius Jordan & Gilbert 1898 -- (Sheiko & Fedorov 2000:26 [ref. 25650], Mecklenburg et al. 2002:442 [ref. 25968], Parin et al. 2002:S68 [ref. 28300], Parin et al. 2014:234 [ref. 33547]). Current status: Valid as Archistes plumarius Jordan & Gilbert 1898. Cottidae. Distribution: Western North Pacific. Habitat: marine.

poecilopus, Cottus Heckel [J. J.] 1839:145, Pl. 8 (figs. 1-2) [Annalen des Wiener Museums der Naturgeschichte v. 2 (no. 1); ref. 2079] Amur mountain brooks of Carpathians at Grosschagendorf near Kaesnark in Upper Hungary [now Slovakia]. Syntypes: NMW 6424 (1), 6729 (3), 78816 (2). See Kottelat 1997:170 [ref. 22952] for locality information. •Valid as Cottus poecilopus [sometimes as pocilopus] Heckel 1839 -- (Berg 1949:1143 [ref. 20662], Sideleva 1982:81 [ref. 14469], Fedorov in Whitehead et al. 1986:1248 [ref. 13677], Lelek 1987:278 [ref. 22962], Kim & Youn 1992:60 [ref. 22287], Zhu 1995:189 [ref. 25213], Chereshnev 1996:606 [ref. 24659], Kottelat 1997:170 [ref. 22952], Kim 1997:402 [ref. 24032], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:152 [ref. 23595], Sheiko & Fedorov 2000:27 [ref. 25650] as cf. poecilopus, Bogutskaya et al. 2001:46 [ref. 26178], Chereshnev et al. 2001:119 [ref. 26736], Pietsch et al. 2001:144 [ref. 25688], Kim & Park 2002:323 [ref. 25913], Youn 2002:262, 573 [ref. 26218], Parin et al. 2002:S73 [ref. 28300], Fedorov et al. 2003:67 [ref. 27579] as cf. poecilopus, Jang et al. 2003:120 [ref. 27681], Hanel 2003:61 [ref. 27894], Bogutskaya & Naseka 2004:188 [ref. 28183] as cf. poecilopus, Knaack 2003:17 [ref. 28219], Fujii et al. 2005:12 [ref. 28191], Freyhof et al. 2005:167 [ref. 28259], Hanel & Lusk 2005:341 [ref. 28804], Kottelat 2006:91 [ref. 28831], Fricke 2007:26 [ref. 30577], Shedko & Miroshnichenko 2007:24 [ref. 29369], Kottelat & Freyhof 2007:515 [ref. 29996], Sideleva & Goto 2009:599 [ref. 30591], Goto et al. 2014:[9] [ref. 33456], Parin et al. 2014:244 [ref. 33547], Sideleva et al. 2015:430 [ref. 33770], Zhang et al. 2016:193 [ref. 34477] dated 1836). Current status: Valid as Cottus poecilopus Heckel 1839. Cottidae. Distribution: Northern Europe. Habitat: freshwater, brackish.

polaris, Cottus Sabine [E.] 1824:213 [ccxiii] [A supplement to the appendix of Captain Parry's voyage; ref. 18649] North Georgia, east side Boothia Peninsula, Northwest Territories, Canada. No types known. •Synonym of Triglopsis quadricornis (Linnaeus 1758), but a valid subspecies -- (Chereshnev 1996:606 [ref. 24659]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (Lee et al. 1980:826 [ref. 22416], Parin et al. 2014:263 [ref. 33547]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

pollicaris, Uranidea Jordan [D. S.] & Gilbert [C. H.] 1882:222 [Proceedings of the United States National Museum v. 5 (no. 277); ref. 18349] Lake Michigan off Racine, Wisconsin, U.S.A. Holotype (unique): USNM 29663. •Synonym of Cottus ricei (Nelson 1876) -- (Scott & Crossman 1998:842 [ref. 24570]). Current status: Synonym of Cottus ricei (Nelson 1876). Cottidae. Habitat: freshwater.

pollux, Cottus Günther [A.] 1873:240 [Annals and Magazine of Natural History (Series 4) v. 12 (no. 69) (art. 31); ref. 2004] River at Otarranai, Japan. Syntypes: BMNH 1873.4.30.1-2 (2). See also Nakabo 2002:1525 [ref. 26001]. •Valid as Cottus pollux Günther 1873 -- (Watanabe 1960:119 [ref. 11959], Okada 1961:693 [ref. 6363], Yabe in Masuda et al. 1984:326 [ref. 6441], Kawanabe & Mizuno 1989:666 [ref. 25598], Zhu 1995:189 [ref. 25213], Nakabo 2000:638 [ref. 25086], Matsuura et al. 2000:207 [ref. 26000], Nakabo 2002:638 [ref. 26001], Goto et al. 2002:318 [ref. 26495], Hagiwara 2003:37 [ref. 27188], Fujii et al. 2005:12 [ref. 28191], Goto et al. 2014:[9] [ref. 33456], Zhang et al. 2016:193 [ref. 34477]). Current status: Valid as Cottus pollux Günther 1873. Cottidae. Distribution: Japan and ? China. Habitat: freshwater.

polyacanthocephalus, Cottus Pallas [P. S.] 1814:133 [Zoographia Rosso-Asiatica v. 3; ref. 3351] No definite locality [= Kodiak, Aleutian Islands]. Holotype (unique): ZMB 4757 (dry). Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. Type information: Svetovidov 1978:27 [ref. 6025]. •Valid as Myoxocephalus polyacanthocephalus (Pallas 1814) -- (Watanabe 1960:66 [ref. 11959], Neyelov 1979:121 [ref. 3152], Eschmeyer & Herald 1983:176 [ref. 9277], Yabe in Masuda et al. 1984:326 [ref. 6441], Lindberg & Krasyukova 1987:210 [ref. 15964], McAllister 1990:200 [ref. 14674], Sokolovsky & Sokolovskaya 1997:57 [ref. 24691], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:645 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:482 [ref. 25968], Nakabo 2002:645 [ref. 26001], Youn 2002:262, 576 [ref. 26218], Parin et al. 2002:S81 [ref. 28300], Fedorov et al. 2003:77 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Mecklenburg et al. 2007:176 [ref. 29448], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:261 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Pietsch & Orr 2015:43 [ref. 34694], Tohkairin et al. 2015:7 [ref. 35331], Mecklenburg et al. 2016:78 [ref. 34440], Kells et al. 2016:184 [ref. 35888], Dyldin & Orlov 2017:66 [ref. 35138], Mecklenburg et al. 2018:215 [ref. 36370]). Current status: Valid as Myoxocephalus polyacanthocephalus (Pallas 1814). Cottidae. Distribution: North Pacific and adjacent Arctic. Habitat: marine.

polyporus, Cottus klamathensis Daniels [R. A.] & Moyle [P. B.] 1984:958 [Copeia 1984 (no. 4); ref. 8977] Klamath River, Siskiyou County, California, U.S.A. Holotype: NYSM 11451. Paratypes: NYSM 11452 (1). Based on 40 specimens; holotype and 1 paratype listed on p. 951; additional specimens are therefore regarded as additional non-type material. •Synonym of Cottus klamathensis Gilbert 1898, but a valid subspecies polyporus Daniels & Moyle as described -- (Moyle 2002:356 [ref. 26449], Scharpf 2008:10 [ref. 30399], Page & Burr 2011:483 [ref. 31215]). Current status: Synonym of Cottus klamathensis Gilbert 1898. Cottidae. Distribution: Klamath River and tributaries, Oregon and California, U.S.A. [subspecies polyporus]. Habitat: freshwater.

porosus, Cottus Valenciennes [A.] in Cuvier & Valenciennes 1832:498 [Histoire naturelle des poissons v. 8; ref. 1000] Baffin Bay, between western Greenland and Baffin Island. Holotype (unique): MNHN 0000-6761. Type catalog: Blanc & Hureau 1968:51 [ref. 20738]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Cottidae. Habitat: brackish, marine.

princeps, Cottus Gilbert [C. H.] 1898:12, Fig. [Bulletin of the U. S. Fish Commission v. 17 (art. 1) [1897]; ref. 12349] Upper Klamath Lake, Oregon, U.S.A. Holotype: USNM 48227. Paratypes: (numerous). •Valid as Cottus princeps Gilbert 1898 -- (Lee et al. 1980:820 [ref. 22416], Page & Burr 1991:248 [ref. 18983], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:472 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus princeps Gilbert 1898. Cottidae. Distribution: Oregon, U.S.A. Habitat: freshwater.

produndulus, Procottus jeittelesi Taliev [D. N.] 1948:113 [Trudy Baikal'skoi Limnologicheskoi Stantsii v. 12; ref. 19628] Not available, appeared as manuscript name in table; apparently not subsequently used by Taliev. Cottidae.

profundorum, Acanthocottus Gilbert [C. H.] 1896:423, Pl. 27 (upper) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bering Sea, north of Unalaska Island, Albatross station 3329, depth 399 fathoms. Syntypes (incl. potential lectotype): USNM 48589 (1), 53030 (1). Lectotype established (as figured specimen) in caption to Pl. 300, p. 3293 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable. •Valid as Zesticelus profundorum (Gilbert 1896) -- (Bolin 1944:95 [ref. 6379], Neyelov 1979:161 [ref. 3152], Eschmeyer & Herald 1983:184 [ref. 9277], Yabe & Okamura 1993:370 [ref. 22281], Yabe 1995:19 [ref. 22666], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:487 [ref. 25968], Parin et al. 2002:S90 [ref. 28300], Fedorov et al. 2003:83 [ref. 27579], Parin et al. 2014:279 [ref. 33547]). Current status: Valid as Zesticelus profundorum (Gilbert 1896). Cottidae. Distribution: Bering Sea and North Pacific. Habitat: marine.

protrusus, Cottus Schultz [L. P.] & Spoor [W. A.] 1933:142 [Copeia 1933 (no. 3); ref. 16522] Small lake on Unalaska Island, about 400 feet above Makushin Bay, Aleutian Islands. Holotype: USNM 104496 [ex UW 2334]. Paratypes: (5) UMMZ 64234 (1). •Synonym of Cottus aleuticus Gilbert 1896 -- (Lee et al. 1980:801 [ref. 22416], Scott & Crossman 1998:820 [ref. 24570], Mecklenburg et al. 2002:470 [ref. 25968]). Current status: Synonym of Cottus aleuticus Gilbert 1896. Cottidae. Habitat: freshwater, brackish.

pugetensis, Artedius Steindachner [F.] 1876:181 [133], Pl. 14 (fig. 2-2a) [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 74 (1. Abth.); ref. 4225] Fox Island near near Steilacom, northern Puget Sound; also San Francisco. Syntypes: (several) NMW 6795-96 (2, 3), 11662-66 (4, 2, 2, 2, 1), 11668 (1), 14783-84 (1, 1), 59720 (?), 75544 (1). On p. 133 of separate. •Valid as Chitonotus pugetensis (Steindachner 1876) -- (Bolin 1944:19 [ref. 6379], Eschmeyer & Herald 1983:164 [ref. 9277], Matarese et al. 1989:388 [ref. 26885], McAllister 1990:196 [ref. 14674], Mecklenburg et al. 2002:437 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32709], Pietsch & Orr 2015:39 [ref. 34694], Kells et al. 2016:166 [ref. 35888]). Current status: Valid as Chitonotus pugetensis (Steindachner 1876). Cottidae. Distribution: Eastern North Pacific: Alaska, U.S.A. to southern Baja California. Habitat: marine.

pulcher, Asprocottus Taliev [D. N.] 1955:69 et seq., 327, Figs. 23, 134-135 [ ref. 4316] Lake Baikal, Russia. Syntypes: not found at ZIN. Appeared without distinguishing features and unavailable as Cottinella pulchra in Taliev 1948:113, 118 [ref. 19628]. Sideleva 2001:67 [ref. 26181] lists a neotype [ZIN 50873] but that is invalid under the code if this is the first time it appeared; same neotype listed in Sideleva 2003:188 [ref. 27815]. •Valid as Asprocottus pulcher Taliev 1955 [usually dated to 1948] -- (Taliev 1955:69 [ref. 4316], Sideleva 1982:29 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702] dating to 1948, Sideleva 1998:156 [ref. 23595], Bogutskaya et al. 2001:47 [ref. 26178] dating to 1948, Kontula et al. 2003:149 [ref. 26791], Sideleva 2003:17, 188 [ref. 27815], Bogutskaya & Naseka 2004:196 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Asprocottus pulcher Taliev 1955. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

punctulatus, Potamocottus Gill [T. N.] 1861:40 [Proceedings of the Boston Society of Natural History v. 8 (1861-1862); ref. 1785] Bridger Pass, Wyoming, U.S.A. Holotype (unique): ?USNM 3251 (not found in 1980). •Synonym of Cottus bairdii Girard 1850 -- (Baxter & Stone 1995:221 [ref. 23374]). •Synonym of Cottus bairdii Girard 1850, but a valid subspecies punctulatus (Gill 1861) -- (Scharpf 2008:8 [ref. 30399]). Current status: Synonym of Cottus bairdii Girard 1850. Cottidae. Distribution: Western U.S.A. : Colorado, Utah, Wyoming and New Mexico [this subspecies]. Habitat: freshwater.

putorania, Paracottus (Paracottus) kneri Koryakov [E. A.] & Sidelev [G. N.] 1976:555 [Voprosy Ikhtiologii v. 16 (no. 3); ref. 9001] Lake Agatha, Enisei River basin, Russia. Syntypes: whereabouts unknown. On p. 500 or English translation. •Synonym of Paracottus knerii (Dybowski 1874), but a valid subspecies putorania Koryakov & Sidelev 1976 as described -- (Sideleva 1982:28 [ref. 14469], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:153 [ref. 23595], Sideleva 2001:59 [ref. 26181]). Current status: Synonym of Paracottus knerii (Dybowski 1874). Cottidae. Habitat: freshwater.

pygmaeus, Cottus Williams [J. D.] 1968:334, Fig. 1 [Copeia 1968 (no. 2); ref. 7934] Spring, 5.7 miles west of Oxford about 500 yards north of U.S. highway 78, tributary of Coosa River, Calhoun County, Alabama, U.S.A. Holotype: UMMZ 187462. Paratypes: ALA 2031 (12), 1138 (39), 1665 (68); CUMB 52064 [ex ALA 1063] (26); TU 34396 (47); UMMZ 187463 (25); USNM 201404 [ex ALA 1063] (26). Type catalog: Suttkus 1970:128 [ref. 20861], Bart & Taylor 1993:67 [ref. 20906]. Objectively invalid; preoccupied by Cottus quadricornis pygmaeus Lönnberg 1932; replaced by Cottus paulus Williams 2000. •Treated as valid as Cottus pygmaeus Williams 1968 -- (Lee et al. 1980:821 [ref. 22416], Page & Burr 1991:252 [ref. 18983], Boschung 1992:102 [ref. 23239], Mettee et al. 1996:495 [ref. 25836]). Current status: Synonym of Cottus paulus Williams 2000. Cottidae. Habitat: freshwater.

pygmaeus, Cottus quadricornis Lönnberg [E.] 1933:9, Pl. 1 (fig. 2) [Arkiv för Zoologi v. 24 A (pt 3) (no. 12) [1932]; ref. 22149] Lake Puruvesi, Finland. Syntypes: MZH. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), subspecies onegensis Berg & Popov 1932 -- (Berg 1949:1135 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

quadratus, Porocottus Bean [T. H.] in Jordan & Evermann 1898:1998 [Bulletin of the United States National Museum No. 47; ref. 2444] Bering Island, Commander Islands. Holotype (unique): USNM 33875. Author is T. H. Bean. Also appeared as Bean in Jordan & Gilbert 1899:466, Pl. 67a [ref. 2478]. •Synonym of Microcottus sellaris (Gilbert 1896) -- (Neyelov 1979:134 [ref. 3152], Lindberg & Krasyukova 1987:218 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:483 [ref. 25968], Parin et al. 2002:S80 [ref. 28300], Parin et al. 2014:259 [ref. 33547], Mecklenburg et al. 2016:75 [ref. 34440]). Current status: Synonym of Microcottus sellaris (Gilbert 1896). Cottidae. Habitat: brackish, marine.

