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Eschmeyer's Catalog of Fishes
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Species in the genus Capoeta: [ 64 ] records


aculeatum, Chondrostoma Valenciennes [A.] in Cuvier & Valenciennes 1844:408 [Histoire naturelle des poissons v. 17; ref. 1010] Iran. Lectotype: MNHN 0000-2357. Paralectotypes: MNHN 1960-0611 [ex MNHN 0000-2357] (5). Type catalog: Bertin & Estève 1948:44 [ref. 19574]. Lectotype selected by Coad & Krupp 1994:63 [ref. 23605]. •Synonym of Capoeta capoeta (Güldenstädt 1773), but a valid subspecies aculeata (Valenciennes 1844) -- (Bianco & Bănărescu 1982:85 [ref. 8508]). •Valid as Capoeta aculeata (Valenciennes 1844) -- (Coad 1991:15 [ref. 15702], Coad & Krupp 1994:63 [ref. 23605], Coad 1995:13 [ref. 23608], Coad 1996:71 [ref. 25297], Turan et al. 2006:154 [ref. 28700], Özuluğ & Freyhof 2008:293 [ref. 29981], Coad 2010:147 [ref. 36620], Esmaeili et al. 2010:368 [ref. 33391], Levin et al. 2012:544 [ref. 33912], Jouladeh-Roudbar et al. 2015:867 [ref. 34062], Teimori et al. 2015:90 [ref. 34095], Zareian et al. 2016:140 [ref. 34227], Esmaeili et al. 2016:42 [ref. 34283], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Esmaeili et al. 2017:38 [ref. 35254], Jouladeh-Roudbar et al. 2017:152 [ref. 35425], Jouladeh-Roudbar et al. 2017:152 [ref. 35538], Zareian et al. 2017:17 [ref. 35636], Esmaeili et al. 2018:18 [ref. 36089], Levin et al. 2019:39 [ref. 36600]). Current status: Valid as Capoeta aculeata (Valenciennes 1844). Cyprinidae: Barbinae. Distribution: Namak Lake and Kavir basins, Iran. Habitat: freshwater.

alborzensis, Capoeta Jouladeh-Roudbar [A.], Eagderi [S.], Ghanavi [H. R.] & Doadrio [I.] 2016:169, Figs. 2-5 [FishTaxa v. 1 (no. 3); ref. 34989] Nam River, tributary of Hableh River, near Harandeh village, Tehran province, Iran, 35°42'41.1"N, 52°40'19.7"E. Holotype: IMNRF-UT 1063-115. Paratypes: IMNRF-UT. •Valid as Capoeta alborzensis Jouladeh-Roudbar, Eagderi, Ghanavi & Doadrio 2016 -- (Esmaeili et al. 2017:38 [ref. 35254], Jouladeh-Roudbar et al. 2017:152 [ref. 35425], Jouladeh-Roudbar et al. 2017:152 [ref. 35538], Levin et al. 2019:40 [ref. 36600]). Current status: Valid as Capoeta alborzensis Jouladeh-Roudbar, Eagderi, Ghanavi & Doadrio 2016. Cyprinidae: Barbinae. Distribution: Kavir basin, Tehran province, Iran. Habitat: freshwater.

amir, Scaphiodon Heckel [J. J.] 1847:258 [Reisen in Europa, Asien und Africa v. 2 (pt 3); ref. 2068] Araxes River, Iran. Syntypes: NMW 16508 (1, dry), 46081 (6); RMNH 2682 [ex NMW] (1). •Synonym of Capoeta damascina (Valenciennes 1842) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391]). •Synonym of Capoeta saadii (Heckel 1847) -- (Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Esmaeili et al. 2015:120 [ref. 35030], Esmaeili et al. 2017:40 [ref. 35254], Zareian & Esmaeili 2017:253 [ref. 36090]). Current status: Synonym of Capoeta saadii (Heckel 1847). Cyprinidae: Barbinae. Habitat: freshwater.

anamisensis, Capoeta Zareian [H.], Esmaeili [H. R.] & Freyhof [J.] 2016:133, Figs. 3-5, 6a, 7a and 8 [Zootaxa 4083 (no. 1); ref. 34227] Moradabad River at Ziarat Ali, Minab River drainage, Hormuzgan Province, Iran, 27°45'47.6"N, 57°14'31.8"E. Holotype: ZM-CBSU Z131. Paratypes: FSJF, ZM-CBSU. •Valid as Capoeta anamisensis Zareian, Esmaeili & Freyhof 2016 -- (Esmaeili et al. 2016:42 [ref. 34283], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Esmaeili et al. 2017:39 [ref. 35254], Esmaeili et al. 2018:18 [ref. 36089]). Current status: Valid as Capoeta anamisensis Zareian, Esmaeili & Freyhof 2016. Cyprinidae: Barbinae. Distribution: Minab and Hasan Langi River drainages, Makran, Iran. Habitat: freshwater.

angorae, Varicorhinus capoeta Hankó [B.] 1925:147, Pl. 3 (fig. 3) [Annales Historico-Naturales Musei Nationalis Hungarici v. 21 (for 29 Dec. 1924); ref. 2035] Bozanti, Asia Minor. Holotype (unique): whereabouts unknown. •Valid as Capoeta angorae (Hankó 1925) -- (Turan et al. 2006:152 [ref. 28700], Fricke et al. 2007:34 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Schöter et al. 2009:235 [ref. 30707], Levin et al. 2012:544 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:72 [ref. 34282]). •Synonym of Capoeta damascina (Valenciennes 1842) -- (Alwan et al. 2016:17 [ref. 35686]). Current status: Synonym of Capoeta damascina (Valenciennes 1842). Cyprinidae: Barbinae. Habitat: freshwater.

antalyensis, Varicorhinus Battalgil [F.] 1944:132, Fig. 4 [Revue de la Faculté des Sciences de l'Université d'Instanbul, Série B: Sciences Naturelles v. 9 (no. 2) (for 1943); ref. 13160] Antalya, southwestern Turkey. •Valid as Capoeta antalyensis (Battalgil 1944) -- (Turan et al. 2006:153 [ref. 28700], Turan et al. 2006:434 [ref. 28752], Fricke et al. 2007:34 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Levin et al. 2012:544 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:72 [ref. 34282], Turan et al. 2017:437 [ref. 35330], Çiçek et al. 2018:3 [ref. 36267], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta antalyensis (Battalgil 1944). Cyprinidae: Barbinae. Distribution: Turkey: Mediterranean tributaries, Antalya Province. Habitat: freshwater.

araxensis, Varicorhinus capoeta Dadikyan [M. G.] 1986:162 [Fishes of Armenia; ref. 16233] Aras River, Armenia. No types known. Spelled Varicorhinus capoëta araxensis in original description. •Synonym of Capoeta sevangi De Filippi 1865 -- (Zareian et al. 2017:37 [ref. 35636]). Current status: Synonym of Capoeta sevangi De Filippi 1865. Cyprinidae: Barbinae. Habitat: freshwater.

asmussii, Scaphiodon Keyserling [E. von] 1861:14 [17], Pl. 6 [Zeitschrift für die Gesammten Naturwissenschaften v. 17 (no. 1); ref. 2601] Near Herat, Afghanistan. No types saved. On p. 17 of separate. •Synonym of Varicorhinus capoeta (Güldenstädt 1773), subspecies heratensis (Keyserling 1861) -- (Berg 1949:681 [ref. 20661]). •Synonym of Capoeta capoeta (Güldenstädt 1773) -- (see Coad 1981:9 [ref. 22348], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391], Esmaeili et al. 2017:39 [ref. 35254], Zareian et al. 2017:22 [ref. 35636]). Current status: Synonym of Capoeta capoeta (Güldenstädt 1773). Cyprinidae: Barbinae. Habitat: freshwater.

aydinensis, Capoeta Turan [D.], Küçük [F.], Kaya [C.], Güçlü [S. S.] & Bektaş [Y.] 2017:437, Figs. 1a-b, 2a-d, 3 [Turkish Journal of Zoology v. 41; ref. 35330] Turkey, Aydin Province, Çine Stream, Büyük Menderes drainage, 37°25'N, 28°08'. Holotype: FFR 01926. Paratypes: FFR, IUSHM. •Valid as Capoeta aydinensis Turan, Küçük, Kaya, Güçlü & Bektaş 2017 -- (Çiçek et al. 2018:3 [ref. 36267], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta aydinensis Turan, Küçük, Kaya, Güçlü & Bektaş 2017. Cyprinidae: Barbinae. Distribution: Turkey: Büyük Menderes basin, western Anatolia. Habitat: freshwater.

baliki, Capoeta Turan [D.], Kottelat [M.], Ekmekçi [F. G.] & Imamoğlu [H. O.] 2006:429, Fig. 7 [Revue Suisse de Zoologie v. 113 (no. 2); ref. 28752] Sakarya River, Kιzιlcahamam Stream, at Kιzιlcahamam, 60 kilometers west of Ankara, 40°29'N, 32°39'E, Ankara, Turkey. Holotype: ESFM-PISI/2004-74. Paratypes: CMK 18541 (10); ESFM-PISI/2004-75 (4); FFR 713 (5), 714 (5), 715 (5), 716 (10). •Valid as Capoeta baliki Turan, Kottelat, Ekmekçi & Imamoğlu 2006 -- (Fricke et al. 2007:34 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Levin et al. 2012:544 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Elp et al. 2018:103 [ref. 35851]). •Synonym of Capoeta tinca (Heckel 1843) -- (Çiçek et al. 2018: 26 [ref. 36267]). Current status: Valid as Capoeta baliki Turan, Kottelat, Ekmekçi & Imamoğlu 2006. Cyprinidae: Barbinae. Distribution: Rivers draining into southwestern Black Sea: Turkey. Habitat: freshwater.

banarescui, Capoeta Turan [D.], Kottelat [M.], Ekmekçi [F. G.] & Imamoğlu [H. O.] 2006:427, Fig. 5 [Revue Suisse de Zoologie v. 113 (no. 2); ref. 28752] Çoruh drainage, stream Tortum, 100 kilometers north of Erzurum 40°34'N, 41°36'E, Tortum District, Artvin, Turkey. Holotype: ESFM-PISI/2004-072. Paratypes: CMK 18474 (5), 18540 (9), 18549 (1); ESFM-PISI/2004-073 (4); FFR 711 (9), 712 (16), 720 (3). •Valid as Capoeta banarescui Turan, Kottelat, Ekmekçi & Imamoğlu 2006 -- (Fricke et al. 2007:34 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Bektaş et al. 2011:65 [ref. 31470], Levin et al. 2012:544 [ref. 33912] as cf. banarescui, Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:72 [ref. 34282], Elp et al. 2018:103 [ref. 35851]). •Synonym of Capoeta tinca (Heckel 1843) -- (Çiçek et al. 2018: 26 [ref. 36267]). Current status: Valid as Capoeta banarescui Turan, Kottelat, Ekmekçi & Imamoğlu 2006. Cyprinidae: Barbinae. Distribution: Çoruh River drainage, Turkey and Georgia. Habitat: freshwater.

barroisi, Capoeta Lortet [L.] in Barrois 1894:308 [Revue biologique du nord de la France. v. 6 (1893-1894); ref. 23598] Lake Homs, at Schoummarieh Village, Orontes drainage, Hims District, Syria; Antioche (Antakya, Turkey). No types known. Syntypes: (several) FMUL uncat. (several, probably lost), MHNL uncat. (1). •Valid as Capoeta barroisi Lortet 1894 -- (Coad 1991:15 [ref. 15702], Coad 1995:13 [ref. 23608], Coad 1996:71 [ref. 25297], Coad 1998:102 [ref. 23869], Turan et al. 2006:153 [ref. 28700], Fricke et al. 2007:34 [ref. 29533], Turan et al. 2008:266 [ref. 29989] with redefined range, Özuluğ & Freyhof 2008:293 [ref. 29981], Küçük et al. 2009:72 [ref. 30468], Coad 2010:149 [ref. 36620], Esmaeili et al. 2010:368 [ref. 33391], Levin et al. 2012:544 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:139 [ref. 34227], Zareian et al. 2016:71 [ref. 34282], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Turan et al. 2017:437 [ref. 35330], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta barroisi Lortet 1894. Cyprinidae: Barbinae. Distribution: Eastern Mediterranean watersheds, Tigris River and Gulf basins: Turkey, Iraq, Iran. Habitat: freshwater.

belayewi, Barbus Menon [M. A. S.] 1960:144, Fig. 2 [Records of the Indian Museum (Calcutta) v. 54 (pts 3-4) (1956); ref. 4961] Tigris River, Bagdad, Iraq. Holotype: ZSI-F1046/2. Paratypes: ZSI F1047/2 (1). Type catalog: Menon & Yazdani 1968:104 [ref. 20743]. See Das 2003:353 [ref. 27706]. •Synonym of Capoeta damascina (Valenciennes 1842) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608]). Current status: Synonym of Capoeta damascina (Valenciennes 1842). Cyprinidae: Barbinae. Habitat: freshwater.

bergamae, Capoeta capoeta Karaman [M. S.] 1969:29, Pl. 1 (fig. 5), Pl. 4 (fig. 3) [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 66; ref. 7823] Southwestern Anatolia, Turkey. Syntypes: ZMH H4146 and 3897 (7), 3597 (5), 3603 (2). Type catalog: Wilkens 1977:157 [ref. 22013]. •Valid as Capoeta bergamae Karaman 1969 -- (Turan et al. 2006:152 [ref. 28700], Fricke et al. 2007:35 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Levin et al. 2012:544 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:72 [ref. 34282], Turan et al. 2017:437 [ref. 35330], Çiçek et al. 2018:3 [ref. 36267], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta bergamae Karaman 1969. Cyprinidae: Barbinae. Distribution: Turkey: Aegean Sea watersheds. Habitat: freshwater.

bergi, Varicorhinus Derjavin [A. N.] 1929:72, Fig. 1 [Bulletin of the Ichthyological Laboratory of Baku. v. 2 (no. 2); ref. 16078] Karaj River, 30 kilometers from Teheran, Iran. Holotype (unique): whereabouts unknown. •Synonym of Capoeta aculeata (Valenciennes 1844) -- (Coad 1991:15 [ref. 15702], Coad & Krupp 1994:64 [ref. 23605], Coad 1995:13 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391], Jouladeh-Roudbar et al. 2015:867 [ref. 34062], Esmaeili et al. 2017:38 [ref. 35254], Zareian et al. 2017:19 [ref. 35636]). Current status: Synonym of Capoeta aculeata (Valenciennes 1844). Cyprinidae: Barbinae. Habitat: freshwater.

birunii, Capoeta Zareian [H.] & Esmaeili [H. R.] 2017:261, Figs. 27-28 [Iranian Journal of Ichthyology v. 4 (no. 3); ref. 36090] Daran River near Daran, Zayandehrud basin, Esfahan Province, Iran, 32°49'25.8"N, 50°25'47.4"E. Holotype: ZM-CBSU Z650. Paratypes: ZM-CBSU. •Valid as Capoeta birunii Zareian & Esmaeili 2017 -- (Esmaeili et al. 2018:18 [ref. 36089]). Current status: Valid as Capoeta birunii Zareian & Esmaeili 2017. Cyprinidae: Barbinae. Distribution: Zayandeh River basin, Esfahan Province, Iran. Habitat: freshwater.