quadricornis, Cottus Linnaeus [C.] 1758:264 [Systema Naturae, Ed. X v. 1; ref. 2787] Baltic Sea. Syntypes: NRM 2491-92 (1, 1). Type catalog: Fernholm & Wheeler 1983:235-236 [ref. 20707] who suggest that the largest syntype would make the best lectotype. •Valid as Triglopsis quadricornis (Linnaeus 1758) -- (Neyelov 1979:133 [ref. 3152], Fedorov in Whitehead et al. 1986:1260 [ref. 13677], Kottelat 1997:170 [ref. 22952], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:154 [ref. 23595], Bogutskaya et al. 2001:46 [ref. 26178], Parin et al. 2002:S89 [ref. 28300], Bogutskaya & Naseka 2004:191 [ref. 28183], Fricke 2007:29 [ref. 30577], Kottelat & Freyhof 2007:519 [ref. 29996]). •Valid as Myoxocephalus quadricornis (Linnaeus 1758) -- (Berg 1949:1128 [ref. 20662], Neyelov 1973:597 [ref. 7219], Lee et al. 1980:826 [ref. 22416], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:504 [ref. 25518], McAllister 1990:200 [ref. 14674], Page & Burr 1991:245 [ref. 18983], Scott & Crossman 1998:842 [ref. 24570], Mecklenburg et al. 2002:477 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Scharpf 2008:11 [ref. 30399], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212], Page & Burr 2011:468 [ref. 31215], Page et al. 2013:124 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:263 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:46, 227 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:80 [ref. 34440], Mecklenburg et al. 2018:217 [ref. 36370]). Current status: Valid as Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Distribution: Circumpolar in Arctic Ocean, in Pacific and north Atlontic. Habitat: freshwater, brackish, marine.

quadrifilis, Porocottus Gill [T. N.] 1859:166 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 11; ref. 1760] Bering Strait. Holotype (unique): USNM 6627 (badly damaged). See Mecklenburg et al. 2002:405 and 484 [ref. 25968] •Valid as Porocottus quadrifilis Gill 1859 -- (Neyelov 1979:136 [ref. 3152], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:484 [ref. 25968], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S84 [ref. 28300], Parin et al. 2014:269 [ref. 33547]). Current status: Valid as Porocottus quadrifilis Gill 1859. Cottidae. Distribution: Probably Bering Strait; known from only one record with uncertain locality. Habitat: marine.

quadriseriatus, Artedius Lockington [W. N.] 1880:330 [Proceedings of the United States National Museum v. 2 (no. 97); ref. 2815] San Francisco, California, U.S.A. Syntypes: USNM 23503 (2). •Valid as Icelinus quadriseriatus (Lockington 1880) -- (Bolin 1944:35 [ref. 6379], Eschmeyer & Herald 1983:173 [ref. 9277], Peden 1984:79 [ref. 5193], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:178 [ref. 35888]). Current status: Valid as Icelinus quadriseriatus (Lockington 1880). Cottidae. Distribution: Eastern North Pacific: northern California to southern Baja California. Habitat: marine.

quiescens, Uranidea DeKay [J. E.] 1842:61, Pl. 5 (fig. 14) [Zoology of New-York; ref. 1098] Round Lake and Lake Pleasant, New York, U.S.A. Syntypes: New York State coll. (whereabouts unknown). •Synonym of Cottus cognatus Richardson 1837 -- (Smith 1986:358 [ref. 22186], Scott & Crossman 1998:835 [ref. 24570]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

raninus, Myoxocephalus Jordan [D. S.] & Starks [E. C.] 1904:277, Fig. 20 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Aomori, Japan. Holotype: CAS-SU 7708. Paratypes: ?CAS 56330 [ex IU 10595] (1), USNM 50920 (13). Additional material: CAS-SU 7591 (15, missing), 7592 (5), 7595 (5), 7938 (now 83); USNM 51285-86 (1, 1), 23815 [ex Iowa Mus.] (1). Type catalog: Böhlke 1953:131 [ref. 12291]. •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Watanabe 1960:82 [ref. 11959], Neyelov 1979:121 [ref. 3152], Lindberg & Krasyukova 1987:207 [ref. 15964], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:480 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Parin et al. 2014:265 [ref. 33547], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Synonym of Myoxocephalus stelleri Tilesius 1811. Cottidae. Habitat: brackish, marine.

rastrinoides, Icelus Taranetz [A. Ya.] 1936:150 [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 4 (pt 13) (No. 3, art. 107); ref. 6509] Cave Cape; Olympyad Bay; and 45°05'N, 138°06'E, depth 230-411 meters, coast of Primorye, Russia. Syntypes: (8) ZIN 21445 (3). See also Nakabo 2002:1525 [ref. 26001]. •Valid as Icelus rastrinoides Taranetz 1936 -- (Nelson 1984:29 [ref. 5391], Lindberg & Krasyukova 1987:193 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670], Parin et al. 2002:S78 [ref. 28300], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:255 [ref. 33547]). Current status: Valid as Icelus rastrinoides Taranetz 1936. Cottidae. Distribution: Western North Pacific. Habitat: marine.

recalvus, Blennicottus Greeley [A. W.] 1899:9, Fig. 1 [Bulletin of the U. S. Fish Commission v. 19 [1899]; ref. 1889] Pacific Grove, California, U.S.A. Holotype: CAS-SU 6068. Paratypes: (4) CAS-SU 4245 (1, discarded). Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Clinocottus recalvus (Greeley 1899) -- (Bolin 1944:79 [ref. 6379], Eschmeyer & Herald 1983:167 [ref. 9277], Matarese et al. 1989:428 [ref. 26885], Nelson et al. 2004:119 [ref. 27807], Page et al. 2013:121 [ref. 32708], Kells et al. 2016:168 [ref. 35888]). Current status: Valid as Clinocottus recalvus (Greeley 1899). Cottidae. Distribution: Eastern North Pacific: southern Oregon (U.S.A.) to central Baja California. Habitat: marine.

regulus, Astrocottus Tsuruoka [O.], Maruyama [S.] & Yabe [M.] 2008:34, Figs. 3C, 6 [Bulletin of the National Museum of Nature and Science (Ser. A) Supplement No. 2; ref. 29568] Off Oumu, Okhotsk coast, Hokkaido, Japan, 44°33.89'N, 143°07.10'E, depth 46 meters. Holotype: HUMZ 190094. Paratypes: CAS 225634 [ex HUMZ 190085, 190087] (2); HUMZ 91981 (1), 91989 (1), 98614-15 (2), 189828-30 (3), 190081-84 (4), 191778 (1), 192515 (1), 195516 (1), 195746 (1), 195747 (1), 195748 (1), 195749-53 (5); NSMT-P 76686-89 [ex HUMZ 195759-62] (4); USNM 384159 [ex HUMZ 190086, 190088] (2). •Valid as Astrocottus regulus Tsuruoka, Maruyama & Yabe 2008 -- (Parin et al. 2014:239 [ref. 33547]). Current status: Valid as Astrocottus regulus Tsuruoka, Maruyama & Yabe 2008. Cottidae. Distribution: Off Usujiri, Okhotsk coast of Hokkaido, and off Miyagi and Fukushima Prefecture, Pacific coast, and off Obira, Hokkaido, Sea of Japan. Habitat: marine.

reinii, Cottus Hilgendorf [F. M.] 1879:105 [Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin 1879; ref. 21467] Hakodate and southern Nippon, Japan. Syntypes: ZMB 10627 (1) s. Nippon, 10629 (3) Hakodate. Type catalog: Paepke & Fricke 1992:283 [ref. 19946]. See also Nakabo 2002:1525 [ref. 26001]. •Valid as Cottus reinii Hilgendorf 1879 -- (Okada 1961:706 [ref. 6363], Yabe in Masuda et al. 1984:326 [ref. 6441], Kawanabe & Mizuno 1989:668 [ref. 25598], Nakabo 2000:638 [ref. 25086], Matsuura et al. 2000:210 [ref. 26000], Nakabo 2002:638 [ref. 26001], Hagiwara 2003:37 [ref. 27188], Fujii et al. 2005:12 [ref. 28191], Goto et al. 2014:[9] [ref. 33456]). Current status: Valid as Cottus reinii Hilgendorf 1879. Cottidae. Distribution: Western North Pacific. Habitat: freshwater.

relicta, Cottus quadricornis var. Lilljeborg [N.] 1891:149 [Sveriges och Norges fauna. Fiskarne. v. 1; ref. 21766] Lake Vättern, Sweden. No types known. •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies relictus (Lilljeborg 1891) -- (Berg 1949:1133 [ref. 20662]). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

retusifrons, Cottus Gronow [L. T.] in Gray 1854:103 [Catalogue of fish collected and described by Laurence Theodore Gronow; ref. 1911] English Sea. No types known. Cottidae.

rhenanus, Cottus Freyhof [J.], Kottelat [M.] & Nolte [A.] 2005:148, Figs. 21-22 [Ichthyological Exploration of Freshwaters v. 16 (no. 2); ref. 28259] Upper reaches of stream Wahnbach at Neunkirchen, 50°51'N, 7°20'E, Germany. Holotype: ZMB 33589. Paratypes: CMK 17844 (20), FSFJ 503 (40), ZMB 33590 (20). Plus non-type material. •Valid as Cottus rhenanus Freyhof, Kottelat & Nolte 2005 -- (Kottelat & Freyhof 2007:516 [ref. 29996], Goto et al. 2014:[11] [ref. 33456], Sideleva et al. 2015:33 [ref. 35276], Sideleva et al. 2018:9 [ref. 36393]). Current status: Valid as Cottus rhenanus Freyhof, Kottelat & Nolte 2005. Cottidae. Distribution: Europe. Habitat: freshwater.

rhodorus, Ascelichthys Jordan [D. S.] & Gilbert [C. H.] 1880:264 [Proceedings of the United States National Museum v. 3 (no. 144); ref. 2367] Wadda Island, Neah Bay, near Cape Flattery, Washington, U.S.A. Syntypes: (about 200) ANSP 12006-07 (2); MNHN A-3274 (4); MTD F 151 [ex USNM] (3); UMMZ 162182 (3); USNM 27331 (25, 4 in skeletal coll.); ZMB 11607 [ex USNM 27331] (3); ZMUB 1295 [ex USNM 27331] (1). Type catalog: Blanc & Hureau 1968:48 [ref. 20738], Frøiland 1979:148 [ref. 21547], Böhlke 1984:65 [ref. 13621], Paepke & Fricke 1992:282 [ref. 19946], Zarske 2003:31 [ref. 27474]. •Valid as Ascelichthys rhodorus Jordan & Gilbert 1880 -- (Bolin 1944:88 [ref. 6379], Eschmeyer & Herald 1983:163 [ref. 9277], Matarese et al. 1989:386 [ref. 26885], McAllister 1990:194 [ref. 14674], Mecklenburg et al. 2002:419 [ref. 25968], Nelson et al. 2004:118 [ref. 27807], Page et al. 2013:121 [ref. 32708], Pietsch & Orr 2015:38 [ref. 34694], Kells et al. 2016:166 [ref. 35888]). Current status: Valid as Ascelichthys rhodorus Jordan & Gilbert 1880. Cottidae. Distribution: Eastern North Pacific: Prince William Sound, northern Gulf of Alaska, to central California, U.S.A. Habitat: marine.

rhothea, Uranidea Smith [R.] 1882:347 [Proceedings of the United States National Museum v. 5 (no. 286); ref. 18726] Falls of Spokane River, Washington, U.S.A. Syntypes: USNM 30737 (2), plus 2 more. •Valid as Cottus rhotheus (Smith 1882) -- (Lee et al. 1980:822 [ref. 22416], McAllister 1990:197 [ref. 14674], Page & Burr 1991:245 [ref. 18983], Scott & Crossman 1998:837 [ref. 24570], Wydoski & Whitney 2003:214 [ref. 27816], Nelson et al. 2004:119 [ref. 27807], McPhail 2007:481 [ref. 31199], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:469 [ref. 31215], Page et al. 2013:122 [ref. 32708], LeMone et al. 2014:249 [ref. 33134], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus rhotheus (Smith 1882). Cottidae. Distribution: Southwestern Canada and northwestern U.S.A. (Washington, Idaho and Montana). Habitat: freshwater.

ricei, Cottopsis Nelson [E. W.] 1876:40 [Bulletin of the Illinois Museum of Natural History v. 1 (no. 1); ref. 3154] Lake Michigan near Evanston, Cook County, Illinois, U.S.A. Holotype (unique): whereabouts unknown, not at INHS. Type catalog: Sabaj et al. 1997:291 [ref. 23225]. •Valid as Cottus ricei (Nelson 1876) -- (Lee et al. 1980:823 [ref. 22416], Smith 1986:358 [ref. 22186], McAllister 1990:197 [ref. 14674], Page & Burr 1991:245 [ref. 18983], Chereshnev 1996:606 [ref. 24659], Scott & Crossman 1998:839 [ref. 24570], Lyons et al. 2000:42 [ref. 26594], Bailey et al. 2004:172 [ref. 27882], Nelson et al. 2004:119 [ref. 27807], McPhail 2007:486 [ref. 31199], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:468 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus ricei (Nelson 1876). Cottidae. Distribution: Canada and northern U.S.A. Habitat: freshwater.

richardsoni, Cottus Agassiz [L.] 1850:300 [Lake Superior; ref. 66] Montreal River, North America. Syntypes: whereabouts unknown. •Synonym of Cottus bairdii Girard 1850 -- (Scott & Crossman 1998:830 [ref. 24570] as bairdi). Current status: Synonym of Cottus bairdii Girard 1850. Cottidae. Habitat: freshwater.

richardsonii, Trachidermus Heckel [J. J.] 1839:162 [Annalen des Wiener Museums der Naturgeschichte v. 2 (no. 1); ref. 2079] Columbia River at Fort Vancouver, Vancouver, southern Washington, U.S.A. Unneeded new name for Cottus asper Richardson. •Synonym of Cottus asper Richardson 1837 -- (Scott & Crossman 1998:825 [ref. 24570]). Current status: Synonym of Cottus asper Richardson 1837. Cottidae. Habitat: freshwater, brackish, marine.

rimensis, Rusciculus Greeley [A. W.] 1899:13, Fig. 3 [Bulletin of the U. S. Fish Commission v. 19 [1899]; ref. 1889] Point Lobos, near Carmel, California, U.S.A. Holotype: CAS-SU 6067. Paratypes: CAS-SU 6067 (1 of 2). Type catalog: Böhlke 1953:132 [ref. 12291]. Holotype established in table on p. 14 if specimen is identifiable. •Valid as Oligocottus rimensis (Greeley 1899) -- (Bolin 1944:63 [ref. 6379], Eschmeyer & Herald 1983:178 [ref. 9277], McAllister 1990:201 [ref. 14674], Mecklenburg et al. 2002:499 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:43 [ref. 34694], Kells et al. 2016:186 [ref. 35888]). Current status: Valid as Oligocottus rimensis (Greeley 1899). Cottidae. Distribution: Eastern North Pacific: southeastern Alaska to northern Baja California. Habitat: marine.

robustus, Clypeocottus Ayres [W. O.] 1854:194 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] California, U.S.A. No types known. Appeared first as above on 13 Oct.; then in Ayres 1854:11 [ref. 18923] where Ayres mentions that Girard's genus Aspicottus and species A. bison are synonyms and have priority. •Synonym of Enophrys bison (Girard 1854) -- (Bolin 1944:91 [ref. 6379], Sandercock & Wilimovsky 1968:844 [ref. 26414]). Current status: Synonym of Enophrys bison (Girard 1854). Cottidae. Habitat: marine.