buhsei, Capoeta Kessler [K. F.] 1877:85 [The Aralo-Caspian Expedition; ref. 2597] Probably Karaj River near Tehran, Iran. Syntypes: ZIN 2330 (2). •Valid as Capoeta buhsei Kessler 1877 -- (Bianco & Bănărescu 1982:88 [ref. 8508], Coad 1995:14 [ref. 23608], Coad 1996:71 [ref. 25297], Coad 1998:102 [ref. 23869], Turan et al. 2006:153 [ref. 28700], Özuluğ & Freyhof 2008:293 [ref. 29981], Esmaeili et al. 2010:368 [ref. 33391], Levin et al. 2012:544 [ref. 33912] one as cf. buhsei, Jouladeh-Roudbar et al. 2015:867 [ref. 34062], Teimori et al. 2015:90 [ref. 34095], Zareian et al. 2016:140 [ref. 34227], Zareian et al. 2016:71 [ref. 34282], Esmaeili et al. 2016:42 [ref. 34283], Alwan et al. 2016:156 [ref. 34334], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Alwan et al. 2016:17 [ref. 35686], Esmaeili et al. 2017:39 [ref. 35254], Jouladeh-Roudbar et al. 2017:152 [ref. 35425], Jouladeh-Roudbar et al. 2017:153 [ref. 35538], Zareian & Esmaeili 2017:243 [ref. 36090], Esmaeili et al. 2018:18 [ref. 36089]). Current status: Valid as Capoeta buhsei Kessler 1877. Cyprinidae: Barbinae. Distribution: Namak Lake and Kavir basin, Iran. Habitat: freshwater.

caelestis, Capoeta Schöter [C.], Özuluğ [M.] & Freyhof [J.] 2009:230, Figs. 1-4 [Ichthyological Exploration of Freshwaters v. 20 (no. 3); ref. 30707] Göksu River at Göksu, below Göksu power station, approximately 80 kilometers northwest of Silifki, 37°02.74'N, 32°44.56'E, Karaman Province, Turkey. Holotype: IUSHM 37930-252. Paratypes: IUSHM 37930-247 (24), FSJF 2304 (22). Type catalog: Özuluğ & Saç 2018:46 [ref. 35782]. •Valid as Capoeta caelestis Schöter, Özuluğ & Freyhof 2009 -- (Levin et al. 2012:545 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:72 [ref. 34282], Alwan et al. 2016:156 [ref. 34334], Alwan et al. 2016:17 [ref. 35686], Turan et al. 2017:437 [ref. 35330], Çiçek et al. 2018:3 [ref. 36267], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta caelestis Schöter, Özuluğ & Freyhof 2009. Cyprinidae: Barbinae. Distribution: Turkey: Göksu River, Antalya and Karaman provinces. Habitat: freshwater.

capoeta, Cyprinus Güldenstädt [J. A. von] 1773:508, Pl. 8 [Novi Commentarii Academiae Scientiarum Imperialis Petropolitanae v. 17 (for 1772); ref. 18173] Tiflis, Caspian Sea. No types known. Species name spelling from title, with u's replacing v's, and p. 521. •Valid as Varicorhinus capoeta (Güldenstädt 1773) -- (Berg 1949:676 [ref. 20661], Gabrielian 1998:229 [ref. 27566]). •Valid as Capoeta capoeta (Güldenstädt 1773) -- (Coad 1981:9 [ref. 22348], Coad 1991:15 [ref. 15702], Talwar & Jhingran 1991:160 [ref. 20764], Coad 1995:14 [ref. 23608], Bănărescu 1999:387 [ref. 24846], Coad & Abdoli 2000:56 [ref. 25296], Coad 1996:71 [ref. 25297], Coad 1998:102 [ref. 23869], Gabrielyan 2001:27 [ref. 25867], Bogutskaya et al. in Bănărescu & Bogutskaya 2003:147 [ref. 27353], Bogutskaya & Naseka 2004:48 [ref. 28183], Turan et al. 2006:152 [ref. 28700], Fricke et al. 2007:34 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Ninua & Japoshvili 2008:165 [ref. 31251], Esmaeili et al. 2010:368 [ref. 33391], Levin et al. 2012:545 [ref. 33912], Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:71 [ref. 34282], Esmaeili et al. 2016:42 [ref. 34283], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Esmaeili et al. 2017:39 [ref. 35254], Jouladeh-Roudbar et al. 2017:152 [ref. 35425], Jouladeh-Roudbar et al. 2017:153 [ref. 35538], Zareian et al. 2017:22 [ref. 35636], Elp et al. 2018:103 [ref. 35851], Levin et al. 2019:39 [ref. 36600]). Current status: Valid as Capoeta capoeta (Güldenstädt 1773). Cyprinidae: Barbinae. Distribution: Eastern Turkey and southern Caspian Sea watersheds of Georgia, Azerbaijan, Armenia and Iran. Habitat: freshwater.

capoetoides, Varicorhinus Pellegrin [J.] 1938:373 [Bulletin de la Société Zoologique de France v. 63; ref. 17213] Questionably the Chad River basin. Holotype (unique): MNHN 1938-0030. Type catalog: Bertin & Estève 1948:47 [ref. 19574]. •Valid as Varicorhinus capoetoides Pellegrin 1938 -- (Lévêque & Daget 1984:337 [ref. 6186], Lévèque et al. 1991:142 [ref. 31367], Getahun et al. 2004:159 [ref. 27830]). •Possibly a synonym of Capoeta trutta (Heckel 1843) -- (Vreven et al. 2016:29 [ref. 34168]). Current status: Uncertain as Capoeta trutta (Heckel 1843). Cyprinidae: Barbinae. Distribution: Africa? Habitat: freshwater.

chebisiensis, Scaphiodon Keyserling [E. Von] 1861:5 [8], Pl. 2 [Zeitschrift für die Gesammten Naturwissenschaften v. 17 (no. 1); ref. 2601] No types saved. On p. 8 of separate. •Synonym of Capoeta damascina (Valenciennes 1842) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391]). •Synonym of Capoeta saadii (Heckel 1847) -- (Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Esmaeili et al. 2015:120 [ref. 35030], Esmaeili et al. 2017:40 [ref. 35254], Zareian & Esmaeili 2017:253 [ref. 36090]). Current status: Synonym of Capoeta saadii (Heckel 1847). Cyprinidae: Barbinae. Habitat: freshwater.

coadi, Capoeta Alwan [N. H.], Zareian [H.] & Esmaeili [H. R.] 2016:158, Figs. 1-4 [ZooKeys No. 572; ref. 34334] Beshar (Bashar) River at Tale Gah village, Karun River drainage, Kohgiluyeh and Boyer Ahmad provinces, Iran, 30°47'27"N, 51°25'13"E. Holotype: ZM-CBSU Z190. Paratypes: ZM-CBSU. Plus non-type specimens. •Valid as Capoeta coadi Alwan, Zareian & Esmaeili 2016 -- (Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Alwan et al. 2016:17 [ref. 35686], Esmaeili et al. 2017:39 [ref. 35254], Jouladeh-Roudbar et al. 2017:153 [ref. 35425], Jouladeh-Roudbar et al. 2017:153 [ref. 35538], Zareian & Esmaeili 2017:245 [ref. 36090], Esmaeili et al. 2018:19 [ref. 36089]). Current status: Valid as Capoeta coadi Alwan, Zareian & Esmaeili 2016. Cyprinidae: Barbinae. Distribution: Karun River drainage, Tigris River basin, Iran. Habitat: freshwater.

damascinus, Gobio Valenciennes [A.] in Cuvier & Valenciennes 1842:314, Pl. 482 [Histoire naturelle des poissons v. 16; ref. 1009] Damascus, Syria. Lectotype: MNHN 0000-4494. Paralectotypes: MNHN 0000-3947 (1, dry), 0000-3948 (1, dry), ?A-789 (1). Type catalog: Bertin & Estève 1948:43 [ref. 19574]. Lectotype selected by Krupp & Schneider 1989:365 [ref. 13651]. •Valid as Capoeta damascina (Valenciennes 1842) -- (Goren 1974:88 [ref. 7502], Krupp & Schneider 1989:365 [ref. 13651], Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608], Coad 1996:4 [ref. 23831], Coad 1996:71 [ref. 25297], Coad 1998:102 [ref. 23869], Turan et al. 2006:154 [ref. 28700], Fricke et al. 2007:35 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Coad 2010:150 [ref. 36620], Esmaeili et al. 2010:368 [ref. 33391], Levin et al. 2012:545 [ref. 33912], Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Esmaeili et al. 2015:120 [ref. 35030], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:71 [ref. 34282], Alwan et al. 2016:156 [ref. 34334], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Alwan et al. 2016:17 [ref. 35686], Esmaeili et al. 2017:40 [ref. 35254], Turan et al. 2017:437 [ref. 35330], Jouladeh-Roudbar et al. 2017:153 [ref. 35538]). Current status: Valid as Capoeta damascina (Valenciennes 1842). Cyprinidae: Barbinae. Distribution: Levant, Mesopotamia and southeastern Anatolia: Turkey, Syria, Iraq, Iran. Habitat: freshwater.

ekmekciae, Capoeta Turan [D.], Kottelat [M.], Kirankaya [S. G.] & Engin [S.] 2006:148, Figs. 1-2, 4a [Ichthyological Exploration of Freshwaters v. 17 (no. 2); ref. 28700] Çoruh River, Borçka, 30 kilometers north of Artvin, 41°22'N, 41°41'E, Artvin Province, Turkey. Holotype: ESFM-PISI/2004-076. Paratypes: CMK 18473 (3), 18554 (7); ESFM-PISI/2004-077 (4); FRR 704 (6). •Valid as Capoeta ekmekciae Turan, Kottelat, Kirankaya & Engin 2006 -- (Fricke et al. 2007:35 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Bektaş et al. 2011:67 [ref. 31470], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:71 [ref. 34282], Zareian et al. 2017:23 [ref. 35636], Çiçek et al. 2018:4 [ref. 36267], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta ekmekciae Turan, Kottelat, Kirankaya & Engin 2006. Cyprinidae: Barbinae. Distribution: Turkey: Çoruh River basin (Black Sea tributary), Artvin Province. Habitat: freshwater.

erhani, Capoeta Turan [D.], Kottelat [M.] & Ekmekçi [F. G.] 2008:264, Figs. 1-2a [Ichthyological Exploration of Freshwaters v. 19 (no. 3); ref. 29989] Ceyhan drainage, Menzelet Reservoir, Geçit stream on road from Kahramanmaraş to Adana, 37°37'N, 36°39'E, Kahramanmaraş Province, Turkey. Holotype: FFR 776. Paratypes: CGE (5); CMK 18473 (3); FFR 768 (14), 769 (12), 770 (4). •Valid as Capoeta erhani Turan, Kottelat & Ekmekçi 2008 -- (Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:139 [ref. 34227], Turan et al. 2017:437 [ref. 35330], Çiçek et al. 2018:4 [ref. 36267], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta erhani Turan, Kottelat & Ekmekçi 2008. Cyprinidae: Barbinae. Distribution: Turkey: Ceyhan River drainage (Mediterranean Sea tributary). Habitat: freshwater.

ferdowsii, Capoeta Jouladeh-Roudbar [A.], Eagderi [S.], Murillo-Ramos [L.], Ghanavi [H. R.] & Doadrio [I.] 2017:138, Figs. 3-6, 8 [FishTaxa v. 2 (no. 3); ref. 35538] Tang-e Shiv River at Bekr sofla village, Zohreh River drainage, Tigris River basin, Fars Province, Iran, 30°25'26"N, 51°21'55"E. Holotype: IMNRF-UT 1111-61. Paratypes: IMNRF-UT. •Valid as Capoeta ferdowsii Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio 2017 -- (, Zareian & Esmaeili 2017:247 [ref. 36090], Esmaeili et al. 2018:19 [ref. 36089]). Current status: Valid as Capoeta ferdowsii Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio 2017. Cyprinidae: Barbinae. Distribution: Zohreh River drainage, Persian Gulf basin, Iran. Habitat: freshwater.

fratercula, Scaphiodon Heckel [J. J.] 1843:1059 [69] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Damascus, Syria. No types at NMW. Illustrated by Heckel 1843: Pl. 5, fig. 2 [ref. 2067]. On p. 69 of separate [ref. 2067]. •Synonym of Capoeta damascina (Valenciennes 1842) -- (Krupp & Schneider 1989:365 [ref. 13651], Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608]). Current status: Synonym of Capoeta damascina (Valenciennes 1842). Cyprinidae: Barbinae. Habitat: freshwater.

fundulus, Cyprinus Güldenstädt [J. A. von] 1787:222 [Reisen durch Rußland und im Kaukasischen Gebürge; ref. 18175] Kura River at Tbilisi. No types known. •Synonym of Varicorhinus capoeta (Güldenstädt 1773) -- (Berg 1949:676 [ref. 20661]). •Synonym of Capoeta capoeta (Güldenstädt 1773) -- (Coad 1995:14 [ref. 23608], Bănărescu 1999:387 [ref. 24846], Zareian et al. 2017:22 [ref. 35636]). Current status: Synonym of Capoeta capoeta (Güldenstädt 1773). Cyprinidae: Barbinae. Habitat: freshwater.

fusca, Capoeta Nikolskii [A. M.] 1897:340 [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 1; ref. 3201] Mondechi and Kuss, Iran, probably Mandehi or Miandehi, 34°53'N, 58°38'E. Syntypes: ZIN 11108-12 (2, ?, 5, 1, 1). •Valid as Capoeta fusca Nikolskii 1897 -- (Coad 1981:9 [ref. 22348], Coad 1995:14 [ref. 23608], Coad 1998:102 [ref. 23869], Coad & Abdoli 2000:56 [ref. 25296], Coad & Keyzer-de Ville 2005:36 [ref. 28505], Turan et al. 2006:153 [ref. 28700], Özuluğ & Freyhof 2008:293 [ref. 29981], Johari et al. 2009:63 [ref. 30505], Esmaeili et al. 2010:368 [ref. 33391], Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Zareian et al. 2016:140 [ref. 34227], Esmaeili et al. 2016:42 [ref. 34283], Alwan et al. 2016:176 [ref. 34334], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Esmaeili et al. 2017:39 [ref. 35254], Jouladeh-Roudbar et al. 2017:153 [ref. 35425], Jouladeh-Roudbar et al. 2017:153 [ref. 35538], Zareian et al. 2017:23 [ref. 35636], Esmaeili et al. 2018:19 [ref. 36089], Levin et al. 2019:40 [ref. 36600]). Current status: Valid as Capoeta fusca Nikolskii 1897. Cyprinidae: Barbinae. Distribution: Hari, Kavir, Bedjestan, Sistan and Lut basins, southern Iran and western Afghanistan. Habitat: freshwater.

gibbosa, Capoeta Nikolskii [A. M.] 1897:344 [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 1; ref. 3201] Bochsani [Bukhsani], se Khorasan, Iran. Syntypes: ZIN 11104 (2). •Synonym of Capoeta capoeta (Güldenstädt 1773) -- (Coad 1981:9 [ref. 22348], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391], Esmaeili et al. 2017:39 [ref. 35254], Zareian et al. 2017:22 [ref. 35636]). Current status: Synonym of Capoeta capoeta (Güldenstädt 1773). Cyprinidae: Barbinae. Habitat: freshwater.

gracilis, Scaphiodon Keyserling [E. Von] 1861:9 [12], Pl. 4 [Zeitschrift für die Gesammten Naturwissenschaften v. 17 (no. 1); ref. 2601] Rivers near Esfahan, central Iran. Syntypes: not saved. On p. 12 of separate. •Synonym of Varicorhinus capoeta (Güldenstädt 1773), but a valid subspecies -- (Berg 1949:680 [ref. 20661]). •Synonym of Capoeta capoeta (Güldenstädt 1773), but a valid subspecies -- (Bianco & Bănărescu 1982:86 [ref. 8508], see Coad 1981:9 [ref. 22348], Coad 1995:14 [ref. 23608], Bănărescu 1999:387, 395 [ref. 24846], Esmaeili et al. 2010:368 [ref. 33391], Esmaeili et al. 2017:39 [ref. 35254]). •Valid as Capoeta gracilis (Keyserling 1861) -- (Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Zareian et al. 2016:71 [ref. 34282], Zareian et al. 2017:26 [ref. 35636], Esmaeili et al. 2018:19 [ref. 36089], Levin et al. 2019:40 [ref. 36600]). Current status: Valid as Capoeta gracilis (Keyserling 1861). Cyprinidae: Barbinae. Distribution: Zayandehrud basin, Iran. Habitat: freshwater.