rondeleti, Cottus Freyhof [J.], Kottelat [M.] & Nolte [A.] 2005:140, Fig. 15 [Ichthyological Exploration of Freshwaters v. 16 (no. 2); ref. 28259] 1 kilometer south of Brissac, 3 kilometers southwest of St. Bauzille-de-Putois, 43°39'N, 3°15'E, creek tributary of Hérault River, Dept. Hérault, France. Holotype: MNHN 2005-0757. Paratypes: CMK 13008 (8), 18516 (3), 18518 (2); FSJF 1655 (2); MNHN 2005-0758 (4). •Valid as Cottus rondeleti Freyhof, Kottelat & Nolte 2005 -- (Kottelat & Freyhof 2007:517 [ref. 29996], Goto et al. 2014:[11] [ref. 33456]). Current status: Valid as Cottus rondeleti Freyhof, Kottelat & Nolte 2005. Cottidae. Distribution: Hérrault drainage, France. Habitat: freshwater.

roseus, Cottus gobio var. Odenwall [E.] 1927:56 [Memoranda Societatis pro Fauna et Flora Fennica. v. 2; ref. 15808] Lappajärvi Sjö, Finland. Syntypes: Zool. Mus. Helsinki. •Synonym of Cottus gobio Linnaeus 1758 -- (Berg 1949:1145 [ref. 20662]). Current status: Synonym of Cottus gobio Linnaeus 1758. Cottidae. Habitat: freshwater, brackish.

rubellio, Eximia Greeley [A. W.] 1899:18, Fig. 5 [Bulletin of the U. S. Fish Commission v. 19 [1899]; ref. 1889] Monterey Bay, California, U.S.A. Holotype: ? CAS-SU 6066. Paratypes: (5) CAS-SU 3642 (1, discarded), 4048 (2, ? missing). Type catalog: Böhlke 1953:129 [ref. 12291]. Type established in table on p. 19 and in Fig. 5 if specimen is identifiable. •Valid as Oligocottus rubellio (Greeley 1899) -- (Bolin 1944:70 [ref. 6379], Eschmeyer & Herald 1983:178 [ref. 9277], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708]). Current status: Valid as Oligocottus rubellio (Greeley 1899). Cottidae. Distribution: Eastern North Pacific: northern California to northern Baja California. Habitat: marine.

sabaudicus, Cottus Sideleva [V. G.] 2009:211, Figs. 2-3 [Journal of Ichthyology v. 49 (no. 3); ref. 30173] Filiere River, upper reaches of the Rhone River, Savoy Alps, Haute-Savoy district, southeastern France. Holotype: MNHN 1960-0373. Paratypes: MNHN 1960-0374 (1), 1960-0375 (1). •Valid as Cottus sabaudicus Sideleva 2009 -- (Goto et al. 2014:[11] [ref. 33456], Sideleva et al. 2015:33 [ref. 35276], Sideleva et al. 2018:9 [ref. 36393]). Current status: Valid as Cottus sabaudicus Sideleva 2009. Cottidae. Distribution: Tributaries of the upper course of the Rhone River, France. Habitat: freshwater.

saburrae, Rusulus Starks [E. C.] & Mann [W. M.] 1911:14, Fig. 2 [University of California Publications in Zoology v. 8 (no. 2); ref. 4199] Ballast Point, off mouth of San Diego Bay, California, U.S.A. Holotype (unique): CAS-SU 23615. Type catalog: Böhlke 1953:132 [ref. 12291]. •Synonym of Clinocottus analis (Girard 1858). Current status: Synonym of Clinocottus analis (Girard 1858). Cottidae. Habitat: marine.

sagamia, Daruma Jordan [D. S.] & Starks [E. C.] 1904:241, Fig. 4 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Sagami Bay, Japan. Holotype: USNM 50914. Paratypes: CAS-SU 7702 (1). Additional material: CAS-SU 7850 (1), 8557 (1). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Daruma sagamia Jordan & Starks 1904 -- (Watanabe 1960:29 [ref. 11959], Yabe in Masuda et al. 1984:325 [ref. 6441], Nakabo 2000:632 [ref. 25086], Nakabo 2002:632 [ref. 26001]). Current status: Valid as Daruma sagamia Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

satoi, Pseudoblennius Watanabe [M.] 1958:391, Figs. 185-186, Pl. 39 [Studies of Japanese cottid fishes; ref. 21357] Toshima-Fukushia, Hokkaido, Japan, depth 20 fathoms. Syntypes: (4). Also described as new in Watanabe 1960:157, Pl. 35 (fig. 2) [ref. 11959]. •Valid as Pseudoblennius satoi Watanabe 1958 -- (Watanabe 1960:157 [ref. 11959]). •Synonym of Pseudoblennius cottoides (Richardson 1848) -- (Nakabo 2002:1526 [ref. 26001]). Current status: Synonym of Pseudoblennius cottoides (Richardson 1848). Cottidae. Distribution: Western North Pacific. Habitat: marine.

scaber, Artediellus Knipowitsch [N. M.] 1907:18, Pl. 1 (figs. 7-12) [Zapiski Imperatorskoj Akademii Nauk po Fiziko-Matematiceskomu Otdelniji = Mémoires de l'Académie Impériale des Sciences de St. Petersbourg, Classe Physico-Mathématique (8 Sér.) v. 18 (no. 5); ref. 15152] Northeastern, eastern and southeastern part of European Arctic Ocean, Kara Sea, to Bering Strait. Lectotype: ZIN 14206. Paralectotypes: (79) ZIN 14205-06 (13, 4); ZMUC 22-23 (2). Type catalog: Nielsen 1974:81 [ref. 9588]. Lectotype selected by Neyelov 1979:154 [ref. 3152]. •Valid as Artediellus scaber Knipowitsch 1907 -- (Neyelov 1973:595 [ref. 7219], Neyelov 1979:154 [ref. 3152], Fedorov in Whitehead et al. 1986:1245 [ref. 13677], Nelson 1986:43 [ref. 5808], McAllister 1990:194 [ref. 14674], Sheiko & Fedorov 2000:27 [ref. 25650], Mecklenburg et al. 2002:491 [ref. 25968], Parin et al. 2002:S71 [ref. 28300], Nelson et al. 2004:118 [ref. 27807], Mecklenburg et al. 2007:174 [ref. 29448], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:121 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:239 [ref. 33547], Mecklenburg & Steinke 2015:163 [ref. 34033], Afonso et al. 2016:41, 100 [ref. 36199], Kells et al. 2016:162 [ref. 35888], Mecklenburg et al. 2016:51 [ref. 34440], Mecklenburg et al. 2018:188 [ref. 36370]). Current status: Valid as Artediellus scaber Knipowitsch 1907. Cottidae. Distribution: Arctic Ocean, North Pacific, North Atlantic: Chukchi and Beaufort seas south to northern Bering Sea, east to Somerset Island, Canada, and west to Barents and Kara seas. Habitat: brackish, marine.

scaturigo, Cottus Freyhof [J.], Kottelat [M.] & Nolte [A.] 2005:152, Figs. 23-24 [Ichthyological Exploration of Freshwaters v. 16 (no. 2); ref. 28259] Timavo spring, southwest of Monfalcone, 45°47.24'N, 13°35.29"E, Province Friuli-Venezia, Italy. Holotype: ZMB 33591. Paratypes: FSJF 181 (12), 219 (10), 1020 (5); CMK 18550 (5). •Synonym of Cottus gobio (Linnaeus 1758) -- (Bianco & Delmastro 2011:6 [ref. 31648] as scaturigio, Bianco 2013:10 [ref. 32897]). •Valid as Cottus scaturigo Freyhof, Kottelat & Nolte 2005 -- (Kottelat & Freyhof 2007:517 [ref. 29996], Sideleva 2009:211 [ref. 30173], Goto et al. 2014:[11] [ref. 33456]). Current status: Valid as Cottus scaturigo Freyhof, Kottelat & Nolte 2005. Cottidae. Distribution: Italy. Habitat: freshwater.

scepticus, Triglops Gilbert [C. H.] 1896:428, Pl. 28 (lower) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Unalaska to Kodiak islands, Alaska, U.S.A., Albatross station 3339, depth 253 meters. Lectotype: USNM 64325. Paralectotypes: BMNH 1900.7.11.6 (1); MCZ 28235 (1), 28353 (1); CAS-SU 2426 (1?), 2427 (1), 2428 (5, 1 + disintegrated specimens), 69686 (5); USNM 125621 (1). Type catalog: Böhlke 1953:133 [ref. 12291]. Lectotype established (as figured specimen from Albatross sta. 3339) in caption to Pl. 288, p. 3291 in Jordan & Evermann 1900 [ref. 2446], which is the same figure as in the original description; Pietsch 1994:358 [ref. 21189] identified that specimen as SU 2426 and designated it as lectotype. •Valid as Triglops scepticus Gilbert 1896 -- (Watanabe 1960:15 [ref. 11959], Yabe in Amaoka et al. 1983:287, 338 [ref. 8379], Yabe in Masuda et al. 1984:325 [ref. 6441], Lindberg & Krasyukova 1987:176 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Pietsch 1994:358 [ref. 21189], Shinohara et al. 1996:176 [ref. 26605], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:628 [ref. 25086], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:425 [ref. 25968], Nakabo 2002:628 [ref. 26001], Youn 2002:263, 579 [ref. 26218], Parin et al. 2002:S89 [ref. 28300], Fedorov et al. 2003:82 [ref. 27579], Nelson et al. 2004:122 [ref. 27807], Shinohara et al. 2011:44 [ref. 31715], Page et al. 2013:125 [ref. 32708], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:278 [ref. 33547], Tohkairin et al. 2015:7 [ref. 35331], Kells et al. 2016:194 [ref. 35888]). Current status: Valid as Triglops scepticus Gilbert 1896. Cottidae. Distribution: North Pacific: Bering Sea to southeastern Alaska and to Okhotsk Sea and Japan Sea to Korea. Habitat: marine.

schitsuumsh, Cottus LeMoine [M.], Young [M. K.], McKelvey [K. S.], Eby [L.], Pilgrim [K. L.] & Schwartz [M. K.] 2014:244, Figs. 4-7 [Zootaxa 3755 (no. 3); ref. 33134] East Fork Steamboat Creek, a tributary of the Coeur d'Alene River at West Fork Road, 47.71821°N, 116.20010°W, Shoshone County, Idaho. Holotype: UW 151670. Paratypes: UW 151671 (3). Plus additional non-type specimens. •Valid as Cottus schitsuumsh LeMoine, Young, McKelvey, Eby, Pilgrim & Schwartz 2014. Current status: Valid as Cottus schitsuumsh LeMoine, Young, McKelvey, Eby, Pilgrim & Schwartz 2014. Cottidae. Distribution: Idaho and Montana, USA. Habitat: freshwater.

schlegelii, Centridermichthys Döderlein [L.] in Steindachner & Döderlein 1884:210 [42] [Denkschriften der Kaiserlichen Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Classe. v. 49 (1. Abth.); ref. 4248] Tokyo, Japan. Syntypes: NMW 6373 (1), ?ZMB 12079 (1). Type catalog: Paepke & Fricke 1992:282 [ref. 19946]. Type information (ZMB) from Amaoka & Fujii 1999:163 [ref. 26515]. Cottidae.

schmidti, Artediellus Soldatov [V. K.] 1915:160, Fig. 2 [Ezhegodnik, Zoologicheskago Muzeya Imperatorskoi Akademii Nauk v. 20 (for 1915); ref. 15068] Tatar Strait and Okhotsk Sea, including Aniva Bay. Syntypes: (many) ZIN 12186-87 (6+, 6+), 12190 (2), 12913 (5). •Valid as Artediellus schmidti Soldatov 1915 -- (Lindberg & Krasyukova 1987:235 [ref. 15964], Parin et al. 2002:S71 [ref. 28300], Parin et al. 2014:239 [ref. 33547]). •Synonym of Artediellus dydymovi Soldatov 1915, but a valid subspecies -- (Neyelov 1979:155 [ref. 3152], Yabe in Masuda et al. 1984:328 [ref. 6441], Nakabo 2000:642 [ref. 25086], Nakabo 2002:642 [ref. 26001]). •Synonym of Artediellus dydymovi Soldatov 1915 -- (Dyldin & Orlov 2017:61 [ref. 35138] with question). Current status: Synonym of Artediellus dydymovi Soldatov 1915. Cottidae. Distribution: Western North Pacific [if valid]. Habitat: marine.

schmidti, Cottiusculus Jordan [D. S.] & Starks [E. C.] 1904:299, Fig. 30 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Kinkazwan [Kinkazan] Island, Matsushima Bay, Japan, depth 79 fathoms. Holotype: USNM 50923. Paratypes: CAS-SU 7714 (4), 7588 (5). Type catalog: Böhlke 1953:128 [ref. 12291]. •Valid as Cottiusculus schmidti Jordan & Starks 1904 -- (Watanabe 1960:168 [ref. 11959], Neyelov 1979:162 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:239 [ref. 15964], im & Youn 1992:72 [ref. 22287], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:643 [ref. 25086], Youn 2002:262, 574 [ref. 26218], Parin et al. 2002:S72 [ref. 28300], Kai & Nakabo 2009:220 [ref. 30299]). Current status: Valid as Cottiusculus schmidti Jordan & Starks 1904. Cottidae. Distribution: Off the Pacific coast of Japan. Habitat: marine.

schraderi, Sclerocottus Fischer [J. G.] 1885:58, Pl. 1 (figs. 3-4) [Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten v. 2; ref. 1333] Reported as South Georgia Island [locality apparently erroneous, see Berg 1940]. Holotype (unique): ZMH H67 [ex 3888]. Type catalog: Ladiges et al. 1958:165 [ref. 19181], Wilkens & Dohse 1993:407 [ref. 21161]. •Synonym of Gymnocanthus tricuspis (Reinhardt 1830) -- (Neyelov 1973:595 [ref. 7219] as Gymnacanthus). Current status: Synonym of Gymnocanthus tricuspis (Reinhardt 1830). Cottidae. Habitat: marine.

scorpioides, Cottus Fabricius [O.] 1780:157 [Fauna groenlandica, systematice sistens animalia Groenlandiae occidentalis, etc.; ref. 17464] Western Greenland. No types known. •Valid as Myoxocephalus scorpioides (Fabricius 1780) -- (Neyelov 1973:599 [ref. 7219], Neyelov 1979:122 [ref. 3152], Fedorov in Whitehead et al. 1986:1254 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:505 [ref. 25518], McAllister 1990:200 [ref. 14674], Yabe in Okamura et al. 1995:165 [ref. 22531], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:479 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:264 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:49, 228 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:82 [ref. 34440], Mecklenburg et al. 2018:219 [ref. 36370]). Current status: Valid as Myoxocephalus scorpioides (Fabricius 1780). Cottidae. Distribution: Arctic Ocean, northwestern Atlantic and North Pacific: eastern Siberian and Chukchi seas to northern Bering Sea, Beaufort Sea to Greenland and Gulf of St. Lawrence. Habitat: brackish, marine.