gueldenstaedti, Capoeta De Filippi [F.] 1865:313 [Note di un viaggio in Persia nel 1862; ref. 13943] Unneeded new name for Cyprinus capoeta Güldenstädt 1773 (see footnote 2, p. 312 in De Filippi 1865). Name originally as Güldenstaedti. Appeared in Tortonese 1940 [ref. 21559] as Capoeta guldenstadtii Filippi. •In the synonymy of Capoeta capoeta (Güldenstädt 1773) -- (Zareian et al. 2017:22 [ref. 35636]). Current status: Synonym of Capoeta capoeta (Güldenstädt 1773). Cyprinidae: Barbinae.

heratensis, Scaphiodon Keyserling [E. von] 1861:11 [15], Pl. 6 [Zeitschrift für die Gesammten Naturwissenschaften v. 17 (no. 1); ref. 2601] Heri-rud at Herat, Afghanistan. No types saved. On p. 15 of separate. •Synonym of Varicorhinus capoeta (Güldenstädt 1773), but a valid subspecies -- (Berg 1949:681 [ref. 20661], Gabrielian 1998:229 [ref. 27566]). •Synonym of Capoeta capoeta (Güldenstädt 1773) -- (Coad 1981:9 [ref. 22348], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391] as a subspecies). •Valid as Capoeta heratensis (Keyserling 1861) -- (Levin et al. 2012:545 [ref. 33912], Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Zareian et al. 2016:71 [ref. 34282], Esmaeili et al. 2016:42 [ref. 34283], Alwan et al. 2016:177 [ref. 34334], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Esmaeili et al. 2017:39 [ref. 35254], Jouladeh-Roudbar et al. 2017:153 [ref. 35425], Jouladeh-Roudbar et al. 2017:153 [ref. 35538], Zareian et al. 2017:29 [ref. 35636], Esmaeili et al. 2018:19 [ref. 36089], Levin et al. 2019:40 [ref. 36600]). Current status: Valid as Capoeta heratensis (Keyserling 1861). Cyprinidae: Barbinae. Distribution: Hari-rud River basin, Turkmenistan, Afghanistan and Iran. Habitat: freshwater.

hohenackeri, Capoeta Kessler [K. F.] 1877:89 [The Aralo-Caspian Expedition; ref. 2597] Probably Kura-Araks river system, Azerbaijan. Holotype (unique): ZIN 2864. •Synonym of Capoeta capoeta (Güldenstädt 1773) -- (Coad 1995:14 [ref. 23608]). Current status: Synonym of Capoeta capoeta (Güldenstädt 1773). Cyprinidae: Barbinae. Habitat: freshwater.

intermedia, Capoeta capoeta Bianco [P. G.] & Bănărescu [P. M.] 1982:86, Figs. 1B, 2B [Cybium 3e série. Bulletin de la Société Française d'Ichtyologie v. 6 (no. 2); ref. 8508] Mand River, near Akbar, southern Iran. Holotype: IZA 7892. Paratypes: CAS 48113 (1), ISBB uncat. (orig. 13), IZA 7893 (62), NMC 82-367 (6), USNM 227935 (1), ?ZMH 6090 (1). Other specimens: IZA 7894 (2). Type catalog: Wilkens & Dohse 1993:412 [ref. 21161]. Objectively invalid; preoccupied by Capoeta intermedia Temminck & Schlegel 1846. •Synonym of Capoeta damascina (Valenciennes 1842) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391]). •Synonym of Capoeta saadii (Heckel 1847) -- (Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Esmaeili et al. 2015:120 [ref. 35030], Esmaeili et al. 2017:40 [ref. 35254], Zareian et al. 2017:39 [ref. 35636]). Current status: Synonym of Capoeta saadii (Heckel 1847). Cyprinidae: Barbinae. Habitat: freshwater.

kaput, Capoeta Levin [B. A.], Prokofiev [A. M.] & Roubenyan [H. R.] 2019:35, Figs. 1a-c, 2, 3a, 3e-f [Inland Water Biology v. 12 (no. 1); ref. 36600] Khoda Afarin Reservoir, Armenia, 39°08'34"N, 46°50'21"E. Holotype: ZMMU P-23837. Paratypes: ZMMU. Plus additional non-type material. •Valid as Capoeta kaput Levin, Prokofiev & Roubenyan 2019. Current status: Valid as Capoeta kaput Levin, Prokofiev & Roubenyan 2019. Cyprinidae: Barbinae. Distribution: Araxes River basin, Armenia. Habitat: freshwater.

kosswigi, Capoeta capoeta Karaman [M. S.] 1969:31, Pl. 1 (fig. 4), Pl. 5 (fig. 1) [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 66; ref. 7823] Lake Van, Turkey. Syntypes: ZMH 4145 and 3565 (9), 3591 (1). Type catalog: Wilkens 1977:157 [ref. 22013]. •Valid as Capoeta kosswigi Karaman 1969 -- (Turan et al. 2006:152 [ref. 28700], Fricke et al. 2007:35 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Levin et al. 2012:545 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:72 [ref. 34282], Alwan et al. 2016:156 [ref. 34334], Çiçek et al. 2018:4 [ref. 36267], Elp et al. 2018:103 [ref. 35851], Şen et al. 2019:66 [ref. 36474]). Current status: Valid as Capoeta kosswigi Karaman 1969. Cyprinidae: Barbinae. Distribution: Turkey: Van Gölü and Van Gölü tributaries endemic, Van and Bitlis provinces. Habitat: freshwater, brackish.

macrolepis, Scaphiodon Heckel [J. J.] 1847:259 [Reisen in Europa, Asien und Africa v. 2 (pt 3); ref. 2068] Confluents of Araxes River at Persepolis, Iran [Sivand River, Fars near Persepolis]. Syntypes: NMW 51653 (2), 55896 (2). •Synonym of Capoeta capoeta (Güldenstädt 1772) -- (Coad 1991:15 [ref. 15702]). •Synonym of Capoeta capoeta (Güldenstädt 1772), but a valid subspecies -- (Bianco & Bănărescu 1982:85 [ref. 8508]). •Synonym of Capoeta aculeata (Valenciennes 1844) -- (Coad & Krupp 1994:64 [ref. 23605], Coad 1995:13 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391], Jouladeh-Roudbar et al. 2015:867 [ref. 34062], Esmaeili et al. 2017:38 [ref. 35254]). •Valid as Capoeta macrolepis (Heckel 1847) -- (Zareian et al. 2017:31 [ref. 35636], Esmaeili et al. 2018:20 [ref. 36089]). Current status: Valid as Capoeta macrolepis (Heckel 1847). Cyprinidae: Barbinae. Distribution: Kor and Tigris River basins, Iran. Habitat: freshwater.

mandica, Capoeta barroisi Bianco [P. G.] & Bănărescu [P. M.] 1982:90, Figs. 1C, 2C [Cybium 3e série. Bulletin de la Société Française d'Ichtyologie v. 6 (no. 2); ref. 8508] Mond River, near Dasht-e-Arzhan, Persian Gulf basin, Iran. Holotype: IZA 7890. Paratypes: ISBB 3123 (5), IZA 7891 (orig. 95), NMC 82-366 (6). •Synonym of Capoeta barroisi Lortet 1894, but a valid subspecies as described -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391]). •Status unclear -- (Turan et al. 2008:268 [ref. 29989]). •Valid as Capoeta mandica Bianco & Bănărescu 1982 -- (Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Teimori et al. 2015:90 [ref. 34095], Zareian et al. 2016:139 [ref. 34227], Esmaeili et al. 2016:42 [ref. 34283], Alwan et al. 2016:176 [ref. 34334], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Esmaeili et al. 2017:40 [ref. 35254], Esmaeili et al. 2018:20 [ref. 36089], Zareian et al. 2018:283 [ref. 36415]). Current status: Valid as Capoeta mandica Bianco & Bănărescu 1982. Cyprinidae: Barbinae. Distribution: Mond River, Persis River drainage, Persian Gulf basin, Iran. Habitat: freshwater.

mauricii, Capoeta Küçük [F.], Turan [D.], Şahin [C.] & Gülle [I.] 2009:73, Figs. 1-2 [Zoology in the Middle East v. 47; ref. 30468] Sanöz Stream, 1 kilometer west of Beyşehir, Lake Beyşehir drainage, Konya Province, Turkey. Holotype: FFR 3800. Paratypes: FFR 3801 (14). Also additional non-type specimens. Species was previously misidentified as Capoeta pestai (non Pietschmann 1933) by authors. •Valid as Capoeta mauricii Küçük, Turan, Şahin & Gülle 2009 -- (Levin et al. 2012:545 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:72 [ref. 34282], Turan et al. 2017:437 [ref. 35330], Çiçek et al. 2018:4 [ref. 36267], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta mauricii Küçük, Turan, Şahin & Gülle 2009. Cyprinidae: Barbinae. Distribution: Turkey: Beyşehir Gölü and Beyşehir Gölü tributaries. Habitat: freshwater.

niger, Scaphiodon Heckel [J. J.] 1847:258 [Reisen in Europa, Asien und Africa v. 2 (pt 3); ref. 2068] Araxes River, Iran. Syntypes: NMW 51654-56 (1, 2, 1). •Synonym of Capoeta damascina (Valenciennes 1842) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391]). •Synonym of Capoeta saadii (Heckel 1847) -- (Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Esmaeili et al. 2017:40 [ref. 35254], Zareian & Esmaeili 2017:253 [ref. 36090]). Current status: Synonym of Capoeta saadii (Heckel 1847). Cyprinidae: Barbinae. Habitat: freshwater.

nikolskii, Varicorhinus Derjavin [A. N.] 1929:74, Fig. 2 [Bulletin of the Ichthyological Laboratory of Baku. v. 2 (no. 2); ref. 16078] Karaj River, 30 kilometers from Teheran, Iran. Syntypes: (11) whereabouts unknown. •Synonym of Capoeta buhsei Kessler 1877 -- (Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391], Jouladeh-Roudbar et al. 2015:867 [ref. 34062], Zareian & Esmaeili 2017:243 [ref. 36090]). Current status: Synonym of Capoeta buhsei Kessler 1877. Cyprinidae: Barbinae. Habitat: freshwater.

nudiventris, Capoeta Nikolskii [A. M.] 1897:342 [Ezhegodnik. Zoologicheskogo Muzeya Akademii Nauk SSSR. v. 1; ref. 3201] Saride, eastern Iran [Sarideh, Iran, 34°22'N, 58°14'E]. Syntypes: ZIN 11105-06 (7, 3). •Synonym of Capoeta fusca Nikolskii 1897 -- (Coad 1981:9 [ref. 22348], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391], Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Esmaeili et al. 2017:39 [ref. 35254], Zareian et al. 2017:23 [ref. 35636]). Current status: Synonym of Capoeta fusca Nikolskii 1897. Cyprinidae: Barbinae. Habitat: freshwater.

oguzelii, Capoeta Elp [M.], Osmanoğlu [M. İ.], Kadak [A. E.] & Turan [D.] 2018:104, Figs. 1-5 [Zoology in the Middle East v. 64 (no. 2); ref. 35851] Ezine Stream at Devrekani, Black Sea drainage, Kastamonu province, Turkey, 41°44'02"N, 33°52'58"E. Holotype: FCME 2017-05a. Paratypes: FCME. •Valid as Capoeta oguzelii Elp, Osmanoğlu, Kadak & Turan 2018 -- (Çiçek et al. 2018:4 [ref. 36267]). Current status: Valid as Capoeta oguzelii Elp, Osmanoğlu, Kadak & Turan 2018. Cyprinidae: Barbinae. Distribution: Turkey: Ezine Stream (southern Black Sea basin). Habitat: freshwater.

peregrinorum, Scaphiodon Heckel [J. J.] 1843:1061 [71] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Aleppo, Syria. Syntypes: NMW 51658 (1), 51660-65 (1, 1, 1, 1, 3, 1); ?RMNH 2681 [ex NMW] (3) Aleppo. On p. 71 of separate [ref. 2067]. •Synonym of Capoeta damascina (Valenciennes 1842) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608]). Current status: Synonym of Capoeta damascina (Valenciennes 1842). Cyprinidae: Barbinae. Habitat: freshwater.

persica, Capoeta barroisi Karaman [M. S.] 1969:35 [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 66; ref. 7823] Mariwan, 120 kilometers west of Sannadaj, Tigris River drainage, Iran. Holotype (unique): ZMH H4119. Type catalog: Wilkens 1977:157 [ref. 22013]. •Status not clear -- (Turan et al. 2008:268 [ref. 29989]). •Synonym of Capoeta barroisi Lortet 1894, but a valid subspecies as described -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608]). •Synonym of Capoeta barroisi Lortet 1894 -- (Esmaeili et al. 2010:368 [ref. 33391]). •Synonym of Capoeta trutta (Heckel 1843) -- (Zareian et al. 2016:139 [ref. 34227]). Current status: Synonym of Capoeta trutta (Heckel 1843). Cyprinidae: Barbinae. Habitat: freshwater.

pestai, Varicorhinus Pietschmann [V.] 1933:21 [1] [Anzeiger der Kaiserlichen Akademie der Wissenschaften, Wien, Mathematisch-Naturwissenschaftliche Classe v. 70; ref. 16453] Egridir Lake [sic, Eğirdir Gölü], Turkey, Asia Minor. Syntypes: (2) MSNM 34 [ex MSNM 4661 and ex NMW] (1) Egirdir Lake; RMNH 24395 (2). Type catalog: Conci & Michelangeli 1974:225 [ref. 21560]. On p. 1 of separate (in 1933), in journal in 1934. •Valid as Capoeta pestai (Pietschmann 1933) -- (Coad 1991:23 [ref. 15702], Bogutskaya 1997:157 [ref. 23277], Turan et al. 2006:154 [ref. 28700], Fricke et al. 2007:35 [ref. 29533], Turan et al. 2008:269 [ref. 29989], Özuluğ & Freyhof 2008:293 [ref. 29981], Küçük et al. 2009:72, 78 [ref. 30468], Schöter et al. 2009:236 [ref. 30707], Çiçek et al. 2015:143 [ref. 35151], Turan et al. 2017:437 [ref. 35330], Çiçek et al. 2018:4 [ref. 36267], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta pestai (Pietschmann 1933). Cyprinidae: Barbinae. Distribution: Turkey: Eğirdir Gölü and Eğirdir Gölü tributaries, Isparta Province. Habitat: freshwater.

platylepida, Capoeta steindachneri var. Kessler [K. F.] 1872:50 [6] [Izvestiia Imperatorskago Obschchestva Liubitelei Estestvozaniia, Antropologii i Etnografii v. 10 (no. 1); ref. 14134] Zeravshan River, Uzbekistan. Syntypes: (5) ZIN 2316 (1, not found in 1996). On p. 6 of separate. Cyprinidae: Barbinae. Habitat: freshwater.

pyragyi, Capoeta Jouladeh-Roudbar [A.], Eagderi [S.], Murillo-Ramos [L.], Ghanavi [H. R.] & Doadrio [I.] 2017:144, Figs. 8-12 [FishTaxa v. 2 (no. 3); ref. 35538] Tire River at Kaghe Village, Sezar River drainage, Tigris River basin, Lorestan Province, Iran, 33°37'06"N, 48°58'13"E. Holotype: IMNRF-UT 1109-141. Paratypes: IMNRF-UT. •Valid as Capoeta pyragyi Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio 2017 -- (, Zareian & Esmaeili 2017:250 [ref. 36090], Esmaeili et al. 2018:20 [ref. 36089]). Current status: Valid as Capoeta pyragyi Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio 2017. Cyprinidae: Barbinae. Distribution: Tireh and Sezar Rivers, Tigris River basin, Persian Gulf basin, Iran. Habitat: freshwater.