scorpius, Cottus Linnaeus [C.] 1758:265 [Systema Naturae, Ed. X v. 1; ref. 2787] European Ocean. Syntypes: BMNH 1853.11.12.17 [Gronovius coll.] (1, skin), LS 17 (left half-skin plus parts), NRM 2357 (2). Type catalog: Goode & Bean 1885:196 [ref. 1841], Wheeler 1958:240 [ref. 13434], Fernholm & Wheeler 1983:234-235 [ref. 20707], Wheeler 1985:46 [ref. 20705]. •Valid as Myoxocephalus scorpius (Linnaeus 1758) -- (Neyelov 1973:598 [ref. 7219], Neyelov 1979:122 [ref. 3152], Fedorov in Whitehead et al. 1986:1254 [ref. 13677], Robins & Ray 1986:283 [ref. 23100], Scott & Scott 1988:507 [ref. 25518], Matarese et al. 1989:402 [ref. 26885], McAllister 1990:200 [ref. 14674], Yabe in Okamura et al. 1995:165 [ref. 22531], Klein-MacPhee in Collette & Klein-MacPhee 2002:353 [ref. 26158], Mecklenburg et al. 2002:478 [ref. 25968], Parin et al. 2002:S82 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Fricke 2007:29 [ref. 30577], Mecklenburg et al. 2007:176 [ref. 29448], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:123 [ref. 31212] with comments. Page et al. 2013:124 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:264 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:49, 229 [ref. 36199], Kells et al. 2016:184 [ref. 35888], Mecklenburg et al. 2016:84 [ref. 34440], Mecklenburg et al. 2018:221 [ref. 36370]). Current status: Valid as Myoxocephalus scorpius (Linnaeus 1758). Cottidae. Distribution: Circumpolar, northern Hemisphere. Habitat: brackish, marine.

scutiger, Icelus Bean [T. H.] 1890:41 [Proceedings of the United States National Museum v. 13 (no. 795); ref. 229] Off Trinity Island, Bering Sea, 56°N, 154°20'W, Albatross station 2853, depth 159 fathoms. Holotype: USNM 45368. Paratypes: MCZ 27860 (1), USNM 227362 (2). •Valid as Rastrinus scutiger (Bean 1890) -- (Nelson 1984:17 [ref. 5391], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:457 [ref. 25968], Parin et al. 2002:S85 [ref. 28300], Nelson et al. 2004:121 [ref. 27807], Stevenson 2006:792 [ref. 28923], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:271 [ref. 33547], Kells et al. 2016:190 [ref. 35888]). Current status: Valid as Rastrinus scutiger (Bean 1890). Cottidae. Distribution: North Pacific: Commander and Aleutian islands to western Gulf of Alaska. Habitat: marine.

sekii, Icelus Tsuruoka [O.], Munehara [H.] & Yabe [M.] 2006:47, Figs. 1-2 [Ichthyological Research v. 53 (no. 1); ref. 28501] 44°02.1'N, 145°13.2'E, off Rausu, Shiretoko Peninsula, Hokkaido Island, Japan, depth 20 meters. Holotype: HUMZ 187896. Paratypes: HUMZ 179902 (1), 187897 (2), 187898 (1), 188505 (1). •Valid as Icelus sekii Tsuruoka, Munehara & Yabe 2006 -- (Parin et al. 2014:256 [ref. 33547]). Current status: Valid as Icelus sekii Tsuruoka, Munehara & Yabe 2006. Cottidae. Distribution: Hokkaido, Japan and Kuril Islands. Habitat: marine.

sellaris, Acanthocottus Gilbert [C. H.] 1896:419 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bristol Bay, Alaska, U.S.A., depth 5-17 fathoms. Syntypes: BMNH 1900.7.11.4-5 (2); MCZ 28321 (4), 28351 (2), 28366 (?); MNHN 1896-0461 [ex USNM] (6); CAS-SU 2846-47 (24, 35); USNM 48575 (3, now 4); ZMUO J1401-02 (1, 3). Type catalog: Böhlke 1953:126 [ref. 12291], Blanc & Hureau 1968:48 [ref. 20738], Pethon 1969:7 [ref. 19268]. •Valid as Microcottus sellaris (Gilbert 1896) -- (Neyelov 1979:134 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:218 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:644 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:483 [ref. 25968], Nakabo 2002:644 [ref. 26001], Parin et al. 2002:S80 [ref. 28300], Yabe & Pietsch 2003:276 [ref. 27215], Fedorov et al. 2003:74 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:259 [ref. 33547], Mecklenburg et al. 2016:74 [ref. 34440], Kells et al. 2016:182 [ref. 35888], Mecklenburg et al. 2018:211 [ref. 36370]). Current status: Valid as Microcottus sellaris (Gilbert 1896). Cottidae. Distribution: North Pacific and adjacent Arctic. Habitat: brackish, marine.

semiscaber, Cottopsis Cope [E. D.] 1872:476 [U. S. Geological Survey Territitories Annual Report v. 5 (pt 4); ref. 923] Fort Hall, Idaho, U.S.A. Syntypes: (3) whereabouts unknown. Type catalog: Böhlke 1984:66 [ref. 13621] as none at ANSP. •Synonym of Cottus bairdii Girard 1850, but a valid subspecies -- (La Rivers 1994:586 [ref. 23389] as bairdi, Scharpf 2008:8 [ref. 30399]). Current status: Synonym of Cottus bairdii Girard 1850. Cottidae. Distribution: Bonneville Basin and upper Snake River, Utah, Nevada and Idaho, U.S.A. [this subspecies]. Habitat: freshwater.

shasta, Cottus Jordan [D. S.] & Starks [E. C.] in Jordan 1896:224 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 2395] McCloud River at Baird, Shasta County, California, U.S.A. Syntypes: BMNH 1897.1.12.14 (1), CAS-SU 4196 (4, now 3). Type catalog: Böhlke 1953:128 [ref. 12291]. •Synonym of Cottus gulosus (Girard 1854) -- (Lee et al. 1980:814 [ref. 22416]). Current status: Synonym of Cottus gulosus (Girard 1854). Cottidae. Habitat: freshwater.

shibusawai, Pseudoblennius Watanabe [M.] 1958:395, Figs. 187-188, Pl. 40 [Studies of Japanese cottid fishes; ref. 21357] No locality [Japan]. Syntypes: (4). Also described as new in Watanabe 1960:160, Pl. 36 (fig. 1) [ref. 11959]. •Valid as Pseudoblennius shibusawai Watanabe 1958 -- (Watanabe 1960:152 [ref. 11959]). •Synonym of Pseudoblennius cottoides (Richardson 1848) -- (Nakabo 2002:1526 [ref. 26001]). Current status: Synonym of Pseudoblennius cottoides (Richardson 1848). Cottidae. Habitat: marine.

sibiricus, Cottus Warpachowski [N. A.] 1889:12 [Zapiski Ak. Nauk St. Petersburg 1889; ref. 19906] Yenisei [Enisei] River near Minusinsk Town. Lectotype: ZIN 6328. Paralectotypes: ZIN 6208 (1), 6330 (1), ZIN 56235 (1). Lectotype designated by Sideleva 2017:828 [ref. 35715]. •Valid as Cottus sibiricus Warpachowski 1889 (author sometimes seen as Kessler) -- (Berg 1949:1148 [ref. 20662], Sideleva 1982:81 [ref. 14469], Chereshnev 1996:606 [ref. 24659], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:152 [ref. 23595], Bogutskaya et al. 2001:46 [ref. 26178], Parin et al. 2002:S73 [ref. 28300], Bogutskaya & Naseka 2004:188 [ref. 28183], Ocock et al. 2006:57 [ref. 28933], Kottelat 2006:91 [ref. 28831], Shedko & Miroshnichenko 2007:24 [ref. 29369], Bogdanov & Knizhin 2007:271 [ref. 31009], Goto et al. 2014:[10] [ref. 33456], Parin et al. 2014:244 [ref. 33547], Romanov et al. 2017:826 [ref. 35687], Sideleva 2017:828 [ref. 35715]). Current status: Valid as Cottus sibiricus Warpachowski 1889. Cottidae. Distribution: Northern Asia: Russia and Mongolia. Habitat: freshwater, brackish.

siemienkiewiczii, Cottocomephorus grewingkii var. Dybowski [B. N.] 1908:559 [24], Fig. 20 [Kosmos Lwów v. 33; ref. 17422] Lake Baikal, Russia. Holotype (unique). Type not in ZIN. •Synonym of Cottocomephorus comephoroides (Berg 1900) -- (Bogdanov 2019:230 [ref. 36814]). Current status: Synonym of Cottocomephorus comephoroides (Berg 1900). Cottidae. Habitat: freshwater.

simus, Cottus Hermann [J.] 1804:302 [Observationes zoologicae; ref. 2146] No locality. Originally as Cottus ? simus. Cottidae.

sinensis, Cottus (Aspicottus) Sauvage [H.-E.] 1873:55 [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris v. 9 (pt 2, Bulletin); ref. 18661] China. Holotype (unique): MNHN 0000-5265. Type catalog: Blanc & Hureau 1968:52 [ref. 20738]. •Valid as Myoxocephalus sinensis (Sauvage 1873) -- (Zhu 1995:189 [ref. 25213]). Current status: Valid as Myoxocephalus sinensis (Sauvage 1873). Cottidae. Distribution: China. Habitat: marine.

sinensis, Lepidobero Qin [K.-J.] & Jin [X.-B.] 1992:1 [4] [Journal of Dalian Fisheries College v. 6 (nos 3-4); ref. 22919] Hei-Shi-Jiao, Dalian. Liaoning Province, China. Holotype: Dalian Fish. Coll. 8600051. Paratypes: Dalian Fish Coll. 8603078 (1, allotype); Shanghai Fish. Univ. 81(X)0010-11 (2), 8603976-77 (2). On p. 4 in English. •Valid as Lepidobero sinensis Qin & Jin 1992. Current status: Valid as Lepidobero sinensis Qin & Jin 1992. Cottidae. Distribution: China. Habitat: freshwater.

smithi, Sigmistes Schultz [L. P.] 1938:190 [Proceedings of the United States National Museum v. 85 (no. 3038); ref. 3951] Igitkin Island, Aleutian Islands, Alaska, U.S.A. Holotype: USNM 105282. Paratypes: USNM 105283 (2). •Valid as Sigmistes smithi Schultz 1938 -- (Yabe et al. 2001:68 [ref. 25246], Mecklenburg et al. 2002:497 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:272 [ref. 33547]). Current status: Valid as Sigmistes smithi Schultz 1938. Cottidae. Distribution: North Pacific: Aleutian Islands and Kuril Islands Intertidal. Habitat: marine.

snyderi, Dialarchus Greeley [A. W.] 1899:14, Fig. 4 [Bulletin of the U. S. Fish Commission v. 19 [1899]; ref. 1889] Pacific Grove, California, U.S.A. Holotype: CAS-SU 5846. Paratypes: CAS-SU 5847 (5). See Oligocottus snyderi Greeley 1898. •Preoccupied by and same as Oligocottus snyderi Greeley 1898. Current status: Synonym of Oligocottus snyderi Greeley 1898. Cottidae. Habitat: marine.

snyderi, Oligocottus Greeley [A. W.] in Jordan & Evermann 1898:2871 [Bulletin of the United States National Museum No. 47; ref. 2445] Pacific Grove, California, U.S.A. Lectotype: CAS-SU 5846. Paralectotypes: CAS-SU 5847 ( 5). Type catalog: Böhlke 1953:131 [ref. 12291]. Based on Oligogottus maculosus of Jordan & Evermann 1896:2013 [ref. 2444]. Lectotype apparently established by Greeley 1899:16 [ref. 1889]. •Valid as Oligocottus snyderi Greeley 1898 -- (Bolin 1944:67 [ref. 6379], Eschmeyer & Herald 1983:178 [ref. 9277], Matarese et al. 1989:432 [ref. 26885], McAllister 1990:201 [ref. 14674], Mecklenburg et al. 2002:501 [ref. 25968], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Pietsch & Orr 2015:43 [ref. 34694], Kells et al. 2016:186 [ref. 35888]). Current status: Valid as Oligocottus snyderi Greeley 1898. Cottidae. Distribution: Eastern North Pacific: western Gulf of Alaska to central Baja California. Habitat: marine.

spatula, Icelus Gilbert [C. H.] & Burke [C. V.] 1912:41, Figs. 3, 3a [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Off Avatcha Bay, Kamchatka, Russia, 52°47'20"N, 158°44'30"E, Albatross station 4794, depth 58-69 fathoms. Holotype: USNM 74367. Paratypes: CAS-SU 40931 (5), USNM 70904-05 (4, 4). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelus spatula Gilbert & Burke 1912 -- (Neyelov 1973:597 [ref. 7219], Nelson 1984:41 [ref. 5391], Fedorov in Whitehead et al. 1986:1251 [ref. 13677], Robins & Ray 1986:282 [ref. 23100], Scott & Scott 1988:499 [ref. 25518], McAllister 1990:199 [ref. 14674], Yabe in Okamura et al. 1995:164 [ref. 22531], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:455 [ref. 25968], Parin et al. 2002:S78 [ref. 28300], Fedorov et al. 2003:72 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2007:176 [ref. 29448], Møller et al. 2010:52 [ref. 31169], Mecklenburg et al. 2011:122 [ref. 31212] with comments, Page et al. 2013:123 [ref. 32708], Dolgov 2013:918 [ref. 33112], Mecklenburg et al. 2014:145 [ref. 33463], Parin et al. 2014:256 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:45, 174 [ref. 36199], Kells et al. 2016:178 [ref. 35888], Mecklenburg et al. 2016:69 [ref. 34440], Mecklenburg et al. 2018:207 [ref. 36370]). Current status: Valid as Icelus spatula Gilbert & Burke 1912. Cottidae. Distribution: Circumpolar, Arctic Ocean, northwestern Atlantic and North Pacific. Habitat: marine.

specus, Cottus Adams [G. L.] & Burr [B. M.] in Adams, Burr, Day & Starkey 2013:488, Figs. 4-5 [Zootaxa 3609 (no. 5); ref. 32429] Mystery Cave, Perry County, Missouri. Holotype: SIUC 88001. Paratypes: SIUC 88002 (2), USNM 409781 (2). Plus additional non-type material. •Valid as Cottus specus Adams & Burr 2013. Current status: Valid as Cottus specus Adams & Burr 2013. Cottidae. Distribution: Southeastern Missouri, U.S.A. Habitat: freshwater.

spilota, Uranidea Cope [E. D.] 1865:82 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 17; ref. 907] Grand Rapids, on the Grand River, Michigan, U.S.A. Lectotype: ANSP 11061 (missing). Paralectotypes: ANSP 11062 (1, missing), ?UMMZ 213813 (9). Type catalog: Böhlke 1984:66 [ref. 13621] as holotype and paratypes; lectotype established by Fowler 1918:51, Pl. 13 [ref. 11665]. •Synonym of Cottus bairdii Girard 1850. Current status: Synonym of Cottus bairdii Girard 1850. Cottidae. Habitat: freshwater.

spiniger, Icelus Gilbert [C. H.] 1896:412, Pl. 24 (top) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Alaska, U.S.A., 54°48'30"N, 165°49'00"W, Albatross station 3225, depth 85 fathoms. Lectotype: ?USNM 164258. Paralectotypes: BMNH 1900.7.11.7 (1?); MCZ 28270 (3); MNHN 1896-0484 to 0486 [ex USNM] (3); CAS-SU 2231 (40), 2266 (2), 2269 (16); ZMUO J1463 (3, poor condition). Type catalog: Böhlke 1953:130 [ref. 12291], Blanc & Hureau 1968:54 [ref. 20738], Pethon 1969:8 [ref. 19268]. Lectotype established (as figured specimen from Albatross sta. 3278) in caption to Pl. 286, p. 3290 in Jordan & Evermann 1900 [ref. 2446] if figured specimen is identifiable; a lectotype (USNM 164248, sta. 3225) was also designated by Nelson 1984:24 [ref. 5391]. •Valid as Icelus spiniger Gilbert 1896 -- (Watanabe 1960:42 [ref. 11959], Eschmeyer & Herald 1983:174 [ref. 9277], Nelson 1984:24 [ref. 5391], Peden & Hughes 1986:7 [ref. 9714], McAllister 1990:199 [ref. 14674], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:451 [ref. 25968], Parin et al. 2002:S78 [ref. 28300], Fedorov et al. 2003:73 [ref. 27579], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:122 [ref. 31212] as cf. spiniger with comments, Page et al. 2013:123 [ref. 32708], Parin et al. 2014:256 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Mecklenburg et al. 2016:71 [ref. 34440] see remarks, Kells et al. 2016:180 [ref. 35888]). Current status: Valid as Icelus spiniger Gilbert 1896. Cottidae. Distribution: North Pacific: northern Bering Sea to southern British Columbia and to Okhotsk Sea. Habitat: marine.

spinosus, Hemilepidotus Ayres [W. O.] 1854:166 [The Pacific [newspaper] v. 3 and 4 (thru no. 6); ref. 157] San Francisco Bay, California, U.S.A. Possible syntypes: BMNH 1863.10.9.103-104 (2?) from Ayres. Originally as Hemilepidotus ? spinosus. Appeared first as above on 25 Aug.; also as Calycilepidotus spinosus in Ayres 1855:76 [ref. 159]. •Valid as Hemilepidotus spinosus Ayres 1854 -- (Peden 1979:18 [ref. 24080], Eschmeyer & Herald 1983:171 [ref. 9277], Matarese et al. 1989:378 [ref. 26885], McAllister 1990:197 [ref. 14674], Mecklenburg et al. 2002:432 [ref. 25968], Tokranov et al. 2003:300 [ref. 26972], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:40 [ref. 34694], Kells et al. 2016:174 [ref. 35888]). Current status: Valid as Hemilepidotus spinosus Ayres 1854. Cottidae. Distribution: Eastern North Pacific: Kachemak Bay, northern Gulf of Alaska to southern California (U.S.A.). Habitat: marine.

spinosus, Porocottus Watanabe [M.] 1958:311, Figs. 132-133, Pl. 24 [Studies of Japanese cottid fishes; ref. 21357] Kurshiro, Hokaidô, Japan, depth 35 fathoms. Syntypes: (3). Also described as new in Watanabe 1960:98, Pl. 3 (fig. 2) [ref. 11959]. •Valid as Porocottus spinosus Watanabe 1958 -- (Watanabe 1960:98 [ref. 11959]). •Synonym of Myoxocephalus jaok (Cuvier 1829) -- (Nakabo 2002:1526 [ref. 26001], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus jaok (Cuvier 1829). Cottidae. Habitat: marine.

spinulosus, Cottus Yashchenko [A. L.] 1896:67 [Trudy St.-Peterburgskogo Obscestva Estestvoispytatelej = Travaux de la Société des Naturalistes de St. Pétersbourg. v. 25 (no. 2); ref. 20230] Not available, no distinguishing features; perhaps refers to Cottus spinulosus Kessler 1872. Cottidae.

spinulosus, Cottus Kessler [K. F.] 1872:47 [3], Pl. 6 (figs. 1-2) [Izvestiia Imperatorskago Obschchestva Liubitelei Estestvozaniia, Antropologii i Etnografii v. 10 (no. 1); ref. 14134] Khodzhent [Leninabad], Syr Dayra River, Tajikistan. Non-types: ZIN 7870 (2, lost). On p. 3 of separate. •Valid as Cottus spinulosus Kessler 1872 -- (Berg 1949:1150 [ref. 20662], Goto et al. 2014:[11] [ref. 33456]). Current status: Valid as Cottus spinulosus Kessler 1872. Cottidae. Distribution: Europe. Habitat: freshwater.

spitzbergensis, Icelus bicornis Schmidt [P. Yu.] 1927:7 [Ezegodnik Zoologiceskogo Muzeja = Annuaire du Musée Zoologique / Akademija Nauk Sojuza Sovetskich Socialisticeskich Respublik. v. 28; ref. 19815] Spitzbergen. Syntypes: ZIN 11867 (3), 11900-02 (1, 4, 6+), 12750-52 (3, 2, 5), 12754-55 (13, 3), 12807-08 (1, 1), 13993-97 (6+, 6+, 1, 1, 1). •Synonym of Icelus bicornis (Reinhardt 1840) -- (Neyelov 1973:596 [ref. 7219], Nelson 1984:36 [ref. 5391], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:253 [ref. 33547]). Current status: Synonym of Icelus bicornis (Reinhardt 1840). Cottidae. Habitat: marine.

stejnegeri, Stelgistrum Jordan [D. S.] & Gilbert [C. H.] in Jordan & Evermann 1898:1921 [Bulletin of the United States National Museum No. 47; ref. 2444] Off Robben Island, Okhotsk Sea [not Bering Sea], Albatross station 3645, depth 10 fathoms. Holotype (unique): USNM 48236. Also appeared in Jordan & Gilbert 1899:456, Pl. 54 [ref. 2478] as steinegeri. •Valid as Stelgistrum stejnegeri Jordan & Gilbert 1898 -- (Watanabe 1960:32 [ref. 11959], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:195 [ref. 15964] as steinegeri, Sokolovskaya et al. 1998:10 [ref. 24670] as steinegeri, Nakabo 2000:636 [ref. 25086], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:447 [ref. 25968], Nakabo 2002:636 [ref. 26001], Parin et al. 2002:S86 [ref. 28300], Fedorov et al. 2003:80 [ref. 27579], Tsuroka et al. 2009:97 [ref. 32107], Shinohara et al. 2014:251 [ref. 33330], Parin et al. 2014:273 [ref. 33547], Dyldin & Orlov 2017:67 [ref. 35138]). Current status: Valid as Stelgistrum stejnegeri Jordan & Gilbert 1898. Cottidae. Distribution: Western North Pacific. Habitat: brackish, marine.

stelleri, Cottus Bloch [M. E.] & Schneider [J. G.] 1801:63 [M. E. Blochii, Systema Ichthyologiae; ref. 471] Mentioned in passing, not available. •In the synonymy of Ceratocottus [now Enophrys] diceraus (Pallas 1787) -- (Watanabe 1960:110 [ref. 11959]). •In the synonymy of Enophrys diceraus (Pallas 1787) -- (Sandercock & Wilimovsky 1968:835 [ref. 26414]). Current status: Synonym of Enophrys diceraus (Pallas 1787). Cottidae.

stelleri, Myoxocephalus Tilesius [W. G. von] 1811:273, Pl. 13 (fig. 1) [Mémoires de l'Académie Impériale des Sciences de Saint Pétersbourg v. 3; ref. 4408] Kamchatka, Russia. No types known. •Valid as Myoxocephalus stelleri Tilesius 1811 -- (Berg 1949:1137 [ref. 20662], Neyelov 1979:121 [ref. 3152], Yabe in Masuda et al. 1984:326 [ref. 6441], Lindberg & Krasyukova 1987:207 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Kim 1997:407 [ref. 24032], Sokolovsky & Sokolovskaya 1997:58 [ref. 24691], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:153 [ref. 23595], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:645 [ref. 25086], Sheiko & Fedorov 2000:28 [ref. 25650], Pietsch et al. 2001:145 [ref. 25688], Zemnukhov et al. 2001:420 [ref. 25412], Kim & Park 2002:327 [ref. 25913], Mecklenburg et al. 2002:480 [ref. 25968], Nakabo 2002:645 [ref. 26001], Youn 2002:262, 576 [ref. 26218], Parin et al. 2002:S82 [ref. 28300], Hagiwara 2003:37 [ref. 27188], Fedorov et al. 2003:77 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Bogutskaya & Naseka 2004:191 [ref. 28183], Page et al. 2013:124 [ref. 32708], Parin et al. 2014:265 [ref. 33547], Kells et al. 2016:186 [ref. 35888], Dyldin & Orlov 2017:66 [ref. 35138]). Current status: Valid as Myoxocephalus stelleri Tilesius 1811. Cottidae. Distribution: North Pacific: Aleutian Islands to southeastern Alaska, and Commander Islands to Okhotsk Sea and Japan Sea to South Korea. Habitat: brackish, marine.

stenosomus, Icelus uncinalis Andriashev [A. P.] 1937:266, Pl. 5 (fig. 3) [Zoologische Jahrbücher, Abteilung für Systematik, Geographie und Biologie der Tiere (Jena) v. 69 (no. 4); ref. 15267] 48°08'30"N, 140°08'30'E, Tatar Strait. Holotype: ZIN 17646. Paratypes: ZIN 17641-42 (6, 2), 17645 (4), 17646A (5, with holotype), 17647 (3), 21906-08 (1, 1, 1). "Typus" given on caption to Pl. 5, specimen noted in catalog in 17646 and regarded as holotype. •Synonym of Icelus uncinalis Gilbert & Burke 1912, but a valid subspecies as described -- (Lindberg & Krasyukova 1987:194 [ref. 15964], Sokolovskaya et al. 1998:9 [ref. 24670]). •Valid as Icelus stenosomus Andriashev 1937 -- (Nelson 1984:53 [ref. 5391], Murai et al. 1994 [ref. 21908], Nakabo 2000:635 [ref. 25086], Nakabo 2002:635 [ref. 26001], Parin et al. 2002:S78 [ref. 28300], Shinohara et al. 2014:250 [ref. 33330], Parin et al. 2014:257 [ref. 33547]). Current status: Valid as Icelus stenosomus Andriashev 1937. Cottidae. Distribution: Western North Pacific. Habitat: marine.

stimpsonii, Triglopsis Hoy [P. R.] 1872:99 [Transactions of the Wisconsin Academy of Sciences Arts and Letters 1870-72; ref. 18119] Lake Michigan, off Racine, Milwaukee, U.S.A. (stomach content). Holotype (unique): whereabouts unknown. Apparently not available; appeared above and also as Gill (MS) in Jordan 1878:50 [ref. 10444] as stimpsoni, both without distingusihing features. Subsequent availability not researched. •In the synonymy of Myoxocephalus quadricornis (Linnaeus 1758) -- (Scott & Crossman 1998:846 [ref. 24570]). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae.

strabo, Icelinus Starks [E. C.] 1896:551 [Proceedings of the California Academy of Sciences (Series 2) v. 6; ref. 4195] Port Ludlow, Washington, U.S.A. Syntypes: BMNH 1897.1.12.19-20 (2), CAS-SU 5045 (about 25, now 19). Type catalog: Böhlke 1953:130 [ref. 12291]. •Synonym of Icelinus borealis Gilbert 1896 -- (Bolin 1936:159 [ref. 506], Rosenblatt & Smith 2004:560 [ref. 27912]). Current status: Synonym of Icelinus borealis Gilbert 1896. Cottidae. Habitat: marine.

subborealis, Triglops pingelii var. Le Danois [E.] 1914:11 [Annales de l'Institut Océanographique Monaco v. 7 (no. 2); ref. 21190] Fair Isle, Shetland Islands, North Sea, 59°26'N, 1°30'W, depth 80 meters. Holotype (unique): whereabouts unknown, not at MNHN. •Synonym of Triglops murrayi Günther 1888 -- (Neyelov 1973:602 [ref. 7219], Pietsch 1994:369 [ref. 21189]). Current status: Synonym of Triglops murrayi Günther 1888. Cottidae. Habitat: marine.

subulatus, Abyssocottus gibbosus var. Dybowski [B. N.] 1908:552 [17], Fig. 12 [Kosmos Lwów v. 33; ref. 17422] Lake Baikal, Russia. Syntypes: whereabouts unknown. •Synonym of Abyssocottus gibbosus Berg 1906 -- (Berg 1949:1157 [ref. 20662]). Current status: Synonym of Abyssocottus gibbosus Berg 1906. Cottidae. Habitat: freshwater.

szanaga, Cottus Dybowski [B. N.] 1869:949, Pl. 14 (fig. 1) [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 19; ref. 1169] Onon River and its tributaries, 51°42'N, 115°50'E, Amur River basin, Russia. Syntypes: ZMB 7107 (1). Type catalog: Paepke & Fricke 1992:284 [ref. 19946]. •Synonym of Cottus poecilopus Heckel 1837 -- (Berg 1949:1143 [ref. 20662] as czanaga, Fedorov in Whitehead et al. 1986:1248 [ref. 13677] as czanaga). •Valid as Cottus szanaga Dybowski 1869 -- (Shedko & Shedko 2003:330 [ref. 27966], Bogutskaya & Naseka 2004:188 [ref. 28183], Kottelat 2006:60 [ref. 28831], Ocock et al. 2006:57 [ref. 28933], Shedko & Miroshnichenko 2007:24 [ref. 29369], Bogutskaya et al. 2008:352 [ref. 30085], Sideleva & Goto 2009:603 [ref. 30591], Goto et al. 2014:[9] [ref. 33456], Parin et al. 2014:241 [ref. 33547], Parin et al. 2014:245 [ref. 33547], Dyldin & Orlov 2017:63 [ref. 35138]). Current status: Valid as Cottus szanaga Dybowski 1869. Cottidae. Distribution: Upper Amur River basin: China, Russia, Mongolia and Siberia. Habitat: freshwater.

taeniopterus, Cottus Kner [R.] 1868:28 [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 58 (nos 1-2); ref. 6074] Decastris Bay near Amur River mouth, northeastern Asia. Holotype (unique): NMW 11612. Appeared in more detail in Kner 1868:310 [p. 18 of separate], Pl. 4 (fig. 10) [ref. 2646]. •Synonym of Myoxocephalus platycephalus (Pallas 1814), but a valid subspecies -- (Berg 1949:1128 [ref. 20662], Zhu 1995:189 [ref. 25213]). •Synonym of Megalocottus platycephalus (Pallas 1814), but a valid subspecies -- (Neyelov 1979:130 [ref. 3152], Lindberg & Krasyukova 1987:212 [ref. 15964], Nikiforov et al. 1997:330 [ref. 24694], Reshetnikov et al. 1997:759 [ref. 24702], Sideleva 1998:153 [ref. 23595], Sokolovskaya et al. 1998:9 [ref. 24670], Bogutskaya & Naseka 2004:190 [ref. 28183]). •Synonym of Megalocottus platycephalus (Pallas 1814) -- (Mecklenburg et al. 2002:475 [ref. 25968], Parin et al. 2002:S79 [ref. 28300], Mecklenburg et al. 2016:73 [ref. 34440]). •Valid as Megalocottus taeniopterus (Kner 1868) -- (Bogutskaya et al. 2008:352 [ref. 30085], Parin et al. 2014:258 [ref. 33547], Dyldin & Orlov 2017:64 [ref. 35138]). Current status: Valid as Megalocottus taeniopterus (Kner 1868). Cottidae. Distribution: Asia/western North Pacific. Habitat: marine.

talievi, Batrachocottus Sideleva [V. G.] 1999:150 [204], Fig. 1 [Voprosy Ikhtiologii v. 39 (no. 2); ref. 24864] Lake Baikal, northern outlet of the Strait of Maloe More, depth 350-370 meters. Holotype: ZIN 13750 [radiograph no. 19614]. Paratypes: ZIN 51524/A (5). On p. 204 of English translation. •Valid as Batrachocottus talievi Sideleva 1999 -- (Bogutskaya et al. 2001:46 [ref. 26178], Sideleva 2001:58 [ref. 26181], Sideleva 2003:15, 159 [ref. 27815], Bogutskaya & Naseka 2004:197 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Batrachocottus talievi Sideleva 1999. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

tallapoosae, Cottus Neely [D. A.], Williams [J. D.] & Mayden [R. L.] 2007:642, Figs. 1A, 2 [Copeia 2007 (no. 3); ref. 29262] 2 kilometers southwest of Red Hill, Gold Banch of Channahatchee Creek at County road 407, 32°40'17"N, 85°57'36"W, Elmore County, Alabama, USA. Holotype: UAIC 13545.03. Paratypes: Many lots at AUM, GMNH, TU, UAIC, UF, UT and USNM. Plus many non-type specimens mentioned in Material Examined. •Valid as Cottus tallapoosae Neely, Williams & Mayden 2007 -- (Scharpf 2008:11 [ref. 30399], Page & Burr 2011:477 [ref. 31215], Page et al. 2013:122 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus tallapoosae Neely, Williams & Mayden 2007. Cottidae. Distribution: Tallapoosa River drainage above the Fall Line in Alabama and Georgia, U.S.A. Habitat: freshwater.