razii, Capoeta Jouladeh-Roudbar [A.], Eagderi [S.], Ghanavi [H. R.] & Doadrio [I.] 2017:144, Figs. 4-7 [ZooKeys No. 682; ref. 35425] Kheyroud River, Caspian Sea basin, Chalus city, Mazandaran Province, Iran, 36°36'35"N, 51°33'45"E. Holotype: IMNRF-UT 1072-9. Paratypes: IMNRF-UT. •Valid as Capoeta razii Jouladeh-Roudbar, Eagderi, Ghanavi & Doadrio 2017 -- (Zareian et al. 2017:33 [ref. 35636], Esmaeili et al. 2018:21 [ref. 36089], Levin et al. 2019:40 [ref. 36600]). Current status: Valid as Capoeta razii Jouladeh-Roudbar, Eagderi, Ghanavi & Doadrio 2017. Cyprinidae: Barbinae. Distribution: Rivers and streams of the central and southern Caspian Sea basin and in the north of the Kavir basin, Iran. Habitat: freshwater.

rostratus, Scaphiodon Keyserling [E. Von] 1861:7 [10], Pl. 3 [Zeitschrift für die Gesammten Naturwissenschaften v. 17 (no. 1); ref. 2601] Holotype (unique): not saved. On p. 10 of separate. •Synonym of Capoeta damascina (Valenciennes 1842) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391]). •Synonym of Capoeta saadii (Heckel 1847) -- (Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Esmaeili et al. 2015:120 [ref. 35030], Esmaeili et al. 2017:40 [ref. 35254], Zareian & Esmaeili 2017:253 [ref. 36090]). Current status: Synonym of Capoeta saadii (Heckel 1847). Cyprinidae: Barbinae. Habitat: freshwater.

saadii, Scaphiodon Heckel [J. J.] 1847:260 [Reisen in Europa, Asien und Africa v. 2 (pt 3); ref. 2068] Persepolis, Pulwar River, Kor River basin, ruins northeast of Shiraz, Iran. Syntypes: NMW 51666 (18), RMNH 3166 [ex NMW] (1). Type catalog: Bianco & Banarescu 1982 [ref. 8508]. •Synonym of Capoeta damascina (Valenciennes 1842) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608], Esmaeili et al. 2010:368 [ref. 33391]). •Valid as Capoeta saadii (Heckel 1847) -- (Bianco & Bănărescu 1982:88 [ref. 8508] as saadi, Levin et al. 2012:545 [ref. 33912] as saadi, Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Teimori et al. 2015:90 [ref. 34095], Esmaeili et al. 2015:119 [ref. 35030], Zareian et al. 2016:140 [ref. 34227], Zareian et al. 2016:71 [ref. 34282], Esmaeili et al. 2016:42 [ref. 34283], Alwan et al. 2016:156 [ref. 34334], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Alwan et al. 2016:17 [ref. 35686], Esmaeili et al. 2017:40 [ref. 35254], Zareian & Esmaeili 2017:253 [ref. 36090], Esmaeili et al. 2018:21 [ref. 36089]). Current status: Valid as Capoeta saadii (Heckel 1847). Cyprinidae: Barbinae. Distribution: Kor, Persis, Hormuz, Kerman and Maharlu Lake basins, southern Iran. Habitat: freshwater.

sevangi, Capoeta De Filippi [F.] 1865:312 [Note di un viaggio in Persia nel 1862; ref. 13943] Lake Sevan (= Goktscha), Armenia. Holotype (unique): MZUT 695. Type catalog: Tortonese 1940:140 [ref. 21559]. •Synonym of Capoeta capoeta (Güldenstädt 1773), but a valid subspecies -- (Bănărescu 1999:387, 395 [ref. 24846], Gabrielian 1998:229 [ref. 27566], Ninua & Japoshvili 2008:165 [ref. 31251]). •Synonym of Varicorhinus capoeta (Güldenstädt 1773), but a valid subspecies -- (Berg 1949:677 [ref. 20661], Gabrielyan 2001:27 [ref. 25867], Turan et al. 2006:154 [ref. 28700]). •Valid as Capoeta sevangi De Filippi 1865 -- (Levin et al. 2012:545 [ref. 33912], Zareian et al. 2016:71 [ref. 34282], Zareian et al. 2017:37 [ref. 35636], Esmaeili et al. 2018:21 [ref. 36089]). Current status: Valid as Capoeta sevangi De Filippi 1865. Cyprinidae: Barbinae. Distribution: Aras, Lake Sevan and Lake Urmia basins, Armenia and Iran. Habitat: freshwater.

shajariani, Capoeta Jouladeh-Roudbar [A.], Eagderi [S.], Murillo-Ramos [L.], Ghanavi [H. R.] & Doadrio [I.] 2017:148, Figs. 8, 14-18 [FishTaxa v. 2 (no. 3); ref. 35538] Gamasiyab River near Doab Village, Tigris River drainage, Hamedan Province, Iran, 34°22'13"N, 47°54'26"E. Holotype: IMNRF-UT 1107-21. Paratypes: IMNRF-UT. •Valid as Capoeta shajariani Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio 2017 -- (Zareian & Esmaeili 2017:255 [ref. 36090], Esmaeili et al. 2018:21 [ref. 36089]). Current status: Valid as Capoeta shajariani Jouladeh-Roudbar, Eagderi, Murillo-Ramos, Ghanavi & Doadrio 2017. Cyprinidae: Barbinae. Distribution: Gamasiyab River drainage, Tigris River basin, Iran. Habitat: freshwater.

sieboldii, Scaphiodon Steindachner [F.] 1864:224 [Verhandlungen der K.-K. zoologisch-botanischen Gesellschaft in Wien v. 14; ref. 13938] Amasya [Amasia], Kizil-Irsen River system, Turkey. Holotype (unique): NMW 55903. •Valid as Varicorhinus sieboldii (Steindachner 1864) -- (Berg 1949:685 [ref. 20661], Gabrielian 1998:229 [ref. 27566] as sieboldy). •Valid as Capoeta sieboldii (Steindachner 1864) -- (Bănărescu & Herzig-Straschil 1999:407 [ref. 24859] as sieboldi, Turan et al. 2006:152 [ref. 28700], Fricke et al. 2007:36 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Ninua & Japoshvili 2008:165 [ref. 31251], Schöter et al. 2009:236 [ref. 30707], Levin et al. 2012:545 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:72 [ref. 34282], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta sieboldii (Steindachner 1864). Cyprinidae: Barbinae. Distribution: Black Sea watersheds, Turkey and Transcaucasia. Habitat: freshwater.

socialis, Scaphiodon Heckel [J. J.] 1843:1061 [71] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Damascus, Syria. Syntypes or holotype: NMW 55855 (1). On p. 71 of separate [ref. 2067]. •Synonym of Capoeta damascina (Valenciennes 1842) -- (Goren 1974:88 [ref. 7502], Krupp & Schneider 1989:365 [ref. 13651], Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608]). Current status: Synonym of Capoeta damascina (Valenciennes 1842). Cyprinidae: Barbinae. Habitat: freshwater.

steindachneri, Capoeta (Scaphiodon) Kessler [K. F.] 1872:49 [5], Pl. 6 (figs. 3-5) [Izvestiia Imperatorskago Obschchestva Liubitelei Estestvozaniia, Antropologii i Etnografii v. 10 (no. 1); ref. 14134] Zeravshan River, Uzbekistan. Syntypes: whereabouts unknown, not ZIN 4493-95 (1, 2, 3). On p. 5 of separate. •Synonym of Varicorhinus capoeta (Güldenstädt 1773) -- (Berg 1949:682 [ref. 20661]). •Synonym of Capoeta capoeta (Güldenstädt 1773) -- (Coad 1981:9 [ref. 22348], Coad 1995:14 [ref. 23608]). •Valid as Capoeta steindachneri Kessler 1872 -- (Levin et al. 2012:545 [ref. 33912], Zareian et al. 2017:32 [ref. 35636], Levin et al. 2019:40 [ref. 36600]). Current status: Synonym of Capoeta capoeta (Güldenstädt 1773). Cyprinidae: Barbinae. Distribution: Turkmenistan. Habitat: freshwater.