taranetzi, Radulinopsis Yabe [M.] & Maruyama [S.] 2001:56, Figs. 3-4 [Ichthyological Research v. 48 (no. 1); ref. 25245] Off Usujiri, 41°54'N, 140°05'E, Pacific coast of southern Hokkaido, Japan, depth 15-20 meters. Holotype: HUMZ 150550. Paratypes: HUMZ 87631 (1), 87770 (1), 92778-79 (2), 92781-83 (3), 92785-88 (4), 113656-61 (6), 121685-86 (2), 121719 (1), 135986 (1), 145741-48 (8), 150549 (1), 150551-52 (2), 150560-61 (2); NSMT-P 57640-41 (1, 1). •Valid as Radulinopsis taranetzi Yabe & Maruyama 2001 -- (Parin et al. 2014:271 [ref. 33547]). Current status: Valid as Radulinopsis taranetzi Yabe & Maruyama 2001. Cottidae. Distribution: Western North Pacific. Habitat: marine.

taurinus, Enophrys Gilbert [C. H.] 1914:135, Pl. 11 (fig. 1) [Proceedings of the United States National Museum v. 47 (no. 2049); ref. 14153] Monterey Bay, near Pacific Grove, California, U.S.A. Holotype: USNM 75064. Paratypes: CAS-SU 57 (1); plus additional material. Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Aspicottus taurinus (Gilbert 1914) -- (Neyelov 1979:148 [ref. 3152]). •Valid as Enophrys taurinus Gilbert 1914 -- (Bolin 1944:93 [ref. 6379]). •Valid as Enophrys taurina Gilbert 1914 -- (Sandercock & Wilimovsky 1968:844 [ref. 26414], Eschmeyer & Herald 1983:169 [ref. 9277], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Kells et al. 2016:170 [ref. 35888]). Current status: Valid as Enophrys taurina Gilbert 1914. Cottidae. Distribution: Eastern North Pacific: California, U.S.A. Habitat: marine.

taylori, Asemichthys Gilbert [C. H.] 1912:215 [Contributions to Canadian Biology being studies from the Marine Biological Stations of Canada 1906-1910 (art. 12); ref. 9276] Departure Bay, Vancouver Island, British Columbia, Canada. Holotype (unique): CAS-SU 23071. Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Radulinus taylori (Gilbert 1912) -- (Eschmeyer & Herald 1983:181 [ref. 9277], Mecklenburg et al. 2002:444 [ref. 25968], Page et al. 2013:121 [ref. 32708], Kells et al. 2016:190 [ref. 35888]). •See Yabe & Maruyama 2001:59 [ref. 25245]. •Valid as Asemichthys taylori Gilbert 1912 -- (McAllister 1990:194 [ref. 14674], Coffie 1998:130 [ref. 25001], Nelson et al. 2004:118 [ref. 27807], Pietsch & Orr 2015:39 [ref. 34694]). Current status: Valid as Asemichthys taylori Gilbert 1912. Cottidae. Distribution: Eastern North Pacific: southeastern Alaska to San Juan Islands, northern Washington. Habitat: marine.

tentaculatus, Cottus Kner [R.] 1868:28 [Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe v. 58 (nos 1-2); ref. 6074] Singapore. Holotype (unique): NMW 11621. Appeared in more detail in Kner 1868:314 [p. 22 of separate], Pl. 5 (fig. 12) [ref. 2646]. •Valid as Porocottus tentaculatus (Kner 1868) -- (Neyelov 1979:137 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:224 [ref. 15964], Kim & Youn 1992:63 [ref. 22287], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:646 [ref. 25086], Nakabo 2002:646 [ref. 26001], Muto et al. 2002:231 [ref. 26331], Parin et al. 2002:S84 [ref. 28300], Fedorov et al. 2003:80 [ref. 27579], Parin et al. 2014:270 [ref. 33547], Dyldin & Orlov 2017:67 [ref. 35138]). Current status: Valid as Porocottus tentaculatus (Kner 1868). Cottidae. Distribution: Western North Pacific. Habitat: brackish, marine.

tenuis, Icelinus Gilbert [C. H.] 1890:86 [Proceedings of the United States National Museum v. 13 (no. 797); ref. 1623] Off southern California, U.S.A., 5 Albatross stations, depth 45-150 fathoms. Syntypes: CAS-SU 78 (3), 28110 (1); USNM 43086 (1). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelinus tenuis Gilbert 1890 -- (Bolin 1944:26 [ref. 6379], Eschmeyer & Herald 1983:173 [ref. 9277], Peden 1984:79 [ref. 5193], McAllister 1990:198 [ref. 14674], Mecklenburg et al. 2002:438 [ref. 25968], Rosenblatt & Smith 2004:560 [ref. 27912], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:41 [ref. 34694], Girard & Smith 2016:556 [ref. 34786], Kells et al. 2016:178 [ref. 35888]). Current status: Valid as Icelinus tenuis Gilbert 1890. Cottidae. Distribution: Eastern North Pacific: southeastern Alaska to central Baja California. Habitat: marine.

tenuis, Uranidea Evermann [B. W.] & Meek [S. E.] 1898:83, Fig. 6 [Bulletin of the U. S. Fish Commission v. 17 (art. 2) [1897]; ref. 14683] Lower end of upper Klamath Lake near Klamath Falls, Oregon, U.S.A. Holotype: USNM 48229. Paratypes or non-types: ANSP 39142 (1, missing), BMNH 1900.9.29.198-199 (2), CAS-SU 5705 (20), USNM 124958 [ex BCF 434] (39). Type catalog: Böhlke 1953:133 [ref. 12291], Böhlke 1984:66 [ref. 13621]. •Valid as Cottus tenuis (Evermann & Meek 1898) -- (Lee et al. 1980:824 [ref. 22416], Page & Burr 1991:248 [ref. 18983], Nelson et al. 2004:119 [ref. 27807], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:473 [ref. 31215], Page et al. 2013:123 [ref. 32708], Goto et al. 2014:[10] [ref. 33456]). Current status: Valid as Cottus tenuis (Evermann & Meek 1898). Cottidae. Distribution: Klamath River basin, Oregon, U.S.A. Habitat: freshwater.

terraenovae, Triglops ommatistius Gilbert [C. H.] 1913:467 [Proceedings of the United States National Museum v. 44 (no. 1963); ref. 14957] Newfoundland, Canada, 45°09'30"N, 49°48'30"W, Albatross station 2445, depth 39 fathoms. Holotype: USNM 46063. Paratypes: (orig. 57, from 11 stations) USNM 46059 (1), 46061-62 (1, 3), 46064-68 (?36), 46069 (1), 15089 (17). •Synonym of Triglops murrayi Günther 1888 -- (Scott & Scott 1988:510 [ref. 25518], Pietsch 1994:368 [ref. 21189], Parin et al. 2014:276 [ref. 33547]). Current status: Synonym of Triglops murrayi Günther 1888. Cottidae. Habitat: marine.

thermalis, Neocottus Sideleva [V. G.] 2002:275, Fig. [Voprosy Ikhtiologii v. 42 (no. 2); ref. 26019] Lake Baikal, depth 430-480 meters. Holotype: ZIN 52196. Paratypes: USNM 360863 (1); ZIN 52197 (4 or 52), ?52219 (23). Sideleva 2001:69 [ref. 26181] indicates that the holotype is ZIN 52197 (52), paratype: ZIN 52219 (23). Appeared first without distinguishing features in Bogutskaya et al. 2001:47 [ref. 26178] and apparently also in Sideleva 2001:69 [ref. 26181]. •Valid as Neocottus thermalis Sideleva 2002 -- (Bogutskaya et al. 2001:47 [ref. 26178] [but not dating to this account], Sideleva 2003:19, 201 [ref. 27815], Bogutskaya & Naseka 2004:199 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Neocottus thermalis Sideleva 2002. Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

thoburni, Paricelinus Gilbert [C. H.] 1896:432, Pl. 30 (middle) [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Off Oregon, U.S.A., Albatross station 3350, depth 75 fathoms. Holotype (unique): USNM 64324. •Synonym of Paricelinus hopliticus Eigenmann & Eigenmann 1889 -- (Bolin 1944:11 [ref. 6379], Mecklenburg et al. 2002:429 [ref. 25968]). Current status: Synonym of Paricelinus hopliticus Eigenmann & Eigenmann 1889. Cottidae. Habitat: marine.

thompsonii, Triglopsis Girard [C. F.] 1851:19 [Proceedings of the Boston Society of Natural History v. 4 (1851-1854); ref. 1816] Lake Ontario off Oswego, New York, U.S.A. Syntypes: MCZ 35979 [ex USNM 3175] (1), USNM 3175 (8, poor condition). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758), but a valid subspecies -- (Scott & Crossman 1998:843 [ref. 24570], Goto et al. 2014: [6] [ref. 33456]). •Valid as Myoxocephalus thompsonii (Girard 1851) -- (Lee et al. 1980:827 [ref. 22416], Smith 1986:359 [ref. 22186], McAllister 1990:200 [ref. 14674] as thompsoni, Page & Burr 1991:244 [ref. 18983] as thompsoni, Bailey et al. 2004:173 [ref. 27882], Nelson et al. 2004:121 [ref. 27807], Murray et al. 2004:642 [ref. 27932], Scharpf 2008:11 [ref. 30399], Page & Burr 2011:467 [ref. 31215], Page et al. 2013:124 [ref. 32708]). Current status: Valid as Myoxocephalus thompsonii (Girard 1851). Cottidae. Distribution: Canada and New York (U.S.A.). Habitat: freshwater.

tilesii, Hemilepidotus Cuvier [G.] in Cuvier & Valenciennes 1829:276, Pl. 85 [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka, Russia. Lectotype: MNHN 0000-7155. Type catalog: Blanc & Hureau 1968:52 [ref. 20738] with 7155 as holotype; based on literature sources and one specimen; MNHN 7155 designated lectotype by Eschmeyer 1998:1679 [ref. 23416]. Apparently based on Cottus hemilepidotus Tilesius and Cottus trachurus Pallas. Type information: Svetovidov 1978:24 [ref. 6025] with 7155 as holotype. •Synonym of Hemilepidotus hemilepidotus (Tilesius 1811) -- (Bolin 1944:17 [ref. 6379], Peden 1979:24 [ref. 24080], Parin et al. 2002:S76 [ref. 28300], Parin et al. 2014:250 [ref. 33547]). Current status: Synonym of Hemilepidotus hemilepidotus (Tilesius 1811). Cottidae. Habitat: marine.

totomius, Pseudoblennius Jordan [D. S.] & Starks [E. C.] 1904:315, Fig. 36 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Off Omi Point, Totomi Bay, Japan, Albatross station 3729, depth 34 fathoms. Holotype (unique): USNM 50928. •Valid as Pseudoblennius totomius Jordan & Starks 1904 -- (Nakabo 2000:649 [ref. 25086], Nakabo 2002:649 [ref. 26001]). Current status: Valid as Pseudoblennius totomius Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

toyamensis, Ricuzenius (Alloricuzenius) Matsubara [K.] & Iwai [T.] 1951:87, Figs. 1-3 [Miscellaneous Reports of the Research Institute for Natural Resources (Tokyo) Nos 19-21; ref. 2909] Off Uozu, Japan. Holotype: FAKU 14450 (not seen in 1997). Paratypes: FAKU 14451-57 (7, not seen in 1997). •Valid as Ricuzenius toyamensis Matsubara & Iwai 1951 -- (Watanabe 1960:22 [ref. 11959], Yabe in Masuda et al. 1984:324 [ref. 6441], Lindberg & Krasyukova 1987:186 [ref. 15964]). •Valid as Icelus toyamensis (Matsubara & Iwai 1951) -- (Nakabo 2000:633 [ref. 25086], Nakabo 2002:633, 1525 [ref. 26001], Tsuruoka et al. 2006:51 [ref. 28501], Shinohara et al. 2011:44 [ref. 31715], Song et al. 2012:304 [ref. 36295], Shinohara et al. 2014:250 [ref. 33330]). Current status: Valid as Icelus toyamensis (Matsubara & Iwai 1951). Cottidae. Distribution: Western North Pacific: southern Korea to Japan. Habitat: marine.

trachurus, Cottus Pallas [P. S.] 1814:138 [Zoographia Rosso-Asiatica v. 3; ref. 3351] Kuril Islands. Lectotype: ZMB 4755. Type information: Svetovidov 1978:26 [ref. 6025]. Lectotype selected by Svetovidov 1978:26 [ref. 6025]. •In part (Kuriles) a synonym of Hemilepidotus gilberti Jordan & Starks 1904 -- (Peden 1979:36 [ref. 24080]). •Synonym of Hemilepidotus hemilepidotus (Tilesius 1811) -- (Bolin 1944:17 [ref. 6379], Peden 1979:24 [ref. 24080] as in part, Parin et al. 2002:S76 [ref. 28300], Parin et al. 2014:250 [ref. 33547]). Current status: Synonym of Hemilepidotus hemilepidotus (Tilesius 1811). Cottidae. Habitat: marine.

transsilvaniae, Cottus Freyhof [J.], Kottelat [M.] & Nolte [A.] 2005:157, Figs. 28-29 [Ichthyological Exploration of Freshwaters v. 16 (no. 2); ref. 28259] River Bratia at bridge on road 73c, about 20 kilometers southwest of village Cimpulung, 45°13'N, 24°47'E, Rumania. Holotype: ZMB 33596. Paratypes: FSFJ 686 (8). •Valid as Cottus transsilvaniae Freyhof, Kottelat & Nolte 2005 -- (Kottelat & Freyhof 2007:518 [ref. 29996], Sideleva 2009:211 [ref. 30173], Goto et al. 2014:[11] [ref. 33456], Sideleva et al. 2015:33 [ref. 35276], Sideleva et al. 2018:9 [ref. 36393]). Current status: Valid as Cottus transsilvaniae Freyhof, Kottelat & Nolte 2005. Cottidae. Distribution: Romania. Habitat: freshwater.

triacis, Orthonopias Starks [E. C.] & Mann [W. M.] 1911:11, Fig. 1 [University of California Publications in Zoology v. 8 (no. 2); ref. 4199] Cortez Bank, off California, U.S.A., depth 11-16 fathoms. Holotype (unique): SIO 60-56. •Valid as Orthonopias triacis Starks & Mann 1911 -- (Bolin 1944:59 [ref. 6379], Eschmeyer & Herald 1983:179 [ref. 9277], Fedorov 1990:745 [ref. 20136], Long 1992:160 [ref. 22238], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708], Kells et al. 2016:188 [ref. 35888]). Current status: Valid as Orthonopias triacis Starks & Mann 1911. Cottidae. Distribution: Eastern North Pacific: northern California (U.S.A.) to central Baja California. Habitat: marine.

tribranchius, Atopocottus Bolin [R. L.] 1936:30, Fig. 6 [Proceedings of the United States National Museum v. 84 (no. 3000); ref. 505] Sea of Japan, 38°14'N, 138°54'E, Albatross station 4816, depth 64 fathoms. Holotype: USNM 102105. Paratypes: CAS-SU 30457 (3), USNM 102106 (7). Type catalog: Böhlke 1953:127 [ref. 12291]. •Valid as Atopocottus tribranchius Bolin 1936 -- (Yabe in Masuda et al. 1984:329 [ref. 6441], Lindberg & Krasyukova 1987:241 [ref. 15964], Nakabo 2000:639 [ref. 25086], Nakabo 2002:639 [ref. 26001], Matsunuma et al. 2017:93 [ref. 35372]). Current status: Valid as Atopocottus tribranchius Bolin 1936. Cottidae. Distribution: Western North Pacific: Japan and East China Sea. Habitat: marine.

tricuspis, Cottus Reinhardt [J. C. H.] 1830:16 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1829-30; ref. 19480] Greenland. Holotype: ZMUC uncat. Type catalog: Nielsen 1974:81 [ref. 9588]. •Valid as Gymnocanthus tricuspis (Reinhardt 1830) -- [sometimes as Gymnacanthus] (Neyelov 1973:595 [ref. 7219], Fedorov in Whitehead et al. 1986:1249 [ref. 13677], Robins & Ray 1986:281 [ref. 23100], Scott & Scott 1988:495 [ref. 25518], Matarese et al. 1989:394 [ref. 26885], McAllister 1990:197 [ref. 14674], Yabe in Okamura et al. 1995:163 [ref. 22531], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Klein-MacPhee in Collette & Klein-MacPhee 2002:347 [ref. 26158], Mecklenburg et al. 2002:464 [ref. 25968], Parin et al. 2002:S75 [ref. 28300] as Gymnacanthus Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2007:174 [ref. 29448], Møller et al. 2010:52 [ref. 31169], Page et al. 2013:123 [ref. 32708], Dolgov 2013:918 [ref. 33112], Parin et al. 2014:249 [ref. 33547], Mecklenburg & Steinke 2015:164 [ref. 34033], Afonso et al. 2016:41, 163 [ref. 36199], Kells et al. 2016:172 [ref. 35888], Mecklenburg et al. 2016:61 [ref. 34440], Mecklenburg et al. 2018:199 [ref. 36370]). Current status: Valid as Gymnocanthus tricuspis (Reinhardt 1830). Cottidae. Distribution: North Atlantic and Arctic Ocean; Circumpolar. Habitat: marine.

trigonocephalus, Cottus Gratzianov [V. I.] 1902:32 [Dnevnik' Zoologicheskogo Otdelenya Obshchestva Lyubiteley Estestvoznaniya, Antropologii i Etnografii, Moskva v. 3 (no. 3); ref. 1868] Ushkan Island, Lake Baikal, Russia, depth 280 feet (40 Russian fathoms). Holotype (unique): ZMMU P-3123. Type catalog: Vasil'eva et al. 2001:80 [ref. 25714]. •Synonym of Cottus kesslerii Dybowski 1874 -- (Berg 1949:1153 [ref. 20662] as kessleri, Sideleva 1982:27 [ref. 14469] as kessleri, Sideleva 1998:151 [ref. 23595]). •Synonym of Leocottus kesslerii (Dybowski 1874) -- (Sideleva 2001:59 [ref. 26181], Sideleva 2003:16, 151 [ref. 27815], Kottelat 2006:60 [ref. 28831]). Current status: Synonym of Leocottus kesslerii (Dybowski 1874). Cottidae. Habitat: freshwater.

tuberculatus, Myoxocephalus Soldatov [V. K.] & Pavlenko [M. N.] 1922:342, Fig. 1 [Ezhegodnik. Zoologicheskogo Muzeya Rossiskoi Akademii Nauk v. 23; ref. 15431] Many localities, near Shantarskiye [Shantar] Islands, western Okhotsk Sea, depth 10-87 meters. Lectotype: ZIN 20615. Paralectotypes: (26) ZIN 19430 (6), 20614 (2, lost), 20615-24 (2, 4, 1, 2, 4, 2, 2, 2, 1, 1). Lectotype selected by Neyelov 1979:122 [ref. 3152]. •Valid as Myoxocephalus tuberculatus Soldatov & Pavlenko 1922 -- (Parin et al. 2014:265 [ref. 33547]). Current status: Valid as Myoxocephalus tuberculatus Soldatov & Pavlenko 1922. Cottidae. Distribution: Western North Pacific. Habitat: marine.

tubulatus, Cottus Hubbs [C. L.] & Schultz [L. P.] 1932:2 [Occasional Papers of the Museum of Zoology University of Michigan No. 242; ref. 16378] A tributary to north fork of the Clearwater River, near Bovill, Idaho, U.S.A. Holotype: UMMZ 92342. Paratypes: UMMZ 92669 (2). •Synonym of Cottus beldingii Eigenmann & Eigenmann 1891 -- (Bailey & Bond 1963:1 [ref. 3934], Lee et al. 1980:806 [ref. 22416], Baxter & Stone 1995:223 [ref. 23374]). Current status: Synonym of Cottus beldingii Eigenmann & Eigenmann 1891. Cottidae. Habitat: freshwater.

typicus, Triglops pingelii var. Le Danois [E.] 1914:11 [Annales de l'Institut Océanographique Monaco v. 7 (no. 2); ref. 21190] Unneeded name for the typical form pingelii described by Reinhardt. •Synonym of Triglops pingelii Reinhardt 1837 -- (Pietsch 1994:374 [ref. 21189]). Current status: Synonym of Triglops pingelii Reinhardt 1837. Cottidae. Habitat: marine.

uchidai, Triglops Watanabe [M.] 1958:201, Figs. 61-62, Pl. 2 [Studies of Japanese cottid fishes; ref. 21357] Mogami-tai, Yamagata Prefecture, Japan, depth 70 fathoms. Holotype (unique): not researched. Also described as new in Watanabe 1960:11, pl. 16 (fig. 2) [ref. 11959]. •Valid as Triglops uchidai Watanabe 1958 -- (Yabe in Masuda et al. 1984:325 [ref. 6441], Lindberg & Krasyukova 1987:179 [ref. 15964]). •Synonym of Triglops scepticus Gilbert 1896 -- (Pietsch 1994:358 [ref. 21189], Nakabo 2002:1525 [ref. 26001], Parin et al. 2014:278 [ref. 33547]). Current status: Synonym of Triglops scepticus Gilbert 1896. Cottidae. Habitat: marine.

uncinalis, Icelus Gilbert [C. H.] & Burke [C. V.] 1912:39, Figs. 2, 2a [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Near Attu Island, Aleutian Islands, Bering Sea, 52°55'40"N, 173°26'E, Albatross station 4784, depth 135 fathoms. Holotype: USNM 74366. Paratypes: CAS-SU 25237 (2); USNM 70855 (1), 70885 (2), 70892 (1), 70897 (2), 70912 (3). Type catalog: Böhlke 1953:130 [ref. 12291]. •Valid as Icelus uncinalis Gilbert & Burke 1912 -- (Nelson 1984:52 [ref. 5391], Kim & Youn 1992:67 [ref. 22287], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:454 [ref. 25968], Youn 2002:262, 576 [ref. 26218], Parin et al. 2002:S79 [ref. 28300], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:123 [ref. 31212], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:257 [ref. 33547]). Current status: Valid as Icelus uncinalis Gilbert & Burke 1912. Cottidae. Distribution: North Pacific: Bering Sea north of Commander Islands and western Aleutian Islands. Habitat: marine.

uncinatus, Cottus Temminck [C. J.] & Schlegel [H.] 1843:38 [Fauna Japonica Parts 2-4; ref. 4371] Japan. Lectotype: RMNH 536a. Paralectotypes: RMNH 536b-c (2). Objectively invalid; preoccupied by Cottus uncinatus Reinhardt 1835. Lectotype selected by Boeseman 1947:48 [ref. 12876]. •Synonym of Trachidermus fasciatus Heckel 1839 -- (Okada 1961:709 [ref. 6363]). Current status: Synonym of Trachidermus fasciatus Heckel 1839. Cottidae. Habitat: freshwater, brackish, marine.

uncinatus, Cottus Reinhardt [J. C. H.] 1834:2-3 [Oversigt over det Kongelige Danske Videnskabernes Selskabs Forhandlinger og dets Medlemmers Arbeider (Kjøbenhavn) 1833-34; ref. 26882] Nanortalik, District of Julianehåb, Greenland. Syntypes: ZMUC (1), plus one syntype lost. Type catalog: Nielsen 1974:81 [ref. 9588]. Also appeared in Kgl. Danske Vidensk. Selsk. Natur. Math. Afhandl., 1837, v. 6:xliv. •Valid as Artediellus uncinatus (Reinhardt 1834) -- (Neyelov 1973:593 [ref. 7219], Neyelov 1979:153 [ref. 3152], Fedorov in Whitehead et al. 1986:1246 [ref. 13677], Nelson 1986:43 [ref. 5808], Robins & Ray 1986:281 [ref. 23100], Scott & Scott 1988:491 [ref. 25518], McAllister 1990:194 [ref. 14674], Yabe in Okamura et al. 1995:162 [ref. 22531], Mecklenburg et al. 2002:403 [ref. 25968], Nelson et al. 2004:118 [ref. 27807] dating to 1835, Møller et al. 2010:51 [ref. 31169] dated to 1835, Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:121 [ref. 32708], Afonso et al. 2016:41, 101 [ref. 36199], Mecklenburg et al. 2018:190 [ref. 36370]). Current status: Valid as Artediellus uncinatus (Reinhardt 1834). Cottidae. Distribution: North Atlantic and Arctic Ocean. Habitat: marine.

uschkani, Batrachocottus Taliev [D. N.] 1955:51 et seq., 263, Figs. 56, 105 [ ref. 4316] Lake Baikal, Russia. Syntypes: none found at ZIN. Appeared without distinguishing features and unavailable in Taliev 1948:113 [ref. 19628] and spelled ushkani. •Synonym of Batrachocottus baicalensis (Dybowski 1874) -- (Sideleva 1982:28 [ref. 14469], Sideleva 1998:149 [ref. 23595], Sideleva 1999:149 [ref. 24864], Sideleva 2001:57 [ref. 26181], Sideleva 2003:15 [ref. 27815] dated 1948). Current status: Synonym of Batrachocottus baicalensis (Dybowski 1874). Cottidae. Habitat: freshwater.

ushkani, Batrachocottus Taliev [D. N.] 1948:113 [Trudy Baikal'skoi Limnologicheskoi Stantsii v. 12; ref. 19628] Not available, manuscript name mentioned in table. In the synonymy of Batrachocottus baicalensis (Dybowskii 1874). •Described in an available way as new in Taliev 1955:263 [ref. 6314] as Batrachocottus uschkani. Current status: Synonym of Batrachocottus baicalensis (Dybowski 1874). Cottidae.

vaenernensis, Cottus quadricornis Lönnberg [E.] 1932:8, Fig. 3 [Arkiv för Zoologi v. 24 A (pt 2) (no. 7); ref. 16380] Lake Vänern, Sweden. Syntypes: NRM 20336-38 (1, 4, 1), 40338 (1). •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:170 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

vandesandei, Gymnocanthus Poll [M.] 1949:234, Figs. 13-14 [Mémoires, Institut Royal des Sciences Naturelles de Belgique 2.Séries, No. 33; ref. 13028] Pulpito Bay, Rio de Oro, Spanish Sahara, depth 10-15 fathoms. Holotype (unique): IRSNB 91. Type catalog: Walschaerts 1987:25 [ref. 20755], Derijst 1991:43 [ref. 21528]. •Valid as Gymnocanthus vandesandei Poll 1949. Current status: Valid as Gymnocanthus vandesandei Poll 1949. Cottidae. Distribution: Eastern Atlantic. Habitat: marine.

variabilis, Cottus Ayres [W. O.] 1842:68 [Proceedings of the Boston Society of Natural History v. 1 (1841-1844); ref. 15926] Brookhaven, Long Island, New York, U.S.A. Appeared first as name only in Ayres 1842:58 [ref. 15927]. Also in Ayres 1842:259 [ref. 15930]. •Synonym of Myoxocephalus scorpius (Linnaeus 1758). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Cottidae. Habitat: brackish, marine.

ventralis, Cottus Cuvier [G.] in Cuvier & Valenciennes 1829:194, Pl. 79 (fig. 1) [Histoire naturelle des poissons v. 4; ref. 998] Kamchatka, Russia. Holotype (unique): BMNH 2011.5.17.3. Type catalog: Blanc & Hureau 1968:60 [ref. 20738]. •Synonym of Gymnocanthus pistilliger (Pallas 1814) -- (Watanabe 1960:56 [ref. 11959], Lindberg & Krasyukova 1987:199 [ref. 15964], Sheiko & Fedorov 2000:27 [ref. 25650] in Gymnacanthus, Dyldin & Orlov 2017:63 [ref. 35138]). Current status: Synonym of Gymnocanthus pistilliger (Pallas 1814). Cottidae. Habitat: marine.

ventricosus, Blepsias Eschscholtz [J. F. von] 1829:4, Pl. 13 [Zoologischer Atlas, enthaltend Abbildungen und Beschreibungen neuer Thierarten, ...; ref. 17461] Norfolk Sound [= Sitka Sound] and Sitka, Alaska. No types known. •Synonym of Hemilepidotus hemilepidotus (Tilesius 1811) -- (Bolin 1944:17 [ref. 6379], Peden 1979:24 [ref. 24080]). Current status: Synonym of Hemilepidotus hemilepidotus (Tilesius 1811). Cottidae. Habitat: marine.

vermelnensis, Cottus quadricornis Nybelin [O.] 1940:77, Fig. 1 [Fauna och Flora (Uppsala) v. 35 (no. 2); ref. 23062] Lake Värmeln, Sweden. Syntypes: whereabouts unknown. •Synonym of Triglopsis quadricornis (Linnaeus 1758) -- (Kottelat 1997:171 [ref. 22952]). •Synonym of Myoxocephalus quadricornis (Linnaeus 1758) -- (based on generic placement of quadricornis). Current status: Synonym of Myoxocephalus quadricornis (Linnaeus 1758). Cottidae. Habitat: freshwater, brackish, marine.

verrucosus, Cottus Bean [T. H.] 1881:152 [Proceedings of the United States National Museum v. 4 (no. 210); ref. 223] Plover Bay, anchorage in Providence Bay, Chukchi Peninsula, northeastern Siberia, Russia, Bering Sea. Holotype (unique): USNM 27547. Non-types: MNHN 1896-0467 (1) station 3232. Type catalog: Blanc & Hureau 1968:52 [ref. 20738]. See nomenclatural comments in Mecklenburg et al. 2007:175-176 [ref. 29448]. •Valid as Myoxocephalus verrucosus (Bean 1881) -- (Sheiko & Fedorov 2000:28 [ref. 25650], Parin et al. 2002:S83 [ref. 28300], Fedorov et al. 2003:78 [ref. 27579], Nelson et al. 2004:121 [ref. 27807], Parin et al. 2014:266 [ref. 33547]). •Synonym of Myoxocephalus scorpius (Linnaeus 1758) -- (Mecklenburg et al. 2002:405 [ref. 25968], Mecklenburg et al. 2007:175 [ref. 29448], Mecklenburg et al. 2011:123 [ref. 31212]). Current status: Synonym of Myoxocephalus scorpius (Linnaeus 1758). Cottidae. Distribution: Western North Pacific (if valid). Habitat: marine.

vicinalis, Icelus Gilbert [C. H.] 1896:413 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] Bristol Bay, Alaska, U.S.A., 53°33'50"N, 167°46'50"W, Albatross station 3324, depth 200 meters. Lectotype: USNM 48737. Paralectotypes: BMNH 1900.7.11.8 (1); CAS 45714 [ex IU 6887] (2); MCZ 28344 (2); MNHN 1896-0487 and 0488 (2) station 3331; CAS-SU 3018 (12), 3484 (1); USNM 48597 (1), 53038 (2), 103605 (2), 233758-59 (1, 2); ZMUO J1464 (2, dry) station 3331. Type catalog: Böhlke 1953:130 [ref. 12291], Blanc & Hureau 1968:54 [ref. 20738], Pethon 1969:8 [ref. 19268]. Lectotype established by Jordan & Evermann 1898:1917 [ref. 2444], but if USNM 48737 contained more than one specimen at that time then lectotype selected by Nelson 1984:30 [ref. 5391]. •Synonym of Icelus euryops Bean 1890 -- (Nelson 1984:30 [ref. 5391], Sheiko & Fedorov 2000:28 [ref. 25650], Mecklenburg et al. 2002:453 [ref. 25968], Parin et al. 2002:S77 [ref. 28300], Parin et al. 2014:254 [ref. 33547]). Current status: Synonym of Icelus euryops Bean 1890. Cottidae. Habitat: marine.