syriacum, Chondrostoma Valenciennes [A.] in Cuvier & Valenciennes 1844:407, Pl. 514 [Histoire naturelle des poissons v. 17; ref. 1010] Abraham's River at foot of Mount Sinai, Egypt. Holotype (unique): MNHN 0000-1945. Type catalog: Bertin & Estève 1948:43-44 [ref. 19574]. •Synonym of Capoeta damascina (Valenciennes 1842) -- (Krupp & Schneider 1989:365 [ref. 13651], Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608]). Current status: Synonym of Capoeta damascina (Valenciennes 1842). Cyprinidae: Barbinae. Habitat: freshwater.

tinca, Scaphiodon Heckel [J. J.] 1843:1021 [31] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Bursa, Nilüfer River basin, Turkey. Lectotype: NMW 55931:1. Paralectotypes: NMW 55931:2-7 (6). On p. 31 of separate [ref. 2067]. Lectotype selected by Bănărescu & Herzig-Straschil in Bănărescu 1999:413 [ref. 24846]. •Valid as Varicorhinus tinca (Heckel 1843) -- (Berg 1949:684 [ref. 20661], Gabrielian 1998:229 [ref. 27566]). •Valid as Capoeta tinca (Heckel 1843) -- (Coad 1996:84 [ref. 25304], Bănărescu & Harzig-Straschil 1999:413 [ref. 24859], Turan et al. 2006:422 [ref. 28752], Fricke et al. 2007:36 [ref. 29533], Özuluğ & Freyhof 2008:293 [ref. 29981], Ninua & Japoshvili 2008:165 [ref. 31251], Schöter et al. 2009:236 [ref. 30707], Bektaş et al. 2011:65 [ref. 31470], Çiçek et al. 2015:143 [ref. 35151], Turan et al. 2017:437 [ref. 35330], Çiçek et al. 2018:4 [ref. 36267], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta tinca (Heckel 1843). Cyprinidae: Barbinae. Distribution: Turkey: Marmara Sea tributaries, Bursa Province. Habitat: freshwater.

trutta, Scaphiodon Heckel [J. J.] 1843:1056 [66] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Aleppo, Syria; Tigris River, Mosul, Iraq. Syntypes: NMW 55926 (1), 55928 (2), 55935-37 (2, 2, 1), 55939-42 (4, 1, 3, 1); ?RMNH 3164 [ex NMW] (1) Aleppo; SMF 923 [ex NMW in 1843] (1), 2567 [ex NMW in 1844] (1); ZMB 8789 (1, dry). Illustrated by Heckel 1843: Pl. 4 (fig. 3) [ref. 2067]. On p. 66 of separate [ref. 2067]. •Valid as Capoeta trutta (Heckel 1843) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608], Coad 1998:102 [ref. 23869], Coad & Keyzer-de Ville 2005:36 [ref. 28505], Turan et al. 2006:153 [ref. 28700], Turan et al. 2008:266 [ref. 29989] with redefined range, Özuluğ & Freyhof 2008:293 [ref. 29981], Küçük et al. 2009:72 [ref. 30468], Schöter et al. 2009:236 [ref. 30707], Coad 2010:153 [ref. 36620], Esmaeili et al. 2010:368 [ref. 33391], Levin et al. 2012:545 [ref. 33912], Jouladeh-Roudbar et al. 2015:868 [ref. 34062], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:140 [ref. 34227], Zareian et al. 2016:71 [ref. 34282], Esmaeili et al. 2016:42 [ref. 34283], Alwan et al. 2016:176 [ref. 34334], Jouladeh-Roudbar et al. 2016:169 [ref. 34989], Esmaeili et al. 2017:40 [ref. 35254], Jouladeh-Roudbar et al. 2017:153 [ref. 35425], Jouladeh-Roudbar et al. 2017:154 [ref. 35538], Elp et al. 2018:103 [ref. 35851], Esmaeili et al. 2018:22 [ref. 36089]). Current status: Valid as Capoeta trutta (Heckel 1843). Cyprinidae: Barbinae. Distribution: Western Asia: Tigris and Euphrates River basins: Syria, Turkey, Iraq, Iran. Habitat: freshwater.

turani, Capoeta Özuluğ [M.] & Freyhof [J.] 2008:290, Figs. 1-3 [Ichthyological Exploration of Freshwaters v. 19 (no. 4); ref. 29981] Stream Çatkit, south of Salbas, lower part of River Pozanti, 37°05.77'N, 35°07.02'E, Adana Province, Turkey. Holotype: IUSHM 36300-100. Paratypes: FSJF 2356 (8); IUSHM 36300-94 (10), 36300-101 (9), 36300-105 (3). Type catalog: Özuluğ & Saç 2018:46 [ref. 35782]. •Valid as Capoeta turani Özuluğ & Freyhof 2008 -- (Küçük et al. 2009:73 [ref. 30468], Schöter et al. 2009:236 [ref. 30707], Levin et al. 2012:545 [ref. 33912], Çiçek et al. 2015:143 [ref. 35151], Zareian et al. 2016:140 [ref. 34227], Zareian et al. 2016:71 [ref. 34282], Turan et al. 2017:437 [ref. 35330], Elp et al. 2018:103 [ref. 35851]). Current status: Valid as Capoeta turani Özuluğ & Freyhof 2008. Cyprinidae: Barbinae. Distribution: Turkey: Seyhan River basin (Mediterranean tributary). Habitat: freshwater.

turcicus, Varicorhinus kemali Hankó [B.] 1925:150, Pl. 3 (fig. 5) [Annales Historico-Naturales Musei Nationalis Hungarici v. 21 (for 29 Dec. 1924); ref. 2035] Eregli, Turkey. Syntypes: (9) MNHN 1928-220 (1). Type catalog: Bertin & Estève 1948:45 [ref. 19574]. Cyprinidae: Barbinae. Habitat: freshwater.

turicica, Capoeta barroisi natio Karaman [S.] 1969:35 [Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut v. 66; ref. 7823] Not available, infrasubspecific; from Aksu, Ceyhan drainage, Turkey. Named erhan by Turan, Kottelat & Ekmekçi 2008. •In the synonymy of Capoeta erhani Turan, Kottelat & Ekmekçi 2008. Current status: Synonym of Capoeta erhani Turan, Kottelat & Ekmekçi 2008. Cyprinidae: Barbinae.

umbla, Scaphiodon Heckel [J. J.] 1843:1060 [70] [Ichthyologie [von Syrien]. In Russegger v. 1 (pt 2); ref. 2066] Tigris River, Mosul, Iraq. Syntypes: NMW 55932-33 (1, 1), 79373-74 (1, 1, both dry); SMF 6777 [ex NMW in 1844] (1, dry). Illustrated by Heckel 1843: Pl. 5 (fig. 3) [ref. 2067]. On p. 70 of separate [ref. 2067]. •Questionably a synonym of Capoeta damascina (Valenciennes 1842) -- (Coad 1991:15 [ref. 15702], Coad 1995:14 [ref. 23608]). •Valid as Capoeta umbla (Heckel 1843) -- (Turan et al. 2006:152 [ref. 28700], Özuluğ & Freyhof 2008:293 [ref. 29981], Schöter et al. 2009:236 [ref. 30707], Çiçek et al. 2015:143 [ref. 35151], Esmaeili et al. 2016:36 [ref. 34283], Alwan et al. 2016:156 [ref. 34334], Alwan et al. 2016:17 [ref. 35686], Esmaeili et al. 2017:40 [ref. 35254], Jouladeh-Roudbar et al. 2017:153 [ref. 35425], Jouladeh-Roudbar et al. 2017:154 [ref. 35538], Zareian & Esmaeili 2017:258 [ref. 36090], Elp et al. 2018:103 [ref. 35851], Esmaeili et al. 2018:22 [ref. 36089]). Current status: Valid as Capoeta umbla (Heckel 1843). Cyprinidae: Barbinae. Distribution: Euphrates and Tigris river basins, Iran, Iraq and Turkey. Habitat: freshwater.