vietleri, Cottus Sauvage [H.-E.] (ex Dybowski) 1878:143 [Nouvelles Archives du Muséum d'Histoire Naturelle, Paris (Série 2) v. 1; ref. 3880] Lake Baikal, Russia. Cottidae.

vinculus, Radulinus (Radulinellus) Bolin [R. L.] 1950:197, Fig. 1 [Copeia 1950 (no. 3); ref. 510] Bechers Bay, Santa Rosa Island, California, U.S.A., depth 15 fathoms. Holotype (unique): CAS-SU 15491. Type catalog: Böhlke 1953:132 [ref. 12291]. •Valid as Radulinus vinculus Bolin 1950 -- (Eschmeyer & Herald 1983:182 [ref. 9277], Yabe & Maruyama 2001:59 [ref. 25245], Nelson et al. 2004:121 [ref. 27807], Page et al. 2013:124 [ref. 32708]). Current status: Valid as Radulinus vinculus Bolin 1950. Cottidae. Distribution: Eastern North Pacific: central to southern California (U.S.A.). Habitat: marine.

virginianus, Cottus Cuvier [G.] (ex Willoughby) 1829:163 [Le Règne Animal (Edition 2) v. 2; ref. 995] Available from footnote as, "(1) C. virginianus, Will. x, 15, ou octodecim spinosus, Mitchill. Trans. New-York, iv, p. 380." •Synonym of Myoxocephalus octodecemspinosus (Mitchill 1814). Current status: Synonym of Myoxocephalus octodecemspinosus (Mitchill 1814). Cottidae. Habitat: brackish, marine.

viscosus, Cottus Haldeman [S. S.] 1840:3 [Suppl. Monogr. Limnea. No. 3; ref. 21965] Susquehanna River, eastern Pennsylvania, U.S.A. No types known. Possible syntypes: MCZ 13614 (1). Original not seen; see Jordan & Evermann 1898:1968 [ref. 2444]. •Synonym of Cottus cognatus Richardson 1837 -- (Smith 1986:358 [ref. 22186]). Current status: Synonym of Cottus cognatus Richardson 1837. Cottidae. Habitat: freshwater, brackish.

volki, Cottus minutus Taranetz [A. Ya.] 1933:84 [2] [Doklady Akademii Nauk SSSR, Ser. A (Comptes Rendus de l'Académie des Sciences de l'URSS), Leningrad v. 2; ref. 16528] Suchan River to Takema River, western coast of the Sea of Japan, Primorye, Russia. Syntypes and/or Taranetz specimens: ZIN 25410 (1) Phusun R. On p. 1 of separate. •Synonym of Cottus poecilopus Heckel 1837, but a valid subspecies as described -- (Berg 1949:1144 [ref. 20662], Reshetnikov et al. 1997:758 [ref. 24702], Sideleva 1998:152 [ref. 23595], Parin et al. 2002:S73 [ref. 28300], Fujii et al. 2005:12 [ref. 28191]). •Synonym of Cottus poecilopus Heckel 1837 -- (Bogutskaya et al. 2001:46 [ref. 26178]). •Valid as Cottus volki Taranetz 1933 -- (Shedko & Shedko 2003:330 [ref. 27966], Bogutskaya & Naseka 2004:188 [ref. 28183], Shedko & Miroshnichenko 2007:24 [ref. 29369], Sideleva & Goto 2009:610 [ref. 30591], Goto et al. 2014:[9] [ref. 33456], Parin et al. 2014:241, 245 [ref. 33547]). Current status: Valid as Cottus volki Taranetz 1933. Cottidae. Distribution: Sea of Japan. Habitat: freshwater, brackish.

werestschagini, Abyssocottus Taliev [D. N.] 1935:59, Pl. 1 (fig. 3) [Trudy Baikal'skoi limnologischeskoi stantsii = Travaux de la Station limnologique du Lac Baikal v. 6; ref. 15741] Off Listvenichnoe, southern Lake Baikal, Russia, depth 877 meters. Syntypes: (5) not at ZIN. Neotype designation [without a number] if first in Sideleva 2001:69 [ref. 26181] is invalid, not in compliance with the Code. Numbered ZIN 46662 in Sideleva 2003:200 [ref. 27815] but still invalid. •Valid as Abyssocottus werestschagini Taliev 1935 -- (Berg 1949:1158 [ref. 20662]). •Valid as Cottinella werestschagini (Taliev 1935) -- (Taliev 1955:344 [ref. 4316]). •Valid as Neocottus werestschagini (Taliev 1935) -- (Sideleva 1982:31 [ref. 14469], Reshetnikov et al. 1997:760 [ref. 24702], Sideleva 1998:158 [ref. 23595], Bogutskaya et al. 2001:47 [ref. 26178], Sideleva 2002:275 [ref. 26019], Sideleva 2003:18, 200 [ref. 27815], Bogutskaya & Naseka 2004:199 [ref. 28183], Goto et al. 2014:[15] [ref. 33456]). Current status: Valid as Neocottus werestschagini (Taliev 1935). Cottidae. Distribution: Lake Baikal, Russia. Habitat: freshwater.

wheeleri, Uranidea Cope [E. D.] 1874:138 [10] [Proceedings of the American Philosophical Society v. 14; ref. 932] Beaver River, southwestern Utah, U.S.A. Syntypes: ANSP 22951 (1); MCZ 13636 (2); USNM 12905 (5), 16973 (5). Type catalog: Böhlke 1984:66 [ref. 13621]. On p. 10 of separate. •Synonym of Cottus bairdii Girard 1850, subspecies semiscaber (Cope 1872) -- (La Rivers 1994:586 [ref. 23389]). Current status: Synonym of Cottus bairdii Girard 1850. Cottidae. Habitat: freshwater.

wilsonii, Cottus Girard [C. F.] 1851:42, Pl. 1 (figs. 3-4) [Smithsonian Contributions to Knowledge v. 3 (art. 3) (for 1850); ref. 18124] Allegheny River, Foxburg, Pennsylvania, U.S.A. Syntypes: ANSP, USNM. •Synonym of Cottus bairdii Girard 1850. Current status: Synonym of Cottus bairdii Girard 1850. Cottidae. Habitat: freshwater.

wosnessenskii, Oligocottus Schmidt [P. Yu.] 1903:518 [16] [Izvestiya Imperatorskago Russkago Geograficheskago Obshchestva v. 38 (pt 5) (for 1902); ref. 3945] On p. 16 of separate. Not available; appeared without distinguishing features as above and also in synonymy in Schmidt 1904:93 [ref. 3946]. •In the synonymy of Oligocottus maculosus Girard 1856 -- (Bolin 1944:65 [ref. 6379], Parin et al. 2002:S83 [ref. 28300]). Current status: Synonym of Oligocottus maculosus Girard 1856. Cottidae.

xenogrammus, Stlegicottus Bolin [R. L.] 1936:36, Fig. 8 [Proceedings of the United States National Museum v. 84 (no. 3000); ref. 505] 150 miles north of Rat Island, Bering Sea, Albatross station 3785, depth 270 fathoms. Holotype (unique): USNM 102108. •Valid as Stlegicottus xenogrammus Bolin 1936 -- (Mecklenburg et al. 2002:458 [ref. 25968]). •Synonym of Rastrinus scutiger (Bean 1890) -- (Stevenson 2006:792 [ref. 28923], Parin et al. 2014:271 [ref. 33547]). Current status: Synonym of Rastrinus scutiger (Bean 1890). Cottidae. Habitat: marine.

xenostethus, Triglops Gilbert [C. H.] 1896:429, Pl. 29 [United States Commission of Fish and Fisheries, Report of the Commissioner v. 19 (for 1893) (art. 6); ref. 1628] North of Unalaska Island, 54°15'N, 165°06'W, Alaska, U.S.A., Albatross station 3220, depth 34 fathoms. Holotype (unique): whereabouts unknown. •Valid as Triglops xenostethus Gilbert 1896 -- (Pietsch 1994:344 [ref. 21189], Sheiko & Fedorov 2000:29 [ref. 25650], Mecklenburg et al. 2002:421 [ref. 25968], Parin et al. 2002:S89 [ref. 28300], Nelson et al. 2004:122 [ref. 27807], Page et al. 2013:125 [ref. 32708], Parin et al. 2014:278 [ref. 33547]). Current status: Valid as Triglops xenostethus Gilbert 1896. Cottidae. Distribution: North Pacific: Aleutian Islands, Commander Islands and Kuril Islands. Habitat: marine.

yesoensis, Myoxocephalus Snyder [J. O.] 1911:538 [Proceedings of the United States National Museum v. 40 (no. 1836); ref. 4152] Hakodate market, Hokkaido, Japan. Holotype (unique): USNM 68238. •Synonym of Myoxocephalus stelleri Tilesius 1811 -- (Berg 1949:1137 [ref. 20662]). •Valid as Myoxocephalus yesoensis Snyder 1911 -- (Watanabe 1960:70 [ref. 11959]). •Synonym of Myxocephalus brandtii (Steindachner 1867) -- (Lindberg & Krasyukova 1987:210 [ref. 15964] as yessoensis and brandti, Parin et al. 2014:261 [ref. 33547], Dyldin & Orlov 2017:65 [ref. 35138]). Current status: Synonym of Myoxocephalus brandtii (Steindachner 1867). Cottidae. Habitat: marine.

zanclus, Bero Snyder [J. O.] 1911:540 [Proceedings of the United States National Museum v. 40 (no. 1836); ref. 4152] Otaru market, Hokkaido, Japan. Holotype (unique): USNM 68239. •Valid as Bero zanclus Snyder 1911 -- (Yabe in Masuda et al. 1984:329 [ref. 6441]). •Synonym of Alcichthys elongatus (Steindachner 1881) -- (Nakabo 2002:1526 [ref. 26001]). Current status: Synonym of Alcichthys elongatus (Steindachner 1881). Cottidae. Habitat: marine.

zanderi, Argyrocottus Herzenstein [S. M.] 1892:219 [49] [Mélanges Biologiques, tirés du Bulletin physico-mathématique de l'Académie Impériale des Sciences de St. Pétersbourg v. 13 (pt 2); ref. 5037] Korsakov, Sakhalin Island, Sea of Okhotsk, Russia. Holotype (unique): ZIN 9679 (lost). On p. 49 of Bull. Acad. Imp. Sci. St. Petersbourg (n. s.) v. 3. •Valid as Argyrocottus zanderi Herzenstein 1892 -- (Watanabe 1960:128 [ref. 11959], Neyelov 1979:141 [ref. 3152], Yabe in Masuda et al. 1984:327 [ref. 6441], Lindberg & Krasyukova 1987:225 [ref. 15964], Sokolovskaya et al. 1998:10 [ref. 24670], Nakabo 2000:644 [ref. 25086], Nakabo 2002:644 [ref. 26001], Parin et al. 2002:S68 [ref. 28300], Fedorov et al. 2003:64 [ref. 27579], Parin et al. 2014:234 [ref. 33547]). Current status: Valid as Argyrocottus zanderi Herzenstein 1892. Cottidae. Distribution: Western North Pacific. Habitat: marine.

zapus, Hemilepidotus Gilbert [C. H.] & Burke [C. V.] 1912:54, Figs. 10, 10a [Bulletin of the Bureau of Fisheries v. 30 (Doc. 754) (for 1910); ref. 1634] Near Attu Island, Bering Sea, Albatross station 4782, depth 57-59 fathoms. Holotype: USNM 74374. Paratypes: CAS-SU 22493 (3); USNM 70866-67 (9, 1), 70881 (1). Type catalog: Böhlke 1953:129 [ref. 12291]. •Valid as Hemilepidotus zapus Gilbert & Burke 1912 -- (Peden 1979:28 [ref. 24080], Matarese et al. 1989:380 [ref. 26885], Mecklenburg et al. 2002:434 [ref. 25968], Parin et al. 2002:S76 [ref. 28300], Tokranov et al. 2003:299 [ref. 26972], Nelson et al. 2004:120 [ref. 27807], Mecklenburg et al. 2011:122 [ref. 31212], Page et al. 2013:123 [ref. 32708], Parin et al. 2014:252 [ref. 33547]). Current status: Valid as Hemilepidotus zapus Gilbert & Burke 1912. Cottidae. Distribution: North Pacific: Aleutian Islands to southwestern Gulf of Alaska and to northern Kuril Islands. Habitat: marine.

zonope, Jordania Starks [E. C.] 1895:410 [Proceedings of the Academy of Natural Sciences of Philadelphia v. 47; ref. 4194] Point Orchard, Puget Sound, Seattle, Washington, U.S.A. Syntypes: (3) C AS-SU 3124 (1) Type catalog: Böhlke 1953:131 [ref. 12291]. •Valid as Jordania zonope Starks 1895 -- (Bolin 1944:9 [ref. 6379], Eschmeyer & Herald 1983:174 [ref. 9277], McAllister 1990:199 [ref. 14674], Mecklenburg et al. 2002:420 [ref. 25968], Nelson et al. 2004:120 [ref. 27807], Page et al. 2013:123 [ref. 32708], Pietsch & Orr 2015:41 [ref. 34694], Kells et al. 2016:180 [ref. 35888]). Current status: Valid as Jordania zonope Starks 1895. Cottidae. Distribution: Eastern North Pacific: northern Gulf of Alaska to central California. Habitat: marine.

zonostigma, Pseudoblennius Jordan [D. S.] & Starks [E. C.] 1904:312, Fig. 35 [Proceedings of the United States National Museum v. 27 (no. 1358); ref. 2528] Misaki, Japan. Holotype: CAS-SU 7718. Paratypes: USNM 50927 (1). Additional material: CAS-SU 7565 (1). Type catalog: Böhlke 1953:131-132 [ref. 12291]. •Valid as Pseudoblennius zonostigma Jordan & Starks 1904 -- (Watanabe 1960:155 [ref. 11959], Nakabo in Masuda et al. 1984:330 [ref. 6441], Kimura et al. 1988:19 [ref. 6692], Kim & Youn 1992:74 [ref. 22287], Nakabo 2000:649 [ref. 25086], Nakabo 2002:649 [ref. 26001], Youn 2002:263, 577 [ref. 26218]). Current status: Valid as Pseudoblennius zonostigma Jordan & Starks 1904. Cottidae. Distribution: Western North Pacific. Habitat: marine.

zopherus, Potamocottus Jordan [D. S.] 1877:320 [Annals of the Lyceum of Natural History of New York v. 11 (nos 11-12) (art. 29); ref. 2373] Small tributaries of the Etowah and Oostanaula rivers near Rome, Georgia, U.S.A. Syntypes: BMNH 1880.1.21.18 (1); MCZ 24377 (1); MNHN A-1295 (1); USNM 17877 (7), 329282 [ex Butler Univ. 379] (6). Type catalog: Blanc & Hureau 1968:54 [ref. 20738]. •Synonym of Cottus carolinae (Gill 1861) -- (Lee et al. 1980:807 [ref. 22416], Robison & Buchanan 1988:476 [ref. 23393]). •Synonym of Cottus carolinae (Gill 1861), but a valid subspecies as described -- (Boschung & Mayden 2004:424 [ref. 27995], Scharpf 2008:8 [ref. 30399], Page & Burr 2011:470 [ref. 31215]). Current status: Synonym of Cottus carolinae (Gill 1861). Cottidae. Distribution: Southeastern U.S.A.: Alabama and Georgia [subspecies zopherus]. Habitat: freshwater.